Untitled - Dutch Birding

Introduction 

I have endeavoured to keep typos, errors, omissions etc in this list to a minimum, however when 

you find more I would be grateful if you could mail the details during 2011 & 2012 to: 

josephobbs@gmail.com. 

Grateful thanks to Ross Ahmed and Tom Shevlin (www.wildlifesnaps.com) for the cover images. All 

images © the photographers. 

Joe Hobbs 

Index 

The general order of species follows the International Ornithological Congress’ World Bird List. 

Version 

Version 1.5 (1 September 2011). 

Cover 

Main image: Paddyfield Warbler. Small Aksuat, Naurzum Nature Reserve, Qostanay Province, 

Kazakhstan. 29 July 2010. Ross Ahmed. 

Vignette: Sedge Warbler. Great Saltee Island, Co. Wexford, Ireland. 7 May 2006. Tom Shevlin. 

Species 

Page No. 

African Reed Warbler [Acrocephalus baeticatus] 22 

Aquatic Warbler [Acrocephalus paludicola] 14 

Australian Reed Warbler [Acrocephalus australis] 10 

Basra Reed Warbler [Acrocephalus griseldis] 4 

Black-browed Reed Warbler [Acrocephalus bistrigiceps] 12 

Blunt-winged Warbler [Acrocephalus concinens] 17 

Blyth's Reed Warbler [Acrocephalus dumetorum] 19 

Bokikokiko [Acrocephalus aequinoctialis] 10 

Cape Verde Warbler [Acrocephalus brevipennis] 5 

Caroline Islands Warbler [Acrocephalus syrinx] 10 

Clamorous Reed Warbler [Acrocephalus stentoreus] 9 

Cook Islands Reed Warbler [Acrocephalus kerearako] 12 

Great Reed Warbler [Acrocephalus arundinaceus] 7 

Greater Swamp Warbler [Acrocephalus rufescens] 5 

Henderson Reed Warbler [Acrocephalus taiti] 12 

Large-billed Reed Warbler [Acrocephalus orinus] 17 

Lesser Swamp Warbler [Acrocephalus gracilirostris] 5 

Madagascar Swamp Warbler [Acrocephalus newtoni] 6 

Manchurian Reed Warbler [Acrocephalus tangorum] 17 

Marsh Warbler [Acrocephalus palustris] 23 

Millerbird [Acrocephalus familiaris] 10 

Moustached Warbler [Acrocephalus melanopogon] 13 

Nauru Reed Warbler [Acrocephalus rehsei] 10 

1

Nightingale Reed Warbler [Acrocephalus luscinius] 10 

Northern Marquesan Reed Warbler [Acrocephalus percernis] 11 

Oriental Reed Warbler [Acrocephalus orientalis] 8 

Paddyfield Warbler [Acrocephalus agricola] 18 

Pitcairn Reed Warbler [Acrocephalus vaughani] 12 

Reed Warbler [Acrocephalus scirpaceus] 20 

Rimatara Reed Warbler [Acrocephalus rimitarae] 12 

Rodrigues Warbler [Acrocephalus rodericanus] 7 

Sedge Warbler [Acrocephalus schoenobaenus] 15 

Seychelles Warbler [Acrocephalus sechellensis] 6 

Southern Marquesan Reed Warbler [Acrocephalus mendanae] 11 

Speckled Reed Warbler [Acrocephalus sorghophilus] 17 

Tahiti Reed Warbler [Acrocephalus caffer] 11 

Tuamotu Reed Warbler [Acrocephalus atyphus] 11 

2

Relevant Publications 

Baker, K. 1997. Warblers of Europe, Asia and North Africa. Christopher Helm, London. 

Cramp, S. & Brooks, D.J. (eds.) 1992. Handbook of the Birds of Europe, the Middle East and North 

Africa, The Birds of the Western Palearctic, Volume 6 - Warblers. Oxford University Press. 

del Hoyo, J. et al (eds.) 2006. Handbook of the Birds of the World Volume 11, Old World Flycatchers 

to Old World Warblers. Lynx Edicions, Barcelona. 

Jenni, L. & Winkler, R. 1994. Moult and Ageing of European Passerines. Academic Press. 

Kennerley, P. & Pearson, D. 2010. Reed and Bush Warblers. Christopher Helm, A & C Black. 

Moore, A. & Wright, B. 1983. A Field Guide to the Warblers of Britain and Europe. Oxford University 

Press. 

Parkin, D.T. & Knox, A.G. 2010. The Status of Birds in Britain & Ireland. Christopher Helm. 

Parmenter, T. & Byers, C. 1991. A Guide to the Warblers of the Western Palearctic. Bruce Coleman 

Books. 

Pratt, H.D. et al 1987. The Birds of Hawaii and the Tropical Pacific. Princeton University Press. 

Sample, G. 2003. Collins Field Guide: Warbler Songs and Calls of Britain and Europe (three CDs). 

Harper Collins. 

Shirihai, H. et al 1996. The Macmillan Birder’s Guide to European and Middle Eastern Birds. 

Macmillan Press, London and Basingstoke. 

Simms, E. 1985. British Warblers. Collins, London. 

Snow, D.W. & Perrins, C.M. (eds.) 1998. The Birds of the Western Palearctic Concise Edition 

Volume 2, Passerines. Oxford University Press. 

Svensson, L. 1992. Identification Guide to European Passerines (4th edition). Privately Published, 

Stockholm. 

van Duivendijk, N. 2010. Advanced Bird ID Guide, The Western Palearctic. New Holland. 

Vinicombe, K. et al. 1989. The Macmillan Field Guide to Bird Identification. Macmillan Press, London 

and Basingstoke. 

Reviews 

Betton, K. 2011. Reviews (Reed and Bush Warblers by Peter Kennerley and David Pearson 2010). 

Birding World 24(1): 44. 

Bradshaw, C. 1993. Reviews (Identification Guide to European Passerines by Lars Svensson). 

Birdwatch 2(2): 50. 

Dowdall, J. 1992. Book reviews (A Guide to the Warblers of the Western Palearctic. By Tim 

Paramenter and Clive Byers, 1991). IWC News 70: 15. 

Fisher, D. 1989. Reviews (The Macmillan Field Guide to Bird Identification. By Alan Harris, Laurel 

Tucker and Keith Vinicombe, 1989). British Birds 82(9): 421. 

Garner, M. 2010. Reviews (Advanced Bird ID Guide by Nils van Duivendijk, 2010). British Birds 

103(11): 680-681. 

Harvey, P. 2010. Reviews (Reed and Bush Warblers by Peter Kennerley and David Pearson, 

illustrated by Brian Small, 2010). British Birds 104(3): 168-170. 

Madge, S. 1996. Book Reviews (The Macmillan Birder’s Guide to European and Middle Eastern Birds 

by Alan Harris, Hadoram Shirihai and David Christie, 1996). Birdwatch 50: 46. 

Madge, S.C. 1984. Reviews (A Field Guide to the Warblers of Britain and Europe. By Alick Moore. 

Illustrated by Bryon Wright, 1983). British Birds 77(12): 643-644. 

Madge, S.C. 1986. Reviews (British Warblers. By Eric Simms, 1985). British Birds 79(5): 270. 

Martins, R. 1988. Reviews (The Birds of Hawaii and the Tropical Pacific. By H. Douglas Pratt, Philip 

L. Bruner and Delwyn G. Berrett, 1987). British Birds 81(2): 81-82. 

Mitchell, D. 1998. Reviews (Warblers of Europe, Asia and North Africa. By Kevin Baker, 1997). 

Birdwatch 68: 42-43. 

Mullarney, K. 1994. Reviews (A good formula - Mould and Ageing of European passerines by Lukas 

Jenni and Raffael Winkler. Academic Press). Birdwatch 29: 46. 

Nikander, P.J. 1996. Arvostelua - Reviews (The Macmillan Birder’s Guide to European and Middle 

Eastern Birds. Harris, A., Shirihai, H. & Christie, D. 1996). Alula 2(4): 183-184. 

Votier, S.C. 2003. Reviews (Collins Field Guide: Warbler Songs and Calls of Britain and Europe 

(three CDs) by Geoff Sample 2003). British Birds 96(7): 352. 

General Notes 

Alström, P. et al 1994. Bestimmung der kleinen fernöstlichen Rohrsänger Acrocephalus 

[Identification of small Eastern Acrocephalus warblers]. Limicola 8(3): 121-131. 

Baker, K. 1996. Ageing passerines: some practical examples. Birding World 9(7): 280-282. 

3

Brown, R.J. et al 1990. Reactions of parasitized and unparasitized populations of Acrocephalus 

warblers to model cuckoo eggs. Ibis 132:109-111. 

Chernetsov, N. 2004. Intraspecific variation of wing pointedness index in juvenile Acrocephalus 

warblers in the southeastern Baltic. Journal of Ornithology 145: 105-108. 

Collinson, M. 2006. Splitting headaches Recent taxonomic changes affecting the British and 

Western Palearctic lists. British Birds 99(6): 306-323 (313-314). 

Fraser, M. 2004. ListCheck (Acrocephalus and Hippolais warblers). Birdwatch 148: 14. 

Fregin, S. et al 2009. Multi-locus phylogeny of the family Acrocephalidae (Aves: Passeriformes)- the 

traditional taxonomy overthrown. Molecular Phylogenetics and Evolution 52: 866-878. 

Gantlett, S. 1998. Bird forms in Britain. Birding World 11(6): 232-239. 

Helbig, A.J. & Seibold, I. 1999. Molecular phylogeny of Palearctic-African Acrocephalus and 

Hippolais warblers (Aves: Sylviidae). Molecular Phylogenetics and Evolution 11: 246-260. 

Holyoak, D.T. 1978. Variable albinism of flight feathers as an adaptation for recognition of individual 

birds in some Polynesian populations of Acrocephalus warblers. Ardea 66: 112-117. 

Kennerley, P.R. & Leader, P.J. 1991. The identification, status and distribution of small 

Acrocephalus warblers in eastern China. Hong Kong Bird Report 1991 pp. 143-187. 

King, J. 1998. OrnithoNews (Redifining Acrocephalus and Hippolais). Birding World 11(2): 42. 

Knox, A.G. et al 2002. Taxonomic recommendations for British birds. Ibis 144: 707-710. 

Leisler, B. & Catchpole, C.K. 1992. The evolution of polygamy in European reed warblers of the 

genus Acrocephalus: a comparative approach. Ethology Ecology & Evolution 4: 225-243. 

Leisler, B. et al 1989. Habitat, behaviour and morphology of Acrocephalus warblers: an integrated 

analysis. Ornis Scandinavica 20: 181-186. 

Leisler, B. et al 1997. Taxonomy and phylogeny of reed warblers (genus Acrocephalus) based on 

mtDNA sequences and morphology. Journal of Ornithology 98: 22-35. 

Leisler, B. et al 2002. Evolution of Breeding Systems in Acrocephaline Warblers. The Auk 119(2): 

379-390. 

Miholcsa, T. et al 2009. Change of timing of autumn migration in Acrocephalus and Locustella genus. 

Acta Zoologica Academiae Scientiarum Hungaricae 55(2): 175-185. 

Mitchell, D. 2011. Birds of Britain: subspecies checklist v1.1. [online PDF]. Available from: 

http://www.birdwatch.co.uk/categories/articleitem.aspcate=22&topic=155&item=800 [Accessed July 

2011]. 

Parkin, D.T. et al 2004. Species limits in Acrocephalus and Hippolais warblers from the Western 

Palearctic. British Birds 97(6): 276-299. 

Sangster, G. 1997. Trends in systematics, Acrocephalus and Hippolais relationships: shaking the 

tree. Dutch Birding 19(6): 294-300. 

Sangster, G. et al 1999. Dutch avifaunal list: species concepts, taxonomic instability, and taxonomic 

changes in 1977-1998. Ardea 87(1): 139-166. 

Sangster, G. et al 2009. Taxonomic recommendations for British birds: Sixth report. Ibis 152: 180- 

186. 

Schulze-Hagen, K. & Barthel, P.H. 1993. Die Bestimmung der europäischen ungestreiften 

Rohrsänger Acrocephalus [The identification of unstreaked European Acrocephalus warblers]. 

Limicola 7(1): 1-34. 

Svensson, E. & Hedenström, A. 1999. A phylogenetic analysis of the evolution of moult 

strategies in Western Palearctic warblers (Aves: Sylviidae). Biological Journal of the Linnean Society 

67: 263-276. 

van den Berg, A.B. 2011. Dutch Birding-vogelnamen [Dutch Birding bird names] [online PDF]. 

Available from: http://www.dutchbirding.nl/page.phppage_id=229 [Accessed March 2011]. 

Basra Reed Warbler 

Acrocephalus griseldis [Hartlaub 1891, Nguru, Kilosa, Tanzania]. 

Lower R Euphrates & R Tigris (E Iraq). Winters S Somalia S to Mozambique & Malawi. 

Ayé, R. 2006. Basra Reed Warblers in Iran in May 2004, with notes on vocalization. Dutch Birding 

28(5): 304-306. 

Fadhel, O. 2007. Days in Iraq with the Basra Reed Warbler Acrocephalus griseldis. Sandgrouse 

29(1): 95-96. 

Fågel, P. 2007. Kuwait - A Birding Destination at the Southeastern Corner of the Western Palearctic. 

Alula 13(4): 166-174 (173, plate 18). 

4

Hansson, B. & Richardson, D.S. 2005. Genetic variation in two endangered Acrocephalus species 

compared to a widespread congener: estimates based on functional and random loci. Animal 

Conservation 8: 83-90. 

Lehto, H. et al 1998. Photonews. Alula 4(2): 70. 

Pearson, D.J. & Backhurst, G.C. 1988. Characters and taxonomic position of Basra Reed Warbler. 

British Birds 81(4): 171-178. 

Perlman, Y. & Geffen, A. 2007. Basra Reed Warbler Acrocephalus griseldis in the Hula Valley, 

Israel, in 2006. Sandgrouse 29(2): 210-214. 

Perlman, Y. & Shanni, I. 2008. Basra Reed Warblers breeding in Israel. Alula 14(1): 46-47. 

Pitches, A. 2007. News and Comment (Warblers withdrawing from Basra). British Birds 100(10): 630. 

Shirihai, H. et al 1995. Identification and taxonomy of large Acrocephalus warblers. Dutch Birding 

17(5): 229-239. 

Tenovuo, J. 2006. Basra Reed Warbler - a little-known Western Palearctic bird. Birding World 19(2): 

66-68. 

van den Berg, A.B. & Symens, P. 1992. Occurrence and identification of Basra Reed Warbler in 

Saudi Arabia. Dutch Birding 14(2): 41-48. 

Cape Verde Warbler 

Acrocephalus brevipennis [Keulemans 1866, São Nicolau, Cape Verde Islands]. 

São Tiago, São Nicolau & Fogo. Probably extinct on Brava (all Cape Verde Islands). 

Other names: Cane Warbler, Cape Verde Cane-warbler, Dohrn’s Warbler. 

Heinrich Wolfgang Ludwig Dohrn (1838-1913), German politician & entomologist. 

Anon. 1998. Cape Verde Warbler rediscovered on São Nicolau. World Birdwatch 20(3): 6. 

Castell, P. 1999. The nest and nestlings of the Cape Verde Cane Warbler Acrocephalus brevipennis. 

Bulletin of the African Bird Club 6(2): 100. 

Bourne, W.R.P. 2009. Letters (The Cape Verde Warbler on Fogo). British Birds 102(7): 403. 

Donald, P.F. et al 2004. Status of the Cape Verde Cane Warbler Acrocephalus brevipennis on São 

Nicolau, with notes on song, breeding behaviour and threats. Malimbus 26(1-2): 34-37. 

Hazevoet, C.J. 1986. Especialidades de Cabo Verde. Dutch Birding 8(4): 134-139 (plate 96). 

Hazevoet, C.J. et al 1999. Rediscovery of the Cape Verde Cane Warbler Acrocephalus brevipennis 

on São Nicolau in February 1998. Bulletin of the British Ornithologists' Club 119: 68-71. 

Hering, J. & Fuchs, E. 2009. The Cape Verde Warbler: distribution, density, habitat and breeding 

biology on the island of Fogo. British Birds 102(1): 17-24. 

Hering, J. & Hering, H. 2005. Discovery of Cape Verde Warbler Acrocephalus brevipennis on Fogo, 

Cape Verde Islands. Bulletin of the African Bird Club 12(2): 147-149. 

Hering, J. 2008. Der Kapverdenrohrsänger auf Fogo [Cape Verde Cane Warbler on Fogo Island]. Der 

Falke 55: 217-221. 

King, J.R. 1999. OrnithoNews (The plight of Cape Verde Cane Warbler). Birding World 12(7): 260. 

Greater Swamp Warbler 

Acrocephalus rufescens [Sharpe and Bouvier 1876, Landana, Cabinda, Angola]. 

[A.r. rufescens] Ghana, S Togo, S Benin, Nigeria, Cameroon, Bioko Island (aka Fernandóo Po), N 

Central African Republic & NW DR Congo. 

[A.r. senegalensis] Senegal & Gambia. 

[A.r. chadensis] Lake Chad and its environs. 

[A.r. ansorgei] NW Angola & S Sudan S to Uganda, Rwanda & adjacent E DR Congo, W Kenya, 

Zambia, NE Namibia, extreme N Botswana & W Zimbabwe. 

Other names: Rufous Swamp Warbler, Rufous Cane-warbler, Long-clawed Papyrus-warbler. 

Aidley, D.J. & Wilkinson, R. 1987. The annual cycle of six Acrocephalus warblers in a Nigerian reedbed. 

Bird Study 34: 226-234. 

Dowsett-Lemaire, F. 1994. The song of the Seychelles Warbler Acrocephalus sechellensis and its 

African relatives. Ibis 136: 489-491. 

Dowsett, R.J. & Moore, A. 1997. Swamp warblers Acrocephalus gracilirostris and Acrocephalus 

rufescens at Lake Chad, Nigeria. Bulletin of the British Ornithologists' Club 117: 48-51. 

Lesser Swamp Warbler 

Acrocephalus gracilirostris [Hartlaub 1864, Liesbeek River, South Africa]. 

[A.g. gracilirostris] SE Zimbabwe & S Mozambique S to S Namibia & South Africa. 

5

[A.g. cunenensis] SW Angola & N Namibia E to N Botswana, SW Zambia & W Zimbabwe. 

[A.g. winterbottomi] E Angola E to N & NW Zambia and SW Tanzania. 

[A.g. leptorhynchus] Tanzania & S Zaire and Angola to Mozambique. 

[A.g. parvus] SW Ethiopia S to Kenya, N Tanzania, Rwanda & Burundi. 

[A.g. jacksoni] W Kenya, Uganda & S Sudan. 

[A.g. tsanae] NW Ethiopia. 

[A.g. neglectus] W Chad. 

Other names: Swamp Warbler, Lesser Swamp Reed-warbler, African Swamp-warbler, Cape Reed 

Warbler. 


Bird Study 34: 226-234. 



Dowsett, R.J. & Moore, A. 1997. Swamp warblers Acrocephalus gracilirostris and Acrocephalus 

rufescens at Lake Chad, Nigeria. Bulletin of the British Ornithologists' Club 117: 48-51. 

Raijmakers, J.M.H. & Raijmakers, J.H.F.A. 2002. Biometrics and moult of the Cape Reed Warbler 

Acrocephalus gracilirostris in southern Gauteng and northern Free State, South Africa. Afring News 

31(1-2): 32-36. 

Madagascar Swamp Warbler 

Acrocephalus newtoni [Hartlaub 1863, near Soamandrikazay, Madagascar]. 

Madagascar. 

Other name: Madagascar Warbler. 

Goodman, S.M. et al 2000. Patterns of morphological and molecular variation in Acrocephalus 

newtoni on Madagascar. Ostrich 71(3-4): 367-370. 

Woog, F. 2008. Madagascar Swamp Warbler Acrocephalus newtoni far from a swamp. Bulletin of the 

African Bird Club 15(1): 85-87. 

Seychelles Warbler 

Acrocephalus sechellensis [Oustalet 1877, Marianne Island, Seychelles]. 

Cousin Island, Aride (where translocated in 1988), Cousine Island (where translocated in 1990) & 

Denis Island (where translocated in 2004) (all Seychelles, W Indian Ocean). 

Other name: Seychelles Brush-Warbler, Seychelles Swamp-warbler. 

Anon. 1988. A new home for the Seychelles Brush Warbler. World Birdwatch 10(3-4): 4. 

Bristol, R. 2005. Conservation introductions of Seychelles Fody and Warbler to Denis Island, 

Seychelles. Re-introduction News 24: 35-36. 

Brouwer, L. et al 2007. Heterozygosity-fitness correlations in a bottlenecked island species: a case 

study on the Seychelles warbler. Molecular Ecology 16(15): 3134-3144. 



Eikenaar, C. et al 2008. Sex biased natal dispersal in a closed, saturated population of Seychelles 

warblers Acrocephalus sechellensis. Journal of Avian Biology 39: 73-80. 

Fraser, M. 2004. Avian revelations (The battle to preserve island wildlife). Birdwatch 143: 10-12. 




Komdeur, J. & Daan, S. 2005. Breeding in the monsoon: semiannual reproduction in the Seychelles 

warbler (Acrocephalus sechellensis). Journal of Ornithology 146: 305-313. 

Komdeur, J. & Edelaar, P. 2001. Evidence that helping at the nest does not result in territory 

inheritance in the Seychelles warbler. Proceedings of the Royal Society London, Biological Sciences 

268(1480): 2007-2012. 

Komdeur, J. & Pels, M. 2005. Rescue of the Seychelles warbler on Cousin Island, Seychelles: The 

rolr of habitat restoration. Biological Conservation 124: 15-26. 

Komdeur, J. 1992. Importance of habitat saturation and territory quality for evolution of cooperative 

breeding in the Seychelles warbler. Nature 358: 493-495. 

Komdeur, J. 1994. Conserving the Seychelles Warbler Acrocephalus sechellensis by translocation 

from Cousin Island to the islands of Aride and Cousine. Biological Conservation 67: 143-152. 

6

Komdeur, J. 2001. Mate guarding in the Seychelles warbler is energetically costly and adjusted to 

paternity risk. Proceedings of the Royal Society London, Biological Sciences 268(1481): 2103-2111. 

Komdeur, J. 2003. Daughters on request: about helpers and egg sexes in the Seychelles warbler. 

Proceedings of the Royal Society London, Biological Sciences 270(1510): 3-11. 

Komdeur, J. et al 1991. Conserving the Seychelles Warbler by translocation: a transfer from Cousin 

Island to Aride Island. Bird Conservation International 1(2): 177-185. 

Komdeur, J. et al 2002. Pre-ovulation control of hatchling sex ratio in the Seychelles warbler. 


Komdeur, J. et al 2004. Experimental evidence that kin discrimination in the Seychelles warbler is 

based on association and not on genetic relatedness. Proceedings of the Royal Society London, 

Biological Sciences 271(1542): 963-969. 

Komdeur, J. et al 2004. Why Seychelles warblers fail to recolonize nearby islands: unwilling or 

unable to fly there Ibis 146: 298-302. 

Komdeur, J. et al 2007. Explicit experimental evidence for the effectiveness of proximity as mateguarding 

behaviour in reducing extra-pair fertilization in the Seychelles warbler. Molecular Ecology 

16(17): 3679-3688. 

Richardson, D.S. et al 2001. Parentage assignment and extra-group paternity in a cooperative 

breeder: the Seychelles warbler (Acrocephalus sechellensis). Molecular Ecology 10(9): 2263-2273. 

Richardson, D.S. et al 2002. Direct benefits and the evolution of female-biased cooperative breeding 

in Seychelles Warblers. Evolution 56(11): 2313-2321. 

Richardson, D.S. et al 2003. Sex-specific associative learning cues and inclusive fitness benefits in 

the Seychelles warbler. Journal of Evolutionary Biology 16(5): 854-861. 

Richardson, D.S. et al 2004. Inbreeding in the Seychelles warbler: environment-dependent maternal 

effects. Evolution 58(9): 2037-2048. 

Richardson, D.S. et al 2005. MHC-based patterns of social and extra-pair mate choice in the 

Seychelles warbler. Proceedings of the Royal Society London, Biological Sciences 272(1564): 759- 

767. 

Richardson, D.S. et al 2006. Translocation of the Seychelles warbler Acrocephalus sechellensis to 

establish a new population on Denis Island, Seychelles. Conservation Evidence 3: 54-57. 

van de Crommenacker, J. et al 2011. Spatio-temporal variation in territory quality and oxidative 

status: a natural experiment in the Seychelles warbler (Acrocephalus sechellensis). Journal of Animal 

Ecology 80(3): 668-680. 

Veen, T. et al 2000. Experimental evidence for innate predator recognition in the Seychelles warbler. 


Rodrigues Warbler 

Acrocephalus rodericanus [A. Newton 1865, Rodrigues Island]. 

Rodrigues Island (Mascarene Islands, W Indian Ocean). 

Other names: Rodrigues Swamp-warbler, Rodrigues Brush Warbler. 

Showler, D. 2002. Encouraging news from Rodrigues. World Birdwatch 24(1): 20-21. 

Showler, D.A. 2002. Short notes (Extension of breeding activity for Rodrigues Warbler Acrocephalus 

rodericanus). Bulletin of the African Bird Club 9(1): 64. 

Showler, D.A. et al 2002. Population census and habitat use of Rodrigues Warbler Acrocephalus 

rodericanus. Bird Conservation International 12: 211-230. 

Great Reed Warbler 

Acrocephalus arundinaceus [Linnaeus 1758, Gdañsk, Poland]. 

[A.a. arundinaceus] NW Africa, continental Europe, Turkey & the Levant E to Caspian Sea & Volga 

basin. Winters Africa S of Sahara. 

[A.a. zarudnyi] N Iraq & Iran and E from the Caspian Sea to Altai, NW Mongolia & W China. Winters 

Africa S of Sahara. 


Bird Study 34: 226-234. 

Beier, J. et al 1996. Erster Nachweis eines Hybriden zwischen Teichrohrsänger Acrocephalus 

scirpaceus und Drosselrohrsänger A. arundinaceus [First record of a hybrid between Reed Warbler 

Acrocephalus scirpaceus and Great Reed Warbler A. arundinaceus]. Limicola 10(6): 304-310. 

Bosschieter, L. et al 2010. Modelling small-scale dispersal of the Great Reed Warbler Acrocephalus 

arundinaceus in a fragmented landscape. Ardea 98(3): 383-394. 

7

Cottridge, D. 1992. Waves of Spring Migrants. Birds Illustrated 1(2): 5-12. 

Dickson, W. & Scott, B. 1997. News and Comment (Just as you thought 'Acros' were getting easier). 

British Birds 90(6): 256. 

Gutiérrez, R. 1998. Notes (Little Bittern's reaction to attack by Great Reed Warbler). British Birds 

91(6): 232. 

Drycz, A. 1977. Polygamy and breeding success among Great Reed Warblers Acrocephalus 

arundinaceus at Milicz, Poland. Ibis 191(1): 73-77. 

Graveland, J. 1998. Reed die-back, water level management and the decline of the Great Reed 

Warbler Acrocephalus arundinaceus in The Netherlands. Ardea 86: 187-201. 

Grim, T. 1999. The food of Great Reed Warbler (Acrocephalus arundinaceus) nestlings. Sylvia 35: 

93-99. 




Hasselquist, D. et al 1995. Low frequency of extrapair paternity in the polygynous great reed warbler, 

Acrocephalus arundinaceus. Behavioral Ecology 6(1): 27-38. 

Hedenstrom, A. et al 1993. Migration, stopover and moult of the Great Reed Warbler Acrocephalus 

arundinaceus in Ghana, West Africa. Ibis 135(2): 177-180. 

Heunks, E. 1999. Leucistische Grote Karekiet bij Tienhoven in juni 1998 [Leucistic Great Warbler 

near Tienhoven in June 1998]. Dutch Birding 21(2): 100. 

Holmbring, J-A. 1973. The Great Reed Warbler Acrocephalus arundinaceus in Sweden in 1971 and 

a review of its earlier status. Vår Fågelvärld 32: 23-31. 

Jackson, W.T. & Stone, D.A. 1983. Notes (Great Reed Warbler attacking Reed Warblers). British 

Birds 76(10): 456. 

King, J.R. 1994. Initiation of remige moult by first-year Great Reed Warblers Acrocephalus a. 

arundinaceus in Europe. Ringing & Migration 15: 123-126. 

Kralj, J. et al 2007. Migration of Central and East European Acrocephalus Warblers at the Eastern 

Adriatic Coast: An Analysis of Recoveries. The Ring 29(1-2): 121-131. 

Lansdown, P. 1992. Mystery Photographs (Great Reed Warbler). British Birds 85(5): 249-250. 

Lekagul, B. et al 1985. Birdwatching in Thailand. British Birds 78(1): 2-39 (25-28). 

Magnani, A. & Serra, L. 1996. Great Reed Warblers Acrocephalus arundinaceus arundinaceus 

performing complete remex moult before post-breeding migration. Avocetta 20(2): 153-154. 

Maragna, P. & Pesente, M. 1997. Complete moult confirmed in a Great Reed Warbler Acrocephalus 

arundinaceus population breeding in northern Italy. Ringing & Migration 18: 57-58. 

Palfery, J. 1989. Notes (Great Reed Warblers feeding from spiders' webs). British Birds 82(8): 373. 

Pearson, D.J. 1975. The timing of complete moult in the Great Reed Warbler Acrocephalus 

arundinaceus. Ibis 117(4): 506-509. 

Riddiford, N. & Potts, P. 1993. Notes (Exceptional claw-wear of Great Reed Warbler). British Birds 

86(11): 572. 

Rooke, S. 2002. Monthly Marathon Solution (Great Reed Warbler). British Birds 95(12): 666-667. 


17(5): 229-239. 

Trnka, A. et al 2009. Interacting effects of vegetation structure and breeding patterns on the survival 

of Great Reed Warbler Acrocephalus arundinaceus nests. Ardea 97(1): 109-116. 

Urano, E. 1990. Factors affecting the cost of polygynous breeding for female Great Reed Warblers 

Acrocephalus arundinaceus. Ibis 132: 584-594. 

Williams, E. 1997. Unusual records of Palaearctic warblers Sylviidae in Ivory Coast. Malimbus 19: 

33-34. 

Oriental Reed Warbler 

Acrocephalus orientalis [Temminck and Schlegel 1847, Japan]. 

C Mongolia & SE Russia S to N & E China, Korea, Sakhalin Island & Japan. Winters mainly SE Asia 

to Greater Sundas, Philippines & Wallacea. 

Other name: Eastern Great Reed Warbler. 

1st WP Record: 28 February 1988. Eilat, Israel. Remained until 13 April (Shirihai 1996). 

Carter, C. 1998. Identification of large Acrocephalus warblers at Candaba Swamp, Luzon, 

Philippines. Oriental Bird Club Bulletin 27: 52-53. 

Dyrcz, A. & Nagata, H. 2002. Breeding ecology of the Eastern Great Reed Warbler at Lake 

Kasumigaura, central Japan. Bird Study 49: 166-171. 

8

Ezaki, Y. 1984. Notes on the Moult of the Eastern Great Reed Warbler Acrocephalus arundinaceus 

orientalis in the Breeding Grounds. Journal of the Yamashina Institute for Ornithology 16: 88-91. 

Harrap, S. 2003. Potential eastern vagrants: a photo gallery. Birding World 16(8): 342-346. 

King, J.R. 1996. Moult of Oriental Great Reed Warbler. Dutch Birding 18(2): 82. 


Nisbet, I.C.T. & Lord Medway 1972. Dispersion, population ecology and migration of Eastern Great 

Reed Warblers Acrocephalus orientalis wintering in Malaysia. Ibis 114(4): 451-494. 

Saitou, T. 1976. Breeding biology of the Eastern Great Reed Warbler, Acrocephalus arundinaceus 

orientalis. Journal of the Yamashina Institute for Ornithology 8(2): 135-156. 

Shirihai, H. 1996. The Birds of Israel. Academic Press. 


17(5): 229-239. 

Clamorous Reed Warbler 

Acrocephalus stentoreus [Hemprich and Ehrenberg 1833, Dumyât, N Egypt]. 

[A.s. stentoreus] Egypt, Sinai & the Levant. 

[A.s. brunnescens] NE Sudan, NW Somalia, Arabian peninsula and SC Kazakhstan S to S Iraq & Iran 

and E to N Indian subcontinent. Winters peninsular India. 

[A.s. meridionalis] S India & Sri Lanka. 

[A.s. amyae] NE India E to N Myanmar & S China. 

[A.s. siebersi] W Java. 

[A.s. harterti] Philippines. 

[A.s. lentecaptus] Borneo, C & E Java and Lombok & Sumbawa (W Lesser Sundas). 

[A.s. celebensis] S Sulawesi. 

[A.s. sumbae] Buru (S Moluccas) and Sumba & Timor (C & E Lesser Sundas) to New Guinea, 

Bismarck Archipelago & Solomon Islands. 

Other names: Indian Great Reed Warbler (brunnescens), Sri Lanka Great Reed Warbler 

(meridionalis), Assam Great Reed Warbler (amyae), Heinroth's Reed Warbler (celebensis). 

Oskar August Heinroth (1871-1945), German doctor & zoologist. 

Carter, C. 1998. Identification of large Acrocephalus warblers at Candaba Swamp, Luzon, 

Philippines. Oriental Bird Club Bulletin 27: 52-53. 

Castell, P. 1999. Clamorous Reed Warbler Acrocephalus stentoreus apparently predating nest of 

Booted Warbler Hippolais caligata in the United Arab Emirates. Sandgrouse 21(2): 177. 

Crumbie, M. 1986. Observations to clarify the status of the reef heron (Egretta gularis), little green 

heron (Butorides striatus) and clamorous reed warbler (Acrocephalus stentoreus) in Abu Dhabi Island, 

1985, with additional notes. Emirates Natural History Group Bulletin 28: 10-28. 

Gantlett, S. 1993. Photo-forum: Clamorous Reed Warbler. Birding World 6(6): 250-251. 

George, P.V. 1961. On the Indian Great Reed Warbler, Acrocephalus stentoreus (Hemprich & 

Ehrenberg) breeding in Kerala. Journal of the Bombay Natural History Society 58(3): 797-798. 

Khoury, F. et al 2006. Observations on the avifauna of the eastern Jordan Valley, during July–August 

2005. Sandgrouse 28(2): 119-126. 

Kinzelbach, R.K. 1986. New records of Goliath Heron, Yellow-billed Stork, Blyth's Reed Warbler and 

Clamorous Reed Warbler in the Middle East. Bulletin of the Ornithological Society of the Middle East 

17: 13-17. 

Laird, W. 1992. Notes (Plumage variation of Clamorous Reed Warblers in Israel). British Birds 85(2): 

83-85. 


Madge, S.C. 1987. Mystery photographs (Clamorous Reed Warbler). British Birds 80(6): 280-282. 

Meadows, B.S. 2009. Status, seasonality and distribution of the Clamorous Reed Warbler 

Acrocephalus stentoreus at Khafrah Marsh, Eastern Province, Saudi Arabia. Sandgrouse 31(2): 182- 

188. 

Morgan, J.H. 2004. Wing lengths of Clamorous Reed Warblers Acrocephalus stentoreus in Israel. 

The Ring 26: 2. 

Parasharya, B.M. et al 1995. The Great Reed Warbler Acrocephalus stentoreus (Hemprich & 

Ehrenberg) feeding on fruits of Salvadora persica. Journal of the Bombay Natural History Society 

92(3): 420-421. 

Roselaar, C.S. 1994. Geographical variation within western populations of Clamorous Reed Warbler. 

Dutch Birding 16(6): 237-239. 

9


17(5): 229-239. 

Australian Reed Warbler 

Acrocephalus australis [Gould 1838, Parramatta, New South Wales, Australia]. 

[A.a. australis] E & SE Australia & Tasmania. 

[A.a. gouldi] SW Australia. 

[A.a. carterae] NW Australia. 

Other name: Long-billed Reed Warbler. 

Lenz, M. 1989. Occupancy of land-based territories by Clamorous Reed-Warblers in Canberra during 

a drought year. Corella 13: 91-92. 

Lenz, M. 1989. Regular double-brooding by Clamorous Reed-Warblers Acrocephalus stentoreus in 

the Canberra region. Australian Bird Watcher 13: 73-77. 

McKean, J.L. 1983. Some notes on the occurrence of the Great Reed Warbler Acrocephalus 

arundinaceus in the Northern Territory. Northern Territory Naturalist 6: 3-8. 

Welbergen, J. et al 2001. Egg discrimination in the Australian reed warbler (Acrocephalus australis): 

rejection response toward model and conspecific eggs depending on timing and mode of artificial 

parasitism. Behavioral Ecology 12(1): 8-15. 

Wooller, S.J. & Wooller, R.D. 1997. Overwintering by Clamorous Reed-Warblers, Acrocephalus 

stentoreus, in Perth, Western Australia. Emu 97(4): 332-334. 

Millerbird 

Acrocephalus familiaris [Rothschild 1892, Laysan Island, Hawaii]. 

[A.f. familiaris] Formally Laysan, Hawaii. Extinct by 1923 following the introduction of the European 

Rabbit to the island. 

[A.f. kingi] Nihoa (Hawaii). 

Other names: Hawaiian Reed-warbler; Laysan Millerbird (familiaris), Nihoa Millerbird (kingi), Nihoa 

Warbler (kingi). 

Nightingale Reed Warbler 

Acrocephalus luscinius [Quoy and Gaimard 1830, Guam]. 

[A.l. luscinius] Saipan & Alamagan Island (N Mariana Islands, W Pacific). 

[A.l. nijoi] Aguijan (aka Agiguan), (N Mariana Islands, W Pacific). 

[A.l. yamashinae] Formally Pagan Island, (N Mariana Islands, W Pacific). Presumed extinct since 

about 1980. 

Reichel, J.D. et al 1992. Island extinctions: the case of the endangered Nightingale Reed-Warbler. 

Wilson Bulletin 104(1): 44-54. 

Nauru Reed Warbler 

Acrocephalus rehsei [Finsch 1883, Nauru]. 

Nauru (Melanesia, W Pacific). 

Other names: Finsch's Reed Warbler, Pleasant Warbler. 

Friedrich Hermann Otto Finsch (1839-1917), German ethnographer, naturalist & traveller. 

Buden, D.W. 2008. The birds of Nauru. Notornis 55(1): 8-19. 

Eds. 2004. News digest (Stamp of approval). Birdwatch 139: 19. 

Finsch, O. 1883. On a new Reed-Warbler from the island of Nawodo, or Pleasant Island, in the 

Western Pacific. Ibis 5(1): 142-144. 

Caroline Islands Warbler 

Acrocephalus syrinx [Kittlitz 1835, Woleai Atoll, Caroline Islands]. 

Woleai, Lamotrek, Truk, Pohnpei, Nukuoro & Kosrae (Caroline Islands, W Pacific). 

Other names: Carolinian Reed Warbler. 

Bokikokiko 

Acrocephalus aequinoctialis [Latham 1790, Kiritimati (Christmas Island), E Polynesia]. 

[A.a. aequinoctialis] Kiritimati (North Line Islands, Kiribati, C Pacific). 

10

[A.a. pistor] Teraina & formally Tabuaeran (North Line Islands, Kiribati, C Pacific). Presumed extinct 

on Tabuaeran. 

Kiritimati formally known as Christmas Island. Teraina formally known as Washington Island. 

Tabuaeran formally known as Fanning Island. Kiribati formally known as the Gilbert Islands. 

Other names: Kiritimati Reed Warbler, Polynesian Reed Warbler, Christmas Island Warbler, Line 

Islands Reed Warbler, Equinoctial Warbler, Kokikokiko. 

IUCN et al 1982. Notes on the nesting behaviour of Acrocephalus aequinoctialis. Bulletin of the British 

Ornithologists' Club 102(1): 20-22. 

Milder, S.L. & Schreiber, R.W. 1989. The vocalizations of the Christmas Island Warbler 

Acrocephalus aequinoctialis, an island endemic. Ibis 131(1): 99-111. 

Pierce, R. et al 2007. Surveys and capacity building in Kiritimati (Christmas Island, Kiribati), June 

2007, to assist in restoration of populations of Bokikokiko and seabirds. A Report for the Wildlife 

Conservation Unit, Kiritimati, Republic of Kiritibati, funded by Invasive Species Specialist Group 

(IUCN, Auckland, New Zealand), Secretariat for the Pacific Regional Environment Programme (Apia, 

Samoa), and the British High Commission (Suva, Fiji). 

Northern Marquesan Reed Warbler 

Acrocephalus percernis [Wetmore 1919, Nuku Hiva, Marquesas Islands]. 

[A.p. percernis] Nuku Hiva (C Marquesas Islands, N French Polynesia, C Pacific). 

[A.p. postremus] Hatutaa (N Marquesas Islands, N French Polynesia, C Pacific). 

[A.p. aquilonis] Eiao (NW Marquesas Islands, N French Polynesia, C Pacific). 

[A.p. idae] Ua Huka (C Marquesas Islands, N French Polynesia, C Pacific). 

Cibois, A. et al 2007. Uniform phenotype conceals double colonization by reed-warblers of a remote 

Pacific archipelago. Journal of Biogeography 34(7): 1150-1166. 

Tahiti Reed Warbler 

Acrocephalus caffer [Sparrman 1786, Tahiti]. 

[A.c. caffer] Tahiti (Windward group of Society Islands, S French Polynesia, SC Pacific). 

[A.c. longirostris] Formally Moorea, now extinct (Windward group of Society Islands, S French 

Polynesia, SC Pacific). 

[A.c. garretti] Formally Huahine, now extinct (Leeward group of Society Islands, S French Polynesia, 

SC Pacific). 

[A.c. musae] Formally Raiatea, now extinct (Leeward group of Society Islands, S French Polynesia, 

SC Pacific). 

Other names: Long-billed Reed Warbler, Tahitian Warbler, Komako, Polynesian Warbler. 

Evans, K. 2004. Home and away (Tahiti). Birding World 17(7): 304. 

Cibois, A. et al 2008. Systematics of the extinct reed warblers Acrocephalus of the Society Islands of 

eastern Polynesia. Ibis 150: 365-376. 

Southern Marquesan Reed Warbler 

Acrocephalus mendanae [Tristram 1883, Hivaoa or Tahuata, Marquesas Islands]. 

[A.m. mendanae] Hiva Oa & Tahuata (SC Marquesas Islands, N French Polynesia, C Pacific). 

[A.m. dido] Ua Pou (C Marquesas Islands, N French Polynesia, C Pacific). 

[A.m. consobrina] Moho Tani (S Marquesas Islands, N French Polynesia, C Pacific). 

[A.m. fatuhivae] Fatu Hiva (S Marquesas Islands, N French Polynesia, C Pacific). 

Previously treated as conspecific with Northern Marquesan Reed Warbler. 

Cibois, A. et al 2007. Uniform phenotype conceals double colonization by reed-warblers of a remote 

Pacific archipelago. Journal of Biogeography 34(7): 1150-1166. 

Tuamotu Reed Warbler 

Acrocephalus atyphus [Wetmore 1919, Fakarava, Tuamotu Archipelago]. 

[A.a. atyphus] Islands in N & W Tuamotu Archipelago (French Polynesia, S Pacific). 

[A.a. ravus] Atolls in SE Tuamotu Archipelago (French Polynesia, S Pacific). 

[A.a. palmarum] Anaa Island (WC Tuamotu Archipelago, French Polynesia, S Pacific). 

[A.a. niauensis] Niau (NW Tuamotu Archipelago, French Polynesia, S Pacific). 

11

[A.a. eremus] Makatea Island (NW Tuamotu Archipelago, French Polynesia, S Pacific). 

[A.a. flavidus] Napuka Island (N Tuamotu Archipelago, French Polynesia, S Pacific). 

Cook Islands Reed Warbler 

Acrocephalus kerearako [Holyoak 1974, Mangaia, Cook Islands]. 

[A.k. kerearako] Mangaia (SE Cook Islands, S Pacific). 

[A.k. kaoko] Mitiaro (SE Cook Islands, S Pacific). 

Other names: Kerearako (kerearako), Kaoko (kaoko). 

Gill, B.J. 1996. Notes on certain Cook Islands birds. Notornis 43(3): 154-158. 

Holyoak, D.T. 1974. Undescribed land birds from the Cook Islands, Pacific Ocean. Bulletin of the 

British Ornithologists' Club 94: 145-150. 

Rimatara Reed Warbler 

Acrocephalus rimitarae [Murphy and Mathews 1929, Rimatara, Austral Islands]. 

Rimatara (Austral Islands, W French Polynesia, CS Pacific). 

Graves, G.R. 1992. The endemic land birds of Henderson Island, southeastern Polynesia: notes on 

natural history and conservation. The Wilson Bulletin 104(1): 32-43 (38). 

Henderson Reed Warbler 

Acrocephalus taiti [Ogilvie-Grant 1913, Henderson Island, Pitcairn Islands]. 

Henderson Island (Pitcairn group, C Pacific). 

This species is named after Henderson Island, which in turn is named after Captain James 

Henderson, master of the vessel Hercules, which visited the island in 1819. 

Bourne, W.R.P. & David, A.C.F. 1983. Henderson Island, central south Pacific, and its birds. 

Notornis 30: 233-252. 

Brooke, M. 2010. Important Bird Areas, Henderson Island. British Birds 103(8): 428-444. 

Brooke, M, deL. & Hartley, I.R. 1995. Nesting Henderson Reed-Warblers (Acrocephalus vaughani 

taiti) studied bu DNA fingerprinting: unrelated coalitions in a stable habitat The Auk 112(1): 77-86. 


natural history and conservation. The Wilson Bulletin 104(1): 32-43 (37-40). 

Ogilvie-Grant, W.R. 1913. On a small collection of birds from Henderson Island, South Pacific. Ibis 

55(3): 343-350. 

Wragg, G.M. & Weisler, M.I. 1994. Extinctions and new records of birds from Henderson Island, 

Pitcairn group, south Pacific Ocean. Notornis 41(1): 61-70. 

Pitcairn Reed Warbler 

Acrocephalus vaughani [Sharpe 1900, Pitcairn Island]. 

Pitcairn Island (C Pacific). 

Sparrow (local name). 

Brooke, M. 2010. Important Bird Areas, Henderson Island. British Birds 103(8): 428-444. 


natural history and conservation. The Wilson Bulletin 104(1): 32-43 (38). 

Black-browed Reed Warbler 

Acrocephalus bistrigiceps [Swinhoe 1860, Xiamen (Amoy), Fujian, China]. 

E Transbaicalia E to S Amurland, Ussuriland and Sakhalin, N Mongolia, C & E China, Korea, 

Hokkaido & Honshu. Winters S China, Indochina, NE India & Bangladesh. 

Other name: Von Schrenck's Reed Warbler. 

Leopold Ivanovich von Schrenck (1826-1894), Russo-German zoologist & geographer. 

Cooper, D. & Kay, B. 2010. Autumn bird migration on Hegura-jima, Japan. Birding World 23(8): 355- 

364 (360, plate 11). 

Cooper, D. & Kay, B. 2010. Hegura-Jima - the Fair Isle of Japan. Birding World 22(12): 506-522 

(516, plate 28). 

12

Hamao, S. & Saito, D.S. 2005. Extrapair fertilization in the black-browed reed warbler (Acrocephalus 

bistrigiceps): effects on mating status and nesting cycle of cuckolded and cuckolder males. The Auk 

122(4): 1086-1096. 

Hamao, S. 2003. Reduction of cost of polygyny by nest predation in the Black-browed Reed Warbler. 

Ornithological Science 2(2): 113-118. 

Hamao, S. 2005. Predation risk and nest-site characteristics of the Black-browed Reed Warbler 

Acrocephalus bistrigiceps: the role of plant strength. Ornithological Science 4(2): 147-153. 

Hamao, S. 2008. Syntactical complexity of songs in the Black-browed Reed Warbler Acrocephalus 

bistrigiceps. Ornithological Science 7(2): 173-177. 

Harrap, S. 2003. Potential eastern vagrants: a photo gallery. Birding World 16(8): 342-346. 


Round, P.D. & Fisher, T.H. 2009. Records of Black-browed Reed Warbler Acrocephalus bistrigiceps 

from Luzon, Philippines. Forktail 25: 159-160. 

Tipling, D. 1992. The great call of China. Birds Illustrated 1(4): 65-71. 

Ueda, K. & Yamaoka, A. 1994. Polygyny in Schrenck's Reed Warbler Acrocephalus bistrigiceps. Ibis 

136: 492-493. 

Moustached Warbler 

Acrocephalus melanopogon [Temminck 1823, near Rome, Italy]. 

[A.m. melanopogon] S Europe, NW Africa & Ukraine. Winters N Africa & the Levant. 

[A.m. mimicus] Transcaspia, S Iran & S Iraq. Winters Azerbaijan NE to Tadzhikistan, E Saudi Arabia 

& NW India. 

[A.m. albiventris] Shores of Sea of Azov, lower R Don & Krasnodar area (SE Ukraine & adjacent S 

Russia). 

Baral, H.S. et al 2004. Moustached Warbler Acrocephalus melanopogon: first record for Nepal. 

Forktail 20: 93. 

Bibby, C.J. 1982. Studies of west Palearctic birds; 184. Moustached Warbler. British Birds 75(8): 

346-359. 

Bonaccorsi, G. 1999. Nouveau cas de reproduction de la Lusciniole à moustaches Acrocephalus 

melanopogon en Corse [A new case of breeding Moustached Warbler Acrocephalus melanopogon in 

Corsica]. Alauda 67(3): 235. 

Bourne, W.R.P. 2007. Letters (The Cambridge 'Moustached Warblers'). British Birds 100(1): 63. 

Bowey, K. 1995. Notes (Apparent female Moustached Warbler singing). British Birds 88(2): 113. 

Bradshaw, C. 2000. From the Rarities Committee's Files: The occurrence of Moustached Warbler in 

Britain. British Birds 93(1): 29-38. 

British Ornithologists’ Union 2006. British Ornithologists’ Union Records Committee: 33rd Report 

(April 2006). Ibis 148: 594. 

Cantera, J-P. 1999. Statut de reproduction de la Lusciniole à moustaches Acrocephalus 

melanopogon en Corse [Reproductive status of the Moustached Warbler Acrocephalus melanopogon 

in Corsica]. Alauda 67(3): 233-234. 

Chytil, J. 1996. Occurrence of Moustached Warbler (Acrocephalus melanopogon) in the Czech 

Republic. Sylvia 32(1): 66-70. 


Eds. 2006. Pigeonhole (Moustached Warbler off at last!). Birding World 19(7): 308. 

Hinde, R.A. & Thom, A.S. Moustached Warblers in Cambridgeshire, pp. 13-18. Found in: Sharrock, 

J.T.R. & Grant, P.J. 1982. Birds new to Britain and Ireland. T. & A.D. Poyser. 

Kleindorfer, S.H. et al 1996. Alarm calls and chick reactions in the Moustached Warbler, 

Acrocephalus melanopogon. Animal Behaviour 51: 1199-1206. 

Kleindorfer, S.H. et al 1998. Clarification of the chick reaction hypothesis. Animal Behaviour 55: 504- 

507. 

Madge, S. 1992. Identification of Moustached Warbler. Birding World 5(8): 299-303. 

Melling, T. 2006. Time to get rid of the Moustache: a review of British records of Moustached 

Warbler. British Birds 99(9): 465-478. 

Oreel, G.J. 1981. Letters (Tail-cocking by Moustached Warblers). British Birds 74(10): 446. 

Payne, D. 2005. Reports (Warbler sings for a select audience. Moustached Warbler: Dodman Point, 

Cornwall, 15-17 May 2005). Birdwatch 157: 58. 

Ranner, A. 1992. Letters (More on Moustached Warbler). Birding World 5(11): 446. 

Vinicombe, K. 2002. British Vagrants (A tale of two warblers). Birdwatch 118: 22-25. 

Wallace, D.I.M. 1981. Letters (Tail-cocking by Moustached Warblers). British Birds 74(10): 446. 

13

Williams, T. 1983. Notes (Song period of Moustached Warbler). British Birds 76(10): 456. 

Aquatic Warbler 

Acrocephalus paludicola [Vieillot, 1817, Lorraine and Picardy, NE France]. 

Fragmented and localised thru E Germany, Poland, Lithuania, Latvia, Belarus, Ukraine & Hungary 

and possibly further E towards the River Ob (Russia). Winters W Africa S of the Sahara. 

Aquatic Warbler Conservation Team 1999. World population, trends and conservation status of the 

Aquatic Warbler Acrocephalus paludicola. Vogelwelt 120: 65-85. 

Atienza, J.C. 2001. Migration and conservation of the Aquatic Warbler Acrocephalus paludicola in 

Spain. Ardeola 48(2): 197-208. 

Bargain, B. et al 2008. Première zone d’hivernage du Phragmite aquatique Acrocephalus paludicola 

découverte en Afrique [First wintering site for Aquatic Warbler Acrocephalus paludicola discovered in 

Africa]. Ornithos 15(6): 411-425. 

Bil, W. 1987. Veel Waterrietzangers bij Makkum in zomer van 1986 [Many Aquatic Warblers near 

Makkum in summer of 1986]. Dutch Birding 9(4): 176. 

Birkenhead, T.R. 1993: Avian mating systems: The Aquatic Warbler is unique. Trends in Ecology & 

Evolution 8: 390-391. 

Boyd, M. 1989. Mystery Photographs (Aquatic Warbler). British Birds 82(2): 68-69. 

Catchpole, C.K. & Leisler, B. 1989: Variation in the song of the Aquatic Warbler Acrocephalus 

paludicola in response to playback of different song structures. Behaviour 108: 125-138. 

Catchpole, C.K. & Leisler, B. 1996: Female Aquatic Warblers (Acrocephalus paludicola) are 

attracted by playback of longer and more complicated song. Behaviour 133: 1153-1164. 

Chadwick, P. 1984. Letters (Aquatic Warbler and streaked Sedge Warbler). British Birds 77(8): 378. 

de By, R.A. 1990. Migration of Aquatic Warbler in western Europe. Dutch Birding 12(4): 165-181. 

Dyrcz, A. & Zdunek, W. 1993. Breeding ecology of the Aquatic Warbler Acrocephalus paludicola on 

the Biebrza marshes, northeast Poland. Ibis 135: 181-189. 

Dyrcz, A. & Zdunek, W. 1993. Breeding statistics of the Aquatic Warbler Acrocephalus paludicola on 

the Biebrza marshes, northeast Poland. Journal of Ornithology 134: 317-323. 

Dyrcz, A. 1989. Polygyny in the Aquatic Warbler Acrocephalus paludicola. Ibis 131: 298-300. 

Dyrcz, A. 1993. Biometrical differences between sexes in the breeding population of Aquatic warbler 

Acrocephalus paludicola. Ringing & Migration 14: 149-151. 

Dyrcz, A. 1993. Nesting biology of the Aquatic Warbler Acrocephalus paludicola in the Biebrza 

marshes (NE Poland). Vogelwelt 114: 2-15. 

Eaton, M.A. et al 2009. Birds of Conservation Concern 3, The population status of birds in the United 

Kingdom, Channel Islands and Isle of Man. British Birds 102(6): 296-341. 

Evans, P.G.H. & Lovegrove, R.R. 1974. The Birds of the South West Irish Islands. Irish Bird Report 

2(9): 33-64. 

Flumm, D. 1997. Sou'west by west. Birdwatch 62: 19-23. 

Flumm, D.S. 1984. Letters (Identification pitfalls: Aquatic Warbler). British Birds 77(8): 377. 

Gorman, G. 2002. Aquatic Warbler. Alula 8(2): 62-65. 

Grantham, M. 2004. Ringing in August. Birding World 17(7): 306-307. 

Heredia, B. 1993. The status and conservation of Aquatic Warbler. Birding World 6(7): 294-295. 

Herroelen, P. 1991. Trek van Waterrietzanger in West-europa [Migration of Aquatic Warbler in 

western Europe]. Dutch Birding 13(2): 65-66. 

Juillard, R. et al 2006. Identifying autumn migration routes for the globally threatened Aquatic 

Warbler Acrocephalus paludicola. Ibis 148: 735-743. 

Kovacs, G. & Vegvari, Z. 1999. Population size and habitat of the Aquatic Warbler Acrocephalus 

paludicola in Hungary. Vogelwelt 120: 121-125. 

Kozulin, A. & Flade, M. 1999. Breeding habitat, abundance and conservation status of the Aquatic 

Warbler Acrocephalus paludicola in Belarus. Vogelwelt 120: 97-111. 

Kozulin, A. et al 1998. Distribution and number of Aquatic Warbler in Belarus. Subbuteo 1: 3-16. 

Kozulin, A. et al 2004. Factors affecting fluctuations of the Aquatic Warbler Acrocephalus 

paludicola population of Byelarussian mires. Acta Ornithologica 39(1): 47-56. 

Kozulin, A.V. et al 1999. Fen mires and the benefit of mobility: a hypothesis for the origin of 

promiscuity in Aquatic Warbler Acrocephalus paludicola. Subbuteo 2: 11-17. 

Mauer, K. 1980. Captures of Aquatic Warbler Acrocephalus paludicola on Putten in 1970-73. Dutch 

Birding 1(4): 122. 

14

Miguélez, D. et al 2009. Habitat, phenology and biometrics of the Aquatic Warbler Acrocephalus 

paludicola during autumn migration through a riverine wetland in Iberia. Ringing & Migration 24: 277- 

279. 

Miskinis, J. 2006. Home and away (The Nemunas Estuary, Lithuania). Birding World 19(8): 343. 

Neto, J.M. et al 2010. Distribution, phenology and condition of aquatic warblers Acrocephalus 

paludicola migrating through Portugal. Ardeola 57(1): 181-189. 

Oreel, G.J. 1980. Mystery bird 2: Aquatic Warbler Acrocephalus paludicola. Dutch Birding 1(4): 120- 

121. 

Pitches, A. 2007. News and Comment (And now we know where Aquatic Warblers go). British Birds 

100(4): 256. 

Porter, R. 1983. Identification pitfalls and assessment problems 6, Aquatic Warbler. British Birds 

76(8): 342-346. 

Poulin, B. et al 2010. Spring stopover of the globally threatened Aquatic Warbler Acrocephalus 

paludicole in Mediterranean France. Ardeola 57(1): 167-173. 

Rumsey, S.J.R. 1984. Letters (Identification pitfalls: Aquatic Warbler). British Birds 77(8): 377. 

Salewski, V. et al 2009. Quest for a phantom - the search for the winter quarters of the Aquatic 

Warbler Acrocephalus paludicola. Bulletin of the African Bird Club 16(1): 61-66. 

Schäffer, N. & Newbery, P. 1999. The elusive Aquatic Warbler. World Birdwatch 21(4): 23-25. 

Schäffer, N. et al 2006. The African migration and wintering grounds of the Aquatic Warbler 

Acrocephalus paludicola. Bird Conservation International 16: 33-56. 

Schulze-Hagen, K. 1989. Bekanntes und weniger Bekanntes vom Seggenrohrsänger Acrocephalus 

paludicola [Notes on Aquatic Warbler Acrocephalus paludicola]. Limicola 3(5): 229-246. 

Schulze-Hagen, K. et al 1999. The breeding system of the Aquatic Warbler Acrocephalus paludicola 

– a review of new results after 1991. Vogelwelt 120: 87-96. 

Topp, A. 1996. Sekalaista - Miscellaneous (Swarovski gives a helping hand for Aquatic Warbler). 

Alula 2(1): 40. 

Vergeichik, L. & Kozulin, A. 2006. Breeding ecology of Aquatic Warblers Acrocephalus paludicola in 

their key habitats in SW Belarus. Acta Ornithologica 41: 2. 

Vinicombe, K. 1999. Migration (Autumn exodus). Birdwatch 87: 6-10. 

Vinicombe, K. 2003. Identification matters (The right stripes). Birdwatch 134: 22-24. 

Walbridge, G. 1991. Identification of Aquatic Warbler in autumn. Birding World 4(7): 237-241. 

Wassink, A. 2009. Birds of Kazakhstan: new and interesting data, part 2. Dutch Birding 31(2): 101- 

110 (108). 

Sedge Warbler 

Acrocephalus schoenobaenus [Linnaeus, 1758, S Sweden]. 

Europe and W & C Asia. Winters Senegal E to Ethiopia and S to E Cape Province & N Namibia. 


Bird Study 34: 226-234. 

Alker, P.J. & Redfern, C.P.F. 1996. Double brooding and polygyny in Sedge Warblers Acrocephalus 

schoenobaenus breeding in north-west England. Bird Study 43: 356-363. 

Bargain, B.R. et al 2002. Importance des marais de la baie dˆAudierne (Bretagne) pour la migration 

du Phragmite des joncs Acrocephalus schoenobaenus [The Audierne bay, an important stop-over site 

for Sedge Warblers in Brittany]. Alauda 70(1): 37-55. 

Bascuitti P, et al 1997. Autumn migration strategies of the Sedge Warbler Acrocephalus 

schoenobaenus in northern Italy. Ringing & Migration 18: 59-67. 

Bermejo, A. & De la Puente, J. 2002. Stopover characteristics of Sedge Warblers (Acrocephalus 

schoenobaenus) in central Iberia. Die Vogelwarte 41: 181-189. 

Bibby, C.J. & Green, R.E. 1981. Autumn migration strategies of Reed and Sedge Warblers. Ornis 

Scandinavica 12: 1-12. 

Bibby, C.J. 1978. Some breeding statistics of Reed and Sedge Warblers. Bird Study 25: 207-222. 

Bibby, C.J. et al 1976. Sedge Warbler migration and reed aphids. British Birds 69: 384-399. 

Bowey, K. 1999. Notes (Sedge Warblers nesting in rape crops). British Birds 92(7): 371. 

Buchanan, K.L. & Catchpole, C.K. 2000. Song as an indicator of male parental effort in the sedge 

warbler. Proceedings of the Royal Society London, Biological Sciences 267: 321-326. 

Catchpole, C.K. 1973. The function of advertising song in the Sedge Warbler (Acrocephalus 

schoenobaenus) and the Reed Warbler (A. scirpaceus). Behaviour 46(1): 1-21. 

Chadwick, P. 1984. Letters (Aquatic Warbler and streaked Sedge Warbler). British Birds 77(8): 378. 

15

Chernetsov, N. & Titov, N. 2001. Movement patterns of European Reed Warblers Acrocephalus 

scirpaceus and Sedge Warblers A. schoenobaenus before and during autumn migration. Ardea 89: 

509-515. 

Chernetsov, N. 1996. Preliminary hypotheses on migration of the Sedge Warbler (Acrocephalus 

schoenobaenus) in the Eastern Baltic. Die Vogelwarte 38: 201-210. 


da Prato, E.S. 1977. Sparrowhawk taking Sedge Warbler from a ringer’s hand. Scottish Birds 9: 381. 

Grandío, J.M. 1998. Comparación del peso y su incremento, del tiempo de estancia y de la 

abundancia del Carricerín Común (Acrocephalus schoenobaenus) entre dos zonas de la marisma de 

Txingudi (N de España) [Comparison of weight and growth, length of stay and plenty of Sedge 

Warbler (Acrocephalus schoenobaenus) between two areas of the marsh Txingudi (N Spain)]. 

Ardeola 45(2): 137-142. 

Grandío, J.M. 1998. Consideraciones sobre la estrategia migratoria postnupcial del Carricerín 

Común Acrocephalus schoenobaenus en la Península Ibérica [Considerations on the autumn 

migration strategy of the Sedge Warbler Acrocephalus schoenobaenus in the Iberian Peninsula]. 

Ardeola 45(2): 213-215. 

Grandío, J.M. 1999. Migración postnupcial diferencial del Carricerín Común (Acrocephalus 

schoenobaenus) en la marisma de Txingudi (N de España) [Differential autumn migration of the 

Sedge Warbler (Acrocephalus schoenobaenus) in the marsh of Txingudi (N Spain)]. Ardeola 46(2): 

171-178. 


Gyurácz, J., & Puskás, J. 1996. The effect of cold weather fronts on the migration activity of the 

Sedge Warbler (Acrocephalus schoenobaenus) in Hungary. Ornis Hungarica 6: 43-45. 

Hałupka, L. 1996. Breeding ecology of the Sedge Warbler (Acrocephalus schoenobaenus) in Biebrza 

marshes (NE Poland). Ornis Hungarica 6: 9-14. 

Harding, B.D. 1988. Notes (Warblers fleeing from attacks by swallowtail butterfly). British Birds 81(5): 

239. 

Harrison, R. 1984. Notes (Sedge warbler feeding on the ground). British Birds 77(3): 115-116. 

Holden, P. 1985. Spring Migration 1985. British Birds 78(9): 427-434. 



Lidster, J. 2007. Monthly Marathon Solution (Sedge Warbler). British Birds 100(12): 684. 

Mellor, M. 1981. Notes (Tail-cocking by Sedge Warbler). British Birds 74(10): 444. 

O'Sullivan, O. 2004. Discover birds (Warblers - Six summer songsters). Wings 33: 24-26. 

Parker, J. 1991. Notes (Warblers fleeing from attacks by swallowtail butterfly). British Birds 84(10): 

444. 

Peach, W.J. et al 1991. Survival of British Sedge Warblers Acrocephalus schoenobaenus in relation 

to west African rainfall. Ibis 133: 300-305. 

Pranty, B. et al 2008. Annual Report of the ABA Checklist Committee 2007-2008. Birding 40(6): 32- 

38. 

Redfern, C.P.F. et al 2004. Fat and pectoral muscle in migrating Sedge Warblers Acrocephalus 

schoenobaenus. Ringing & Migration 22: 24-34. 

Sharrock, J.T.R. 1985. Mystery Photographs (Hybrid Sedge x Reed Warbler). British Birds 78(9): 

434-437. 

Shennan, N.M. 1985. Relationships between morphology and habitat selection by male Sedge 

Warblers Acrocephalus schoenobaenus. Ringing & Migration 6: 97-102. 

Thomas, D.K. 1984. Aspects of habitat selection in the Sedge Warbler. Bird Study 31: 187-194. 

Trocinska, A. et al 2001. Field studies of directional preferences of the Reed Warbler (Acrocephalus 

scirpaceus) and the Sedge Warbler (A. schoenobaenus) on autumn migration along the eastern and 

southern coast of the Baltic Sea and in western part of Ukraine. The Ring 23(1-2): 109-117. 

van Eerde, K.A. 1999. Hybride Rietzanger x Kleine Karekiet te Makkum in augustus 1997 [Hybrid 

Sedge x European Reed Warbler at Makkum in August 1997]. Dutch Birding 21(1): 34-37. 


Vinicombe, K. 2002. British Vagrants (A tale of two warblers). Birdwatch 118: 22-25. 

Vinicombe, K. 2003. Identification matters (The right stripes). Birdwatch 134: 22-24. 

Wassink, A. 1980. Sedge Warbler Acrocephalus schoenobaenus with Square-terminating 

Supercilium. Dutch Birding 1(4): 122. 

Yosef, R. & Chernetsov, N. 2004. Stopover ecology of migratory Sedge Warblers (Acrocephalus 

schoenobaenus) at Eilat, Israel. Ostrich 75(1-2): 52-56. 

16

Speckled Reed Warbler 

Acrocephalus sorghophilus [Swinhoe 1863, Xiamen (Amoy), Fujian, China]. 

Range largely unknown, possibly Liaoning & Hebei (NE China) and adjacent parts of Russian Far 

East. Winters Philippines. 

Other names: Streaked Reed Warbler, Chinese Reed Warbler. 

Blunt-winged Warbler 

Acrocephalus concinens [Swinhoe 1870, Beijing, China]. 

[A.c. concinens] N China. Winters S China, S Myanmar, Thailand & C Laos. 

[A.c. haringtoni] N Afghanistan, N Pakistan & Kashmir. Winters not clearly established. 

[A.c. stevensi] Plains of the R Brahmaputra (Assam, NE India) & adjacent parts of Bangladesh. 

Other names: Blunt-winged Paddyfield Warbler, Swinhoe's Reed Warbler. 

Robert Swinhoe (1836-1877) Anglo-Indian diplomat and naturalist. 


Mey, E. 1997. Records of Blunt-winged Warbler Acrocephalus concinens in north Vietnam. Forktail 

12: 166-167. 

Manchurian Reed Warbler 

Acrocephalus tangorum [La Touche 1912, Chin-wang-tao, NE Hebei, China]. 

Manchuria (NE China) & Ussuriland (SE Russia). Winters SE Asia. 

Other names: Manchurian Paddyfield Warbler, North China Reed-warbler. 

Alström, P. et al 1991. The taxonomic status of Acrocephalus agricola tangorum. Forktail 6: 3-13. 

Round, P.D. 1993. Winter records of the Manchurian Reed-Warbler, Acrocephalus (agricola) 

tangorum, from Thailand. Forktail 9: 83-88. 

Tipling, D. 1992. The great call of China. Birds Illustrated 1(4): 65-71. 

van der Spek, V. 2006. Cambodia - a rare bird bonanza. Alula 12(1): 26-37. 

Large-billed Reed Warbler 

Acrocephalus orinus [Oberholser 1905, Himachal Pradesh, India]. 

Riparian bushland along the Wakhan & Pamir Rivers in NE Afghanistan and adjacent SE Tajikistan. 

Winters not clearly established, possibly parts of the Indian subcontinent & SE Asia. 

Other name: Hume's Large-billed Warbler. 

Allan Octavian Hume (1826-1901), English theosophist, poet and writer on Indian birds. 

Ayé, R. et al 2010. Discovery of a breeding area of the enigmatic large-billed reed warbler 

Acrocephalus orinus. Journal of Avian Biology 41: 452-459. 

Banwell, A. 2009. World Birding roundup. Birding World 22(8): 351. 

Bensch, S. & Pearson, D. 2002. The large-billed reed warbler revisited. Ibis 144: 259-267. 

Eds. 2007. Pigeonhole. Birding World 20(3): 132. 

Eds. 2010. Frontispage (Long-billed Warbler, Zebak, Afghanistan, June 2009 - Wildlife Conservation 

Society Afghanistan). Birding World 23(1): 1. 

Pitches, A. 2007. News and Comment (Large-billed Reed Warbler rediscovered). British Birds 

100(4): 257. 

Pitches, A. 2010. News and Comment (And Acros just got a lot more interesting…). British Birds 

103(2): 134. 

Raju, D. et al 2009. A possible record of Large-billed Reed-warbler Acrocephalus orinus from Kanha 

Tiger Reserve, central India. Indian Birds 4(4): 130-133. 

Round, P.D. & Kennerley, P.R. 2007. Large-billed Reed Warbler Acrocephalus orinus back from the 

dead. BirdingASIA 7: 55-60. 

Round, P.D. et al 2007. Lost and found: the enigmatic large-billed reed warbler Acrocephalus orinus 

rediscovered after 139 years. Journal of Avian Biology 38(2): 133-138. 

Sangster, G. 2002. DB Actueel (Large-billed Reed Warbler is valid species). Dutch Birding 24(4): 

256. 

Svensson, L. et al 2008. Discovery of ten new specimens of large-billed reed warbler Acrocephalus 

orinus, and new insights into its distributional range. Journal of Avian Biology 39 (6): 605-610. 

Timmins, R.J. et al 2009. The discovery of Large-billed Reed Warblers Acrocephalus orinus in northeastern 

Afghanistan. BirdingASIA 12: 42-45. 

17

Timmins, R.J. et al 2010. New information on the Large-billed Reed Warbler Acrocephalus orinus, 

including its song and breeding habitat in north-eastern Afghanistan. Forktail 26: 9-23. 

Wassink, A. 2009. Birds of Kazakhstan: new and interesting data, part 2. Dutch Birding 31(2):101- 

110 (105). 

Paddyfield Warbler 

Acrocephalus agricola [Jerdon 1845, near Nellore, Tamil Nadu, India]. 

[A.a. agricola] Area surrounding Caspian & Aral Seas and W Mongolia, NW China, Tajikistan, E Iran 

& N Afghanistan. Winters Indian subcontinent. 

[A.a. septima] Black Sea coast from Bulgaria & Romania to S Ukraine. Winters Indian subcontinent. 

Other name: Jerdon's Reed Warbler. 

Thomas Claverhill Jerdon (1811-1872), British physician, amateur botanist & wildlife author. 

Boon, L. 1994. Finding Paddyfield Warblers in Bulgaria. Birding World 7(11): 444-445. 

Bradshaw, C. & Steele, J. 1995. Mystery Photographs (probable rama Booted Warbler). British Birds 

88(11): 561-564. 

Bradshaw, C. 1997. From the Rarities Committee's Files: Identification of pale Paddyfield Warbler. 


Butterfield, D. 1988. Paddyfield Warbler in Suffolk. Birding World 1(6): 203-205. 

Cederroth, C. 1996. Feldrohrsänger Acrocephalus agricola erstmals in diesem Jahrhundert in 

Deutschland [First 20th century record of Paddyfield Warbler Acrocephalus agricola in Germany]. 

Limicola 10(2): 83-84. 

de Jong, J. 1985. Veldrietzanger te Makkum in oktober 1984 [Paddyfield Warbler at Makkum in 

October 1984]. Dutch Birding 7(4): 140-141. 

de Knijff, P. & van Swelm, N.D. 1998. Letter (Mystery warblers in Tselinograd: an attempt to solve 

part of the problem). British Birds 91(5): 190-192. 

de Vries, J.A. 1987. Veldrietzanger ook al in oktober 1982 te Makkum [Paddyfield Warbler at 

Makkum already in October 1982]. Dutch Birding 9(1): 28. 

Dernjatin, P. 2005. Chokpak - Central Asian songbirds. Alula 11(2): 50-58. 

Dubois, P.J. 1983. Letters (Status of Paddyfield Warbler in Romania). British Birds 76(12): 585. 

Gorman, G. 2000. The Birds of the Danube Delta. Alula 6(4): 154-159 (159). 

Grantham, M. 2004. Ringing in September. Birding World 17(8): 348-349. 

Harvey, P. & Bradshaw, C. 1999. From the Rarities Committee's files: Booted Warblers with long 

supercilia: the 1976 Fair Isle bird. British Birds 92(9): 477-479. 

Hirschfeld, E. & Stawarczyk, T. 1991. First record of Paddyfield Warbler Acrocephalus agricola in 

Bahrain. Sandgrouse 13(2): 110-112. 


Lidster, J. 2005. Monthly Marathon Solution (Paddyfield Warbler). British Birds 98(3): 168-169. 

McAdams, D. & Jännes, H. 2000. British Vagrants: Paddyfield Warbler (Odd one out). Birdwatch 99: 

26-30. 

McAdams, D.G. 1994. Paddyfield Warbler in County Cork. Irish Birds 5(2): 192-195. 

Merne, O.J. & Walsh, A. 1988. Paddyfield Warbler in County Wexford - a species new to Ireland. 

Irish Birds 3(4): 592. 

Nadler, T. & Ihle, U. 1988. Beobachtungen am Feldrohrsänger Acrocephalus agricola in Bulgarien 

[Observations on Paddyfield Warbler Acrocephalus agricola in Bulgaria]. Limicola 2(6): 205-217. 

Osborn, K. 1987. Notes (Moulting on upperparts of a Paddyfield Warbler). British Birds 80(12): 634. 

Oudebeek, W.H.P. 1989. Veldrietzanger bij Zuidland in september 1987 [Paddyfield Warbler at 

Zuidland in September 1987]. Dutch Birding 11(1): 29-30. 

Pollock, C.M. et al 1994. A Paddyfield Warbler off the south Irish coast. Irish Birds 5(2): 195-197. 

Riddington, R. & Reid, J. 1997. Classic Fair Isle: September 1997. Birding World 10(10): 387-389. 

Riddington, R. 1996. The Paddyfield Warbler on Fair Isle. Birding World 9(10): 388. 

Rooke, S. 1998. Secret lives (A plainly challenging warbler). Birdwatch 75: 22-23. 

Svensson, L. 1997. Notes (Mystery warblers in Tselinograd). British Birds 90(4): 152-158. 

Svensson, L. 1999. Letters (The Kazakh 'mystery warblers' once again). British Birds 92(9): 481-482. 

Terpstra, K. 1988. Veldrietzanger op Vlieland in september 1986 [Paddyfield Warbler on Vlieland in 

September 1986]. Dutch Birding 10(2): 91. 

Tolvanen, P. et al 2005. Birding in the steppes, wetlands and forests of Kostanay region, Kazakstan. 

Alula 11(2): 64-72 (70). 

van der Have, T.M. & van der Berk, V.M. 1995. Paddyfield Warbler in Göksü delta, southern Turkey, 

in September 1991. Dutch Birding 17(1): 20-21. 

18

van der Veen, L. & Ebels, E.B. 1996 Veldrietzanger op Vlieland in september 1994 [Paddyfield 

Warbler on Vlieland in September 1994]. Dutch Birding 18(1): 13-16. 

van IJzendoorn, E.J. & Westhof, J.H.P. 1985. Veldrietzanger uit 1971 in ere hersteld [Paddyfield 

Warbler from 1971 rehabilitated]. Dutch Birding 7(4): 121-128. 

van Ree, T. 2007. Vier Veldrietzangers op Vlieland [Paddyfield Warblers in Vlieland]. Dutch Birding 

29(5): 343-344. 

Votier, S. 1997. Identification (Acrocephalus now). Birdwatch 64: 27-32. 

Wraithmell, A. 2003. Reports (Paddyfield Warbler: Dungeness, Kent, 9 September 2003). Birdwatch 

137: 51. 

Blyth's Reed Warbler 

Acrocephalus dumetorum [Blyth 1849, India]. 

S Finland E thru Russia to C Siberia and E to W Lake Baikal region & NW Mongolia and S to S 

Russia, Azerbaijan, E Iran, N Afghanistan & SW Siberia. Winters Pakistan, India & Sri Lanka. 

Edward Blyth (1810-1873), English zoologist, author and Curator of the Asiatic Society of Bengal 

between 1842 and 1864. 

Ashby, V. & Annenkova, S. 2004. Birding Kazakhstan. Birding World 17(6): 242-253. 

Bradshaw, C. 2000. Letters (Separating Acrocephalus and Hippolais warblers). British Birds 93(6): 

277. 

Bradshaw, C. 2001. From the Rarities Committee's files: Blyth's Reed Warbler: problems and pitfalls. 


Brandes, S. & Focks, O. 1994. Ein singender Buschrohrsänger Acrocephalus dumetorum in 

Schleswig-Holstein [A singing Blyth’s Reed Warbler Acrocephalus dumetorum in Schleswig-Holstein]. 

Limicola 8(1): 1-6. 

Breek, C.J. & van den Berg, A.B. 1992. Struikrietzanger te Lelystad in juni 1990 [Blyth's Reed 

Warbler at Lelystad in June 1990]. Dutch Birding 14(4): 121-126. 

Butterfield, D. 2000. Blyth’s Reed Warbler at Nigg Ferry, Highland, 7th-25th June 2000. Birding 

Scotland 3: 127-128. 

Butterfield, D. 2000. The Blyth's Reed Warbler in Highland. Birding World 13(7): 277-278. 



Ellis, P. et al 1994. The Blyth's Reed Warbler in Shetland. Birding World 7(6): 227-230. 

Fry, C.H. et al 1974. A new subspecies of Acrocephalus baeticatus from Lake Chad and a taxonomic 

reappraisal of Acrocephalus dumetorum. Ibis 116(3): 340-346. 

Golley, M. & Millington, R. 1996. Identification of Blyth's Reed Warbler in the field. Birding World 

9(9): 351-353. 

Grantham, M. 2004. Ringing in October. Birding World 17(9): 393-395. 

Haitjema, T. 2009. Letters (Blyth's Reed Warblers in Estonia). British Birds 102(1): 30. 

Harrap, H. 1989. The difficulties of Reed, Marsh and Blyth's Reed Warbler identification. Birding 

World 2(9): 318-324. 

Harvey, P.V. & Shaw, K.D. 1999. Dealing with difficult rarities - their identification and assessment. 

Number three: Blyth’s Reed Warbler, Fair Isle, September 1996. Birding Scotland 2: 14-17. 

Harvey, W.G. & Porter, R.F. 1984. Field identification of Blyth's Reed Warbler. British Birds 77(9): 

393-411. 

Harvey, W.G. & Porter, R.F. 1986. Mystery Photographs (Blyth's Reed Warbler). British Birds 79(8): 

392-396. 

Hottola, P. 1993. Alula colour of Blyth's Reed Warbler in spring. Dutch Birding 15(2): 71-72. 

Kapanen, M. 1996. Finish birds 3. Blyth's Reed Warbler - Viitakerttunen. Alula 2(3): 134-135. 

Kikkert, J. 1997. Struikrietzanger te Walem in Juni-Juli 1996 [Blyth's Reed Warbler at Walem in June- 

July 1996]. Dutch Birding 19(6): 273-276. 

Kinzelbach, R.K. 1986. New records of Goliath Heron, Yellow-billed Stork, Blyth's Reed Warbler and 

Clamorous Reed Warbler in the Middle East. Bulletin of the Ornithological Society of the Middle East 

17: 13-17. 

Lewington, I. et al 1988. The Welches Dam Acrocephalus Warbler. Birding World 1(5): 180-183. 

Lindholm, A. et al 2007. Hybrid Marsh x Blyth's Reed Warbler with mixed song in Finland in June 

2003. Dutch Birding 29(4): 223-231. 

Mädlow, W. 1992. Ein Buschrohrsänger Acrocephalus dumetorum in Berlin [Blyth’s Reed Warbler 

Acrocephalus dumetorum in Berlin]. Limicola 6(6): 292-296. 

Mullarney, K. 2007. Reports (Shy Blyth's joins the Irish list). Birdwatch 186: 67. 

19

Osborn, K. & Donald, C. 1995. Letters (Blyth's Reed Warbler identification). Birding World 8(2): 77. 

Raju, K.S.R.K. et al 1972. Movement of Blyth's Reed Warbler (Acrocephalus dumetorum Blyth) 

through Point Calimere. Journal of the Bombay Natural History Society 69(1): 186-187. 

Shaw, D. 1999. The Blyth's Reed Warbler on Fair Isle. Birding World 12(6): 238. 

Topp, A. 1996. Photonews. Alula 2(3): 141. 

van Loon, A.J. & Keijl, G.O. 2001. Blyth's Reed Warbler at Nieuwegein in June-July 1998. Dutch 

Birding 23(2): 83-85. 

Vinicombe, K. 2001. Letters (The Unst Blyth's Reed Warbler). British Birds 94(6): 291-295. 

Vinicombe, K. 2002. Identification matters (Agrocephalus). Birdwatch 124: 27-30. 

Votier, S. & Riddington, R. 1996. The two Blyth's Reed Warblers on Fair Isle. Birding World 9(6): 

221-223. 


Wallace, D.I.M. 2001. Letters (The identification of an Acrocephalus warbler). British Birds 94(9): 441. 

Reed Warbler 

Acrocephalus scirpaceus [Hermann 1804, Alsace, France]. 

[A.s. scirpaceus] NW Africa & Europe E to C European Russia, Crimea & W Asia Minor. Winters 

Africa S of Sahara. 

[A.s. fuscus] E & N Caspian Sea regions, Cacusus, E Asia Minor, Cyprus & the Levant. Winters Africa 

S of Sahara. 

[A.s. avicenniae] Coasts of S Red Sea, W Saharan Egypt & E Saharan Libya. 

Other names: Eurasian Reed Warbler, Caspian Reed Warbler (fuscus), Mangrove Reed Warbler 

(avicenniae). 

1st WP Record (A.s. avicenniae): November 2009. Near Ain Safi, W Saharan Egypt. Jens Hering, 

Stefan Brehme, Elmar Fuchs and Hans Winkler. Definate ID confirmed by using DNA analysis (Hering 

et al 2010). 


Bird Study 34: 226-234. 

Åkesson, S. et al 2001. Reed warbler orientation: initiation of nocturnal migratory flights in relation to 

visibility of celestial cues at dusk. Animal Behaviour 61: 181-189. 

Åkesson, S. et al 2002. Nocturnal migratory flight initiation in reed warblers Acrocephalus 

scirpaceus: effect of wind on orientation and timing of migration. Journal of Avian Biology 33: 349- 

357. 

Amezian, M. et al 2010. Complete moult of an undescribed resident taxon of the Reed Warbler 

Acrocephalus scirpaceus / baeticatus complex in the Smir marshes, Northern Morocco. Ardea 98(2): 

225-234. 

Andrews, J. 1991. The management and creation of reedbeds - especially for rare birds. British 

Wildlife 3(2): 81-91 (86). 

Arizaga, J. et al 2010. Short Notes (Non-breeding reed warblers Acrocephalus scirpaceus (Hermann, 

1804) in June in southern Europe: local or still migrating birds). Belgian Journal of Zoology 140(1): 

74-76. 

Ash, J.S. et al 1989. The mangrove reed warblers of the Red Sea and Gulf of Aden coasts, with 

description of a new subspecies of the African reed warbler Acrocephalus baeticatus. Bulletin of the 


Bargain, B.R. & Henry, J. 2000. Biologie de reproduction de la Rousserolle effarvatte Acrocephalus 

scirpaceus en baie d'Audierne (Bretagne, France) [Reproductive biology of the Reed Warbler 

Acrocephalus scirpaceus in Audierne Bay (Brittany, France)]. Alauda 68(2): 95-108. 

Beier, J. et al 1996. Erster Nachweis eines Hybriden zwischen Teichrohrsänger Acrocephalus 

scirpaceus und Drosselrohrsänger A. arundinaceus [First record of a hybrid between Reed Warbler 

Acrocephalus scirpaceus and Great Reed Warbler A. arundinaceus]. Limicola 10(6): 304-310. 

Bibby, C.J. & Green, R.E. 1981. Autumn migration strategies of Reed and Sedge Warblers. Ornis 

Scandinavica 12: 1-12. 

Bibby, C.J. 1978. Some breeding statistics of Reed and Sedge Warblers. Bird Study 25: 207-222. 

Bruce, K. 1997. Reed Warblers breeding in south west Scotland. Scottish Birds 19: 119-120. 

Bundy, G. 1975. Reed Warblers breeding in Shetland. British Birds 68: 210-211. 

Calvert, M. 2005. Notes (Abnormal Reed Warbler chicks). British Birds 98(8): 433. 

Calvert, M. 2005. Notes (Apparent loss of pigmentation in an aged Reed Warbler). British Birds 98(2): 

101. 

20

Calvert, M. 2005. Reed Warblers at Rostherne Mere: Working Today for Nature Tomorrow. English 

Nature, Shrewsbury. 

Cantelo, J. 1984. Notes (Identification of singing Reed and Marsh Warblers by mouth colour). British 

Birds 77(5): 214. 

Catchpole, C.K. 1973. The function of advertising song in the Sedge Warbler (Acrocephalus 

schoenobaenus) and the Reed Warbler (A. scirpaceus). Behaviour 46(1): 1-21. 

Chandler, R.J. et al 1981. Bird Photograph of the Year. British Birds 74(5): 215-218. 

Chernetsov, N. & Titov, N. 2001. Movement patterns of European Reed Warblers Acrocephalus 

scirpaceus and Sedge Warblers A. schoenobaenus before and during autumn migration. Ardea 89: 

509-515. 

Ciach, M. 2005. Notes (Reed Warbler with abnormal body-feathers). British Birds 98(8): 433. 

Clarke, R. 1989. Notes (Early Reed Warbler in Hen Harrier pellet). British Birds 82(1): 31. 


Dennis, M. 1996. Notes (Reed Warbler sunning). British Birds 89(7): 318. 


Dickson, W. & Scott, B. 1997. News and Comment (Just as you thought 'Acros' were getting easier). 


Dowsett-Lemaire, F. & Dowsett, R.J. 1987. European and African Reed Warblers, Acrocephalus 

scirpaceus and A. baeticatus: vocal and other evidence for a single species. Bulletin of the British 


Dowsett-Lemaire, F. & Dowsett, R.J. 1987. European Reed and Marsh Warblers in Africa: migration 

patterns, moult and habitat. Ostrich 58(2): 65-85. 

Dunn, P. 2001. The putative Caspian Reed Warbler in North Yorkshire. Birding World 14(8): 329-332. 

García-Peiró, I. 2003. Intraspecific variation in the wing shape of the long-distance migrant Reed 

Warbler Acrocephalus scirpaceus: effects of age and distance of migration. Ardeola 50(1): 31-37. 

Gordón, L.M.R. et al 2010. Alloparental behaviour of Eurasian Reed Warbler towards European 

Goldfinch chicks. Dutch Birding 32(1): 46-48. 


Grantham, M. 2005. Ringing in January. Birding World 18(1): 41-42. 

Harding, B.D. 1988. Notes (Warblers fleeing from attacks by swallowtail butterfly). British Birds 81(5): 

239. 

Harrap, H. 1989. The difficulties of Reed, Marsh and Blyth's Reed Warbler identification. Birding 

World 2(9): 318-324. 

Hering, J. et al 2010. African Reed Warblers and Mangrove Reed Warblers in Libya & Egypt - both 

new to the Western Palearctic. Birding World 23(5): 218-219. 

Idrissi, H.R. et al 2002. Premiers cas de nidification de la Rousserolle effarvate Acrocephalus 

scirpaceus a Sidi Bou Ghaba (Littoral Nord-Atlantique du Maroc) [First record of nesting Reed 

Warbler Acrocephalus scirpaceus in Sidi Bou Ghaba (North Atlantic Coast of Morocco)]. Alauda 70: 

223-225. 

Jackson, W.T. & Stone, D.A. 1983. Notes (Great Reed Warbler attacking Reed Warblers). British 

Birds 76(10): 456. 

Jiguet, F. et al 2010. Undescribed reed warbler breeding in Morocco. Dutch Birding 32(1): 29-36. 

Kehoe, C. 2006. Racial identification and assessment in Britain: a report from the RIACT 

subcommittee. British Birds 99(12): 619-645 (636). 

Kelsey, M.G. 1985. Notes (Mouth and gape colours of singing Marsh and Reed Warblers). British 

Birds 78(6): 297. 

Kok, D. & van Duivendijk, N. 1998. Masters of Mystery Solution (Caspian Reed Warbler). Dutch 

Birding 20(1): 36-37. 

Komen, J. 1988. Identity crisis: African Marsh, European Reed and European Marsh Warbler. 

Bokmakierie 40(4): 106-110. 

Korner-Nievergelt, F. et al 2002. How does age and body condition affect migratory restlessness 

and orientation in Reed Warblers Acrocephalus scirpaceus Ardeola 49(1): 29-37. 



Lewington, I. et al 1988. The Welches Dam Acrocephalus Warbler. Birding World 1(5): 180-183. 

Marsh, P. 1982. Notes (Grey-and-white juvenile Reed Warbler). British Birds 75(1): 35. 

Meakin, K. et al 2005. Monitoring birds, reptiles and butterflies in the St Katherine Protectorate, 

Egypt. Egyptian Journal of Biology 7: 66-95 (76 & 83). 

Morgan, J. 1998. Wing formula of Reed Warblers Acrocephalus scirpaceus from Israel – a cautionary 

note. Ringing & Migration 19: 57-58. 

21

O'Sullivan, O. 2004. Discover birds (Warblers - Six summer songsters). Wings 33: 24-26. 

Ozarowska, A. et al 2011. Autumn and spring migration of the Reed Warbler Acrocephalus 

scirpaceus in Egypt – some interesting aspects and questions. Ostrich 82(1): 49-56. 

Parker, J. 1991. Notes (Warblers fleeing from attacks by swallowtail butterfly). British Birds 84(10): 

444. 

Pearson, D.J. 1981. Letters (Identification of first-winter Marsh and Reed Warblers). British Birds 

74(10): 445-446. 

Pearson, D.J. et al 2002. Eurasian Reed Warbler: the characters and variation associated with the 

Asian form fuscus. British Birds 95(2): 42-61. 

Radford, A.P. 1982. Notes (Reed Warbler singing at Magpie and Cuckoo). British Birds 75(8): 383. 

Radford, A.P. 2007. Notes (Reed Warbler apparently using willow-bark pieces in nest construction). 


Raijmakers, J.M.H. & Raijmakers, J.H.F.A. 2002. More records of the European Reed Warbler 

Acrocephalus scirpaceus in south Africa. Afring News 31(1-2): 17-18. 

Rguibi Idrissi, I. et al 2002. Premiers cas de nidification de la Rousserolle effarvatte Acrocephalus 

scirpaceus à Sidi Bou Ghaba (littoral nord-atlantique du Maroc) [Breeding records of the Reed 

Warbler at Sidi Bou Ghaba (Northern Atlantic Morocco)]. Alauda 70(1): 223-225. 

Robertson, D. 2003. Eurasian Reed Warblers in Scotland: a review of probable breeding records. 

Scottish Birds 24 (1): 36-39. 

Rumsey, S.J.R. 1984. Notes (Warblers migrating in family groups). British Birds 77(10): 487. 

Schaub, M. et al 1999. Does tape-luring of migrating Eurasian Reed Warblers increase number of 

recruits or capture probability The Auk 166: 1047-1053. 

Sharrock, J.T.R. 1985. Mystery Photographs (Hybrid Sedge x Reed Warbler). British Birds 78(9): 

434-437. 

Sharrock, J.T.R. 1986. Notes (Reed Warbler singing in oil-seed rape field). British Birds 79(9): 432. 

Shaw, D. et al 2000. The Caspian Reed Warbler on Fair Isle. Birding World 13(8): 315-317. 

Smiddy, P. & O'Mahony, B. 1998. The status of the Reed Warbler Acrocephalus scirpaceus in 

Ireland. Irish Birds 6(1): 23-28. 

Thaxter, C.B. et al 2006. Survival rates of adult Reed Warblers Acrocephalus scirpaceus at a 

northern and southern site in England. Ringing & Migration 23: 65-79. 

Trocinska, A. et al 2001. Field studies of directional preferences of the Reed Warbler (Acrocephalus 

scirpaceus) and the Sedge Warbler (A. schoenobaenus) on autumn migration along the eastern and 

southern coast of the Baltic Sea and in western part of Ukraine. The Ring 23(1-2): 109-117. 

van Eerde, K.A. 1999. Hybride Rietzanger x Kleine Karekiet te Makkum in augustus 1997 [Hybrid 

Sedge x European Reed Warbler at Makkum in August 1997]. Dutch Birding 21(1): 34-37. 


Vinicombe, K. 2001. Letters (The Unst Blyth's Reed Warbler). British Birds 94(6): 291-295. 



Votier, S.C. & Riddington, R. 2005. Notes (Tail pattern of Reed Warblers). British Birds 98(2): 99. 

Voous, K.H. 1975. An aberrant Reed Warbler, or: on the inequality of genera in birds. Ardeola 9: 77- 

85. 


Warden, D. 1998. Notes (Two Reed Warblers laying in same nest). British Birds 91(8): 329. 

Warden, D. 2003. Notes (Reed Warblers reusing nest). British Birds 96(6): 303. 

Warden, D. 2005. Reviews (Reed Warblers at Rostherne Mere by Malcolm Calvert, 2005). British 

Birds 98(8): 437-438. 

Westwood, N.J. 2005. Notes (Reed Warblers reusing nests). British Birds 98(2): 101. 

Yosef, R. & Chernetsov, N. 2005. Longer is fatter: body mass changes of migrant Reed Warblers 

(Acrocephalus scirpaceus) staging at Eilat, Israel. Ostrich 76(3-4): 142-147. 

African Reed Warbler 

Acrocephalus baeticatus [Vieillot 1817, Knysna district, SE Western Cape, South Africa]. 

[A.b. baeticatus] S,C & E South Africa to Zimbabwe & E Botswana. 

[A.b. guiersi] Senegal & perhaps the River Niger delta in Mali. 

[A.b. cinnamomeus] S Senegal, S Mali, E Niger, Nigeria, Cameroon, Gabon, Chad, W & S Sudan E to 

Ethiopia & S Somalia and S to DR Congo, E Zambia, Malawi & Mozambique and possibly this race in 

Libya. 

[A.b. suahelicus] Coastal Tanzania S to Mozambique & KwaZulu-Natal (South Africa). 

[A.b. hallae] SW Angola, E to SW Zambia and S to Namibia, NW Botswana & W South Africa. 

22

Other names: African Marsh Warbler, Rufescent Reed Warbler, Cinnamon Reed Warbler, Tropical 

African Reed Warbler, South African Reed-warbler (baeticatus). 

1st WP Record: May 2008. Western Lakes, (near) Benghazi, Libya. Jens Hering, Stefan Brehme, 

Elmar Fuchs and Hans Winkler. Birds found to be breeding; definate ID by DNA analysis (Hering et al 

2010). 


Bird Study 34: 226-234. 

Amezian, M. et al 2010. Complete moult of an undescribed resident taxon of the Reed Warbler 

Acrocephalus scirpaceus / baeticatus complex in the Smir marshes, Northern Morocco. Ardea 98(2): 

225-234. 

Ash, J.S. et al 1989. The mangrove reed warblers of the Red Sea and Gulf of Aden coasts, with 

description of a new subspecies of the African reed warbler Acrocephalus baeticatus. Bulletin of the 


Colston, P.R. & Morel, G.J. 1984. A new subspecies of the African Reed Warbler Acrocephalus 

baeticatus from Senegal. Bulletin of the British Ornithologists' Club 104(1): 3-5. 

Devillers, P. & Dowsett-Lemaire, F. 1978. African Reed Warblers (Acrocephalus baeticatus) in 

Kaouar (Niger). Le Gerfaut 68: 211-213. 

Dowsett-Lemaire, F. & Dowsett, R.J. 1987. European and African Reed Warblers, Acrocephalus 

scirpaceus and A. baeticatus: vocal and other evidence for a single species. Bulletin of the British 


Eising, C.M. et al 2001. Islands in a desert: breeding ecology of the African Reed Warbler 

Acrocephalus baeticatus in Namibia. Ibis 143(4): 482-493. 

Fry, C.H. et al 1974. A new subspecies of Acrocephalus baeticatus from Lake Chad and a taxonomic 

reappraisal of Acrocephalus dumetorum. Ibis 116(3): 340-346. 

Jiguet, F. et al 2010. Undescribed reed warbler breeding in Morocco. Dutch Birding 32(1): 29-36. 

Hering, J. et al 2009. Zimtrohrsänger Acrocephalus baeticatus und “Mangroverohrsänger“ A. 

scirpaceus avicenniae erstmals in der Paläarktis – Irritierendes aus den Schilfröhrichten Nordafrikas 

[African Reed Warbler Acrocephalus baeticatus and “Mangrove Reed Warbler“ A. scirpaceus 

avicenniae recorded in the Palearctic – confusing discoveries in North African reed beds]. Limicola 

23(3): 202-232. 

Hering, J. et al 2010. African Reed Warblers and Mangrove Reed Warblers in Libya & Egypt - both 

new to the Western Palearctic. Birding World 23(5): 218-219. 

Hering, J. et al 2010. Neues zum Vorkommen und zur Brutbiologie von Zimtrohrsänger Acrocephalus 

baeticatus und Teichrohrsänger A. scirpaceus in Libyen [New information on the occurrence and 

breeding biology of the African Reed Warbler Acrocephalus baeticatus and Eurasian Reed Warbler A. 

scirpaceus in Libya]. Limicola 24(2): 117-139. 


Bokmakierie 40(4): 106-110. 

Wilkinson, R. & Aidley, D.J. 1983. African Reed Warblers in northern Nigeria; morphometrics and 

the taxonomic debate. Bulletin of the British Ornithologists' Club 103: 135-138. 

Marsh Warbler 

Acrocephalus palustris [Bechstein 1798, Thuringia, Germany]. 

C Europe including C European Russia. Winters E & SE Africa. 

Adam, R.G. & Meek, E.R. 1994. Did Marsh Warblers Acrocephalus palustris breed in Orkney in 

1993 Orkney Bird Report 1993 pp. 73-76. 

Cantelo, J. 1984. Notes (Identification of singing Reed and Marsh Warblers by mouth colour). British 

Birds 77(5): 214. 


Dowsett-Lemaire, F. & Dowsett, R.J. 1987. European Reed and Marsh Warblers in Africa: migration 

patterns, moult and habitat. Ostrich 58(2): 65-85. 

Eaton, M.A. et al 2009. Birds of Conservation Concern 3, The population status of birds in the United 

Kingdom, Channel Islands and Isle of Man. British Birds 102(6): 296-341. 


Harrap, S. 1989. Letters (Marsh Warbler Identification). Birding World 2(10): 369. 

Harrap, S. 1989. The difficulties of Reed, Marsh and Blyth's Reed Warbler identification. Birding 

World 2(9): 318-324. 

23

Karakaş, R. 2009. Short Communication (Autumn records of Marsh Warbler in Southeastern 

Anatolia, Turkey). Podoces 4(1): 64-66. 

Kelly, A.G. 1997. Marsh Warbler Acrocephalus palustris singing in County Wexford. Irish Birds 6(1): 

45-47. 

Kelsey, M.G. 1985. Notes (Mouth and gape colours of singing Marsh and Reed Warblers). British 

Birds 78(6): 297. 

Kelsey, M.G. et al 1989. Marsh Warblers in Britain. British Birds 82(6): 239-256. 


Bokmakierie 40(4): 106-110. 

Maher, M. et al 2003. Marsh Warblers breeding at Baltasound: first Shetland breeding record. 

Shetland Bird Report 2002 p. 105. 

Meek, E.R. & Adam, R.G. 1997. Letter (Marsh Warbler breeding in Orkney: first Scottish breeding 

record). British Birds 90(6): 230. 

Mobakken, G. 2000. Letters (Marsh Warbler breeding on Utsira). British Birds 93(6): 279. 

Pearson, D.J. 1981. Letters (Identification of first-winter Marsh and Reed Warblers). British Birds 

74(10): 445-446. 

Smiddy, P. 1992. Marsh Warbler in Co. Cork - a species new to Ireland. Irish Birding News 3(1): 28- 

29. 

Smiddy, P. 1996. Marsh Warbler in County Cork: a species new to Ireland. Irish Birds 5(4): 442-444. 

Stein, H. 2004. Teilalbinismus beim Sumpfrohrsänger Acrocephalus palustris [Partial albinistic Marsh 

Warbler Acrocephalus palustris]. Limicola 18(4): 209-210. 

Syvertsen, Ø. 1992. Letters (Status of Marsh Warbler in Norway). British Birds 85(2): 89. 


Vinicombe, K. 2005. Marsh Warbler. Birdwatch 155: 30-32. 



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