R. Knapp_2014_Index Ferns Fern Allies Taiwan_KBCC

Indexto 

Fernsand 

FernAlies 

ofTaiwan 

RalfKnapp

Indexto 

Fernsand 

FernAlies 

ofTaiwan 

RalfKnapp

Index to Ferns and Fern Allies of Taiwan 

Author: 

Ralf Knapp 

Steigestrasse 78 

69412 Eberbach, Germany 

E-mail: ralf.knapp@gmail.com 

Published by: 

Dr. Cecilia Koo Botanic Conservation Center (KBCC Press) 

No. 31, Tongsing Rd., Gaoshu Township, Pingtung County, Taiwan 

http://www.kbcc.org.tw 

This publication can be cited as: 

Knapp, Ralf. 2014. Index to Ferns and Fern Allies of Taiwan. KBCC Press. 

All rights reserved. It is allowed to print a single copy of the electronic version 

of this document on paper for personal use only. All other cases of 

reproduction or transmission in any form by any means require the prior 

written permission of the copyright owner. 

© Copyright 2014 by KBCC Press. 

All rights reserved. 

Publication date: August 2014. 

ISBN 978-986-87098-3-6 

A PDF version of this book can be downloaded from the KBCC website at 

http://www.kbcc.org.tw.

iii 

F O R E W O R D 

Since the beginning of my botanical studies in Taiwan I have been much 

inspired by the Manual of Taiwan vascular plants Vol. 6 published in the year 

2002, where Yang, Y. P. & Liu, H. Y. provided their list of accepted and 

uncertain taxa, along with synonyms and misapplied names, the respective 

publications, and in some cases as well type information. On numerous 

occasions, the information contained within this volume have proven to be 

very useful when studying local taxa. 

Much has happened since its publication, so I thought it would be 

good to amend it, and also to revise in areas where this has become 

necessary. 

The structure of this index has been chosen in such way that it complements 

my earlier published books. It provides an alphabetically sorted list of genera 

and lower taxa. For each species (and lower level taxon) a range of 

information is provided that allows further intensive study. My other 

publications on the same topic typically do not display data in this detail. 

My books aim to provide tools to allow for the safe identification of taxa in the 

field and in herbaria, with a focus on species and infra-specific levels. 

To display Taiwan's pteridoflora in a practical way I chose a 

taxonomic system that focuses on morphology, classifying taxa based on 

observable structural features. The system proposed in Kramer, K. U. & 

Green, P. S. 1990 appeared best suited for my purposes. However, in recent 

years great advances have been made in understanding evolutionary 

relationships, and I therefore also provide alternative systems based on 

phylogenetic data, and corresponding taxa names where applicable. 

Limitations and advantages of various recent and modern systems are 

discussed in Christenhusz, M. J. M. & Chase, M. W. 2014 and Fraser-Jenkins, 

C. R. 2009. 

In order to cover the local pteridophyte flora as completely as it is possible I 

included undetermined species, and even putative or proven hybrids, even if 

they are still lacking the publication of a name. 

I also looked into the origin of scientific names of genera and lower taxa. In 

almost all cases their roots are Greek or Latin. Often it is clear why an author

iv 

chose this name when publishing the taxon. Occasionally relevant literature 

even contained conflicting information requiring guesswork in some cases. 

Supplementary information is available in chapter 3 Comments where I for 

example point out areas that require further study, or I indicate different 

opinions such as provided in the new Flora of China. 

Moreover, I summarize numerous synonyms and names misapplied to local 

taxa. However, many cases remain where the identity has not been 

satisfactorily resolved. 

Finally, I provide a short summary for taxa that are assumed endemic, 

meaning present in Taiwan only and nowhere else. This publication concludes 

with colour photographs of the most endemic taxa. 

This publication contains an enormous amount of information, raising the 

possibility of errors. In addition, some data is subject to ongoing change. 

Therefore, I would be very grateful of receiving feedback concerning errors, 

inaccurate or missing information, or advances in science relevant to my 

studies and publications. 


Taipei, August 2014

v 

C O N T E N T S 

Foreword 

iii 

1 Introduction 

General remarks 1 

Taxonomic systems 7 

2 List 9 

3 Comments 

Comments regarding selected taxa of the list 163 

Flora of China: additional taxa for Taiwan 166 

Flora of China: name changes for taxa of Taiwan 181 

4 Synonyms and misapplied names 189 

5 References 297 

6 Acknowledgements 301 

7 Appendix A (Taxa endemic to Taiwan) 303 

8 Appendix B (Taxonomic systems) 323

1 

1 I N T R O D U C T I O N 

The present document is closely linked to my publications Ferns and fern 

allies of Taiwan (2011) and Ferns and fern allies of Taiwan - Supplement 

(2013). In only very few cases was it necessary to modify taxon names as 

they were introduced in 2011 and 2013. This consistency allows for 

unencumbered cross-referencing between the three publications. 

The following taxa are very recent additions and not covered in Knapp, R. 

2011 and Knapp, R. 2013: 

• Grammitis nutans (comb. ined., validly published as Prosaptia 

nutans) 

Introduced for Taiwan in FOC. Known at present only from two very 

small areas in SE Taiwan. 

• Huperzia changii 

Only known from three sites in tiny populations in E and SE Taiwan. 

Wild populations might have already disappeared at the time of its 

first description in 2012. Based on its morphology and distribution, 

this taxon might be a hybrid of Huperzia carinata and Huperzia 

salvinioides. 

• Polystichum attenuatum 

Discovered in a small population in the mountains of C Taiwan. 

• Pteris kawabatae 

So far found in half a dozen locations in N and NE Taiwan plus one in 

SC Taiwan in rather individual poor sub-populations. 

General remarks 

For all taxa of pteridophytes in Taiwan recognized presently by me the below 

set of data is provided. It is important to note that not all data categories are 

presented for all taxa simply because the data category may not apply. For 

example, for a yet undetermined species I cannot produce information like 

reference publication or type details as it is simply not yet known. 

Scientific names: 

Each taxon is introduced with its binomial name: the name of the genus to 

which it belongs followed by the name of the species within this genus. If the 

species name is not yet known it is shown as "sp.". Hybrids for which names 

have not been published are shown using a hybrid formula: the binomial 

names of the parent taxa with a multiplication sign "x" placed between them.

2 

Where applicable ranks below the species level are provided. This third part 

of the (trinomial) name is preceded with the rank: "subsp." for subspecies, 

"var." for variety and "f." for form. 

In a few cases where a corresponding species epithet could not be found in 

the respective genus a provisional name is coined and amended by "comb. 

ined." (=combinatio ineditus, not formally published combination). These 

combinations are provided only to facilitate reference to the other publications. 

They are not accepted here and should therefore not be considered valid 

names in the sense of the International Code of Nomenclature for algae, 

fungi, and plants (ICN) articles 33 and 36. In addition a valid name, typically 

that following a recent phylogenetic system (see Appendix B), is given. 

Stress of scientific names: 

In order to support a standardized pronunciation and stressing of scientific 

names of genera, species and infraspecific taxa, vowels (including letter “y”) 

and diphthongs of stressed syllables are underlined in chapters 2 List and 

3 Comments. Following general rule is applied: 

In Latin words of two syllables, the stress is on the first syllable. In words of 

three or more syllables, the stress is on the penultimate syllable if this is 

heavy, otherwise on the antepenultimate syllable. 

Names of authors: 

For the writing of plant name authors reference was made to the 

International Plant Names Index (IPNI, http://www.ipni.org/index.html). Plant 

name authors in chapter 2 List are given in capital letters. 

An asterisk (*) following the scientific and, where applicable, author name 

indicates that additional information is provided in chapter 3 Comments. 

IUCN threat classification: 

Where applicable, threat levels as classified in IUCN Red List categories and 

criteria: version 3.1 (2 nd ed.) (IUCN Species Survival Commission. IUCN, 

Gland, Switzerland. 2012) are provided in square brackets: 

EX Extinct No known individuals remaining 

EW Extinct Wild Known only to survive in cultivation 

CR Critically Endangered Extremely high risk of extinction 

EN Endangered High risk of extinction in the wild 

VU Vulnerable High risk of endangerment in the wild 

NT Near Threatened Likely to become endangered in the 

future 

LC Least Concern Lowest risk, not qualifying for any 

higher category

DD Data Deficient Not enough data to make an 

assessment of its risk of extinction 

(though in many cases category NT 

or even higher is probable) 

Data are derived from A preliminary Red List of Taiwanese vascular plants by 

Wang, J. C. et al. 2012. In cases where I arrive at different classifications I 

provide my opinion in parentheses. Moreover, some taxa were not included in 

the Preliminary Red List of Taiwanese vascular plants partly due to disputed 

taxonomy or being recent new records of obviously very rare and (potentially) 

threatened taxa. In such cases I provide my classification in parentheses with 

a preceeding hyphen (-). Hybrids are not considered. Fortunately, there has 

been, until now, no case of observed extinction (EX) of any pteridophyte in 

Taiwan. 

Reference publication (for an accepted name) in row "Pub.": 

The title of the name giving publication like journal or book is provided with 

its publication year in parentheses. The reference is typically provided in an 

abbreviated form. No information is given for taxa with pending determination, 

or those lacking a valid publication (nom. nud., or not published hybrids). 

Relevant data were often retrieved from the Internet databases Tropicos 

(http://www.tropicos.org/) which is maintained by the Missouri Botanical 

Garden (continuously updated), and Checklist of Ferns and Lycophytes of the 

World, initiated and continuously updated by M. Hassler and B. Schmitt 

(http://worldplants.webarchiv.kit.edu/ferns/). 

Local or vernacular name of a taxon in row "Loc.": 

Primarily, Chinese plant names as listed in the Flora of Taiwan Checklist 

project of the eFloras online database are used (http://www.efloras.org/). For 

the genus Athyrium (s. str.) names as assigned in Fern flora of Taiwan: 

Athyrium (Liu, Y. C. et al. 2009) are adopted. In cases of newly recorded taxa 

any available, including tentatively assigned Chinese names are provided for 

reference. 

The Hanyu pinyin system was used for the romanization of the Chinese plant 

names (more details available e. g. in the English Wikipedia entry "pinyin"). 

For easier readability each Chinese character starts with a capital letter in the 

transcribed form. 

Basionym plus protologue reference in row "Bas.": 

In case the accepted name is not equivalent with the basionym (a previously 

published valid and namebringing synonym from which a new name is formed 

for a taxon of different rank or position, see Terms used in bionomenclature 

by Hawksworth, D. L. 2010), the basionym is provided along with a reference 

of the protologue (in abbreviated form) and publication year in parentheses. 

Also these data were often retrieved from the database Checklist of Ferns and 

3

4 

Lycophytes of the World (M. Hassler and B. Schmitt). 

Origin of taxon names in row "Ety.": 

Etymology analyzes the component parts of words making up of the 

scientific taxon name. Origins in other languages, typically Greek (G) and 

Latin (L), are identified. The following publications were of particular 

importance: Jaeger, E. C. 1955, Stearn, W. 2007, Stewart, R. R. et al. 1983, 

Genaust, H. 1996 and Lellinger, D. B. 2007. In addition, I checked a large 

number of protologues, not all of which provided useful clues however. 

Alternative name based on molecular systems in row "Alt.": 

Christenhusz, M. J. M., Zhang, X. C. & Schneider, H. 2011, Christenhusz, M. 

J. M. & Schneider, H. 2011 and Rothfels, C. J. et al. 2012 propose 

comprehensive taxonomic systems for ferns respective pteridophytes based 

on phylogenetic data. Their concept results in differences for several families 

and also genera, which then affects the species name of my system. In cases 

where the name of the selected molecular system does not differ, this row is 

not provided. 

Type information in row "Type": 

If available, information concerning type material is provided, such as name 

of collector(s), collection number, collection date, location, country, herbarium 

(shown as acronym) where the type specimen(s) are stored along with 

identification of the kind of type. In many cases information was retrieved from 

the following sources: 

• The Manual of Taiwan vascular plants Vol. 6 (MAN, see hereafter) 

• Internet database Tropicos (http://www.tropicos.org/) 

• Web site Ferns of Thailand, Laos and Cambodia (http://rbgweb2.rbge.org.uk/thaiferns/) 

which is managed by S. Lindsay and D. 

Middleton of the Royal Botanic Garden Edinburgh (continuously 

updated) 

First publication of a taxon (name) for Taiwan in row "TW": 

Reference to that publication where a taxon was presumably first published 

under this name or a combination of it with occurance in Taiwan with the year 

of publication shown in parentheses. Here, any combination of the name is 

recognized, including the basionym. In many cases the very taxa have been 

published earlier for Taiwan, but under a different epithet. Such information is 

not reflected in the list. 

The following is a chronological overview of floras and lists for Taiwan that 

include pteridophytes (where relevant with the total number of ferns and allied 

taxa recognized in parentheses), details of publications can be found in 

chapter 5 References:

5 

1863 Swinhoe, R. Lists of plants from the island of 

Formosa, or Taiwan 

1885 Baker, J. G. Ferns collected in North 

Formosa by Mr. William 

Hancock 

(33 taxa, 

only N 

Taiwan) 

(92 taxa, 

only N 


1896 Henry, A. A list of plants from Formosa (149 taxa) 

1902 Yabe, Y. A note of ferns from the island of 

Koto (Botel-Tobago) 

1906 Matsumara, 

J. & 

Hayata, B. 

Enumeratio Plantarum: in insula 

formosa sponte crescentium 

hucusque rite cognitarum 

adjectis descriptionibus et 

figuris specierum pro regione 

novarum 

1908 Hayata, B. Flora montana formosae 

1909 Hayata, B. Some ferns from the mountain 

regions of Formosa 

1910 Kawakami, 

T. 

A list of plants of Formosa 

1911 Hayata, B. Materials for a flora of Formosa 

1914-21 Hayata, B. Iconum plantarum 

formosanarum 4-8 & 10 

1917 Hayata, B. General index to the flora of 

Formosa, supplement to 

icones plantarum 

formosanarum 6 (1916) 

(272 taxa) 

(309 taxa) 

(527 taxa) 

1928 Ito, T. Illustrations of Taiwan ferns 

1928-40 Ogata, M. Icones filicum japoniae 1-8 

1928 Sasaki, S. Lists of plants of Formosa (550 taxa) 

1928 Yamamoto, 

Y. 

Supplementa iconum plantarum 

formosanarum 4 

1932 Yamamoto, 

Y. 

Supplementa iconum plantarum 

formosanarum 5 

1936 Masamune, A short flora of Formosa 

(598 taxa) 

G. 

1940-49 Tagawa, M. Studies on Formosan ferns 1-7 

1975 Li, H. L. et al. Flora of Taiwan (1 st ed.) Vol. 1 (598 taxa) 

1976 Liew, F. S. A list of ferns and fern allies 

found in Orchid Island, Taiwan 

1981 Ito, H. Icones filicum japoniae 9 

1985 Kuo, C. M. Taxonomy and phytogeography 

of Taiwanese pteridophytes 

(617 taxa) 

1994 Huang, T. C. Flora of Taiwan (2 nd ed.) Vol. 1 (597 taxa)

6 

1994 Huang, T. C. 

et al. 

The flora of Taipingtao (Aba Itu 

Island) (part 1) 

1997 Kuo, C. M. Manual of Taiwan vascular (650 taxa) 

plants Vol. 1 

2002 Yang, Y. P. & Manual of Taiwan vascular (678 taxa) 

Liu, H. Y. plants Vol. 6 

2003 Huang, T. C. Flora of Taiwan (2 nd ed.) Vol. 6 (653 taxa) 

2005 Lu, S. G. & The checklist of Taiwanese (677 taxa) 

Yang, T. Y. 

A. 

pteridophytes following 

Ching’s system 

2011 Knapp, R. Ferns and fern allies of Taiwan (729 taxa) 

2011 Lu, F. Y. Flora of Kinmen 

2013 Knapp, R. Ferns and fern allies of Taiwan - (747 taxa) 

Supplement 

2013 Wu, Z. Y. et Flora of China, Vol. 2-3 

(710 taxa) 

al. 

(Pteridophytes) 

2014 Knapp, R. Index to ferns and fern allies 

of Taiwan (present document) 

(751 taxa) 

List of relevant abbreviations: 

(details of publications can be found in chapter 5 References) 

Alt. 

Alternative taxon name when classified according to 

molecular systems (used in chapter 2 List) 

Bas. Basionym plus protologue reference (used in chapter 2 

List) 

C, E, N, S, 

W 

CHK Lu, S. G. & Yang, T. Y. A. 2005 

comb. ined. 

Directions E=East, N=North, S=South, W=West plus 

C=Central (also used in combinations) 

Combinatio ineditus, used for taxa where corresponding 

species epithet could not be found; a temporary 

scientific name is coined, followed by the closest 

relevant validly published scientific name 

Origin of taxon name (etymology) (used in chapter 2 List) 

Ety. 

f. Forma, form (taxonomic rank below species) 

FOC Wu, Z. Y. et al. 2013 

FOJ Iwatsuki, K. et al. 1995 

FOT 1975 Li, H. L. et al. 1975 

FOT 1994 Huang, T. C. 1994 

FOT 2003 Huang, T. C. 2003 

G 

Greek (as used in chapter 2 List) 

HT Holotype (see Hawksworth, D. L. 2010) 

ICN 

International Code of Nomenclature for algae, fungi, and 

plants (see McNeill, J. et al. 2012) 

ILT Isolectotype (see Hawksworth, D. L. 2010)

INT Isoneotype (see Hawksworth, D. L. 2010) 

IST Isosyntype (see Hawksworth, D. L. 2010) 

IT Isotype (see Hawksworth, D. L. 2010) 

IUCN Threat classification according to criteria outlined by the 

International Union for Conservation of Nature 

L 

Latin (as used in chapter 2 List) 

Loc. Local or vernacular name of a taxon (used in chapter 2 

List) 

LT Lectotype (see Hawksworth, D. L. 2010) 

MAN Yang, Y. P. & Liu, H. Y. 2002 

MT Merotype (see Hawksworth, D. L. 2010) 

NT Neotype (see Hawksworth, D. L. 2010) 

nom. cons. Nomen conservandum, conserved name (see Hawksworth, 

D. L. 2010) 

nom. nov. Nomen novum, new name (see Hawksworth, D. L. 2010) 

nom. nud. Nomen nudum, used for scientific names that have not 

Pub. 

yet been validly published 

Reference publication for an accepted name (used in 

chapter 2 List) 

s. l. Sensu lato, in a broad sense 

s. n. Sine numero, used after a collector’s name to indicate the 

absence of a collector’s number 

sp. 

Species, used for taxa where the relevant scientific name 

is not known or not yet published 

s. str. Sensu stricto, in a strict sense (of the circumscription of a 

taxon) 

ST Syntype (see Hawksworth, D. L. 2010) 

subsp. 

TW 

var. 

Subspecies (taxonomic rank below species) 

First publication of a taxon (name) for Taiwan (used in 

chapter 2 List) 

Varietas, variety (taxonomic rank below species) 

Herbarium acronyms follow the Index Herbariorum: 

http://sweetgum.nybg.org/ih/, continuously updated. 

7 

Taxonomic systems 

The system of Kramer and Green (1990) 

The classification of pteridophytes by Kramer, K. U. & Green, P. S. 1990 in 

Kubitzki's The families and genera of vascular plants recognizes four classes: 

• Fern allies: Psilotatae, Lycopodiatae, Equisetatae 

• Real ferns: Filicatae

8 

Their system is mainly the result of morphological analysis and makes it thus 

most useful for classifying taxa during field and herbarium work. The system 

is applied, without alteration, for this document. For details see Appendix B. 

The system of Christenhusz et al. (2011) 

In contrast, the system proposed by Christenhusz, M. J. M., Zhang, X. C. & 

Schneider, H. 2011, Christenhusz, M. J. M. & Schneider, H. 2011 and 

Rothfels, C. J. et al. 2012 based on phylogenetic results contains: 

• Lycophytes 

• Monilophytes (ferns) 

For details see Appendix B. Consequent alternative combinations are shown 

in chapter 2 List. 

The system of Christenhusz and Chase (2014) 

A recently refined system is proposed in Christenhusz, M. J. M. & Chase, M. 

W. 2014. It covers: 

• Lycopodiophyta (lycopods) 

• Polypodiophyta (ferns) 

For details see Appendix B. 

Several changes result in the lumping and also splitting of genera, which 

require then scientific names in different combinations for affected species. 

However, it is obvious that still many adjustments in the system are necessary 

and many taxa require further research. Therefore, I do not reflect at this point 

consequences of this system in chapter 2 List.

9 

2 L I S T 

ACROPHORUS (Dryopteridaceae) 

Ety.: akros (G: highest), phoreo (G: to bear); relating to the position of sori 

at the apex of veins 

(1) Acrophorus macrocarpus CHING & S. H. WU 

Pub.: Acta Phytotax. Sin. 21(4): 381-382, pl. 2, f. 1 (1983) 

Loc.: 大孢孢魚鱗蕨蕨 - DaBaoYuLinJue 

Ety.: makros (G: large), karpos (G: fruit); relating to the large-sized 

sorus 

Type: Yunnan, Deqen, Cizhong, K. M. Feng 5512 (HT: PE) // China 

TW: Kuo, C. M. 1997 (1997) 

(2) Acrophorus paleolatus PIC. SERM. 

Pub.: Webbia 31(1): 252 (1977) 

Loc.: 魚鱗蕨蕨 - YuLinJue 

Ety.: palea (L: scale) and latus (L: bearing, carrying); relating to the 

scales of the lamina (see protologue) 

Type: Darjiling, Birch Hill, H. C. Levinge 99 (HT: FI) // India 

TW: Kuo, C. M. 1985 (1985) 

ACROSTICHUM (Pteridaceae) 

Ety.: akros (G: highest), stichos (G: order); relating to the position of fertile 

pinnae at the frond apex 

(3) Acrostichum aureum L. 

[IUCN: VU] 

Pub.: Sp. Pl. 2: 1069 (1753) 

Loc.: 鹵蕨蕨 - LuJue 

Ety.: aureum (L: golden, splendid); relating to the young sori that 

colour the adaxial pinna brownish-yellow 

Type: (LT: LINN), designated by Proctor, Fl. Lesser Antilles 2: 152 

(1977) 

TW: Swinhoe, R. (1863) 

ACYSTOPTERIS (Dryopteridaceae) 

Ety.: a (G: not), kystos (G: cyst), pteris (relates to genus Pteris, or ferns in 

general); relating to the similarity to members of fern genus 

Cystopteris (see there) 

(4) Acystopteris taiwaniana (TAGAWA) A. LÖVE & D. LÖVE 

Pub.: Taxon 26(2-3): 326 (1977) 

Loc.: 台台灣亮毛蕨蕨 - TaiWanLiangMaoJue 

Bas.: Cystopteris japonica var. taiwaniana TAGAWA // Acta Phytotax. 

Geobot. 4(2): 57 (1935) 

Ety.: relating to Taiwan, the type location of this taxon 

Type: Arisan, Tagawa 526 (HT: KYO) // Taiwan 

TW: Acta Phytotax. Geobot. 4(2): 57 (1935)

10 

(5) Acystopteris tenuisecta (BLUME) TAGAWA 

Pub.: Acta Phytotax. Geobot. 7: 73 (1938) 

Loc.: 禾稈亮毛蕨蕨 - HeGanLiangMaoJue 

Bas.: Aspidium tenuisectum BLUME // Enum. Pl. Javae 170 (1828) 

Ety.: tenuisecta (L: finely divided); relating to the lamina habit 

Type: Java // Indonesia 

TW: Ito, H. 1944 (1944) 

ADIANTUM (Pteridaceae) 

Ety.: adiantos (G: unwettable); relating to the observation when fronds are 

plunged in water they are coming out dry 

(6) Adiantum capillus-junonis RUPR. [IUCN: CR (EN)] 

Pub.: Beitr. Pflanzenk. Russ. Reiches 3: 49 (1845) 

Loc.: 團羽鐵線蕨蕨 - TuanYuTieXianJue 

Ety.: capillus (L: hair), junonis (L: of Juno, Roman Goddess); 

relating to the axes, which resemble dark women hairs 

Type: China, Hebei, Peking, Bunge s. n. // China 

TW: Henry, A. (1896) 

(7) Adiantum capillus-veneris L. (f. capillus-veneris) 

Pub.: Sp. Pl. 2: 1096 (1753) 

Loc.: 鐵線蕨蕨 - TieXianJue 

Ety.: capillus (L: hair), veneris (L: of Venus, Roman Goddess); 

relating to the axes, which resemble dark women hairs 

Type: South Europe, France (without data), Magnol s. n. (LT: LINN), 

lectotypified by Pichi-Sermolli, Webbia 12: 678 (1957) // 

France 


(8) Adiantum capillus-veneris f. lanyuanum W. C. SHIEH [IUCN: - (VU)] 

Pub.: Quart. J. Chin. Forest. 2(1): 165 (1968) 

Loc.: 蘭蘭嶼鐵線蕨蕨 - LanYuTieXianJue 

Ety.: relating to LanYu, an islet of SE Taiwan and the type location 

Type: Lanyu Isl., Shieh no. 388, Oct-1967 (HT: TCB) // Taiwan 

TW: Quart. J. Chin. Forest. 2(1): 165 (1968) 

(9) Adiantum caudatum L. 

Pub.: Mant. Pl. 308 (1771) 

Loc.: 鞭葉鐵線蕨蕨 - BianYeTieXianJue 

Ety.: caudatum (L: having a tail); relating to the frequently prolonged 

axis 

Type: Burman // Sri Lanka 

TW: Henry, A. (1896) 

(10) Adiantum diaphanum BLUME 

Pub.: Enum. Pl. Javae 215 (1828) 

Loc.: 長尾尾鐵線蕨蕨 - ChangWeiTieXianJue 

Ety.: dia (G: much), phanos (G: light); relating to the thin lamina 

texture which is quite transparent

Type: Java (L) // Indonesia 

TW: J. Bot. 23: 103 (1885) 

(11) Adiantum edgeworthii HOOK. 

Pub.: Sp. Fil. 2: 14-15, pl. 81B (1851) 

Loc.: 愛氏鐵線蕨蕨 - AiShiTieXianJue 

Ety.: in honor of M. P. Edgeworth (Irish botanist, and collector of 

type specimen) 

Type: Adah Valley in northwest, 1600 m, M. P. Edgeworth s. n., Sep- 

1838 (?K, ?OXF) // India 

TW: FOT 1975 (1975) 

(12) Adiantum flabellulatum L. 

Pub.: Sp. Pl. 2: 1095 (1753) 

Loc.: 扇葉鐵線蕨蕨 - ShanYeTieXianJue 

Ety.: diminutive of flabellum (L: a small fan), latum (L: suffix, added 

to nouns to form adjectives meaning "provided with"); relating 

to the lamina shape 

Type: Osbeck s. n. (LT: LINN) // China 

TW: J. Bot. 23: 103 (1885) 

(13) Adiantum formosanum TAGAWA [IUCN: VU] 

Pub.: J. Jap. Bot. 14: 315-316 (1938) 

Loc.: 深山鐵線蕨蕨 - ShenShanTieXianJue 

Ety.: relating to Formosa (=Taiwan), the type location of this taxon 

Type: Noko-goe, Ohwi 3268 (HT: KYO) // Taiwan 

TW: J. Jap. Bot. 14: 315-316 (1938) 

(14) Adiantum hispidulum SW. 

Pub.: J. Bot. (Schrader) 1800(2): 82 (1801) 

Loc.: 毛葉鐵線蕨蕨 - MaoYeTieXianJue 

Ety.: hispidus (L: spiny, rough); relating to the stiff-hairy lamina 

Type: (HT: S) // Australia 

TW: Henry, A. (1896) 

(15) Adiantum malesianum J. GHATAK 

Pub.: Bull. Bot. Surv. India 5: 73, f. 4, 6, 7 (1963) 

Loc.: 馬來鐵線蕨蕨 - MaLaiTieXianJue 

Ety.: relating to Malesia, the phytogeographical region, rather than 

to Malaysia, the political entity; of SE Asian distribution 

Type: China 

TW: Kuo, C. M. 1985 (1985) 

(16) Adiantum meishanianum F. S. HSU ex Y. C. LIU & W. L. CHIOU [IUCN: CR] 

Pub.: Novon 19(1): 59-61 (2009) 

Loc.: 梅山鐵線蕨蕨 - MeiShanTieXianJue 

Ety.: relating to MeiShan, a village in SC Taiwan, the type (and 

only) location of this taxon 

Type: Kaohsiung, Meishankou, B. J. Wang 3025, 30-Oct-1983 (HT: 

HAST) // Taiwan 

TW: Novon 19(1): 59-61 (2009) 

11

12 

(17) Adiantum monochlamys D. C. EATON [IUCN: EN (VU)] 

Pub.: Proc. Amer. Acad. Arts 4: 110 (1858) 

Loc.: 石長生 - ShiChangSheng 

Ety.: monos (G: one-), chlamys (G: mantle); relating to the singular 

soral flap 

Type: Simoda, Wright s. n. (US) // Japan 

TW: Masamune, G. (1936) 

(18) Adiantum myriosorum BAKER [IUCN: VU] 

Pub.: Bull. Misc. Inform. Kew 1898: 230 (1898) 

Loc.: 灰背鐵線蕨蕨 - HuiBeiTieXianJue 

Ety.: myrio (G: countless), soros (G: sorus); relating to the large 

number of sori of a fertile frond 

Type: Yunnan, 2000 m, Henry 9266 (HT: K; IT: NY, US) // China 

TW: FOT 1975 (1975) 

(19) Adiantum philippense L. 

Pub.: Sp. Pl. 2: 1094 (1753) 

Loc.: 半月鐵線蕨蕨 - BanYueTieXianJue 

Ety.: filipino (Spanish: found in the Philippines); relating to the type 

location of this taxon 

Type: (LT: Petiver, Gazophyllacii Naturae et Artis decas prima t. 4, f. 

4. 1702), designated by Pichi-Sermolli, Webbia 12 (1957) // 

Philippines 

TW: Sasaki, S. (1928) 

(20) Adiantum soboliferum WALL. ex HOOK. [IUCN: VU] 

Pub.: Numer. List n. 74 (1828) ex Hooker, Sp. Fil. 2: 13, pl. 74 A 

(1851) 

Loc.: 翅柄鐵線蕨蕨 - ChiBingTieXianJue 

Ety.: soboles (L: root- or stem-like structure), -fer (L: carrying); 

relating to the development of plantlets at the tip of frond 

Type: Wallich 74 // India 

TW: Ito, H. 1944 (1944) 

(21) Adiantum taiwanianum TAGAWA [IUCN: VU (NT)] 

Pub.: Acta Phytotax. Geobot. 4: 92-93 (1935) 

Loc.: 台台灣鐵線蕨蕨 - TaiWanTieXianJue 


Type: Kwarenko, inter Sekigohara & Gokwan, 3000 m, Tagawa 820 

(HT: KYO) // Taiwan 

TW: Acta Phytotax. Geobot. 4: 92-93 (1935) 

(Polypodiaceae) 

aglaios (G: splendid), morphe (G: shape); relating to morphological 

observation ("splendid form") of author, with reference to one of 

the Graces of Greek mythology named Aglaia 

AGLAOMORPHA 

Ety.: 

(22) Aglaomorpha coronans (WALL. ex METT.) COPEL. 

Pub.: Univ. Calif. Publ. Bot. 16: 117 (1929)

Loc.: 崖薑薑蕨蕨 - YaiJiangJue 

Bas.: Polypodium coronans WALL. // Numer. List n. 288 (1828), ex 

Mettenius, Abh. Senckenberg. Naturf. Ges. 1: 121, pl. 3, f. 

40-41 (1856) 

Ety.: coronans (L: crowning); relating to its epiphytic growth habit 

Type: Wallich 288 (HT: B; IT: BM, BR, G, GH, K, L, P, US, W) // 

Nepal 


(23) Aglaomorpha meyeniana SCHOTT [IUCN: NT] 

Pub.: Gen. Fil. 4: pl. 19 (1836) 

Loc.: 連珠蕨蕨 - LianZhuJue 

Ety.: in honor of F. J. F. Meyen (German botanist) 

Type: Luzon, Cuming 49 (B, BM, K, L) // Philippines 

TW: Henry, A. (1896) 

ANGIOPTERIS (Marattiaceae) 

Ety.: angeion (G: case, capsule), pteris (relates to genus Pteris, or ferns 

in general); relating to the incipient synangia 

(24) Angiopteris itoi (W. C. SHIEH) J. M. CAMUS 

Pub.: Proc. Int. Symp. Syst. Pterid. (1988): 35 (1989) 

Loc.: 伊藤藤氏原原始觀音座蓮 - YiTengShiYuanShiGuanYinZuoLian 

Bas.: Archangiopteris itoi W. C. SHIEH // J. Jap. Bot. 45: 165, f. 2-3 

(1970) 

Ety.: in honor of H. Ito (Japanese botanist) 

Type: Nantou, Lienhwachi, Shieh 1128 (HT: TAIF (transferred from 

TCB) // Taiwan 

TW: J. Jap. Bot. 45: 165, f. 2-3 (1970) 

(25) Angiopteris lygodiifolia ROSENST. 

Pub.: Meded. Rijks-Herb. 31: 2 (1917) 

Loc.: 觀音座蓮 - GuanYinZuoLian 

Ety.: lygodium relating to a fern genus (see there), folium (L: leaf); 

relating to the shape of lamina 

Type: ?Bürge s. n. // Japan 

TW: FOT 1975 (1975) 

(26) Angiopteris palmiformis (CAV.) C. CHR. [IUCN: - (NT)] 

Pub.: Dansk Bot. Ark. 9(3): 30, pl. 3 (7-8) (1937) 

Loc.: 蘭蘭嶼觀音座蓮 - LanYuGuanYinZuoLian 

Bas.: Clementea palmiformis CAV. // Descr. Pl. 554 (1802) 

Ety.: palma (L: hand), forma (L: shape, appearance); relating to the 

lamina shape 

Type: L. Nee s. n., 1801 (BM, MA) // Philippines 

TW: FOT 1975 (1975) 

(27) Angiopteris somae (HAYATA) MAKINO & NEMOTO [IUCN: EN (VU)] 

Pub.: Fl. Japan 1563 (1925) 

Loc.: 台台灣原原始觀音座蓮 - TaiWanYuanShiGuanYinZuoLian 

13

14 

Bas.: Archangiopteris somae HAYATA // Icon. Pl. Formosan. 5: 256 

(1915) 

Ety.: relating to T. Soma (Japanese botanist, and collector of the 


Type: Rahao, T. Soma s. n., Dec-1910 (HT: TI) // Taiwan 

TW: Icon. Pl. Formosan. 5: 256 (1915) 

ANOGRAMMA (Pteridaceae) 

Ety.: ano (G: upward), gramme (G: line); relating to the elongated sori that 

are on terminal segments 

(28) Anogramma leptophylla (L.) LINK [IUCN: NT] 

Pub.: Fil. Sp. 137 (1841) 

Loc.: 翠蕨蕨 - CuiJue 

Bas.: Polypodium leptophyllum L. // Sp. Pl. 2: 1092 (1753) 

Ety.: leptos (G: thin); phyllon (G: leaf); relating to the thin lamina 

Type: Lusitania, Galloprovincia (LT: LINN), designated by Morton, 

Amer. Fern J. 60: 101 (1970) // Spain (or Portugal) 

TW: FOT 1975 (1975) 

ANTROPHYUM (Vittariaceae) 

Ety.: antron (G: a cave), phyein (G: to grow); relating to the habitat 

requirements ("cave dweller") 

(29) Antrophyum alatum BRACK. [IUCN: - (CR)] 

Pub.: U. S. Expl. Exped., Filic. 16: 64 (1854) 

Loc.: 有翅車前蕨蕨 - YouChiCheQianJue 

Ety.: alatum (L: winged); relating to the attenuated lamina base (see 

protologue) 

Type: Herb US South Pacific Exped., Wilkes et al. s. n., 1838-42 

(HT: B) // Fiji 

TW: Knapp, R. 2011 (2011) 

(30) Antrophyum castaneum H. ITO [IUCN: - (NT)] 

Pub.: J. Jap. Bot. 12: 473, pl. 7, f. 1 (1936) 

Loc.: 阿里山車前蕨蕨 - ALiShanCheQianJue 

Ety.: castaneum (L: of chestnut colour); probably relating to the 

colour of soral lines 

Type: Nimandaira mont. Arisan, Ito & Hayata s. n. (HT: TI) // Taiwan 

TW: J. Jap. Bot. 12: 473, pl. 7, f. 1 (1936) 

(31) Antrophyum formosanum HIERON. 

Pub.: Hedwigia 67: 210-212 (1915) 

Loc.: 台台灣車前蕨蕨 - TaiWanCheQianJue 


Type: Kushaku, Faurie 675 (LT: B; ILT: KYO), annotated by C. M. 

Kuo on 27-Sep-1983 // Taiwan 

TW: Hedwigia 67: 210-212 (1915)

(32) Antrophyum henryi HIERON. [IUCN: - (EN)] 

Pub.: Hedwigia 67: 208-210 (1915) 

Loc.: 亨利氏車前蕨蕨 - HengLiShiCheQianJue 

Ety.: in honor of A. Henry (Irish botanist, collector of type specimen) 

Type: Yunnan, 4500 ft., Henry 11517A (HT: P; IT: E, MO) // China 


(33) Antrophyum obovatum BAKER 

Pub.: Bull. Misc. Inform. Kew 1898: 233 (1898) 

Loc.: 長柄車前蕨蕨 - ChangBingCheQianJue 

Ety.: ob (L: over), ovatus (L: egg-shaped); relating to the obovate 

form of lamina 

Type: Yunnan, Henry 9153A (HT: K; IT: ?B) // China 

TW: Ito, H. 1944 (1944) 

(34) Antrophyum parvulum BLUME [IUCN: VU] 


Loc.: 無柄車前蕨蕨 - WuBingCheQianJu 

Ety.: parvulum (L: small); relating to the lamina size 


TW: FOT 1975 (1975) 

(35) Antrophyum sessilifolium (CAV.) SPRING [IUCN: NT (VU)] 

Pub.: Syst. Veg. 4: 67 (1827) 

Loc.: 蘭蘭嶼車前蕨蕨 - LanYuCheQianJue 

Bas.: Hemionitis sessilifolia CAV. // Descr. Pl. 261 (1802) 

Ety.: sessilis (L: of sitting), folium (L: leaf); relating to the short stipe 

Type: L. Nee s. n. (HT: MA) // Philippines 

TW: FOT 1975 (1975) 

ARACHNIODES (Dryopteridaceae) 

Ety.: arachnion (G: spider's web), odes (G: having the form of); perhaps 

relating to the resemblance of one of the more finely dissected 

pentagonal species to a spider's web 

(36) Arachniodes aristata (G. FORST.) TINDALE 

Pub.: Contr. New South Wales Natl. Herb. 3: 89 (1961) 

Loc.: 細葉複葉耳蕨蕨 - XiYeFuYeErJu 

Bas.: Polypodium aristatum G. FORST. // Fl. Ins. Austr. 82 (1786) 

Ety.: aristata (L: awned, bearded); relating to the presence of stiff, 

bristle-like points at apex of pinnules 

Type: a: Otahiete, G. Forsters Herbarium 294 (448) (LT: BM, 

selected by Morton) // French Polynesia 

b: without locality and collector information (LT: S, selected by 

Tindale (1961)) 

TW: Matsumara, J. & Hayata, B. (1906) 

(37) Arachniodes caudata CHING [IUCN: - (CR)] 

Pub.: Acta Phytotax. Sin. 9: 384 (1964) 

Loc.: 尾尾形複葉耳蕨蕨 - WeiXingFuYeErJue 

15

16 

Ety.: caudata (L: having a tail); relating to the presence of long and 

narrow projections at apex of pinnules 

Type: Honsyu, Hidue prope Singu, Prov. Kii, Tasiro-Z. s. n., 22-Mar- 

1931 (HT: KYO), type of Polystichum simplicius var. majus 

Tagawa in Acta Phytotax. 1(1): 90-91 (1932), selected by 

Ching in his publication for "nom. et stat. nov." // Japan 


(38) Arachniodes chinensis (ROSENST.) CHING [IUCN: - (VU)] 

Pub.: Acta Bot. Sin. 10: 257 (1962) 

Loc.: 中華複葉耳蕨蕨 - ZhongHuaFuYeErJu 

Bas.: Polystichum amabile var. chinense ROSENST. // Repert. Spec. 

Nov. Regni Veg. 13: 130 (1914) 

Ety.: relating to China, the type location 

Type: Kuy-tcheu, Gan-chuen, P. Cavalerie 4010, 1912 (IT: BM) // 

China 

TW: TW J. of Biodivers. 14(1-2): 75-80 (2012) 

(39) Arachniodes festina (HANCE) CHING 


Loc.: 台台灣兩面複葉耳蕨蕨 - TaiWanLiangMianFuYeErJue 

Bas.: Aspidium festinum HANCE // J. Bot. 21: 269 (1883) 

Ety.: two alternatives: (1) festina (L: swift, quick, rapid); relationship 

unclear to me, perhaps relating to the nature of development 

of young fronds; (2) festa (L: used for festivals, ornamental): 

perhaps relating to the decorative habit 

Type: Guangdong, Faber s. n., Sep-1882 (HT: BM, Herb. Hance 

22146) // China 

TW: Ito, H. 1944 (1944) 

(40) Arachniodes globisora (HAYATA) CHING [IUCN: NT] 


Loc.: 台台灣複葉耳蕨蕨 - TaiWanFuYeErJue 

Bas.: Polystichum globisorum HAYATA // Icon. Pl. Formosan. 4: 193- 

195, f. 131 (1914) 

Ety.: globus (L: ball), sorus (L: sorus); relating to the shape of sori 

Type: Arisan, ad 7000-8000 ped. alt., B. Hayata & S. Sasaki s. n., 

Jan-1912 (HT: TI) // Taiwan 

TW: Icon. Pl. Formosan. 4: 193-195, f. 131 (1914) 

(41) Arachniodes nigrospinosa (CHING) CHING [IUCN: NT (VU)] 


Loc.: 黑鱗複葉耳蕨蕨 - HeiLinFuYeErJue 

Bas.: Polystichum nigrospinosum CHING // Bull. Fan Mem. Inst. Biol. 

2: 191-192, pl. 6 (1931) 

Ety.: niger (L: black), spinosa (L: spiny); relating to the black scales 

on axes 

Type: Guangdong, Tai-mo Shan, opposite to Hongkong, Matthew s. 

n. (HT: PE) // China

TW: FOT 1975 (1975) 

(42) Arachniodes pseudoaristata (TAGAWA) OHWI 

Pub.: J. Jap. Bot. 37: 76 (1962) 

Loc.: 小小葉複葉耳蕨蕨 - XiaoYeFuYeErJue 

Bas.: Polystichum pseudoaristatum TAGAWA // Acta Phytotax. 

Geobot. 1: 91 (1932) 

Ety.: pseudos (G: false), aristata relating to the taxon with epithet 

"aristata"; relating to the similarity to taxon Arachniodes 

aristata (see there) 

Type: Tasiro-Z. s. n. (HT: KYO; IT: UC) // Japan 


(43) Arachniodes quadripinnata (HAYATA) SERIZ. 

Pub.: J. Jap. Bot. 61(2): 53 (1986) 

Loc.: 毛孢擬複葉耳蕨 - MaoBaoNiFuYeErJue 

Bas.: Microlepia quadripinnata HAYATA // J. Coll. Sci. Imp. Univ. 

Tokyo 30: 434-435 (1911) 

Ety.: quadra (L: fourfold), pinnata (L: pinnate); relating to the lamina 

Type: 

shape (being four-times pinnate) 

Nanto, Mushazan, Kawakami & Mori 2390, Aug-1906 (HT: TI) 

// Taiwan 

TW: J. Coll. Sci. Imp. Univ. Tokyo 30: 434-435 (1911) 

(44) Arachniodes rhomboidea (SCHOTT) CHING 

Pub.: Acta Phytotax. Sin. 9: 383-384 (1964) 

Loc.: 斜方複葉耳蕨蕨 - XieFangFuYeErJue 

Bas.: Polystichum rhomboideum SCHOTT // Gen. Fil. ad., pl. 9 (1834) 

Ety.: rhomboidea (L: shape of a rhombus); relating to the shape of 

pinnules 

Type: Wallich 364, 1821 (IT: US) // Nepal 

TW: a: Matsumara, J. & Hayata, B. (1906) as Aspidium 

rhomboideum WALL., synonym of Polystichum amabile SM. 

b: FOT 1975 (1975) 

(45) Arachniodes yakusimensis (H. ITO) T. NAKAIKE 

Pub.: New Fl. Japan, Pterid. 839 (1992) 

Loc.: 屋屋久複葉耳蕨蕨 - WuJiuFuYeErJue 

Bas.: Rumohra amabilis var. yakusimensis H. ITO // J. Jap. Bot. 11: 

574-575, pl. 7, f. 14; f. 3 (1935) 

Ety.: relating to Yakushima Island (Japan, type location) 

Type: Kyusyui, Prov. Oosumi, Yakusima, T. Makino s. n., Jul-1909 

(HT: TI) // Japan 


ARAIOSTEGIA (Davalliaceae) 

Ety.: araios (G: thin), stege (G: roof, covering); relating to the thin 

indusium 

17

18 

(46) Araiostegia parvipinnula (HAYATA) COPEL. 

Pub.: Philipp. J. Sci. 34(3): 241 (1927) 

Loc.: 鳞鳞轴小小膜盖蕨蕨 - LinZhouXiaoMoGaiJue 

Bas.: Davallia parvipinnula HAYATA // J. Coll. Sci. Imp. Univ. Tokyo 

30: 431-432 (1911) 

Ety.: parva (L: small), pinnula (L: little feather); relating to the small 

size of ultimate lamina dissection level 

Alt.: Davallodes parvipinnula (comb. ined.) 

Type: Mt. Morrison, Kawakami & Mori 1823, 21-Oct-1906 (HT: TI) // 

Taiwan 


ARTHROMERIS (Polypodiaceae) 

Ety.: arthron (G: joint), meris (G: part); relating to the pinnae being 

articulate with rachis 

(47) Arthromeris lehmannii (METT.) CHING 

Pub.: Contr. Inst. Bot. Natl. Acad. Peiping 2: 96-97 (1933) 

Loc.: 肢節蕨蕨 - ZhiJieJue 

Bas.: Polypodium lehmannii METT. // Abh. Senckenberg. Naturf. 

Ges. 1: 117, pl. 3, f. 35 (1856) 

Ety.: in honor of J. G. C. Lehmann (German botanist), collector (or 

at least the sender) of the type specimen 

Type: sent by Lehmann to Mettenius // ?India 

TW: Hayata, B. 1909 (1909) 

ARTHROPTERIS (Oleandraceae) 

Ety.: arthron (G: joint), pteris (relates to genus Pteris, or ferns in general); 

relating to the frond and pinnae being articulate 

(48) Arthropteris palisotii (DESV.) ALSTON 

Pub.: Bol. Soc. Brot., ser. 2, 30: 6 (1956) 

Loc.: 藤藤蕨蕨 - TengJue 

Bas.: Aspidium palisotii DESV. // Mag. Neuesten Entdeck. 

Gesammten Naturk. Ges. Naturf. Freunde Berlin 5: 320 

(1811) 

Ety.: in honor of A. M. F. J. Palisot de Beauvois (French botanist, 

and collector of type specimen) 

Type: Owaria Africes, Palisot de Beauvois s. n. (HT: P) // West Africa 

TW: FOT 1975 (1975) 

ASPLENIUM (Aspleniaceae) 

Ety.: a (G: privative), splen (G: spleen); relating to medical qualities 

(49) Asplenium adiantum-nigrum L. 

Pub.: Sp. Pl. 2: 1081 (1753) 

Loc.: 深山鐵角蕨蕨 - ShenShanTieJiaoJue

Ety.: adiantum relates to genus Adiantum (see there), niger (L: 

dark, black); probably relating to the dark coloured stipe 

Type: LT designated by Fernandes, Bol. Soc. Brot. 56: 64 (1983) 

TW: Hayata, B. 1914-21 (1914) 

(50) Asplenium antiquum MAKINO 

Pub.: J. Jap. Bot. 6: 32 (1929) 

Loc.: 山蘇蘇花 - ShanSuHua 

Ety.: antiquum (L: old); relationship unclear to me, perhaps relating 

to the perception of the dark-green adaxial lamina of plants 

growing in dull places, or from a hypothesis that the plant 

corresponds to "Mitsuna-gasahiwa", a plant appearing in 

ancient Japanese myths 

Type: Japan 

TW: FOT 1975 (1975) 

(51) Asplenium apogamum N. MURAK. & S. I. HATAN. 

Pub.: J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 14(3): 193, f. 6 (1988) 

Loc.: 無配鐵角蕨蕨 - WuPeiTieJiaoJue 

Ety.: ap (G: away from), gamos (G: marriage); relating to the 

apogamous reproduction 

Alt.: Hymenasplenium apogamum (N. MURAK. & S. I. HATAN.) T. 

Type: 

NAKAIKE // New Fl. Japan, Pterid. 841 (1992) 

Ryukyus, Okinawa, Mt. Sedake, Nackejima & Iha 13718 (HT: 

TI) // Japan 

TW: FOJ (1995) 

(52) Asplenium australasicum (J. SM.) HOOK. 

Pub.: Fil. Exot. pl. 88 (1854) 

Loc.: 南洋山蘇蘇花 - NanYangShanSuHua 

Bas.: Neottopteris australasica J. SM. // Cult. Ferns 49 (1857) 

Ety.: australasia (L: Southern Asia); relating to its distribution (as 

known to its author) and includes Australia, the type location 

Type: cult. Hort. Bot. Kew., origin Australia (BM, in herb. J. Sm.) // 

Australia 

TW: a: Matsumara, J. & Hayata, B. (1906), as synonym of 

Asplenium nidus L. 

b: Kuo, C. M. 1997 (1997) 

(53) Asplenium boreale (OHWI ex SA. KURATA) T. NAKAIKE 

Pub.: New Fl. Japan Pterid. 839 (1992) 

Loc.: 北方倒掛鐵角蕨蕨 - BeiFangDaoGuaTieJiaoJue 

Bas.: Asplenium normale var. boreale OHWI ex SA. KURATA // J. 

Geobot. 11: 100 (1963) 

Ety.: boreale (L: northern); relating to its distribution 

Type: Tochigi, Mt. Kogashi, H. Sekimoto s. n., 1-Aug-1923 (HT: TNS) 

// Japan 


19

20 

(54) Asplenium bullatum WALL. ex METT. [IUCN: NT (LC)] 

Pub.: Numer. List n. 215 (1828) ex Mettenius, Abh. Senckenberg. 

Naturf. Ges. 6: 106 (1859) 

Loc.: 大鐵角蕨蕨 - DaTieJiaoJue 

Ety.: bullatum (L: blistered); relating to the uneven surface of 

adaxial fertile lamina (little raised swollen-looking lines over 

the sorus) 

Type: Wallich 215 (US, Z) // Nepal 

TW: FOT 1975 (1975) 

(55) Asplenium capillipes MAKINO 

Pub.: Bot. Mag. (Tokyo) 17: 77 (1903) 

Loc.: 姬鐵角蕨蕨 - JiTieJiaoJue 

Ety.: 

Type: 

capillus (L: hair), pes (L: foot); relating to thin stipe 

Iyo, Mt. Fudzinoishi-goe, Y. & T. Yoshinaga s. n., 5-Aug-1890 

(HT: TI; IST: TNS) // Japan 


(56) Asplenium cataractarum ROSENST. 

Pub.: Hedwigia 56(4): 334 (1915) [non Blume 1828] 

Loc.: 

Ety.: 

單邊鐵角蕨蕨 - DanBianTieJiaoJue 

cataracta (L: waterfall); relating to its habitat, typically growing 

close to water 

Hymenasplenium cataractarum (comb. ined.) 

Tamsui, Faurie 151 (HT: S; IT: TI, UC) // Taiwan 

Alt.: 

Type: 

TW: Hedwigia 56(4): 334 (1915) [non Blume 1828] 

(57) Asplenium cheilosorum KUNZE ex METT. 

Pub.: Abh. Senckenberg. Naturf. Ges. 6: 133, t. 5, f. 12-13 (1859) 

Loc.: 薄薄葉孔雀鐵角蕨蕨 - BoYeKongQueTieJiaoJue 

Ety.: cheilos (G: margin), soros (G: sorus); relating to the position of 

sori 

Alt.: Hymenasplenium cheilosorum (KUNZE ex METT.) TAGAWA // 

Acta Phytotax. Geobot. 7: 84 (1938) 

Gardner 27 (BM) // Sri Lanka 

Type: 


(58) Asplenium cuneatiforme CHRIST 

Pub.: Bull. Herb. Boissier, ser. 2, 4(7): 613 (1904) 

Loc.: 大蓬萊鐵角蕨蕨 - DaPengLaiTieJiaoJue 

Ety.: cuneatus (L: wedge-shape), forma (L: shape, appearance); 

relating to the shape of pinnae 

Type: Kushaku, Faurie 661 (IT: B, KYO) // Taiwan 

TW: Bull. Herb. Boissier, ser. 2, 4(7): 613 (1904) 

(59) Asplenium ensiforme WALL. ex HOOK. & GREV. 

Pub.: Numer. List n. 200 (1828) ex Hooker & Greville, Icon. Filic. 1: t. 

71 (1828) 

Loc.: 

劍葉鐵角蕨蕨 - JianYeTieJiaoJue

Ety.: ensis (L: sword), forma (L: shape, appearance); relating to the 

frond shape 

Type: Wallich 200 (IT: UC) // Nepal 

TW: Ito, H. 1944 (1944) 

(60) Asplenium excisum C. PRESL 

Pub.: Epimel. Bot. 74 (1851) 

Loc.: 剪葉鐵角蕨蕨 - JianYeTieJiaoJue 

Ety.: excisum (L: cut-out, cut-off); relating to the unsymmetrical 

shape of (dimidiate) pinna base 

Alt.: Hymenasplenium excisum (C. PRESL) S. LINDS. // Thai Forest 

Bull., Bot. 37: 69 (2009) 

Type: Luzon, Cuming 110 (HT: E; IT: L) // Philippines 

TW: FOT 1975 (1975) 

(61) Asplenium falcatum LAM. 

Pub.: Encycl. 2: 306 (1786) 

Loc.: 革葉鐵角蕨蕨 - GeYeTieJiaoJue 

Ety.: falcatum (L: sickle-shaped), relating to the shape of pinna 

Type: LT: Burman, Thesaurus Zeylanicus t. 43 (1737), designated in 

Fern Gaz. 19(6): 223 (2013) // Sri Lanka 

TW: Henry, A. (1896) 

(62) Asplenium filipes COPEL. 

Pub.: Philipp. J. Sci. 3: 34 (1908) 

Loc.: 複齒鐵角蕨蕨 - FuChiTieJiaoJue 

Ety.: fili (L: thread-like), pes (L: foot); relating to thin stipe 

Alt.: Hymenasplenium filipes (comb. ined.) 

Type: Mt. Mariveles, 1200 m, Copeland P. P. E. 73 (HT: MICH; IT: L, 

S, UC, US) // Philippines 

TW: (a) FOJ (1995) but placing Asplenium obliquissimum (Hayata) 

Sugim. et Sa. Kurata into synonymy, which might be not 

correct 

(b) Kuo, C. M. 1997 (1997) 

(63) Asplenium formosae CHRIST 

Pub.: Bull. Herb. Boissier, ser. 2, 4(7): 613 (1904) 

Loc.: 南海鐵角蕨蕨 - NanHaiTieJiaoJue 


Type: Kushaku, Faurie 669 (P) // Taiwan 

TW: Bull. Herb. Boissier, ser. 2, 4(7): 613 (1904) 

(64) Asplenium formosae CHRIST x Asplenium griffithianum HOOK. 


(65) Asplenium griffithianum HOOK. 

Pub.: Icon. Pl. pl. 928 (1854) 

Loc.: 叢葉鐵角蕨蕨 - CongYeTieJiaoJue 

Ety.: in honor of W. Griffith (British botanist, and collector of type 

specimen) 

Type: Mishmee, E. Indies, W. Griffith s. n. (K) // ?India 

21

22 

TW: Ogata, M. (1936) 

(66) Asplenium incisum THUNB. 

Pub.: Trans. Linn. Soc. London 2: 342 (1794) 

Loc.: 縮羽鐵角蕨蕨 - SuoYuTieJiaoJue 

Ety.: incisum (L: cut), relating to the pinna margin 

Type: Japan 

TW: FOT 1975 (1975) 

(67) Asplenium lacinioides FRASER-JENK., PANGTEY & KHULLAR 

Pub.: Indian Fern J. 27(1-2): 211-212, fig. 36-37 (2011) 

Loc.: 鱗柄鐵角蕨蕨 - LinBingTieJiaoJue 

Ety.: laciniatum (L: fringed) relating to the taxon with epithet 

"laciniatum", oides (L: suffix, denoting likeness of form); 

relates to the similarity to taxon Asplenium laciniatum D. DON 

Type: Sikkim, N. Sikkim District, Singhik, 4000 ft., S. S. Bir 1121, 30- 

Jul-1958 (HT: PAN) // India 

TW: Indian Fern J. 27(1-2): 211-212, fig. 36-37 (2011) 

(68) Asplenium neolaserpitiifolium TARDIEU & CHING [IUCN: - (VU)] 

Pub.: Notul. Syst. (Paris) 5: 153-154, pl. 6, f. 1-2 (1936) 

Loc.: 大黑柄鐵角蕨蕨 - DaHeiBingTieJiaoJue 

Ety.: neos (G: new), laserpitiifolium relating to the taxon with epithet 

"laserpitiifolium" (laserpitium relates to a flowering plant 

genus (Apiaceae) with similar leaves); relating to the 

similarity to taxon Asplenium laserpitiifolium LAM. 

Type: Hainan, Katsumata 6706 (HT: P) // China 

TW: Kuo, C. M. 1985 (1985) 

(69) Asplenium nidus L. 

Pub.: Sp. Pl. 2: 1079 (1753) 

Loc.: 台台灣山蘇蘇花 - TaiWanShanSuHua 

Ety.: 

Type: 

nidus (L: nest); relating to its appearance 

Java, Osbeck s. n. (LT: LINN), designated by Holttum in Gard. 

Bull. Singapore 27: 147 (1974) // Indonesia 

TW: J. Bot. 23: 104 (1885) 

(70) Asplenium normale D. DON 

Pub.: Prodr. Fl. Nepal. 7 (1825) 

Loc.: 生芽鐵角蕨蕨 - ShengYaTieJiaoJue 

Ety.: normalis (L: according to the rule, in a straight way or line); 

probably relating to the habit of frond 

Type: Narainhetty, Buchanan-Hamilton s. n. (HT: BM) // Nepal 

TW: J. Bot. 23: 104 (1885) 

(71) Asplenium obscurum BLUME * 

[IUCN: NT (LC)] 

Pub.: Enum. Pl. Javae 181-182 (1828) 

Loc.: 綠柄剪葉鐵角蕨蕨 - LüBingJianYeTieJiaoJue 

Ety.: obscurum (L: dusky, dark); relating to the lamina colour 

Alt.: Hymenasplenium obscurum (BLUME) TAGAWA // Acta Phytotax. 

Geobot. 7: 83 (1938)

Type: Java, Blume s. n. (B, L) // Indonesia 

TW: Ogata, M. (1930) 

(72) Asplenium oldhami HANCE 

Pub.: Ann. Sci. Nat., Bot., ser. 5, 5: 256-257 (1866) 

Loc.: 俄氏鐵角蕨蕨 - EShiTieJiaoJue 

Ety.: in honor of R. Oldham (British botanist) 

Type: Tamsui, Oldham s. n., Apr-1864 // Taiwan 

TW: Ann. Sci. Nat., Bot., ser. 5, 5: 256-257 (1866) 

(73) Asplenium pekinense HANCE [IUCN: DD (LC)] 

Pub.: J. Bot. 5: 262 (1867) 

Loc.: 北京鐵角蕨蕨 - BeiJingTieJiaoJue 

Ety.: relates to BeiJing ("Peking") in China, the type location 

Type: Beijing, S. W. Williams 12404, summer of 1865 // China 

TW: Kuo, C. M. 1985 (1985) 

(74) Asplenium prolongatum HOOK. 

Pub.: Sec. Cent. Ferns pl. 42 (1861) 

Loc.: 長生鐵角蕨蕨 - ChangShengTieJiaoJue 

Ety.: prolongatum (L: prolonged); relating to the rachis, which is 

often prolonged and carries plantlets 

Type: Japan 

TW: Hayata, B. 1917 (1917) 

(75) Asplenium pseudolaserpitiifolium CHING ex TARDIEU & CHING 

(LC)] 

[IUCN: DD 

Pub.: Notul. Syst. (Paris) 5: 150-153, pl. 8 (1936) 

Loc.: 黑鱗鐵角蕨蕨 - HeiLinTieJiaoJue 

Ety.: pseudos (G: false), laserpitiifolium relating to the taxon with 

epithet "laserpitiifolium" (laserpitium relates to a flowering 

plant genus (Apiaceae) with similar leaves); relating to the 

similarity to taxon Asplenium laserpitiifolium LAM. 

Type: Massif du Pia Ouac, Petelot 660, Jul-1922 (HT: P; IT: MICH) // 

Vietnam 

TW: Kuo, C. M. 1997 (1997) 

(76) Asplenium pulcherrimum (BAKER) CHING ex TARDIEU 

Pub.: Asplen. Tonkin 52 (1932) 

Loc.: 細葉鐵角蕨蕨 - XiYeTieJiaoJue 

Bas.: Davallia pulcherrima BAKER // Bull. Misc. Inform. Kew 1895: 53 

(1895) 

Ety.: superlative of pulcher (L: beautiful); relating to the habit 

Type: Yunnan, near Mongtse, 6000 ft, Hancock 105 (HT: K) // China 

TW: Tagawa, M. 1940-49 (1940) 

(77) Asplenium ritoense HAYATA 

Pub.: Icon. Pl. Formosan. 4: 226-227, f. 156 (1914) 

Loc.: 尖尖葉鐵角蕨蕨 - JianYeTieJiaoJue 

Ety.: relating to Rito, a mountain in Taiwan and type location 

23

24 

Type: Mt. Ritozan, ad 6500 ped. alt., T. Kawakami s. n., Jul-1913 

(HT: TI) // Taiwan 


(78) Asplenium ruta-muraria L. 

[IUCN: VU (LC)] 

Pub.: Sp. Pl. 2: 1081 (1753) 

Loc.: 銀杏葉鐵角蕨蕨 - YinXingYeTieJiaoJue 

Ety.: ruta relates to a flowering plant genus (Rutaceae, "rue"), 

murus (L: wall); relating to the shape of lamina resembling 

leaves of Ruta, and the distribution (frequently found growing 

on walls) 

Type: (LT: LINN), designated by Viane, Nordic J. Bot. 14: 149 (1994) 


(79) Asplenium scolopendrium L. 

[IUCN: VU] 

Pub.: Sp. Pl. 2: 1079 (1753) 

Loc.: 對對開蕨蕨 - DuiKaiJue 

Ety.: skolopendrion relates to a Greek name of an Asplenium, 

probably this taxon, relating to the shape of sori resembling a 

centipede (scolopender) 

Type: (LT: LINN), designated by Reichstein, Bot. Helv. 91: 106 

(1981) 

TW: Kuo, C. M. 1997 (1997) 

(80) Asplenium septentrionale (L.) HOFFM. [IUCN: NT] 

Pub.: Deutschl. Fl. 2: 12-13 (1795) 

Loc.: 線葉鐵角蕨蕨 - XianYeTieJiaoJue 

Bas.: Acrostichum septentrionale L. // Sp. Pl. 2: 1068 (1753) 

Ety.: septentrionalis (L: northern); relating to the distribution 

(northernmost taxon of genus Acrostichum, into which 

Linnaeus combined this taxon) 

Type: (LT: UPS), designated by Jonsell & Jarvis, Nordic J. Bot. 14: 

150 (1994) 

TW: a: Masamune, G. (1936) as Amesium septentrionale var. 

sasakii (HAYATA) MASAM. 

b: Tagawa, M. 1940-49 (1941) 

(81) Asplenium serricula FEE [IUCN: NT (VU)] 

Pub.: Mem. Foug. 5: 196 (1852) 

Loc.: 蘭蘭嶼鐵角蕨蕨 - LanYuTieJiaoJue 

Ety.: serra (L: saw), cula (L: suffix added to form diminutive form); 

relating to the acroscopic pinna margin 

Type: Gardner 30 // Sri Lanka 

TW: Kuo, C. M. 1985 (1985) 

(82) Asplenium x shikokianum MAKINO 

Pub.: Bot. Mag. (Tokyo) 13: 13-14 (1899) 

Loc.: 四國鐵角蕨蕨 - SiGuoTieJiaoJue 

Ety.: relating to the island of Shikoku (Japan), probably the type 

location ("Suginokawa in Hayama" and "Usa")

Type: Tosa, Suginokawa (ST: MAK, TI) // Japan 

TW: Tagawa, M. 1940-49 (1941) 

(83) Asplenium sp. 

[IUCN: - (VU)] 

Alt.: Hymenasplenium sp. 


(84) Asplenium subnormale COPEL. [IUCN: DD (NT)] 

Pub.: Fragm. Fl. Philipp. 183, pl. 4, f. B (1905) 

Loc.: 小小鐵角蕨蕨 - XiaoTieJiaoJue 

Ety.: sub (L: similar to), normale relating to the taxon with epithet 

"normale"; relating to the similarity to taxon Asplenium 

normale (see there) 

Alt.: Hymenasplenium subnormale (COPEL.) T. NAKAIKE // New Fl. 

Japan 841 (1992) 

Type: Luzon, Copeland 236 (HT: PNH, lost; IT: B, MICH) // 

Philippines 

TW: Kuo, C. M. 1997 (1997) 

(85) Asplenium tenerum G. FORST. [IUCN: NT] 

Pub.: Fl. Ins. Austr. 80 (1786) 

Loc.: 鈍齒鐵角蕨蕨 - DunChiTieJiaoJue 

Ety.: tenerum (L: soft); probably relating to the lamina texture 

Type: Australia 


(86) Asplenium tenuicaule HAYATA 


Loc.: 小小葉鐵角蕨蕨 - XiaoYeTieJiaoJue 

Ety.: 

Type: 

tenuis (L: thin), caulis (L: stem); relating to the thin stipe 

Arisan, ad 6000-7000 ped. alt., B. Hayata & S. Sasaki s. n., 

Jan-1912 (HT: TI) // Taiwan 


(87) Asplenium tenuifolium D. DON 


Loc.: 

Ety.: 

Type: 

薄薄葉鐵角蕨蕨 - BoYeTieJiaoJue 

tenuis (L: thin), folium (L: leaf); relating to the quality of lamina 

Wallich 216 (LT: BR; ILT: B; UC), verified by C. V. Morton on 

1-Jul-1970 // Nepal 

TW: Ito, H. 1944 (1944) 

(88) Asplenium trichomanes L. 

Pub.: Sp. Pl. 2: 1080 (1753) 

Loc.: 

Ety.: 

Type: 

25 

鐵角蕨蕨 - TieJiaoJue 

two alternatives: (1) thrix (G: hair), mainomai (G: to be furious); 

relating to ferns with black, hair-like stipes pointing in all 

directions; (2) resembling Trichomanes, a fern genus (see 

there) 

"Habitat in Europae fissuris rupium" (LT: LINN), designated by 

Bobrov in Novosti Sist. Vyssh. Rast. 21 : 15 (1984) // Europe

26 

TW: Hayata, B. 1908 (1908) 

(89) Asplenium tripteropus NAKAI 


Loc.: 三翅鐵角蕨蕨 - SanChiTieJiaoJue 

Ety.: treis (G: three), pteron (G: wing), pous (G: foot); relating to the 

three wings on the stipe 

Type: Honsyu, Mt. Nachi, T. Nakai s. n. (ST: TI) // Japan 


(90) Asplenium viride HUDS. 

Pub.: Fl. Angl. 385 (1762) 

Loc.: 綠柄鐵角蕨蕨 - LüBingTieJiaoJue 

Ety.: 

Type: 

viridis (L: green); relating to the entirely green rachis 

in rupibus humidis in montibus Walliæ et in comitatibus 

Eboracensi et Westmorlandico // United Kingdom 

TW: Ogata, M. (1940) 

(91) Asplenium x wangii C. M. KUO 

Pub.: Bot. Bull. Acad. Sin. 29(2): 109-111 (1988) 

Loc.: 

Ety.: 

王氏鐵角蕨蕨 - WangShiTieJiaoJue 

in honor of B. J. Wang (Taiwanese botanist, collector of the 


Chiayi, Fushan, Wang 9600 (HT: TAI; IT: Z) // Taiwan 

Type: 

TW: Bot. Bull. Acad. Sin. 29(2): 109-111 (1988) 

(92) Asplenium wilfordii METT. ex KUHN 

Pub.: Linnaea 36: 94-95 (1870) 

Loc.: 

Ety.: 

Type: 

TW: 

威氏鐵角蕨蕨 - WeiShiTieJiaoJue 

in honor of C. Wilford (British Botanist) 

Tsus-sima, C. Wilford 783, May-1859 (K) // China 

a: Hayata, B. 1914-21 (1919) as Asplenium wilfordii var. 

densum ROSENST. 

b: Sasaki, S. (1928) 

(93) Asplenium wrightii EATON ex HOOK. 

Pub.: Sp. Fil. 3: 113-114, pl. 182 (1859-1860) 

Loc.: 萊氏鐵角蕨蕨 - LaiShiTieJiaoJue 

Ety.: in honor of C. Wright (US botanist and collector of the type 

specimen) 

Type: Honshu, C. Wright s. n. (BM, GH, US) // Japan 


(94) Asplenium yoshinagae MAKINO 

Pub.: Phan. Pterid. Jap. Icon. 1: pl. 64 (1900) 

Loc.: 斜葉鐵角蕨蕨 - XieYeTieJiaoJue 

Ety.: relates to E. Yoshinaga (Japanese botanist, collector of type 

specimen) 

Type: ?Mt. Tebako, Yoshinaga s. n. (TI) // Japan 

TW: Ito, H. 1944 (1944)

(95) Asplenium yunnanense FRANCH. [IUCN: DD (VU)] 

Pub.: Bull. Soc. Bot. France 32: 28 (1885) 

Loc.: 雲南鐵角蕨蕨 - YunNanTieJiaoJue 

Ety.: relates to YunNan (China, type location) 

Type: Yunnan, in monte Che-tcho-tze, supra Ta-pin-tze, prope Tali, 

ca. 3000 m, Delavay 72, 23-Aug-1883 (HT: P) // China 

TW: Kuo, C. M. 1985 (1985) 

ATHYRIUM (incl. Rhachidosorus) (Dryopteridaceae) 

Ety.: athyros (G: doorless); relating to the sporangia only tardily pushing 

back the outer edge of indusium 

Rhachidosorus: rhachis (G: central axes of a compound lamina), 

soros (G: sorus); perhaps relating to the position of sori near the 

rachis 

(96) Athyrium anisopterum CHRIST 

Pub.: Bull. Herb. Boissier 6: 962-963 (1898) 

Loc.: 宿宿蹄蓋蕨蕨 - SuTiGaiJue 

Ety.: anisos (G: unequal), pteron (G: wing); relating to the pinna 

shape 

Type: Yunnan, Henry 10109 (HT: P; IT: MO, US) // China 

TW: Ito, H. 1944 (1944) 

(97) Athyrium arisanense (HAYATA) TAGAWA 


Loc.: 阿里山蹄蓋蕨蕨 - ALiShanTiGaiJue 

Bas.: Diplazium arisanense HAYATA // Icon. Pl. Formosan. 4: 212- 

213, f. 144 (1914) 

Ety.: relating to ALiShan ("Arisan", Taiwan, the type location) 

Type: Arisan, Hayata & Sasaki s. n., Jan-1912 (HT: TI) // Taiwan 


(98) Athyrium atkinsonii BEDD. 

Pub.: Suppl. Ferns S. Ind. 11, pl. 359 (1876) 

Loc.: 大葉假冷蕨蕨 - DaYeJiaLengJue 

Ety.: in honor of W. S. Atkinson (British botanist and collector of the 


Type: Sikkim, Atkinson s. n. (K) // India 

TW: FOT 1975 (1975) as Pseudocystopteris atkinsonii (Bedd.) 

Ching 

(99) Athyrium auriculatum SERIZ. [IUCN: VU] 

Pub.: J. Jap. Bot. 46: 280-281 (1971) 

Loc.: 耳垂蹄蓋蕨蕨 - ErChuiTiGaiJue 

Ety.: auriculatum (L: furnished with ear-like appendages); relating to 

the shape of pinna base 

Type: Kaohsiung Co., Terngjy, Daigobo 634 (HT: TNS) // Taiwan 

TW: J. Jap. Bot. 46: 280-281 (1971) 

27

28 

(100) Athyrium cryptogrammoides HAYATA 

Pub.: Icon. Pl. Formosan. 6: 156-157 (1916) 

Loc.: 合合歡山蹄蓋蕨蕨 - HeHuanShanTiGaiJue 

Ety.: cryptogramma relating to the fern genus Cryptogramma (see 

there), oides (L: suffix, denoting likeness of form); relates to 

the similarity of fronds to that of genus Cryptogramma 

Type: Hokwanzan, Yada s. n., Dec-1914 (HT: ?TI) // Taiwan 

TW: Icon. Pl. Formosan. 6: 156-157 (1916) 

(101) Athyrium cumingianum (C. PRESL) MILDE [IUCN: NT (EN)] 

Pub.: Bot. Zeitung (Berlin) 28: 353 (1870) 

Loc.: 安安蕨蕨 - AnJue 

Bas.: Anisocampium cumingianum C. PRESL // Epimel. Bot. 59 

(1851) 

Ety.: in honor of H. Cuming (British botanist, and collector of the 


Alt.: Anisocampium cumingianum C. PRESL // Epimel. Bot. 59 

(1851) 

Type: Luzon, Cuming 239 (HT: K) // Philippines 

TW: Taiwania 45(2): 182 (2000) 

(102) Athyrium delavayi CHRIST var. delavayi * 

Pub.: Bull. Soc. Bot. France 52, Mem. 1: 47 (1905) 

Loc.: 溪谷蹄蓋蕨蕨 - XiGuTiGaiJue 

Ety.: in honor of P. J. M. Delavay (French botanist, and collector of 

the type specimen) 

Type: Mt. Omei, Delavay 5035, 5123 (MT: PE), 5166; Faber s. n. 

(ST: P) // China 

TW: Kuo, C. M. 1985 (1985) 

(103) Athyrium delavayi var. subrigescens (HAYATA) Y. C. LIU 

Pub.: Fern Fl. Taiwan Athyrium 18 (2009) 

Loc.: 姬蹄蓋蕨蕨 - JiTiGaiJue 

Bas.: Diplazium subrigescens HAYATA // Icon. Pl. Formosan. 4: 219- 

220, f. 149 (1914) 

Ety.: sub (L: similar to), rigescens (L: growing stiff, stand up) relating 

to the taxon with epithet "rigescens"; relating to the similarity 

to taxon Asplenium rigescens 

Type: Arisan, ad 7000 ped. Alt., B. Hayata & S. Sasaki s. n., Jan- 

1912 (HT: TI) // Taiwan 


(104) Athyrium epirachis (CHRIST) CHING [IUCN: DD] 

Pub.: Index Filic., Suppl. 3: 41 (1934) 

Loc.: 軸果蹄蓋蕨蕨 - ZhouGuoTiGaiJue 

Bas.: Diplazium epirachis CHRIST // Bull. Soc. Bot. France 52, Mem. 

1: 51-52 (1905)

Ety.: epi (G: on, upon, upright), rachia (G: literally a ridge, here 

relating to the rachis); relating to the abundant hairs on the 

rachis 

Type: Kweichow, Bodinier 1706 (HT: P) // China 

TW: Kuo, C. M. 1985 (1985) 

(105) Athyrium erythropodum HAYATA 


Loc.: 紅柄蹄蓋蕨蕨 - HongBingTiGaiJue 

Ety.: erythros (G: red, reddish), pous (G: foot); relating to the colour 

of stipe 

Type: Monte Arisan, B. Hayata & S. Sasaki s. n., Jan-1912 (HT: TI) // 



(106) Athyrium iseanum ROSENST. 

Pub.: Repert. Spec. Nov. Regni Veg. 13: 124-125 (1913) 

Loc.: 細葉蹄蓋蕨蕨 - XiYeTiGaiJue 

Ety.: relates to Ise, a place in Japan and the type location 

Type: Ise, in silvis profundis, Sakurai 42, 17-Aug-1913 (S) // Japan 

TW: Kuo, C. M. 1985 (1985) 

(107) Athyrium leiopodum (HAYATA) TAGAWA 


Loc.: 小小葉蹄蓋蕨蕨 - XiaoYeTiGaiJue 

Bas.: Diplazium leiopodum HAYATA // Icon. Pl. Formosan. 4: 217- 

Ety.: 

219, f. 148 (1914) 

leios (G: smooth), pous (G: foot); relating to the surface of 

stipe 

Type: Arisan, B. Hayata & S. Sasaki s. n., Jan-1912 (ST: TI) // 



(108) Athyrium minimum CHING [IUCN: EN (VU)] 

Pub.: Acta Bot. Boreal.-Occid. Sin. 6(3): 151 (1986) 

Loc.: 七星山蹄蓋蕨蕨 - QiXingShanTiGaiJue 

Ety.: minimum (L: smallest); relating to the habit (the type is 

extremely small, not typical for this taxon) 

Type: Tamsui, Hancock s. n., Dec-1881 (HT: PE) // Taiwan 

TW: Acta Bot. Boreal.-Occid. Sin. 6(3): 151 (1986) 

(109) Athyrium mupinense CHRIST 

Pub.: Bot. Gaz. 51: 355-356 (1911) 

Loc.: 

Ety.: 

細裂蹄蓋蕨蕨 - XiLieTiGaiJue 

relates to Mupin (BaoXing in SiChuan Province), a place in 

China and the type location 

Sichuan, Wilson 2610 (HT: P; IT: US) // China 

Type: 

TW: Liu, Y. C. 2009 (2009) 

(110) Athyrium nakanoi MAKINO 


29

30 

Loc.: 紅苞蹄蓋蕨蕨 - HongBaoTiGaiJue 

Ety.: in honor of Y. Nakano (Japanese botanist) 

Type: Yakushima Island, Kyusyu, T. Makino s. n., Sep-1909 (HT: TI; 

IT: KYO) // Japan 


(111) Athyrium niponicum (METT.) HANCE [IUCN: NT] 

Pub.: J. Linn. Soc., Bot. 13: 92-93 (1873) 

Loc.: 日本蹄蓋蕨蕨 - RiBenTiGaiJue 

Bas.: Asplenium niponicum METT. // Ann. Mus. Bot. Lugduno-Batavi 

2: 240 (1866) 

Ety.: relating to Nippon (=Japan), the type location 

Alt.: Anisocampium niponicum (METT.) Y. C. LIU, W. L. CHIOU & M. 

KATO // Taxon 60(3): 828 (2011) 

Type: (ST: L) // Japan 

TW: FOT 1975 (1975) 

(112) Athyrium oppositipennum HAYATA var. oppositipennum * 

Pub.: J. Coll. Sci. Imp. Univ. Tokyo 30: 441-442 (1911) 

Loc.: 對對生蹄蓋蕨蕨 - DuiShengTiGaiJue 

Ety.: oppositum (L: opposite), penn (L: pinna); relating to the lamina 

habit 

Type: Mt. Morrison, Kawakami & Mori 1863 (HT: ?TI) // Taiwan 


(113) Athyrium oppositipennum var. pubescens (TAGAWA) TAGAWA * 

Pub.: Acta Phytotax. Geobot. 4(3): 142 (1935) 

Loc.: 逆葉蹄蓋蕨蕨 - NiYeTiGaiJue 

Bas.: Athyrium taiwanense var. pubescens TAGAWA // Acta Phytotax. 

Geobot. 2: 18 (1933) 

Ety.: pubescens (L: short hairy); relating to the axes of frond 

Type: Arisan, 2500 m alt., Faurie 566, Jun-1914 (HT: KYO; IT: KYO) 


TW: Acta Phytotax. Geobot. 2: 18 (1933) 

(114) Athyrium otophorum (MIQ.) KOIDZ. 

Pub.: Fl. Symb. Orient.-Assiat. 40 (1930) 

Loc.: 光蹄蓋蕨蕨 - GuangTiGaiJue 

Bas.: Asplenium otophorum MIQ. // Ann. Mus. Bot. Lugduno-Batavi 

3: 175 (1867) 

Ety.: otos (G: ear shaped), phero (G: to bear); relating to the 

acroscopic auricle at the base of pinnules 

Type: Buerger s. n. (LT: L) // Japan 

TW: Kuo, C. M. 1985 (1985) 

(115) Athyrium palustre SERIZ. * 

[IUCN: - (EN)] 

Pub.: J. Jap. Bot. 45: 264 (1970) 

Loc.: 沼生蹄蓋蕨蕨 - ZhaoShengTiGaiJue 

Ety.: palustre (L: marshy); relating to the habitat, only found in 

wetland

Type: Kyushu, Kagoshima Prefecture, Yaku Island (=Yakushima), 

near Mt. Ishizuka, S. Serizawa s. n., Nov-1968 (HT: TNS) // 

Japan 

TW: TW J. of Biodivers. 16(3): 263-272 (2014) 

(116) Athyrium pubicostatum CHING & Z. Y. LIU 

Pub.: Bull. Bot. Res., Harbin 4(3): 7-8, f. 9 (1984) 

Loc.: 假軸果蹄蓋蕨蕨 - JiaZhouGuoTiGaiJue 

Ety.: pubes (L: hairs (of adolescence)), costatum (L: provided with a 

costa); relating to the presence of hairs on costa 

Type: Sichuan, Z. Y. Liu 4168 (HT: PE) // China 

TW: Liu, Y. C. 2008 (2008) 

(117) (Athyrium pulchrum comb. ined.) Rhachidosorus pulcher (TAGAWA) 

CHING [IUCN: NT] 

Pub.: Acta Phytotax. Sin. 9: 74-75 (1964) 

Loc.: 花莲蹄盖蕨蕨 - HuaLianTiGaiJue 

Bas.: Diplazium pulchrum TAGAWA // Acta Phytotax. Geobot. 4: 144- 

145 (1935) 

Ety.: pulcher (L: beautiful); relating to the habit 

Alt.: Rhachidosorus pulcher (TAGAWA) CHING // Acta Phytotax. Sin. 

9: 74 (1964) 

Type: Hualien, between Batakan and Tabito, Tagawa 702 (HT: KYO; 

IT: MICH, TAI) // Taiwan 


(118) Athyrium puncticaule (BLUME) T. MOORE [IUCN: VU (?EN)] 

Pub.: Index. Fil. 186 (1860) 

Loc.: 密密腺蹄蓋蕨蕨 - MiXianTiGaiJue 

Bas.: Aspidium puncticaule BLUME // Enum. Pl. Javae 159 (1828) 

Ety.: punctus (L: puncture, spotted), caulis (L: stem); relating to the 

presence of dark spots on stipe (based on protologue) 

Type: Java, anonymous (HT: L) // Indonesia 

TW: Taiwania 51(4): 293-297 (2006) 

(119) Athyrium sheareri (BAKER) CHING [IUCN: - (CR)] 


Loc.: 華東安安蕨蕨 - HuaDongAnJue 

Bas.: Nephrodium sheareri BAKER // J. Bot. 13: 200 (1875) 

Ety.: in honor of J. Shearer (British botanist, collector of type 

specimen) 

Alt.: Anisocampium sheareri (BAKER) CHING // Acta Bot. Yunnan. 

7(3): 314 (1985) 

Type: 

TW: MAN (2002) 

(120) Athyrium silvicola TAGAWA 

Jiangxi, Lushan, Shearer s. n. 1873 (HT: K) // China 


Loc.: 高山蹄蓋蕨蕨 - GaoShanTiGaiJue 

31

32 

Ety.: silva (L: forest), colo (L: to inhabit); relating to the growth habit 

(typically growing in forests, though this is not too diagnostic 

for members of this genus) 

Type: Mt. Arisan, Faurie 564 (HT: KYO; IT: KYO) // Taiwan 


(121) Athyrium sp. 

TW: Liu, Y. C. 2008 (2008) 

(122) Athyrium strigillosum (T. MOORE ex E. J. LÖWE) SALOMON 

Pub.: Nomencl. Gefässkrypt. 112 (1883) 

Loc.: 生芽蹄蓋蕨蕨 - ShengYaTiGaiJue 

Bas.: Asplenium strigillosum T. MOORE ex E. J. LÖWE // Ferns 5: 107- 

108, pl. 36 (1858) 

Ety.: strigillosum (L: strigillose, with stiff, slender bristles); relating to 

the prominent spines on adaxial costules and pinnule midribs 

Type: Nepal 

TW: Kuo, C. M. 1985 (1985) 

(123) Athyrium tozanense (HAYATA) HAYATA 


Loc.: 蓬萊蹄蓋蕨蕨 - PengLaiTiGaiJue 

Bas.: Asplenium tozanense HAYATA // J. Coll. Sci. Imp. Univ. Tokyo 

30: 440-441 (1911) 

Ety.: relating to Toza, a place in Taiwan and type location 

Type: Tozan, Nakahara 1907 (ST: TI) // Taiwan 


(124) Athyrium tripinnatum TAGAWA 


Loc.: 三回蹄盖蕨蕨 - SanHuiTiGaiJue 

Ety.: tri (L: three), pinnatum (L: pinnate); relating to the lamina 

Type: 

shape (being three-times pinnate) 

Tainan, inter Numanohira et Tataka, M. Tagawa 398 (HT: 

KYO; IT: L) // Taiwan 


(125) Athyrium vidalii (FRANCH. & SAV.) NAKAI 


Loc.: 山蹄蓋蕨蕨 - ShanTiGaiJue 

Bas.: Asplenium vidalii FRANCH. & SAV. // Enum. Pl. Jap. 2: 229, 624- 

625 (1877) 

Ety.: in honor of S. Vidal (Spanish botanist) 

Type: Japan 

TW: Ito, H. 1944 (1944) 

AZOLLA (Azollaceae) 

Ety.: azo (G: to dry), olluo (G: to kill); this aquatic fern is presumably killed 

by drying

(126) Azolla filiculoides LAM. 

Pub.: Encycl. 1: 343 (1783) 

Loc.: 細葉滿江紅 - XiYeManJiangHong 

Ety.: filiculoides (L: like a small fern); relating to the appearance of 

the plant 

Type: Straits of Magellan, Commerson s. n. (HT: P) // Argentina 


(127) Azolla pinnata R. BR. 

[IUCN: DD (VU)] 

Pub.: Prodr. 167 (1810) 

Loc.: 滿江紅 - ManJiangHong 

Ety.: pinnata (L: with feathers); relating to the outline of the plant 

Type: Richmond, Hawkesbury, R. Brown 134, Oct-1804 (LT: BM; 

ILT: E, K) // Australia 

TW: a: Matsumara, J. & Hayata, B. (1906) as Azolla pinnata var. 

africana BAKER 

b: FOT 1975 (1975) 

BELVISIA (Polypodiaceae) 

Ety.: in honor of A. M. F. J. Palisot de Beauvois (L: Belvisius) 

(128) Belvisia mucronata (FEE) COPEL. [IUCN: CR (EN)] 

Pub.: Gen. Fil. (Copeland) 192 (1947) 

Loc.: 尖尖嘴蕨蕨 - JianZuiJue 

Bas.: Hymenolepis mucronata FEE // Mem. Foug. 5: 82, pl. 6B, f. 1 

(1852) 

Ety.: mucronata (L: pointed); relating to the frond habit 

Alt.: Lepisorus mucronatus (FEE) LI WANG // Bot. J. Linn. Soc. 

162(1): 35 (2010) 

Type: Luzon, Cuming 92 (HT: P; IT: BM, G, P, UC, US, W) // 

Philippines 

TW: Ito, H. 1944 (1944) 

BLECHNUM (Blechnaceae) 

Ety.: blechnon (G: an ancient name for ferns in general) 

(129) Blechnum eburneum CHRIST [IUCN: VU] 

Pub.: Bull. Acad. Int. Geogr. Bot. 11: 233 (1902) 

Loc.: 天長烏毛蕨蕨 - TianChangWuMaoJue 

Ety.: eburneum (L: of ivory); relating to the colour of abaxial (sterile) 

pinna 

Type: Guizhou, Guiyang, Bodinier 2000 (HT: P; IT: P) // China 

TW: a: Tagawa, M. 1940-49 (1940) as Spicantopsis eburnea var. 

obtusa TAGAWA 

b: FOT 1975 (1975) 

(130) Blechnum fraseri (A. CUNN.) LUERSS. [IUCN: CR (EN)] 

Pub.: Flora 59: 292 (1876) 

Loc.: 假桫欏 - JiaSuoLuo 

33

34 

Bas.: Lomaria fraseri A. CUNN. // Companion Bot. Mag. 2: 364 (1836) 

Ety.: in honor of C. Fraser (British botanist and collector of the type 

specimen) 

Alt.: Diploblechnum fraseri (A. CUNN.) DEVOL // Fl. Taiwan 1: 153 

(1975) 

Type: North Island, East coast, Fraser s. n., 1825 (LT: K; ILT: E), 

designated by Chambers & Farrant, Blumea 46: 306 (2001) // 

New Zealand 

TW: a: Ogata, M. (1940), as Blechnum fraseri var. philippinense 

CHRIST 

b: FOT 1975 (1975) 

(131) Blechnum hancockii HANCE 

Pub.: J. Bot. 21: 267 (1883) 

Loc.: 韓氏烏毛蕨蕨 - HanShiWuMaoJue 

Ety.: in honor of W. Hancock (British botanist, collector of the type 

specimen) 

Type: Tamsui, Hancock 87, Nov-1881 (HT: BM) // Taiwan 

TW: J. Bot. 21: 267 (1883) 

(132) Blechnum melanopus HOOK. 

Pub.: Sp. Fil. 3: 64, pl. 161 (1859) 

Loc.: 雉尾尾烏毛蕨蕨 - ZhiWeiWuMaoJue 

Ety.: melaina (G: black), pous (G: foot); relating to the black 

coloured lower half of abaxial rachis (based on protologue) 

Type: Khasya Hills, Simons 78 // India 

TW: Ito, H. 1944 (1944) 

(133) Blechnum orientale L. 

Pub.: Sp. Pl. 2: 1077 (1753) (occidentale ex err.), Sp. Pl. (ed. 2) 2: 

1535 (1763) 

Loc.: 烏毛蕨蕨 - WuMaoJue 

Ety.: orientalis (L: oriental); probably relating to the distribution as 

known to its author ("habitat in China") 

Type: without locality, Osbeck s. n. (LINN) // China 

TW: Henry, A. (1896) 

BOLBITIS (Lomariopsidaceae) 

Ety.: bolbition (G: diminutive of bolbos, bulb); relates to the small bulbs 

(gemmae) borne on the lamina of many species of this genus 

(134) Bolbitis angustipinna (HAYATA) H. ITO 

Pub.: J. Jap. Bot. 14: 443 (1938) 

Loc.: 細葉實實蕨蕨 - XiYeShiJue 

Bas.: Leptochilus angustipinnus HAYATA // Icon. Pl. Formosan. 5: 

297-298, f. 119 (1915) 

Ety.: angusta (L: narrow), pinna (L: wing, here relating to pinna); 

relating to the pinna habit

Type: near Hokkokel, Owatari s. n., Jan-1898 (LT: TI), designated by 

Hennipman, Leid. Bot. Ser. 2: 152 (1977) // Taiwan 


(135) Bolbitis appendiculata (WILLD.) K. IWATS. 


Loc.: 刺蕨蕨 - CiJue 

Bas.: Acrostichum appendiculatum WILLD. // Sp. Pl. 5: 114 (1810) 

Ety.: appendiculata (L: with small appendages or projections); 

relating to the conspicuous spines in the sinuses of the 

lamina margin 

Type: India orientalis, Klein 912, 1803 (HT: B) // India 

TW: Henry, A. (1896) 

(136) Bolbitis heteroclita (C. PRESL) CHING 


Loc.: 尾尾葉實實蕨蕨 - WeiYeShiJue 

Bas.: Acrostichum heteroclitum C. PRESL // Reliq. Haenk. 1: 15, pl. 2 

(1825) 

Ety.: heterokliton (G: different); relating to a (very) variable frond 

and lamina shape 

Type: Insula Sorzogon and Luzon, Haenke s. n. (HT: PR; IT: B, BR, 

W) // Philippines 

TW: Hayata, B. 1917 (1917) 

(137) Bolbitis heteroclita (C. PRESL) CHING x Bolbitis subcordata (COPEL.) 

CHING in C. CHR. * 


(138) Bolbitis x laxireticulata K. IWATS. 

Pub.: Acta Phytotax. Geobot. 18: 50-52, f. 7-8 (1959) 

Loc.: 網脈刺蕨蕨 - WangMaiCiJue 

Ety.: laxa (L: wide, loose, spacious), reticulata (L: made like a net); 

relating to the venation of sterile pinna 

Type: Ryukyus, Tagawa & Iwatsuki 2918 (HT: KYO; IT: KYO, L) // 

Japan 

TW: FOT 1975 (1975) 

(139) Bolbitis x nanjenensis C. M. KUO 

Pub.: Bot. Bull. Acad. Sin. n. s. 31(4): 308 (1990) 

Loc.: 南仁刺蕨蕨 - NanRenCiJue 

Ety.: relates to NanRenShan ("Nanjen"), a place in Taiwan and the 

type location 

Type: Pingtung, Tsu-lao-shyu-shan, Kuo & Yu 14856 (HT: TAI) // 


TW: Bot. Bull. Acad. Sin. n. s. 31(4): 308 (1990) 

(140) Bolbitis rhizophylla (KAULF.) HENNIPMAN [IUCN: NT] 

Pub.: Blumea 18: 148 (1970) 

Loc.: 大刺蕨蕨 - DaCiJue 

Bas.: Gymnogramma rhizophylla KAULF. // Enum. Filic. 78 (1824) 

35

36 

Ety.: rhiza (G: root), phyllon (G: leaf); relating to the habit (plantlets 

being developed on sterile lamina) 

Type: Manila, Chamisso s. n. (HT: ?LE; IT: B) // Philippines 

TW: FOT 1975 (1975) 

(141) Bolbitis scalpturata (FEE) CHING [IUCN: - (NT)] 


Loc.: 紅柄實實蕨蕨 - HongBingShiJue 

Bas.: Heteroneuron scalpturatum FEE // Mem. Foug. 2: 95 (1845) 

Ety.: scalpturata (L: engraved); relating to the grooved rachis 

Type: Manila, Gaudichaud s. n., Nov-1836 (LT: B; ILT: B, BM, P) // 

Philippines 

TW: Kuo, C. M. 1985 (1985) 

(142) Bolbitis subcordata (COPEL.) CHING in C. CHR. 


Loc.: 海南實實蕨蕨 - HaiNanShiJue 

Bas.: Campium subcordatum COPEL. // Philipp. J. Sci. 37: 369-370, 

pl. 23, f. 1 (1928) 

Ety.: sub (L: almost), cordata (L: with two equal rounded lobes at 

base); relating to base of sterile pinna 

Type: Hainan, McClure 9436 (IT: BISH, BM, C, MO, P) // China 

TW: FOT 1975 (1975) 

(143) Bolbitis virens var. compacta HENNIPMAN [IUCN: - (EN)] 

Pub.: Blumea 18: 149 (1970) 

Loc.: 厚厚葉實實蕨蕨 - HouYeShiJue 

Ety.: virens (L: becoming green), compacta (L: compact, close 

together); probably relating to the frond habit 

Type: Trang, Khao Chong, 600-1100 m alt., Tagawa 6802, 27-Jan- 

1966 (HT: L; IT: KYO) // Thailand 


BOTRYCHIUM (Ophioglossaceae) 

Ety.: botrys (G: bunch, cluster); relating to the fertile section appearing 

like a bunch of grapes 

(144) Botrychium formosanum TAGAWA [IUCN: NT (LC)] 


Loc.: 薄薄葉大陰地蕨蕨 - BoYeDaYinDiJue 


Type: Prov. Taito, near Baribugai, Taito-gun, Tagawa 2912, 5-Feb- 

1940 (HT: KYO) // Taiwan 


(145) Botrychium lanuginosum WALL. ex HOOK. & GREV. [IUCN: VU] 

Pub.: Icon. Filic. 1(4): pl. 79 (1828) 

Loc.: 阿里山蕨蕨萁 - ALiShanJueQi 

Ety.: lanuginosum (L: woolly); relating to the stipe base 

Type: Wallich 48 (BM, E, US) // Nepal

37 


(146) Botrychium lunaria (L.) SW. 

[IUCN: NT (LC)] 


Loc.: 扇羽陰地蕨蕨 - ShanYuYinDiJue 

Bas.: Osmunda lunaria L. // Sp. Pl. 2: 1064 (1753) 

Ety.: lunatus (L: like a half-moon); relating to the shape of sterile 

pinnae 

Type: Herb. Clifford 472, Osmunda n. 1 (LT: BM) 


(147) Botrychium ternatum (THUNB.) SW. 

[IUCN: EN] 


Loc.: 大陰地蕨蕨 - DaYinDiJue 

Bas.: Osmunda ternata THUNB. // Syst. Veg. (ed. 14): 927 (1784) 

Ety.: ternatum (L: consisting of threes); relating to the trophophore 

habit 

Type: Thunberg s. n. // Japan 

TW: Hayata, B. 1909 (1909) 

BRAINEA (Blechnaceae) 

Ety.: in honour of C. Braine (British botanist) 

(148) Brainea insignis (HOOK.) J. SM. 

[IUCN: VU] 

Pub.: Cat. Ferns Gard. Kew 5 (1856) 

Loc.: 蘇蘇鐵蕨蕨 - SuTieJue 

Bas.: Bowringia insignis HOOK. // Hooker's J. Bot. Kew Gard. Misc. 

5: 237-238, pl. 2 (1853) 

Ety.: insignis (L: outstanding, noted); relating to its habit (a splendid 

and outstanding plant) 

Type: Khasya // India 


CALYMMODON (Grammitidaceae) 

Ety.: kalymma (G: cover), odous (G: tooth); relating to the dents of pinnae 

being folded like a hood, enclosing the sporangia 

(149) Calymmodon cucullatus (NEES & BLUME) C. PRESL [IUCN: VU] 

Pub.: Tent. Pterid. 204 (1836) 

Loc.: 姬荷包蕨蕨 - JiHeBaoJue 

Bas.: Polypodium cucullatum NEES & BLUME // Nova Acta Regiae 

Soc. Sci. Upsal. 11: 121, pl. 12, f. 3 (1823) 

Ety.: cucullatus (L: hooded); relating to the shape of fertile pinnae 

Type: Java, Ripanna, Blume s. n. (LT: L) // Indonesia 

TW: Hayata, B. 1909 (1909) 

(150) Calymmodon gracilis (FEE) COPEL. [IUCN: NT] 

Pub.: Philipp. J. Sci. 34: 266-267 (1927) 

Loc.: 疏毛荷包蕨蕨 - ShuMaoHeBaoJue 

Bas.: Plectopteris gracilis FEE // Gen. 230, t. 19B (1850-52)

38 

Ety.: gracilis (L: slender); relating to the shape of frond 

Type: Luzon, Cuming 206 (B, BM, CGE, E, K, L, OXF) // Philippines 

TW: FOT 1975 (1975) 

CEPHALOMANES (Hymenophyllaceae) 

Ety.: kephale (G: head), manes (G: cup); relates to the shape of indusium 

(151) Cephalomanes apiifolium (C. PRESL) K. IWATS. 

Pub.: Acta Phytotax. Geobot. 35(4-6): 176 (1984) 

Loc.: 毛桿蕨蕨 - MaoGanJue 

Bas.: Trichomanes apiifolium C. PRESL // Hymenophyllaceae 16, 44 

(1843) 

Ety.: apium relating to flowering plant genus Apium (Apiaceae, 

"marshworts"), folium (L: leaf); relating to the similarity of 

fronds to leaves of members of genus Apium 

Alt.: Callistopteris apiifolia (C. PRESL) COPEL. // Philipp. J. Sci. 67: 

65 (1938) 

Type: Luzon, Cuming 137 (GH, K, MICH) // Philippines 

TW: Ito, H. 1944 (1944) 

(152) Cephalomanes clathratum (TAGAWA) K. IWATS. [IUCN: CR (EN)] 

Pub.: J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 13(5): 547 (1985) 

Loc.: 厚厚葉蕨蕨 - HouYeJue 

Bas.: Trichomanes clathratum TAGAWA // Acta Phytotax. Geobot. 8: 

164-165 (1939) 

Ety.: clathratum (L: latticed); relating to the venation of lamina 

Alt.: Abrodictyum clathratum (TAGAWA) EBIHARA & K. IWATS. // 

Blumea 51(2): 243 (2006) 

Type: Sinuiei, Tagawa 2293, 11-Feb-1939 (HT: KYO) // Taiwan 


(153) Cephalomanes cumingii (C. PRESL) K. IWATS. [IUCN: NT] 


Loc.: 長片蕨蕨 - ChangPianJue 

Bas.: Abrodictyum cumingii C. PRESL // Hymenophyllaceae 63, pl. 7 

(1843) 

Ety.: in honor of H. Cuming (British botanist and collector of type 

specimen) 

Alt. Abrodictyum cumingii C. PRESL // Hymenophyllaceae 63, pl. 7 

(1843) 

Type: Luzon, Cuming 208 (ST: B, E, BM, K, L) and Cuming 358 (ST: 

E, GH) // Philippines 

TW: Tagawa, M. 1940-49 (1940) 

(154) Cephalomanes grande (COPEL.) K. IWATS. [IUCN: - (CR)] 


Loc.: 大球毛桿蕨蕨 - DaQiuGanMaoJue 

Bas.: Trichomanes grande COPEL. // Philipp. J. Sci. 6: 70-71 (1911) 

Ety.: grande (L: large); relating to its habit

Alt.: Crepidomanes grande (COPEL.) K. IWATS. // Blumea 51(2): 239 

(2006) 

Type: San Ramon, Mindanao, alt. 3600 ft., Copeland 1739, Apr-1905 

(IT: P) // Philippines 

TW: FOT 1975 (1975) 

(155) Cephalomanes javanicum var. asplenioides (C. PRESL) K. IWATS. 

[IUCN: NT] 


Loc.: 菲律賓厚厚葉蕨蕨 - FeiLüBinHouYeJue 

Bas.: Trichomanes javanicum var. asplenioides (C. PRESL) C. CHR. // 

Index Filic. 635 (1906); Trichomanes asplenioides C. PRESL // 

Hymenophyllaceae 37-38 (1843) is illegitimate as it is a later 

homonym of Trichomanes asplenioides SW. in Prodr. 136 

(1788) 

Ety.: javanicum is relating to Java (Indonesia, type location), 

asplenium is relating to a fern genus (see there), oides (L: 

suffix, denoting likeness of form); relating to the similarity of 

fronds to that of members of genus (Hymen-) Asplenium (see 

there) 

Type: Java, Blume s. n. (IT: L) // Indonesia 

TW: a: Matsumara, J. & Hayata, B. (1906) (not as var. 

asplenioides) 

b: FOJ (1995) 

(156) Cephalomanes obscurum (BLUME) K. IWATS. 


Loc.: 線片長筒蕨蕨 - XianPianChangTongJue 

Bas.: Trichomanes obscurum BLUME // Enum. Pl. Javae 227 (1828) 

Ety.: obscurum (L: dusky, dark); relating to the lamina colour 

Alt.: Abrodictyum obscurum (BLUME) EBIHARA & K. IWATS. // Blumea 

51(2): 244 (2006) 

Type: Java, Zippel s. n. (LT: L), designated by Morton, 1974 // 

Indonesia 

TW: Kawakami, T. (1910) 

(157) Cephalomanes thysanostomum (MAKINO) K. IWATS. [IUCN: VU] 


Loc.: 球毛桿蕨蕨 - QiuMaoGanJue 

Bas.: Trichomanes thysanostomum MAKINO // Bot. Mag. (Tokyo) 13: 

46-47 (1899) 

Ety.: thysanos (G: fringe, tassel), stoma (G: mouth); relating to the 

involucre 

Alt.: Crepidomanes thysanostomum (MAKINO) EBIHARA & K. IWATS. 

// Blumea 51(2): 240 (2006) 

Type: Ryukyus, Yaeyama, Y. Tashiro s. n., Apr-1887 (LT: TI; ILT: TI), 

designated by Iwatsuki, J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 

13: 546 (1985) // Japan 

39

40 

TW: Yabe, Y. (1902) 

CERATOPTERIS (Pteridaceae) 

Ety.: keras (G: horn), pteris (relates to genus Pteris, or ferns in general); 

relating to the observed morphology ("horned fern") 

(158) Ceratopteris thalictroides (L.) BRONGN. 

Pub.: Bull. Sci. Soc. Philom. Paris 8: 186 (1821) 

Loc.: 水蕨蕨 - ShuiJue 

Bas.: Acrostichum thalictroides L. // Sp. Pl. 2: 1070 (1753) 

Ety.: thalictroides (G: like Thalictrum, a genus of flowering plants, 

Ranunculaceae, "meadow-rue"); relating to the shape of 

lamina 

Type: (LT: BM, Hermann Herb.), designated by Pichi-Sermolli, 

Webbia 12 (1957) // Sri Lanka 

TW: Henry, A. (1896) 

CHEILANTHES (Pteridaceae) 

Ety.: cheilos (G: lip), anthos (G: flower); relating to the position of 

sporangia beneath the lip-like false indusium 

(159) Cheilanthes agetae (SAIKI) C. M. KUO [IUCN: - (DD)] 

Pub.: Taiwania 30: 57 (1985) 

Loc.: 深山粉背蕨蕨 - ShenShanFenBeiJue 

Bas.: Aleuritopteris agetae SAIKI // J. Phytogeogr. Taxon. 32(1): 1 

(1984) 

Ety.: in honor of H. Ageta (Japanese botanist, who first suspected 

this to be a distinct taxon) 

Alt.: Aleuritopteris agetae SAIKI // J. Phytogeogr. Taxon. 32(1): 1 

(1984) 

Type: Alishan, Faurie 506 (HT: KYO) // Taiwan 

TW: J. Phytogeogr. Taxon. 32(1): 1 (1984) 

(160) Cheilanthes argentea (GMEL.) KUNZE 

Pub.: Linnaea 23: 242 (1850) 

Loc.: 長柄粉背蕨蕨 - ChangBingFenBeiJue 

Bas.: Pteris argentea GMEL. // Novi Comment. Acad. Sci. Imp. 

Petrop. 12: 519, pl. 12, f. 2 (1768) 

Ety.: argentea (L: silvery); relating to the farina on the abaxial 

lamina 

Alt.: Aleuritopteris argentea (GMEL.) FEE // Mem. Foug. 5: 154 

(1852) 

Type: Siberia // Russia 

TW: Hayata, B. 1917 (1917) 

(161) Cheilanthes chusana HOOK. 

Pub.: Sp. Fil. 2: 95, pl. 106B (1852) 

Loc.: 細葉碎米蕨蕨 - XiYeSuiMiJue 

Ety.: relating to "Chusan", a place in China and the type location

Type: Zhejiang, Chusan, Alexander s. n. (K) // China 


Cheilanthes mysurensis WALL. 

B: Kuo, C. M. 1985 (1985) 

(162) Cheilanthes concolor (LANGSD. & FISCH.) R. M. TRYON & A. F. TRYON 

Pub.: Rhodora 83(833): 133 (1981) 

Loc.: 黑心蕨蕨 - HeiXinJue 

Bas.: Pteris concolor LANGSD. & FISCH. // Icon. Filic. 19, pl. 21 (1810) 

Ety.: concolor (L: of the same colour); relating to similar colours of 

both sides of lamina 

Alt.: Doryopteris concolor (LANGSD. & FISCH.) KUHN // v. Decken, 

Reis. 3(3), Bot. 19 (1879) 

Type: Archipelagi Marquesas dicti insula Nucahiva, Langsdorf s. n. 

(HT: LE; IT: BM) // French Polynesia 

TW: Hayata, B. 1917 (1917) 

(163) Cheilanthes formosana HAYATA 

Pub.: Enum. Pl. Form. 612 (1906) 

Loc.: 台台灣粉背蕨蕨 - TaiWanFenBeiJue 


Alt.: Aleuritopteris formosana (HAYATA) TAGAWA // Acta Phytotax. 

Geobot. 14: 191 (1952) 

Type: Rakurakusha, Nakahara 203, Aug-1905 (HT: TI) // Taiwan 

TW: Enum. Pl. Form. 612 (1906) 

(164) Cheilanthes krameri FRANCH. & SAV. [IUCN: DD (LC)] 

Pub.: Enum. Pl. Jap. 2: 212, 619-620 (1879) 

Loc.: 粉背蕨蕨 - FenBeiJue 

Ety.: in honor of a person with family name Kramer (possibly C. 

Kramer, 1843-82, German plant collector) 

Alt.: Aleuritopteris krameri (FRANCH. & SAV.) CHING // Hong Kong 

Naturalist 10: 202 (1941) 

Type: Savatier 2234 (HT: P; IT: P) // Japan 

TW: Ogata, M. (1940) 

(165) Cheilanthes nitidula HOOK. 

Pub.: Sp. Fil. 2: 112 (1852) 

Loc.: 擬旱蕨蕨 - NiHanJue 

Ety.: 

Alt.: 

nitidula (L: shiny); relating to the stipe 

Mildella nitidula (HOOK.) C. C. HALL & LELLINGER // Amer. Fern 

J. 57: 127 (1967) 

Type: Kashmir, alt. 10000-11000 ped., T. Thomson s. n. (ILT: B) // 

India 

TW: Ito, H. 1944 (1944) 

(166) Cheilanthes nudiuscula (R. BR.) T. MOORE [IUCN: NT] 

Pub.: Index Fil. 249 (1861) 

Loc.: 毛碎米蕨蕨 - MaoSuiMiJue 

Bas.: Pteris nudiuscula R. BR. // Prodr. 155 (1810) 

41

42 

Ety.: nudiusculus (L: somewhat naked); relating to being less hairy 

than other species (as considered by the author) 

Type: N. Coast, R. Brown 60 (E) // Australia 

TW: MAN (2002) 

(167) Cheilanthes subargentea (CHING) C. M. KUO [IUCN: DD (LC)] 


Loc.: 擬長柄粉背蕨蕨 - NiChangBingFenBeiJue 

Bas.: Aleuritopteris subargentea CHING // Acta Phytotax. Sin. 19(1): 

65-66, pl. 2, f. 4-5 (1981) 

Ety.: sub (L: similar to), argentea (L: silvery); relating to the similarity 

to taxon Cheilanthes argentea (see there) 

Alt.: Aleuritopteris subargentea CHING // Acta Phytotax. Sin. 19(1): 

65-66, pl. 2, f. 4-5 (1981) 

Type: Yunnan, Feng 6059 (HT: PE) // China 

TW: Kuo, C. M. 1985 (1985) 

(168) Cheilanthes tenuifolia (BURM.) SW. 

Pub.: Syn. Fil. 129, 332 (1806) 

Loc.: 薄薄葉碎米蕨蕨 - BoYeSuiMiJue 

Bas.: Trichomanes tenuifolium BURM. // Fl. Indica 237 (1768) 

Ety.: tenuis (L: thin), folium (L: leaf); relating to the quality of lamina 

Type: Plantae Leylansia, Burman s. n. (G) // ?India 

TW: Henry, A. (1896) 

CHEIROPLEURIA 

Ety.: 

(Cheiropleuriaceae) 

cheir (G: hand), pleuron (G: rib); relating to the palmately-veined 

sterile fronds 

(169) Cheiropleuria integrifolia (D. C. EATON ex HOOK.) M. KATO, Y. YATABE, 

SAHASHI & N. MURAK. 

Pub.: Blumea 46(3): 522 (2001) 

Loc.: 燕尾尾蕨蕨 - YanWeiJue 

Bas.: Acrostichum bicuspe var. integrifolia D. C. EATON ex HOOK. // 

Sp. Fil. 5: 272 (1864) 

Ety.: integer (L: entire), folium (L: leaf); relating to the shape of 

lamina 

Type: Ryukyus, Wright s. n. (K) // Japan 

TW: J. Bot. 23: 107 (1885) 

CIBOTIUM (Dicksoniaceae) 

Ety.: kibotos (G: small chest or casket); relating to the form of the 

indusium 

(170) Cibotium barometz (L.) J. SM. 

Pub.: Lond. J. Bot. 1: 437 (1842) 

Loc.: 金狗毛蕨蕨 - JinGouMaoJue 

Bas.: Polypodium barometz L. // Sp. Pl. 2: 1092 (1753)

Ety.: baranets (Russian: diminutive of baran, lamb); relating to the 

woolly hairs covering plant parts 

Type: Linnaeus // China 

TW: a: Henry, A. (1896), based on material from Northern Taiwan 

thus probably a mis-identification of Cibotium taiwanense 

b: Matsumara & Hayata (1906), with one location from 

C Taiwan cited, however, I have not seen relevant material 

c: Hayata, B. 1917 (1917) 

(171) Cibotium taiwanense C. M. KUO 

Pub.: Taiwania 30: 56-57 (1985) 

Loc.: 台台灣金狗毛蕨蕨 - TaiWanJinGouMaoJue 


Type: Hsinchu, Chu-tong, Kuo 1703 (HT: TAI; IT: TAI) // Taiwan 

TW: Taiwania 30: 56-57 (1985) 

COLYSIS (Polypodiaceae) 

Ety.: kolysis (G: separation); relating to the discontinuous rows of sori 

(172) Colysis elliptica (THUNB.) CHING 

Pub.: Bull. Fan Mem. Inst. Biol. 4: 333 (1933) 

Loc.: 橢圓線蕨蕨 - TuoYuanXianJue 

Bas.: Polypodium ellipticum THUNB. // Syst. Veg. (ed. 14) 935 (1784) 

Ety.: elliptica (L: elliptic); relating to the shape of pinna 

Alt.: Leptochilus ellipticus (THUNB.) NOOT. // Blumea 42: 283-285, 

pl. 1 (f. 3), pl. 8 (f. 11 & 14) (1997) 


TW: J. Bot. 23: 103 (1885) 

(173) Colysis hemionitidea (C. PRESL) C. PRESL 

Pub.: Epimel. Bot. 147 (1849) 

Loc.: 斷線蕨蕨 - DuanXianJue 

Bas.: Selliguea hemionitidea C. PRESL // Tent. Pterid. 216, pl. 9, f. 17 

(1836) 

Ety.: two alternatives: (1) hemionitis relates to a fern genus 

(Pteridaceae); perhaps with fronds resembling members of 

genus Hemionitis (as understood by the author); (2) hemionitidea 

(L: barren, see Gledhill, D. 2008); relationship unclear 

to me 

Alt.: Leptochilus hemionitideus (C. PRESL) NOOT. // Blumea 42: 285 

(1997) 

Type: Wallich 284 (K, L, P) // Nepal 


(174) Colysis henryi (BAKER) CHING 

Pub.: Bull. Fan Mem. Inst. Biol. 4: 325-326 (1933) 

Loc.: 亨利氏線蕨蕨 - HengLiShiXianJue 

Bas.: Gymnogramma henryi BAKER // J. Bot. 25: 171 (1887) 


43

44 

Alt.: Leptochilus henryi (BAKER) X. C. ZHANG // Lycophytes Ferns 

China: 654 (2012) 

Type: Prov. Hupeh, Henry 7880, 1885-88 (IT: HUH) // China 

TW: Tagawa, M. 1940-49 (1942) 

(175) Colysis x shintenensis (HAYATA) H. ITO 

Pub.: J. Jap. Bot. 11: 90 (1935) 

Loc.: 新店線蕨蕨 - XinDianXianJue 

Bas.: Polypodium shintenense HAYATA // Icon. Pl. Formosan. 8: 154- 

155, f. 85-86 (1919) 

Ety.: relating to XinDian ("Shinten"), a place in Taiwan and type 

location 

Alt.: Leptochilus x shintenensis (HAYATA) X. C. ZHANG & NOOT. // Fl. 

China 2-3: 835 (2013) 

Type: Shinten, U. Faurie 215, Jul-1914 (HT: TI; IT: KYO) // Taiwan 

TW: Icon. Pl. Formosan. 8: 154-155, f. 85-86 (1919) 

(176) Colysis wrightii (HOOK.) CHING 


Loc.: 萊氏線蕨蕨 - LaiShiXianJue 

Bas.: Gymnogramma wrightii HOOK. // Sp. Fil. 5: 160, pl. 303 (1864) 

Ety.: in honor of C. H. Wright (US botanist and collector of the type 

specimen) 

Alt.: Leptochilus wrightii (HOOK.) X. C. ZHANG // Lycophytes Ferns 

China: 656 (2012) 

Type: Okinawa Prefecture; Ryukyu islands, C. H. Wright s. n. (IT: 

US) // Japan 

TW: J. Bot. 23: 103 (1885) 

CONIOGRAMME (Pteridaceae) 

Ety.: konis (G: dust), gramme (G: line); relating to the sporangia, which 

are not in round sori but in distinct brown lines following the veins 

(177) Coniogramme fraxinea (D. DON) FEE ex DIELS 

Pub.: Nat. Pflanzenfam. 1(4): 262 (1899) 

Loc.: 全緣鳳鳳ㄚ蕨蕨 - QuanYuanFengAJue 

Bas.: Diplazium fraxineum D. DON // Prodr. Fl. Nepal. 12-13 (1825) 

Ety.: fraxinea (L: ash-like); relating to the fronds resembling the 

leaves of the flower plant genus Fraxinus (Oleaceae, "ash") 

Type: Hamilton s. n. // Nepal 

TW: a: Matsumara, J. & Hayata, B. (1906), as Neurogramme 

fraxinea CHRIST, synonym of Cryptogramme aurata PRANTL 

b: Hayata, B. 1908 (1908) 

(178) Coniogramme intermedia HIERON. 

Pub.: Hedwigia 57: 301-307 (1916) 

Loc.: 華鳳鳳ㄚ蕨蕨 - HuaFengAJue

Ety.: intermedia (L: intermediate); relating to the habit (intermediate 

morphology between Coniogramme pubescens HIERON. and 

Coniogramme indica FEE) 

Type: Hakodate, Albrecht s. n., 1862 (LT: B), designated by T. 

Nakaike (1978) // Japan 

TW: FOT 1975 (1975) 

(179) Coniogramme japonica (THUNB.) DIELS 


Loc.: 日本鳳鳳ㄚ蕨蕨 - RiBenFengAJue 

Bas.: Hemionitis japonica THUNB. // Syst. Veg. (ed. 14) 932 (1784) 

Ety.: relating to Japan, the type location of this taxon 

Type: Japan 

TW: Hayata, B. 1911 (1911) 

(180) Coniogramme procera WALL. ex FEE 

Pub.: 

Mem. Soc. Sci. Nat. Strasbourg 6(1): 22 (1865), based on 

Grammitis procera WALL. (nom. nud.) in Numer. List n. 3 

(1828) 

Loc.: 高山鳳鳳ㄚ蕨蕨 - GaoShanFengAJue 

Ety.: procera (L: very tall, high); relating to its habit 

Type: Sheopore, Apr-1821 (HT: B) // Nepal 

TW: FOT 1975 (1975) 

CORNOPTERIS (Dryopteridaceae) 

Ety.: cornu (L: horn), pteris (relates to genus Pteris, or ferns in general); 

relating to the horn-like growths in the rachis groove where the 

pinnae are inserted 

(181) Cornopteris banajaoensis (C. CHR.) K. IWATS. & M. G. PRICE 

Pub.: SouthE. Asian Stud. 14(4): 564 (1977) 

Loc.: 大葉貞蕨蕨 - DaYeZhenJue 

Bas.: Dryopteris banajaoensis C. CHR. // Index Filic., Suppl. 1: 30 

(1913) 

Ety.: relating to "Banahao", a mountain in the Philippines and the 

type location 

Type: Mt. Banahao, Luzon, Copeland 2142 (HT: PNH, lost; IT: 

MICH) // Philippines 

TW: Kuo, C. M. 1985 (1985) 

(182) Cornopteris decurrentialata (HOOK.) NAKAI var. decurrentialata * [IUCN: 

- (DD)] 


Loc.: 貞蕨蕨 - ZhenJue 

Bas.: Gymnogramma decurrentialata HOOK. // Sp. Fil. 5: 142-143, pl. 

294 (1864) 

Ety.: decurrentis (L: running down), alata (L: winged); relating to the 

pinnae where a wing connects pinnules 

Type: Okosima, Oldham 374, 1863 (K) // Japan 

45

46 


(183) Cornopteris decurrentialata var. pilosella H. ITO 


Loc.: 毛葉貞蕨蕨 - MaoYeZhenJue 

Ety.: pilosella (L: tomentose, finely pelted with soft hairs); relating to 

the habit of axes 

Type: unclear, lectotypification necessary as protologue contains an 

ambiguous reference to two taxa: Cornopteris musashiensis 

NAKAI (type: Mt. Takao, Nakai s. n. (HT: TI) // Japan) and 

Cornopteris christenseniana TAGAWA (type: Isl. Quelpaert, 

Taquet 3933 (HT: KYO) // Korea), the latter being considered 

by several botanists as recognizable and distinct taxon 

TW: a: Hayata, B. 1917 (1917), no variety described but probably 

for var. pilosella 

b: FOT 1975 (1975) not describing explicitely any variety, but 

as abaxial axes are pilose hairy (as shown on Plate 161) 

described taxon is var. pilosella 

c: Knapp, R. 2013 (2013) 

(184) Cornopteris opaca (D. DON) TAGAWA 


Loc.: 黑葉貞蕨蕨 - HeiYeZhenJue 

Bas.: Hemionitis opaca D. DON // Prodr. Fl. Nepal. 13 (1825) 

Ety.: opaca (L: darkened, dull, not shining); relating to the adaxial 

lamina 

Type: Wallich // Nepal 

TW: Kuo, C. M. 1985 (1985) 

(185) Cornopteris philippinensis M. KATO [IUCN: DD] 

Pub.: Acta Phytotax. Geobot. 30(4-6): 110, f. 6 (1979) 

Loc.: 菲律賓貞蕨蕨 - FeiLüBinZhenJue 

Ety.: relating to the Philippines (type location) 

Type: Mindanao, Mt. Matutum, Copeland s. n., May-1917 (HT: 

MICH) // Philippines 

TW: Kuo, C. M. 1985 (1985) 

CREPIDOMANES (incl. Vandenboschia) (Hymenophyllaceae) 

Ety.: crepis (G: slipper), manes (G: cup); relating to the shape of sorus 

Vandenboschia: in honor of R. B. van den Bosch (Dutch botanist) 

(186) Crepidomanes acutum (C. PRESL) K. IWATS. [IUCN: - (CR)] 


Loc.: 稀毛毛葉蕨蕨 - XiMaoMaoYeJue 

Bas.: Trichomanes acutum C. PRESL // Hymenophyllaceae 134 

(1843) 

Ety.: acutum (L: pointed); relating to the apex of lobes (based on 

protologue)

Alt.: Hymenophyllum acutum (C. PRESL) EBIHARA & K. IWATS. // 

Blumea 51(2): 232 (2006) 

Type: Luzon, H. Cuming 219 (HT: E; IT: L, P, US) // Philippines 


(187) Crepidomanes auriculatum (BLUME) K. IWATS. 


Loc.: 瓶蕨蕨 - PingJue 

Bas.: Trichomanes auriculatum BLUME // Enum. Pl. Javae 225 

(1828) 

Ety.: auriculatum (L: furnished with ear-like appendages); relating to 


Alt.: Vandenboschia auriculata (BLUME) COPEL. // Philipp. J. Sci. 67: 

55 (1938) 

Type: Java, Blume s. n. (L) // Indonesia 

TW: Henry, A. (1896) 

(188) Crepidomanes bipunctatum (POIR.) COPEL. 


Loc.: 圓唇假脈蕨蕨 - YuanChunJiaMaiJue 

Bas.: Trichomanes bipunctatum POIR. // Encycl. 8: 69 (1808) 

Ety.: bi (L: two), punctatum (L: dotted, with spots); relating to the 

involucre (based on protologue) 

Type: Petit-Thouars s. n. (IT: ?P) // Madagascar 


(189) Crepidomanes birmanicum (BEDD.) K. IWATS. 


Loc.: 南海瓶蕨蕨 - NanHaiPingJue 

Bas.: Trichomanes birmanicum BEDD. // Suppl. Ferns Brit. Ind. 3, pl. 

349 (1876) 

Ety.: relating to Myanmar ("Birma", ancient form), the type location 

Alt.: Vandenboschia birmanica (BEDD.) CHING // Ching & Wang, 

Acta Phytotax. Sin. 8: 135 (1959) 

Type: Moulmein, Parish 181 (K) // Myanmar 

TW: Kuo, C. M. 1985 (1985) 

(190) Crepidomanes humile (G. FORST.) BOSCH 

Pub.: Versl. Meded. Kon. Akad. Wetensch., Afd. Natuurk. 9: 16, t. 11 

(1861) 

Loc.: 厚厚邊蕨蕨 - HouBianJue 

Bas.: Trichomanes humile G. FORST. // Fl. Ins. Austr. 8: 84 (1786) 

Ety.: humilis (L: small, on the ground); relating to the growth habit 

Type: Society Isl., Forster (BM, FR, K) // French Polynesia 

TW: Yabe, Y. (1902) 

(191) (Crepidomanes kalamocarpum comb. ined.) Vandenboschia 

kalamocarpa (HAYATA) EBIHARA 


Loc.: 華東瓶蕨蕨 - HuaDongPingJue 

47

48 

Bas.: Trichomanes kalamocarpum HAYATA // Icon. Pl. Formosan. 5: 

260, f. 93 (1915) 

Ety.: kalamos (G: stalk, reed), karpos (G: fruit); relating to the 

involucre habit 

Alt.: Vandenboschia kalamocarpa (HAYATA) EBIHARA // Acta 

Phytotax. Geobot. 60(1): 32 (2009) 

Type: Mt. Arisan, B. Hayata & S. Sasaki s. n., Jan-1912 (HT: TI; IT: 

TAIF) // Taiwan 

TW: Icon. Pl. Formosan. 5: 260, f. 93 (1915) 

(192) Crepidomanes kurzii (BEDD.) TAGAWA & K. IWATS. [IUCN: NT] 


Loc.: 假脈蕨蕨 - JiaMaiJue 

Bas.: Trichomanes kurzii BEDD. // Ferns Brit. India pl. 286 (1866) 

Ety.: in honor of W. S. Kurz (German botanist and collector of the 


Type: S. Andaman Isl., Kurz s. n. (BM, K) // India 


(193) Crepidomanes latealatum (BOSCH) COPEL. 


Loc.: 翅柄假脈蕨蕨 - ChiBingJiaMaiJue 

Bas.: Didymoglossum latealatum BOSCH // Ned. Kruidk. Arch. 5: 138 

(1863) 

Ety.: late (L: wide, broad), alatum (L: winged); relating to the 

obvious rachis wing 

Type: Assam, Griffith s. n. (K) // India 

TW: Ito, H. 1944 (1944) 

(194) Crepidomanes latemarginale (EATON) COPEL. [IUCN: NT] 


Loc.: 闊邊假脈蕨蕨 - KuoBianJiaMaiJue 

Bas.: Trichomanes latemarginale EATON // Proc. Amer. Acad. Arts 4: 

111 (1858) 

Ety.: late (L: wide, broad), marginalis (L: marginal); relating to the 

presence of an intramarginal vein pretending a broad lamina 

margin 

Type: Hongkong, Wright s. n. (B, GH, K, L, PE) // China 

TW: FOT 1975 (1975) 

(195) Crepidomanes latifrons (BOSCH) CHING [IUCN: DD (LC)] 

Pub.: Fl. Reipubl. Popularis Sin. 2: 172 (1959) 

Loc.: 寬寬葉瓶蕨蕨 - KuanYePingJue 

Bas.: Trichomanes latifrons BOSCH // Ned. Kruidk. Arch. 5: 209 

(1863) 

Ety.: lati (L: wide, broad), frons (L: leaf, frond); relating to the frond 

habit 

Type: Khasya, Anon., Herb. Hooker (K) // India 

TW: FOT 1975 (1975)

(196) Crepidomanes maximum (BLUME) K. IWATS. 


Loc.: 大葉瓶蕨蕨 - DaYePingJue 

Bas.: Trichomanes maximum BLUME // Enum. Pl. Javae 228 (1828) 

Ety.: maximum (L: greatest); relating to its habit 

Alt.: Vandenboschia maxima (BLUME) COPEL. // Philipp. J. Sci. 67: 

54 (1938) 

Type: Java, Blume s. n. (HT: L; IT: ?P) // Indonesia 

TW: Hayata, B. 1914-21 (1914) 

(197) Crepidomanes minutum (BLUME) K. IWATS. 

Pub.: J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 13(5): 524 (1985) [fide 

Fraser-Jenkins (1997)] 

Loc.: 團扇蕨蕨 - TuanShanJue 

Bas.: Trichomanes minutum BLUME // Enum. Pl. Javae 223 (1828) 

Ety.: minutus (L: very small, minute); relating to its habit 


TW: Ito, H. 1944 (1944) 

(198) Crepidomanes pallidum (BLUME) K. IWATS. [IUCN: VU] 


Loc.: 毛葉蕨蕨 - MaoYeJue 

Bas.: Trichomanes pallidum BLUME // Enum. Pl. Javae 225-226 

(1828) 

Ety.: pallidum (L: pale); relating to the lamina colour 

Alt.: Hymenophyllum pallidum (BLUME) EBIHARA & K. IWATS. // 

Blumea 51(2): 232 (2006) 

Type: Java, Blume s. n. (HT: L) // Indonesia 

TW: Henry, A. (1896) 

(199) Crepidomanes palmifolium (HAYATA) DEVOL [IUCN: DD (LC)] 

Pub.: Taiwania 14: 108-109, pl. 6 (1968) 

Loc.: 變葉假脈蕨蕨 - BianYeJiaMaiJue 

Bas.: Trichomanes palmifolium HAYATA // Icon. Pl. Formosan. 4: 

138-140, f. 78 (1914) 

Ety.: palma (L: palm of hand), folium (L: leaf); relating to the shape 

of lamina 

Type: Arisan, B. Hayata et S. Sasaki s. n., Jan-1912 (HT: TI) // 



(200) Crepidomanes parvifolium (BAKER) K. IWATS. [IUCN: - (CR)] 


Loc.: 纖小小單葉假脈蕨蕨 - XianXiaoShanYeJiaMaiJue 

Bas.: Hymenophyllum parvifolium BAKER // J. Linn. Soc., Bot. 9: 340, 

pl. 8, f. E (1867) 

Ety.: parva (L: little), folium (L: leaf); relating to the size of lamina 

Type: Moulmein, C. Parish s. n., Feb-1862 (IT: E) // Myanmar 


49

50 

(201) Crepidomanes rupicola (RACIB.) COPEL. [IUCN: - (VU)] 


Loc.: 石生假脈蕨蕨 - ShiShengJiaMaiJue 

Bas.: Trichomanes rupicola RACIB. // Pteridoph. Buitenzorg 24 

(1898) 

Ety.: rupes (L: rock), colo (L: to inhabit); relating to the growth habit 

(typically growing on rocks) 

Type: Java, Pantjar, M. Raciborski s. n. (IT: L, P) // Indonesia 


(202) Crepidomanes sp. 1 

[IUCN: - (CR)] 

Alt.: Vandenboschia sp. 


(203) Crepidomanes sp. 2 



(204) Crepidomanes vitiense (BAKER) BOSTOCK [IUCN: VU] 

Pub.: Fl. Australia 48: 706 (1998) 

Loc.: 斐濟假脈蕨蕨 - FeiJiJiaMaiJue 

Bas.: Trichomanes vitiense BAKER // J. Linn. Soc., Bot. 9: 338, t. 8, f. 

D (1866) 

Ety.: relating to the Fijis (type location) 

Type: Milne // Fiji 

TW: Taxon 53(4): 946 (2004) 

CRYPTOGRAMMA 

Ety.: 

(Pteridaceae) 

kryptos (G: hidden), gramme (G: line); relating to the soral line 

hidden by the reflexed lamina margin 

(205) Cryptogramma brunoniana WALL. ex HOOK. & GREV. 

Pub.: Numer. List n. 396 (1828), ex Hooker & Greville, Icon. Filic. 2: 

pl. 158 (1829) 

Loc.: 

Ety.: 

Type: 

TW: Hayata, B. 1909 (1909) 

(206) Cryptogramma stelleri (S. G. GMEL.) PRANTL 

Pub.: Bot. Jahrb. Syst. 3: 413 (1882) 

高山珠蕨蕨 - GaoShanZhuJue 

in honor of R. Brown (British botanist) 

Alpes summa Kamson, Wallich 396, 1826 (US) // India 

Loc.: 疏葉珠蕨蕨 - ShuYeZhuJue 

Bas.: Pteris stelleri S. G. GMEL. // Novi Comment. Acad. Sci. Imp. 

Petrop. 12: 519, pl. 12, f. 1 (1768) 

Ety.: in honor of G. W. Steller (German biologist) 

Type: Siberia // Russia 

TW: Tagawa, M. 1940-49 (1940) 

CTENITIS (Dryopteridaceae) 

Ety.: kteis (G: comb); relating to the narrow fertile pinna lobes that 

resemble the teeth of a comb

(207) Ctenitis eatonii (BAKER) CHING 

Pub.: Bull. Fan Mem. Inst. Biol., Bot. 8: 291 (1938) 

Loc.: 愛德氏肋毛蕨蕨 - AiDeShiLeMaoJue 

Bas.: Nephrodium eatonii BAKER // Syn. Fil. 276 (1867) 

Ety.: in honor of D. C. Eaton (US botanist) 

Type: Ryukyu Island, Wright 1853-56 // Japan 

TW: Yabe, Y. (1902) 

(208) Ctenitis subglandulosa (HANCE) CHING 


Loc.: 肋毛蕨蕨 - LeMaoJue 

Bas.: Alsophila subglandulosa HANCE // Ann. Sci. Nat., Bot., ser. 5, 

6: 253 (1866) 

Ety.: sub (L: almost, litle), glandulosa (L: glandular); relating to the 

lamina (see protologue) 

Type: Tamsui, Oldham 6 (HT: BM, Herb. Hance 11186; IT: GH) // 


TW: Ann. Sci. Nat., Bot., ser. 5, 6: 253 (1866) 

CYATHEA (Cyatheaceae) 

Ety.: kyathos (G: wine cup); relating to the shape of indusium 

(209) Cyathea fenicis COPEL. [IUCN: - (NT)] 


Loc.: 蘭蘭嶼筆筒樹 - LanYuBiTongShu 

Ety.: in honor of E. Fenix (Philippine botanist and collector of the 


Alt.: Alsophila fenicis (COPEL.) C. CHR. // Index Filic., Suppl. 5 

(1913) 

Type: Batan Isl., Fenix BS 3797 (HT: US; IT: P) // Philippines 

TW: FOT 1975 (1975) 

(210) Cyathea hancockii COPEL. [IUCN: NT] 


Loc.: 韓氏桫欏 - HanShiSuoLuo 


specimen) 

Alt.: Alsophila denticulata BAKER // J. Bot. 23: 102-103 (1885) 

Type: near Tamsui, Hancock (HT: K; IT: ?US), type of Alsophila 

denticulata BAKER in J. Bot. 23: 102 (1885) // Taiwan 

TW: a: J. Bot. 23: 102-103 (1885) for Alsophila denticulata BAKER 

b: Philipp. J. Sci. 4: 37 (1909) 

(211) Cyathea lepifera (J. SM. ex HOOK.) COPEL. 


Loc.: 筆筒樹 - BiTongShu 

Bas.: Alsophila lepifera J. SM. ex HOOK. // Sp. Fil. 1: 54-55 (1846) 

Ety.: lepis (G: scale), fer (suffix, meaning -carrying); relating to axes 

of fronds, which are all very scaly 

51

52 

Alt.: Sphaeropteris lepifera (J. SM. ex HOOK.) R. M. TRYON // Contr. 

Gray Herb. 200: 21 (1970) 

Type: Luzon, Cuming 180 (HT: K; IT: A, BM, BO, F, K) // Philippines 


(212) Cyathea loheri CHRIST [IUCN: NT] 

Pub.: Bull. Herb. Boissier, ser. 2, 6: 1007-1008 (1906) 

Loc.: 南洋桫欏 - NanYangSuoLuo 

Ety.: in honor of A. Loher (German botanist and collector of the type 

specimen) 

Alt.: Alsophila loheri (CHRIST) R. M. TRYON // Contr. Gray Herb. 200: 

32 (1970) 

Type: Luzon, Mt. Banajao, Loher s. n., 7-Jan-1906 (BM, S) // 

Philippines 

TW: FOT 1975 (1975) 

(213) Cyathea metteniana (HANCE) C. CHR. & TARDIEU 

Pub.: Bull. Mus. Natl. Hist. Nat., ser. 2, 6: 450 (1934) 

Loc.: 台台灣樹蕨蕨 - TaiWanShuJue 

Bas.: Alsophila metteniana HANCE // J. Bot. 6: 175 (1868) 

Ety.: in honor of G. H. Mettenius (German botanist) 

Alt.: Alsophila metteniana HANCE // J. Bot. 6: 175 (1868) 

Type: Prov. Tokien, C. de Grijs s. n. (B) // China 

TW: FOT 1975 (1975) 

(214) Cyathea podophylla (HOOK.) COPEL. 


Loc.: 鬼桫欏 - GuiSuoLuo 

Bas.: Alsophila podophylla HOOK. // Hooker's J. Bot. Kew Gard. 

Misc. 9: 334 (1857) 

Ety.: pous (G: foot), phyllon (G: leaf); possibly relating to the habit 

(with fronds developing at the apical stem, though this is 

characteristic for a large number of taxa of this genus), or the 

frond shape 

Alt.: Alsophila podophylla HOOK. // Hooker's J. Bot. Kew Gard. 

Type: 

Misc. 9: 334 (1857) 

(a) Chusan, T. Alexander s. n. (ST: ?K) // ?Korea 

(b) Hong Kong, J. C. Bowring s. n. // China 


(215) Cyathea spinulosa WALL. ex HOOK. 

Pub.: Sp. Fil. 1: 25, pl. 12C (1846) 

Loc.: 台台灣桫欏 - TaiWanSuoLuo 

Ety.: spinulosa (L: with spines); relating the the stipe base and trunk 

Alt.: Alsophila spinulosa (WALL. ex HOOK.) R. M. TRYON // Contr. 

Gray Herb. 200: 32 (1970) 

Type: Wallich 178 (E) // Nepal 

TW: Yabe, Y. (1902)

CYCLOSORUS (incl. Dictyocline) (Thelypteridaceae) * 

Ety.: kyklos (G: circular), soros (G: sorus); relating to the shape of sorus 

Dictyocline: dictyon (G: net), kline (G: bed); relating to the sporangia 

that rest upon the surface of the netted veins 

(216) Cyclosorus acuminatus (HOUTT.) NAKAI 

Pub.: Misc. Pap. Japan. Pl. 15 (1935) 

Loc.: 毛蕨蕨 - MaoJue 

Bas.: Polypodium acuminatum HOUTT. // Nat. Hist. 2: 181, pl. 99, f. 2 

(1783) 

Ety.: acuminatus (L: acuminate); relating to the pinna apex 

Type: Thunberg s. n. (UPS) // Japan 

TW: Ito, H. 1944 (1944) 

(217) Cyclosorus aridus (D. DON) TAGAWA [IUCN: NT (LC)] 

Pub.: Bull. Fan Mem. Inst. Biol., Bot. 8: 194-196 (1938) 

Loc.: 擬毛蕨蕨 - NiMaoJue 

Bas.: Aspidium aridum D. DON // Prodr. Fl. Nepal. 4 (1825) 

Ety.: aridus (L: dry, withered); relating to the typical habitat, or the 

sensation when touching the rough lamina 

Type: Wallich s. n. (?BM or BR) // Nepal 

TW: Ito, T. (1928) 

(218) Cyclosorus auriculatus (J. SM.) C. M. KUO 

Pub.: Taiwania 47(2): 171 (2002) 

Loc.: 耳羽鉤毛蕨蕨 - ErYuGouMaoJue 

Bas.: Phegopteris auriculata J. SM. // Hist. Fil. 233 (1875) 

Ety.: auriculatus (L: furnished with ear-like appendages); relating to 


Type: Wallich 314 (K) // Nepal 

TW: FOT 1975 (1975) 

(219) Cyclosorus dentatus (FORSSK.) CHING 

Pub.: Bull. Fan Mem. Inst., Bot. 8: 206-209 (1938) 

Loc.: 野毛蕨蕨 - YeMaoJue 

Bas.: Polypodium dentatum FORSSK. // Fl. Aegypt.-Arab. 185 (1775) 

Ety.: dentatus (L: toothed); relating to the lamina (see protologue) 

Type: Forsskal s. n. (HT: C) // Yemen 

TW: FOT 1975 (1975) 

(220) Cyclosorus dictyoclinoides (CHING) C. M. KUO [IUCN: EN (VU)] 


Loc.: 溪邊蕨蕨 - XiBianJue 

Bas.: Stegnogramma dictyoclinoides CHING // Sinensia 7: 92-93, t. 1 

(1936) 

Ety.: dictyocline relating to a fern genus (Thelypteridaceae), oides 

(L: suffix, denoting likeness of form); relating to the similarity 

of fronds to that of genus Dictyocline (see under Cyclosorus) 

Type: Yunnan // China 

TW: Tagawa, M. 1940-49 (1940) 

53

54 

(221) Cyclosorus ensifer (TAGAWA) W. C. SHIEH [IUCN: DD (LC)] 

Pub.: J. Sci. Engin. 13: 42 (1976) 

Loc.: 廣葉毛蕨蕨 - GuangYeMaoJue 

Bas.: Dryopteris ensifera TAGAWA // Acta Phytotax. Geobot. 6: 89-90 

(1937) 

Ety.: ensis (L: sword), fer (suffix, meaning -carrying); relating to the 

Type: 

habit of lamina 

Kusukusu in Takao Province, Tagawa 943, 21-Sep-1934 (HT: 

KYO; IT: KYO) // Taiwan 


(222) Cyclosorus erubescens (WALL. ex HOOK.) C. M. KUO 


Loc.: 

Bas.: 

Ety.: 

方桿蕨蕨 - FangGanJue 

Polypodium erubescens WALL. // Numer. List n. 330 (1828), ex 

Hooker, Sp. Fil. 4: 236 (1862) 

erubescens (L: reddening); relating to the colour of axes (see 

protologue) 

Wallich 330 (IT: UC) // Nepal 

Type: 

TW: Hayata, B. 1914-21 (1915) 

(223) Cyclosorus esquirolii (CHRIST) C. M. KUO 


Loc.: 假毛蕨蕨 - JiaMaoJue 

Bas.: Dryopteris esquirolii CHRIST // Bull. Acad. Int. Geogr. Bot. 144- 

145 (1907) 

Ety.: in honor of J. H. Esquirol (French botanist and collector of the 


Type: Guizhou, Guiyang, Esquirol 903 (P) // China 

TW: FOT 1975 (1975) 

(224) Cyclosorus griffithii (T. MOORE) C. M. KUO 


Loc.: 聖蕨蕨 - ShengJue 

Bas.: Dictyocline griffithii T. MOORE // Index Fil. 59 (1857) 

Ety.: in honor of W. Griffith (British botanist) 

Type: Assam, Griffith s. n. (K) // India 

TW: J. Bot. 23: 107 (1885) 

(225) Cyclosorus gymnopteridifrons (HAYATA) C. M. KUO 


Loc.: 大羽新月蕨蕨 - DaYuXinYueJue 

Bas.: Dryopteris gymnopteridifrons HAYATA // Icon. Pl. Formosan. 8: 

148-149, f. 75-76 (1919) 

Ety.: gymnopteris relating to a fern genus (Pteridaceae), frons (L: 

leaf, frond); relating to the similarity of fronds to that of some 

taxa of genus Gymnopteris (G: gymnos, naked, and pteris 

relating to genus Pteris, or ferns in general) 

Type: 

Kusukusu, T. Soma s. n., 1912 (HT: TI; IT: TAI) // Taiwan


(226) Cyclosorus insularis (K. IWATS.) C. M. KUO [IUCN: DD] 


Loc.: 變葉新月蕨蕨 - BianYeXinYueJue 

Bas.: Abacopteris insularis K. IWATS. // Acta Phytotax. Geobot. 18: 6- 

8, f. 5 (1959) 

Ety.: insularis (L: pertaining to islands); relating to habitat (based on 

protologue) 

Type: Ryukyu, Isl. Okinoerabu, Ooyama, ca. 200 m, Tagawa & 

Iwatsuki 2248 (HT: KYO; IT: E, U, UC) // Japan 

TW: FOT 1975 (1975) 

(227) Cyclosorus interruptus (WILLD.) H. ITO 


Loc.: 鐵毛蕨蕨 - TieMaoJue 

Bas.: Pteris interrupta WILLD. // Phytographia 13, pl. 10, f. 1 (1794) 

Ety.: interruptus (L: interrupted); relating to the absence of sori at 

base of segments (based on protologue) 

Type: S. India, Klein s. n. (B) // India 

TW: FOT 1975 (1975) 

(228) Cyclosorus jaculosus (CHRIST) H. ITO 

Pub.: Bot. Mag. (Tokyo) 51: 725, f. 4 (1937) 

Loc.: 小小毛蕨蕨 - XiaoMaoJue 

Bas.: Aspidium jaculosum CHRIST // Bull. Herb. Boissier, ser. 2, 4: 

615 (1904) 

Ety.: jaculus (L: javelin); perhaps relating to the long narrow frond 

tapered to both ends 

Type: Kushaku, Faurie 646 (LT: P) // Taiwan 

TW: Bull. Herb. Boissier, ser. 2, 4: 615 (1904) 

(229) Cyclosorus latipinnus (BENTH.) TARDIEU [IUCN: DD (NT)] 

Pub.: Notul. Syst. (Paris) 7: 73 (1938) 

Loc.: 微縮小小毛蕨蕨 - WeiSuoXiaoMaoJue 

Bas.: Aspidium molle var. latipinnum BENTH. // Fl. Hongk. 455 (1861) 

Ety.: lati (L: wide, broad), pinnus (L: pinna); relating to the pinna 

habit 

Type: Hongkong, Hance 135 (LT: K, Morton 1974) // China 


(230) Cyclosorus liukiuensis (CHRIST) MASAM. 

Pub.: Sc. Rep. Kanazawa Univ. 1: 195 (1951) 

Loc.: 頂芽新月蕨蕨 - DingYaXinYueJue 

Bas.: Meniscium liukiuense CHRIST // Bot. Mag. (Tokyo) 24: 240 

(1910) 

Ety.: relating to the Ryukyu Islands (Japan) 

Type: Liukiu, Onnah, Matsamura 150 (HT: P) // Japan 

TW: Ito, H. 1944 (1944) 

55

56 

(231) Cyclosorus longipetiolatus (K. IWATS.) C. M. KUO [IUCN: CR (EN)] 


Loc.: 長柄新月蕨蕨 - ChangBingXinYueJue 

Bas.: Abacopteris longipetiolata K. IWATS. // Acta Phytotax. Geobot. 

18: 11-13, f. 6 (1959) 

Ety.: longus (L: long), petiolatus (L: with stipe); relating to the 

presence of obvious stipe at pinna base (see protologue) 

Type: Tagawa 2593 (HT: KYO; IT: KYO) // Taiwan 

TW: Acta Phytotax. Geobot. 18: 11-13, f. 6 (1959) 

(232) Cyclosorus megacuspis (BAKER) TARDIEU & C. CHR. [IUCN: - (EN)] 


Loc.: 微紅新月蕨蕨 - WeiHongXinYueJue 

Bas.: Polypodium megacuspe BAKER // J. Bot. 28: 266 (1890) 

Ety.: megas (G: great), cuspis (L: point); relating to the long caudate 

pinna apex 

Type: Tonkin: Mt. Bani, Balansa 47, 1890 (HT: K; IT: P) // Vietnam 


(233) Cyclosorus omeiensis (BAKER) C. M. KUO 


Loc.: 狹基鉤毛蕨蕨 - XiaJiGouMaoJue 

Bas.: Polypodium omeiense BAKER // J. Bot. 26: 229 (1888) 

Ety.: relating to "Omei", a mountain in China and the type location 

Type: Sichuan, Omei Shan, Faber 1059 (HT: K; IT: BM, MO) // China 

TW: Tagawa, M. 1940-49 (1940) 

(234) Cyclosorus papilio (C. HOPE) CHING [IUCN: DD (LC)] 


Loc.: 薄薄葉梳毛蕨蕨 - BoYeShuMaoJue 

Bas.: Nephrodium papilio C. HOPE // J. Bombay Nat. Hist. Soc. 12: 

625-628, pl. 12 (1899) 

Ety.: 

Type: 

papilio (L: butterfly); relating to the shape of basal pinnae 

NE India, below Darjeeling, 1200 m, Levinge s. n., 1880 (LT: 

K), designated by Holttum, Kew Bulletin 31(2): 322 (1976) // 

India 

TW: Tagawa, M. 1940-49 (1940) 

(235) Cyclosorus parasiticus (L.) FARW. 

Pub.: Amer. Midl. Naturalist 12: 259 (1931) 

Loc.: 密密毛毛蕨蕨 - MiMaoMaoJue 

Bas.: Polypodium parasiticum L. // Sp. Pl. 2: 1090 (1753) 

Ety.: parasiticus (L: parasitic); relating to the description of growth 

habit in the protologue ("on trees") 

Type: Guangdong, Guangzhou, Osbeck s. n. (HT: S; IT: S) // China 


(236) Cyclosorus parishii (BEDD.) TARDIEU ex TARDIEU & C. CHR. 


Loc.: 羽葉新月蕨蕨 - YuYeXinYueJue

Bas.: Meniscium parishii BEDD. // Ferns Brit. India pl. 184 (1866) 

Ety.: in honor of C. S. Parish (collector of the type specimen) 

Type: Moulmein, Parish 135 (K) // Myanmar 

TW: FOT 1975 (1975) 

(237) Cyclosorus pozoi (LAG.) C. M. KUO [IUCN: EN] 


Loc.: 非洲伏蕨蕨 - FeiZhouFuJue 

Bas.: Hemionitis pozoi LAG. // Gen. Sp. Pl. 33 (1816) 

Ety.: in honor of D. G. del Pozo (Spanish botanist, and collector of 


Type: N. Spain, D. G. del Pozo (HT: SPA) // Spain 

TW: FOT 1975 (1975) 

(238) Cyclosorus productus (KAULF.) CHING [IUCN: - (NT)] 


Loc.: 蘭蘭嶼大葉毛蕨蕨 - LanYuDaYeMaoJue 

Bas.: Aspidium productum KAULF. // Enum. Fil. Chamisso: 237-238 

(1824) 

Ety.: productus (L: extended, elongated); relating to the attenuated 

frond shape 

Type: Luzon, Cuming 10 (K) // Philippines 

TW: Kuo, C. M. 1985 (1985) 

(239) Cyclosorus proliferus (RETZ.) TARDIEU 


Loc.: 星毛蕨蕨 - XingMaoJue 

Bas.: Hemionitis prolifera RETZ. // Observ. Bot. 6: 36, 38 (1791) 

Ety.: proles (L: offsprings), fero (L: to bear); relating to the 

development of plantlets on the rachis 

Type: S. India, Koenig s. n. (?S-P) // India 

TW: Hayata, B. 1914-21 (1914) 

(240) Cyclosorus x pseudoliukiuensis (SERIZ.) RALF KNAPP 

Pub.: Ferns Fern Allies Taiwan 445 (2011) 

Loc.: 擬琉球新月蕨蕨 - NiLiuQiuXinYueJue 

Bas.: Thelypteris x pseudoliukiuensis SERIZ. // J. Phytogeogr. Taxon. 

29(1): 25 (1981) 

Ety.: pseudos (G: false), liukiuensis relating to a taxon with epithet 

"liukiuensis"; relating to the similarity to taxon Cyclosorus 

liukiuensis (see there) 

Type: Mt. Komi-dake, alt. ca. 90 m, Serizawa 14678, 9-Aug-1971 

(HT: AICH) // Japan 


(241) Cyclosorus simplex (HOOK.) COPEL. 

Pub.: Gen. Fil. (Copeland) 143 (1947) 

Loc.: 單葉新月蕨蕨 - DanYeXinYueJue 

Bas.: Meniscium simplex HOOK. // London. J. Bot. 1: 294, pl. 11 

(1842) 

57

58 

Ety.: simplex (L: simple, undivided); relating to the lamina 

Type: Hongkong, Hinds s. n., 1841 (HT: K) // China 

TW: Henry, A. (1896) 

(242) Cyclosorus taiwanensis (C. CHR.) H. ITO 

Pub.: Bot. Mag. (Tokyo) 51: 728, f. 2 (1937) 

Loc.: 台台灣毛蕨蕨 - TaiWanMaoJue 

Bas.: Dryopteris taiwanensis C. CHR. // Index Filic. 297 (1905) 


Type: replaces Aspidium lobulatum Christ in Bull. Herb. Boissier, ser. 

2, 4(7): 614-615 (1904): U. Faurie 642 (P) // Taiwan 

TW: Index Filic. 297 (1905) 

(243) Cyclosorus tottoides (H. ITO) C. M. KUO 


Loc.: 尾尾葉伏蕨蕨 - WeiYeFuJue 

Bas.: Leptogramma tottoides H. ITO // Bot. Mag. (Tokyo) 49: 434-435 

(1935) 

Ety.: totta is relating to the taxon with epithet "totta" (refering to the 

Khoikhoi, SW-African natives, labelled by the Dutch the 

"Hottentots"), oides (L: suffix, denoting likeness of form); 

relating to the similarity to Leptogramma totta (SCHLTDL.) J. 

SM. 

Type: Mt. Arisan, Hayata & Sasaki s. n., 1912 (TI) // Taiwan 

TW: Bot. Mag. (Tokyo) 49: 434-435 (1935) 

(244) Cyclosorus triphyllus (SW.) TARDIEU 


Loc.: 三葉新月蕨蕨 - SanYeXinYueJue 

Bas.: Meniscium triphyllum SW. // J. Bot. (Schrader) 1800(2): 16 

(1801) 

Ety.: tri (L: three), phyllon (G: leaf); relating to the lamina shape 

Type: (SPA) // China 

TW: J. Bot. 23: 103 (1885) 

(245) Cyclosorus truncatus (POIR.) FARW. 


Loc.: 大葉毛蕨蕨 - DaYeMaoJue 

Bas.: Polypodium truncatum POIR. // Encycl. 5: 534 (1804) 

Ety.: truncatus (L: truncate); relating to the habit of segment apex 

Type: "Brezil", no collector cited (P) 

TW: Henry, A. (1896) 

(246) Cyclosorus tylodes (KUNZE) PANIGRAHI [IUCN: NT (EN)] 

Pub.: Res. J. Pl. Environm. 9: 67 (1993) 

Loc.: 疣狀假毛蕨蕨 - YouZhuangJiaMaoJue 

Bas.: Aspidium tylodes KUNZE // Linnaea 24: 281 (1851) 

Ety.: tylos (G: knot, knob); relating to the projections 

(pneumatophores) at pinna base 

Type: S. India, Nilgiri Hills, Schmid-Koch 11, 63, 89, 115 // India

TW: Kuo, C. M. 1985 (1985) 

(247) (Cyclosorus wilfordii comb. ined.) Dictyocline wilfordii (HOOK.) J. SM. 

Pub.: Hist. Fil. 149 (1875) 

Loc.: 威氏聖蕨蕨 - WeiShiShengJue 

Bas.: Hemionitis wilfordii HOOK. // Fil. Exot. pl. 93 (1859) 

Ety.: in honor of C. Wilford (British Botanist and collector of the type 

specimen) 

Type: Wilford s. n. // Taiwan 

TW: Fil. Exot. pl. 93 (1859) 

CYSTOPTERIS (Dryopteridaceae) 

Ety.: kystos (G: bladder), pteris (relates to genus Pteris, or ferns in 

general); relating to the inflated indusium 

(248) Cystopteris fragilis (L.) BERNH. 

Pub.: Neues J. Bot. 1: 26, t. 2, f. 9 (1805) 

Loc.: 冷蕨蕨 - LengJue 

Bas.: Polypodium fragile L. // Sp. Pl. 2: 1091 (1753) 

Ety.: fragilis (L: fragile); relating to the quality of frond 

Type: "Habitat [in collibus Europae frigidioris.]", J. Ammann 51 (LT: 

LINN), designated by Copeland in Univ. Calif. Publ. Bot. 16 : 

56 (1929) // Europe 

TW: Ito, H. 1944 (1944) 

(249) Cystopteris moupinensis FRANCH. 

Pub.: Nouv. Arch. Mus. Hist. Nat., ser. 2, 10: 111 (1887) 

Loc.: 寬寬葉冷蕨蕨 - KuanYeLengJue 

Ety.: relating to Moupine (BaoXing in SiChuan Province), a place in 

China and the type location 

Type: Sichuan, M. David s. n., 1870 (HT: PE) // China 


DAVALLIA (Davalliaceae) 

Ety.: in honour of E. Davall (Swiss botanist) 

(250) Davallia chrysanthemifolia HAYATA [IUCN: NT] 


Loc.: 阿里山陰石蕨蕨 - ALiShanYinShiJue 

Ety.: chrysanthemum relating to a flowering plant genus 

(Asteraceae, "chrysanth"), folium (L: leaf); relating to the 

shape of lamina 

Type: Mt. Arisan, inter Mingetzu et Senninbora, T. Ito 27, Apr-1914 

(HT: TI) // Taiwan 


(251) Davallia cumingii HOOK. 

Pub.: Sp. Fil. 1: 155, pl. 45B (1846) 

Loc.: 鱗葉陰石蕨蕨 - LinYeYinShiJue 

59

60 

Ety.: in honor of H. Cuming (British botanist and collector of the type 

specimen 

Type: Samar, Cuming 138 (A, BM, K, L, P) // Philippines 


(252) Davallia formosana HAYATA 


Loc.: 大葉骨碎補 - DaYeGuSuiBu 


Type: Taichu, Kashigatani, Nakahara s. n., Feb-1907 (HT: TI) // 



(253) Davallia griffithiana HOOK. 

Pub.: Sp. Fil. 1: 168-169, pl. 49B (1846) 

Loc.: 杯狀蓋陰石蕨蕨 - BeiZhuangGaiYiShiJue 

Ety.: in honor of W. Griffith (British botanist, collector of type 

specimen) 

Type: Mishmee, Griffith 910 (HT: K; IT: GH, L) // India 

TW: J. Bot. 23: 103 (1885) 

(254) Davallia pectinata J. SM. 

[IUCN: EN (?CR)] 

Pub.: Mem. Acad. Roy. Sci. (Turin) 5: 415 (1793) 

Loc.: 馬來陰石蕨蕨 - MaLaiYinShiJue 

Ety.: pectinata (L: with narrow, close divisions like a comb); relating 

to the shape of lamina 

Type: India orientalis, Hurloch 1786 // India 

TW: Liew, F. S. (1976) 

(255) Davallia repens (L. F.) KUHN [IUCN: NT] 

Pub.: Filic. Afr. 27 (1868) 

Loc.: 陰石蕨蕨 - YinShiJue 

Bas.: Adiantum repens L. F. // Suppl. Pl. 446 (1781) 

Ety.: repens (L: creeping); relating to the habit 

Type: Mascareignes, Ile de France, Sonnerat par Thouin 

(Commerson) 74 (HT: P; IT: L) // Mascarene Islands 

TW: Henry, A. (1896) 

(256) Davallia solida (G. FORST.) SW. 


Loc.: 闊葉骨碎補 - KuoYeGuSuiBu 

Bas.: Trichomanes solidum G. FORST. // Fl. Ins. Austr. 86 (1786) 

Ety.: solida (L: solid, firm); relating to the lamina texture 

Type: Pacific Islands, Forster 308 (HT: BM; IT: P) 

TW: Henry, A. (1896) 

(257) Davallia trichomanoides BLUME 


Loc.: 海州骨碎補 - HaiZhouGuSuiBu

61 

Ety.: trichomanes relating to a fern genus (Hymenophyllaceae), 

oides (L: suffix, denoting likeness of form); relating to the 

frond habit 


TW: MAN (2002) 

(258) Davallia tyermanii (T. MOORE) BAKER [IUCN: - (VU)] 

Pub.: Syn. Fil. (ed. 2): 467 (1874), as 'tyermanni' 

Loc.: 圓蓋陰石蕨蕨 - YuanGaiYinShiJue 

Bas.: Humata tyermanii T. MOORE // Gard. Chron. 1871: 870-871 

(1871) 

Ety.: in honor of J. S. Tyerman (British botanist), collector of the 

type specimen 

Type: ?Herb. Moore (K) 

TW: Kuo, C. M. 1997 (1997) 

DENNSTAEDTIA (Dennstaedtiaceae) 

Ety.: in honour of A. W. Dennstaedt (German botanist) 

(259) Dennstaedtia hirsuta (THUNB. ex SW.) METT. ex MIQ. 

Pub.: Ann. Mus. Bot. Lugduno-Batavi 3: 181 (1867) 

Loc.: 細毛碗蕨蕨 - XiMaoWanJue 

Bas.: Davallia hirsuta SW. // J. Bot. (Schrader) 1800(2): 87 (1801), 

replacing the illeg. Trichomanes hirsutum THUNB. in 

Thunberg, Fl. Jap. 339 (1784) 

Ety.: hirsuta (L: hairy); relating to the presence of hairs on all plant 

parts 


TW: Ito, H. 1944 (1944) 

(260) Dennstaedtia scabra (WALL. ex HOOK.) T. MOORE 


Loc.: 碗蕨蕨 - WanJue 

Bas.: Dicksonia scabra WALL. // Numer. List n. 2173 (1829), ex 

Hooker, Sp. Fil. 1: 80, pl. 28B (1844) 

Ety.: scabra (L: rough); relating to the quality of adaxial lamina 

Type: Wallich 2173 (BM) // Nepal 

TW: J. Bot. 23: 103 (1885) 

(261) Dennstaedtia scandens (BLUME) T. MOORE 

Pub.: Parker's Cat. (1858) 

Loc.: 刺柄碗蕨蕨 - CiBingWanJue 

Bas.: Dicksonia scandens BLUME // Enum. Pl. Javae 240 (1828) 

Ety.: scandens (L: climbing); relating to the growth habit 

Type: Java, Blume s. n. (ST: L) // Indonesia 


(262) Dennstaedtia smithii (HOOK.) T. MOORE 

Pub.: Index. Fil. 308 (1861) 

Loc.: 司司氏碗蕨蕨 - SiShiWanJue

62 

Bas.: Dicksonia smithii HOOK. // Sp. Fil. 1: 80-81, pl. 28D (1844) 

Ety.: in honor of J. Smith (British botanist) 

Type: Luzon, Manilla, Cuming 108, 145, 222 (ST: BM, L; IST: E, Z) // 

Philippines 


DEPARIA (Dryopteridaceae) 

Ety.: depas (G: cup or basin); relating to the form of indusium 

(263) Deparia allantodioides (BEDD.) M. KATO 


Loc.: 亞蹄蓋蕨蕨 - YaTiGaiJue 

Bas.: Athyrium allantodioides BEDD. // Ferns Brit. India pl. 221 (1866) 

Ety.: allantodia (from allantos, G: small sausage; referring to the 

indusium shape) relating to a fern genus (Dryopteridaceae), 


frond shape 

Type: Himalaya 

TW: Kuo, C. M. 1997 (1997) 

(264) Deparia dawuense C. M. KUO (nom. nud.) 

[IUCN: - (DD)] 

Loc.: 大武蹄蓋蕨蕨 - DaWuTiGaiJue 

Ety.: relating to the DaWu area (Taiwan), the location where this 

taxon was first observed 


(265) Deparia lancea (THUNB.) FRASER-JENK. 

Pub.: New Sp. Syndr. Indian Pteridol. 101 (1997) 

Loc.: 單葉雙蓋蕨蕨 - DanYeShuangGaiJue 

Bas.: Asplenium lanceum THUNB. // Fl. Jap. 333-334 (1784) 

Ety.: lancea (L: small spear); relating to frond habit 

Type: Japan 

TW: J. Bot. 23: 104 (1885) 

(266) Deparia longipes (CHING) SHINOHARA [IUCN: DD (LC)] 

Pub.: Amer. Fern J. 96(3): 99 (2006) 

Loc.: 昆明假蹄蓋蕨蕨 - KunMingJiaTiGaiJue 

Bas.: Athyriopsis longipes CHING // Acta Phytotax. Sin. 9: 68-69 

(1964) 

Ety.: 

Type: 

longus (L: long), pes (L: foot); relating to the habit of stipe 

Kunming, Sih-shan, Hua-ting-tze, T. N. Lion 14278, 6-Jul-1945 

(HT: PE) // China 

TW: Amer. Fern J. 96(3): 96-99 (2006) 

(267) Deparia petersenii (KUNZE) M. KATO 


Loc.: 假蹄蓋蕨蕨 - JiaTiGaiJue 

Bas.: Asplenium petersenii KUNZE // Analecta Pteridogr. 24 (1837) 

Ety.: in honor of C. W. Petersen, collector of type specimen 

Type: Guangdong, Petersen s. n. (LT: C) // China

TW: Kuo, C. M. 1997 (1997) 

(268) Deparia sp. 1 


(269) Deparia sp. 2 



(270) Deparia subfluvialis (HAYATA) M. KATO 


Loc.: 波氏蹄蓋蕨蕨 - PoShiTiGaiJue 

Bas.: Dryopteris subfluvialis HAYATA // Icon. Pl. Formosan. 5: 288- 

289, f. 113a-b (1915) 

Ety.: sub (L: similar to), fluvialis (L: of rivers, of flows) is relating to a 

taxon with epithet "fluvialis"; relating to the similarity of 

another taxon in Hayata's genus Dryopteris, D. fluvialis 

HAYATA (=Cornopteris banajaoensis, see there) 

Type: Mt. Arisan, inter Karapin et Funkiko, T. Ito & B. Hayata s. n., 

29-Mar-1914 (HT: TI) // Taiwan 

TW: Icon. Pl. Formosan. 5: 288-289, f. 113a-b (1915) 

(271) Deparia tomitaroana (MASAM.) R. SANO [IUCN: DD (VU)] 

Pub.: J. Pl. Res. 113: 162 (2000) 

Loc.: 裂葉雙蓋蕨蕨 - LieYeTiGaiJue 

Bas.: Diplazium tomitaroanum MASAM. // Journ. Soc. Trop. Agric. 2: 

33 (1930) 

Ety.: in honor of Tomitaro Makino, the author of the taxon which it 

replaces (Diplazium lanceum var. crenatum MAKINO) 

Type: replaces Diplazium lanceum var. crenatum MAKINO in Bot. 

Mag. (Tokyo) 27: 253 (1913): Liukiu, Y. Tashiro s. n. (HT: TI) 

// Japan 

TW: Journ. Sci. Engin. 13: 59-60 (1976) 

(272) Deparia unifurcata (BAKER) M. KATO [IUCN: VU (EN)] 


Loc.: 擬大蹄蓋蕨蕨 - NiDaTiGaiJue 

Bas.: Nephrodium unifurcatum BAKER // J. Bot. 26: 228 (1888) 

Ety.: uni (L: one), furcata (L: forked); relating to the veinlets of 

pinnules 

Type: Sichuan, Mt. Omei, Faber 1051 (HT: K; IT: KYO) // China 

TW: Tagawa, M. 1940-49 (1940) 

DICRANOPTERIS (Gleicheniaceae) 

Ety.: dikranos (G: forked), pteris (relates to genus Pteris, or ferns in 

general); relating to the fronds being repeatedly forked 

(273) Dicranopteris linearis (BURM. F.) UNDERW. 

Pub.: Bull. Torrey Bot. Club 34: 250 (1907) 

Loc.: 芒萁 - MangQi 

Bas.: Polypodium lineare BURM. F. // Fl. Indica 235, pl. 67, f. 2 (1768) 

[non Thunb. (1784)] 

63

64 

Ety.: linearis (L: linear); relating to habit of pinnules or segments, or 

both (see protologue) 

Type: Herb. Delessert (G) // Sri Lanka 


(274) Dicranopteris subpectinata (CHRIST) C. M. KUO [IUCN: DD] 


Loc.: 賽芒萁 - SaiMangQi 

Bas.: Gleichenia subpectinata CHRIST // Bot. Tidsskr. 24: 111-112 

(1901) 

Ety.: sub (L: similar to), pectinata (L: with narrow, close divisions 

like a comb); relating to the similarity to taxon Gleichenia 

pectinata C. PRESL (see protologue) 

Type: Koh Chang Isl., Schmidt s. n. ( HT: P) // Thailand 

TW: FOT 1975 (1975) 

(275) Dicranopteris taiwanensis CHING & CHIU [IUCN: DD (LC)] 


Loc.: 台台灣芒萁 - TaiWanMangQi 


Type: without locality, Tsu s. n. (HT: PE; IT: PE) // Taiwan 

TW: Fl. Reipubl. Popularis Sin. 2: 346 (1959) 

(276) Dicranopteris tetraphylla (ROSENST.) C. M. KUO 


Loc.: 蔓芒萁 - ManMangQi 

Bas.: Gleichenia linearis var. tetraphylla ROSENST. // Repert. Spec. 

Nov. Regni Veg. 13: 213 (1914) 

Ety.: tetra (G: four), phyllon (G: leaf); relating to the habit of ultimate 

forks 

Type: Sumatra, Batakerland, Winkler 136 (HT: S; IT: UC) // 

Indonesia 

TW: FOT 1975 (1975) 

DICTYODROMA (Dryopteridaceae) 

Ety.: diktyon (G: net), droma (G: running); relating to the reticulate veins 

having a flowing pattern 

(277) Dictyodroma formosana (ROSENST.) CHING 

Pub.: Acta Phytotax. Sin. 9: 60, pl. 5, f. 1-8 (1964) 

Loc.: 假腸蕨蕨 - JiaChangJue 

Bas.: Diplazium formosanum ROSENST. // Hedwigia 56: 337-338 

(1915) 

Alt.: Deparia formosana (ROSENST.) R. SANO // Acta Phytotax. 

Geobot. 51(1): 17 (2000) 


Type: Urai, Faurie 188 (IT: L, S, UC) // Taiwan 

TW: Hedwigia 56: 337-338 (1915)

65 

DIPLAZIOPSIS (Dryopteridaceae) 

Ety.: diplazios (G: double), opsis (G: like); relating to the similarity to 

members of genus Diplazium (see there) 

(278) Diplaziopsis javanica (BLUME) C. CHR. 

Pub.: Index Filic. 227 (1905) 

Loc.: 腸蕨蕨 - ChangJue 

Bas.: Asplenium javanicum BLUME // Enum. Pl. Javae 175-176 

(1828) 

Ety.: relating to Java (Indonesia), the type location 



DIPLAZIUM (Dryopteridaceae) 

Ety.: diplazios (G: double); relating to the occasional presence of indusia 

on both sides of a vein 

(279) Diplazium aphanoneuron OHWI 

Pub.: J. Jap. Bot. 31: 137-138 (1956) 

Loc.: 隱脈雙蓋蕨蕨 - YinMaiShuangGaiJue 

Ety.: aphanes (G: invisible), neuron (G: nerve); relating to the 

obscured veins of abaxial pinna 

Type: Kyusyu, Ins. Yakusima, 6-Sep-1909 (HT: TNS) // Japan 

TW: a: FOT (1974) as synonym of Diplazium donianum 

b: FOJ (1995) 

(280) Diplazium chinense (BAKER) C. CHR. [IUCN: EN] 

Pub.: Index. Filic. 229 (1905) 

Loc.: 中華雙蓋蕨蕨 - ZhongHuaShuangGaiJue 

Bas.: Asplenium chinense BAKER // Syn. Fil. 237 (1867) 


Type: Shanghai // China 

TW: Kuo, C. M. 1985 (1985) 

(281) Diplazium conterminum CHRIST 

Pub.: J. Bot. (Morot) 19: 67-68 (1905) 

Loc.: 邊生雙蓋蕨蕨 - BianShengShuangGaiJue 

Ety.: conterminum (L: bordering upon); relating to the position of 

sori 

Type: Annam, Cadiere 88, 1903 (LT: P), designated by Pacheco & 

Moran in Amer. Fern J. 93(2): 91 (2003) // Vietnam 

TW: Kuo, C. M. 1997 (1997) 

(282) Diplazium crassiusculum CHING [IUCN: VU (EN)] 

Pub.: Lingnan Sci. J. 15: 279-281 (1936) 

Loc.: 厚厚葉雙蓋蕨蕨 - HouYeShuangGaiJue 

Ety.: crassiusculum (L: moderately thick); relating to lamina texture 

Type: Kwangtung, Lungt’au-shan near Iu, Y. K. Wang 31699 (HT: 

NAS; IT: SYS) // China 

TW: Tagawa, M. 1940-49 (1941)

66 

(283) Diplazium dilatatum BLUME 


Loc.: 廣葉鋸齒雙蓋蕨蕨 - GuangYeJuChiShuangGaiJue 

Ety.: dilatatum (L: broadened, widened); relating to the lamina habit 

(great wide deltate frond with wide pinnules) 



Diplazium latifolium (D. Don) T. MOORE 

b: FOT 1975 (1975) 

(284) Diplazium doederleinii (LUERSS.) MAKINO 


Loc.: 德氏雙蓋蕨蕨 - DeShiShuangGaiJue 

Bas.: Asplenium doederleinii LUERSS. // Bot. Jahrb. Syst. 4: 358-359 

(1883) 

Ety.: in honor of L. H. P. Döderlein (German scientist and collector 

of the type specimen) 

Type: Liu-kiu, Amami-Osima, Döderlein s. n. // Japan 


(285) Diplazium donianum (METT.) TARDIEU 

Pub.: Aspl. Tonkin 58, pl. 5, f. 1-2 (1932) 

Loc.: 雙蓋蕨蕨 - ShuangGaiJue 

Bas.: Asplenium donianum METT. // Fil. Lechl. 2: 177, n. 198b (1859) 

Ety.: in honor of D. Don (Scottish Botanist) 

Type: N. India, Himalaya // India 

TW: Tagawa, M. 1940-49 (1941) 

(286) Diplazium esculentum (RETZ.) SW. 


Loc.: 過溝菜蕨蕨 - GuoGouCaiJue 

Bas.: Hemionitis esculenta RETZ. // Observ. Bot. 6: 38 (1791) 

Ety.: esculentum (L: good to eat); relating to the use of plant as food 

Type: Indonesia 

TW: J. Bot. 23: 105 (1885) 

(287) Diplazium incomptum TAGAWA [IUCN: EN] 


Loc.: 翅柄雙蓋蕨蕨 - ChiBingShuangGaiJue 

Ety.: incomptum (L: disheveled, untidy, unpolished, inelegant); 

relating to the appearance of lamina 

Type: Prov. Taihoku, Agyoku, J. Ohwi 681, 1933 (HT: KYO) // 



(288) Diplazium kawakamii HAYATA var. kawakamii * 


Loc.: 川上氏雙蓋蕨蕨 - ChuanShangShiShuangGaiJue 

Ety.: in honor of T. Kawakami (Japanese botanist), (co-) collector of 

type specimen

Type: Tozan, Kawakami & Nakahara s. n., 1906 (HT: TI) // Taiwan 


(289) Diplazium kawakamii var. subglabratum TAGAWA 

Pub.: J. Jap. Bot. 12: 538 (1936) 

Loc.: 光柄川上氏雙蓋蕨蕨 - GuangBingChuanShuangGaiJue 

Ety.: sub (L: almost), glabratum (L: to make smooth); relating to the 

stipe habit 

Type: Kwarenko Prov., between Seraoka and Hituroku, M. Tagawa 

773, 9-Sep-1934 (HT: KYO; IT: MICH) // Taiwan 

TW: J. Jap. Bot. 12: 538 (1936) 

(290) Diplazium latifrons ALDERW. 

Pub.: Malayan Ferns Fern Allies, Suppl. I: 271 (1917) 

Loc.: 廣葉深山雙蓋蕨蕨 - GuangYeShenShanShuangGaiJue 

Ety.: lati (L: wide, broad), frons (L: leaf, frond); relating to the frond 

habit 

Type: replaces Athyrium platyphyllum COPEL. in Philipp. J. Sci. 3(5): 

292-293 (1908): Luzon, Mindanao, Mt. Data, Copeland s. n., 

29-Oct-1905 (HT: MICH) // Philippines 

TW: Kuo, C. M. 1985 (1985) 

(291) Diplazium laxifrons ROSENST. [IUCN: DD (LC)] 

Pub.: Hedwigia 56: 337 (1915) 

Loc.: 疏葉雙蓋蕨蕨 - ShuYeShuangGaiJue 

Ety.: laxi (L: loose), frons (L: leaf, frond); relating to lamina habit 

Type: Bankinsing, Faurie 172 (IT: KYO) // Taiwan 

TW: Hedwigia 56: 337 (1915) 

(292) Diplazium lobatum (TAGAWA) TAGAWA [IUCN: NT (LC)] 

Pub.: Acta Phytotax. Geobot. 20: 215, f. 2 (1962) 

Loc.: 裂葉雙蓋蕨蕨 - LieYeShuangGaiJue 

Bas.: Diplazium donianum var. lobatum TAGAWA // Acta Phytotax. 

Ety.: 

Geobot. 10: 290 (1941) 

lobatum (L: lobed); relating to the presence of lobes at the 

apical lamina 

Type: Ryukyus, Okinawa, G. Koidzumi s. n., May-1923 (HT: KYO) // 

Japan 

TW: FOT 1975 (1975) 

(293) Diplazium maonense CHING 

Pub.: Hongkong Nat. 7: 88 (1936) 

Loc.: 馬鞍山雙遛 - MaAnShanShuangLiu 

Ety.: relating to "Maon", a mountain in Hong Kong, type location 

Type: Hong Kong, New Territories, Maonshan, C. G. Matthew s. n., 

6-Apr-1906 (K) // China 

TW: Taiwan J. For. Sci. 17(1): 113-118 (2002) 

(294) Diplazium megaphyllum (BAKER) CHRIST 

Pub.: Bull. Herb. Boissier 6: 961 (1898) 

Loc.: 大葉雙蓋蕨蕨 - DaYeShuangGaiJue 

[IUCN: VU (NT)] 

67

68 

Bas.: Asplenium megaphyllum BAKER // J. Bot. 28: 264 (1890) 

Ety.: megas (G: great), phyllon (G: leaf); relating to the pinna shape 

Type: Tonkin, 800 m, Balansa 1836, 21-Jul-1886 (LT: P; ILT: K, P), 

designated by Pacheco & Moran in Amer. Fern J. 93(2): 91 

(2003) // China 

TW: Tagawa, M. 1940-49 (1940) 

(295) Diplazium mettenianum (MIQ.) C. CHR. 


Loc.: 深山雙蓋蕨蕨 - ShenShanShuangGaiJue 

Bas.: Asplenium mettenianum MIQ. // Ann. Mus. Bot. Lugduno- 

Batavi 3: 174-175 (1867) 

Ety.: in honor of G. H. Mettenius (German botanist) 

Type: (L) // Japan 

TW: Hayata, B. 1917 (1917) 

(296) Diplazium okinawaense TAGAWA 

Pub.: Acta Phytotax. 2: 199 (1933) 

Loc.: 琉球雙蓋蕨蕨 - LiuQiuShuangGaiJue 

Ety.: relating to Okinawa (Japan), the type location 

Type: Ryukyus, Okinawa, G. Koidzumi s. n. (HT: KYO, fragment in 

US) // Japan 

TW: Tagawa, M. 1940-49 (1940) 

(297) Diplazium okudairae MAKINO [IUCN: DD (CR)] 


Loc.: 假耳羽雙蓋蕨蕨 - JiaErYuShuangGaiJue 

Ety.: relating to K. Okudaira (Japanese botanist, collector of type 

specimen) 

Type: Prov. Iyo, Mt. Iwaya in Kamiukena-gori, K. Okudaira s. n., 28- 

Oct-1905 (HT: TI) // Japan 

TW: Kuo, C. M. 1997 (1997) 

(298) Diplazium proliferum (LAM.) THOUARS [IUCN: - (DD)] 

Pub.: Esquisse Fl. Tristan D'Acugna 35 (1808) 

Loc.: 多生菜蕨蕨 - DuoShengCaiJue 

Bas.: Asplenium proliferum LAM. // Encycl. 2: 307 (1786) 


Type: 

development of plantlets on the adaxial rachis 

(a) P. Commerson s. n. (ST: MPU) // Reunion 

(b) P. Commerson s. n. (ST: MPU) // Mauritius 

TW: Copeland, E. B. 1960 (1960) 

(299) Diplazium pseudodoederleinii HAYATA 

Pub.: Icon. Pl. Formosan. 8: 145-146, f. 71-72 (1919) 

Loc.: 

Ety.: 

Type: 

擬德氏雙蓋蕨蕨 - NiDeShiShuangGaiJue 

pseudos (G: false), doederleinii relating to a taxon with epithet 

"doederleinii"; relating to the similarity to taxon Diplazium 

doederleinii (see there) 

Ushoko, B. Hayata s. n., Apr-1916 (HT: TI) // Taiwan


(300) Diplazium pullingeri (BAKER) J. SM. 

Pub.: Ferns Brit. For. (ed. 2): 315 (1877) 

Loc.: 樸氏雙蓋蕨蕨 - PuShiShuangGaiJue 

Bas.: Asplenium pullingeri BAKER // Gard. Chron. n. s. 4: 484 (1875) 

Ety.: in honor of Mr. Pullinger (first collector) 

Type: Hong Kong, Pullinger s. n. // China 

TW: Ogata, M. (1940) 

(301) Diplazium sikkimense (C. B. CLARKE) C. CHR. [IUCN: DD (?EW)] 

Pub.: Contr. U. S. Natl. Herb. 26: 304 (1931) 

Loc.: 錫金短腸蕨蕨 - XiJinDuanChangJue 

Bas.: Asplenium sikkimense C. B. CLARKE // Trans. Linn. Soc. 

London, Bot. 1: 500-501, pl. 65, f. 1 (1880) 

Ety.: 

Type: 

relating to Sikkim (India), the type location 

Sikkim, Hooker s. n. (LT: K), designated by Pacheco & Moran 

in Amer. Fern J. 93(2): 92 (2003) // India 


(302) Diplazium squamigerum (METT.) MATSUM. * 

Pub.: Index Pl. Jap. 1: 304 (1904) 

Loc.: 

Bas.: 

Ety.: 

長苞雙蓋蕨蕨 - ChangBaoShuangGaiJue 

Asplenium squamigerum METT. // Ann. Mus. Bot. Lugduno- 

Batavi 2: 239 (1866) 

squama (L: scale), ger (L: carrying, bearing, used as suffix); 

relating to the stipe and rachis habit 

Japan 

Type: 

TW: FOT 1975 (1975) 

(303) Diplazium taiwanense TAGAWA 


Loc.: 

Ety.: 

Type: 

台台灣雙蓋蕨蕨 - TaiWanShuangGaiJue 

relating to Taiwan, the type location of this taxon 

Prov. Taihoku, Mt. Sitisei, Tagawa 624, 28-Aug-1934 (HT: 

KYO; IT: MICH) // Taiwan 


(304) Diplazium virescens KUNZE 


Loc.: 

Ety.: 

Type: 

刺柄雙蓋蕨蕨 - CiBingShuangGaiJue 

virescens (L: becoming green); relating to the lamina 

(a) Goering s. n. (ST: ?LZ) // Japan 

(b) Hb. Zollinger & Moritzi no. 19 (ST: L) // Japan 

TW: Ito, H. 1944 (1944) 

(305) Diplazium wichurae (METT.) DIELS 


Loc.: 

Bas.: 

鋸齒雙蓋蕨蕨 - JuChiShuangGaiJue 

Asplenium wichurae METT. // Ann. Mus. Bot. Lugduno-Batavi 

2: 237 (1866) 

69

70 

Ety.: in honor of M. E. Wichura (German botanist, collector of (a 

syn-) type specimen) 

Type: (a) Siebold s. n. (ST: L) // Japan 

(b) Buerger s. n. (ST: L) // Japan 

(c) Wichura s. n. (ST: ?B) // Japan 

TW: J. Bot. 23: 105 (1885) 

DIPLOPTERYGIUM (Gleicheniaceae) 

Ety.: diploos (G: double), pterygium (G: little wing); relating to two pinnae 

of a frond resembling wings 

(306) Diplopterygium blotianum (C. CHR.) NAKAI 

Pub.: Bull. Natl. Sci. Mus. 29: 49 (1950) 

Loc.: 逆羽裏白 - NiYuLiBai 

Bas.: Gleichenia blotiana C. CHR. // Bull. Mus. Natl. Hist. Nat., ser. 2, 

6: 103-104 (1934) 

Ety.: probably in honor of M. L. Tardieu-Blot (French botanist) 

Type: Tonkin, near Chapa, Petelot 3900 (HT: BM, fragment in PE; IT: 

P) // Vietnam 

TW: Tagawa, M. 1940-49 (1940) 

(307) Diplopterygium chinense (ROSENST.) DEVOL 

Pub.: Fl. Taiwan 1: 92, pl. 28 (1975) 

Loc.: 中華裏白 - ZhongHuaLiBai 

Bas.: Gleichenia chinensis ROSENST. // Repert. Spec. Nov. Regni 

Veg. 13: 120-121 (1913) 


Type: Pin-fa, Cavalerie 3860 (HT: P) // China 

TW: FOT 1975 (1975) 

(308) Diplopterygium glaucum (THUNB.) NAKAI 

Pub.: Bull. Nation. Sci. Mus. 29: 51 (1950) 

Loc.: 裏白 - LiBai 

Bas.: Polypodium glaucum THUNB. // Nat. Hist. 2(14): 177 (1783) 

Ety.: glaucos (G: bluish-green); relating to the colour of abaxial 

lamina 

Type: Thunberg s. n. (BM, ?S) // Japan 


(309) Diplopterygium laevissimum (CHRIST) NAKAI 


Loc.: 鱗芽裏白 - LinYaLiBai 

Bas.: Gleichenia laevissima CHRIST // Bull. Acad. Int. Geogr. Bot. 11: 

268-269 (1902) 

Ety.: superlative of laevis (L: smooth); relating to the habit (entirely 

glabrous, see protologue) 

Type: Kouyyang, Bodinier 2095 (HT: P) // China 

TW: FOT 1975 (1975)

71 

DIPTERIS (Dipteridaceae) 

Ety.: di (G: two), pteris (relates to genus Pteris, or ferns in general); 

relating to the fronds that are evenly divided into two flabellate 

parts 

(310) Dipteris conjugata REINW. 

Pub.: Syll. Pl. Nov. 2: 3 (1828) 

Loc.: 雙扇蕨蕨 - ShuangShanJue 

Ety.: conjugata (L: united); relating to the lamina habit 

Type: Java (HT: ?L) // Indonesia 

TW: J. Bot. 23: 106 (1885) 

DRYMOTAENIUM (Polypodiaceae) 

Ety.: drymos (G: forest), taenia (G: ribbon); relating to an epiphyte with 

ribbon-like fronds 

(311) Drymotaenium miyoshianum (MAKINO) MAKINO [IUCN: NT] 


Loc.: 二條線蕨蕨 - ErTiaoXianJue 

Bas.: Taenitis miyoshiana MAKINO // Bot. Mag. (Tokyo) 12: 26-27 

(1899) 

Ety.: in honor of M. Miyoshi (Japanese botanist, collector of (a syn-) 


Alt.: Lepisorus miyoshianus (MAKINO) FRASER-JENK. & SUBH. 

CHANDRA // Taxon. Revis. Indian Subcontinental 

Pteridophytes 37 (2008) 

Type: (a) Gifu (Hida), Kuragari, Ochiai-mura in Masuda-gori, Y. 

Nawa & M. Miyoshi s. n., 15-Aug-1889 (ST: TI) // Japan 

(b) Prov. Mino, Kuragari-dani, Shiro-yama, Iwamura in Enagori, 

S. Mori, 19-Jul-1890 (ST: MAK) // Japan 

(c) Shirasu-hora in Shimo, Shimoharada-mura, Ena-gori, M. 

Iino, 1890 (ST: MAK) // Japan 

TW: Ito, H. 1944 (1944) 

DRYNARIA (Polypodiaceae) 

Ety.: dryinos (G: of oaks), aris (G: like) or aria (G: substantive ending); 

relating to the oak leaf shape of the humus collecting sterile leaves 

(312) Drynaria fortunei (KUNZE ex METT.) J. SM. 

Pub.: Bot. Voy. Herald 425-426 (1857) [not T. Moore (1855)] 

Loc.: 槲蕨蕨 - HuJue 

Bas.: Polypodium fortunei KUNZE ex METT. // Abh. Senckenberg. 

Naturf. Ges. 1: 121, pl. 3, f. 42-45 (1856) 

Ety.: in honor of R. Fortune (British botanist and collector of the type 

specimen) 

Type: Fuo-chou-fou, Fortune 34 (IST: B) // China 

TW: Matsumara, J. & Hayata, B. (1906)

72 

DRYOPSIS (Dryopteridaceae) 

Ety.: a combination of the two generic names Dryopteris and Ctenitis, 

which it each resembles in parts (derived from the protologue) 

(313) Dryopsis apiciflora (WALL. ex METT.) HOLTT. & P. J. EDWARDS 

Pub.: Kew Bull. 41(1): 189 (1986) 

Loc.: 頂囊肋毛蕨蕨 - DingNangLeMaoJue 

Bas.: Aspidium apiciflorum WALL. // Numer. List n. 345 (1828), ex 

Mettenius, Abh. Senckenberg. Naturf. Ges. 4: 54 (1858) 

Ety.: apex (L: tip), flos (L: flower); relating to the position of sori on 

apical veins 

Type: Wallich 345 (IT: K, L, S) // Nepal 

TW: Hayata, B. 1914-21 (1914) 

(314) Dryopsis x fauriei HOLTT. & P. J. EDWARDS 

Pub.: Kew Bull. 41(1): 198-199 (1986) 

Loc.: 傅氏擬鱗毛蕨蕨 - FuShiNiLinMaoJue 

Ety.: in honor of U. J. Faurie (French botanist, collector of type 

specimen) 

Type: Arisan, Faurie 381 (HT: P; IT: S) // Taiwan 

TW: Kew Bull. 41(1): 198-199 (1986) 

(315) Dryopsis kawakamii (HAYATA) HOLTT. & P. J. EDWARDS 

Pub.: Kew Bull. 41(1): 186 (1986) 

Loc.: 川上氏擬鱗毛蕨蕨 - ChuanShangShiNiLinMaoJue 

Bas.: Dryopteris kawakamii HAYATA // J. Coll. Sci. Imp. Univ. Tokyo 

30: 416-417 (1911) 

Ety.: in honor of T. Kawakami (Japanese botanist) 

Type: Randaisan, Mori 7134, Aug-1908 (HT: TAI) // Taiwan 


(316) Dryopsis maximowicziana (MIQ.) HOLTT. & P. J. EDWARDS [IUCN: VU (EN)] 

Pub.: Kew Bull. 41(1): 197 (1986) 

Loc.: 白鱗擬鱗毛蕨蕨 - BaiLinNiLinMaoJue 

Bas.: Aspidium maximowiczianum MIQ. // Ann. Mus. Bot. Lugduno- 

Batavi 3: 178-179 (1867) 

Ety.: in honor of K. J. Maximowicz (Russian botanist) 

Type: Buerger s. n. (HT: L) // Japan 

TW: Ito, H. 1944 (1944) 

(317) Dryopsis transmorrisonensis (HAYATA) HOLTT. & P. J. EDWARDS 

Pub.: Kew Bull. 41(1): 191 (1986) 

Loc.: 玉山擬鱗毛蕨蕨 - YuShanNiLinMaoJue 

Bas.: Polystichum transmorrisonense HAYATA // J. Coll. Sci. Imp. 

Univ. Tokyo 30: 427-428 (1911) 

Ety.: trans (L: across), morrisonensis is relating to YuShan ("Mt. 

Morrison"), the highest mountain of Taiwan; name relates to 

the type location 

Type: in monte Morrison, ad 12000 ped. alt., T. Kawakami & U. Mori 

s. n., Oct-1906 (HT: TI) // Taiwan

73 


DRYOPTERIS (Dryopteridaceae) * 

Ety.: drys (G: oak), pteris (relates to genus Pteris, or ferns in general); 

relating to a fern growing among oaks 

(318) Dryopteris alpestris TAGAWA 


Loc.: 腺鱗毛蕨蕨 - XianLinMaoJue 

Ety.: alpes (L: the Alps), relating to the alpine habitat (though not 

present in the Alps, Europe) 

Type: Taitung, Mt. Daisuikutusan, Ohwi 3801, 1933 (HT: KYO) // 



(319) Dryopteris atrata (WALL. ex KUNZE) CHING 

Pub.: Sinensia 3: 326 (1933) 

Loc.: 暗鱗鱗毛蕨蕨 - AnLinLinMaoJue 

Bas.: Aspidium atratum WALL. ex KUNZE // Linnaea 24: 279 (1851) 

Ety.: atrata (L: clothed in black); relating to the scales on axes 

Type: S. India, Schmid & Koch 10, 25, 91, 147 (UC) // India 

TW: FOT 1975 (1975) 

(320) Dryopteris cacaina TAGAWA [IUCN: - (NT)] 


Loc.: 蓬萊鱗毛蕨蕨 - PengLaiLinMaoJue 

Ety.: cacaina (L: chocolate-brown); relating to the colour of the stipe 

base 

Type: Prov. Tainan, inter Numanohira et Tataka, M. Tagawa 397, 15- 

Aug-1934 (HT: KYO) // Taiwan 


(321) Dryopteris championii (BENTH.) C. CHR. ex CHING [IUCN: VU] 

Pub.: Sinensia 3: 327 (1933) 

Loc.: 闊鱗鱗毛蕨蕨 - KuoLinLinMaoJue 

Bas.: Aspidium championii BENTH. // Fl. Hongk. 456 (1861) 

Ety.: in honor of J. G. Champion (British botanist and collector of the 

Type: 

(syn-) type specimen) 

(a) Hongkong, Champion s. n. // China 

(b) Hongkong, Urquhart s. n. // China 


(322) Dryopteris costalisora TAGAWA 


Loc.: 能高鱗毛蕨蕨 - NengGaoLinMaoJue 

Ety.: costalis (L: costal), sorus (L: sorus); relating to the costal 

position of sori 

Type: Prov. Taityu, in monte Noko-goe, J. Ohwi 3251, 1933 (HT: 

KYO) // Taiwan 

TW: Acta Phytotax. Geobot. 3: 88-89 (1934)

74 

(323) Dryopteris cycadina (FRANCH. & SAV.) C. CHR. 


Loc.: 桫欏鱗毛蕨蕨 - SuoLuoLinMaoJue 

Bas.: Aspidium cycadinum FRANCH. & SAV. // Enum. Pl. Jap. 2: 236, 

630 (1879) 

Ety.: cycas relates to gymnosperm genus Cycas (Cycadaceae), ina 

(L: suffix, denoting likeness); relating to the frond habit 

Type: sine loco spec., S. Ono s. n. (HT: P) // Japan 

TW: Kuo, C. M. 1985 (1985) 

(324) Dryopteris decipiens (HOOK.) KUNTZE [IUCN: VU] 

Pub.: Revis. Gen. Pl. 2: 812 (1891) 

Loc.: 迷人鱗毛蕨蕨 - MiRenLinMaoJue 

Bas.: Nephrodium decipiens HOOK. // Sp. Fil. 4: 86-87, pl. 243 

(1862) 

Ety.: decipiens (L: to deceive, to trap); perhaps relating to the 

difficulties experienced by the author when classifying this 

taxon (see protologue) 

Type: Foo-chow-foo, Alexander s. n. // China 

TW: Ito, T. (1928) 

(325) Dryopteris dickinsii (FRANCH. & SAV.) C. CHR. [IUCN: DD (EN)] 


Loc.: 遠軸鱗毛蕨蕨 - YuanZhouLinMaoJue 

Bas.: Aspidium dickinsii FRANCH. & SAV. // Enum. Pl. Jap. 2: 236, 

629-630 (1879) 

Ety.: in honor of F. V. Dickins (British botanist and collector of the 


Type: Hakone, Dickins, Herb. Savatier (LT: P; ILT: K, PE), 

designated by Fraser-Jenkins, Bull. Brit. Mus. (Nat. Hist.), 

Bot. 18: 337 (1989) // Japan 

TW: Fraser-Jenkins, C. R. 1989 (1989) 

(326) Dryopteris diffracta (BAKER) C. CHR. 


Loc.: 彎柄假複葉耳蕨蕨 - WanBingJiaFuYeErJue 

Bas.: Nephrodium diffractum BAKER // Bull. Misc. Inform. Kew 1898: 

230 (1898) 

Ety.: diffracta (L: shattered); relating to the zigzag shaped rachis 

Alt.: Acrorumohra diffracta (BAKER) H. ITO // Nov. Fl. Jap. 4: 101 

(1939) 

Type: Yunnan, Mengtze, Henry 9028 (HT: K) // China 


(327) Dryopteris edwardsii FRASER-JENK. 

Pub.: New Sp. Syndr. Indian Pteridol. 138-140 (1997) 

Loc.: 愛德華鱗毛蕨蕨 - AiDeHuaLinMaoJue 

Ety.: in honor of P. J. Edwards (British botanist)

Type: Uttarkhand, Chamoli Garhwal, 2800-3000 m, C. R. Fraser- 

Jenkins 622, 29-Sep-1996 (HT: BM; IT: K) // India 


(328) Dryopteris enneaphylla (BAKER) C. CHR. 


Loc.: 頂羽鱗毛蕨蕨 - DingYuLinMaoJue 

Bas.: Nephrodium enneaphyllum BAKER // J. Bot. 25: 170 (1887) 

Ety.: ennea (G: nine), phyllon (G: leaf); relating to the average 

number of pinna of a frond 

Type: Hubei, Ichang, Henry 3217 (HT: K) // China 

TW: FOT 1975 (1975) 

(329) Dryopteris expansa (C. PRESL) FRASER-JENK. & JERMY 

Pub.: Fern Gaz. 11(5): 338 (1977) 

Loc.: 闊葉鱗毛蕨蕨 - KuoYeLinMaoJue 

Bas.: Nephrodium expansum C. PRESL // Reliq. Haenk. 1: 38 (1825) 

Ety.: expansa (L: expanded); relating to the wide lamina shape 

Type: British Columbia, Nootka-Sund, Haenke s. n. (LT: PRC) // 

Canada 

TW: Kuo, C. M. 1985 (1985) 

(330) Dryopteris formosana (CHRIST) C. CHR. 


Loc.: 台台灣鱗毛蕨蕨 - TaiWanLinMaoJue 

Bas.: Aspidium formosanum CHRIST // Bull. Herb. Boissier, ser.2, 4: 

615-616 (1904) 


Type: Taitung, Faurie 687 (HT: P) // Taiwan 

TW: Bull. Herb. Boissier, ser.2, 4: 615-616 (1904) 

(331) Dryopteris fructuosa (CHRIST) C. CHR. 


Loc.: 深山鱗毛蕨蕨 - ShenShanLinMaoJue 

Bas.: Aspidium varium var. fructuosum CHRIST // Bull. Herb. Boissier 

6: 967 (1898) 

Ety.: fructuosa (L: fruitful, abounding in fruit); relating the abundance 

of sori 

Type: Yunnan, Manmei, Henry 10095 (LT: P; ILT: K), designated by 

Fraser-Jenkins, Bull. Brit. Mus. (Nat. Hist.), Bot. 18: 402 

(1989) // China 


(332) Dryopteris fuscipes C. CHR. [IUCN: - (VU)] 


Loc.: 黑足鱗毛蕨蕨 - HeiZuLinMaoJue 

Ety.: fuscus (L: brown), pes (L: foot); relating to the colour of the 

stipe 

75

76 

Type: replaces Dryopteris bipinnata C. CHR. in Cat. Plant. Yun-Nan 

102-103 (1916): Kouy-Tcheou (=Guizhou), J. Cavalerie s. n. 

(HT: BM) // China 

TW: Ito, H. 1944 (1944) 

(333) Dryopteris hasseltii (BLUME) C. CHR. 


Loc.: 哈氏假複葉耳蕨蕨 - HaShiJiaFuYeErJue 

Bas.: Polypodium hasseltii BLUME // Fl. Javae 195-196, pl. 92 (1829) 

Ety.: in honor of J. C. v. Hasselt (Dutch botanist) 

Alt.: Acrorumohra hasseltii (BLUME) CHING // Acta Phytotax. Sin. 9: 

385 (1964) 

Type: Java, Karanjo Marat, Zippel s. n. (HT: L) // Indonesia 

TW: Ito, H. 1944 (1944) 

(334) Dryopteris integriloba C. CHR. [IUCN: DD (VU)] 

Pub.: Bull. Dept. Biol. Sun Yatsen Univ. 6: 5 (1933) 

Loc.: 蓬萊紅苞鱗毛蕨蕨 - PengLaiHongBaoLinMaoJue 

Ety.: integer (L: entire), lobus (L: lobe); relating to the margin of 

pinnules 

Type: Guangdong, 22-Sep-1918 (US) // China 

TW: MAN (2002) 

(335) Dryopteris komarovii KOSSINSKY [IUCN: DD (LC)] 

Pub.: Not. Syst. Hort. Petr. 2: 1-2 (1921) 

Loc.: 近多鱗鱗毛蕨蕨 - JinDuoLinLinMaoJue 

Ety.: in honor of V. L. Komarov (Russian botanist and collector of 


Type: Tienshan, Chschartob, Komarov s. n., 20-Aug-1892 (HT: LE) // 

Russia 


(336) Dryopteris kwanzanensis TAGAWA [IUCN: - (NT)] 


Loc.: 擬倒鱗鱗毛蕨蕨 - NiDaoLinLinMaoJue 

Ety.: relating to GuanShan ("Kwanzan") in Taiwan, the type location 

Type: S. Ogamoto s. n., 8-Oct-1938 (HT: KYO) // Taiwan 


(337) Dryopteris lacera (THUNB.) KUNTZE [IUCN: VU] 


Loc.: 二型鱗毛蕨蕨 - ErXingLinMaoJue 

Bas.: Polypodium lacerum THUNB. // Fl. Jap. 337 (1784) 

Ety.: lacero (L: to tear); relating to the stipe scale shape 

Type: 

TW: Kuo, C. M. 1985 (1985) 

(338) Dryopteris lepidopoda HAYATA 

Thunberg s. n. (IT: ?L) // Japan 


Loc.: 厚厚葉鱗毛蕨蕨 - HouYeLinMaoJue

Ety.: lepidotos (G: scaly), podos (G: foot); relating to the scaly stipe 

base and rachis 

Type: Arisan, B. Hayata & S. Sasaki s. n., Jan-1912 (HT: TI) // 



(339) Dryopteris marginata (C. B. CLARKE) CHRIST 


Loc.: 三角葉鱗毛蕨蕨 - SanJiaoYeLinMaoJue 

Bas.: Nephrodium filix-mas var. marginatum C. B. CLARKE // Trans. 

Linn. Soc. London 1: 521, pl. 71 (1880) 

Ety.: margo (L: edge, border); relationship unclear to me, probably 

Type: 

relating to the position of sori as illustrated in the protologue 

Wallich 391 (LT: K, Wallich Herbarium; ILT: K), designated by 

Fraser-Jenkins, Bull. Brit. Mus. (Nat. Hist.), Bot. 18: 420 

(1989) // Nepal 

TW: Kuo, C. M. 1985 (1985) 

(340) Dryopteris melanocarpa HAYATA 


Loc.: 黑苞鱗毛蕨蕨 - HeiBaoLinMaoJue 

Ety.: melaina (G: black), karpos (G: fruit); relating to the colour of 

sorus 

Type: Arisan, B. Hayata & S. Sasaki s. n., Jan-1912 (HT: ?TI; ST: 



(341) Dryopteris namegatae (SA. KURATA) SA. KURATA [IUCN: DD (CR)] 

Pub.: J. Geobot. 17: 87, 89 (1969) 

Loc.: 黑鳞鳞远轴鳞鳞毛蕨蕨 - HeiLinYuanZhouLinMaoJue 

Bas.: Dryopteris dickinsii var. namegatae SA. KURATA // J. Geobot. 7: 

115-116 (1958) 

Ety.: in honor of T. Namegata (Japanese botanist) 

Type: Honshu, Kurata 1099, 5-Jun-1958 (IT: KYO) // Japan 

TW: Kuo, C. M. 1985 (1985) 

(342) Dryopteris polita ROSENST. 

Pub.: Repert. Spec. Nov. Regni Veg. 13: 218-219 (1914) 

Loc.: 台台東鱗毛蕨蕨 - TaiDongLinMaoJue 

Ety.: polita (L: polished); relating to the glossy adaxial lamina 

Type: Sumatra, H. J. P. Winkler 51-b (IT: MICH, NY) // Indonesia 

TW: Ito, H. 1944 (1944) 

(343) Dryopteris pseudolunanensis TAGAWA [IUCN: - (VU)] 


Loc.: 擬桫欏鱗毛蕨蕨 - NiSuoLuoLinMaoJue 

Ety.: pseudos (G: false), lunanensis relating to the taxon with 

epithet "lunanensis" (which is referring to LuNan, a region in 

China); relating to the similarity to taxon Dryopteris 

lunanensis (CHRIST) C. CHR. 

77

78 

Type: Prov. Taito, near Asahi, Kwanzan-gun, ca. 1700 m, Tagawa 

3108, 6-Mar-1940 (HT: KYO; IT: KYO, MICH) // Taiwan 


(344) Dryopteris pseudosieboldii HAYATA [IUCN: NT (LC)] 


Loc.: 大頂羽鱗毛蕨蕨 - DaDingYuLinMaoJue 

Ety.: pseudos (G: false), sieboldii relating to the taxon with epithet 

"sieboldii" (in honor of P. F. Siebold, German botanist); 

relating to the similarity to taxon Dryopteris sieboldii (HOUTT. 

ex METT.) KUNTZE 




(345) Dryopteris redactopinnata S. K. BASU & PANIGR. 

Pub.: Indian J. Forest. 3(3): 270, t. 2 (1980) 

Loc.: 藏藏布鱗毛蕨蕨 - CangBuLinMaoJue 

Ety.: redacta (L: reduced), pinnata (L: pinnate); relating to the basal 

lamina habit 

Type: Kashmir, Gulmarg, 7000 ft (2135 m), 21-Jul-1891, G. A. 

Gammie (HT: CAL; IT: CAL) // India 


(346) Dryopteris reflexosquamata HAYATA 


Loc.: 逆鱗鱗毛蕨蕨 - NiLinLinMaoJue 

Ety.: reflexa (L: reflexed), squamata (L: furnished with scales); 

relating to the reflexed scales at stipe base 

Type: Arisan, ad 8000 ped. alt., B. Hayata & S. Sasaki s. n., Jan- 

1912 (HT: TI; IT: TAIF) // Taiwan 


(347) Dryopteris ryo-itoana SA. KURATA [IUCN: - (VU)] 

Pub.: J. Geobot. 15: 84 (1967) 

Loc.: 阔羽鳞鳞毛蕨蕨 - KuanYuLinMaoJue 

Ety.: in honor of R. Ito (Japanese scientist) 

Type: Mie (Kii), Kitamuro-gun, Miyama-machi, Uotori, collected by S. 

Kurata in May-1955, cult. in Kurata Fern Garden, S. Kurata s. 

n., Jun-1966 (HT: TOFO) // Japan 

TW: Taiwan J. For. Sci. 28: 43-49 (2013) 

(348) Dryopteris scottii (BEDD.) CHING ex C. CHR. 

Pub.: Bull. Dept. Biol. Sun Yatsen Univ. 6: 3 (1933) 

Loc.: 史史氏鱗毛蕨蕨 - ShiShiLinMaoJue 

Bas.: Polypodium scottii BEDD. // Ferns Brit. India pl. 345 (1866) 

Ety.: in honor of J. Scott (British botanist and collector of the type 

specimen) 

Type: Sikkim, Darjeeling, Rungit, Scott s. n. (?CAL) // India 

TW: Ito, H. 1944 (1944)

(349) Dryopteris serratodentata (BEDD.) HAYATA 

Pub.: Icon. Pl. Formosan. 4: 179, f. 116 (1914) 

Loc.: 鋸齒葉鱗毛蕨蕨 - JuChiYeLinMaoJue 

Bas.: Lastrea filix-mas var. serratodentata BEDD. // Handb. Suppl. 55 

(1892) 

Ety.: serrata (L: serrated), dentata (L: toothed); relating to the 

lamina segments 

Type: Sikkim, Beddome s. n. (LT: BM), designated by Fraser- 

Jenkins, Bull. Brit. Mus. (Nat. Hist.), Bot. 18: 377 (1989) // 

India 


(350) Dryopteris sordidipes TAGAWA 


Loc.: 落鱗鱗毛蕨蕨 - LuoLinLinMaoJue 

Ety.: sordidus (L: dirty looking), pes (L: foot); relating to the colour of 

the stipe 

Type: Kyusyu, Yakushima, Mt. Nagata-dake, M. Tagawa 702, Aug- 

1933 (HT: KYO; IT: P, US) // Japan 

TW: Ito, H. 1944 (1944) 

(351) Dryopteris sp. 



(352) Dryopteris sparsa (D. DON) KUNTZE 


Loc.: 長葉鱗毛蕨蕨 - ChangYeLinMaoJue 

Bas.: Nephrodium sparsum D. DON // Prodr. Fl. Nepal. 6 (1825) 

Ety.: sparsa (L: sparse, scattered); relating to the small number of 

sori developed (see protologue) 

Type: Suembu, Buchmann s. n. (HT: BM) // Nepal 

TW: Hayata, B. 1911 (1911) 

(353) Dryopteris stenolepis (BAKER) C. CHR. 


Loc.: 狹鱗鱗毛蕨蕨 - XiaLinLinMaoJue 

Bas.: Polypodium stenolepis BAKER // Bull. Misc. Inform. Kew 1898: 

231 (1898) 

Ety.: stenos (G: narrow), lepis (G: scale); relating to the narrow 

Type: 

rachis and stipe scales 

Yunnan, Mengtze, Henry 9038 (LT: K; ILT: E, NY, US), 

designated by Fraser-Jenkins, Bull. Brit. Mus. (Nat. Hist.), 

Bot. 18: 336 (1989) // China 

TW: Tagawa, M. 1940-49 (1941) 

(354) Dryopteris subatrata TAGAWA 


Loc.: 細葉鱗毛蕨蕨 - XiYeLinMaoJue 

79

80 

Ety.: 

Type: 

sub (L: similar to), atrata relating to the taxon with epithet 

"atrata"; relating to the similarity to taxon Dryopteris atrata 

(see there) 

Prov. Takao, between Keinanzan and Isiyama, Kizan-gun, ca. 

1500-1700 m, Tagawa 1615, 25-Dec-1938 (HT: KYO) // 



(355) Dryopteris subexaltata (CHRIST) C. CHR. 


Loc.: 早田氏鱗毛蕨蕨 - ZaoTianShiLinMaoJue 

Bas.: Aspidium subexaltatum CHRIST // Bull. Herb. Boissier, ser.2, 4: 

616 (1904) 

Ety.: sub (L: almost), exaltata (L: very tall); relating to the habit 

Type: sylvis Saitouka, U. Faurie 651, 22-Jan-1901 (IT: KYO) // 


TW: Bull. Herb. Boissier, ser.2, 4: 616 (1904) 

(356) Dryopteris sublacera CHRIST [IUCN: DD (CR)] 


Loc.: 半育鳞鳞毛蕨蕨 - BanYuLinMaoJue 

Ety.: 

Type: 

sub (L: similar to), lacera relating to the taxon with epithet 

"lacera"; relating to the similarity to taxon Dryopteris lacera 

(see there) 

Yunnan, Yunnan Sen, Tchong Chan, F. Ducloux 3347, 4-Oct- 

1905 (LT: P; ILT: P), designated by Fraser-Jenkins, Bull. Brit. 

Mus. (Nat. Hist.), Bot. 18: 389 (1989) // China 

TW: Kuo, C. M. 1985 (1985) 

(357) Dryopteris subreflexipinna M. OGATA 

Pub.: J. Jap. Bot. 11: 30, f. 2 (1935) 

Loc.: 

Ety.: 

Alt.: 

微彎柄複葉耳蕨蕨 - WeiWanBingFuYeErJue 

sub (L: similar to), reflexipinna (L: reflexed pinna) relating to 

the taxon with epithet "reflexipinna"; relating to the similarity 

to taxon Dryopteris reflexipinna HAYATA (=D. diffracta, see 

there) 

Acrorumohra subreflexipinna (M. OGATA) H. ITO // Nov. Fl. Jap. 

4: 102 (1939) 

Type: Prov. Taityu, Keitau, M. Ogata s. n., 9-Dec-1928 (ST: KYO) // 


TW: J. Jap. Bot. 11: 30, f. 2 (1935) 

(358) Dryopteris subtriangularis (C. HOPE) C. CHR. 


Loc.: 紅苞鱗毛蕨蕨 - HongBaoLinMaoJue 

Bas.: Nephrodium subtriangulare C. HOPE // J. Bot. 28: 327-328 

(1890) 

Ety.: sub (L: similar to), triangularis (L: triangular); relating to the 

shape of lamina

Type: above Laukot, Khasi Hills, 800 ft, G. Mann s. n., Sep-1888 (LT: 

P; ILT: E), designated by Fraser-Jenkins, Bull. Brit. Mus. 

(Nat. Hist.), Bot. 18: 430 (1989) // India 


(359) Dryopteris tenuicula C. G. MATTHEW & CHRIST 


Loc.: 疏葉鱗毛蕨蕨 - ShuYeLinMaoJue 

Ety.: tenuis (L: slender, slim), cula (L: suffix, diminutive form); 

relating to a very small, thin narrow fern (which are 

characteristics not typically displayed in local populations of 

this taxon) 

Type: Kwangtung, North River, Matthew 54 (?K) // China 


(360) Dryopteris tenuipes (ROSENST.) SERIZ. [IUCN: DD (VU)] 

Pub.: J. Jap. Bot. 46: 19 (1971) 

Loc.: 落葉鱗毛蕨蕨 - LuoYeLinMaoJue 

Bas.: Dryopteris erythrosora var. tenuipes ROSENST. // Hedwigia 56: 

341 (1915) 

Ety.: tenuis (L: slender, slim), pes (L: foot); probably relating to the 

slender stipe (protologue describes that this taxon, described 

originally as variety of Dryopteris erythrosora (D. C. EATON) 

KUNTZE, is smaller in several parts) 

Type: Taipei, Faurie 63 (HT: HUH; IT: KYO) // Taiwan 

TW: Hedwigia 56: 341 (1915) 

(361) Dryopteris toyamae TAGAWA [IUCN: EN] 


Loc.: 外山氏鱗毛蕨蕨 - WaiShanShiLinMaoJue 

Ety.: in honor of S. Toyama (Japanese botanist and collector of a 

Type: 

paratype) 

cultivated, Ryukyu origin: Prov. Hizen, Mt. Kokuzo-yama, 

Tagawa 2552 (HT: KYO; IT: US) // Japan 

TW: Tagawa, M. 1940-49 (1940) 

(362) Dryopteris varia (L.) KUNTZE 


Loc.: 南海鱗毛蕨蕨 - NanHaiLinMaoJue 

Bas.: Polypodium varium L. // Sp. Pl. 2: 1090 (1753) 

Ety.: varia (L: variable); probably relating to the lamina morphology 

Type: Osbeck s. n. // China 


(363) Dryopteris wallichiana subsp. nepalensis FRASER-JENK. 

Pub.: New Sp. Syndr. Indian Pteridol. 134-136 (1997) 

Loc.: 尼尼泊爾鱗毛蕨蕨 - NiBoErLinMaoJue 

Ety.: relating to Nepal, the type location 

Type: C. Nepal, Bagmati Zone, Kathmandu District, c. 2500 m, C. R. 

Fraser-Jenkins 15793, 16-Nov-1989 (HT: BM) // Nepal 

81

82 


(364) Dryopteris wallichiana (SPRENG.) HYL. subsp. wallichiana * 

Pub.: Bot. Not. 1953: 352 (1953) 

Loc.: 瓦氏鱗毛蕨蕨 - WaShiLinMaoJue 

Bas.: Aspidium wallichianum SPRENG. // Syst. Veg. 4: 104 (1827) 

Ety.: in honor of N. Wallich (Danish botanist, collector of the type 

Type: 

specimen) 

Wallich 340 (LT: BM; ILT: BM, G, K, L, P, UC, W), designated 

by Fraser-Jenkins, Bull. Brit. Mus. (Nat. Hist.), Bot. 18: 354 

(1989) // Nepal 

TW: FOT 1975 (1975) 

(365) Dryopteris woodsiisora HAYATA 


Loc.: 

Ety.: 

擬岩蕨蕨 - NiYanJue 

woodsia is relating to fern genus Woodsia (see there), soros 

(G: sorus); probably relating to the sorus having a habit like 

members of Woodsia 

Type: in rupibus Arisan, ad 2500 m alt., U. Faurie 518, Jun-1914 

(HT: TI; IT: KYO) // Taiwan 

TW: Icon. Pl. Formosan. 6: 158-159 (1916) 

(366) Dryopteris xanthomelas (CHRIST) C. CHR. [IUCN: - (VU)] 


Bas.: Aspidium xanthomelas CHRIST // Bull. Acad. Int. Geogr. Bot. 

16: 117 (1906) 

Ety.: xanthos (G: various shades of yellow), melas (G: black); 

relating to the colour of sori (changing from yellow when 

young, to black) 

Type: W. China (HT: P) // China 


(367) Dryopteris yoroii SERIZ. [IUCN: EN] 

Pub.: J. Jap. Bot. 46: 20-21, f. 1b (1971) 

Loc.: 玉山假複葉耳蕨蕨 - YuShanJiaFuYeErJue 

Ety.: 

in honor of R. Yoroi (Japanese botanist, collector of type 

specimen) 

Alt.: Acrorumohra yoroii (SERIZ.) W. C. SHIEH // J. Sci. Engin. 12: 

301 (1975) 

Type: Mt. Morrison, Yoroi s. n., 8-Aug-1968 (HT: TNS) // Taiwan 

TW: J. Jap. Bot. 46: 20-21, f. 1b (1971) 

(368) Dryopteris zayuensis CHING & S. K. WU [IUCN: - (VU)] 

Pub.: Fl. Xizang. 1: 255-256 (1983) 

Ety.: relating to "Zayu" (the Chinese name for Cha Yul, a place in 

SE Tibet, and the type location) 

Type: Zayu, alt. 3700 m, Qinghai-Xizang (Tibet) Complex Exped. 373 

(HT: PE; IT: PE) // China 

TW: Knapp, R. 2011 (2011)

83 

ELAPHOGLOSSUM (Lomariopsidaceae) 

Ety.: elaphos (G: stag), glossa (G: tongue); relating to the shape of fronds 

(like a "stag's tongue") 

(369) Elaphoglossum angulatum (BLUME) T. MOORE 


Loc.: 爪哇舌蕨蕨 - ZhuaWaSheJue 

Bas.: Acrostichum angulatum BLUME // Enum. Pl. Javae 101 (1828) 

Ety.: angulatum (L: angled); relating to the stipe scales 


TW: FOT 1975 (1975) 

(370) Elaphoglossum callifolium (BLUME) T. MOORE [IUCN: VU] 


Loc.: 銳頭舌蕨蕨 - RuiTouSheJue 

Bas.: Acrostichum callifolium BLUME // Enum. Pl. Javae 100-101 

(1828) 

Ety.: callus (L: hard skin), folium (L: leaf); relating to the lamina 

texture 

Type: Java, Blume (HT: L) // Indonesia 

TW: FOT 1975 (1975) 

(371) Elaphoglossum commutatum (METT. ex KUHN) ALDERW. [IUCN: CR (EN)] 


Loc.: 大葉舌蕨蕨 - DaYeSheJue 

Bas.: Acrostichum commutatum METT. ex KUHN // Ann. Mus. Bot. 

Lugduno-Batavi 4: 292 (1869) 

Ety.: commutatum (L: changed); relationship unclear to me, it might 

be relating to its previous inclusion in other taxa (see 

protologue) 

Type: Thwaites 1310 (HT: B; IT: K, ?P) // Sri Lanka 

TW: Kuo, C. M. 1985 (1985) 

(372) Elaphoglossum luzonicum COPEL. [IUCN: VU] 

Pub.: Leafl. Philipp. Bot. 1: 235 (1907) 

Loc.: 台台灣舌蕨蕨 - TaiWanSheJue 

Ety.: relating to Luzon (Philippines), the type location 

Type: Elmer 9036 (HT: MICH; IT: BO, K, L, P, US, Z) // Philippines 

TW: Kuo, C. M. 1985 (1985) 

(373) Elaphoglossum marginatum (WALL. ex FEE) T. MOORE 


Loc.: 垂葉舌蕨蕨 - ChuiYeSheJue 

Bas.: Acrostichum marginatum WALL. // Numer. List n. 17 (1828) ex 

Fee, Mem. Foug. 1: 31 (1845) [non L. (1759) nec Schkuhr 

(1809)] 

Ety.: marginatum (L: with a distinct margin, marginated); probably 

relating to the conspicuous lamina margin (see protologue) 

Type: Wallich 17 (K) // Nepal

84 

TW: Kuo, C. M. 1985 (1985) 

(374) Elaphoglossum sp. 



(375) Elaphoglossum yoshinagae (YATABE) MAKINO 

Pub.: Phan. Pterid. Jap. Icon. 3: pl. 51-52 (1901) 

Loc.: 舌蕨蕨 - SheJue 

Bas.: Acrostichum yoshinagae YATABE // Bot. Mag. (Tokyo) 5: 109- 

112, pl. 23 (1891) 

Ety.: relates to E. Yoshinaga (Japanese botanist, collector of type 

specimen) 

Type: Kochi (Tosa), Mt. Honokawa-yama, E. Yoshinaga, Aug-1887 

(TI) // Japan 

TW: Yamamoto, Y. (1932) 

EQUISETUM (Equisetaceae) 

Ety.: equus (L: horse), seta (L: seta, bristle); relating to the resemblance 

of shoots of some species ("horse's tail") 

(376) Equisetum ramosissimum subsp. debile (ROXB. ex VAUCHER) HAUKE 


Loc.: 台台灣木賊 - TaiWanMuZei 

Bas.: Equisetum debile ROXB. ex VAUCHER // Mem. Soc. Phys. 

Geneve 1: 387 (1822) 

Ety.: debile (L: weak); relating to the stem 

Type: Roxburgh s. n. (BM, ?LINN) // India 

TW: Henry, A. (1896) 

(377) Equisetum ramosissimum DESF. subsp. ramosissimum * 

Pub.: Fl. Atlant. 2: 398-399 (1799) 

Loc.: 木賊 - MuZei 

Ety.: ramosissimum (L: much branched); relating to the plant habit 

Type: Mt. Zaghouan [Mt. Zowran], Desfontaine s. n. (P) // Tunisia 


GRAMMITIS (incl. Chrysogrammitis, Dasygrammitis, Prosaptia, 

Tomophyllum, Xiphopterella) (Grammitidaceae) 

Ety.: gramme (G: line); relating to the sori being elongate to linear (which 

is actually not true for many taxa in this genus) 

Chrysogrammitis: chryso (G: gold), grammitis is relating to genus 

Grammitis (see there); relating to the colour of glands 

Dasygrammitis: dasys (G: hairy), grammitis is relating to genus 

Grammitis (see there); relating to the genus Grammitis in a broad 

sense 

Prosaptia: prosaptien (G: to insert or immerse); relating to the sori, 

which are sunken in the lamina 

Tomophyllum: tomos (G: slice), phyllon (G: leaf); relating to the leaf 

division of the type specimen

Xiphopterella: xiphopteris refers to a fern genus (Grammitidaceae), 

ella (G: diminutive form of genus name Xiphopteris); relating to the 

similarity to members of genus Xiphopteris (xiphos (G: sword), 

pteris relates to genus Pteris, or ferns in general) 

(378) Grammitis adspersa BLUME 

Pub.: 

[IUCN: VU (EN)] 

Fl. Javae Filic. 115, pl. 48, f. 2 (1828); Polypodium adspersum 

BLUME in Enum. Pl. Javae 2: 113 (1828) [non SCHRAD. 

(1818)] is a alter homonym, thus an illegitimate name 

Loc.: 無毛禾葉蕨蕨 - WuMaoHeYeJue 

Ety.: adspersa (L: scattered); relating to the spacing of sori on the 

lamina 

Alt.: Oreogrammitis adspersa (BLUME) PARRIS // Gard. Bull. 

Singapore 58(2): 255 (2007) 

Type: Java, Blume s. n. (LT: L; ILT: L) // Indonesia 

TW: FOT 1975 (1975) 

(379) Grammitis alepidota M. G. PRICE [IUCN: EN] 

Pub.: Philipp. Agric. 57: 34 (1973) 

Loc.: 無鱗禾葉蕨蕨 - WuLinHeYeJue 

Ety.: a (L: without), lepidota (L: covered with small scales); relating 

to the absence of scales on the entire plant 

Alt.: Radiogrammitis alepidota (M. G. PRICE) PARRIS // Gard. Bull. 

Singapore 58(2): 241 (2007) 

Type: Luzon, Mt. Banahaw, Price (K, L, PNH, US) // Philippines 

TW: Moore, S. J. et al. (2009) 

(380) (Grammitis celebica comb. ined.) Prosaptia celebica (BLUME) TAGAWA 

& K. IWATS. [IUCN: - (EN)] 


Loc.: 南亞穴子蕨蕨 - NanYaXueZiJue 

Bas.: Polypodium celebicum BLUME // Enum. Pl. Javae 127 (1828) 

Ety.: relating to Celebes (Indonesia), the type location 

Type: Celebes (L) // Indonesia 


(381) Grammitis congener BLUME 


Loc.: 大武禾葉蕨蕨 - DaWuHeYeJue 

Ety.: congener (L: member of the same genus); relationship unclear 

to me 

Alt.: Oreogrammitis congener (BLUME) PARRIS // Gard. Bull. 

Singapore 58(2): 257 (2007) 

Type: Java, Blume s. n. (LT: L) // Indonesia 

TW: Kuo, C. M. 1985 (1985) 

(382) (Grammitis contigua comb. ined.) Prosaptia contigua (G. FORST.) C. 

PRESL 


Loc.: 穴子蕨蕨 - XueZiJue 

85

86 

Bas.: Trichomanes contiguum G. FORST. // Fl. Ins. Austr. 84 (1786) 

Ety.: contigua (L: adjoining); relating to the pinnae 

Type: "1 Huahine" [Society Islands], Forster, 2 Otaheite [Society 

Islands], Dav. Nelson (LT: BM) // French Polynesia 

TW: Hayata, B. 1917 (1917) 

(383) (Grammitis curtisii comb. ined.) Themelium curtisii (BAKER) PARRIS 

Pub.: Kew Bull. 59(2): 225 (2004) 

Loc.: 蒿蕨蕨 - HaoJue 

Bas.: Polypodium curtisii BAKER // J. Bot. 19: 367-368 (1881) 

Ety.: in honor of Mr. Curtis, collector of the type specimen 

Type: Sumatra, Padang, Curtis 98 [sphalm. "28"] (HT: K) // Indonesia 

TW: FOT 1975 (1975) 

(384) (Grammitis devolii comb. ined.) Xiphopterella devolii S. J. MOORE, 

PARRIS & W. L. CHIOU [IUCN: - (CR)] 

Pub.: Bot. Stud. 54: 24 (2013) 

Loc.: 棣慕氏姬梳葉蕨蕨 - DiMuShiJiShuYeJue 

Ety.: in honor of C. E. DeVol (American botanist) 

Type: Ilan County, Sunglo Lake, S. J. Moore 24567, 20-Jul-2000 

(HT: TAIF; IT: HAST, K, L, TNS, US) // Taiwan 

TW: Bot. Stud. 54: 24 (2013) 

(385) (Grammitis doniana comb. ined.) Tomophyllum donianum (SPRENG.) 

FRASER-JENK. & PARRIS [IUCN: NT (VU)] 

Pub.: Taxon. Revis. Indian Subcontinental Pteridophytes 75 (2008) 

Loc.: 虎虎尾尾蒿蕨蕨 - HuWeiHaoJue 

Bas.: Polypodium donianum SPRENG. // Syst. Veg. (ed. 16) 4(1): 54 

(1827), nom. nov. for Polypodium tenellum D. DON 

Ety.: in honor of D. Don (British botanist) 

Type: Polypodium tenellum D. DON in Podr. Fl. Nepal. 2 (1824): 

Wallich s. n., 1818 (BM) // Nepal 


(386) Grammitis fenicis COPEL. [IUCN: - (CR)] 

Pub.: Univ. Calif. Publ. Bot. 18: 224 (1942) 

Loc.: 短柄禾葉蕨蕨 - DuanBingHeYeJue 

Ety.: in honor of E. Fenix (Philippine botanist) 

Alt.: Oreogrammitis fenicis (COPEL.) PARRIS // Gard. Bull. Singapore 

58(2): 259 (2007). Based on information in FOC is the taxon 

from SE Taiwan a member of genus Radiogrammitis: 

Radiogrammitis moorei PARRIS & RALF KNAPP // Fl. China 2-3: 

842 (2013). 

Type: Batan Isl., Mt. Iraya, Ramos 80147, 17-Apr-1930 (HT: MICH; 

IT: L, NY, UC) // Philippines 

TW: Kuo, C. M. 1985 (1985) 

(387) (Grammitis glandulosa comb. ined.) Chrysogrammitis glandulosa (J. 

SM.) PARRIS [IUCN: EN] 

Pub.: Kew Bull. 53(4): 912-914 (1998)

Loc.: 擬虎虎尾尾蒿蕨蕨 - NiHuWeiGaoJue 

Bas.: Polypodium glandulosum HOOK. [non DESV. (1811)] // Sp. Fil. 

4: 193-194, pl. 276a (1863), validated as Ctenopteris 

glandulosa J. SM. in Hist. Fil. 185 (1875) 

Ety.: glandulosa (L: glandular); relating to the glands of the frond 

Type: Gardner & Thwaites 1289 (LT: K; ILT: B, BM, CGE, GH, K, 

PDA) // Sri Lanka 

TW: Kuo, C. M. 1985 (1985) 

(388) Grammitis intromissa (CHRIST) PARRIS * 


Pub.: Fern Gaz. 12(3): 180 (1981) 

Loc.: 大禾葉蕨蕨 - DaHeYeJue 

Bas.: Polypodium intromissum CHRIST // Verh. Naturf. Ges. Basel 

11: 440-441 (1896) 

Ety.: intromissa (L: introduced into); relating to the intermediate 

position between two taxa as discussed in the protologue 

Alt.: Radiogrammitis setigera (BLUME) PARRIS // Gard. Bull. 

Singapore 58(2): 244 (2007), based on Polypodium 

setigerum BLUME in Enum. Pl. Javae 123 (1828), type: Blume 

s. n. (LT: L; ILT: K, L) // Indonesia 

Type: Celebes, Wawokaraeng, Sarasin 1368 (BAS) // Indonesia 

TW: Kuo, C. M. 1985 (1985) 

(389) Grammitis jagoriana (METT. ex KUHN) COPEL. [IUCN: DD (VU)] 


Loc.: 擬禾葉蕨蕨 - NiHeYeJue 

Bas.: Polypodium jagorianum METT. ex KUHN // Linnaea 36: 128 

(1869) 

Ety.: in honor of A. F. Jagor (German botanist and collector of the 


Alt.: Radiogrammitis jagoriana (METT. ex KUHN) PARRIS // Gard. 

Bull. Singapore 58(2): 242 (2007). Based on information in 

FOC is this name misapplied, and is to be substituted (at 

least) by the following name: Radiogrammitis taiwanensis 

PARRIS & RALF KNAPP // Fl. China 2-3: 843 (2013). 

Type: Luzon, F. Jagor 835, 1861 (HT: B) // Philippines 

TW: Tagawa, M. 1940-49 (1941) 

(390) (Grammitis mollicoma comb. ined.) Dasygrammitis mollicoma (NEES & 

BLUME) PARRIS [IUCN: - (EN)] 

Pub.: Gard. Bull. Singapore 58(2): 239 (2007) 

Loc.: 南洋蒿蕨蕨 - NanYangHaoJue 

Bas.: Polypodium mollicomum NEES & BLUME // Nova Acta 11: 121, 

pl. 12, f. 2 (1823) 

Ety.: mollis (L: soft), coma (L: hair-tufted); relating to the frond hairs 

Type: Java, Blume s. n. (L, NY) // Indonesia 

TW: Kuo, C. M. 1985 (1985) 

87

88 

(391) Grammitis nuda TAGAWA * 

[IUCN: DD (EN)] 


Loc.: 長孢孢禾葉蕨蕨 - ChangBaoHeYeJue 

Ety.: 

Alt.: 

nuda (L: naked); relating to the absence of lamina hairs 

Oreogrammitis nuda (TAGAWA) PARRIS // Gard. Bull. Singapore 

58(2): 264 (2007) 

Type: Prov. Takao, between Daizyurin and Taito border, ca. 1000 m, 

Tagawa 2092, 22-Jan-1939 (HT: KYO) // Taiwan 


(392) (Grammitis nutans comb. ined.) Prosaptia nutans (BLUME) METT. * 


Pub.: Reise Novara 1: 214 (1870) 

Loc.: 俯垂穴子蕨蕨 - FuChuiXueZiJue 

Bas.: Polypodium nutans BLUME // Enum. Pl. Javae 128 (1828) 

Ety.: nutans (L: nodding); relating to the frond shape 


TW: FOC (2013) 

(393) Grammitis obliquata (BLUME) HASSK. 

Pub.: Cat. Hort. Bot. Bogor. 4 (1844) 

Loc.: 密密毛蒿蕨蕨 - MiMaoHaoJue 

Bas.: Polypodium obliquatum BLUME // Enum. Pl. Javae: 128 (1828) 

Ety.: obliquata (L: obliquely positioned); relating to the sori 

Alt.: Prosaptia obliquata (BLUME) METT. // Reise Novara 1: 214 

(1870) 



(394) Grammitis okuboi (YATABE) CHING 


Loc.: 梳葉蕨蕨 - ShuYeJue 

Bas.: Polypodium okuboi YATABE // Bot. Mag. (Tokyo) 5: 35-38, pl. 

21 (1891) 

Ety.: in honor of S. Okubo (Japanese botanist and collector of the 


Alt.: Micropolypodium okuboi (YATABE) HAYATA // Bot. Mag. (Tokyo) 

42: 341 (1928) 

Type: Honsyu, near Ashinoyu, S. Okubo, 1888 (HT: TI) // Japan 

TW: Tagawa, M. 1940-49 (1941) 

(395) Grammitis reinwardtii BLUME [IUCN: NT (VU)] 

Pub.: Enum. Pl. Javae, Add. 2 (1828) 

Loc.: 毛禾葉蕨蕨 - MaoHeYeJue 

Ety.: in honor of C. G. C. Reinwardt (Dutch botanist) 

Alt.: Oreogrammitis reinwardtii (BLUME) PARRIS // Gard. Bull. 

Singapore 58(2): 266 (2007) 

Type: Celebes, G. Klabat, Reinwardt 104 (LT: L) // Indonesia 

TW: FOT 1975 (1975)

(396) Grammitis tenuisecta (BLUME) CHING [IUCN: VU] 


Loc.: 細葉蒿蕨蕨 - XiYeHaoJue 

Bas.: Polypodium tenuisectum BLUME // Enum. Pl. Javae 134 (1828) 

Ety.: tenuisectum (L: finely divided); relating to the lamina 

Alt.: Themelium tenuisectum (BLUME) PARRIS // Kew Bull. 52(3): 740 

(1997) 


TW: Tagawa, M. 1940-49 (1941) 

(397) (Grammitis urceolaris comb. ined.) Prosaptia urceolaris (HAYATA) 

COPEL. [IUCN: NT (EN)] 


Loc.: 台台灣穴子蕨蕨 - TaiWanXueZiJue 

Bas.: Polypodium urceolare HAYATA // Icon. Pl. Formosan. 5: 324- 

Ety.: 

332, f. 136-137a-f (1915) 

urceolus (L: pitcher), aris (L: resembling); relating to the shape 

of involucre 

Type: Mt. Arisan, prope Senninbora, T. Ito, 2-Apr-1914 (HT: TI) // 


TW: Icon. Pl. Formosan. 5: 324-332, f. 136-137a-f (1915) 

GYMNOCARPIUM (Dryopteridaceae) 

Ety.: gymnos (G: naked), karpos (G: fruit); relating to the sori lacking 

indusia 

(398) Gymnocarpium oyamense (BAKER) CHING [IUCN: CR] 

Pub.: Contr. Biol. Lab. Sci. Soc. China 9: 40, f. 3 (1933) 

Loc.: 羽節蕨蕨 - YuJieJue 

Bas.: Polypodium oyamense BAKER // J. Bot. 15: 366 (1877) 

Ety.: relating to Oyama, a place in Japan and the type location 

Type: Japan 

TW: FOT 1975 (1975) 

(399) Gymnocarpium remotepinnatum (HAYATA) CHING 

Pub.: Icon. Fil. Sin. 4: pl. 172 (1937) 

Loc.: 細裂羽節蕨蕨 - XiLieYuJieJue 

Bas.: Dryopteris remotepinnata HAYATA // Icon. Pl. Formosan. 6 

(suppl.): 108 (1917) 

Ety.: remotum (L: distant), pinnatum (L: pinnate); relating to the 

lamina shape (bearing spaced pinnae) 

Type: Mt. Morrison, Nakahara s. n., Oct-1905 // Taiwan 

TW: Icon. Pl. Formosan. 6 (suppl.): 108 (1917) 

HELMINTHOSTACHYS (Ophioglossaceae) 

Ety.: helminthos (G: worm), stachys (G: spike); relating to the fertile 

section where sporangia are in compact spikes, worm-like 

89

90 

(400) Helminthostachys zeylanica (L.) HOOK. [IUCN: CR (EN)] 

Pub.: Gen. Fil. (Hooker) pl. 47 (1840) 

Loc.: 錫蘭蘭七指蕨蕨 - XiLanQiZhiJue 

Bas.: Osmunda zeylanica L. // Sp. Pl. 2: 1063 (1753) 

Ety.: relating to Sri Lanka ("Ceylon"), the type location 

Type: (BM) // Sri Lanka 

TW: Yabe, Y. (1902) 

HISTIOPTERIS (Dennstaedtiaceae) 

Ety.: histion (G: sail), pteris (relates to genus Pteris, or ferns in general); 

relating to the sail-like basal pinnae 

(401) Histiopteris incisa (THUNB.) J. SM. 

Pub.: Hist. Fil. 295 (1875) 

Loc.: 栗蕨蕨 - LiJue 

Bas.: Pteris incisa THUNB. // Prodr. Pl. Cap. 171 (1800) 

Ety.: incisa (L: cut in deeply and sharply); relating to the lamina 

habit 

Type: Cape of Good Hope (Cap. Bon. Spei) Thunberg s. n., 1774 

(HT: UPS; IT: S) // South Africa 

TW: J. Bot. 23: 103 (1885) 

HUPERZIA (Lycopodiaceae) 

Ety.: in honor of J. P. Huperz (German botanist) 

(402) Huperzia appressa (DESV.) A. LÖVE & D. LÖVE [IUCN: VU] 

Pub.: Bot. Not. 114: 34 (1961) 

Loc.: 小小杉葉石松 - XiaoShanYeShiSong 

Bas.: Lycopodium selago var. appressum DESV. // Mem. Soc. Linn. 

Paris 6: 180 (1827) 

Ety.: appressa (L: kept down); relating to the orientation of 

trophophylls 

Type: St. Pierre et T. N., B. de La Pilaye s. n. (P) 

TW: FOT 1975 (1975) 

(403) Huperzia carinata (DESV. ex POIR.) TREV. [IUCN: EN] 

Pub.: Atti Soc. Ital. Sci. Nat. 17: 247 (1874) 

Loc.: 覆葉石松 - FuYeShiSong 

Bas.: Lycopodium carinatum DESV. ex POIR. // Encycl., Suppl. 3: 555 

(1813 [1814]) 

Ety.: carinata (L: keeled); relating to the leaves 

Type: Jussieu 639 (HT: P; IT: B) // Sri Lanka 

TW: Henry, A. (1896) 

(404) Huperzia changii (T. Y. HSIEH) RALF KNAPP (comb. nov.) * 

[IUCN: - (?EW)] 

Pub.: Index Ferns Fern Allies Taiwan 164 (2014) 

Loc.: 張氏馬尾尾杉 - ZhangShiMaWeiShan 

Bas. Phlegmariurus changii T. Y. HSIEH // Amer. Fern J. 102(4): 

284-287, f. 1-2 (2012)

Ety.: in honor of L. R. Chang (Taiwanese botanist who first 

observed this taxon) 

Type: Hualien County, Wanrong Township, Hsilin Village, Tung-Yu 

Hsieh 516, 6-Apr-2006 (HT: TAI) // Taiwan 

TW: Amer. Fern J. 102(4): 284-287, f. 1-2 (2012) 

(405) Huperzia cryptomerina (MAXIM.) R. D. DIXIT [IUCN: DD (EN)] 

Pub.: J. Bombay Nat. Hist. Soc. 77(3): 541 (1981) 

Loc.: 柳杉葉蔓石松 - LiuShanYeManShiSong 

Bas.: Lycopodium cryptomerinum MAXIM. // Bull. Acad. Imp. Sci. 

Saint-Petersbourg 15: 231 (1870) 

Ety.: cryptomeria relating to a gymnosperm genus (Taxodiaceae, 

"Japanese cedar"); relating to the similarity of branches to 

that of genus Cryptomeria 

Type: Naga-yama, 1862 (LT: LE), designated by J. A. Ivanenko // 

Japan 

TW: FOT 1975 (1975) 

(406) Huperzia cunninghamioides (HAYATA) HOLUB * 

[IUCN: CR] 

Pub.: Folia Geobot. Phytotax. 20(1): 72 (1985) 

Loc.: 寬寬葉石松 - KuanYeShiSong 

Bas.: Lycopodium cunninghamioides HAYATA // Icon. Pl. Formosan. 

4: 131 (1914) 

Ety.: cunninghamia relating to a gymnosperm genus (Taxodiaceae, 

"China-fir"), oides (L: suffix, denoting likeness of form); 

relating to the similarity of branches to that of genus 

Cunninghamia 

Type: Urai, T. Soma no. 1, Jun-1910 (ST: TI) // Taiwan 


(407) Huperzia fargesii (HERTER) HOLUB [IUCN: EN (VU)] 

Pub.: Folia Geobot. Phytotax. 20: 72 (1985) 

Loc.: 銳葉石松 - RuiYeShiSong 

Bas.: Lycopodium fargesii HERTER // Bot. Jahrb. Syst. 43, Beibl. 98: 

48-49 (1909) 

Ety.: in honor of P. P. G. Farges (French botanist and the collector 


Type: Sichuan, Farges 1160 (HT: P; IT: L, MO) // China 

TW: FOT 1975 (1975) 

(408) Huperzia fordii (BAKER) DIXIT 

Pub.: Census Indian Pterid. 7 (1984) 

Loc.: 福氏石松 - FuShiShiSong 

Bas.: Lycopodium fordii BAKER // Handb. Fern-Allies 17 (1887) 

Ety.: in honor of C. Ford (British botanist and collector of the type 

specimen) 

Type: Guangdong, Lofo Shan, Ford 4 (HT: K) // China 

TW: FOT 1975 (1975) 

91

92 

(409) Huperzia phlegmaria (L.) ROTHM. [IUCN: EN] 

Pub.: Feddes Repert. Spec. Nov. Regni Veg. 54: 62 (1944) 

Loc.: 垂枝石松 - ChuiZhiShiSong 

Bas.: Lycopodium phlegmaria L. // Sp. Pl. 2: 1101 (1753) 

Ety.: phlegma (G: mucus); probably relating to its medical usage 

Type: Dillenius, Hist. Musc. Tab. 61, figs. A-C (1741), LT chosen by 

Oellgaard (1989) // India (fide Field, A. R. & Bostock, P. D. 

2013) 

TW: Hayata, B. 1911 (1911) 

(410) Huperzia quasipolytrichoides (HAYATA) CHING 

Pub.: Acta Bot. Yunnan. 3(3): 299 (1981) 

Loc.: 反反捲葉石杉 - FanJuanYeShiShan 

Bas.: Lycopodium quasipolytrichoides HAYATA // Icon. Pl. Formosan. 

5: 252-254, f. 89 (1915) 

Ety.: quasi (L: as if, nearly), polytrichum (a moss genus, 

Polytrichaceae, "haircap moss"), oides (L: suffix, denoting 

likeness of form); relating to the similarity of branches to that 

of genus Polytrichum (G: poly, many, and thrix, hair) 

Type: Mt. Arisan, ad 7500 ped. alt., T. Ito & B. Hayata s. n., 4-Apr- 

1914 (HT: TI; IT: TAIF) // Taiwan 


(411) Huperzia salvinioides (HERTER) HOLUB [IUCN: EN] 


Loc.: 小小垂枝石松 - XiaoChuiZhiShiSong 

Bas.: Urostachys salvinioides HERTER // Philipp. J. Sci. 22(1): 67 

(1923) 

Ety.: salvinia relating to a fern genus (see there), oides (L: suffix, 

denoting likeness of form); relating to the similarity of 

branches to plants of genus Salvinia 

Type: Herter lists many syntypes, no lectotype might have ever been 

designated // Philippines 

TW: FOT 1975 (1975) 

(412) Huperzia serrata (THUNB.) TREV. [IUCN: DD (LC)] 

Pub.: Atti Soc. Ital. Sci. Nat. 17: 247-248 (1875) 

Loc.: 千層塔 - QianCengTa 

Bas.: Lycopodium serratum THUNB. // Syst. Veg. (ed. 14) 944 (1784) 

Ety.: serrata (L: serrated); relating to the margin of pinna 

Type: Thunberg s. n. (HT: UPS) // Japan 


(413) Huperzia sieboldii (MIQ.) HOLUB [IUCN: EN (VU)] 


Loc.: 鱗葉石松 - LinYeShiSong 

Bas.: Lycopodium sieboldii MIQ. // Ann. Mus. Bot. Lugduno-Batavi 3: 

184 (1867)

Ety.: in honor of P. F. Siebold (German botanist and the collector of 


Type: Honshu, P. F. Siebold s. n. (IT: L, fragment in US) // Japan 


(414) Huperzia somae (HAYATA) CHING 

Pub.: Acta Bot. Yunnan. 3: 301 (1981) 

Loc.: 相馬氏石杉 - XiangMaShiShiShan 

Bas.: Lycopodium somae HAYATA // Icon. Pl. Formosan. 5: 255, f. 91 

(1915) 

Ety.: relating to T. Soma (Japanese botanist, and collector of the 


Type: Mt. Arisan, T. Soma s. n., 1915 (HT: TI) // Taiwan 


(415) Huperzia squarrosa (G. FORST.) TREVIS. [IUCN: EN] 

Pub.: Atti Soc. Ital. Sci. Nat. 17: 247 (1875) 

Loc.: 杉葉石松 - ShanYeShiSong 

Bas.: Lycopodium squarrosum G. FORST. // Fl. Ins. Austr. 86 (1786) 

Ety.: squarrosa (L: rough with objects projecting outwards); relating 

to the leaves 

Type: J. & G. Forster s. n. (IST: BM, P) // Tahiti 

TW: Henry, A. (1896) 

HYMENOPHYLLUM (Hymenophyllaceae) 

Ety.: hymen (G: membrane), phyllon (G: leaf); relating to texture of fronds 

("filmy-leaf fern") 

(416) Hymenophyllum alishanense DeVol 


Pub.: Taiwania 15: 283-284, f. 6 (1970) 

Loc.: 阿里山膜蕨蕨 - ALiShanMoJue 

Ety.: relating to ALiShan, a mountain range in Taiwan, and the type 

location 

Type: Alishan, Ito 25 (HT: TAI; IT: TAI) // Taiwan 

TW: Taiwania 15: 283-284, f. 6 (1970) 

(417) Hymenophyllum badium HOOK. & GREV. 

Pub.: Icon. Filic. 1: pl. 76 (1828) 

Loc.: 蕗蕨蕨 - LuJue 

Ety.: badium (L: chestnut-coloured, brown); relating to the colour of 

dried fronds 

Type: Sikkim, Herb. Schlaymhreit (K) // India 

TW: Ito, H. 1944 (1944) 

(418) Hymenophyllum barbatum (BOSCH) BAKER 

Pub.: Syn. Fil., ed. 2: 68 (1868) 

Loc.: 華東膜蕨蕨 - HuaDongMoJue 

Bas.: Leptocionium barbatum BOSCH // Ned. Kruidk. Arch. 5: 416 

(1863) 

Ety.: barbatum (L: bearded); relating to the lamina margin 

93

94 

Type: Tsus-Sima, Wilford 846 (HT: K; IT: B, GH, L) // Japan 


(419) Hymenophyllum blandum RACIB. [IUCN: NT] 

Pub.: Pteridoph. Buitenzorg 20 (1898) 

Loc.: 爪哇厚厚壁蕨蕨 - ZhuaWaHouBiJue 

Ety.: blandum (L: smooth); probably relating to a lamina without 

hairs and wrinkles 

Type: Java, Raciborski s. n. (BO, K, L) // Indonesia 


(420) Hymenophyllum denticulatum SW. 

[IUCN: NT] 


Loc.: 厚厚壁蕨蕨 - HouBiJue 

Ety.: denticulatum (L: with very small teeth); relating to the lamina 

margin 

Type: Thunberg s. n., Java // Indonesia 


(421) Hymenophyllum devolii M. J. LAI [IUCN: DD (NT)] 


Loc.: 棣氏膜蕨蕨 - LiShiMoJue 


Type: Taitung, Chingshuiying, Liew 45 (HT: TAI; IT: TAI) // Taiwan 

TW: Taiwania 21: 82 (1976) 

(422) Hymenophyllum fimbriatum J. SM. 

[IUCN: VU] 

Pub.: J. Bot. (Hooker) 3: 418 (1841) 

Loc.: 叢葉蕗蕨蕨 - CongYeLuJue 

Ety.: fimbriatum (L: fimbriate), relating to the involucre margin 

Type: Luzon, Cuming 218 (IT: B, BM, K, L) // Philippines 

TW: Tagawa, M. 1940-49 (1940) 

(423) Hymenophyllum holochilum (BOSCH) C. CHR. 


Loc.: 南洋厚厚壁蕨蕨 - NanYangHouBiJue 

Bas.: Didymoglossum holochilum BOSCH // Pl. Jungh. 1: 561-566 

(1856) 

Ety.: holos (G: entire, whole), cheilos (G: margin); relating to the 

involucre margin 

Type: Java, Hasskarl s. n. (L) // Indonesia 

TW: Ito, H. 1944 (1944) 

(424) Hymenophyllum javanicum SPRENG. [IUCN: NT (VU)] 

Pub.: Syst. Veg. 4: 132 (1827) 

Loc.: 爪哇蕗蕨蕨 - ZhuaWaLuJue 


Type: Java, Nees s. n. (?L) // Indonesia 

TW: Hayata, B. 1914-21 (1914) 

(425) Hymenophyllum oligosorum MAKINO 

Pub.: Bot. Mag. (Tokyo) 13: 44-45 (1899)

Loc.: 

Ety.: 

Type: 

95 

長毛蕗蕨蕨 - ChangMaoLuJue 

oligos (G: few), soros (G: sorus); relating to the shape of 

lamina (bearing only relatively few sori) 

Mt. Kiyosumi-yama, Makino s. n., 6-Apr-1896 (LT: TI), 

designated by Iwatsuki, J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 

13: 520 (1985) // Japan 


(426) Hymenophyllum paniculiflorum C. PRESL 

Pub.: Hymenophyllaceae 55 (1843) 

Loc.: 

Ety.: 

圓錐孢孢膜蕨蕨 - YuanZhuiBaoMoJue 

panicula (L: tuft), flos (L: flower); relating to the fertile section 

of lamina 

Cuming 214 (B, BM, HUH, P) // Philippines 

Type: 

TW: Kuo, C. M. 1985 (1985) 

(427) Hymenophyllum polyanthos (SW.) SW. 


Loc.: 細葉蕗蕨蕨 - XiYeLuJue 

Bas.: Trichomanes polyanthos SW. // Prodr. 137 (1788) 

Ety.: poly (G: many), anthos (L: flower); relating to the abundance of 

sori 

Type: Swartz s. n. (HT: S; IT: BM) // Jamaica 

TW: J. Bot. 23: 103 (1885) 

(428) Hymenophyllum productum KUNZE [IUCN: DD] 

Pub.: Bot. Zeitung (Berlin) 6: 305-306 (1848) 

Loc.: 南洋蕗蕨蕨 - NanYangLuJue 

Ety.: productum (L: lengthened); relating to the terminal lamina 

segment, typically longer than other segments 

Type: Java, Zollinger 74, 363 (IST: P) // Indonesia 

TW: Tagawa, M. 1940-49 (1940) 

(429) Hymenophyllum simonsianum HOOK. [IUCN: DD] 

Pub.: Sec. Cent. Ferns pl. 13 (1861) 

Loc.: 寬寬片膜蕨蕨 - KuanPianMoJue 

Ety.: in honor of C. J. Simons (British botanist and collector of the 


Type: Khasya Hills, Simons s. n. (LT: K) // India 

TW: Hayata, B. 1914-21 (1915) 

(430) Hymenophyllum taiwanense (TAGAWA) C. V. MORTON [IUCN: DD] 

Pub.: Contr. U. S. Nat. Herb. 38: 167 (1968) 

Loc.: 台台灣蕗蕨蕨 - TaiWanLuJue 

Bas.: Mecodium taiwanense TAGAWA // Acta Phytotax. Geobot. 9: 

141 (1940) 


Type: Prov. Taito, near Kakayo, Kwanzan-gun, ca. 1100 m, Tagawa 

3144, 3-Mar-1940 (HT: KYO) // Taiwan 

TW: Acta Phytotax. Geobot. 9: 141 (1940)

96 

HYPODEMATIUM (Dryopteridaceae) 

Ety.: hypodemation (G: small slipper); relating to the shape of indusium 

(431) Hypodematium crenatum (FORSSK.) KUHN & DECKEN 

Pub.: Reisen Ost-Afr. 3: 37, f. a (1879) 

Loc.: 腫足蕨蕨 - ZhongZuJue 

Bas.: Polypodium crenatum FORSSK. // Fl. Aegypt.-Arab. 185 (1775) 

Ety.: crenatum (L: notched); relating to the margin of segments 

Type: Bolghose, Forsskal s. n. (?type lost) // Yemen 

TW: Hayata, B. 1914-21 (1914) 

HYPOLEPIS (Dennstaedtiaceae) 

Ety.: hypo (G: under), lepis (G: scale); relating to the sporangia under 

scale-like leaf margin or false indusium 

(432) Hypolepis alpina (BLUME) HOOK. 

Pub.: Sp. Fil. 2: 63 (1852) 

Loc.: 台台灣姬蕨蕨 - TaiWanJiJue 

Bas.: Cheilanthes alpina BLUME // Enum. Pl. Javae 138 (1828) 

Ety.: alpina (L: alpine), relating to the alpine growth habit (described 

from the summit area of Mt. Gede in Indonesia, ca. 2900 m 

altitude) 


TW: Blumea 32: 227-276 (1987) 

(433) Hypolepis pallida (BLUME) HOOK. [IUCN: - (CR)] 

Pub.: Sp. Fil. 2: 64 (1852) 

Loc.: 灰姬蕨蕨 - HuiJiJue 

Bas.: Cheilanthes pallida BLUME // Enum. Pl. Javae 139 (1828) 

Ety.: pallida (L: pale); relating to the colour of lamina (see 

protologue) 

Type: Java, C. L. Blume s. n. (LT: L) // Indonesia 

TW: Blumea 32: 227-276 (1987) 

(434) Hypolepis polypodioides (BLUME) HOOK. 

Pub.: Sp. Fil. 2: 63 (1852) 

Loc.: 無腺姬蕨蕨 - WuXianJiJue 

Bas.: Cheilanthes polypodioides BLUME // Enum. Pl. Javae 139 

(1828) 

Ety.: polypodium relates to a fern genus, oides (L: suffix, denoting 

likeness of form); relating to the similarity to a - at that time 

broadly defined - genus Polypodium (including many ferns 

without indusia, as well large compound species) 

Type: Java, Tjiruk tjipanna, C . L. Blume (LT: L) // Indonesia 

TW: Blumea 32: 227-276 (1987) 

(435) Hypolepis punctata (THUNB.) METT. ex KUHN 

Pub.: Filic. Afr. 120 (1868) 

Loc.: 姬蕨蕨 - JiJue

Bas.: Polypodium punctatum THUNB. // Fl. Jap. 337-338 (1784) 

Ety.: punctata (L: spotted); relating to the stipe (see protologue) 



(436) Hypolepis tenuifolia (G. FORST.) BERNH. 

Pub.: Neues J. Bot. 1(2): 34 (1806) 

Loc.: 細葉姬蕨蕨 - XiYeJiJue 

Bas.: Lonchitis tenuifolia G. FORST. // Flora Ins. Austr. 80 (1786) 

Ety.: tenuis (L: thin), folium (L: leaf); relating to the texture of lamina 

Type: Pacific Islands, Forster s. n. (LT: BM), designated by 

Brownsey & Chinnock (1987) 

TW: a: J. Bot. 23: 103 (1885), probably a misidentification only 

b: Henry, A. (1896), second location refers to "Bankinsing, S. 

Cape" in S Taiwan and might be thus correctly identified 

ISOETES 

Ety.: 

(Isoetaceae) 

isos (G: equal), etes (G: year); relating to that leaves are green all 

the year (in many species) 

(437) Isoetes taiwanensis var. kinmenensis F. Y. LU, H. H. CHEN & Y. L. 

HSUEH [IUCN: - (CR)] 

Pub.: Flora of Kinmen 26 (2011) 

Loc.: 

Ety.: 

Type: 

金門水韭 - JinMenShuiJiu 

relating to JinMen (Kinmen), small archipelago of islands 

administered by Taiwan, the type (and only) location of this 

taxon 

Kinmen, inter mont. Taiwu, W. L. Hsieh et al. H2747, 5-Mar- 

2010 (HT: CHIA) // Taiwan 

TW: Flora of Kinmen 26 (2011) 

(438) Isoetes taiwanensis DEVOL var. taiwanensis * 

[IUCN: CR] 

Pub.: Taiwania 17(1): 1-7, f. 1 (1972) 

Loc.: 台台灣水韭 - TaiWanShuiJiu 

Ety.: relating to Taiwan, the type (and only) location of this taxon 

Type: Chihsingshan, Hsu & Chang 1715 (HT: TAI; IT: BM, TAI, TAIF) 


TW: Taiwania 17(1): 1-7, f. 1 (1972) 

LASTREOPSIS (Dryopteridaceae) 

Ety.: lastrea is in honour of C. J. L. de Lastre (French botanist), opsis (G: 

like); relating to the similarity to members of fern genus Lastrea 

(Thelypteridaceae) 

(439) Lastreopsis tenera (R. BR.) TINDALE [IUCN: NT] 

Pub.: Victoria Naturalist 73: 181 (1957) 

Loc.: 金毛蕨蕨 - JinMaoJue 

Bas.: Nephrodium tenerum R. BR. // Prodr. 149 (1810) 

Ety.: tener (L: soft, tender); probably relating to the texture of lamina 

97

98 

Type: Queensland, R. Brown 23 (BM, ?E, K) // Australia 

TW: FOT 1975 (1975) 

LEMMAPHYLLUM (Polypodiaceae) 

Ety.: lemma (G: scale), phyllon (G: leaf); relating to the sori containing 

scales (paraphyses) 

(440) Lemmaphyllum microphyllum C. PRESL 

Pub.: Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6: 623 (1851) 

Loc.: 抱樹蕨蕨 - BaoShuJue 

Ety.: mikros (G: small), phyllon (G: leaf); relating to the habit of 

(sterile) fronds 

Type: von Siebold s. n. (L, PRC) // Japan 


(441) Lemmaphyllum rostratum (BEDD.) TAGAWA 

Pub.: Fl. E. Himalaya 493 (1966) 

Loc.: 骨牌蕨蕨 - GuPaiJue 

Bas.: Pleopeltis rostrata BEDD. // Ferns Brit. India pl. 159 (1866) 

Ety.: rostratum (L: beaked); relating to the lamina apex 

Type: Khasya, Thomson s. n. (K, W) // India 

TW: Hayata, B. 1914-21 (1914) 

LEPISORUS (Polypodiaceae) 

Ety.: lepis (G: scale), soros (G: sorus); relating to the presence of scales 

(paraphyses) in the sorus 

(442) Lepisorus clathratus (C. B. CLARKE) CHING 


Loc.: 網眼瓦葦 - WangYanWaWei 

Bas.: Polypodium clathratum C. B. CLARKE // Trans. Linn. Soc. 

London, Bot. 1: 559, pl. 82, f. 1 (1880) 

Ety.: clathratus (L: latticed); relating to the scale habit 

Type: Kashmir, Pir Pinjul (Pir Panjal Range), alt. 11,000-12,000 feet, 

Clarke s. n. (?K) // Pakistan 

TW: Ito, T. (1928) 

(443) Lepisorus kawakamii (HAYATA) TAGAWA 


Loc.: 鱗瓦葦 - LinWaWei 

Bas.: Polypodium kawakamii HAYATA // Bot. Mag. (Tokyo) 23: 77 

(1909) 

Ety.: in honor of T. Kawakami (Japanese botanist, and (co-) 

collector of type specimen) 

Type: Toroku, Nanshikiaku, Kawakami & Mori 2346, Nov-1906 (LT: 

TI), designated by M. J. Zink (1993) // Taiwan 

TW: Bot. Mag. (Tokyo) 23: 77 (1909) 

(444) Lepisorus kuchenensis (Y. C. WU) CHING [IUCN: DD] 

Pub.: Bull. Fan Mem. Inst. Biol. 4: 69-70 (1933)

Loc.: 

Bas.: 

Ety.: 

Type: 

猺山瓦葦 - YaoShanWaWei 

Polypodium kuchenense Y. C. WU // Bull. Dept. Biol. Sun 

Yatsen Univ. 3: 276, pl. 129 (1932) 

relating to "Ku-chen" (China, type location) 

Guangxi, Yao Shan, Ku-chen, Sin & Wang 2121 (HT: PE; IT: 

IBK) // China 

TW: Tagawa, M. 1940-49 (1942) 

(445) Lepisorus megasorus (C. CHR.) CHING 


Loc.: 長柄瓦葦 - ChangBingWaWei 

Bas.: Polypodium megasorum C. CHR. // Index Filic. 544 (1906), 

based on later homonym P. macrosorum BAKER [not FEE 

(1850)] 

mega (G: big), soros (G: sorus); relating to the size of sorus 

Tamsui, Hancock 31 (HT: K; IT: GH, US) // Taiwan 

Ety.: 

Type: 

TW: Index Filic. 544 (1906) 

(446) Lepisorus monilisorus (HAYATA) TAGAWA 


Loc.: 

Bas.: 

Ety.: 

擬芨瓦葦 - NiJiWaWei 

Polypodium lineare var. monilisorum HAYATA // Icon. Pl. 

Formosan. 4: 248-249, f. 173 (1914) 

monilis (G: necklace), soros (G: sorus); relating to fertile 

lamina habit (sori arranged like in a necklace) 

Type: monte Arisan, B. Hayata & S. Sasaki s. n., Jan-1912 (HT: TI) // 



(447) Lepisorus morrisonensis (HAYATA) H. ITO 

Pub.: J. Jap. Bot. 11: 92 (1935) 

Loc.: 玉山瓦葦 - YuShanWaWei 

Bas.: Polypodium morrisonense HAYATA // Bot. Mag. (Tokyo) 23: 77- 

78 (1909) 

Ety.: relating to YuShan ("Mt. Morrison"), the highest mountain of 

Taiwan and type location 

Type: in monte Morrison, ad 9000 ped alt., Kawakami & Mori 1819, 

18-Oct-1906 (HT: TI) // Taiwan 

TW: Bot. Mag. (Tokyo) 23: 77-78 (1909) 

(448) Lepisorus obscurevenulosus (HAYATA) CHING 


Loc.: 

Bas.: 

Ety.: 

奧瓦葦 - AoWaWei 

Polypodium obscurevenulosum HAYATA // Icon. Pl. Formosan. 

5: 322-323, f. 134A/B (1915) 

obscurus (L: dusky, dark), venulosus (L: with a network of 

veinlets); relating to the venation habit of lamina 

Type: Taichu, Kashigatam, Nakahara 5926, Feb-1907 (HT: TI) // 


99

100 

TW: Icon. Pl. Formosan. 5: 322-323, f. 134A/B (1915) 

(449) Lepisorus pseudoussuriensis TAGAWA 


Loc.: 擬烏蘇蘇里瓦葦 - NiWuSuLiWaWei 

Ety.: pseudos (G: false), ussuriensis (refering to the Ussuri River in 

China) relating to the taxon with epithet "ussuriensis"; relating 

to the similarity to taxon Lepisorus ussuriensis (REGEL & 

MAACK) CHING 

Type: Mt. Arisan, Tozan, Tagawa 530, 20-Aug-1934 (HT: KYO; IT: P) 



(450) Lepisorus suboligolepidus CHING 


Loc.: 擬鱗瓦葦 - NiLinWaWei 

Ety.: sub (L: similar to), oligolepidus (G: oligos, few, and lepidos, 

scale) relating to taxon with the epithet "oligolepidus"; relating 

to the similarity to taxon Lepisorus oligolepidus (BAKER) 

CHING 

Type: Yunnan, A. Henryi 10088A (?K, ?US, PE) // China 

TW: Tagawa, M. 1940-49 (1942) 

(451) Lepisorus thunbergianus (KAULF.) CHING 


Loc.: 瓦葦 - WaWei 

Bas.: Pleopeltis thunbergiana KAULF. // Wesen Farrenk. 113 (1827), 

based on Polypodium lineare THUNB. in Syst. Veg. (ed. 14): 

934 (1784) [not N. F. Burman (1768)] 

Ety.: in honor of C. P. Thunberg (Swedish naturalist) 

Type: Kosido s. n. (HT: UPS) // Japan 

TW: Ito, H. 1944 (1944) 

(452) Lepisorus tosaensis (MAKINO) H. ITO 

Pub.: J. Jap. Bot. 11: 93 (1935) 

Loc.: 擬瓦葦 - NiWaWei 

Bas.: Polypodium tosaense MAKINO // Bot. Mag. (Tokyo) 27: 127 

(1913) 

Ety.: 

Type: 

relating to Tosa (Japan), the type location 

Prov. Tosa, Mt. Honokawa, Makino s. n., Aug-1889 (ST: MAK, 

IST: TI) // Japan 

TW: Tagawa, M. 1940-49 (1942) 

LEPTOCHILUS (Polypodiaceae) 

Ety.: leptos (G: slender), cheilos (G: lip); relating to the lamina, which 

ends in a slender curled tip 

(453) Leptochilus decurrens BLUME 


Loc.: 萊蕨蕨 - LaiJue

Ety.: decurrens (L: running down); relating to the basal lamina of 

sterile frond 



LEUCOSTEGIA (Davalliaceae) 

Ety.: leukos (G: white), stege (G: cover); relating to the pale-coloured 

(almost white) indusium 

(454) Leucostegia truncata (D. DON) FRASER-JENK. * 


Loc.: 大膜蓋蕨蕨 - DaMoGaiJue 

Bas.: Davallia truncata D. DON // Prodr. Fl. Nepal.: 10 (1825) 

Ety.: truncata (L: truncate); relating to the habit of segment apex 

Type: Wallich 256 (LT: BM), designated by C. R. Fraser-Jenkins, 

Taxon. Revis. Indian Subcontinental Pteridophytes 348 

(2008) // Nepal 

TW: Index Ferns Fern Allies Taiwan (2014) (present document) 

101 

LINDSAEA (Dennstaedtiaceae) 

Ety.: in honor of J. Lindsay (a Jamaican botanist) 

(455) Lindsaea chienii CHING 

Pub.: Sinensia 1: 4-5 (1929) 

Loc.: 錢氏鱗始蕨蕨 - QianShiLinShiJue 

Ety.: in honor of S. S. Chien (Chinese botanist) 

Type: Kwangsi, R. C. Ching 7184, 27-Aug-1928 (IT: UC, US) // China 

TW: Ito, H. 1981 (1981) 

(456) Lindsaea commixta TAGAWA 

Pub.: Acta Phytotax. Geobot. 6: 37-38, f. 3H-J (1937) 

Loc.: 海島鱗始蕨蕨 - HaiDaoLinShiJue 

Ety.: commixta (L: mixed together); relating to lamina habit being 

intermediate between two taxa (see protologue) 

Type: Kyushu, Osumi, Isl. Tane-ga-Shima, Hidaka s. n. (HT: KYO) // 

Japan 

TW: FOT 1975 (1975) 

(457) Lindsaea cultrata (WILLD.) SW. 

[IUCN: VU] 

Pub.: Syn. Fil. 119 (1806) 

Loc.: 網脈陵齒蕨蕨 - WangMaiLingChiJue 

Bas.: Adiantum cultratum WILLD. // Phytographia 14, pl. 10, f. 2 

(1794) 

Ety.: cultrata (L: shaped like a knife-blade); relating to the lamina 

shape (as described and illustrated in the protologue, though 

this is little representative of fully grown specimens of this 

taxon) 

Type: Malabaria, Anon. (B, Willd. Herb.) // India

102 

TW: a: Swinhoe, R. (1863), probably a misidentification only as this 

tropical taxon appears to be confined to SE Taiwan 

b: Yabe, Y. (1902) 

(458) Lindsaea ensifolia SW. 

[IUCN: NT] 


Loc.: 箭葉陵齒蕨蕨 - JianYeLingChiJue 

Ety.: ensis (L: sword), folium (L: leaf); relating to the habit of pinna 

Type: Mauritius, Anon. (SPA) // Mauritius 

TW: Henry, A. (1896) 

(459) Lindsaea heterophylla DRYAND. [IUCN: DD] 

Pub.: Trans. Linn. Soc. London 3: 41, pl. 8, f. 1 (1797) 

Loc.: 異葉陵齒蕨蕨 - YiYeLingChiJue 

Ety.: heteros (G: different), phyllon (G: leaf); relating to the variable 

lamina morphology, probably specifically to the dimorphism 

of sterile and fertile laminae 

Type: Malacca, Robertson s. n., Sep-1772 (HT: BM) // Malaysia 

TW: Ogata, M. (1930) 

(460) Lindsaea japonica (BAKER) DIELS 


Loc.: 日本陵齒蕨蕨 - RiBenLingChiJue 

Bas.: Lindsaea cultrata var. japonica BAKER // Syn. Fil. (ed. 2) 105 

(1867) 


Type: Oldham s. n. (K) // Japan 

TW: Ito, H. 1944 (1944) 

(461) Lindsaea javanensis BLUME 


Loc.: 爪哇鱗始蕨蕨 - ZhuaWaLinShiJue 



TW: Kuo, C. M. 1985 (1985) 

(462) Lindsaea kawabatae SA. KURATA [IUCN: - (VU)] 

Pub.: J. Geobot. 13: 100 (1965) 

Loc.: 細葉鱗始蕨蕨 - XiYeLinShiJue 

Ety.: in honor of M. Kawabata (Japanese botanist, collector of the 


Type: Kagoshima, Isl. Yakushima, Nakabase, Kawabata 991 (TOFO) 

// Japan 


(463) Lindsaea lucida BLUME [IUCN: CR (VU)] 

Pub.: Enum. Pl. Javae 216-217 (1828) 

Loc.: 方柄鱗始蕨蕨 - FangBingLingChiJue 

Ety.: lucida (L: shining); relating to the adaxial glossy lamina 


TW: Kuo, C. M. 1985 (1985)

103 

(464) Lindsaea obtusa J. SM. ex HOOK. [IUCN: NT] 

Pub.: Sp. Fil. 1: 224 (1846) 

Loc.: 鈍齒鱗始蕨蕨 - DunChiLinShiJue 

Ety.: obtusa (L: blunt); relating to the pinna teeth apex 

Type: Malacca, Cuming 394 (HT: K; IT: B, E, GH, P, W) // Malaysia 

TW: Kuo, C. M. 1985 (1985) 

(465) Lindsaea odorata ROXB. 

Pub.: Calcutta J. Nat. Hist. 4: 511 (1844) 

Loc.: 陵齒蕨蕨 - LingChiJue 

Ety.: odorata (L: having a smell); relating to the scent released by 

dry and drying plants (see protologue) 

Type: Garrow Hills // India 

TW: FOT 1975 (1975) 

(466) Lindsaea orbiculata (LAM.) METT. ex KUHN 

Pub.: Miq., Ann. Mus. Bot. Lugduno-Batavi 4: 279 (1869) 

Loc.: 圓葉陵齒蕨蕨 - YuanYeLingChiJue 

Bas.: Adiantum orbiculatum LAM. // Encycl. 1: 41 (1783) 

Ety.: orbiculata (L: round); relating to the pinna shape 

Type: near Malacca, Sonnerat s. n. (P) // Malaysia 

TW: Hayata, B. 1917 (1917) 

(467) Lindsaea yaeyamensis TAGAWA 

Pub.: Acta Phytotax. Geobot. 6: 31-33, f. 2C/D (1937) 

Loc.: 攀緣陵齒蕨 - PanYuanLingChiJue 

Ety.: relating to Yaeyama, the largest of the Iriomote islands (Japan, 

type location) 

Type: Ryukyu, Koidzumi s. n., Jul-1923 (HT: KYO; IT: L) // Japan 

TW: Ito, H. 1944 (1944) 

LOMARIOPSIS (Lomariopsidaceae) 

Ety.: lomaria relates to a fern genus (Blechnaceae), opsis (G: like); 

relating to the similarity to members of genus Lomaria (G: loma, 

border, and aris, substantive ending) 

(468) Lomariopsis spectabilis (KUNZE) METT. [IUCN: NT] 

Pub.: Fil. Hort. Bot. Lips. 22 (1856) 

Loc.: 羅蔓藤藤蕨蕨 - LuoManTengJue 

Bas.: Lomaria spectabilis KUNZE // Bot. Zeitung (Berlin) 6: 144 

(1848) 

Ety.: spectabilis (L: remarkable); relating to the plant habit 

Type: Java, Zollinger 395 (HT: L) // Indonesia 

TW: FOT 1975 (1975) 

LOXOGRAMME (Polypodiaceae) 

Ety.: loxos (G: oblique), gramme (G: line); relating to the shape and 

oblique arrangement of sori to the midrib

104 

(469) Loxogramme biformis TAGAWA [IUCN: EN (NT)] 


Loc.: 二形劍蕨蕨 - ErXingJianJue 

Ety.: bi (L: two), forma (L: shape, appearance); relating to the 

dimorphism of fertile and sterile laminae 

Type: Prov. Sintiku, Kyurin, Simada 5123, 1932 (HT: KYO) // Taiwan 


(470) Loxogramme chinensis CHING [IUCN: DD (CR)] 

Pub.: Sinensia 1: 13 (1929) 

Loc.: 中國劍蕨蕨 - ZhongGuoJianJue 


Type: Guangxi (IT: NY) // China 

TW: Kuo, C. M. 1985 (1985) 

(471) Loxogramme formosana NAKAI 


Loc.: 台台灣劍蕨蕨 - TaiWanJianJue 


Type: Arisan, Nimandaira, Hayata & Ito s. n., 2-Apr-1914 (HT: TI) // 



(472) Loxogramme grammitoides (BAKER) C. CHR. [IUCN: NT] 


Loc.: 小小葉劍蕨蕨 - XiaoYeJianJue 

Bas.: Gymnogramma grammitoides BAKER // J. Bot. 27: 178 (1889) 

Ety.: grammitis relates to a fern genus (see there), oides (L: suffix, 

denoting likeness of form); relating to the similarity of fronds 

to that of (simple fronded members of) genus Grammitis 

Type: Henry 5451 (ST: E, K) // China 

TW: Tagawa, M. 1940-49 (1940) 

(473) Loxogramme remotefrondigera (HAYATA) HAYATA 

Pub.: Ind. Filic, Suppl. 2: 28 (1917) 

Loc.: 長柄劍蕨蕨 - ChangBingJianJue 

Bas.: Polypodium remote-frondigerum HAYATA // Icon. Pl. Formosan. 

5: 323, f. 135A/B (1915); Loxogramme remote-frondigerum 

HAYATA introduced in same publication as synonym of former 

Ety.: remotus (L: distant), frons (L: leaf, frond), ger (L: carrying, 

bearing, used as suffix); relating to the rhizome with spaced 

fronds 

Type: Mt. Arisan, inter Funkiko et Taroyen, T. Ito & B. Hayata s. n., 

29-Mar-1914 (HT: TI) // Taiwan 

TW: Icon. Pl. Formosan. 5: 323, f. 135A/B (1915) 

(474) Loxogramme salicifolia (MAKINO) MAKINO 


Loc.: 柳葉劍蕨蕨 - LiuYeJianJue

105 

Bas.: Gymnogramma salicifolia MAKINO // Phan. Pterid. Jap. Icon. 1: 

pl. 34 (1899) 

Ety.: salix relates to a flowering plant genus (Salicaceae, "willow"), 

folium (L: leaf); relating to the shape of lamina resembling 

leaves of Salix 

Type: Japan 

TW: FOT 1975 (1975) 

LYCOPODIELLA (Lycopodiaceae) 

Ety.: diminutive of Lycopodium (a lycopod genus, see there) 

(475) Lycopodiella cernua (L.) PIC. SERM. 

Pub.: Webbia 23(1): 166 (1968) 

Loc.: 過山龍 - GuoShanLong 

Bas.: Lycopodium cernuum L. // Sp. Pl. 2: 1103 (1753) 

Ety.: cernuum (L: slightly drooping); relating to the strobilus 

Type: Tropical Asia, Anon. (LINN) // India 


LYCOPODIUM (Lycopodiaceae) 

Ety.: lykos (G: wolf), pous (G: foot); relating to the branch tips resembling 

a wolf's paw 

(476) Lycopodium annotinum L. 

[IUCN: VU] 

Pub.: Sp. Pl. 2: 1103 (1753) 

Loc.: 杉葉蔓石松 - ShanYeManShiSong 

Ety.: annotinum (L: of the previous year); relating to developing 

every year new branches 

Type: Plukenet, L. in Phytographia 3: pl. 205, fig. 5 (1692), LT 

designated by Jonsell & Jarvis, Nordic J. Bot. 14: 147 (1994) 

TW: FOT 1975 (1975) 

(477) Lycopodium casuarinoides SPRING 

Pub.: Monogr. Lycop. 1: 94 (1842) 

Loc.: 木賊葉石松 - MuZeiYeShiSong 

Ety.: casuarina relating to a flowering plant genus (Casuarinaceae), 


plant habit 

Type: 

TW: 

Cuming 2346 (BM, L) // Philippines 

a: Masamune, G. (1936) as Lycopodium casuarinoides var. 

japonicum NAKAI 

b: FOT 1975 (1975) 

(478) Lycopodium japonicum THUNB. 

Pub.: Fl. Jap. 341 (1784) 

Loc.: 日本石松 - RiBenShiSong 



TW: Sasaki, S. (1928)

106 

(479) Lycopodium juniperoideum SW. 

Pub.: Syn. Fil. 401 (1806) 

Loc.: 玉柏 - YuBo 

Ety.: juniperus relating to a gymnosperm genus (Cupressaceae, 

"juniper"), oideum (L: suffix, denoting likeness of form); 

relating to the similarity of branches to that of genus 

Juniperus 

Type: Siberia, E. Laxmann s. n. // Russia 

TW: Kuo, C. M. 1985 (1985) 

(480) Lycopodium multispicatum J. H. WILCE 

Pub.: Nova Hedwigia 3(1): 103-107 (1961) 

Loc.: 地刷子 - DiShuaZi 

Ety.: multi (L: many), spicatum (L: bearing a spike); relating to the 

fertile branch habit equipped with a large number of strobili 

Type: Mount Santo Tomas, Prov. Union, Northern Luzon, R. S. 

Williams 1603, 12-Oct-1914 (NY, UC, US) // Philippines 

TW: Kuo, C. M. 1985 (1985) 

(481) Lycopodium veitchii CHRIST 

Pub.: Bull. Acad. Int. Geogr. Bot. 16: 141 (1906) 

Loc.: 玉山石松 - YuShanShiSong 

Ety.: in honor of J. H. Veitch (British nurseryman) 

Type: Western China, Wilson 5409 (HT: P) // China 

TW: FOT 1975 (1975) 

(482) Lycopodium yueshanense C. M. KUO 


Loc.: 玉山地刷 - YuShanDiShua 

Ety.: relating to YuShan, the highest mountain of Taiwan and type 

location 

Type: Chiayi, Paiyunshanchuang to Yushanchienshan, Hsu s. n., 

Nov-1968 (HT: TAI) // Taiwan 


LYGODIUM (Schizaeaceae) 

Ety.: lygodes (G: flexibel); relating to the plant morphology, a climbing 

plant with flexuous rachis 

(483) Lygodium japonicum (THUNB.) SW. 


Loc.: 海金沙 - HaiJinSha 

Bas.: Ophioglossum japonicum THUNB. // Syst. Veg. (ed. 14) 926 

(1784) 


Type: Thunberg s. n., 1777 (?S) // Japan 


(484) Lygodium microphyllum (CAV.) R. BROWN [IUCN: VU] 

Pub.: Prodr. 162 (1810)

107 

Loc.: 小小葉海金沙 - XiaoYeHaiJinSha 

Bas.: Ugena microphyllum CAV. // Icon. 6: 76, pl. 595, f. 2 (1801) 

Ety.: mikros (G: small), phyllon (G: leaf); relating to the habit of 

segments 

Type: Luzon, Nee s. n. (HT: MA) // Philippines 


MACROTHELYPTERIS (Thelypteridaceae) 

Ety.: makros (G: large), thelypteris relates to another fern genus 

(Thelyperidaceae); relating to a large thelypterid fern 

(485) Macrothelypteris polypodioides (HOOK.) HOLTT. [IUCN: EN] 

Pub.: Blumea 17: 29-30 (1969) 

Loc.: 桫欏大金星蕨蕨 - SuoLuoDaJinXingJue 

Bas.: Alsophila polypodioides HOOK. // Nightingale Oceanic 

Sketches 131 (1835) 

Ety.: polypodium relates to a fern genus, oides (L: suffix, denoting 

likeness of form); relating to the similarity to a - at that time 

broadly defined - genus Polypodium (including many ferns 

without indusia, as well large compound species) 

Type: Nightingale s. n. (LT: K) // French Polynesia 

TW: FOT 1975 (1975) 

(486) Macrothelypteris torresiana (GAUD.) CHING 


Loc.: 大金星蕨蕨 - DaJinXingJue 

Bas.: Polystichum torresianum GAUD. // Voy. Uranie 8: 333 (1828) 

Ety.: probably named in honor of after D. L. Torres (vice-governor of 

the Marianas in the early 19 th century) 

Type: Mariana Islands, Gaudichaud s. n. (HT: P; IT: G) 

TW: FOT 1975 (1975) 

MARATTIA (Marattiaceae) 

Ety.: in honour of G. F. Maratti (Italian botanist) 

(487) Marattia pellucida C. PRESL [IUCN: CR (?EN)] 

Pub.: Suppl. Tent. Pterid. 10 (1845) 

Loc.: 觀音座蓮舅 - GuanYinZuoLianJiu 

Ety.: pellucida (L: clear, transparent); relating to the veins (see 

protologue) 

Alt.: Ptisana pellucida (C. PRESL) MURDOCK // Taxon 57: 747 (2008) 

Type: Luzon, Prov. Camarines, Cuming 177 (HT: ?L, ?NY fragm.; IT: 

BM, P, S, US) // Philippines 

TW: FOT 1975 (1975) 

MARSILEA (Marsileaceae) 

Ety.: in honor of L. F. Marsigli (Italian botanist)

108 

(488) Marsilea minuta L. 


Pub.: Mant. Pl. Alt. 308 (1771) 

Loc.: 田字草 - TianZiCao 

Ety.: minuta (L: small); relating to the plant habit 

Type: (LT: LINN), designated by Launert in Senckenberg. Biol. 49: 

273-315 (1968) // India 

TW: Henry, A. (1896) 

MICROLEPIA (Dennstaedtiaceae) 

Ety.: mikros (G: small), lepis (G: scale); relating to the small indusia 

(489) Microlepia x bipinnata (MAKINO) Y. SHIMURA 

Pub.: J. Phytogeogr. Taxon. 27(1): 41 (1979) 

Loc.: 台台北鱗蓋蕨蕨 - TaiBeiLinGaiJue 

Bas.: Microlepia marginata var. bipinnata MAKINO // J. Jap. Bot. 

3(12): 47 (1926) 

Ety.: bi (L: two), pinnatum (L: pinnate); relating to the lamina shape 

(being twice pinnate) 

Type: T. Makino s. n., 12-Dec-1926 // Japan 

TW: Tagawa, M. 1940-49 (1941) 

(490) Microlepia calvescens (WALL. ex HOOK.) C. PRESL 


Loc.: 光葉鳞鳞盖蕨蕨 - GuangYeLinGaiJue 

Bas.: Davallia calvescens WALL. ex HOOK. // Sp. Fil. 1: 172-173, pl. 

48B (1846) 

Ety.: calvescens (L: becoming bald); relating to the indument of 

several plant parts, especially that of veins and involucres 

(see protologue) 

Type: Wallich 2983 (K) // Nepal 


Microlepia marginalis HANCE [pub.: Monsunia 1: 87 (1900)] 

b: Tagawa, M. 1940-49 (1941) 

(491) Microlepia calvescens (WALL. ex HOOK.) C. PRESL x Microlepia 

trichosora CHING 


(492) Microlepia hookeriana (WALL. ex HOOK.) C. PRESL 


Loc.: 虎虎克氏鱗蓋蕨蕨 - HuKeShiLinGaiJue 

Bas.: Davallia hookeriana WALL. // Numer. List n. 2684 (1829), ex 

Hooker, Sp. Fil. 1: 172, t. 47B (1846) 

Ety.: in honor of W. J. Hooker (British botanist) 

Type: 


(493) Microlepia intramarginalis (TAGAWA) SERIZ. 

Assam, in montibus Silhet et Kamoun, Wallich 2684 (K) // India 

Pub.: J. Jap. Bot. 47: 48 (1972) 

Loc.: 羽裂鱗蓋蕨蕨 - YuLieLinGaiJue

109 

Bas.: Microlepia strigosa var. intramarginalis TAGAWA // Acta 

Phytotax. Geobot. 10: 202 (1941) 

Ety.: intra (L: inwardly), marginalis (L: marginal); relating to the 

position of sorus 

Type: Prov. Takao, between Ogidaira and Nanhozan, Kizan-gun, 

Tagawa 1482, 11-Dec-1938 (HT: KYO; IT: KYO, MICH) // 



(494) Microlepia krameri C. M. KUO 


Loc.: 克氏鱗蓋蕨蕨 - KeShiLinGaiJue 

Ety.: in honor of K. U. Kramer (Dutch botanist) 

Type: Taipei, Toutingli, Kuo 12254 (HT: TAI) // Taiwan 

TW: Taiwania 30: 59 (1985) 

(495) Microlepia marginata (HOUTT.) C. CHR. 


Loc.: 邊缘鱗蓋蕨蕨 - BianYuanLinGaiJue 

Bas.: Polypodium marginatum HOUTT. // Vollst. Pflanzensyst. 13: 

199 (1786) 

Ety.: marginata (L: with a distinct margin, marginated); related to the 

position of sori ("at margin of both sides of pinna", see 

protologue) 

Type: Thunberg s. n. (K) // Japan 

TW: Hayata, B. 1917 (1917) 

(496) Microlepia obtusiloba HAYATA 


Loc.: 團羽鱗蓋蕨蕨 - TuanYuLinGaiJue 

Ety.: obtusa (L: blunt), loba (L: lobe); relating to habit of apical 

segment 

Type: Koshun, Botanrokusha, Kawakami & Nakahara 896, 908, Dec- 

1905 (ST: TI) // Taiwan 


(497) Microlepia platyphylla (D. DON) J. SM. 

[IUCN: DD (CR)] 

Pub.: Lond. J. Bot. 1: 427 (1842) 

Loc.: 阔葉鱗蓋蕨蕨 - KuoYeLinGaiJue 

Bas.: Davallia platyphylla D. DON // Prodr. Fl. Nepal. 10 (1825) 

Ety.: platy (G: broad), phyllon (G: leaf); relating to the shape of 

lamina 

Type: Wallich s. n. (BM) // Nepal 

TW: FOT 1975 (1975) 

(498) Microlepia rhomboidea (WALL. ex KUNZE) PRANTL 

Pub.: Arbeiten Königl. Bot. Gart. Breslau 1: 31 (1892) 

Loc.: 斜方鱗蓋蕨蕨 - XieFangLinGaiJue 

Bas.: Davallia rhomboidea WALL. // Numer. List n. 257 (1828), ex 

Kunze, Bot. Zeitung (Berlin) 8: 158 (1850)

110 

Ety.: rhomboidea (L: shape of a rhombus); relating to the shape of 

segments 

Type: Wallich 257 (K, NY) // Nepal 

TW: a: Henry, A. (1896), probably only a mis-identification of a 

similar taxon growing at lower altitude 

b: Kuo, C. M. 1985 (1985) 

(499) Microlepia sinostrigosa CHING [IUCN: DD] 


Loc.: 中華鱗蓋蕨蕨 - ZhongHuaLinGaiJue 

Ety.: sino relates to China, the type location, strigosus (L: covered 

with short, appressed hairs); relating to the similarity to taxon 

Microlepia strigosa (see there) 

Type: Sichuan, Tianchuan Xian, W. P. Fang 3473 (HT: PE) // China 

TW: Kuo, C. M. 1985 (1985) 

(500) Microlepia speluncae (L.) T. MOORE 


Loc.: 熱帶鱗蓋蕨蕨 - ReDaiLinGaiJue 

Bas.: Polypodium speluncae L. // Sp. Pl. 2: 1093-1094 (1753) 

Ety.: spelunca (L: cave); relating to the place of first record 

("spelunca rupia innascens", L: born in a cave in the rocks; 

see Pluk. Alm. 255 (1696)) 

Type: Herb. Hermann: 3, 41, No. 384 (LT: BM) // Sri Lanka 


(501) Microlepia strigosa (THUNB.) C. PRESL 


Loc.: 粗毛鱗蓋蕨蕨 - CuMaoLinGaiJue 

Bas.: Trichomanes strigosum THUNB. // Syst. Veg. (ed. 14) 941 

(1784) 

Ety.: strigosus (L: covered with short, appressed hairs); relating to 

the axes of frond and lamina 

Type: Thunberg s. n. (?S) // Japan 

TW: Henry, A. (1896) 

(502) Microlepia substrigosa TAGAWA 


Loc.: 亞粗毛鱗蓋蕨蕨 - YaCuMaoLinGaiJue 

Ety.: sub (L: similar to), strigosus (L: covered with short, appressed 

hairs); relating to the similarity to taxon Microlepia strigosa 

(see there) 

Type: Taihoku, Rato-gun, near Doba, Tagawa 140, 29-Jul-1934 (HT: 



(503) Microlepia tenera CHRIST 

Pub.: Notul. Syst. (Paris) 1: 53-54 (1909) 

Loc.: 嫩鱗蓋蕨蕨 - NenLinGaiJue 

Ety.: tener (L: soft, tender); relating to the texture of lamina

111 

Type: Yunnan, Henryi 13155 (& 13155A) (E, P) // China 

TW: FOT 1975 (1975) 

(504) Microlepia trichocarpa HAYATA [IUCN: NT] 


Loc.: 毛果鱗蓋蕨蕨 - MaoGuoLinGaiJue 

Ety.: thrix (G: hair), karpos (G: fruit); relating to the presence of hair 

at the sorus 




(505) Microlepia trichosora CHING 

Pub.: Fl. Reipubl. Popularis Sin. 2: 358-359 (1959) 

Loc.: 毛囊鱗蓋蕨蕨 - MaoNangLinGaiJue 

Ety.: thrix (G: hair), soros (G: sorus); relating to the presence of hair 

at the sorus 

Type: Yunnan, western Tengchong and southern Burma border, 

Rock 7316 (HT: PE; IT: US) // China 

TW: Kuo, C. M. 1985 (1985) 

MICROSORUM (Polypodiaceae) 

Ety.: mikros (G: small), soros (G: sorus); relating to the small sori 

(506) Microsorum henryi (CHRIST) C. M. KUO 


Loc.: 大星蕨蕨 - DaXingJue 

Bas.: Polypodium henryi CHRIST // Bull. Herb. Boissier 6: 873 (1898) 

[non Diels (1899)] 


Type: Yunnan, Mengzi, Henry 9780 (HT: P; IT: MO, Z) // China 

TW: Kuo, C. M. 1985 (1985) 

(507) Microsorum insigne (BLUME) COPEL. 


Loc.: 箭葉星蕨蕨 - JianYeXingJue 

Bas.: Polypodium insigne BLUME // Enum. Pl. Javae 127 (1828) 

Ety.: insignis (L: remarkable, outstanding); relating to the plant habit 

Type: Java, Zippelius s. n. (LT: L) // Indonesia 

TW: MAN (2002) 

(508) Microsorum membranaceum (D. DON) CHING 

Pub.: Bull. Fan Mem. Inst. Biol. 4: 309 (1933) 

Loc.: 膜葉星蕨蕨 - MoYeXingJue 

Bas.: Polypodium membranaceum D. DON // Prodr. Fl. Nepal. 2 

(1825) 

Ety.: membranaceum (L: thin); relating to the lamina texture 

Type: Wallich s. n. (BM) // Nepal 

TW: Hayata, B. 1917 (1917)

112 

(509) Microsorum pteropus (BLUME) COPEL. 


Loc.: 三叉葉星蕨蕨 - SanChaYeXingJue 

Bas.: Polypodium pteropus BLUME // Enum. Pl. Javae, Add. 3 (1828) 

Ety.: pteron (G: wing), pous (G: foot); relating to the shape of lamina 

(decurrent at base) 


TW: J. Bot. 23: 103 (1885) 

(510) Microsorum punctatum (L.) COPEL. 


Loc.: 星蕨蕨 - XingJue 

Bas.: Acrostichum punctatum L. // Sp. Pl. (ed. 2): 1524 (1763) 

Ety.: punctatum (L: dotted, with spots); relating to the small sori 

Type: Fothergill s. n. (original material absent in LINN, and the name 

appears to need a neotype) // China 


(511) Microsorum steerei (HARR.) CHING [IUCN: EN] 


Loc.: 廣葉星蕨蕨 - GuangYeXingJue 

Bas.: Polypodium steerei HARR. // J. Linn. Soc., Bot. 16: 32 (1877) 

Ety.: in honor of W. C. Steere (US botanist and collector of the type 

specimen) 

Type: Takow, Apes Hill, Steere s. n., 1876 (HT: MICH; IT: GH, MO, 

P) // Taiwan 

TW: J. Linn. Soc., Bot. 16: 32 (1877) 

(512) Microsorum superficiale (BLUME) CHING 


Loc.: 表面星蕨蕨 - BiaoMianXingJue 

Bas.: Polypodium superficiale BLUME // Fl. Javae 136, pl. 56, f. 1 

(1828) 

Ety.: superficialis (L: occurring on the surface); relating to the 

positioning of sori on lamina 

Type: Yunnan, Mengzi, 5000 ft., Hancock 166 (?E, L) // China 

TW: Henry, A. (1896) 

MONACHOSORUM (Monachosoraceae) 

Ety.: monachos (G: solitary), soros (G: sorus); relating to the usually 

solitary sori, which are terminal on veins 

(513) Monachosorum henryi CHRIST 

Pub.: Bull. Herb. Boissier 6: 869 (1898) 

Loc.: 稀子蕨蕨 - XiZiJue 


Type: Mengtze, 6000 ft., Henry 10457 (IT: NY, US, ?P) // China 

TW: FOT 1975 (1975)

113 

(514) Monachosorum maximowiczii (BAKER) HAYATA 


Loc.: 岩穴蕨蕨 - YanXueJue 

Bas.: Polypodium maximowiczii BAKER // Syn. Fil. (ed. 2) 504 (1874) 

Ety.: in honor of K. J. Maximowicz (Russian botanist and collector of 


Type: Nagasaki, Kundsho-san, K. J. Maximowicz 17 b, 1863 (S) // 

Japan 

TW: a: Hayata, B. 1914-21 (1916) as Monachosorum maximowiczii 

var. melanocaulon HAYATA 

b: Ito, T. (1928) 

MONOGRAMMA (Vittariaceae) 

Ety.: monos (G: one), gramme (G: line); relating to the sori arranged in 

one line on the lamina, and there is a single vascular bundle per 

frond (in the simpler taxa) 

(515) Monogramma paradoxa (FEE) BEDD. [IUCN: NT] 

Pub.: Suppl. Ferns S. Ind. 24 (1876) 

Loc.: 連孢一條線蕨蕨 - LianBaoYiTiaoXianJue 

Bas.: Pleurogramme paradoxa FEE // Mem. Foug. 3: 38, pl. 4, f. 4 

(1851) 

Ety.: paradoxa (L: strange); relating to the overall plant habit (which 

is confusing, as described in the protologue) 

Type: Qulan, K. H. Mertens s. n. (HT: ?LE) // Polynesia 


(516) Monogramma trichoidea (FEE) J. SM. ex HOOK. [IUCN: DD] 

Pub.: J. Bot. (Hooker) 3: 394 (1841) 

Loc.: 一條線蕨蕨 - YiTiaoXianJue 

Bas.: Vaginularia trichoidea FEE // Mem. Foug. 3: 34 (1851) 

Ety.: thrix (G: hair), oides (L: suffix, denoting likeness of form); 

relates to the frond habit 

Type: Cuming 160 (B, L, MICH, Z) // Philippines 

TW: Ogata, M. (1933) 

NEOCHEIROPTERIS (Polypodiaceae) 

Ety.: neos (G: new), cheiropteris relating to a fern genus (Polypodiaceae); 

relating to the similarity to members of genus Cheiropteris (G: 

cheiro meaning band, pteris relating to genus Pteris, or ferns in 

general) 

(517) Neocheiropteris ensata (THUNB.) CHING 


Loc.: 扇蕨蕨 - ShanJue 

Bas.: Polypodium ensatum THUNB. // Trans. Linn. Soc. London, Bot.) 

2: 341 (1794) 

Ety.: ensata (L: sword-like); relating to the lamina habit

114 



NEPHROLEPIS (Nephrolepidaceae) 

Ety.: nephros (G: kidney), lepis (G: scale); relating to kidney-shaped 

scale-like indusium 

(518) Nephrolepis biserrata (SW.) SCHOTT. 

Pub.: Gen. Fil., ad. pl. 3 (1834) 

Loc.: 長葉腎蕨蕨 - ChangYeShenJue 

Bas.: Aspidium biserratum SW. // J. Bot. (Schrader) 1800(2): 32 

(1801) 

Ety.: bi (L: two-), serrata (L: serrated); relating to the margin of 

pinna 

Type: Grondal s. n. (LT: S; ILT: S) // Mauritius 


Nephrolepis acuta C. PRESL 

b: Kawakami, T. (1910) 

(519) Nephrolepis brownii (DESV.) HOVENKAMP & MIYAM. 

Pub.: Blumea 50(2): 293 (2005) 

Loc.: 毛葉腎蕨蕨 - MaoYeShenJue 

Bas.: Nephrodium brownii DESV. // Mem. Soc. Linn. Paris 6: 252 

(1827) 

Ety.: in honor of R. Brown (British botanist and collector of the type 

specimen) 

Type: R. Brown 20 (BM, E, K) // Australia 

TW: Blumea 50: 279-322 (2005) 

(520) Nephrolepis x copelandii W. H. WAGNER 

Pub.: Contr. Univ. Michigan Herb. 22: 185 (1999) 

Loc.: 科氏腎蕨蕨 - KeShiShenJue 

Ety.: in honor of E. B. Copeland (US botanist) 

Type: Hawaii, Wagner et al. 91044 (HT: MICH) // USA 


(521) Nephrolepis cordifolia (L.) C. PRESL 

Pub.: Tent. Pterid. 79 (1836), nom. cons. 

Loc.: 腎蕨蕨 - ShenJue 

Bas.: Polypodium cordifolium L. // Sp. Pl. 2: 1089 (1753) 

Ety.: cor (L: heart), folium (L: leaf); relating to the pinna base habit 

Type: Prov. De Azua, San Jose de Ocoa, Loma del Rancho, H. 

Ekman 11627 (HT: K; IT: B, LD, S, UPS) // Dominican 

Republic 

TW: J. Bot. 23: 105 (1885) 

(522) Nephrolepis x hippocrepicis MIYAM. * 

Pub.: Blumea 50(2): 312 (2005) 

Loc.: 馬蹄腎蕨蕨 - MaTiShenJue

115 

Ety.: hippos (G: horse), krepis (G: shoe); relating to the shape of 

indusium, resembling a horseshoe 

Type: Ryukyu Islands, Miyamoto & Nakayama s. n., 5-Mar-1985 (HT: 

TNS; IT: B, BISH, BM, K, MICH, NY, P, TAI, TNS) // Japan 


(523) Nephrolepis x pseudobiserrata MIYAM. 

Pub.: Blumea 50(2): 313 (2005) 

Loc.: 耳羽長葉腎蕨蕨 - ErYuChangYeShenJue 

Ety.: pseudos (G: false), biserrata relating to the taxon with epithet 

"biserrata"; relating to the similarity to taxon Nephrolepis 

biserrata (see there) 

Type: Ryukyu Islands, Miyamoto & Nakayama 1444, 11-Mar-1985 

(HT: TNS; IT: BISH, BM, K, MICH, NY, TAI) // Japan 


NOTHOPERANEMA (Dryopteridaceae) 

Ety.: nothos (G: false), peranema relates to a fern genus; relating to the 

similarity to members of genus Peranema (see there) 

(524) Nothoperanema hendersonii (BEDD.) CHING 


Loc.: 小小苞鱗毛蕨蕨 - XiaoBaoLinMaoJue 

Bas.: Lastrea hendersonii BEDD. // Suppl. Ferns S. Ind. 17, pl. 377 

(1876) 

Ety.: in honor of F. Henderson (British botanist and collector of the 


Type: Khasya, Shillong Hill, 6,000 feet elev., Capt. Henderson s. n. // 

India 

TW: Ito, H. 1944 (1944) 

(525) Nothoperanema squamiseta (HOOK.) CHING 


Loc.: 阿里山肉刺蕨蕨 - ALiShanRouCiJue 

Bas.: Nephrodium squamisetum HOOK. // Sp. Fil. 4: 140-141, pl. 268 

(1862) 

Ety.: squama (L: scale), seta (L: bristle); relating to the prominent 

setae above the pinnules and scales on the axes 

Type: Fernando Po, Clarence Peak, G. Mann 380 (HT: K) // Sierra 

Leone 

TW: FOT 1975 (1975) 

ODONTOSORIA 

Ety.: 

(Dennstaedtiaceae) 

odontos (G: of a tooth), soros (G: sorus); relating to the sori, which 

are attached at the extreme end of a small vein that traverses a 

marginal denticulation 

(526) Odontosoria biflora (KAULF.) C. CHR. 

Pub.: Index Filic. 464 (1906)

116 

Loc.: 闊片烏蕨蕨 - KuoPianWuJue 

Bas.: Davallia biflora KAULF. // Enum. Filic. 221-222 (1824) 

Ety.: bi (L: two-), flos (L: flower); relating to the presence of two sori 

at the apex of ultimate segment 

Type: Luzon, Manila, Chamisso s. n. (HT or IT: B) // Philippines 

TW: FOT 1975 (1975) 

(527) Odontosoria biflora (KAULF.) C. CHR. x Odontosoria chinensis (L.) J. 

SM. 


(528) Odontosoria chinensis (L.) J. SM. 

Pub.: Bot. Voy. Herald 430 (1857) 

Loc.: 烏蕨蕨 - WuJue 

Bas.: Trichomanes chinense L. // Sp. Pl. 2: 1099 (1753) 


Type: Osbeck s. n. (SPA) // China 

TW: Hayata, B. 1917 (1917) 

(529) Odontosoria chinensis (L.) J. SM. x Odontosoria gracilis (TAGAWA) 

RALF KNAPP 


(530) Odontosoria gracilis (TAGAWA) RALF KNAPP [IUCN: - (VU)] 


Loc.: 小小烏蕨蕨 - XiaoWuJue 

Bas.: Stenoloma gracile TAGAWA // Acta Phytotax. Geobot. 6: 227 

(1937) 

Ety.: gracilis (L: slender); relating to the shape of frond 

Type: Okinawa, Isl. Iriomote, Koidzumi s. n., 20-Jul-1923 (HT: KYO; 

IT: KYO) // Japan 


OLEANDRA (Oleandraceae) 

Ety.: resemblance of flowering plant oleander (genus Nerium, family 

Apocynaceae) 

(531) Oleandra wallichii (HOOK.) C. PRESL 


Loc.: 蓧蕨蕨 - TiaoJue 

Bas.: Aspidium wallichii HOOK. // Exot. Fl. 1: 5, t. 5 (1823) 

Ety.: in honor of N. Wallich (Danish botanist and collector of the 


Type: Wallich s. n. (BM, GH, PH) // Nepal 

TW: Hayata, B. 1911 (1911) 

ONOCLEA (Dryopteridaceae) 

Ety.: onos (G: vessel), kleiein (G: to close); relating to the rolled-up fertile 

pinnae enclosing the sori

(532) Onoclea orientalis (HOOK.) HOOK. [IUCN: CR] 

Pub.: Sp. Fil. 4: 161-162 (1862) 

Loc.: 東方莢果蕨蕨 - DongFangJiaGuoJue 

Bas.: Struthiopteris orientalis HOOK. // Sec. Cent. Ferns 4, pl. 4 

Ety.: 

(1861) 

orientalis (L: oriental); relating to the distribution as known to 

its author (see protologue) 

Type: (a) Sikkim, Himalaya, 12,000 ft., Hooker & Thomson s. n. // 

India 

(b) Assam, Simons s. n. // India 

(c) Hakodadi, C. Wright s. n. // Japan 

TW: Ogata, M. (1935) 

117 

ONYCHIUM (Pteridaceae) 

Ety.: onychion (G: little nail); relating to the resemblance of the ultimate 

lamina segments 

(533) Onychium japonicum (THUNB.) KUNZE 


Loc.: 日本金粉蕨蕨 - RiBenJinFenJue 

Bas.: Trichomanes japonicum THUNB. // Syst. Veg. (ed. 14) 941 

(1784) 


Type: Japan 

TW: J. Bot. 23: 103 (1885) 

(534) Onychium lucidum (D. DON) SPRENG. 

Pub.: Syst. Veg. 4: 66 (1827) 

Loc.: 高山金粉蕨蕨 - GaoShanJinFenJue 

Bas.: Leptostegia lucida D. DON. // Prodr. Fl. Nepal. 14-15 (1825) 

Ety.: lucidum (L: shining); relating to the adaxial lamina habit 

Type: Narainhetty, Buchanan-Hamilton s. n., 22-Feb-1803 (HT: BM) 

// Nepal 

TW: a: Swinhoe, R. (1863), probably a misidentified for O. 

japonicum 

b: Matsumara, J. & Hayata, B. (1906), as synonym of 

Cryptogramme japonica PRANTL 

c: Kuo, C. M. 1985 (1985) 

(535) Onychium siliculosum (DESV.) C. CHR. [IUCN: VU] 


Loc.: 金粉蕨蕨 - JinFenJue 

Bas.: Pteris siliculosa DESV. // Mag. Neuesten Entdeck. Gesammten 

Ety.: 

Naturk. Ges. Naturf. Freunde Berlin 5: 324 (1811) 

siliculosum (L: form of a silicle); relating to the habit of fertile 

segments 

Type: America australi, Anon. (HT: P) 

TW: Hayata, B. 1917 (1917)

118 

OPHIOGLOSSUM (Ophioglossaceae) 

Ety.: ophis (G: a serpent), glossa (G: tongue); relating to the shape of 

fertile section (long and slender like a snake) 

(536) Ophioglossum austroasiaticum M. NISHIDA [IUCN: - (VU)] 


Loc.: 高山瓶爾小小草 - GaoShanPingErXiaoCao 

Ety.: australis (L: southern), asiaticum (L: Asian); relating to its 

distribution 

Type: inter Tataka et Mt. Arisan, Tagawa 339, 15-Aug-1934 (HT: 


TW: Bull. Natl. Sci. Mus. 4: 329 (1959) 

(537) Ophioglossum pendulum L. 

Pub.: Sp. Pl (ed. 2) 2: 1518 (1763) 

Loc.: 帶狀瓶爾小小草 - DaiZhuangPingErXiaoCao 

Ety.: pendulum (L: hanging down); relating to the plant habit 

Type: India 

TW: Henry, A. (1896) 

(538) Ophioglossum petiolatum HOOK. 

Pub.: Exot. Fl. 1: 56, pl. 56 (1823) 

Loc.: 銳頭瓶爾小小草 - RuiTouPingErXiaoCao 

Ety.: petiolatus (L: stalked); probably relating to the characteristic 

long peduncle (see protologue) 

Type: cultivated, Antilles origin, Anon. (K) // Antilles 

TW: FOT 1975 (1975) 

(539) Ophioglossum reticulatum L. 

[IUCN: DD] 

Pub.: Sp. Pl. 2: 1063 (1753) 

Loc.: 網脈瓶爾小小草 - WangMaiPingErXiaoCao 

Ety.: reticulatum (L: marked with a network); relating to the venation 

of trophophyll 

Type: 

TW: 

South America 

a: Sasaki, S. (1928), though he probably misapplied this name 

to another taxon of this genus 

b: Kuo, C. M. 1985 (1985) 

(540) Ophioglossum sp. 



(541) Ophioglossum thermale KOMAROV [IUCN: NT] 

Pub.: Repert. Spec. Nov. Regni Veg. 13: 85 (1914) 

Loc.: 狹葉瓶爾小小草 - XiaYePingErXiaoCao 

Ety.: 

Type: 

thermale (L: relating to warm springs or water); relating to the 

collection site of the type ("ad fontes calidioris in cratero 

vulcani Uzon") 

Kamtchatka, V. Komarov s. n., 24-Aug-1909 (S) // Russia

119 

TW: 

a: Ito, H. 1981 (1981), as Ophioglossum thermale var. 

nipponicum (MIYABE & KUDO) NISHIDA ex TAGAWA 

b: FOJ (1995) 

OSMUNDA (Osmundaceae) 

Ety.: perhaps relating to Osmunder, a Saxon divinity; another alternative 

is os (L: bone), munda (L: clean, cure); perhaps relating to the past 

medical usage of roots of osmunds 

(542) Osmunda banksiifolia (C. PRESL) KUHN 

Pub.: Ann. Mus. Bot. Lugduno-Batavi 4: 299 (1869) 

Loc.: 粗齒革葉紫萁 - CuChiGeYeZiQi 

Bas.: Nephrodium banksiifolium C. PRESL // Reliq. Haenk. 1: 34 

(1825) 

Ety.: banksia relating to a flowering plant genus (Proteaceae, 

"Banksia"), folium (L: leaf); relating to the shape of lamina 

Type: Hab. in insula Sorzogon, Haenke s. n. (HT: FR; IT: FR) // 

Philippines 

TW: a: Matsumara, J. & Hayata, B. (1906) as Ophioglossum 

javanica BLUME and Ophioglossum banksiifolia as synonym 

b: FOT 1975 (1975) 

(543) Osmunda cinnamomea L. 

[IUCN: VU (EN)] 

Pub.: Sp. Pl. 2: 1066 (1753) 

Loc.: 假紫萁 - JiaZiQi 

Ety.: cinnamomea (L: cinnamon-coloured); relating to the colour of 

fertile frond 

Alt.: Osmundastrum cinnamomeum (L.) C. PRESL // Gefässbündel 

Farrn 18 (1847) 

Type: Maryland, Kalm s. n. (LT: LINN) // USA 

TW: Yamamoto, Y. (1932) 

(544) Osmunda claytoniana L. 

[IUCN: VU] 

Pub.: Sp. Pl. 2: 1066 (1753) 

Loc.: 絨假紫萁 - RongJiaZiQi 

Ety.: in honor of J. Clayton (US botanist and collector of the type 

specimen) 

Type: Virginia, Clayton s. n. (LT: BM, designated by Reveal & al. in 

Huntia 7: 229 (1987)) // USA 


(545) Osmunda japonica THUNB. 

Pub.: Nova Acta Regiae Soc. Sci. Upsal. 2: 209 (1780) 

Loc.: 紫萁 - ZiQi 


Type: Japan 

TW: Sasaki, S. (1928)

120 

PAESIA (Dennstaedtiaceae) 

Ety.: in honor of F. D. Paes Leme (Portuguese scientist) 

(546) Paesia radula (BAKER) C. CHR. [IUCN: CR] 


Loc.: 曲軸蕨蕨 - QuZhouJue 

Bas.: Pteris radula BAKER // J. Bot. 18: 211 (1880) 

Ety.: 

radula (L: rough); relating to the minutely rough stipe and 

rachis (see protologue) 

Type: Sumatra, Mt. Singalan 1700 m, Odoardo Beccari 417 (BM) // 

Indonesia 

TW: Kuo, C. M. 1985 (1985) 

PARACETERACH (incl. Parahemionitis) (Pteridaceae) 

Ety.: para (G: near to), ceterach is the name of a genus (Aspleniaceae); 

relating to the similarity to members of genus Ceterach (derived 

from the Arabian sjetrak, an old name aplied to this taxon) 

Parahemionitis: para (G: beside), hemionitis relates to genus 

Hemionitis (Pteridaceae); relates to the similarity of members of 

this genus to that of genus Hemionitis (G: hemionos, mule, relating 

to the usage of the plant in the past) 

(547) (Paraceterach cordata comb. ined.) Parahemionitis cordata (ROXB. ex 

HOOK. & GREV.) FRASER-JENK. [IUCN: VU] 

Pub.: New Sp. Syndr. Indian Pteridol. 187 (1997) 

Loc.: 澤瀉蕨蕨 - ZeXieJue 

Bas.: Hemionitis cordata ROXB. ex HOOK. & GREV. // Icon. Filic. 1: pl. 

64 (1828) 

Ety.: cordata (L: with two equal rounded lobes at base); relating to 

base of pinna 

Alt.: Hemionitis cordata ROXB. ex HOOK. & GREV. // Icon. Filic. 1: pl. 

64 (1828) 

Type: Prope H. B. Calcutta, Herb. East India Company no. 44 (LT: K; 

ILT: BM) // India 

TW: Henry, A. (1896) 

(548) Paraceterach vestita (HOOK.) R. M. TRYON [IUCN: EN] 


Loc.: 金毛裸蕨蕨 - JinMaoLuoJue 

Bas.: Gymnogramma vestita HOOK. // Icon. Pl. 2: pl. 115 (1837) 

Ety.: vestita (L: clothed); relating to the dense cover of hairs 

Alt.: Paragymnopteris vestita (HOOK.) G. M. ZHANG & RANKER // 

Indian Fern J. 10(1-2): 230 (1993, publ. 1994) 

Type: Kumoon, Wallich no. 12 (IT: US, UC) // Nepal 

TW: FOT 1975 (1975)

PERANEMA (Dryopteridaceae) 

Ety.: pera (L: pouch), nema (L: thread); relating to the pouch-like sorus 

born on slender thread like stalk 

(549) Peranema aspidioides (BLUME) METT. [IUCN: VU] 

Pub.: Fil. Lechl. 2: 33 (1859) 

Loc.: 紅線蕨蕨 - HongXianJue 

Bas.: Diacalpe aspidioides BLUME // Enum. Pl. Javae 241 (1828) 

Ety.: aspidium relating to a fern genus (Dryopteridaceae, name of 

genus now obsolete), oides (L: suffix, denoting likeness of 

form); relating to the similarity in habit to members of genus 

Aspidium (based on protologue) 

Alt.: Diacalpe aspidioides BLUME // Enum. Pl. Javae 241 (1828) 


TW: Kuo, C. M. 1985 (1985) 

(550) Peranema cyatheoides D. DON 


Loc.: 柄囊蕨蕨 - BingNangJue 

Ety.: cyathea relating to a fern genus (Cyatheaceae), oides (L: 

suffix, denoting likeness of form); relating to the similarity in 

indusium to members of genus Cyathea (see there) 

Type: Wallich 183 (IT: S) // Nepal 

TW: Ogata, M. (1940) 

PHEGOPTERIS (Thelypteridaceae) 

Ety.: phegos (G: beech), pteris (relates to genus Pteris, or ferns in 

general); relating to a fern growing under beech trees 

(551) Phegopteris connectilis (MICHX.) WATT [IUCN: NT] 

Pub.: Canad. Naturalist Geol., n. s., 3: 159 (1866) 

Loc.: 長柄假金星蕨蕨 - ChangBingJiaJinXingJue 

Bas.: Polypodium connectile MICHX. // Fl. Bor.-Amer. 2: 271-272 

(1803) 

Ety.: connectilis (L: connecting); relating to the rachis wing 

connecting the pinnae 

Type: Canada 

TW: FOT 1975 (1975) 

(552) Phegopteris decursivepinnata (H. C. HALL) FEE 

Pub.: Mem Foug. 5: 242, pl. 20A, f. 1 (1852) 

Loc.: 翅軸假金星蕨蕨 - ChiZhouJiaJinXingJue 

Bas.: Polypodium decursivepinnatum H. C. HALL // Nieuwe Verh. 

Eerste Kl. Kon. Ned. Inst. Wetensch. Amsterdam 5: 204 

(1836) 

Ety.: decursiva (L: running down), pinnata (L: pinnate); relating to 

the lamina shape 

Type: S. China, van Hall (L) // China 

TW: J. Bot. 23: 105 (1885) 

121

122 

PHYMATOSORUS (Polypodiaceae) 

Ety.: phymatos (G: swelling), soros (G: sorus); relating to the sori that are 

immersed in the lamina 

(553) Phymatosorus longissimus (BLUME) PIC. SERM. [IUCN: EN (CR)] 

Pub.: Webbia 28: 459 (1973) 

Loc.: 水社擬茀蕨蕨 - ShuiSheNiFuJue 

Bas.: Polypodium longissimum BLUME // Enum. Pl. Javae 127 (1828) 

Ety.: superlative of longus (L: long); relating to the habit 

Type: Java, Blume 35 (LT: L) // Indonesia 

TW: Henry, A. (1896) 

(554) Phymatosorus membranifolius (R. BR.) S. G. LU [IUCN: VU] 

Pub.: Guihaia 19(1): 27 (1999) 

Loc.: 薄薄葉擬茀蕨蕨 - BoYeNiFuJue 

Bas.: Polypodium membranifolium R. BR. // Prodr. 147 (1810) 

Ety.: membrana (L: skin), folium (L: leaf); relating to the lamina 

texture 

Type: New Holland, W. J. Hooker s. n. (PH) // Australia 

TW: Kuo, C. M. 2001 (2001) 

(555) Phymatosorus scolopendria (BURM. F.) PIC. SERM. 

Pub.: Webbia 28: 457, 460 (1973) 

Loc.: 海岸擬茀蕨蕨 - HaiAnNiFuJue 

Bas.: Polypodium scolopendria BURM. F. // Fl. Indica 232 (1768) 

Ety.: scolopendria (L: scolopender, centipede; used as noun in 

apposition); relating to the lamina habit 

Type: “India”, Java, Pryon s. n. (LT: G) // Indonesia 

TW: Ito, H. 1944 (1944) 

PITYROGRAMMA (Pteridaceae) 

Ety.: pityron (G: scurf, bran), gramme (G: line); relating to the lower 

surface of lamina being covered in a scurfy farina, and the 

sporangia are arranged in lines along the veins 

(556) Pityrogramma calomelanos (L.) LINK 

Pub.: Handbuch 3: 20 (1833) 

Loc.: 粉葉蕨蕨 - FenYeJue 

Bas.: Acrostichum calomelanos L. // Sp. Pl. 2: 1072 (1753) 

Ety.: kalos (G: beautiful), melaina (G: black); relating to the black 

stipe (see protologue) 

Type: Anon. (LT: LINN), designated by Panigrahi, Kew Bull. 30: 657- 

667 (1976) 

TW: FOT 1975 (1975) 

PLAGIOGYRIA (Plagiogyriaceae) 

Ety.: plagios (G: oblique), gyros (G: circle); relating to the oblique annulus 

of sporangium

123 

(557) Plagiogyria adnata (BLUME) BEDD. 

Pub.: Ferns Brit. India pl. 51 (1866) 

Loc.: 瘤足蕨蕨 - LiuZuJue 

Bas.: Lomaria adnata BLUME // Enum. Pl. Javae 205 (1828) 

Ety.: adnata (L: born to, grow to, joined by having grown together); 

relating to the lamina (pinna) habit 


TW: J. Bot. 23: 103 (1885) 

(558) Plagiogyria euphlebia (KUNZE) METT. 

Pub.: Abh. Senckenb. Naturf. Ges. 2: 274 (1858) 

Loc.: 華中瘤足蕨蕨 - HuaZhongLiuZuJue 

Bas.: Lomaria euphlebia KUNZE // Bot. Zeitung (Berlin) 6: 521 (1848) 

Ety.: eu (G: well, good), phlebia (G: veined); relating to the wellvisible 

veins of abaxial lamina (see protologue) 

Type: Zollinger s. n. (LT: L) // Japan 

TW: Hayata, B. 1911 (1911) 

(559) Plagiogyria falcata COPEL. 

Pub.: Philipp. J. Sci. 2: 133, t. 1, f. B (1907) 

Loc.: 倒葉瘤足蕨蕨 - DaoYeLiuZuJue 

Ety.: falcata (L: sickle-shaped); relating to the shape of pinna 

Type: Mindoro, Mt. Halcon, Merrill 5960 (HT: MICH; IT: P) // 

Philippines 

TW: Hayata, B. 1914-21 (1914) 

(560) Plagiogyria glauca (BLUME) METT. 

Pub.: Abh. Senckenberg. Naturf. Ges. 2: 273 (1858) 

Loc.: 台台灣瘤足蕨蕨 - TaiWanLiuZuJue 

Bas.: Lomaria glauca BLUME // Enum. Pl. Javae 204 (1828) 

Ety.: glaucos (G: bluish-green); relating to the colour of abaxial 

lamina 

Type: Java, Salak, Blume s. n. (LT: L) // Indonesia 

TW: a: Matsumara, J. & Hayata, B. (1906) for Plagiogyria glauca 

var. philippinensis CHRIST 

b: MAN (2002) 

(561) Plagiogyria japonica NAKAI [IUCN: VU] 


Loc.: 華東瘤足蕨蕨 - HuaDongLiuZuJue 


Type: Izu, Yahatano, Nakai s. n. Apr 1928 (LT: TI) // Japan 

TW: FOT 1975 (1975) 

(562) Plagiogyria koidzumii TAGAWA [IUCN: CR (EN)] 


Loc.: 小小泉氏瘤足蕨蕨 - XiaoQuanShiLiuZuJue 

Ety.: in honor of G. Koidzumi (Japanese botanist and collector of 

the type specimen)

124 

Type: Iromote Isl., G. Koidzumi s. n., 20-Jul-1923 (HT: KYO; IT: BM, 

KYO, L) // Japan 

TW: FOT 1975 (1975) 

(563) Plagiogyria stenoptera (HANCE) DIELS 


Loc.: 耳形瘤足蕨蕨 - ErXingLiuZuJue 

Bas.: Blechnum stenopterum HANCE // J. Bot. 21: 268 (1883) 

Ety.: stenos (G: narrow), pteron (G: wing); relating to the basal 

lamina habit 

Type: Tamsui, Hancock 39 (HT: K; IT: BM) // Taiwan 

TW: J. Bot. 21: 268 (1883) 

PLEOCNEMIA (Dryopteridaceae) 

Ety.: pleos (G: full), knemia (G: rays); relating to the venation, where free 

included veinlets radiate in all directions 

(564) Pleocnemia rufinervis (HAYATA) NAKAI 


Loc.: 網脈突齒蕨蕨 - WangMaiTuChiJue 

Bas.: Dryopteris rufinervis HAYATA // J. Coll. Sci. Imp. Univ. Tokyo 

30: 420-421 (1911) 

Ety.: rufus (L: reddish), nervus (L: nerve); relating to the colour of 

costae 

Type: Chioran, G. Nakahara 274, Aug-1905 (HT: TI; IT: TAI) // 



(565) Pleocnemia submembranacea (HAYATA) TAGAWA & K. IWATS. 


Loc.: 黃腺羽蕨蕨 - HuangXianYuJue 

Bas.: Aspidium submembranaceum HAYATA // Icon. Pl. Formosan. 4: 

188-189, f. 126 (1914) 

Ety.: sub (L: almost), membranacea (L: thin and very flexible); 

relating to the lamina texture 

Type: Urai, B. Hayata & S. Sasaki s. n., Jul-1912 (HT: TI) // Taiwan 


POLYPODIUM (Polypodiaceae) 

Ety.: poly (G: many), podion (G: foot); relating to the frequently branching 

rhizome resembling many feet 

(566) Polypodium amoenum WALL. ex METT. 

Pub.: Abh. Senckenb. Naturf. Ges. (Polypodium): 80 (1857) 

Loc.: 阿里山水龍骨 - ALiShanShuiLongGu 

Ety.: amoena (L: pleasing, lovely); relating to the plant habit 

Alt.: Goniophlebium amoenum (WALL. ex METT.) J. SM. ex BEDD. // 

Ferns Brit. India pl. 5 (1866) 

Type: Wallich 290, 1820 (BM, BR, PH) // Nepal

TW: Henry, A. (1896) 

(567) Polypodium formosanum BAKER 

Pub.: J. Bot. 23: 105-106 (1885) 

Loc.: 台台灣水龍骨 - TaiWanShuiLongGu 


Alt.: Goniophlebium formosanum (BAKER) RÖDL-LINDER // Blumea 

34(2): 406 (1990) 

Type: Tamsui, Hancock 50 (HT: K; IT: US) // Taiwan 

TW: J. Bot. 23: 105-106 (1885) 

(568) Polypodium mengtzeense CHRIST 

Pub.: Bull. Herb. Boissier 6: 869-670 (1898) 

Loc.: 箭葉水龍骨 - JianYeShuiLongGu 

Ety.: 

Alt.: 

125 

relating to Mengtze, mountains in China and the type location 

Goniophlebium mengtzeense (CHRIST) RÖDL-LINDER // Philipp. 

J. Sci. 116(2): 154 (1987) 

Type: Yunnan, Mengtze mountains, Henry 10964B (LT: P; ILT: K) // 

China 

TW: Kuo, C. M. 2001 (2001) 

(569) Polypodium microrhizoma C. B. CLARKE ex BAKER 

Pub.: Syn. Fil. (ed. 2) 511 (1874) 

Loc.: 栗柄水龍骨 - LiBingShuiLongGu 

Ety.: 

[IUCN: VU (NT)] 

mikros (G: small), rhizoma (G: mass of roots, here rhizome); 

relating to the habit of rhizome 

Alt.: Goniophlebium microrhizoma (C. B. CLARKE ex BAKER) C. B. 

CLARKE ex BEDD. // Suppl. Ferns S. India 21, pl. 384 (1876) 

Type: Sikkim, Lachong, Elwes s. n., Dec-1872, p. p. (LT: K) 

TW: Kuo, C. M. 1985 (1985) 

(570) Polypodium persicifolium DESV. * 

[IUCN: - (DD)] 

Pub.: Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. 

Gesammten Naturk. 5: 316 (‘persicaefolium’) (1811) 

Loc.: 棱脈蕨蕨 - LengMaiJue 

Ety.: persica relating to a flowering plant (Rosaceae, "peach"), 

folium (L: leaf); relating to the similarity of pinna with leaves 

of the peach tree 

Alt.: 

Goniophlebium persicifolium (DESV.) BEDD. // Suppl. Ferns Brit. 

Ind. 21, pl. 79 (1876) 

Java, Jussieu s. n. (?P) // Indonesia 

Type: 

TW: TW J. of Biodivers. 14(3-4): 81-90 (2012) 

(571) Polypodium raishaense ROSENST. 

Pub.: Hedwigia 56: 346 (1915) 

Loc.: 大葉水龍骨 - DaYeShuiLongGu 

Ety.: relating to Raisha, a place in Taiwan and the type location 

Alt.: Goniophlebium raishaense (ROSENST.) C. M. KUO (?nom. nud.) 

Type: Raisha, 1200 m, Faurie 219, Mar-1914 (ST: B, NY, S, TI) // 

Taiwan

126 

TW: Hedwigia 56: 346 (1915) 

(572) Polypodium transpianense YAMAMOTO 

Pub.: J. Soc. Trop. Agric. 3: 236 (1931) 

Loc.: 疏毛水龍骨 - ShuMaoShuiLongGu 

Ety.: trans (L: across), pian relating to Mt. Pianon in Taiwan; name 

relates to the type location 

Alt.: Goniophlebium transpianense (YAMAMOTO) C. M. KUO (?nom. 

nud.) 

Type: Taihoku, Mt. Pianon, Rato-gun, Yamamoto s. n., 04-Sep-1925 

(TAI) // Taiwan 

TW: J. Soc. Trop. Agric. 3: 236 (1931) 

POLYSTICHUM (incl. Cyrtomium) (Dryopteridaceae) 

Ety.: poly (G: many), stichos (G: row); relating to the distribution of sori in 

many regular rows on the pinnae of the type specimen 

Cyrtomium: kyrtoma (G: arch); relating to the veins anastomosing 

into arches 

(573) Polystichum acanthophyllum (FRANCH.) CHRIST 

Pub.: Bull. Soc. Bot. France, Mem. 1: 30 (1905) 

Loc.: 針葉耳蕨蕨 - ZhenYeErJue 

Bas.: Aspidium acanthophyllum FRANCH. // Bull. Soc. Bot. France 32: 

28-29 (1885) 

Ety.: akantha (G: thorn), phyllon (G: leaf); relating to the lamina 

habit 

Type: Yunnan (HT: P) // China 

TW: Kuo, C. M. 1985 (1985) 

(574) Polystichum acutidens CHRIST 

Pub.: Bull. Acad. Int. Geogr. Bot. 11: 259-260 (1902) 

Loc.: 台台東耳蕨蕨 - TaiDongErJue 

Ety.: acutum (L: sharp, pointed), dens (L: tooth); relating to the 

pinna margin 

Type: Guizhou, Bodinier 1836 (P) // China 

TW: FOT 1975 (1975) 

(575) Polystichum anomophyllum (ZENKER) NAKAI [IUCN: - (DD)] 


Loc.: 奇葉貫眾蕨蕨 - QiYeGuanZhongJue 

Bas.: Aspidium anomophyllum ZENKER // Pl. Ind. 1, pl. 1 (1835) 

Ety.: anomos (G: irregular, unequal), phyllon (G: leaf); relating to the 

habit of pinna base 

Alt.: Cyrtomium anomophyllum (ZENKER) FRASER-JENK. // Taxon. 

Revis. Indian Subcontinental Pteridophytes 330 (2008) 

Type: Utacamund, B. Schmid s. n. (LT: B) // India 

TW: a: Fraser-Jenkins, C. R. 2008 (2008) with uncertainty 

statement 

b: Knapp, R. 2013 (2013)

127 

(576) Polystichum atkinsonii BEDD. 

Pub.: Suppl. Ferns S. India 14, pl. 362 (1876) 

Loc.: 小小芽孢孢耳蕨蕨 - XiaYaBaoErJue 

Ety.: in honor of W. S. Atkinson (British botanist and collector of the 


Type: Sikkim, Atkinson s. n. // India 


(577) Polystichum attenuatum TAGAWA & K. IWATS. * 


Pub.: Acta Phytotax. Geobot. 23: 113-114, f. 9 (1968) 

Loc.: 長羽芽苞耳蕨蕨 - ZhangYuYaBaoErJue 

Ety.: attenuatum (L: reduced); relating to the tapering lamina apex 

Type: Loei (=Loey) Prov., Mt. Phu Luang, M. Tagawa et al. T1487 

(HT: KYO; IT: L, US) // Thailand 

TW: TW J. of Biodivers. 16(3): 263-272 (2014) 

(578) Polystichum biaristatum (BLUME) T. MOORE 


Loc.: 二尖尖耳蕨蕨 - ErJianErJue 

Bas.: Aspidium biaristatum BLUME // Enum. Pl. Javae 164-165 

(1828) 

Ety.: bi (L: two-), aristatum (L: awned, bearded); relating to the 

margin of pinnules (see protologue) 


TW: Tagawa, M. 1940-49 (1941) 

(579) Polystichum capillipes (BAKER) DIELS [IUCN: CR (?EN)] 


Loc.: 小小耳蕨蕨 - XiaoErJue 

Bas.: Aspidium capillipes BAKER // J. Bot. 26: 228 (1888) 

Ety.: capillus (L: hair), pes (L: foot); relating to thin stipe 

Type: Szechuen, Faber 1086 (MO, NY) // China 

TW: a: Fraser-Jenkins, C. R. 1997 (1997) with uncertainty 

statement 

b: Knapp, R. 2011 (2011) 

(580) Polystichum caryotideum (WALL. ex HOOK. & GREV.) DIELS 


Loc.: 細齒貫眾蕨蕨 - XiChiGuanZhongJue 

Bas.: Aspidium caryotideum WALL. // Numer. List n. 376 (1828) 

(nom. nud.) ex Hooker & Greville, Icon. Filic. 1: pl. 69 (1828) 

Ety.: caryota relates to a flowering plant genus (Arecaceae, "fishtailpalm"); 

probably relating to the shape of pinnae which are 

similar to the leaves of (some) members of genus Caryota 

Alt.: Cyrtomium caryotideum (WALL. ex HOOK. & GREV.) C. PRESL // 

Tent. Pterid. 86, pl. 2, f. 26 (1836) 

Type: Wallich 376, 1828 (MICH, PH) // Nepal 

TW: Hayata, B. 1909 (1909)

128 

(581) Polystichum chunii CHING [IUCN: - (EN)] 

Pub.: Sinensia 1: 2-3 (1929) 

Loc.: 陳氏耳蕨蕨 - ChenShiErJue 

Ety.: in honor of W. Y. Chun (Chinese botanist) 

Type: Guangxi, Luchen, Dar-Young Kiang, R. C. Ching 6256, 27- 

Jun-1928 (HT: PE; IT: LBG, US) // China 

TW: Taiwania 59: 86-92 (2014) 

(582) Polystichum deltodon (BAKER) DIELS 


Loc.: 對對生耳蕨蕨 - DuiShengErJue 

Bas.: Aspidium deltodon BAKER // Gard. Chr., n. s., 14: 494 (1880) 

Ety.: deltodon (L: having a triangular tooth); relating to the teeth at 

pinna apex 

Type: Nanto, Schang (BM) // China 

TW: J. Bot. 23: 105 (1885) 

(583) Polystichum devexiscapulae KOIDZ. 


Loc.: 披針貫眾蕨蕨 - PiZhenGuanZhongJue 

Ety.: devexus (L: sloping), scapula (L: shoulder blade); relating to 

the lamina habit 

Alt.: Cyrtomium devexiscapulae (KOIDZ.) CHING // Bull. Chin. Bot. 

Soc. 2: 96-97 (1936) 

Type: insualae Quelpaert, U. Faurie 35, 25-Oct-1906 (HT: KYO) // 

Korea 

TW: Tagawa, M. 1940-49 (1940) 

(584) Polystichum falcatum (L. F.) DIELS 

Pub.: Nat. Pflanzenfam. 1(4): 194 (1899) [not Fee (1850)] 

Loc.: 全緣貫眾蕨蕨 - QuanYuanGuanZhongJue 

Bas.: Polypodium falcatum L. F. // Suppl. Pl. 446 (1781) 

Ety.: falcatum (L: sickle-shaped), relating to the shape of pinna 

Alt.: Cyrtomium falcatum (L. F.) C. PRESL // Tent. Pterid. 86 (1836) 


TW: J. Bot. 23: 105 (1885) 

(585) Polystichum formosanum ROSENST. 

Pub.: Hedwigia 56: 338-339 (1915) 

Loc.: 台台灣耳蕨蕨 - TaiWanErJue 


Type: Bunkikyo, 2000 m, Faurie 363, May-1914 (HT: TI; IT: S) // 


TW: Hedwigia 56: 338-339 (1915) 

(586) Polystichum fortunei (J. SM.) NAKAI [IUCN: - (VU)] 


Loc.: 貫眾蕨蕨 - GuanZhongJue 

Bas.: Cyrtomium fortunei J. SM. // Ferns Brit. For. 286 (1866) 

Ety.: in honor of R. Fortune (British botanist)

129 

Alt.: Cyrtomium fortunei J. SM. // Ferns Brit. For. 286 (1866) 

Type: Cultivated at Kew, originated from Japan, Anon. s. n. (BM) // 

Japan 

TW: Hayata, B. 1917 (1917) 

(587) Polystichum fraxinellum (CHRIST) DIELS [IUCN: VU] 


Loc.: 網脈耳蕨蕨 - WangMaiErJue 

Bas.: Aspidium fraxinellum CHRIST // Bull. Herb. Boissier 7: 15-16 

(1899) 

Ety.: fraxinea (L: relating to a flowering plant genus, Oleaceae, 

"ash"), ella (L: diminutive suffix); probably relating to the frond 

habit 

Alt.: 

Cyrtogonellum fraxinellum (CHRIST) CHING // Bull. Fan Mem. 

Inst. Biol., Bot. 8: 329, pl. 7, f. 1 (1938) 

Type: Yunnan, Mengtze, A. Henry 11550 (HT: P; IT: MO, US) // 

China 

TW: Tagawa, M. 1940-49 (1940) 

(588) Polystichum glaciale CHRIST [IUCN: CR] 

Pub.: Bull. Soc. Bot. France, Mem. 1: 28-29 (1905) 

Loc.: 玉龍蕨蕨 - YuLongJue 

Ety.: glacialis (L: frozen); relating to the habitat of first collection 

(verified on type annotation) 

Type: Delavay 45 (HT: P; IT: P) // China 

TW: Kuo, C. M. 2001 (2001) 

(589) Polystichum grandifrons C. CHR. [IUCN: DD (CR)] 


Loc.: 九州耳蕨蕨 - JiuZhouErJue 

Ety.: grandis (L: large, great, tall), frons (L: leaf, frond); relating to 

the plant habit 

Type: Yunnan, Mengzi, A. Henry 13686 (HT: PE; IT: PE) // China 

TW: MAN (2002) 

(590) Polystichum hancockii (HANCE) DIELS 


Loc.: 韓氏耳蕨蕨 - HanShiErJue 

Bas.: Ptilopteris hancockii HANCE // J. Bot. 22: 139 (1884) 


specimen) 

Type: Tam-shui, 3500 ft., W. Hancock 41, 11-Dec-1881 (B, BM) // 


TW: J. Bot. 22: 139 (1884) 

(591) Polystichum hecatopterum DIELS 

Pub.: Bot. Jahrb. Syst. 29: 193 (1900) 

Loc.: 

Ety.: 

鋸齒葉耳蕨蕨 - JuChiYeErJue 

hekaton (G: a hundred), pteron (G: wing); relating to the 

lamina habit (number and shape of pinnae)

130 

Type: Sichuan, Rosthorn 1021 (HT: BM) // China 

TW: Ito, H. 1944 (1944) 

(592) Polystichum integripinnum HAYATA 


Loc.: 狹葉貫眾蕨蕨 - XiaYeGuanZhongJue 

Ety.: integer (L: entire), pinna (L: wing, here relating to pinna); 

relating to the pinna habit 

Alt.: Cyrtomium integripinnum (HAYATA) COPEL. // Philipp. J. Sci. 38: 

136 (1929) 

Type: Arisan, B. Hayata & S. Sasaki s. n., Jan-1912 (HT: KYO) // 



(593) Polystichum lachenense (HOOK.) BEDD. 


Loc.: 高山耳蕨蕨 - GaoShanErJue 

Bas.: Aspidium lachenense HOOK. // Sp. Fil. 4: 8, pl. 212 (1862) 

Ety.: relating to Lachen, a place in China and the type location 

Type: Sikkim-Himalaya, Lachen, elev. 13-16,000 ft. (K) // India 

TW: Ogata, M. (1929) 

(594) Polystichum lepidocaulon (HOOK.) J. SM. 

Pub.: Ferns Brit. For. 286-287 (1866) 

Loc.: 鞭葉耳蕨蕨 - BianYeErJue 

Bas.: Aspidium lepidocaulon HOOK. // Sp. Fil. 4: 12, pl. 217 (1862) 

Ety.: lepidotos (G: scaly), caulon (G: stalk, stem); relating to the 

stipe habit 

Alt.: Cyrtomidictyum lepidocaulon (HOOK.) CHING // Bull. Fan Mem. 

Inst. Biol., Bot. 10: 182-183 (1940) 

Type: Tsu-shima, Wilford 565 // Japan 


(595) Polystichum macrophyllum (MAKINO) TAGAWA [IUCN: EN (VU)] 


Loc.: 大葉貫眾蕨蕨 - DaYeGuanZhongJue 

Bas.: Aspidium falcatum var. macrophyllum MAKINO // Bot. Mag. 

(Tokyo) 16: 90 (1902) 

Ety.: 

Alt.: 

makros (G: large), phyllon (G: leaf); relating to the pinna habit 

Cyrtomium macrophyllum (MAKINO) TAGAWA (var. 

macrophyllum) // Acta Phytotax. Geobot. 3: 62-63 (1934) 

Type: Kii, Mt. Mikoshi-toge, M. Miyoshi s. n., 22-Aug-1887 (HT: TI) // 

Japan 

TW: Ito, T. (1928) 

(596) Polystichum manmeiense (CHRIST) T. NAKAIKE 

Pub.: Misc. Publ. Nat. Sci. Mus. Tokyo: 141, pl. 4, f. 1 (1982) 

Loc.: 鐮葉耳蕨蕨 - LianYeErJue 

Bas.: Aspidium manmeiense CHRIST // Bull. Herb. Boissier 6: 965 

(1898)

131 

Ety.: relating to Manmei, a place in China and the type location 

Type: Yunnan, Henry 10097 (HT: P) // China 

TW: Kuo, C. M. 1985 (1985) 

(597) Polystichum mucronifolium (BLUME) C. PRESL 


Loc.: 兒玉氏耳蕨蕨 - ErYuShiErJue 

Bas.: Aspidium mucronifolium BLUME // Enum. Pl. Javae 164 (1828) 

Ety.: mucro (L: sharp point), folium (L: leaf); relating to the pinna 

margin 



(598) Polystichum neolobatum NAKAI [IUCN: EN] 

Pub.: Bot. Mag. (Tokyo) 39: 118 (1925), replacing Aspidium 

(Polystichum) lobatum var. chinense CHRIST in Nuovo 

Giornale Botanico Italiano, n. s., 4: 92 (1897) 

Loc.: 硬葉耳蕨蕨 - YingYeErJue 

Ety.: neos (G: new), lobatum (L: lobed) relating to the taxon with 

epithet "lobatum"; relating to the similarity to taxon 

Polystichum lobatum (HUDSON) C. PRESL 

Type: Sichuan // China 

TW: a: Tagawa, M. 1940-49 (1940) as var. brevipinnum TAGAWA 

b: FOT 1975 (1975) 

(599) Polystichum nepalense (SPRENG.) C. CHR. 

Pub.: Index Filic. 84 (1905), 585 (1906) 

Loc.: 軟骨耳蕨蕨 - RuanGuErJue 

Bas.: Aspidium nepalense SPRENG. // Syst. Veg. 4: 97 (1827) 

Ety.: relating to Nepal, the type location 

Type: Nepal 

TW: Ito, H. 1944 (1944) 

(600) Polystichum obliquum (D. DON) T. MOORE [IUCN: VU] 


Loc.: 知本耳蕨蕨 - ZhiBenErJue 

Bas.: Aspidium obliquum D. DON // Prodr. Fl. Nepal. 3 (1825) 

Ety.: obliquum (L: slanting); relating to the lamina habit 

Type: Wallich s. n. (IT: ?UC) // Nepal 


(601) Polystichum parvipinnulum TAGAWA 


Loc.: 尖尖葉耳蕨蕨 - JianYeErJue 

Ety.: parvum (L: small), pinnula (L: little feather); relating to the 

small size of pinnules 

Type: Prov. Tainan, Tozan in Arisan, Tagawa 552, 20-Aug-1934 (HT: 



132 

(602) Polystichum piceopaleaceum TAGAWA 


Loc.: 黑鱗耳蕨蕨 - HeiLinErJue 

Ety.: piceum (L: pitch black), palea (L: scale); relating to the colour 

of stipe base scale 

Type: Tagawa 554, 20-Aug-1934 (HT: KYO; IT: KYO) // Taiwan 


(603) Polystichum prescottianum (WALL. ex METT.) T. MOORE [IUCN: DD (EN)] 


Loc.: 南湖耳蕨蕨 - NanHuErJue 

Bas.: Aspidium prescottianum WALL. ex METT. // Abh. Senckenberg. 

Naturf. Ges. 2: 332-333 (1858) 

Ety.: in honor of J. D. Prescott (British botanist) 

Type: Wallich 363 (US, UC) // India 

TW: FOT 1975 (1975) 

(604) Polystichum prionolepis HAYATA 


Loc.: 鋸葉耳蕨蕨 - JuYeErJue 

Ety.: prion (G: saw), lepis (G: scale), relating to stipe and rachis 

scale margin 

Type: Arisan, ad 8000 ped. alt., B. Hayata & S. Sasaki s. n., Jan- 



(605) Polystichum pseudostenophyllum TAGAWA [IUCN: - (NT)] 


Loc.: 擬芽孢孢耳蕨蕨 - NiYaBaoErJue 

Ety.: pseudos (G: false), stenophyllum relating to the taxon with 

epithet "stenophyllum"; relating to the similarity to taxon 

Polystichum stenophyllum (see there) 

Type: Prov. Taityu, in monte Nokogoe, Ohwi 3228, 1933 (HT: KYO) 



(606) Polystichum scariosum (ROXB.) C. V. MORTON 

Pub.: Contr. U. S. Nat. Herb. 38: 359 (1974) 

Loc.: 阿里山耳蕨蕨 - ALiShanErJue 

Bas.: Polypodium scariosum ROXB. // Calcutta J. Nat. Hist. 4: 494 

(1844) 

Ety.: scariosus (L: thin); relating to the stipe hair habit (see 

protologue) 

Type: Amboyna, Roxburgh s. n. (LT: BR) // Indonesia 


(607) (Polystichum simadae comb. ined.) Cyrtomium macrophyllum var. 

simadae TAGAWA [IUCN: EN] 


Loc.: 尖尖葉貫眾蕨蕨 - JianYeGuanZhongJue

133 

Ety.: in honor of Y. Simada (Japanese botanist, collector of the type 

specimen) 

Alt.: Cyrtomium macrophyllum var. simadae TAGAWA // Acta 

Phytotax. Geobot. 3(2): 63 (1934) 

Type: Mt. Kiraisya-zan, Simada 5163B, Oct-1918 (HT: KYO) // 


TW: Acta Phytotax. Geobot. 3(2): 63 (1934) 

(608) Polystichum sinense (CHRIST) CHRIST 


Loc.: 福山氏耳蕨蕨 - FuShanShiErJue 

Bas.: Aspidium prescottianum var. sinense CHRIST // Bull. Soc. Bot. 

Ital. 10: 289-290 (1901) 


Type: Yunnan, Delavay 2173, 15-Jul-1886 (P) // China 

TW: MAN (2002) 

(609) Polystichum stenophyllum CHRIST 

Pub.: Bull. Soc. Bot. France, Mem. 1: 27-28 (1905) 

Loc.: 芽孢耳蕨蕨 - YaBaoErJue 

Ety.: stenos (G: narrow, straight), phyllon (G: leaf); relating to the 

lamina shape 

Type: Sichuan, David 1870 (HT: P) // China 

TW: Hayata, B. 1911 (1911) 

(610) Polystichum taiwanianum (TAGAWA) C. M. KUO [IUCN: VU (EN)] 


Loc.: 台台灣貫眾蕨蕨 - TaiWanGuanZhongJue 

Bas.: Cyrtomium taiwanianum TAGAWA // Acta Phytotax. Geobot. 7: 

78 (1938) 


Alt.: Cyrtomium taiwanianum TAGAWA // Acta Phytotax. Geobot. 7: 

78 (1938) 

Type: Prov. Taito, inter Kiriyama et Tippon-zan, Tagawa 1076, 4-Oct- 

1934 (HT: KYO) // Taiwan 


(611) Polystichum taizhongense H. S. KUNG [IUCN: - (NT)] 

Pub.: Acta Phytotax. Sin. 35(5): 461-462, f. 1 (1997) 

Loc.: 台台中耳蕨蕨 - TaiZhongErJue 

Ety.: relating to TaiZhong County (Taiwan, type location) 

Type: Taizhong, Nanhudashan, alt. 3500 m, S. J. Moore 12815, 12- 

Aug-1991 (HT: PYU; IT: TAIF) // Taiwan 

TW: Acta Phytotax. Sin. 35(5): 461-462, f. 1 (1997) 

(612) Polystichum tenuius (CHING) L. B. ZHANG * 


Pub.: Phytotaxa 60: 59 (2012) 

Loc.: 離脈柳葉蕨蕨 - LiMaiLiuYeJue 

Bas.: Cyrtogonellum tenuius CHING // Bull. Fan Mem. Inst. Biol., Bot. 

8: 329, pl. 7(2) (1938)

134 

Ety.: comparative of tenuis (L: thin, fine, slender); probably relating 

to the lamina texture 

Alt.: Cyrtogonellum tenuius CHING // Bull. Fan Mem. Inst. Biol., Bot. 

8: 329, pl. 7(2) (1938) 

Type: Guangxi, Lan Lon, 1133 m, 2-Aug-1928, R. C. Ching 6649 (LT: 

PE; ILT: LBG, PE) // China 

TW: Taiwania 59: 86-92 (2014) 

(613) Polystichum thomsonii (HOOK.) BEDD. [IUCN: NT] 


Loc.: 尾尾葉耳蕨蕨 - WeiYeErJue 

Bas.: Aspidium thomsonii HOOK. // Sec. Cent. Ferns pl. 25 (1861) 

Ety.: in honor of T. Thomson (British botanist) 

Type: Sikkim (K) // India 

TW: FOT 1975 (1975) 

(614) Polystichum tsussimense (HOOK.) J. SM. 

Pub.: Hist. Fil. 219 (1875) 

Loc.: 馬祖耳蕨蕨 - MaZuErJue 

Bas.: Aspidium tsus-simense HOOK. // Sp. Fil. 4: 16, pl. 220 (1862) 

Ety.: relating to Tsushima Island (Japan, type location) 

Type: Tsus Sima Island, C. Wilford s. n. (HT: K; IT: ?US) // Japan 

TW: Hayata, B. 1914-21 (1914) 

(615) Polystichum xiphophyllum (BAKER) DIELS [IUCN: EN] 


Loc.: 關山耳蕨蕨 - GuanShanErJue 

Bas.: Aspidium xiphophyllum BAKER // J. Bot. 26: 227 (1888) 

Ety.: xiphos (G: sword), phyllon (G: leaf); relating to the shape of 

lamina 

Type: Sichuan, Faber 1040 (HT: K; IT: MO, NY) // China 

TW: FOT 1975 (1975) 

PSEUDOPHEGOPTERIS (Thelypteridaceae) 

Ety.: pseudos (G: false), phegopteris relates to a fern genus; relating to 

the similarity to members of genus Phegopteris (see there) 

(616) Pseudophegopteris hirtirachis (C. CHR.) HOLTT. 

Pub.: Blumea 17: 22-23 (1969) 

Loc.: 毛囊紫柄蕨蕨 - MaoNangZiBingJue 

Bas.: Dryopteris hirtirachis C. CHR. // Fl. Kouy-Tcheou 49 (1915) 

Ety.: hirta (L: rough, hairy), rachia (G: literally a ridge, here relating 

to the rachis); relating to the rachis indument 

Type: Guizhou, Pinfa, Cavalerie 1810, replacing Dryopteris christii 

LEVEILLE (1915) [non C. CHR. (1905)] // China 

TW: FOT 1975 (1975) 

(617) Pseudophegopteris levingei (C. B. CLARKE) CHING [IUCN: NT] 


Loc.: 星毛紫柄蕨蕨 - XingMaoZiBingJue

135 

Bas.: Gymnogramma aurita var. levingei C. B. CLARKE // Trans. Linn. 

Soc. London, Bot. 1: 568 (1880) 

Ety.: in honor of H. C. Levinge (Irish botanist, and collector of type 

specimen) 

Type: Sikkim: Kashmir, Jhelum, and Chittapani Valleys, H. C. 

Levinge s. n. // India 

TW: Kuo, C. M. 1985 (1985) 

(618) Pseudophegopteris subaurita (TAGAWA) CHING 


Loc.: 光囊紫柄蕨蕨 - GuangNangZiBingJue 

Bas.: Dryopteris subaurita TAGAWA // Acta Phytotax. 1: 157-158 

(1932) 

Ety.: sub (L: similar to), aurita (L: eared); relating to the similarity to 

taxon Pseudophegopteris aurita (HOOK.) CHING 

Type: Pai-wan-sia, Ito s. n. (HT: KYO) // Taiwan 

TW: Acta Phytotax. 1: 157-158 (1932) 

PSILOTUM (Psilotaceae) 

Ety.: psilos (G: naked, smooth); relating to aerial stems being smooth, 

bearing naked sporangia 

(619) Psilotum nudum (L.) P. BEAUV. 

Pub.: Prodr. Aetheogam. 106, 112 (1805) 

Loc.: 松葉蕨蕨 - SongYeJue 

Bas.: Lycopodium nudum L. // Sp. Pl. 2: 1100-1101 (1753) 

Ety.: nudum (L: naked); relating to the lack of leaves 

Type: LINN. No. 1257.1 (LT: LINN) // India 


PTERIDIUM (Dennstaedtiaceae) 

Ety.: probably a derivative form of pteris, which relates to genus Pteris, or 

ferns in general 

(620) Pteridium latiusculum (DESV.) HIERON. ex FRIES 

Pub.: Wiss. Erg. D. Schwed. Rhod.-Kongo-Exped. 1: 7 (1914) 

Loc.: 蕨蕨 - Jue 

Bas.: Pteris latiuscula DESV. // Mem. Soc. Linn. Paris 6: 303 (1827) 

Ety.: latiusculum (L: somewhat broad); relating to the wide laminar 

segments 

Type: America boreali, Anon. (LT: P), designated by R. M. Tryon, 

Rhodora 43: 43 (1941) 

TW: Tagawa, M. 1940-49 (1941) 

(621) Pteridium revolutum (BLUME) NAKAI 


Loc.: 巒大蕨蕨 - LuanDaJue 

Bas.: Pteris revoluta BLUME // Enum. Pl. Javae 214 (1828) 

Ety.: revolutum (L: rolled back); relating to the segment margin

136 


TW: Kuo, C. M. 1985 (1985) 

PTERIDRYS (Dryopteridaceae) 

Ety.: a combination of the two generic names Pteris and Dryopteris (see 

there), which it each resembles in parts (height and venation of 

Pteris quadriaurita RETZ., stelar structure, sori and indusia of 

Dryopteris filix-mas (L.) SCHOTT) 

(622) Pteridrys cnemidaria (CHRIST) C. CHR. & CHING [IUCN: CR (EN)] 

Pub.: Bull. Fan Mem. Inst. Biol., Bot. 5: 136-138, pl. 12 (1934) 

Loc.: 長柄牙蕨蕨 - ChangBingYaJue 

Bas.: Dryopteris cnemidaria CHRIST // Bull. Acad. Int. Geogr. Bot. 20: 

140 (1910) 

Ety.: perhaps referring to Cnemidaria, a genus of small tree ferns 

(now lumped into Cyathea), and relating to the similarity of 

fronds to that of genus Cnemidaria (knemis (G: spoke of a 

wheel), aris (G: like); the lowest veins radiate like wheel 

spokes) 

Type: Cavalerie 3382 (E) // China 

TW: FOT 1975 (1975) 

PTERIS (Pteridaceae) 

Ety.: pteryx (G: wing, feather); relating to an old Greek name for ferns, 

derived from the resemblance of the pinnae to wings or feathers 

(623) Pteris amoena BLUME 


Loc.: 紅桿鳳鳳尾尾蕨蕨 - HongGanFengWeiJue 

Ety.: amoena (L: pleasing, lovely); relating to the plant habit 


TW: Kuo, C. M. 1985 (1985) 

(624) Pteris angustipinna TAGAWA [IUCN: CR] 


Loc.: 細葉鳳鳳尾尾蕨蕨 - XiYeFengWeiJue 

Ety.: angusta (L: narrow), pinna (L: wing, here relating to pinna); 

relating to the pinna habit 

Type: Taityu, inter Hattukwan et Tompo, Tagawa 478, 17-Aug-1934 

(HT: KYO; IT: KYO, MICH) // Taiwan 


(625) Pteris arisanensis TAGAWA 


Loc.: 三角脈鳳鳳尾尾蕨蕨 - SanJiaoMaiFengWeiJue 

Ety.: relating to ALiShan ("Arisan", Taiwan, the type location) 

Type: Prov. Tainan, Arisan, 2500 m, Faurie 603, Jun-1914 (HT: 



137 

(626) Pteris aspericaulis WALL. ex HIERON. 

Pub.: Recens. Spec. Pter. 22 (1839) 

Loc.: 紅柄鳳鳳尾尾蕨蕨 - HongBingFengWeiJue 

Ety.: asper (L: rough), caulis (L: stem); relating to the stipe habit 

Type: Wallich 107, 1820 (E, US) // Nepal 


(627) Pteris bella TAGAWA [IUCN: NT (LC)] 


Loc.: 長柄鳳鳳尾尾蕨蕨 - ChangBingFengWeiJue 

Ety.: bella (L: neat, handsome); relating to the habit 

Type: Prov. Takeo, between Daizyurin and Sinsuiei, Tagawa 2056, 

22-Jan-1939 (HT: KYO; IT: KYO, L, MICH, P) // Taiwan 


(628) Pteris biaurita L. 

Pub.: Sp. Pl. 2: 1076 (1753) 

Loc.: 弧脈鳳鳳尾尾蕨蕨 - HuMaiFengWeiJue 

Ety.: bi (L: two-), aurita (L: eared); relating to the lamina habit, 

probably the two elongated first basiscopic pinnules of basal 

most pair of pinnae 

Type: "Habitat in Domingo, Martinica, Jamaica" (LT: LINN), 

designated by R. M. Tryon in Contr. Gray Herb. 194: 201 

(1964) // Jamaica 


(629) Pteris cadieri CHRIST [IUCN: DD (LC)] 

Pub.: J. Bot. (Morot) 19: 72 (1905) 

Loc.: 二形鳳鳳尾尾蕨蕨 - ErXingFengWeiJue 

Ety.: in honor of P. L. M. Cadiere (French botanist and collector of 


Type: Cadiere 86 (HT: P; IT: BM, P) // Vietnam 

TW: Tagawa, M. 1940-49 (1940) 

(630) Pteris cretica L. subsp. cretica * 

Pub.: Mant. Pl. 1: 130 (1767) 

Loc.: 大葉鳳鳳尾尾蕨蕨 - DaYeFengWeiJue 

Ety.: relates to Crete (Greece, type location) 

Type: Crete, Anon. (LT: LINN), designated by R. M. Tryon, Contr. 

Gray Herb. 194: 192 (1964) // Greece 

TW: Henry, A. (1896) 

(631) Pteris cretica subsp. laeta (WALL. ex ETTINGSH.) FRASER-JENK. 


Loc.: 粗糙鳳鳳尾尾蕨蕨 - CuCaoFengWeiJue 

Bas.: Pteris laeta WALL. ex ETTINGSH. // Farnkr. Jetztw. 96, pl. 57 (f. 

8, 11), pl. 58 (f. 4, 12) (1865) 

Ety.: laeta (L: abundant, pleasing, vivid, bright); relationship not 

clear to me, perhaps relating to the lamina colour 

Type: Wallich 95 (NY, PH) // Nepal

138 

TW: FOT 1975 (1975) 

(632) Pteris dactylina HOOK. 

Pub.: Sp. Fil. 2: 160-161, pl. 130A (1858) 

Loc.: 掌鳳鳳尾尾蕨蕨 - ZhangFengWeiJue 

Ety.: daktylos (G: finger), ina (L: suffix, denoting likeness); relating 

to the lamina habit 

Type: Sri Lanka 

TW: Hayata, B. 1914-21 (1914) 

(633) Pteris deltodon BAKER 

Pub.: J. Bot. 26: 226 (1888) 

Loc.: 岩鳳鳳尾尾蕨蕨 - YanFengWeiJue 

Ety.: deltodon (L: having a triangular tooth); relating to the teeth at 

(sterile) pinna apex (see protologue) 

Type: Szechuan, Mt. Omei, Faber 1010 (IT: MO) // China 

TW: FOT 1975 (1975) 

(634) Pteris dispar KUNZE 


Loc.: 天草鳳鳳尾尾蕨蕨 - TianCaoFengWeiJue 

Ety.: dispar (L: unequal, different); relating to the pinna habit 

Type: Goering s. n. // Japan 


(635) Pteris ensiformis BURM. F. 

Pub.: Fl. Indica 230 (1768) 

Loc.: 箭葉鳳鳳尾尾蕨蕨 - JianYeFengWeiJue 


pinna shape 

Type: "Zeylona et Java" (a) Sri Lanka 

(b) Indonesia 

TW: J. Bot. 23: 103 (1885) 

(636) Pteris fauriei HIERON. 

Pub.: Hedwigia 55: 345-348 (1914) 

Loc.: 傅氏鳳鳳尾尾蕨蕨 - FuShiFengWeiJue 

Ety.: in honor of U. J. Faurie (French botanist) 

Type: Tamsui, Henry 1427 (B) // Taiwan 

TW: Hedwigia 55: 345-348 (1914) 

(637) Pteris formosana BAKER 

Pub.: J. Bot. 23: 103 (1885) 

Loc.: 台台灣鳳鳳尾尾蕨蕨 - TaiWanFengWeiJue 


Type: Hancock 83 (HT: K; IT: US) // Taiwan 

TW: J. Bot. 23: 103 (1885) 

(638) Pteris grevilleana WALL. ex J. AGARDH 

Pub.: Recens. Spec. Pter. 23-24 (1839) 

Loc.: 翅柄鳳鳳尾尾蕨蕨 - ChiBingFengWeiJue 

Ety.: in honor of R. K. Greville (British botanist)

Type: Kamoon, Wallich 2680 (HT: K) // India 

TW: Henry, A. (1896) 

(639) Pteris kawabatae SA. KURATA * 


Pub.: J. Geobot. 17: 59 (1969) 

Loc.: 無柄鳳鳳尾尾蕨蕨 - WuBingFengWeiJue 

Ety.: in honor of M. Kawabata (Japanese botanist, collector of the 


Type: Kagoshima, Yaku Isl., Tachu, M. Kawabata s. n., 7-Apr-1961 

(TOFO) // Japan 


(640) Pteris kidoi SA. KURATA [IUCN: NT] 

Pub.: J. Geobot. 13: 8-9 (1964) 

Loc.: 城戶氏鳳鳳尾尾蕨蕨 - ChengHuShiFengWeiJue 

Ety.: in honor of M. Kido (Japanese botanist, collector of the type 

specimen) 

Type: Kumamoto, Ashikitagun, Ashikitamachi, Shiraishi, M. Kido 

5880 (TOFO) // Japan 

TW: FOT 1975 (1975) 

(641) Pteris longipes D. DON 

Pub.: Prodr. Fl. Nepal. 15-16 (1825) 

Loc.: 蓬萊鳳鳳尾尾蕨蕨 - PengLaiFengWeiJue 

Ety.: longa (L: long), pes (L: foot); relating to the habit of stipe 

Type: Nepal 


(642) Pteris longipinna HAYATA 


Loc.: 長葉鳳鳳尾尾蕨蕨 - ChangYeFengWeiJue 

Ety.: longa (L: long), pinna (L: wing, here relating to pinna); relating 

to the pinna habit 

Type: Mt. Morrison, Kawakami & Mori 1875, Oct-1906 (HT: TI) // 



(643) Pteris multifida POIR. 

Pub.: Encycl. 5: 714 (1804) 

Loc.: 鳳鳳尾尾蕨蕨 - FengWeiJue 

Ety.: multi (L: many), fidi (L: indicative perfect of "findere", split); 

Type: 

139 

relating to the divided lamina habit 

Anon., cultivated at Hort. Herb. Paris, origin unknown (P, Herb. 

Lamarck, 2 sheets) 


(644) Pteris x namegatae KURATA 

Pub.: J. Geobot. 17: 60 (1969) 

Loc.: 

Ety.: 

行方氏鳳鳳尾尾蕨蕨 - XingFangShiFengWeiJue 

in honor of T. Namegata (Japanese botanist, (co-) collector of 

the type specimen)

140 

Type: Kagoshima (Satsuma), Ibusuki, T. Namegata & S. Kurata s. n., 

Dec-1960 (TOFO) // Japan 


(645) Pteris nipponica W. C. SHIEH [IUCN: DD] 


Loc.: 日本鳳鳳尾尾蕨蕨 - RiBenFengWeiJue 


Type: Nagasaki, Isl. Iki, Siwara, H. Ito s. n., 22-Jul-1962 (HT: TNS) // 

Japan 

TW: FOT 1975 (1975) 

(646) Pteris pellucida C. PRESL [IUCN: EN] 

Pub.: Reliq. Haenk. 1: 55 (1825) 

Loc.: 爪哇鳳鳳尾尾蕨蕨 - ZhuaWaFengWeiJue 

Ety.: pellucida (L: clear, transparent); relating to the pinna margin 

Type: Luzon, Haenke s. n. (PRC) // Philippines 

TW: a: Hayata, B. 1914-21 (1914), misidentification, probably 

corresponding to Pteris sp. in this list 


(647) Pteris pellucidifolia HAYATA [IUCN: NT] 

Pub.: Icon. Pl. Formosan. 6 (suppl.): 114 (1917) 

Loc.: 闊葉鳳鳳尾尾蕨蕨 - KuoYeFengWeiJue 

Ety.: two alternatives, the first being more likely based on relevant 

literature: (1) pellucida is probably relating to the taxon in the 

same genus with epithet "pellucida", folium (L: leaf); probably 

relating to the habit of (sterile) pinna, similar to those of Pteris 

pellucida (see there); (2) pellucida (L: clear, transparent), 

folium (L: leaf); relating to the quality of pinna 

Type: Arisan, Hayata & Sasaki s. n., Jan-1912 (?TI) // Taiwan 

TW: Icon. Pl. Formosan. 6 (suppl.): 114 (1917) 

(648) Pteris plumbea CHRIST [IUCN: DD (LC)] 

Pub.: Notul. Syst. (Paris) 1: 49-50 (1909) 

Loc.: 栗柄鳳鳳尾尾蕨蕨 - LiBingFengWeiJue 

Ety.: plumbea (L: leaden, lead-coloured); relating to the lamina 

colour 

Type: Luzon, Sablan, Elmer 6160, 21-Oct-1907 (B, P) // Philippines 

TW: Tagawa, M. 1940-49 (1940) 

(649) Pteris ryukyuensis TAGAWA [IUCN: NT (VU)] 


Loc.: 琉球鳳鳳尾尾蕨蕨 - LiuQiuFengWeiJue 

Ety.: relating to the Ryukyu Islands (Japan, type location) 

Type: Ryukyus, Z. Tasiro s. n., 11-Jan-1924 (HT: KYO; IT: KYO) // 

Japan 

TW: DeVol, C. E. (1979) 

(650) Pteris semipinnata L. 

Pub.: Sp. Pl. 2: 1076 (1753)

Loc.: 

Ety.: 

半邊羽裂鳳鳳尾尾蕨蕨 - BanBianYuLieFengWeiJue 

semi (L: half), pinnatum (L: pinnate); relating to the lamina 

dissection (being pinnate only on the lower side of each 

pinna, entire acroscopically) 

Osbeck s. n. (LINN) // China 

Type: 

TW: J. Bot. 23: 103 (1885) 

(651) Pteris setulosocostulata HAYATA 


Loc.: 

Ety.: 

141 

有刺鳳鳳尾尾蕨蕨 - YouCiFengWeiJue 

setulosa (L: with small bristles), costulatus (L: having small riblike 

ridges); relating to the costule habit 

Type: Mt. Arisan, B. Hayata & S. Sasaki s. n., Jan-1912 (HT: TI) // 



(652) Pteris sp. 

[IUCN: - (NT)] 


(653) Pteris terminalis WALL. ex J. AGARDH 

Pub.: Recens. Spec. Pter. 20 (1839) 

Loc.: 

Ety.: 

Type: 

溪鳳鳳尾尾蕨蕨 - XiFengWeiJue 

terminalis (L: terminal); relating to a prominent long terminal 

segment 

Wallich 101, 1829 (LT: K), designated by Fraser-Jenkins in 

Taxon. Revis. Indian Subcontinental Pteridophytes 416 

(2008) // Nepal 


(654) Pteris tripartita SW. 

[IUCN: DD] 


Loc.: 三腳鳳鳳尾尾蕨蕨 - SanJiaoFengWeiJue 

Ety.: tri (L: three), partita (L: divided); relating to the lamina division 

Type: Java, Thunberg s. n. (UPS) // Indonesia 

TW: a: Matsumara, J. & Hayata, B. (1906), as synonym of Pteris 

marginata BORY. ["Hab. Tamsui (ex Henry)"] 

b: Hayata, B. 1914-21 (1914) 

(655) Pteris vittata L. 

Pub.: Sp. Pl. 2: 1074 (1753) 

Loc.: 蜈蚣草 - WuGongCao 

Ety.: vittata (L: striped); relating to the lamina habit 

Type: Osbeck s. n. (HT: LINN) // China 

TW: Hayata, B. 1917 (1917) 

(656) Pteris wallichiana J. AGARDH 

Pub.: Recens. Spec. Pter. 69-70 (1839) 

Loc.: 

Ety.: 

Type: 

瓦氏鳳鳳尾尾蕨蕨 - WaShiFengWeiJue 

in honor of N. Wallich (Danish botanist, collector of type 

specimen) 

Wallich s. n. (?LD) // India

142 


(657) Pteris wulaiensis C. M. KUO [IUCN: CR (VU)] 

Pub.: Bot. Bull. Acad. Sin. 30(2): 143-145 (1989) 

Loc.: 烏來鳳鳳尾尾蕨蕨 - WuLaiFengWeiJue 

Ety.: relates to WuLai, a place in Taiwan and the type location 

Type: Taipei County, Tataoshan, Wulai, alt. 500 m, S. J. Moore 

4383, 14-Jun-1988 (HT: TAI; IT: Z) // Taiwan 

TW: Bot. Bull. Acad. Sin. 30(2): 143-145 (1989) 

PYRROSIA (Polypodiaceae) 

Ety.: pyr (G: fire); relating to the tawny indument of the type specimen 

(658) Pyrrosia angustissima (GIESENH. ex DIELS) TAGAWA & K. IWATS. [IUCN: VU] 


Loc.: 捲葉蕨蕨 - JuanYeJue 

Bas.: Niphobolus angustissimus GIESENH. ex DIELS // Nat. 

Pflanzenfam. 1(4): 326 (1899) 

Ety.: superlative of angusta (L: narrow); relating to the frond habit 

Type: Patung, Henry 5137 (HT: K) 

TW: FOT 1975 (1975) 

(659) Pyrrosia lanceolata (L.) FARWELL. 


Loc.: 抱樹石葦 - BaoShuShiWei 

Bas.: Acrostichum lanceolatum L. // Sp. Pl. 2: 1067 (1753) 

Ety.: lanceolata (L: lance-like); relating to the lamina habit 

Type: Herb. Hermann (LT: BM, designated by Tardieu-Blot in 

Aubreville (ed.), Fl. Gabon 8 : 201 (1964)) // Sri Lanka 

TW: MAN (2002) 

(660) Pyrrosia linearifolia (HOOK.) CHING 

Pub.: Bull. Chin. Bot. Soc. 1: 48 (1935) 

Loc.: 絨毛石葦 - RongMaoShiWei 

Bas.: Niphobolus linearifolius HOOK. // Sec. Cent. Ferns pl. 58 (1861) 

Ety.: linearis (L: linear), folium (L: leaf); relating to the lamina habit 

Type: Tsus-sima, Wilford s. n. (HT: K; IT: B, BM) // Japan 

TW: J. Bot. 23: 106 (1885) 

(661) Pyrrosia lingua (THUNB.) FARWELL 


Loc.: 石葦 - ShiWei 

Bas.: Acrostichum lingua THUNB. // Syst. Veg. (ed. 14) 928 (1784) 

Ety.: lingua (L: tongue); relating to the lamina habit 

Type: Thunberg s. n. (HT: UPS; IT: BM) // Japan 


(662) Pyrrosia matsudae (HAYATA) TAGAWA [IUCN: DD (LC)] 

Pub.: J. Jap. Bot. 24: 119-120 (1949) 

Loc.: 松田氏石葦 - SongTianShiShiWei

143 

Bas.: Cyclophorus matsudai HAYATA // Icon. Pl. Formosan. 10: 73, f. 

48 (1921) 

Ety.: in honor of Y. Matsuda (Japanese botanist, collector of the 


Type: Prov. Takao, Ariko-banti, Thabogan-goe, Matsuda s. n., Mar- 



(663) Pyrrosia polydactyla (HANCE) CHING 

Pub.: Bull. Chin. Bot. Soc. 1: 48 (1935) 

Loc.: 槭葉石葦 - CuYeShiWei 

Bas.: Polypodium polydactylon HANCE // J. Bot. 21: 269-270 (1883) 

Ety.: poly (G: many), daktylos (G: finger); relating to the lamina habit 

Type: Hancock 33 (HT: BM, Herb. Hance 22168; IT: US) // Taiwan 

TW: J. Bot. 21: 269-270 (1883) 

(664) Pyrrosia porosa (C. PRESL) HOVENKAMP [IUCN: DD (LC)] 

Pub.: Blumea 30(1): 208 (1984) 

Loc.: 中國石葦 - ZhongGuoShiWei 

Bas.: Niphobolus porosus C. PRESL // Tent. Pterid. 202 (1836) 

Ety.: porosa (L: porous, with pores); relating to the hydathodes on 

adaxial lamina 

Type: India Orientalis, Wallich 266 (HT: PRC; IT: B, BM, BR, K, M, P, 

US) 

TW: MAN (2002) 

(665) Pyrrosia sheareri (BAKER) CHING 

Pub.: Bull. Chin. Bot. Soc. 1: 64-65 (1935) 

Loc.: 盧山石葦 - LuShanShiWei 

Bas.: Polypodium sheareri BAKER // J. Bot. 13: 201 (1875) 

Ety.: in honor of J. Shearer (British botanist, collector of type 

specimen) 

Type: Shearer 282 (HT: K) // China 

TW: Ito, H. 1944 (1944) 

SALVINIA (Salviniaceae) 

Ety.: in honor of A. M. Salvini (Italian botanist) 

(666) Salvinia natans (L.) ALL. [IUCN: CR] 

Pub.: Fl. Pedem. 2: 289 (1785) 

Loc.: 槐葉蘋蘋 - HuaiYePing 

Bas.: Marsilea natans L. // Sp. Pl. 2: 1099 (1753) 

Ety.: natans (L: floating); relating to the growth habit, floating on 

water 

Type: (LT: LINN), designated by A. E. Bobrov in Conspectus 

filicarum Asiae Mediae et Kazakhstaniae. Novosti Sist. 

Vyssh. Rast. 21: 20 (1984) 

TW: Hayata, B. 1917 (1917)

144 

SCHIZAEA (Schizaeaceae) 

Ety.: schizein (G: to split); relating to the fan-shaped fronds being split into 

narrow lobes 

(667) Schizaea dichotoma (L.) J. SM. 

[IUCN: CR] 

Pub.: Mem. Acad. Sci. Turin 5: 422, t. 9, f. 9 (1793) 

Loc.: 分枝莎草蕨蕨 - FenZhiShaCaoJue 

Bas.: Acrostichum dichotomum L. // Sp. Pl. 2: 1068 (1753) 

Ety.: dichotoma (L: dividing into two equal parts); relating to the 

fronds 

Type: Herb. Sloane s. n. (BM) // China 

TW: FOT 1975 (1975) 

(668) Schizaea digitata (L.) SW. 

[IUCN: VU] 

Pub.: Syn. Fil. 150 (1806) 

Loc.: 莎草蕨蕨 - ShaCaoJue 

Bas.: Acrostichum digitatum L. // Sp. Pl. 2: 1068 (1753) 

Ety.: digitata (L: radiating like fingers); relating to the arrangement of 

sorophores 

Type: Herb. Hermann s. n. (LT: BM, designated by Holttum in Fl. 

Malesiana, ser. II, 1: 41 (1959) // Sri Lanka 

TW: Ito, T. (1928) 

SCLEROGLOSSUM (Grammitidaceae) 

Ety.: skleros (G: hard), glossa (G: tongue); relating to the strap-shaped 

and leathery fronds 

(669) Scleroglossum sulcatum (METT. ex KUHN) ALDERW. [IUCN: VU (EN)] 

Pub.: Bull. Jard. Bot. Buitenzorg, ser. 2, 7: 37, pl. 5, f. 3-4 (1912) 

Loc.: 革舌蕨蕨 - GeSheJue 

Bas.: Vittaria sulcata METT. ex KUHN // Linnaea 36: 68 (1869) 

Ety.: sulcatum (L: grooved); relating to the midrib of lamina 

Type: Thwaites 3807 (LT: B; ILT: BM, CGE, GH, K, P, PDA) // Sri 

Lanka 

TW: Parris, B. S. et al. (2010) 

SELAGINELLA (Selaginellaceae) 

Ety.: selago relates to the lycopod taxon Lycopodium selago L., see Sp. 

Pl. 2: 1102 (1753), followed by ella (L: diminutive suffix); relating to 

the resemblance of a "small Selago" 

(670) Selaginella aristata SPRING 

Pub.: Bull. Acad. Roy. Sci. Bruxelles 10: 232 (1843) 

Loc.: 膜葉卷卷柏 - MoYeJuanBo 

Ety.: aristata (L: awned, bearded); relating to the presence of a long 

bristle at the apex of pinnules 

Type: P. Commerson 110 (HT: P; IST: E) // Philippines 

TW: Chang, H. M. et al. (2012)

145 

(671) Selaginella boninensis BAKER [IUCN: NT] 

Pub.: J. Bot. 23: 178 (1885) 

Loc.: 小小笠原原卷卷柏 - XiaoLiYuanJuanBo 

Ety.: relating to the Bonin Islands (Japan), the type location 

Type: Bonin Islands, C. Wright 317, 1853-56 (HT: K, IT: US) // Japan 

TW: Tagawa, M. 1940-49 (1941) 

(672) Selaginella ciliaris (RETZ.) SPRING [IUCN: VU] 


Loc.: 緣毛卷卷柏 - YuanMaoJuanBo 

Bas.: Lycopodium ciliare RETZ. // Observ. Bot. 5: 32 (1789) 

Ety.: 

Type: 

ciliata (L: bearing fine hairs); relating to the leaves 

eastern Ceylon, König s. n. (LT: LD; ILT: K), designated by 

Alston (1934) // Sri Lanka 

TW: Tagawa, M. 1940-49 (1941) 

(673) Selaginella delicatula (DESV. ex POIR.) ALSTON 

Pub.: J. Bot. 70: 282 (1932) 

Loc.: 全緣卷卷柏 - QuanYuanJuanBo 

Bas.: Lycopodium delicatulum DESV. ex POIR. // Encycl., Suppl. 3: 

554 (1813, publ. 1814) 

Ety.: delicata (L: dainty or pleasant); relating to the habit 

Type: (a) l'Amerique meridionale (without any further details) (HT: P) 

(b) Pisang Isl., near New Guinea, C. Gaudichaud 13 (NT: G; 

INT: P), designated by Alston (1945) // Indonesia 


(674) Selaginella devolii H. M. CHANG, P. F. LU & W. L. CHIOU [IUCN: - (EN)] 

Pub.: Blumea 56(1): 21-23, f. 1 (2011) 

Loc.: 棣氏卷柏 - DiShiJuanBo 


Type: Nantou County, Chushan, Fuchouli Village, Fangliao, H. M. 

Chang 6635, 23-Oct-2009 (HT: TAIF; IT: L, TAIE) // Taiwan 

TW: Blumea 56(1): 21-23, f. 1 (2011) 

(675) Selaginella doederleinii HIERON. 

Pub.: Hedwigia 43(1): 41-42 (1904) 

Loc.: 生根卷卷柏 - ShengGenJuanBo 

Ety.: in honor of L. H. P. Döderlein (German scientist and collector 


Type: Ryukyu, Amami-Oshima, Doederlein s. n. (LT: B), designated 

by Alston (1934) // Japan 


(676) Selaginella helvetica subsp. pseudonipponica (TAGAWA) H. M. 

CHANG, W. L. CHIOU & J. C. WANG [IUCN: DD (LC)] 

Pub.: Fl. Taiwan, Selaginellaceae 38 (2012) 

Loc.: 擬日本卷卷柏 - NiRiBenJuanBo 

Bas.: Selaginella pseudonipponica TAGAWA // Acta Phytotax. 

Geobot. 25: 177 (1973)

146 

Ety.: helvetica (L: of Switzerland) relating to the taxon with epithet 

"helvetica" (not present in Taiwan), pseudos (G: false), 

nipponica relating to the taxon with epithet "nipponica"; 

relating to the similarity to taxon Selaginella nipponica (see 

there) 

Type: Hualien, between Shinjo and Tabito, Ohwi 1126 (HT: KYO) // 



(677) Selaginella heterostachys BAKER 

Pub.: J. Bot. 23: 177 (1885) 

Loc.: 姬卷卷柏 - JiJuanBo 

Ety.: heteros (G: different), stachys (G: spike); probably relating to 

the two different kinds of bracts in the spike (see protologue) 

Type: Hong Kong, C. Wright s. n., 1853-56 (LT: K; ILT: GH, NY, P, 

US), designated by Alston (1934) // China 


(678) Selaginella involvens (SW.) SPRING 


Loc.: 密密葉卷卷柏 - MiYeJuanBo 

Bas.: Lycopodium involvens SW. // Syn. Fil. 182 (1806) 

Ety.: involvens (L: rolling up, wrapping up); relating to the branchlets 

Type: (HT: S; IT: B) // Japan 

TW: Henry, A. (1896) 

(679) Selaginella labordei HIERON. ex CHRIST 

Pub.: Bull. Acad. Int. Geogr. Bot. 11: 272-273 (1902) 

Loc.: 玉山卷卷柏 - YuShanJuanBo 

Ety.: in honor of J. Laborde, (co-) collector of the type specimen 

Type: Guizhou (Kouy-Tcheou), Laborde & Bodinier 2758, 11-Sep- 

1899, (LT: E; ILT: P), designated by Alston (1934) // China 

TW: Tagawa, M. 1940-49 (1941) 

(680) Selaginella lutchuensis KOIDZ. [IUCN: EN] 


Loc.: 琉球卷卷柏 - LiuQiuJuanBo 

Ety.: relating to the Ryukyu Islands (Japan) 

Type: Ryukyu, Amami-oshima Island, U. Faurie 4655, Jul-1900 (HT: 

KYO; IT: P) // Japan 


(681) Selaginella matsuensis C. M. KUO (nom. nud.) 


Loc.: 馬祖卷卷柏 - MaZuJuanBo 

Ety.: relating to the MaZu Islands (Taiwan), the location where this 

taxon was first observed 


(682) Selaginella moellendorffii HIERON. 

Pub.: Nat. Pflanzenfam. 1(4): 680 (1901, publ. 1902) 

Loc.: 異葉卷卷柏 - YiYeJuanBo

147 

Ety.: in honor of O. F. Möllendorff (German botanist, collector of the 


Type: Jiangxi, prope Kiukiang ad fluvium Yangtse, O. Möllendorff 26 

(LT: B), designated by Alston (1934) // China 


(683) Selaginella nipponica FRANCH. & SAV. [IUCN: CR] 

Pub.: Enum. Pl. Jap. 2: 199 (1877), 615 (1878) 

Loc.: 日本卷卷柏 - RiBenJuanBo 


Type: near Yokoska, Savatier 2415 (LT: P; ILT: K), selected by H. M. 

Chang et al., Flora Taiwan Selaginellaceae 67 (2012) // 

Japan 

TW: FOT 1975 (1975), in part including Selaginella helvetica subsp. 

pseudonipponica 

(684) Selaginella remotifolia SPRING 

Pub.: Pl. Jungh. 3: 276 (1854) 

Loc.: 疏葉卷卷柏 - ShuYeJuanBo 

Ety.: remota (L: distant), folium (L: leaf); relating to the trophophylls, 

which are well spaced along the stem 

Type: Sumatra, F. W. Junghuhn s. n. (HT: ?H) // Indonesia 

TW: Tagawa, M. 1940-49 (1941) 

(685) Selaginella repanda (DESV. ex POIR.) SPRING 

Pub.: Voy. Bonite, Bot. 3: 329 (1846) 

Loc.: 高雄卷卷柏 - GaoXiongJuanBo 

Bas.: Lycopodium repandum DESV. ex POIR. // Encycl., Suppl. 3: 558 

(1813, publ. 1814) 

Ety.: repanda (L: with a slightly uneven margin); relating to the 

leaves 

Type: in insula Manila, ?N. A. Desvaux 124 (HT: ?P) // Philippines 

TW: Tagawa, M. 1940-49 (1941) 

(686) Selaginella sp. 



(687) Selaginella stauntoniana SPRING 

Pub.: Mem. Acad. Roy. Sci. Belgique 24: 71 (1850) 

Loc.: 擬密密葉卷卷柏 - NiMiYeJuanBo 

Ety.: in honor of G. L. Staunton (Irish botanist, collector of the type 

specimen) 

Type: Jiangxi, G. L. Staunton s. n. (LT: FI; ILT: BM), designated by 

Alston (1934) // China 


(688) Selaginella tama-montana SERIZ. [IUCN: - (EN)] 

Pub.: J. Jap. Bot. 53(8): 242 (1978) 

Loc.: 山地卷卷柏 - ShanDiJuanBo 

Ety.: relating to the type location in Japan, the Tama Mountains

148 

Type: Honshu, Pref. Tokyo, near Hacho-bashi, alt. ca. 680 m (HT: 

AICH; IT: KYO, TI, TNS) // Japan 

TW: Kuo, C. M. 2010 (2010) 

(689) Selaginella tamariscina (P. BEAUV.) SPRING 


Loc.: 萬年松 - WanNianSong 

Bas.: Stachygynandrum tamariscinum P. BEAUV. // Mag. Encycl. 

Ety.: 

9(5): 483 (1804) 

probably relating to tamarix, a flowering plant genus 

(Tamaricaceae, "tamarisk"), ina (L: suffix, denoting likeness), 

and suggesting its resemblance to the tamarisk tree 

Type: India orientalis, A. M. F. J. Palisot de Beauvois s. n. (LT: B; 

ILT: P), designated by Alston (1934) // India 


SELLIGUEA (Polypodiaceae) 

Ety.: in honor of A. F. Gilles, called Selligue (French instrument maker) 

(690) Selliguea echinospora (TAGAWA) FRASER-JENK. 

Pub.: Taxon. Revis. Indian Subcontinental Pteridophytes 46-47 

(2008) 

Loc.: 大葉玉山茀蕨蕨 - DaYeYuShanFuJue 

Bas.: Phymatodes echinospora TAGAWA // Acta Phytotax. Geobot. 3: 

95-96 (1934) 

Ety.: echinos (G: hedge-hog), spora (G: seed); relating to the 

ornamentation of spore 

Type: Mt. Tugitaka, alt. 2100 m, Ito s. n., 1925 (HT: KYO) // Taiwan 


(691) Selliguea engleri (LUERSS.) FRASER-JENK. 


Loc.: 恩氏茀蕨蕨 - EnShiFuJue 

Bas.: Polypodium engleri LUERSS. // Bot. Jahrb. Syst. 4: 361-362 

(1883) 

Ety.: in honor of H. G. A. Engler (German botanist) 

Type: Kawo-nabe, L. Döderlein s. n., Mar-1880 (HT: B) // Japan 

TW: Hayata, B. 1911 (1911) 

(692) Selliguea falcatopinnata (HAYATA) H. OHASHI & K. OHASHI [IUCN: VU] 

Pub.: J. Jap. Bot. 84(5): 306 (2009) 

Loc.: 掌葉茀蕨蕨 - ZhangYeFuJue 

Bas.: Polypodium falcatopinnatum HAYATA // Icon. Pl. Formosan. 4: 

247-248, f. 172 (1914) 

Ety.: falcata (L: sickle-shaped), pinnata (L: pinnate); relating to the 

lamina shape (pinnae being falcate) 

Type: Kotosho, T. Kawakami & S. Sasaki s. n., Jul-1912 (HT: TI) // 


TW: Icon. Pl. Formosan. 4: 247-248, f. 172 (1914)

(693) Selliguea hastata (THUNB.) FRASER-JENK. 

Pub.: Taxon. Revis. Indian Subcontinental Pteridophytes 44-45 

(2008) 

Loc.: 三葉茀蕨蕨 - SanYeFuJue 

Bas.: Polypodium hastatum THUNB. // Fl. Jap. 335 (1784) 

Ety.: hastata (L: spear-shaped); relating to the lamina habit 

Type: Nagasaki, Thunberg s. n. (HT: UPS) // Japan 


(694) Selliguea okamotoi (TAGAWA) RALF KNAPP [IUCN: DD (VU)] 


Loc.: 岡本氏茀蕨蕨 - GangBenShiFuJue 

Bas.: Phymatodes okamotoi TAGAWA // Acta Phytotax. Geobot. 7: 

Ety.: 

189 (1938) 

in honor of S. Okamoto (Japanese botanist, collector of type 

specimen) 

Type: Prov. Takao, between Takimi and Hinode, ca. 2000 m, S. 

Okamoto s. n., 24-Sep-1937 (HT: KYO) // Taiwan 


(695) Selliguea quasidivaricata (HAYATA) H. OHASHI & K. OHASHI 

Pub.: J. Jap. Bot. 84(5): 307 (2009) 

Loc.: 

Bas.: 

Ety.: 

149 

玉山茀蕨蕨 - YuShanFuJue 

Polypodium quasidivaricatum HAYATA // J. Coll. Sci. Imp. Univ. 

Tokyo 30: 446 (1911) 

quasi (L: as if, nearly), divaricatum (L: spreading apart); 

relating to the way the basal pinnae are more or less 

spreading 

Type: Mt. Morison, Kawakami & Mori 1871 // Taiwan 

TW: J. Coll. Sci. Imp. Univ. Tokyo 30: 446 (1911) 

(696) Selliguea taiwanensis (TAGAWA) H. OHASHI & K. OHASHI [IUCN: DD (LC)] 

Pub.: J. Jap. Bot. 84(5): 308 (2009) 

Loc.: 台台灣茀蕨蕨 - TaiWanFuJue 

Bas.: Phymatodes taiwanensis TAGAWA // Acta Phytotax. Geobot. 

11: 310 (1942) 


Type: Prov. Taihoku, Mt. Nanko-taisan, J. Ohwi 2607 (HT: KYO) // 



(697) Selliguea yakushimensis (MAKINO) FRASER-JENK. [IUCN: NT] 


Loc.: 姬茀蕨蕨 - JiFuJue 

Bas.: Polypodium engleri var. yakushimense MAKINO // Bot. Mag. 

(Tokyo) 23: 248 (1909) 

Ety.: relating to Yakushima Island (Japan, type location) 

Type: Prov. Oshima, Yaku-shima, Makino s. n., Sep-1909 (HT: TI) // 

Japan

150 

TW: Ito, T. (1928) 

SPHAEROCIONIUM (Hymenophyllaceae) 

Ety.: sphaera (G: sphere), kionion (G: small column); relating to the 

sporangia which are born on a small pedicel 

(698) (Sphaerocionium digitatum comb. ined.) Hymenophyllum digitatum 

(SW.) FOSBERG [IUCN: EN (CR?)] 

Pub.: Smithsonian Contr. Bot. 45: 1 (1980) 

Loc.: 指裂細口團扇蕨蕨 - ZhiLieXiKouTuanShanJue 

Bas.: Trichomanes digitatum SW. // Syn. Fil. 370, 422 (1806) 

Ety.: digitatum (L: radiating like fingers); relating to the arrangement 

of lamina segments 

Alt.: Hymenophyllum digitatum (SW.) FOSBERG // Smithsonian 

Contr. Bot. 45: 1 (1980) 

Type: "In Ins. Franciae et Borboniae", collector unknown (HT: ?S) // 

Mauritius and Reunion 


(699) Sphaerocionium nitidulum (BOSCH) K. IWATS. [IUCN: NT] 


Loc.: 細口團扇蕨蕨 - XiKouTuanShanJue 

Bas.: Trichomanes nitidulum BOSCH // Pl. Jungh. 1: 547 (1856) 

Ety.: nitidulum (L: bright, glittering); relating to lamina habit 

Alt.: Hymenophyllum nitidulum (BOSCH) EBIHARA & K. IWATS. // 

Taxon 53(4): 941 (2004) 

Type: Java, Junghuhn s. n. (HT: L; IT: P) // Indonesia 

TW: Ito, H. 1981 (1981) 

(700) Sphaerocionium pilosissimum (C. CHR.) COPEL. [IUCN: - (EN)] 


Loc.: 尾尾毛膜蕨蕨 - WeiMaoMoJue 

Bas.: Hymenophyllum pilosissimum C. CHR. // Gard. Bull. Straits 

Settlem., ser. 3, 7: 213 (1934) 

Ety.: superlative of pilosum (L: pilose); relating to the abundant 

presence of hairs 

Alt.: Hymenophyllum pilosissimum C. CHR. // Gard. Bull. Straits 

Settlem., ser. 3, 7: 213 (1934) 

Type: Borneo, Kinabalu, Burbidge s. n. (BM, K, UC) // Malaysia 

TW: Amer. Fern J. 100(3): 180-187 (2010) 

TAPEINIDIUM (Dennstaedtiaceae) * 

Ety.: tapeinos (G: small, dwarf), idium (G: a diminutive suffix); relationship 

unknown to me, perhaps relating to the plant habit or the small sori 

size 

(701) Tapeinidium biserratum (BLUME) ALDERW. [IUCN: VU] 


Loc.: 二羽達邊蕨蕨 - ErYuDaBianJue

Bas.: Davallia biserrata BLUME // Enum. Pl. Javae 232 (1828) 

Ety.: bi (L: two-), serratum (L: serrated); relating to the margin of 

pinna 

Type: Java, Blume s. n. (?Reinwardt s. n.) (L) // Indonesia 

TW: FOT 1975 (1975) 

(702) Tapeinidium pinnatum (CAV.) C. CHR. 

Pub.: Index Filic. 213 (1905), 631 (1906) 

Loc.: 達邊蕨蕨 - DaBianJue 

Bas.: Davallia pinnata CAV. // Descr. Pl. 277 (1802) 

Ety.: pinnatum (L: pinnate); relating to the lamina shape (being 

pinnate) 

Type: Nee s. n. (MA) // Philippines 

TW: a: Matsumara, J. & Hayata, B. (1906), as Microlepia pinnata 

var. gracilis BAKER 

b: Sasaki, S. (1928) 

TECTARIA (Dryopteridaceae) 

Ety.: tectum (L: roof), aria (L: a substantive suffix); relating to the roof-like 

indusium in some species 

(703) Tectaria coadunata (J. SM.) C. CHR. 


Loc.: 陰地三叉蕨蕨 - YinDiSanChaJue 

Bas.: Sagenia coadunata J. SM. // J. Bot. (Hooker) 4: 184 (1842), 

based on Aspidium coadunatum WALL. ex HOOK. & GREV., 

Icon. Filic. t. 202 (1831) [not KAULFUSS (1824)] 

Ety.: coadunata (L: united); relating to the lamina habit (uniting of 

pinnae at rachis) 

Type: Wallich 377 (HT: E; IT: UC) // Nepal 


(704) Tectaria decurrens (C. PRESL) COPEL. 

Pub.: Leafl. Philipp. Bot. 1: 234 (1907) 

Loc.: 翅柄三叉蕨蕨 - ChiBingSanChaJue 

Bas.: Aspidium decurrens C. PRESL // Reliq. Haenk. 1: 28 (1825) 

Ety.: decurrens (L: running down); relating to the habit of basal 

pinnae at rachis 

Type: Luzon, Haenke s. n. (PR) // Philippines 

TW: J. Bot. 23: 105 (1885) 

(705) Tectaria devexa (KUNZE ex METT.) COPEL. 


Loc.: 薄薄葉三叉蕨蕨 - BoYeSanChaJue 

Bas.: Aspidium devexum KUNZE ex METT. // Ann. Mus. Bot. 

Lugduno-Batavi 1: 237 (1864) 

Ety.: devexa (L: sloping); relating to the sloping pinnae 

Type: Java, Zollinger 2717 (HT: LZ, lost; IT: B, G, L) // Indonesia 


151

152 

(706) Tectaria dissecta (G. FORST.) LELLINGER 

Pub.: Amer. Fern J. 58: 156 (1968) 

Loc.: 南洋三叉蕨蕨 - NanYangSanChaJu 

Bas.: Polypodium dissectum G. FORST. // Fl. Ins. Austr. 81 (1786) 

Ety.: dissectus (L: dissected); relating to the lamina habit 

Type: G. Forster s. n. (HT: BM; IT: K) // Pacific Islands 


(707) Tectaria dubia (C. B. CLARKE & BAKER) CHING [IUCN: VU] 

Pub.: Sinensia 2: 23, f. 5 (1931) 

Loc.: 大葉叉蕨蕨 - DaYeChaJue 

Bas.: Nephrodium cicutarium var. dubia C. B. CLARKE & BAKER // J. 

Linn. Soc. 24: 417 (1888) 

Ety.: dubia (L: uncertain); relating to the intermediate morphology 

between two taxa (of Nephrodium, see protologue) 

Type: Assam, Luckimpore, C. B. Clarke 37819 (?K, ?NY) // India 


(708) Tectaria fauriei TAGAWA 

Pub.: J. Jap. Bot. 14: 102-104 (1938) 

Loc.: 傅氏三叉蕨蕨 - FuShiSanChaJue 

Ety.: in honor of U. J. Faurie (French botanist and collector of type 

specimen) 

Type: Prov. Takao, Bankinsing, alt. 800 m, U. Faurie 67, Feb-1914 

(HT: KYO; IT: KYO) // Taiwan 

TW: J. Jap. Bot. 14: 102-104 (1938) 

(709) Tectaria fuscipes (WALL. ex BEDD.) C. CHR. 


Loc.: 屏屏東擬肋毛蕨蕨 - PingDongNiLeMaoJue 

Bas.: Aspidium fuscipes WALL. // Numer. List n. 361 (ex parte) 

(1827) (nom. nud.) ex Beddome, Suppl. Ferns Brit. India 15, 

pl. 366 (1876) 

Ety.: fuscus (L: brown), pes (L: foot); relating to the colour of the 

stipe 

Type: C. B. Clarke 7050, Cachar, Assam (K) // India 

TW: FOT 1975 (1975) 

(710) Tectaria griffithii (BAKER) C. CHR. 


Loc.: 葛氏三叉蕨蕨 - GeShiSanChaJue 

Bas.: Nephrodium griffithii BAKER // Syn. Fil. 300 (1867) 

Ety.: in honor of W. Griffith (British botanist and collector of type 

specimen) 

Type: 

TW: Hayata, B. 1917 (1917) 

(711) Tectaria harlandii (HOOK.) C. M. KUO 

Sikkim, Griffith s. n. (K) // ?Myanmar 


Loc.: 沙皮蕨蕨 - ShaPiJue

153 

Bas.: Acrostichum harlandii HOOK. // Sp. Fil. 5: 274 (1864) 

Ety.: in honor of W. A. Harland (British botanist and collector of type 

specimen) 

Type: Hongkong, Mt. Gough, Harland s. n. // China 

TW: Henry, A. (1896) 

(712) Tectaria harlandii (HOOK.) C. M. KUO x Tectaria subtriphylla (HOOK. & 

ARN.) COPEL. 


(713) Tectaria impressa (FEE) HOLTT. 

Pub.: Kew Bull. 43(3): 483 (1988) 

Loc.: 變葉三叉蕨蕨 - BianYeSanChaJue 

Bas.: Phlebiogonium impressum FEE // Mem. Foug. 5: 314, pl. 24A, 

f. 2 (1852) 

Ety.: impressa (L: pressed into); relating to the sori position on 

lamina 

Type: Griffith s. n. (HT: RB) 


(714) Tectaria kusukusensis (HAYATA) LELLINGER 

Pub.: Amer. Fern J. 58: 157 (1968) 

Loc.: 高士佛三叉蕨蕨 - GaoShiFoSanChaJue 

Bas.: Dryopteris kusukuensis HAYATA // Icon. Pl. Formosan. 4: 157- 

158, f. 98 (1914) 

Ety.: relating to Kusukusu, a place in Taiwan and the type location 

Type: Kusukusu, B. Hayata & S. Sasaki s. n., Jul-1912 (TI) // Taiwan 


(715) Tectaria multicaudata (C. B. CLARKE) CHING [IUCN: CR] 

Pub.: Sinensia 2(2): 20 (1931) 

Bas.: Nephrodium multicaudatum C. B. CLARKE // Trans. Linn. Soc. 

London, Bot. 1(8): 540, t. 77 (1880) 

Ety.: multi (L: many), caudata (L: having a tail); relating to the 

lamina habit 

Type: Sylhet, Clarke 18427, 30-Nov-1887 (HT: K) // Bangladesh 

TW: a: Kuo, C. M. 1985 (1985) for Tectaria griffithii auct. non 

(BAKER) C. CHR. 

B: Knapp, R. 2014 (2014) 

(716) Tectaria phaeocaulis (ROSENST.) C. CHR. 


Loc.: 蛇脈三叉蕨蕨 - SheMaiSanChaJue 

Bas.: Aspidium phaeocaule ROSENST. // Hedwigia 56: 345 (1915) 

Ety.: phaios (G: dark, dusky), caulon (G: stalk, stem); relating to the 

dark coloured frond axes (see protologue) 

Type: Urai, Faurie 33 (IT: KYO) // Taiwan 

TW: Hedwigia 56: 345 (1915) 

(717) Tectaria polymorpha (WALL. ex HOOK.) COPEL. 

Pub.: Philipp. J. Sci. 2: 413 (1907)

154 

Loc.: 

Bas.: 

Ety.: 

南投三叉蕨蕨 - NanTouSanChaJue 

Aspidium polymorphum WALL. // Numer. List n. 382 (1828), ex 

Hooker, Sp. Fil. 4: 54-55 (1862) 

poly (G: many), morphe (G: shape, form); relating to the 

variable shape of lamina 

Wallich 382 (HT: K; IT: BM, E) // Nepal 

Type: 

TW: Henry, A. (1896) 

(718) Tectaria simonsii (BAKER ) CHING 

Pub.: Sinensia 2: 32-33, f. 13 (1931) 

Loc.: 紫柄三叉蕨蕨 - ZiBingSanChaJue 

Bas.: Nephrodium simonsii BAKER // Syn. Fil. (ed. 2) 504 (1874) 

Ety.: in honor of C. J Simons (British botanist), collector of the type 

specimen 

Type: Sikkim // India 

TW: Kuo, C. M. 1985 (1985) 

(719) Tectaria subfuscipes (TAGAWA) C. M. KUO 

Pub.: Stud. Ferns Ken-Ting Nation. Park 66 (1986) 

Loc.: 排灣三叉蕨蕨 - PaiWanSanChaJue 

Bas.: Ctenitopsis subfuscipes TAGAWA // Acta Phytotax. Geobot. 8: 

168-169 (1939) 

Ety.: sub (L: similar to), fuscipes relating to the taxon with epithet 

"fuscipes"; relating to the similarity to taxon Tectaria fuscipes 

(see there) 

Type: Prov. Takao, Kizan-gun, between Sansaro and Oohara, M. 

Tagawa 2389, 13-Dec-1938 (HT: KYO) // Taiwan 


(720) Tectaria subtriphylla (HOOK. & ARN.) COPEL. 


Loc.: 三叉蕨蕨 - SanChaJue 

Bas.: Polypodium subtriphyllum HOOK. & ARN. // Bot. Beechey Voy. 

256, pl. 50 (1841) 

Ety.: sub (L: similar to), triphylla (L: three-leaved); relating to the 

shape of lamina 

Type: Guangdong // China 

TW: Henry, A. (1896) 

(721) Tectaria sulitii COPEL. [IUCN: - (EN)] 


Loc.: 多羽三叉蕨蕨 - DuoYuSanChaJue 

Ety.: in honor of M. D. Sulit (Philippine botanist, collector of the type 

specimen) 

Type: Mindanao, Bukidnon Prov., Mt. Katanglad, M. D. Sulit 8727, 

10-Mar-1949 (HT: MICH) // Philippines 

TW: TW J. of Biodivers. 13: 175-178 (2011) 

(722) Tectaria zeilanica (HOUTT.) SLEDGE 

Pub.: Kew Bull. 27: 422-423 (1972)

155 

Loc.: 地耳蕨蕨 - DiErJue 

Bas.: Ophioglossum zeilanicum HOUTT. // Nat. Hist. 2: 43, pl. 94, f. 1 

(1783) 

Ety.: relating to Sri Lanka ("Ceylon"), the type location 

Type: Sri Lanka 


THELYPTERIS (Thelypteridaceae) 

Ety.: thelys (G: female), pteris (relates to genus Pteris, or ferns in 

general); relates to an old Greek name for a fern more delicate 

than the male fern (Dryopteris filix-mas (L.) SCHOTT) 

(723) Thelypteris adscendens CHING 


Loc.: 微毛金星蕨蕨 - WeiMaoJinXingJue 

Ety.: adscendens (L: ascend, rise); relating to the pinna habit 

Type: Guangxi, vicinity of Wuzhou, Wang 5171 (HT: PE) // China 

TW: FOT 1975 (1975) 

(724) Thelypteris angulariloba CHING 


Loc.: 鈍頭金星蕨蕨 - DunTouJinXingJue 

Ety.: angularis (L: having corners), loba (L: lobe); probably relating 

to the segment apex habit 

Type: Guangdong, Jaoping, Chun 42644 (HT: PE; IT: MO) // China 

TW: FOT 1975 (1975) 

(725) Thelypteris angustifrons (MIQ.) CHING 


Loc.: 小小梯葉副金星蕨蕨 - XiaoTiYeFuJinXingJue 

Bas.: Aspidium angustifrons MIQ. // Ann. Mus. Bot. Lugduno-Batavi 

3: 178 (1867) 

Ety.: angustus (L: narrow), frons (L: leaf, frond); relating to the frond 

habit 

Type: Buerger s. n. (L) // Japan 

TW: FOT 1994 (1994) 

(726) Thelypteris beddomei (BAKER) CHING 


Loc.: 縮羽金星蕨蕨 - SuoYuJinXingJue 

Bas.: Nephrodium beddomei BAKER // Syn. Fil. 267 (1867) 

Ety.: in honor of R. H. Beddome (British botanist, collector of the 


Type: Travancore, Beddome s. n. (K) // India 

TW: Hayata, B. 1909 (1909) 

(727) Thelypteris glanduligera (KUNZE) CHING 


Loc.: 密密腺金星蕨蕨 - MiXianJinXingJue

156 

Bas.: Aspidium glanduligerum KUNZE // Analecta Pteridogr. 44 

(1837) 

Ety.: glandula (L: gland), ger (L: carrying, bearing, used as suffix); 

relating to the abaxial surface of segments 

Type: Guangdong, Guangzhou, C. W. Peterson s. n. (HT: B, Herb. 

Lehmann) // China 


(728) Thelypteris gracilescens (BLUME) CHING 


Loc.: 光葉金星蕨蕨 - GuangYeJinXingJue 

Bas.: Aspidium gracilescens BLUME // Enum. Pl. Javae 155 (1828) 

Ety.: gracilescens (L: becoming slender); relating to the graceful 

lamina appearance 


TW: Henry, A. (1896) 

(729) Thelypteris japonica (BAKER) CHING 


Loc.: 栗柄金星蕨蕨 - LiBingJinXingJue 

Bas.: Nephrodium japonicum BAKER // Ann. Bot. (Oxford) 5: 318 

(1891) 


Type: Nagasaki, Capt. Blomfield s. n. (K) // Japan 

TW: Ogata, M. (1929) 

(730) Thelypteris laxa (FRANCH. & SAV.) CHING 


Loc.: 柔葉金星蕨蕨 - RouYeJinXingJue 

Bas.: Aspidium laxum FRANCH. & SAV. // Enum. Pl. Jap. 2: 237, 631- 

632 (1879) 

Ety.: laxa (L: spaced); relating to the prominent vertical distance 

between the lateral pinnae 

Type: Yokoska, Savatier s. n. (P) // Japan 

TW: Ito, H. 1944 (1944) 

(731) Thelypteris uraiensis (ROSENST.) CHING 


Loc.: 烏來金星蕨蕨 - WuLaiJinXingJue 

Bas.: Dryopteris uraiensis ROSENST. // Hedwigia 56: 341 (1915) 

Ety.: relates to WuLai ("Urai"), a place in Taiwan and the type 

location 

Type: Urai, Faurie 22, Jul-1914 (HT: S; IT: KYO, MICH, P) // Taiwan 

TW: Hedwigia 56: 341 (1915) 

TRICHOMANES (Hymenophyllaceae) 

Ety.: thrix (G: hair), manes (G: cup); relating to the hair-like receptacle 

protruding from the cup-shaped involucre

157 

(732) Trichomanes bimarginatum (BOSCH) BOSCH [IUCN: EN (?CR)] 

Pub.: Ned. Kruidk. Arch. 5(2): 143-144 (1860) 

Loc.: 叉脈單葉假脈蕨蕨 - ChaMaiDanYeJiaMaiJue 

Bas.: Microgonium bimarginatum BOSCH // Hymenophyll. Javan. 7 

(1860) 

Ety.: bi (L: two-), margo (L: border), atum (L: suffix, added to nouns 

to form adjectives meaning "provided with"); relating to the 

presence of an intramarginal false vein (a second margin) 

Alt.: Didymoglossum bimarginatum (BOSCH) EBIHARA & K. IWATS. // 

Blumea 51(2): 236 (2006) 

Type: Thwaites 2986 (HT: L; IT: B, BM, GH, K, L, P) // Sri Lanka 

TW: Hayata, B. 1917 (1917) 

(733) Trichomanes motleyi (BOSCH) BOSCH [IUCN: NT] 

Pub.: Ned. Kruidk. Arch. 5(2): 145 (1860) 

Loc.: 短柄單葉假脈蕨蕨 - DuanBingDanYeJiaMaiJue 

Bas.: Microgonium motleyi BOSCH // Hymenophyll. Javan. 5 (1860) 

Ety.: in honor of J. Motley (British botanist, collector of the type 

specimen) 

Alt.: Didymoglossum motleyi (BOSCH) EBIHARA & K. IWATS. // 

Blumea 51(2): 236 (2006) 

Type: Borneo, Motley 203 (HT: L; IT: GH, K) 

TW: Yabe, Y. (1902) 

(734) Trichomanes sublimbatum C. MÜLL. [IUCN: - (?CR)] 


Loc.: 亞緣單葉假脈蕨蕨 - YaYuanDanYeJiaMaiJue 

Ety.: sub (L: prefix, meaning under, below, near), limbatum (L: 

bordered); probably relating to the presence of an 

intramarginal vein 

Alt.: Didymoglossum sublimbatum (C. MÜLL.) EBIHARA & K. IWATS. // 

Type: 

Blumea 51(2): 236 (2006) 

Java, Zollinger Fil. Jav. 865, 1899 & 3500 (ST: K, fragment in 

US) // Indonesia 


(735) Trichomanes tahitense NADEAUD 

Pub.: Enum. Pl. Tahiti 18 (1873) 

Loc.: 盾型單葉假脈蕨蕨 - DunXingDanYeJiaMaiJue 

Ety.: relating to Tahiti (French Polynesia), the type location 

Alt.: Didymoglossum tahitense (NADEAUD) EBIHARA & K. IWATS. // 

Blumea 51(2): 236 (2006) 

Type: Nadeaud s. n. (HT: P) // French-Polynesia 

TW: Kuo, C. M. 1985 (1985)

158 

VITTARIA (incl. Haplopteris) (Vittariaceae) 

Ety.: vitta (L: ribbon, band), aris (L: resembling); relating to the shape of 

narrow fronds 

Haplopteris: haplous (G: simple, single), pteris (relates to genus 

Pteris, or ferns in general); relating to the leaves, which are narrow 

without reticulate veins 

(736) Vittaria anguste-elongata HAYATA 

Pub.: Icon. Pl. Formosan. 6: 161 (1916) 

Loc.: 姬書帶蕨蕨 - JiShuDaiJue 

Ety.: angustus (L: narrow), elongata (L: lengthened out, elongated); 

relating to the narrowly elongated fronds 

Alt.: Haplopteris anguste-elongata (HAYATA) E. H. CRANE // Syst. 

Bot. 22: 514 (1998) 

Type: Urai, Suisha, Kelung, Giran, Soseikyaku (ST: TI; IST: TAIF) // 



(737) Vittaria ensiformis SW. 

[IUCN: - (NT)] 

Pub.: Ges. Naturf. Freunde Berlin Neue Schriften 2: 134, pl. 7, f. 1 

(1799) 

Loc.: 劍葉書帶蕨蕨 - JianYeShuDaiJue 


pinna shape 

Alt.: Haplopteris ensiformis (SW.) E. H. CRANE // Syst. Bot. 22: 514 

(1998) 

Type: 

TW: 

Sonnerat s. n. (?K, ?S) // Mauritius 

a: FOJ (1995) as synonym of Vittaria anguste-elongata 

HAYATA 


(738) Vittaria flexuosa FEE 

Pub.: Mem. Foug. 3: 16 (1851) 

Loc.: 書帶蕨蕨 - ShuDaiJue 

Ety.: flexuosa (L: curved); relating to the frond habit 

Alt.: Haplopteris flexuosa (FEE) E. H. CRANE // Syst. Bot. 22: 514 

(1998) 

Type: Kumaon, Wallich 144 (B, BM) // India 

TW: Ito, H. 1944 (1944) 

(739) (Vittaria heterophylla comb. ined.) Haplopteris heterophylla C. W. 

CHEN, Y. H. CHANG & Y. C. LIU [IUCN: - (CR)] 

Pub.: Syst. Bot. 38(4): 901-905, f. 1-4, 5J (2013) 

Loc.: 異葉書帶蕨蕨 - YiYeShuDaiJue 

Ety.: heteros (G: different), phyllon (G: leaf); relating to the 

dimorphism of young sterile fronds and fertile fronds 

Alt.: Haplopteris heterophylla C. W. CHEN, Y. H. CHANG & Y. C. LIU 

// Syst. Bot. 38(4): 901-905, f. 1-4, 5J (2013)

159 

Type: Taipei, Junjianyan, 150 m alt., Wade 2147, 22-Mar-2012 (HT: 


TW: Syst. Bot. 38(4): 901-905, f. 1-4, 5J (2013) 

(740) Vittaria mediosora HAYATA [IUCN: VU (EN)] 

Pub.: Icon. Pl. Formosan. 5: 346-347, f. 149g-i (1915) 

Loc.: 細葉書帶蕨蕨 - XiYeShuDaiJue 

Ety.: medius (L: middle), sorus (L: sorus); relating to the position of 

soral lines on abaxial lamina 

Alt.: Haplopteris mediosora (HAYATA) X. C. ZHANG // Ann. Bot. Fenn. 

Type: 

40: 460 (2003) 

Mt. Arizan, Tozan ad 7800 ped. alt., S. Sasaki s. n., Mar-1913 

(HT: TI; IT: TAIF) // Taiwan 

TW: Icon. Pl. Formosan. 5: 346-347, f. 149g-i (1915) 

(741) Vittaria taeniophylla COPEL. 

Pub.: Philipp. J. Sci. 1 (Suppl. 2): 157 (1906) 

Loc.: 廣葉書帶蕨蕨 - GuangYeShuDaiJue 

Ety.: taenia (G: band, ribbon), phyllon (G: leaf); relating to the shape 

of lamina 

Alt.: Haplopteris taeniophylla (COPEL.) E. H. CRANE // Syst. Bot. 22: 

Type: 

514 (1998) 

Luzon, Province Benguet, Copeland 1936, 7-Nov-1905 (B, 

?BM, P) // Philippines 

TW: Ito, H. 1944 (1944) 

(742) Vittaria zosterifolia WILLD. 

Pub.: Sp. Pl. 6: 406 (1810) 

Loc.: 

Ety.: 

垂葉書帶蕨蕨 - ChuiYeShuDaiJue 

zostera relates to a flowering plant genus (Zosteraceae, 

"eelgrass"); relating to the shape of lamina 

Alt.: Haplopteris zosterifolia (WILLD.) E. H. CRANE // Syst. Bot. 22: 

514 (1998) 

Type: "in insulae Borboniae", B. de St. Vincent s. n. (B, BM, FI, P) // 

Reunion 

TW: Ito, H. 1944 (1944) 

WOODSIA (Dryopteridaceae) 

Ety.: in honour of J. Wood (British botanist) 

(743) Woodsia andersonii (BEDD.) CHRIST [IUCN: - (CR)] 


Loc.: 蜘蛛岩蕨蕨 - ZhiZhuYanJue 

Bas.: Gymnogramma andersonii BEDD. // Ferns Brit. India pl. 190 

(1866) 

Ety.: in honor of T. Anderson (British botanist, collector of the type 

specimen) 

Type: Kumaon, Soondadunga Valley, 13,000 ft., Anderson s. n. // 

India

160 


(744) Woodsia okamotoi TAGAWA [IUCN: CR (EN)] 


Loc.: 岡本氏岩蕨蕨 - GangBenShiYanJue 

Ety.: in honor of S. Okamoto (Japanese botanist, collector of the 


Type: Prov. Takao, Mt. Kwanzan, S. Okamoto s. n., 7-Oct-1937 (HT: 



(745) Woodsia polystichoides D. C. EATON 

Pub.: Proc. Amer. Acad. Arts 4: 110 (1858) 

Loc.: 岩蕨蕨 - YanJue 

Ety.: polystichum relates to a fern genus (see there), oides (L: 

suffix, denoting likeness of form); relates to the appearance 

of lamina 

Type: Hokkaido, C. Wright s. n. (NY) // Japan 

TW: Hayata, B. 1914-21 (1916) 

(746) Woodsia sp. 


Loc.: 王氏岩蕨蕨 - WangShiYanJue 


WOODWARDIA (Blechnaceae) 

Ety.: in honour of T. J. Woodward (British botanist) 

(747) Woodwardia harlandii HOOK. [IUCN: NT] 

Pub.: Fil. Exot. pl. 7 (1857) 

Loc.: 哈氏狗脊蕨蕨 - HaShiGouJiJue 

Ety.: in honor of W. A. Harland (British botanist, collector of the type 

specimen) 

Type: Guangdong, W. A. Harland 78 (E) // China 

TW: Hayata, B. 1914-21 (1915) 

(748) Woodwardia japonica (L. F.) J. SM. 

Pub.: Mem. Acad. Sci. Turin 5: 411 (1793) 

Loc.: 日本狗脊蕨蕨 - RiBenGouJiJue 

Bas.: Blechnum japonicum L. F. // Suppl. Pl. 445 (1781) 




(749) Woodwardia kempii COPEL. [IUCN: VU] 


Loc.: 細葉狗脊蕨蕨 - XiYeGouJiJue 

Ety.: in honor of E. Kemp (British botanist, and collector of the type 

specimen) 

Type: Guangdong, 31-Mar-1924 (PE) // China 

TW: FOT 1975 (1975)

(750) Woodwardia prolifera HOOK. & ARN. 

Pub.: Bot. Beechey Voy. 275, pl. 56 (1841) 

Loc.: 東方狗脊蕨蕨 - DongFangGouJiJue 


development of plantlets on the adaxial lamina 

Type: Okinawa, Loo choo, Lay s. n., Jun-1827 (E, K) // Japan 


Woodwardia radicans SM. 

B: Sasaki, S. (1928) 

(751) Woodwardia unigemmata (MAKINO) NAKAI 


Loc.: 生芽狗脊蕨蕨 - ShengYaGouJiJue 

Bas.: Woodwardia radicans var. unigemmata MAKINO // J. Jap. Bot. 

2: 7 (1918) 

Ety.: uni (L: one), gemmata (L: to put forth with buds); relates to the 

presence of a singular bud near the apex of rachis 

Type: Shizuoka, Fall Joren, K. Hisauchi, 11-Aug-1917 (MAK) // 

Japan 


161

162

163 

3 C O M M E N T S 

Comments regarding selected taxa of the list 

The following is a list of comments and supplementary information regarding 

taxa with relevance for Taiwan. Omitted is information directly related to the 

Flora of China which are provided in two lists hereafter. 

Athyrium delavayi 

var. delavayi 

Athyrium oppositipennum 

var. 

oppositipennum 

Athyrium oppositipennum 

var. 

pubescens 

Athyrium palustre 

Bolbitis heteroclita 

x Bolbitis subcordata 

Cornopteris 

decurrentialata 

var. decurrentialata 

Cyclosorus 

Based on ICN article 26.1 are autonyms not followed by 

an author citation, not as incorrectly provided in Knapp, 

R. 2011. 



R. 2011. 

In Knapp, R. 2011, Y. C. Liu is shown as author of this 

plant name, following a publication in Liu, Y. C. et al. 

2009: 36-38. However, Tagawa published this combination 

earlier, see Acta Phytotax. Geobot. 4: 142-143 

(1935), basing on Athyrium taiwanense var. pubescens 

Tagawa in Acta Phytotax. Geobot. 2: 18 (1933). 

Only known from the TaiBei area. This name replaces 

Athyrium sp. 2 as used in Knapp, R. 2011. Regarding 

the threat category according to IUCN, I follow the 

opinion of Chang, Y. H. et al. 2014. 

One parent of this hybrid was mistyped in Knapp, R. 

2013 (captions of fig. 4.21 and fig. 4.22), which is 

corrected here as Bolbitis heteroclita. 



R. 2013. 

Additional taxon C. ferox (Blume) Ching (Bull. Fan Mem. 

Inst. Biol., Bot. 8: 167 (1938); basionym: Aspidium ferox 

Blume // Enum. Pl. Javae 153 (1828); type: West Java, 

Blume 908 (L) // Indonesia), with unconfirmed status for 

Taiwan. This taxon is a large member of the subgenus 

Chingia. First published for Taiwan in Kuo, C. M. 2010 

from LanYu, it has been cultivated at the Taitung District 

Agricultural Research and Extension Station. One 

hypothesis is that plants developed from spores in the 

soil brought from LanYu to the research facility in 

TaiDong City. There have been no field reports yet for 

gametophytes or sporophytes from Taiwan.

164 

Diplazium 

kawakamii var. 

kawakamii 

Diplazium 

squamigerum 

Dryopteris 

Dryopteris 

wallichiana 

subsp. 

wallichiana 

Equisetum 

ramosissimum 

subsp. ramosissimum 

Grammitis 

intromissa 

Grammitis nuda 

Grammitis nutans 

Huperzia changii 



R. 2011. 

In Knapp, R. 2011, C. Hope is shown as author of this 

plant name. However, in J. Bombay Nat. Hist. Soc. 14: 

259, pl. 26 (1902), C. Hope described this taxon as 

Asplenium, and he treated Diplazium only as subgenus. 

Thus, it was J. Matsumura who validly published this 

name in Index Pl. Jap. 1: 304 (1904). 

A revision of Section Hirtipedes is needed for Taiwan 

and could result in additional taxa and confirm the status 

of others. 



R. 2013. 



R. 2011. 

The name used for this taxon is Radiogrammitis setigera 

(Blume) Parris according to recent phylogenetic 

systems, which is based on Polypodium setigerum 

Blume. It is earlier than Polypodium intromissum Christ, 

but the name could not be used in Grammitis because it 

was was predated by G. setigera J. Sm. (B. S. Parris, 

pers. comm. in 2014). 

Confirmed for one location in N Taiwan based on the 

outcome of herbaria revision. 

Hsu, T. C. et al. 2014 describe this epiphyte from warmtemperate 

rain forests at an elevation range of ca. 1200- 

1500 m, existing in only a few small populations in 

mountains of SE Taiwan. As IUCN threat category they 

propose EN, which I support. 

As genus Phlegmariurus is not accepted by many taxonomists 

(see e. g. Appendix B), I provide a new 

combination for this taxon in (the here broadly defined) 

genus Huperzia: 

Huperzia changii (T. Y. Hsieh) Ralf Knapp, comb. 

nov., basionym: Phlegmariurus changii T. Y. Hsieh in 

Amer. Fern J. 102(4): 284 (2012). 

Huperzia cunninghamioides 

This taxon was formerly known from Taiwan and Japan, 

but Kadoya, T. et al. (2014) consider it as being extinct 

now in Japan. In Taiwan, this taxon survives in very

165 

Isoetes 

taiwanensis var. 

taiwanensis 

Leucostegia 

truncata 

Nephrolepis x 

hippocrepicis 

Polypodium 

persicifolium 

Polystichum 

attenuatum 

Polystichum 

tenuius 

Pteris cretica 

subsp. cretica 

Pteris kawabatae 

scattered and small populations in N, C, E, SE and S 

parts. It is now presumed endemic to Taiwan. 



R. 2013. 

Fraser-Jenkins, C. R. 2008: 348-349 shows that this 

name is older and thus replacing L. immersa (Wall. ex 

Hook.) C. Presl, published in Tent. Pterid. 95, t. 4, f. 1 1 

(1836), based on Davallia immersa Wall. // Numer. List 

n. 256 (1828), ex Hook., Sp. Fil. 1: 156 (1846). 

Specimens of Wallich's collection 256 are in BM, K, L 

and P. 

Putative hybrid of N. biserrata and N. cordifolia. In 

Knapp, R. 2011 the epithet has been misspelt 

("hipocrepicis", see pages 292, 293, 467, 488, 811, 812, 

1002). This was caused by an incorrect spelling on the 

holotype and isotype and several paratype sheets (all in 

TNS). 

Known from a small population of a few plants only. 

IUCN category CR D was proposed by Chang, Y. H. et 

al. 2012. However, as the present status (including 

distribution and history of arrival in Taiwan) is not 

confirmed, I propose category DD. 

Only known from one small population from C Taiwan. 

Regarding the threat category according to IUCN, I 

follow the opinion of Chang, Y. H. et al. 2014. 

Currently only known from two locations of the limestone 

areas of HuaLian County. This name replaces 

Cyrtogonellum caducum and Polystichum sp. 2 as used 

in Knapp, R. 2013. 



R. 2011. 

Chao, Y. S. et al. 2013 report that this taxon is 

distributed at altitudes of less than 1000 m. In SC 

Taiwan I have found a small population at 1500 m 

altitude, whereas in N Taiwan it grows typically at an 

elevation range of ca. 300-700 m. 

In June of 2013 volumes 2 and 3 of the Flora of China (=FOC) were published 

(as a single book), which contain pteridophytes of Mainland China and Taiwan. 

There are several cases where information provided differs from my data. In 

the following tables I will focus on differences in the used species or infraspecific 

epithet, and on taxa reported with distribution in Taiwan but not 

recognized in the previous chapter in this document. Taxon names are given

166 

as used in FOC, and numbers after the taxon name refer to the relevant 

pages in FOC. 

Flora of China: additional taxa for Taiwan 

Taxa reported in FOC with occurrence in Taiwan but not accepted in local 

literature (i. e. Knapp, R. 2011 and Knapp, R. 2013): 

Adiantum 

erythrochlamys 

Diels 

Asplenium 

caucasicum 

(Fraser-Jenk. & 

Lovis) Viane 

Asplenium 

holosorum Christ 

248 • Taxon not recognized in Knapp, R. 2011 and 

Knapp, R. 2013 

• Taiwan included in FOC with uncertainty 

statement 

• Protologue: Bot. Jahrb. Syst. 29: 201 (1900) 

• Re-evaluation of populations of the similar A. 

monochlamys in C Taiwan (at higher elevations, 

above 2000 m) might result in a 

confirmation of this taxon for Taiwan 

• Not recognized by me for the time being for 





statement 

• Basionym: A. septentrionale subsp. caucasicum 

Fraser-Jenk. & Lovis // Notes Roy. Bot. 

Gard. Edinburgh 38: 281 (1980) 

• A. sasakii (Hayata) Makino & Nemoto // Acta 

Phytotax. Geobot. 10: 204-205 (1941) might 

be an earlier name for A. caucasicum if 

Taiwanese plants turn out to be diploid. 

However, at present, no chromosome counts 

or flow-cytometric data are available for 

Taiwanese plants (FOC). 





• Protologue: Bull. Herb. Boissier 7: 10 (1899) 

• In morphology similar to A. ensiforme; until 

now I have neither found field populations nor 

specimens collected in Taiwan that would 

allow recognition of a second taxon

167 

Asplenium 

quadrivalens (D. 

E. Meyer) 

Landolt 

Asplenium 

sublaserpitiifolium 

Ching 

Athyrium 

foliolosum T. 

Moore ex R. Sim 

Athyrium kuratae 

Seriz. 

Athyrium 

mengtzeense 

Hieron. 

Botrychium 

daucifolium Wall. 

ex Hook. & Grev. 



• Basionym: A. trichomanes subsp. quadrivalens 

D. E. Meyer // Ber. Deutsch. Bot. Ges. 74: 456 

(1962) 

• Due to its morphological similarity to A. 

trichomanes, cytological studies are necessary 





• Protologue: Notul. Syst. (Paris) 5: 146 (1936) 

• FOC lumps A. neolaserpitiifolium into A. 

pseudolaserpitiifolium. As type material and 

protologues of such highly dissected 

Aspleniums are of inadequate quality, it might 

be very difficult to successfully resolve 

taxonomy 


Taiwan; a monographic revision of tripinnate 

Asplenium is badly needed 



• Protologue: Priced Cat. Ferns 6: 22 (1859) 

• Not recognized in the monograph of Liu, Y. C. 

et al. 2009 



• Protologue: J. Jap. Bot. 45: 117 (1970) 

• Not recognized in the monograph of Liu, Y. C. 

et al. 2009 



• Protologue: Hedwigia 59: 319 (1918) 

• Considered as synonym of A. arisanense in 

FOT 1994 






statement 

• Protologue: Icon. Filic. 2: pl. 161 (1830) 

• No data available that proofs existence in

168 

Botrychium 

japonicum 

(Prantl) Underw. 

Cheilanthes 

opposita Kaulf. 

Cheiropleuria 

bicuspis (Blume) 

C. Presl 







statement 

• Basionym: B. daucifolium var. japonicum 

Prantl // Jahrb. Königl. Bot. Gart. Berlin 3: 340 

(1884) 

• At this point I assume that names B. 

daucifolium, B. japonicum and B. formosanum 

were all used locally to describe a single 

taxon, namely B. formosanum (Knapp, R. 

2011, Kuo, C. M. 1985, MAN) or B. 

daucifolium (FOT 1994 and CHK). 





• Protologue: Enum. Filic. 211 (1824) 

• Shaffer-Fehre in her Revised handbook to the 

Flora of Ceylon XV (B): 404-405 (2006) states 

that all records of C. opposita from most part 

of China, Taiwan, Japan and Philippines might 

be C. chusana. Kuo, C. M. 1985 explicitly 

rejected the presence of C. opposita in 

Taiwan. Study of material from Taiwan (field 

images, specimens) does not allow me 

recognizing two taxa. 






statement 

• Basionym: Polypodium bicuspe Blume // 

Enum. Pl. Javae 125 (1828) 

• I have no evidence for existence in Taiwan. 

Occasionally there are populations where 

some fronds develop forked sterile laminae - 

but not like in material I have seen from 

Malaysia (in SING). Therefore, I recognize 

only one taxon for Taiwan, namely C. 

integrifolia.

169 

Cyclosorus 

procurrens 

(Mett.) Copel. 

Cyclosorus 

siamensis 

(Tagawa & K. 

Iwats.) Panigrahi 

Cyclosorus 

subacutus Ching 

Cyrtomium 

atropunctatum 

Sa. Kurata 

Cyrtomium 

omeiense Ching 

& K. H. Shing 

378 • This taxon is not recognized in any recent 

literature for Taiwan (CHK, FOT 2003, Kuo, C. 

M. 1985, MAN, Knapp, R. 2011 and Knapp, R. 

2013) 

• Basionym: Aspidium procurrens Mett. // Ann. 

Mus. Bot. Lugduno-Batavi 1: 231 (1864) 



378 • Taxon not recognized in Knapp, R. 2011 

(comments provided there) and Knapp, R. 

2013 


statement 

• Basionym: Thelypteris siamensis Tagawa & K. 

Iwats. // Acta Phytotax. Geobot. 22: 101 (1967) 






2013) 


statement 

• Protologue: Fl. Fujian. 1: 598 (1982) 





• Protologue: Sci. Rep. Yokosuka City Mus. 8: 

46 (1964) 

• Only known from two specimens in HAST and 

TAIF revised by S. Matsumoto. Only NC 

Taiwan, ca. 2000 m altitude. 

• A revision of Taiwanese members of 

Cyrtomium is required. Until then I do not 

recognize this taxon for Taiwan. 



• Protologue: Acta Phytotax. Sin., Addit. 1: 36 

(1965) 

• Only known from few specimens in HAST 

revised by S. Matsumoto. Only C Taiwan, ca. 

1400-1700 m altitude. 

• A revision of Taiwanese members of

170 

Cyrtomium 

tukusicola 

Tagawa 

Cystopteris 

dickieana R. Sim 

Cystopteris 

montana (Lam.) 

Bernhardi ex 

Desv. 

Dennstaedtia 

scabra var. 

glabrescens 

(Ching) C. Chr. 





• Protologue: Acta Phytotax. Geobot. 7(2): 79 

(1938) 

• Only known from a few specimens in HAST 

and TAI revised by S. Matsumoto. Only C 

Taiwan, ca. 1800-2600 m altitude. 





Knapp, R. 2013 (comments provided there) 


statement 

• Protologue: Gard. Farmers’ J. 2: 308 (1848) 






statement 

• Basionym: Polypodium montanum Lam. // Fl. 

Franc. 1: 23 (1779) 

• I have no data for this taxon for Taiwan. This 

taxon is not recognized in any recent literature 

for Taiwan (CHK, FOT 2003, Kuo, C. M. 1985, 

MAN, Knapp, R. 2011 and Knapp, R. 2013). 





• Basionym: D. glabrescens Ching // Bull. Dept. 

Biol. Sun Yatsen Univ. 6: 24 (1933) 

• I have difficulties to conduct an infraspecific 

treatment for material in Taiwan. Local plants 

are mostly conspicuously hairy on both 

surfaces, but often not very densely so. All 

material I have seen so far seems to have 

(almost) glabrous indusia. In the broad sense, 

this species is widespread in Taiwan in an 

altitude range of ca. 400-3400 m. 

• No infra-specific taxonomy accepted for the 

time being for Taiwan

171 

Deparia conilii 

(Franch. & Sav.) 

M. Kato 

Deparia japonica 

(Thunb.) M. Kato 

Dicranopteris 

pedata (Houtt.) 

T. Nakaike 

Diplaziopsis 

brunoniana 

(Wall.) W. M. 

Chu 

Diplazium 

amamianum 

Tagawa 



• Basionym: Asplenium conilii Franch. & Sav. // 

Enum. Pl. Jap. 2: 227 (1877) 





• Basionym: Asplenium japonicum Thunb. in 

Murray // Syst. Veg. (ed. 14): 934 (1784) 



111 • FOC lumps all taxa of D. linearis s. l. from 

Taiwan into this name except for D. 

taiwanensis, which is not accepted by me 

• This taxon/name is not recognized in any 

recent literature for Taiwan (CHK, FOT 2003, 

Kuo, C. M. 1985, MAN, Knapp, R. 2011 and 

Knapp, R. 2013) 

• Basionym: Polypodium pedatum Houtt. // Nat. 

Hist. 14: 174 (1783) 



318 • Taxon as recognized in Knapp, R. 2011 and 

Knapp, R. 2013 corresponds to D. brunoniana 

in FOC 

• Basionym: Allantodia brunoniana Wall. // Pl. 

Asiat. Rar. 1: 44 (1830) 

• I agree with the majority of authors (i. e. 

Christenhusz et al. 2011: 48) and lump D. 

brunoniana into D. javanica. Differences as 

stated in FOC regarding morphological 

characters are variable, and not necessarily 

correlated. Study of Taiwan material showed 

that particularly lamina base width and pinna 

apex shape are not linked to each other. I do 

not know which voucher specimen of Taiwan 

("Gaoxiong") was consulted, but expect it to be 

within the morphological variation range of this 

taxon which is rather wide-spread in Taiwan. 



2013 

• Protologue: Acta Phytotax. Geobot. 3: 35 

(1934)

172 

Diplazium forrestii 

(Ching ex Z. R. 

Wang) Fraser- 

Jenk. 

Diplazium uraiense 

Rosenst. 

Dryopteris 

annamensis 

(Tagawa) L. B. 

Zhang 

Dryopteris 

decipiens var. 

diplazioides 

(Christ) Ching 

Dryopteris labordei 

(Christ) C. Chr. 

Dryopteris nodosa 

(C. Presl) L. B. 

Zhang 





2013 

• Basionym: Allantodia forrestii Ching ex Z. R. 

Wang // Acta Phytotax. Sin. 32: 82 (1994) 





2013 

• Protologue: Hedwigia 56: 336 (1915) 






2013) 

• Basionym: Diacalpe annamensis Tagawa // 

Acta Phytotax. Geobot. 14: 46 (1950) 





2013 

• Basionym: Polystichum diplazioides Christ // 

Bull. Acad. Int. Geogr. Bot. 1902: 260 (1902) 

• No infra-specific taxa recognized for the time 

being for Taiwan 



• Basionym: Aspidium labordei Christ // Bull. 

Soc. Bot. France, Mem. 1: 40 (1905) 

• Most collections from Taiwan determined as D. 

labordei are actually D. subtriangularis or D. 

tenuicula. Further studies are necessary prior 

to revising subgenus Erythrovariae section 

Erythrovariae. 





• Basionym: Acrophorus nodosus C. Presl //

173 

Haplopteris 

plurisulcata 

(Ching) X. C. 

Zhang 

Hymenasplenium 

obliquissimum 

(Hayata) 

Sugimoto 


obscurum 

(Blume) Tagawa 

Tent. Pterid. 94 (1836), based on Aspidium 

nodosum Blume, Enum. Pl. Javae 171 (1828) 

[not Willd. (1810)] 

• At present do all specimens and field 

observations support the acceptance of one 

taxon only, namely Acrophorus paleolatus, 

with mostly entirely glabrous rachis. 





• Basionym: Vittaria plurisulcata Ching // 

Sinensia 1: 186 (1931) 

• Probably a misidentification of V. flexuosa (or 

V. taeniophylla). My doubts (shared by C. W. 

Chen, pers. comm. in 2012) are included in 

FOC. 



• Basionym: Asplenium unilaterale var. obliquissimum 

Hayata // Icon. Pl. Formosan. 4: 230 

(1914) 

• Asplenium "filipes" in Taiwan requires a 

detailed study, including cytology (R. Viane, 

pers. comm. in 2012 and 2013), and at least 

two recognizable taxa are currently lumped 

into this name. Forms from a small area in the 

mountains of the TaiBei - YiLan border region 

appear to correspond to the morphology of A. 

obliquissimum. 

• This taxon is probably recognizable and 

present, e. g. in NE Taiwan. However, a 

revision of Taiwanese members of Hymenasplenium 

is certainly still necessary. 

313 • Taxon recognized in Knapp, R. 2011 and 

Knapp, R. 2013 as Asplenium obscurum is 

treated in FOC as H. pseudobscurum 


statement 

• Basionym: Asplenium obscurum Blume // 



Hymenasplenium is required. Until then I 

continue to recognize A. obscurum for Taiwan.

174 

Hymenophyllum 

wrightii Bosch 

Hypodematium 

taiwanense 

Ching ex K. H. 

Shing 

Lemmaphyllum 

diversum 

(Rosenst.) 

Tagawa 

Lemmaphyllum 

microphyllum 

var. obovatum 

(Harr.) C. Chr. 

Lepisorus 

scolopendrium 

(Buch.-Ham. ex 

Ching) Mehra & 

Bir 

106 • Excluded taxon in FOC 

• Taxon not recognized in Knapp, R. 2011 


2013; reference missing in FOC so it appears 

as if pteridologists in Taiwan still widely accept 

this taxon to occur locally, which is not the 

case 

• Protologue: Nederl. Kruidk. Arch. 4: 391 

(1859) 



• Protologue: Fl. Reipubl. Popularis Sin. 4(1): 

318 (1999) 

• H. taiwanense may fall within the range of 

variation of H. crenatum, which is rather 

widespread in Taiwan, but further study is 

needed to ascertain whether or not the two 

taxa are distinct 





• Basionym: Polypodium diversum Rosenst. // 

Hedwigia 56: 346 (1915) 

• The Lemmaphyllum taxon in Taiwan with 

round and discrete sori is accepted by most as 

L. rostratum, which is widely distributed in 

Taiwan, growing on trees and rocks at an 

altitude up to 2000 m (or even more) 





2013 

• Basionym: Drymoglossum carnosum var. 

obovatum Harr. // J. Linn. Soc., Bot. 16: 33 

(1877) 

• No infra-specific taxa recognized for the time 

being for Taiwan 




2013) 


statement

175 

Leptochilus 

ellipticus var. 

flexilobus (Christ) 

X. C. Zhang 

Leptochilus 

ellipticus var. 

pothifolius 

(Buch.-Ham. ex 

D. Don) X. C. 

Zhang 

Lycopodium 

complanatum L. 

• Basionym: L. excavatus var. scolopendrium 

Buch.-Ham. ex Ching // Bull. Fan Mem. Inst. 

Biol. 4: 69 (1933), based on Polypodium 

scolopendrium Buchanan-Hamilton ex D. Don 

// Prodr. Fl. Nepal. 1 (1825), not P. 

scolopendria N. L. Burman // Fl. Indica 232 

(1768) 






2013) 

• Basionym: Polypodium flexilobum Christ // 

Bull. Acad. Int. Geogr. Bot. 1904: 107 (1904) 

• No infraspecific splitting in Knapp, R. 2011 and 

by most other local authors. This taxon is 

variable, and morphology changes with slight 

habitat changes (sunny/shaded, terrestrial/on 

rock, moist/dry) within a population at the 

same spot. No clear cut-lines in morphology 

observed in Taiwan. 





2013 

• Basionym: Hemionitis pothifolia Buch.-Ham. ex 

D. Don // Prodr. Fl. Nepal. 13 (1825) 





• FOC treats Diphasiastrum wilceae provisionally 

as synonym, and includes Taiwan in its 

distribution 

• Protologue: Sp. Pl. 2: 1104 (1753) 

• Comparing with L. complanatum from several 

sites in Germany, Taiwanese material is not 

conspecific. Material of Taiwan can be 

separated into two taxa (data e. g. in Kuo, C. 

M. 1985 or Knapp, R. 2011): 

(1) L. multispicatum (a name as well accepted 

in other countries such as Thailand): common, 

often in grass on steep slopes near forest

176 

Lygodium 

longifolium 

(Willd.) Sw. 

Lygodium 

salicifolium C. 

Presl 

Macrothelypteris 

oligophlebia var. 

elegans (Koidz.) 

Ching 


setigera (Blume) 

Ching 

Microlepia hancei 

Prantl 

edges. Altitude: ca. 1600-2100 m (the range is 

certainly wider, but I have no particular data on 

hand). 

(2) L. yueshanense: rather rare, subalpine 

bamboo grassland. Generally at higher altitude 

as L. multispicatum: ca. 2700-3200 m. 




2013) 

• Basionym: Hydroglossum longifolium Willd. // 

Abh. Kurfüstl.- Mainz. Akad. Nützl. Wiss. Erfurt 

2(4): 22 (1802) 






2013) 

• Protologue: Suppl. Tent. Pterid. 102 (1845) 






2013) 

• Basionym: Dryopteris elegans Koidz. // Bot. 

Mag. (Tokyo) 38: 108 (1924) 



340 • Only reported from Taiwan in CHK, not 

recognized in FOT 2003, Kuo, C. M. 1985, 

MAN, Knapp, R. 2011 (see comments there) 

and Knapp, R. 2013 

• Basionym: Cheilanthes setigera Blume // 


• Presumably based on identifications of S. G. 

Lu (in CHK) 





2013 

• Protologue: Arbeiten Königl. Bot. Gart. Breslau 

1: 35 (1892)

177 

Microlepia 

marginata var. 

villosa (C. Presl) 

Y. C. Wu 

Microlepia 

trapeziformis 

(Roxb.) Kuhn 

Osmunda 

angustifolia 

Ching 

Parathelypteris 

caoshanensis 

Ching ex K. H. 

Shing 


castanea 

(Tagawa) Ching 






2013) 

• Basionym: M. villosa C. Presl // Epimel. Bot. 

95 (1851) 

• In Taiwan, many transitions for hairiness 

towards typical M. marginata (var. marginata) 

exist. I therefore follow the opinion of M. G. 

Gilbert who is of the opinion that M. marginata 

var. villosa is only arbitrarily distinguished from 

M. marginata var. marginata and that it would 

be better to merge these two varieties (see 

FOC 161). 





• Basionym: Davallia trapeziformis Roxb. // 

Calcutta J. Nat. Hist. 4: 516 (1844) 






2013) 


statement 

• Protologue: Acta Phytotax. Sin. 8: 160 (1959) 






2013) 

• Protologue: Fl. Reipubl. Popularis Sin. 4(1): 

50, 321 (1999) 





• Basionym: Dryopteris castanea Tagawa // Acta 

Phytotax. Geobot. 4: 132 (1935)

178 


cystopteroides 

(D. C. Eaton) 

Ching 


grammitoides 


Phlegmariurus 

taiwanensis (C. 

M. Kuo) L. B. 

Zhang 

Polypodiodes 

chinensis (Christ) 

S. G. Lu 

Polystichum 

sozanense Ching 

ex H. S. Kung & 

• This name was used by some authors to 

describe Thelypteris japonica. There is no data 

to support the existence of both, T. japonica 

and T. castanea (if distinct at all) in Taiwan. 






2013) 

• Basionym: Athyrium cystopteroides D. C. 

Eaton // Proc. Amer. Acad. Arts 4: 110 (1858) 






2013) 

• Basionym: Aspidium grammitoides Christ // 

Bull. Herb. Boissier 6: 193 (1898) 





2013 

• Basionym: Lycopodium taiwanense C. M. Kuo 

// Taiwania 30: 51 (1985) 

• Several local botanists (including me) did not 

succeed to separate this taxon from Huperzia 

cryptomerina. Hence, I only accept one taxon, 

namely Huperzia cryptomerina. 



• Basionym: Polypodium subamoenum var. 

chinense Christ // Nuovo Giorn. Bot. Ital., n. s., 

4: 99 (1897) 

• I agree with Fraser-Jenkins, C. R. 2008: 43 

where he suggested that this taxon merely 

represents more hairy populations of P. 

amoena not worthy of formal recognition. 





• Protologue: Acta Phytotax. Sin. 33: 309 (1995)

179 

L. B. Zhang • Morphology of this taxon is very similar to P. 

parvipinnulum. Further study, particularly of 

populations in N Taiwan (type location!) is 

necessary. 

• Not recognized for the time being 

Pseudophegopteris 

pyrrhorhachis 

(Kunze) Ching 

Pteris 

confertinervia 

Ching 

Pteris taiwanensis 

Ching 

Pyrrosia 

adnascens (Sw.) 

Ching 

Salvinia molesta D. 

S. Mitch. 

Selaginella 

uncinata (Desv. 

ex Poir.) Spring 

348 • Taxon not recognized in Knapp, R. 2011 (see 

comments there) and Knapp, R. 2013 

• Basionym: Polypodium pyrrhorhachis Kunze // 

Linnaea 24: 257 (1851) 


Pseudophegopteris is required. Until then I do 

not recognize this taxon in addition to P. 

hirtirachis. 



• Protologue: Acta Bot. Austro Sin. 1: 4 (1983) 

• Fraser-Jenkins, C. R. 2008: 101 regarded P. 

confertinervia as a synonym of P. cretica 

subsp. cretica 





• Protologue: Acta Bot. Austro Sin. 1: 16 (1983) 

• No information available, considered as synonym 

of P. semipinnata in MAN 



789 • Basionym: Polypodium adnascens Sw. // Syn. 

Fil. 25, 222 (1806) 

• In Taiwan, this taxon is included in a broadly 

defined P. lanceolata. Based on the key in 

FOC is material from Taiwan matching P. 

adnascens, though the name P. lanceolata is 

used in MAN and Knapp, R. 2011. In any case 

there is only one taxon present in Taiwan. 

• Name not recognized by me for the time being 

for Taiwan 

125 • Taxon only recognized in Knapp, R. 2011 


2013 as naturalized species; this is 

insufficiently reflected in FOC 

46 • Taxon only recognized in Knapp, R. 2011 


2013 as naturalized species; this is

180 

Selliguea 

pellucidifolia 

(Hayata) S. G. 

Lu, Hovenkamp 

& M. G. Gilbert 

Tectaria rockii C. 

Chr. 

Tectaria subpedata 

(Harr.) Ching 

Woodwardia 

orientalis Sw. 

insufficiently reflected in FOC 




2013) 

• Basionym: Polypodium pellucidifolium Hayata 

// Icon. Pl. Formosan. 4: 250 (1914) 

• Not recognized for the time being, as 

differences to S. engleri are unclear 



• Protologue: Contr. U. S. Natl. Herb. 26: 331 

(1931) 






2013) 

• Basionym: Nephrodium subpedatum Harr. // J. 

Linn. Soc., Bot. 16: 30 (1877) 

• The type of T. subpedata from Taiwan is 

confirmed to be a young frond of T. 

polymorpha. The similar T. morsei (Baker) P. 

J. Edwards ex S. Y. Dong (based on 

Nephrodium morsei Baker in Bull. Misc. 

Inform. Kew 1906: 11 (1906) with type from 

China) represents a separate species and is 

not present in Taiwan, see Systematic Botany 

35(2): 235-243 (2010). 

415 • Taiwan included in FOC with uncertainty 

statement 

• Protologue: J. Bot. (Schrader) 1800(2): 76 

(1801) 

• Recent literature for Taiwan accept all only 

one taxon (W. prolifera (FOT 2003, Kuo, C. M. 

1985, MAN, Knapp, R. 2011 and Knapp, R. 

2013, except for CHK: W. orientalis var. 

formosana) 


Taiwan

181 

Flora of China: name changes for taxa of Taiwan 

Taxa recognized for Taiwan in FOC but with change in (infra-) specific epithet 

compared with local literature (i. e. Knapp, R. 2011 and Knapp, R. 2013): 

Alsophila 

denticulata Baker 

Angiopteris evecta 

(Forst.) Hoffm. 

Arachniodes 

amabilis (Blume) 

Tindale 

Arachniodes 

speciosa (D. 

Don) Ching 

Araiostegia 

perdurans 

(Christ) Copel. 

Asplenium 

exiguum Bedd. 

Asplenium indicum 

Sledge 

137 • Name in Knapp, R. 2011: Cyathea hancockii 

• Protologue: J. Bot. 23: 102 (1885) 

• This epithet is necessary if combined in genus 

Alsophila 

86 • Name in Knapp, R. 2011: A. palmiformis 

• Corresponding basionym: Polypodium evectum 

G. Forster // Fl. Ins. Austr. 81 (1786) 

• Further study is necessary prior to adopting 

change 

554 • Name in Knapp, R. 2011: A. yakusimensis 

• Corresponding basionym: Aspidium amabile 

Blume // Enum. Pl. Javae 165 (1828) 


change, and see also comment in Knapp, R. 

2011 

553 • Name in Knapp, R. 2011: A. pseudoaristata 

• Corresponding basionym: Aspidium speciosum 



change 

752 • Name in Knapp, R. 2011: A. parvipinnula 

• Corresponding basionym: Davallia perdurans 

Christ // Bull. Herb. Boissier 6: 970 (1898) 

• See comments in Knapp, R. 2013. Further 

study is necessary prior to adopting change. 

293 • Name in Knapp, R. 2011: A. yunnanensis 

• Protologue: Ferns South. India pl. 146 (1864) 

• Fraser-Jenkins combined A. yunnanense as 

subspecies into A. exiguum in Indian Fern J. 

27: 178-215 (2011). Until now I have no 

evidence that the typical A. exiguum (subsp. 

exiguum) is present in Taiwan. 

• Name not recognized by me for the time being 


286 • Name in Knapp, R. 2011: A. yoshinagae 

• Protologue: Bull. Brit. Mus. (Nat. Hist.) Bot. 3: 

264 (1965) 


change

182 

Asplenium 

komarovii 

Akasawa 

Asplenium 

matsumurae 

Christ ex 

Matsum. 

Athyrium clivicola 

Tagawa 

Athyrium 

drepanopterum 

(Kunze) A. Br. ex 

Milde 

Athyrium nigripes 

(Blume) T. Moore 

Calymmodon 

ordinatus Copel. 

Cephalomanes 

javanicum 

(Blume) C. Presl 

Cibotium cumingii 

Kunze 

Coniogramme 

intermedia var. 

glabra Ching 

274 • Name in Knapp, R. 2011: A. scolopendrium 

• Protologue: Bull. Kochi Wom. Univ. 10: 26 

(1962) 


change 

283 • Name in Knapp, R. 2011: A. serricula 

• Protologue: Bot. Mag. (Tokyo) 24: 241 (1910) 


change 

481 • Name in Knapp, R. 2011: A. auriculatum 


(1934) 

• No change necessary based on monograph by 

Liu, Y. C. et al. 2009 

463 • Name in Knapp, R. 2011: A. mupinense 

• Corresponding basionym: Polypodium drepanopterum 

Kunze // Linnaea 23: 278, 318 

(1850) 


Liu, Y. C. et al. 2009 

493 • Name in Knapp, R. 2011: A. tozanense 

• Corresponding basionym: Aspidium nigripes 



Liu, Y. C. et al. 2009 

844 • Name in Knapp, R. 2011: C. cucullatus 

• Protologue: Philipp. J. Sci. 34: 259-269, t. 1-6 

(1927) 


change 

95 • Name in Knapp, R. 2011: C. javanicum var. 

asplenioides 

• Corresponding basionym: Trichomanes javanicum 



change 

133 • Name in Knapp, R. 2011: C. taiwanense 

• Protologue: Farrnkräuter 1: 64 (1841) 


change 

177 • Name in Knapp, R. 2011: no recognition of 

infraspecific taxa 

• Protologue: Icon. Filic. Sin. 3: t. 143 (1935) 

• Further study is necessary prior to adopting

183 

Cyrtomium 

yamamotoi 

Tagawa 

Davallia divaricata 

Blume 

Deparia boryana 

(Willd.) M. Kato 

Deparia 

jiulungensis var. 

albosquamata 

(M. Kato) Z. R. 

Wang 

Diplaziopsis 

brunoniana 

(Wall. ex Bedd.) 

W. M. Chu 

Diplazium asperum 

Blume 

Diplazium 

kappanense 

Hayata 

Diplazium petrii 

Tardieu 

change or recognizing infra-specific taxa 

569 • Name in Knapp, R. 2013: C. macrophyllum 

var. simadae (synonymy of name and distribution 

with uncertainty statement) 


(1938) 


change 

753 • Name in Knapp, R. 2011: D. formosana 

• Protologue: Enum. Pl. Javae 237 (1828) 


change 

425 • Name in Knapp, R. 2011: D. subfluvialis 

• Corresponding basionym: Aspidium boryanum 

Willd. // Sp. Pl. 5: 285 (1810) 


change 

434 • In Knapp, R. 2011 probably corresponding to: 

D. sp. 1 (uncertainty expressed whether conspecific 

with D. orientalis (Z. R. Wang & J. J. 

Chien) T. Nakaike) 

• Corresponding basionym: D. pycnosora var. 

albosquamata M. Kato // J. Fac. Sci. Univ. 

Tokyo, Sect. 3, Bot. 13(4): 396 (1984) 

• A revision of Taiwanese members of Deparia 

is required prior to adopting change 

318 • Name in Knapp, R. 2011: no splitting into two 

taxa, only name D. javanica used 

• Corresponding basionym: Allantodia brunoniana 

Wall. // Pl. Asiat. Rar. 1: 44-45 (1830) 


change 

528 • Name in Knapp, R. 2013: D. sikkimense 

• Protologue: Enum. Pl. Javae 195 (1828) 

• See data in Knapp, R. 2013, no change 

necessary 

531 • Name in Knapp, R. 2011: D. taiwanense 

• Protologue: Icon. Pl. Formosan. 8: 143, f. 69- 

70 (1919) 


change 

526 • Name in Knapp, R. 2011: D. latifrons 

• Protologue: Asplen. Tonkin 67, 181, t. 9, f. 1-2 

(1932)

184 

Diplazium 

viridissimum 

Christ 

Dryopteris x 

holttumii L. B. 

Zhang 

Dryopteris 

pseudocaenopteris 

(Kunze) L. 

B. Zhang 

Dryopteris 

wuzhaohongii L. 

B. Zhang 

Dryopteris 

zhuweimingii L. 

B. Zhang 

Haplopteris 

elongata (Sw.) E. 

H. Crane 


change 

532 • Name in Knapp, R. 2011: D. pseudodoederleinii 

• Protologue: Notul Syst. (Paris) 1: 45 (1909) 


change, see also comments in Knapp, R. 2011 

616 • Name in Knapp, R. 2011: Dryopsis x fauriei 

• Protologue: Flora of China, Vol. 2-3 (Pteridophytes) 

616 (2013) 

• This name replaces Dryopsis x fauriei Holtt. & 

P. J. Edwards // Kew Bull. 41: 198 (1986) and 

its synonym Dryopteris x fauriei (Holtt. & P. J. 

Edwards) L. B. Zhang // Phytotaxa 71: 21 

(2012), not D. fauriei Kodama // Icon. Pl. 

Koisikav. 2: 11 (1914) 

• No change necessary when using the 

taxonomic system of Kramer & Green 

624 • Name in Knapp, R. 2011: Peranema 

aspidioides 

• Corresponding basionym: Diacalpe pseudocaenopteris 

Kunze // Bot. Zeitung (Berlin) 4: 

457 (1846) 



622 • Name in Knapp, R. 2011: Acrophorus 

macrocarpus 

• Protologue: Taxon 61(6): 1208 (2012) 



627 • Name in Knapp, R. 2011: Peranema 

cyatheoides 

• Protologue: Taxon 61: 1211 (2012) 

• In FOC Peranema cyatheoides is split into two 

taxa: D. peranema (with Peranema 

cyatheoides as synonym, reported not to be 

present in Taiwan) and D. zhuweimingii 

(Taiwan included in distribution list). 


change 

255 • Name in Knapp, R. 2011: Vittaria zosterifolia 

• Corresponding basionym: Vittaria elongata 

Sw. // Syn. Fil. 109: 302 (1806) 

• Further study is necessary prior to adopting

185 


adiantifrons 

(Hayata) Viane & 

S. Y. Dong 


murakamihatanakae 

T. 

Nakaike 


pseudobscurum 

Viane 

Hymenophyllum 

riukiuense Christ 

Lepisorus albertii 

(Regel) Ching 

Loxogramme 

duclouxii Christ 

Lycopodium 

verticale L. B. 

change 

312 • Name in Knapp, R. 2011: Asplenium filipes (in 

part) 

• Corresponding basionym: Asplenium resectum 

f. adiantifrons Hayata // Icon. Pl. Formosan. 4: 

226, f. 155 (1914) 


change 

313 • Name in Knapp, R. 2011 and Knapp, R. 2013: 

Asplenium cataractarum 

• Protologue: New Fl. Jap. Pterid. 841 (1992) 


change 

315 • Name in Knapp, R. 2011: Asplenium obscurum 

• Protologue: Flora of China, Vol. 2-3 

(Pteridophytes) 315 (2013) 

• Type: Hainan, Ledong County (“Kan-en 

District”), Jianfeng Ling (“Chim Fung Ling”), 

“near Sam Mo Watt village, Shan Mong,” S.-K. 

Lau 3841, 23-Apr-1934 (HT: PE) // China 


Hymenasplenium is required. Until then I 

continue to recognize A. obscurum only for 

Taiwan. 

103 • Name in Knapp, R. 2011: H. taiwanense 

(synonymy of name and distribution with 

uncertainty statement) 

• Protologue: Annuaire Conserv. Jard. Bot. 

Geneve 4: 208 (1900) 


change 

822 • Name in Knapp, R. 2011: L. clathratus 

• Corresponding basionym: Polypodium albertii 

Regel // Acta Hort. Petrop. 7: 622 (1881) 


change 

764 • Name in Knapp, R. 2011: L. remotefrondigera 

• Protologue: Bull. Acad. Int. Geogr. Bot. 17: 

140 (1907) [non Polypodium duclouxii Christ 

(1909)] 

• Lumping is incorrect (see comment in Knapp, 

R. 2011), no change necessary 

29 • Name in Knapp, R. 2011: L. juniperoideum 

• Protologue: Flora of China, Vol. 2-3 (Pterido-

186 

Zhang phytes) 29 (2013) 

• Type: Sichuan, Leibo County, Xining, 

Wayaoping, Xian-Xu Kong (H. S. Kung) 5642, 

3-Aug-1978 (HT: CDBI-302; IT: CDBI-303) // 

China 


change 

Neolepisorus 

fortunei (T. 

Moore) Li Wang 

Onychium 

cryptogrammoides 

Christ 

Paesia taiwanensis 

W. C. Shieh 

Pleocnemia 

leuzeana (Gaud.) 

C. Presl 

Pleocnemia winitii 

Holtt. 

Polypodioides 

niponica (Mett.) 

Ching 

Polystichum 

hookerianum (C. 

Presl) C. Chr. 

807 • Name in Knapp, R. 2011: Microsorum henryi 

• Corresponding basionym: Drynaria fortunei T. 

Moore // Gard. Chron. 1855: 708 (1855) 


change, see also comment in Knapp, R. 2011 

214 • Name in Knapp, R. 2011: O. lucidum 

• Protologue: Notul. Syst. (Paris) 1: 52 (1909) 



151 • Name in Knapp, R. 2011: P. radula 

• Protologue: J. Jap. Bot. 45: 161 (1970) 


change 

731 • Name in Knapp, R. 2011: P. rufinervis 

• Corresponding basionym: Polypodium leuzeanum 

Gaud. // Tent. Pterid. 184 (1836) 

• Further study including a revision on genus 

level is necessary prior to adopting change 

731 • Name in Knapp, R. 2011: P. submembranacea 

• Protologue: Reinwardtia 1: 181 (1951) 

• Further study including a revision on genus 

level is necessary prior to adopting change 

801 • Name in Knapp, R. 2011: Polypodium transpianense 

• Corresponding basionym: Polypodium niponicum 

Mett. // Ann. Mus. Bot. Lugduno-Batavi 2: 

222 (1866) 



690 • Name in Knapp, R. 2011: P. integripinnum 

• Corresponding basionym: Lastrea hookeriana 

C. Presl // Tent. Pterid. 77 (1836), nom. nov. 

for Aspidium caducum Wall. 

• Further study as to whether lumping of P. 

integripinnum into P. hookerianum is correct is 

necessary prior to adopting change

187 

Polystichum 

pseudodeltodon 

Tagawa 

Pronephrium 

cuspidatum 

(Blume) Holtt. 

Pteris scabristipes 

Tagawa 

Pteris venusta 

Kunze 

Pyrrosia davidii 

(Baker) Ching 

Radiogrammitis 

moorei Parris & 



setigera (Blume) 

Parris 


taiwanensis 

702 • Name in Knapp, R. 2011: P. deltodon 


(1937) 


change 

391 • Name in Knapp, R. 2011: Cyclosorus 

liukiuensis 

• Corresponding basionym: Meniscium cuspidatum 



necessary 

206 • Name in Knapp, R. 2011: P. aspericaulis 


(1936) 



192 • Name in Knapp, R. 2011: P. pellucida 

• Protologue: Bot. Zeitung (Berlin) 6: 195 (1868) 



795 • Name in Knapp, R. 2011: P. matsudae 

• Corresponding basionym: Polypodium davidii 

Baker // Ann. Bot. (Oxford) 5: 472 (1891) [non 

Franch. (1887)] 


change 


Grammitis fenicis 


842 (2013) 

• B. S. Parris pointed out that material from 

Taiwan is not conspecific with Grammitis 

fenicis from the Philippines. Therefore, a 

change of name is necessary and will be 

conducted when the situation for Grammitis 

jagoriana (see Radiogrammitis taiwanensis 

below) has been clarified. 

843 • Name in Knapp, R. 2011: Grammitis intromissa 

• Corresponding basionym: Polypodium setigerum 


• This epithet is necessary if combined in genus 



Grammitis jagoriana

188 

Parris & Ralf 

Knapp 

Selaginella 

leptophylla Baker 

Selliguea 

rhynchophylla 

(Hook.) Fraser- 

Jenk. 

Selliguea taeniata 

(Sw.) Parris 

Struthiopteris 

eburnea var. 

obtusa (Tagawa) 

Tagawa 

Themelium 

blechnifrons 

(Hayata) Parris 

Vandenboschia 

striata (D. Don) 

Ebihara 


843 (2013) 

• A more detailed study of this and allied taxa is 

necessary prior to adopting change as the 

name Grammitis jagoriana is in Taiwan applied 

to several recognizable taxa 

57 • Name in Knapp, R. 2013: S. aristata following 

Chang, H. M. et al. 2012 

• Protologue: J. Bot. 23: 157 (1885) 

• No change necessary, see Knapp, R. 2013 

775 • Name in Knapp, R. 2011: S. okamotoi 

• Corresponding basionym: Polypodium rhynchophyllum 

Hook. // Hooker's Icon. Pl. 10: pl. 

954 (1854) 


necessary as lumping is incorrect 

780 • Name in Knapp, R. 2013: S. falcatopinnata 

• Corresponding basionym: Polypodium taeniatum 

Sw. // J. Bot. (Schrader) 1800(2): 26 

(1801) 

• No change necessary, see Knapp, R. 2013 

413 • Name in Knapp, R. 2011: Blechnum 

eburneum, no recognition of infraspecific taxa 

• Corresponding basionym: Spicantopsis eburnea 

var. obtusa Tagawa // Acta Phytotax. 

Geobot. 9: 88 (1940) 


change 

849 • Name in Knapp, R. 2011: Grammitis 

(Ctenopteris) curtisii 

• Corresponding basionym: Polypodium decrescens 

var. blechnifrons Hayata // Icon. Pl. 

Formosan. 4: 245, f. 171 (1914) 

• Epithet change probably necessary 

109 • Name in Knapp, R. 2011: Vandenboschia 

birmanica 

• Corresponding basionym: Trichomanes striatum 


• With reference to Knapp, R. 2011: 442 a 

change of name appears necessary

189 

4 S Y N O N Y M S A N D M I S A P P L I E D N A M E S 

Following is a collection of scientific names applied to Taiwanese 

pteridophytes as used in various literatures. The names provided to the left of 

the equals sign (=) are either considered as synonyms, or as names in 

alternative combinations, or wrongly applied to taxa in Taiwan. The names to 

the right of the equals sign correspond to those listed in chapter 2 List. 

For better traceability references are provided for that literature, in which the 

name to the left of the equals sign was lumped into that provided after it. 

Example: 

Abacopteris =Cyclosorus simplex MAN 

simplex (Hook.) 

Ching 

The name to the left of the equals sign was placed into 

synonymy with the name to the right in MAN. 

In some cases was the lumping the result of two consequent steps, in which 

case both literature references are given. 

Example: 

Davallia multiflora 

Roxb. 

=Nephrolepis brownii MAN/Knapp, R. 

2011 

The name Davallia multiflora was considered a 

synonym of Nephrolepis multiflora in MAN. Afterwards, in 

Knapp, R. 2011 Nephrolepis multiflora was treated as synonym 

of Nephrolepis brownii. 

Please note that the listed literature reference might not be the first instance - 

even in local literature - where the respective lumping was done. 

Abacopteris aspera auct. non =Cyclosorus 

FOT 2003/MAN 

(C. Presl) Ching 

gymnopteridifrons 

Abacopteris cuspidata auct. 

non (Blume) Ching: MAN 

=Cyclosorus liukiuensis MAN/Knapp, R. 

2011 

Abacopteris gymnopteridifrons =Cyclosorus 

MAN 

(Hayata) Ching 


Abacopteris insularis K. Iwats. =Cyclosorus insularis MAN 

Abacopteris liukiuensis (Christ) 

Tagawa 

=Cyclosorus liukiuensis FOJ/Knapp, R. 

2011 

Abacopteris longipetiolata K. 

Iwats. (type: Taiwan) 

=Cyclosorus 

longipetiolatus 

MAN

190 

Abacopteris prolifera (Retz.) 

Shieh 

Abacopteris simplex (Hook.) 

Ching 

Abacopteris triphylla var. 

parishii (Bedd.) Ching 

Abacopteris triphylla (Sw.) 

Ching (var. triphylla) 

Abacopteris urophylla auct. 

non (Wall. ex Hook.) Ching 

Abrodictyum cumingii C. Presl 

Acrophorus hookeri T. Moore 

Acrophorus immersus (C. 

Presl) T. Moore 

Acrophorus stipellatus T. 

Moore (nom. nud.) 

Acrophorus stipellatus var. 

macrostegius Tagawa (type: 


Acrorumohra diffracta (Baker) 

H. Ito 

Acrorumohra hasseltii (Blume) 

Ching 

Acrorumohra subreflexipinna 

(Ogata) H. Ito 

Acrorumohra yoroii (Seriz.) W. 

C. Shieh 

Acrostichum angulatum Blume 

Acrostichum appendiculatum 

Willd. 

Acrostichium bicuspe var. 

integrifolia D. C. Eaton ex 

Hook. 

Acrostichum callifolium Blume 

Acrostichum calomelanos L. 

Acrostichum commutatum 

Mett. ex Kuhn 

=Cyclosorus proliferus 

=Cyclosorus simplex 

FOT 2003/MAN 

MAN 

=Cyclosorus parishii MAN/Knapp, R. 

2011 

=Cyclosorus triphyllus MAN 

=Cyclosorus 


=Cephalomanes 

cumingii 

=Araiostegia 

parvipinnula 

=Leucostegia truncata 

=Acrophorus paleolatus 

=Acrophorus 

macrocarpus 

=Dryopteris diffracta 

=Dryopteris hasseltii 

=Dryopteris 

subreflexipinna 

=Dryopteris yoroii 

=Elaphoglossum 

angulatum 

=Bolbitis appendiculata 

=Cheiropleuria 

integrifolia 


callifolium 

=Pityrogramma 

calomelanos 


commutatum 

Acta Phytotax. 

Geobot. 18(1): 11 

(1959)/ Knapp, R. 

2011 

MAN 

MAN 

MAN/Fraser- 

Jenkins, C. R. 

2008 

MAN 

Knapp, R. 2011; 

published in: Acta 

Phytotax. Geobot. 

8: 230 (1939) 

MAN 

MAN 

MAN 

MAN 

MAN 

MAN 

MAN 

MAN 

MAN 

MAN

Acrostichum conforme Sw. =Elaphoglossum FOT 2003/MAN 

marginatum 

Acrostichum contaminans Wall. =Bolbitis angustipinna FOT 2003/MAN 

(nom. nud.) 

Acrostichum dichotomum L. =Schizaea dichotoma MAN 

Acrostichum digitatum L. =Schizaea digitata MAN 

Acrostichum harlandii Hook. =Tectaria harlandii MAN 

Acrostichum heteroclitum C. =Bolbitis heteroclita MAN 

Presl 

Acrostichum lanceolatum L. =Pyrrosia lanceolata MAN 

Acrostichum lingua Thunb. =Pyrrosia lingua MAN 

Acrostichum marginatum Wall. =Elaphoglossum MAN 

ex Fee 

marginatum 

Acrostichum punctatum L. =Microsorum 

MAN 

punctatum 

Acrostichum septentrionale L. =Asplenium 

MAN 

septentrionale 

Acrostichum spectabile =Lomariopsis 

MAN 

(Kunze) Racib. 

spectabilis 

Acrostichum thalictroides L. =Ceratopteris 

MAN 

thalictroides 

Acrostichum virens auct. non =Bolbitis subcordata FOT 

Wall. 

Acrostichum yoshinagae =Elaphoglossum MAN 

Yatabe 

yoshinagae 

Actinostachys digitata (L.) Wall. =Schizaea digitata MAN 

ex J. Sm. 

Acystopteris japonica var. 

taiwaniana (Tagawa) W. C. 

Shieh 

Adiantum caudatum var. 

edgeworthii (Hook.) Bedd. 

ADIANTACEAE 

=Acystopteris 

taiwaniana 

=Adiantum edgeworthii 

=PTERIDACEAE 

(Subfamily 

Adiantoideae) 

=Odontosoria chinensis 

MAN 

MAN 

Adiantum chinense (L.) Burm. 

MAN 

f. 

Adiantum chusanum L. =Odontosoria chinensis MAN 

Adiantum cultratum Willd. =Lindsaea cultrata MAN 

Adiantum edentulum auct. non =Adiantum 

FOT 2003 

Christ: FOT 1975 

formosanum 

Adiantum monochlamys var. =Adiantum 

MAN 

simozawai Masam. (type: monochlamys 


Adiantum orbiculatum Lam. =Lindsaea orbiculata MAN 

Adiantum pedatum auct. non L. =Adiantum myriosorum FOT 2003 

-- 

191

192 

Adiantum pubescens Schkuhr =Adiantum hispidulum FOC 

Adiantum repens L. f. =Davallia repens MAN 

Adiantum roborowskii var. =Adiantum 

MAN 

taiwanianum (Tagawa) W. C. 

Shieh 

taiwanianum 

Adiantum wangii C. M. Kuo =Adiantum caudatum MAN 

(nom. nud.) 

Aleuritopteris agetae Saiki =Cheilanthes agetae MAN 

(type: Taiwan) 

Aleuritopteris formosana =Cheilanthes 

MAN 

(Hayata) Tagawa 

formosana 

Aleuritopteris subargentea =Cheilanthes 

MAN 

Ching 

subargentea 

Allantodia agyokuensis =Diplazium latifrons this publication 

(Tagawa) C. M. Kuo 

Allantodia amamiana (Tagawa) 

W. M. Chu & Z. R. He 

=Diplazium 

FOC 

amamianum (present 

in Taiwan?), 

?=Diplazium laxifrons 

=Diplazium sikkimense MAN/Knapp, R. 

2013 

Allantodia aspera (Blume) 

Ching 

Allantodia brunoniana Wall. =Diplaziopsis javanica MAN/Christenhusz 

et al. 2011 

Allantodia chinensis (Baker) =Diplazium chinense MAN 

Ching 

Allantodia contermina (Christ) =Diplazium 

MAN 

Ching 

conterminum 

Allantodia dilatata (Blume) =Diplazium dilatatum MAN 

Ching 

Allantodia doederleinii 

=Diplazium doederleinii MAN 

(Luerss.) Ching 

Allantodia incompta (Tagawa) =Diplazium incomptum MAN 

Ching 

Allantodia javanica (Blume) =Diplaziopsis javanica MAN 

Trevis. 

Allantodia kappanensis ?=Diplazium 

FOC 


taiwanense 

Allantodia kawakamii (Hayata) =Diplazium kawakamii MAN 

Ching 

Allantodia latifrons (Alderw.) =Diplazium latifrons alternative 

Ching 

combination 

Allantodia laxifrons (Rosenst.) 

Ching 

Allantodia megaphylla (Baker) 

Ching 

=Diplazium laxifrons 

=Diplazium 

megaphyllum 

(nom. nud.?) 

MAN 

MAN

Allantodia metteniana (Miq.) 

Ching 

Allantodia metteniana var. 

fauriei (Christ) Ching 

Allantodia okinawaensis 

(Tagawa) C. M. Kuo 

=Diplazium 

mettenianum 

=Diplazium 

mettenianum 

=Diplazium 

okinawaense 

MAN 

MAN 

Allantodia okudairae (Makino) =Diplazium okudairae 

Ching 

Allantodia petrii (Tardieu) =Diplazium latifrons MAN 

Ching 

Allantodia phaeolepis 

=Diplazium 


pseudodoederleinii 

Allantodia pseudodoederleinii =Diplazium 

MAN 



Allantodia squamigera (Mett.) =Diplazium 

MAN 

Ching 

squamigerum 

Allantodia taiwanensis 

=Diplazium taiwanense MAN 


Allantodia uraiensis (Rosenst.) 

Ching 

Allantodia virescens (Kunze) 

Ching 

Allantodia virescens var. =Diplazium 

MAN 

okinawaensis (Tagawa) W. okinawaense 

M. Chu 

Allantodia wichurae (Mett.) =Diplazium wichurae FOC 

Ching 

Allosorus argenteus C. Presl =Cheilanthes argentea MAN 

Alsophila acaulis Makino =Cyathea hancockii MAN 

Alsophila denticulata Baker =Cyathea hancockii MAN 

Alsophila fauriei Christ =Cyathea spinulosa MAN 

alternative 

combination 

(nom. nud.?) 

MAN 

Kuo, C. M. 1985 

=Diplazium dilatatum MAN/Knapp, R. 

2011 

=Diplazium virescens MAN 

Alsophila fenicis (Copel.) C. =Cyathea fenicis MAN 

Chr. 

Alsophila formosana Baker =Cyathea metteniana MAN 

Alsophila fujiana Nakai (type: =Cyathea fenicis MAN 


Alsophila lepifera J. Sm. ex =Cyathea lepifera MAN 

Hook. 

Alsophila loheri (Christ) R. M. =Cyathea loheri MAN 

Tryon 

Alsophila metteniana Hance =Cyathea metteniana MAN 

Alsophila podophylla Hook. =Cyathea podophylla MAN 

Alsophila polypodioides Hook. =Macrothelypteris MAN 

polypodioides 

Alsophila pustulosa Christ =Cyathea lepifera MAN 

193

194 

Alsophila spinulosa (Wall. ex =Cyathea spinulosa MAN 

Hook.) R. M. Tryon 

Alsophila subglandulosa Hance =Ctenitis 

MAN 


subglandulosa 

Alsophila taiwaniana Nakai =Cyathea spinulosa FOT 2003 

Alsophylla podophylla Hook. =Cyathea podophylla FOT 2003 

Amesium sasakii Hayata (type: =Asplenium 

MAN 



Ampelopteris prolifera (Retz.) =Cyclosorus proliferus MAN 

Copel. 

Anapausia bonii (Christ) Nakai =Tectaria harlandii FOT 2003/MAN 

Anapausia decurrens (Blume) =Leptochilus decurrens FOT 2003 

C. Presl 

Anapausia harlandii (Hook.) =Tectaria harlandii FOT 2003/MAN 

Nakai 

Angiopteris angustifolia C. =Angiopteris 

MAN 

Presl 

palmiformis 

Angiopteris durvilleana de =Angiopteris 

MAN 

Vriese 

palmiformis 

Angiopteris fauriei var. 

formosana Hieron. (type: 


? (MAN) published 

in: Hedwigia 61: 

274 (1919) 

Angiopteris formosana Ching 


Angiopteris henryi Hieron. 


Angiopteris lobulata Ching 


Angiopteris oldhamii Hieron. 


Angiopteris rahaoensis Ching 


Angiopteris sakuraii Hieron. 


Angiopteris suboppositifolia 

auct. non de Vriese 


in: Fl. Reipubl. 

Popularis Sin. 2: 

335 (1959) 



260 (1919) 




334 (1959) 

=Angiopteris lygodiifolia 

(MAN) published 


265 (1919) 




333 (1959) 



280 (1919) 

=Angiopteris lygodiifolia FOT 2003

195 

Angiopteris taiwanensis Ching 


Anisocampium cumingianum 

C. Presl 

Anisocampium sheareri 

(Baker) Ching in Y. T. Hsieh 

Anisogonium esculentum 

(Retz.) C. Presl 

Antrophyum cumingii auct. non 

Fee: Masam. & Hayata 

Antrophyum cumingii Fee 

Antrophyum grevillei auct. non 

Balf. 

Antrophyum japonicum Makino 

Antrophyum latifolium var. 

obovatum (Baker) C. Chr. 

Antrophyum plantagineum 

auct. non Kaulf. 

Antrophyum reticulatum auct. 

non Kaulf. 

Antrophyum reticulatum var. 

parvulum (Blume) Bedd. 

Arachniodes arisanica 

(Rosenst.) Ching 

Arachniodes diffracta (Baker) 

Ching 

Arachniodes dimorphophyllum 


Arachniodes exilis (Hance) 

Ching 

Arachniodes fujian(g)ensis 

auct. non Ching 

Arachniodes hasseltii (Blume) 

Ching 

Arachniodes rhomboidea var. 

yakusimensis (H. Ito) W. C. 

Shieh 

Arachniodes sp. in Knapp, R. 

2011 

Arachniodes subreflexipinna 

(Ogata) Ching 




334 (1959) 

=Athyrium 

MAN 

cumingianum 

=Athyrium sheareri MAN 

=Diplazium esculentum 

=Antrophyum 

formosanum 

=Antrophyum 

sessilifolium 

=Antrophyum 

formosanum 

=Antrophyum 

obovatum 

=Antrophyum 

obovatum 

=Antrophyum 

obovatum 

=Antrophyum 

formosanum 

=Antrophyum parvulum 

=Arachniodes globisora 

=Dryopteris diffracta 

=Polystichum deltodon 

MAN 

FOT 2003 

FOT 2003 

FOT 2003 

FOT 2003 

MAN 

FOT 2003 

FOT 2003 

MAN 

MAN 

FOC 

MAN 

=Arachniodes aristata Knapp, R. 2011 

=Arachniodes caudata Knapp, R. 2011 

=Dryopteris hasseltii 

=Arachniodes 

yakusimensis 

=Arachniodes 

chinensis 

=Dryopteris 

subreflexipinna 

MAN 

FOC/Knapp, R. 

2011 


MAN

196 

Arachniodes tripinnata 

(Goldm.) Sledge 

Araiostegia clarkei (Baker) 

Copel. 

Araiostegia divaricata var. 

formosana (Hayata) M. Kato 

Araiostegia hookeri (T. Moore) 

Ching 

Araiostegia parvipinnata 

(Hayata) Copel. 

Araiostegia perdurans (Christ) 

Copel. 

Archangiopteris henryi var. 

somai (Hayata) Tagawa 

Archangiopteris itoi W. C. 

Shieh (type: Taiwan) 

Archangiopteris somai Hayata 


Arthromeris pinnata (Hayata) 

Ching 

Arthropteris obliterata (R. 

Brown) J. Sm. 

ASPIDIACEAE 

Aspidium acanthophyllum 

Franch. 

Aspidium amabile Blume 

Aspidium angustifrons Miq. 

=Arachniodes 

pseudoaristata 

=Araiostegia 

parvipinnula 

=Davallia formosana 

=Araiostegia 

parvipinnula 

typo, =Araiostegia 

parvipinnula 

=Araiostegia 

parvipinnula 

=Angiopteris somae 

=Angiopteris itoi 

=Angiopteris somae 

=Arthromeris lehmannii 

Kuo, C. M. 

1985/MAN 

MAN 

MAN 

MAN 

MAN 

MAN 

MAN 

MAN 

MAN 

MAN 

=Arthropteris palisotii FOT 2003 

=DRYOPTERIDACEAE 

(Subfamily 

Dryopteridoideae / 

Tribe Tectarieae) 

=Polystichum 

acanthophyllum 

=Arachniodes 

rhomboidea 

=Thelypteris 

angustifrons 

=Dryopsis apiciflora 

-- 

MAN 


2011 

MAN 

Aspidium apiciflorum Wall. ex 

MAN 

Mett. 

Aspidium aridum D. Don =Cyclosorus aridus MAN 

Aspidium atratum Wall. =Dryopteris atrata FOT 2003 

Aspidium auriculatum (L.) Sw. =Nephrolepis cordifolia MAN 

Aspidium auriculatum var. 

stenophyllum (Baker) 

Matthew 

=Polystichum 

hecatopterum 

FOT 2003 

Aspidium biaristatum Blume =Polystichum 

MAN 

biaristatum 

Aspidium biserratum Sw. =Nephrolepis biserrata MAN 

Aspidium boryanum Willd. =Deparia subfluvialis FOT 2003/MAN

Aspidium brachiatum auct. non 

Zoll. & Mor.: MAN 

=Tectaria impressa 

Fraser-Jenkins, 

C. R. 2008/ 


MAN 

197 

Aspidium caryotideum Wall. ex 

Hook. & Grev. 

=Polystichum 

caryotideum 

Aspidium coadunatum Wall. ex =Tectaria coadunata MAN 

Hook. & Grev. 

Aspidium crenatum (Forssk.) =Hypodematium MAN 

Kuhn 

crenatum 

Aspidium cycadinum Franch. & =Dryopteris cycadina MAN 

Sav. 

Aspidium decurrens C. Presl =Tectaria decurrens MAN 

Aspidium deltodon Baker =Polystichum deltodon MAN 

Aspidium devexum Kunze ex =Tectaria devexa MAN 

Mett. 

Aspidium dickinsii Franch. & =Dryopteris dickinsii MAN 

Sav. 

Aspidium diffractum (Baker) =Dryopteris diffracta FOC 

Christ 

Aspidium discretum auct. non =Polystichum 


Buch.-Ham. ex D. Don: MAN piceopaleaceum C. R. 2008 

Aspidium drepanopterum auct. =Athyrium mupinense MAN/Liu, Y. C. et 

non (Kunze) Mett.: MAN 

al. 2009 

Aspidium dubium (C. B. Clarke =Tectaria dubia Knapp, R. 2013 

& Baker) Bedd. 

Aspidium duthiei auct. non C. =Polystichum glaciale MAN/Knapp, R. 

Hope: MAN 

2011 

Aspidium exile Hance =Arachniodes aristata MAN/Knapp, R. 

2011 

Aspidium falcatum (L. f.) Sw. =Polystichum falcatum MAN 

Aspidium falcatum var. =Polystichum 

MAN 

macrophyllum Makino 

macrophyllum 

Aspidium festinum Hance =Arachniodes festina MAN 

Aspidium filix-mas var. 

=Dryopteris 

FOT 2003/MAN 

chrysocoma Christ 

woodsiisora 

Aspidium formosanum Christ =Dryopteris formosana MAN 


Aspidium fraxinellum Christ =Polystichum 

MAN 

fraxinellum 

Aspidium fuscipes Wall. ex =Tectaria fuscipes MAN 

Bedd. 

Aspidium glanduligerum Kunze =Thelypteris 

MAN 

glanduligera 

Aspidium gongylodes Schkuhr =Cyclosorus interruptus FOT 2003 

Aspidium gracilescens Blume =Thelypteris 

gracilescens 

MAN

198 

Aspidium griffithii (T. Moore) =Cyclosorus griffithii FOT 2003/MAN 

Diels 

Aspidium hancockii (Hance) =Polystichum hancockii MAN 

Baker 

Aspidium hokutoense Hayata =Tectaria subtriphylla FOT 2003 


Aspidium jaculosum Christ =Cyclosorus jaculosus MAN 


Aspidium kwanonense Hayata =Tectaria coadunata MAN 


Aspidium kwarenkoensis =Tectaria simonsii MAN 

Hayata (type: Taiwan) 

Aspidium labordei auct. non 

Christ: FOT 2003 

=Dryopteris tenuicula FOT 2003/Knapp, 

R. 2011 

Aspidium lachenense Hook. =Polystichum 

MAN 

lachenense 

Aspidium latipinnum (Bentham) =Cyclosorus latipinnus FOC 

Hance 

Aspidium laxum Franch. & Sav. =Thelypteris laxa MAN 

Aspidium lepidocaulon Hook. =Polystichum 

MAN 

lepidocaulon 

Aspidium leucostipes (Baker) =Ctenitis eatonii MAN 

Christ 

Aspidium lobulatum Christ =Cyclosorus 

FOT 2003 

taiwanensis 

Aspidium macrocarpon Blume =Athyrium puncticaule Liu, Y. C. et al. 

2009 

Aspidium manmeiense Christ =Polystichum 

MAN 

manmeiense 

Aspidium marginatum Wall. =Dryopteris marginata FOT 2003 

(nom. nud.) 

Aspidium matsumurae Makino =Dryopsis 

FOT 2003 

maximowicziana 

Aspidium maximowiczianum =Dryopsis 

MAN 

Miq. 


Aspidium melanocaulon auct. =Tectaria simonsii FOT 2003/MAN 

non Blume 

Aspidium membranifolium (C. =Tectaria fuscipes FOT 2003/MAN 

Presl) Hook. 

Aspidium molle var. latipinnum =Cyclosorus latipinnus MAN 

Benth. 

Aspidium nantoense Hayata =Tectaria polymorpha MAN 


Aspidium nepalense Spreng. =Polystichum 

nepalense 

MAN

199 

Aspidium nigripes auct. non 

Blume: MAN 

=Athyrium tozanense MAN/Liu, Y. C. et 

al. 2009 

Aspidium obliquum D. Don =Polystichum obliquum MAN 

Aspidium oligophlobium =Macrothelypteris FOT 2003/MAN 

(Baker) Christ 

torresiana 

Aspidium pachinense Hayata =Tectaria simonsii MAN 


Aspidium palisotii Desv. =Arthropteris palisotii MAN 

Aspidium paludosum auct. non 

Blume: FOT 2003 

=Pseudophegopteris 

hirtirachis 

FOT 2003/Knapp, 

R. 2011 

Aspidium parasiticum (L.) Sw. =Cyclosorus 

MAN 

parasiticus 

Aspidium patens (Sw.) Sw. =Cyclosorus 

FOT 2003 

parasiticus 

Aspidium phaeocaulon =Tectaria phaeocaulis FOC 

Rosenst. 

Aspidium polymorphum Wall. =Tectaria polymorpha MAN 

ex Hook. 

Aspidium prescottianum Wall. =Polystichum 

MAN 

ex Mett. 

prescottianum 

Aspidium prescottianum var. =Polystichum sinense MAN 

sinense Christ 

Aspidium productum Kaulf. =Cyclosorus productus MAN 

Aspidium pteropus Kunze =Tectaria devexa MAN 

Aspidium puncticaule Blume =Athyrium puncticaule Liu, Y. C. et al. 

2009 

Aspidium rhomboideum =Arachniodes 

MAN 

(Schott) Wall. ex Mett. 

rhomboidea 

Aspidium rufinerve Hayata =Pleocnemia rufinervis MAN 


Aspidium simonsii (Baker) =Tectaria simonsii MAN 

Bedd. 

Aspidium subexaltatum Christ =Dryopteris subexaltata MAN 

Aspidium submembranaceum =Pleocnemia 

MAN 


submembranacea 

Aspidium subpubescens auct. =Cyclosorus jaculosus FOT 2003 

non Blume 

Aspidium subtriphyllum (Hook. =Tectaria subtriphylla MAN 

& Arn.) Hook. 

Aspidium subtriphyllum f. 

cuspidatopinnatum Hayata 

=Tectaria impressa FOT 2003/Knapp, 

R. 2011 

(type: Taiwan), nom. nud.? 

Aspidium subtriphyllum var. =Tectaria simonsii MAN 

ebenosum Nakai 

Aspidium subtripinnatum Miq. 

=Ctenitis 

subglandulosa 

FOT 2003

200 

Aspidium tacticopterum Kunze =Polystichum 

mucronifolium 

MAN/Knapp, R. 

2011 

Aspidium tenuisectum Blume =Acystopteris 

MAN 

tenuisecta 

Aspidium thomsonii Hook. f. =Polystichum thomsonii MAN 

Aspidium trifolium Alderw. =Tectaria polymorpha MAN 

Aspidium tsus-simense Hook. =Polystichum 

MAN 

tsussimense 

Aspidium uliginosa (Kunze) =Macrothelypteris FOT 2003/MAN 

Ching 

torresiana 

Aspidium variolosum Wall. ex 

Hook. 

=Tectaria impressa Fraser-Jenkins, 

C. R. 2008/ 


Aspidium varium (L.) Sw. =Dryopteris varia FOC 

Aspidium varium var. 

=Dryopteris fructuosa MAN 

fructuosum Christ 

Aspidium wallichianum Spreng. =Dryopteris wallichiana MAN 

subsp. wallichiana 

Aspidium wallichii Hook. =Oleandra wallichii MAN 

Aspidium xiphophyllum Baker =Polystichum 

MAN 

xiphophyllum 

Aspidium yunnanense (Baker) 

Christ 

=Tectaria griffithii MAN/this 

publication 

Asplenium achilleifolium auct. =Asplenium 

FOC 

non (Lam.) C. Chr. 

prolongatum 

Asplenium adiantoides auct. =Asplenium falcatum MAN 

non (L.) C. Chr. 

Asplenium anceps var. =Asplenium tripteropus MAN 

proliferum Nakai 

Asplenium arifolium Burm. f. =Paraceterach cordata MAN/Knapp, R. 

2013 

Asplenium arisanense Tagawa =Asplenium 

MAN 


cuneatiforme 

Asplenium atkinsonii (Bedd.) C. =Athyrium atkinsonii MAN 

B. Clarke 

Asplenium auritum var. =Asplenium obscurum MAN 

obscurum (Blume) E. Forn. 

Asplenium austrochinense =Asplenium wilfordii FOT 1975 

auct. non Ching 

Asplenium baibarense 

Tatewaki & Tagawa (type: 


=Asplenium 

griffithianum 

FOT 2003 

Asplenium bicuspe Hayata =Asplenium ensiforme MAN 

Asplenium billetii Christ =Asplenium 

FOC 

pulcherrimum

201 

Asplenium calcicola Tagawa =Asplenium 

MAN 


pulcherrimum 

Asplenium centrochinense =Asplenium wrightii FOC 

Christ 

Asplenium chinense Baker =Diplazium chinense MAN 

Asplenium chlorophyllum 

Baker (type: Taiwan) 


in: J. Bot. 5: 104 

(1885) 

Asplenium conilli auct. non =Deparia petersenii FOT 2003 

Franch. & Sav. 

Asplenium cuneatum auct. =Asplenium 

MAN 

non. Lam. 

neolaserpitiifolium 

Asplenium davallioides Hook. =Asplenium ritoense FOT 2003 

Asplenium doederleinii Luerss. =Diplazium doederleinii MAN 

Asplenium donianum Mett. =Diplazium donianum MAN 

Asplenium ensiforme f. bicuspe =Asplenium ensiforme MAN 

(Hayata) Ching ex S. H. Wu 

Asplenium ensiforme var. =Asplenium ensiforme FOC 

bicuspe (Hayata) Tagawa 

Asplenium exiguum auct. non =Asplenium 

FOT 2003 

Bedd. 

yunnanense 

Asplenium formosanum Baker =Asplenium oldhamii FOC 

Asplenium gueinzianum Mett. =Asplenium lacinioides Knapp, R. 2013 

ex Kuhn 

Asplenium hancockii Baker =Asplenium oldhamii FOC 

Asplenium hancockii Maxim. =Diplazium pullingeri MAN 


Asplenium heterocarpum Wall. =Asplenium 

FOC 

ex Hook. 

cheilosorum 

Asplenium holophyllum Baker =Asplenium 

FOC 


griffithianum 

Asplenium iridiphyllum Hayata =Asplenium 

MAN 


griffithianum 

Asplenium japonicum auct. non =Deparia petersenii FOT 2003 

Thunb. 

Asplenium javanicum Blume =Diplaziopsis javanica MAN 

Asplenium laciniatum auct. non 

D. Don 

=Asplenium lacinioides MAN /Knapp, R. 

2013 

Asplenium lanceum Thunb. =Deparia lancea MAN 

Asplenium laserpitiifolium auct. =Asplenium 


non Lam. 

neolaserpitiifolium 

Asplenium laserpitiifolium var. =Asplenium wilfordii MAN 

morrisonense Hayata 

(syntype: Taiwan) 

Asplenium latecuneatum Christ =Asplenium x 

shikokianum 

FOT 2003

202 

Asplenium lobulatum auct. non =Asplenium 


Mett. ex Kuhn: MAN 

cuneatiforme 

Asplenium loriceum var. =Asplenium formosae MAN 

karapinense (Hayata) 

Tagawa 

Asplenium makinoi Hayata =Asplenium formosae MAN 


Asplenium matsumurae Christ =Asplenium serricula MAN 


Asplenium megaphyllum Baker =Diplazium 

MAN 

megaphyllum 

Asplenium mettenianum Miq. =Diplazium 

FOT 2003 

mettenianum 

Asplenium morrisonense =Asplenium wilfordii MAN 

(Hayata) Hayata 

Asplenium muricatum auct. 

non Mett.: MAN 

=Diplazium kawakamii MAN/Knapp, R. 

2011 

Asplenium nakanoanum =Asplenium 

MAN 

Makino 

griffithianum 

Asplenium nigripes auct. non 

(Blume) Hook. 

=Athyrium tozanense FOC/Knapp, R. 

2011 

Asplenium niponicum Mett. =Athyrium niponicum Liu, Y. C. et al. 

2009 

Asplenium normale var. 

boreale Ohwi ex Sa. Kurata 

=Asplenium boreale J. Geobot. 11: 

100 (1963) 

Asplenium obscurum var. 

angustum Tagawa (type: 



in: Acta Phytotax. 

Geobot. 5: 194- 

195 (1936) 

Asplenium otophorum Miq. =Athyrium otophorum Liu, Y. C. et al. 

2009 

Asplenium petersenii Kunze =Deparia petersenii MAN 

Asplenium planicaule Wall. ex 

Hook. 

Asplenium planicaule var. 

yoshinagae (Makino) Tagawa 

Asplenium polyodon auct. non 

G. Forst 

Asplenium pseudofalcatum var. 

subintegrum Rosenst. (type: 


=Asplenium 

MAN 

yoshinagae 

=Asplenium 

MAN 

yoshinagae 

=Asplenium falcatum Knapp, R. 2013 



334 (1915) 

Asplenium pullingeri Baker =Diplazium pullingeri MAN 

Asplenium rahaoense (Y. Yabe =Asplenium excisum MAN 

ex Hayata) H. Ito 

Asplenium ramosum L. =Asplenium viride MAN

Asplenium resectum var. 

adiantifrons Hayata (type: 


Asplenium resectum var. 

rahaoense Y. Yabe ex 

Hayata (syntype: Taiwan) 

Asplenium robustum auct. non 

Blume 

203 


in Icon. Pl. 

Formosan. 4: 226 

(1914) 

=Asplenium excisum MAN 

=Asplenium 

pseudolaserpitiifolium 

misidentification 

of a specimen 

from S Taiwan (C. 

M. Kuo no. 1886 

in PH) 

FOT 2003 

Asplenium rutaefolium auct. 

non Kunze 

=Asplenium 

prolongatum 

Asplenium sarelii var. 

=Asplenium pekinense MAN 

pekinense (Hance) C. Chr. 

Asplenium sasakii Hayata =Asplenium 

MAN 


Asplenium scolopendrifrons =Asplenium 

MAN 


griffithianum 

Asplenium setoi N. Murak. & eventually correct Knapp, R. 2013 

Seriz. 

name and replacing 

Asplenium 

australasicum in 


Asplenium sinense Baker =Athyrium arisanense FOT 2003 

Asplenium squamigerum Mett. =Diplazium 

MAN 

squamigerum 

Asplenium strigillosum T. =Athyrium strigillosum Liu, Y. C. et al. 

Moore ex E. J. Löwe 

2009 

Asplenium subsinuatum Wall. =Deparia lancea MAN 


Asplenium subvarians Ching =Asplenium tenuicaule FOT 2003 

Asplenium taiwanense Ching =Asplenium wrightii MAN 

ex S. H. Wu 

Asplenium tenuissimum =Asplenium tenuifolium MAN 


Asplenium tozanense Hayata =Athyrium tozanense Liu, Y. C. et al. 


2009 

Asplenium trichomanes subsp. =Asplenium tripteropus MAN 

tripteropus (Nakai) A. Love & 

D. Love

204 

Asplenium unilaterale auct. non 

Lam. 

Asplenium unilaterale var. 

obliquissimum Hayata (type: 


Asplenium unilaterale var. 

rahaoense (Y. Yabe ex 

Hayata) Hayata 

Asplenium varians auct. non 

Wall. ex Hook. & Grev. 

Asplenium vidalii Franch & 

Sav. 

Asplenium viridissimum Hayata 


Asplenium vulcanicum auct. 

non. Blume 

misapplied for: 

MAN 

Asplenium 

apogamum, 

Asplenium 

cataractarum, 

Asplenium filipes, 

Asplenium 

subnormale 

? published in Icon. 

Pl. Formosan. 4: 

230 (1914) 

=Asplenium excisum MAN 

=Asplenium tenuicaule 

=Athyrium vidalii 

=Asplenium bullatum 

=Asplenium serricula 

MAN 

Liu, Y. C. et al. 

2009 

MAN 

MAN 

Asplenium wichurae Mett. =Diplazium wichurae MAN 

Asplenium wilfordii var. densa 

Rosenst. (type: Taiwan) 

Asplenium wrightii var. aristoserrulatum 

Hayata (type: 


ATHYRIACEAE 



334 (1915) 

?=Asplenium wrightii 

=DRYOPTERIDACEAE 

(Subfamily 

Athyrioideae / Tribe 

Physematieae) 

=Deparia petersenii 

published in: Icon. 


232, fig. 4 (1914) 

Athyriopsis japonica auct. non 

MAN 

(Thunb.) Ching: FOT 1975 

Athyriopsis petersenii (Kunze) =Deparia petersenii MAN 

Christ 

Athyrium acrostichoides auct. =Deparia allantodioides FOT 2003 

non (Sw.) Diels 

Athyrium allanticarpum =Diplazium kawakamii MAN/Knapp, R. 


2011 

Athyrium allantodioides Bedd. =Deparia allantodioides MAN 

Athyrium asperum (Blume) =Diplazium sikkimense MAN/Knapp, R. 

Milde 

2013 

Athyrium boryanum auct. non =Deparia subfluvialis MAN 

(Willd.) Tagawa: FOT 2003 

--

205 

Athyrium brunonianum (Wall.) 

Milde 

=Diplaziopsis javanica MAN/Christenhusz 

et al. 2011 

Athyrium clivicola auct. non 

Tagawa: Acta Phytotax. Sin. 

=Athyrium auriculatum Liu, Y. C. et al. 

2009 

34: 182 

Athyrium commixtum Koidz. =Athyrium vidalii FOT 2003 

Athyrium conilii auct. non =Deparia petersenii FOT 2003 

(Franch. & Sav.) Tagawa 

Athyrium dawuense C. M. Kuo 

(nom. nud.) 

?=Athyrium arisanense (Kuo, C. M. 1997) 


Athyrium decurrenti-alatum =Cornopteris 

MAN 

(Hook.) Copel. 


Athyrium dilatatum (Blume) =Diplazium dilatatum MAN 

Milde 

Athyrium doederleinii (Luerss.) =Diplazium doederleinii MAN 

Ohwi 

Athyrium drepanopterum auct. 

non (Kunze) A. Braun ex 

=Athyrium mupinense Liu, Y. C. et al. 

2009 

Milde: FOT 1975 

Athyrium elegans Tagawa =Athyrium delavayi var. Liu, Y. C. et al. 

Athyrium elegans var. 

purpurascens Tagawa 

subrigescens 2009 


in: Acta Phytotax. 

Geobot. 3: 33-34 

(1934) 

Athyrium esculentum (Retz.) =Diplazium esculentum MAN 

Copel. 

Athyrium fimbriatum auct. non 

(Hook.) T. Moore 

=Athyrium tripinnatum Liu, Y. C. et al. 

2009 

Athyrium fluviale (Hayata) C. =Cornopteris 

MAN 

Chr. 

banajaoensis 

Athyrium foliolosum auct. non 

T. Moore ex R. Sim.: Kuo, C. 

=Athyrium tripinnatum Liu, Y. C. et al. 

2009 

M. 1985 

Athyrium formosanum 

(Rosenst.) Copel. 

=Dictyodroma 

formosana 


2011 

Athyrium fragile auct. non (L.) 

Spreng.: MAN 

=Athyrium pulchrum MAN/Knapp, R. 

2011 

Athyrium fragile Tardieu =Athyrium pulchrum MAN/Knapp, R. 

2011 

Athyrium goeringianum auct. 

non. (Kunze) T. Moore: FOT 

1975 

=Athyrium iseanum Liu, Y. C. et al. 

2009 

Athyrium x hohuanshanense 

Yoshik. 

?=Athyrium 

cryptogrammoides 

(MAN) Knapp, R. 

2011

206 

Athyrium iseanum f. angustisectum 

(Tagawa) Sa. Kurata 

Athyrium iseanum var. 

angustisectum Tagawa 

Athyrium japonicum auct. non 

(Thunb.) Copel.: FOT 2003 

Athyrium kawakamii (Hayata) 

C. Chr. 

Athyrium kuankaoense C. M. 

Kuo (nom. nud.) 

Athyrium kuratae auct. non 

Seriz.: FOT 1975 

=Athyrium iseanum 

=Athyrium iseanum 


=Diplazium kawakamii 

Hokuriku Journ. 

Bot. 6: 42. 

1957/Liu, Y. C. et 

al. 2009 


2009 (only taxon 

which is present 

in Taiwan according 

to Liu, Y. C. et 

al. 2009) 

MAN 

MAN 

? (MAN) 

=Athyrium arisanense 

in part, =Athyrium 

delavayi in part, 

=Athyrium otophorum 

in part, =Athyrium 

pubicostatum in part 

=Athyrium anisopterum 


2009 

Athyrium macrocarpum 

(Blume) Bedd. 

Athyrium mengtzeense Hieron. =Athyrium arisanense FOT 2003 

Athyrium mettenianum (Miq.) =Diplazium 

MAN 

Ohwi 

mettenianum 

Athyrium micans Tagawa =Athyrium mupinense 


Athyrium monticola Rosenst. =Athyrium atkinsonii FOT 2003 

Athyrium nigripes auct. non. =Athyrium tozanense 

(Blume) T. Moore: Kuo, C. M. 

1985 

Athyrium obtusifolium Rosenst. 


Athyrium obtusifolium f. 

pumilum Rosenst. (type: 


Athyrium okudairae (Makino) 

Ohwi 

Athyrium opacum (D. Don) 

Copel. 

Athyrium oppositipinnum 

Hayata (mis-spelling of 

species epithet) 

=Athyrium nakanoi 

=Athyrium nakanoi 

=Diplazium okudairae 

=Cornopteris opaca 

=Athyrium 

oppositipennum var. 



2009 


2009 


2009 


2009 


2009 

MAN 

MAN 


2009

=Athyrium oppositipennum var. 

pubescens (Tagawa) Y. C. 

Liu 

Athyrium oreopteris Copel. 

=Athyrium 


pubescens (Tagawa) 

Tagawa 

=Athyrium 


Athyrium oshimense Christ =Deparia petersenii FOC 

Athyrium petrii (Tardieu) Ohwi =Diplazium latifrons MAN 

Athyrium procerum auct. non 

(Wall. ex C. B. Clarke) Milde 


var. kawakamii 

Athyrium procerum var. 

subglabratum (Tagawa) 

Tagawa 

Athyrium pseudo-arisanense 

C. M. Kuo (nom. nud.) 

Athyrium x pseudocryptogrammoides 

Yoshikawa 

Athyrium pseudo-delavayi C. 

M. Kuo (nom. nud.) 

Athyrium pseudo-kuratae C. M. 


Athyrium pseudo-leiopodum C. 


Athyrium pseudo-mearnsianum 


Athyrium pseudo-niponicum C. 


Athyrium pseudo-silvicola C. 


Athyrium pullingeri (Baker) 

Copel. 

Athyrium x purpurascens 

(Tagawa) Kurata 

Athyrium pycnosorum auct. 

non Christ: FOT 2003 

Athyrium reflexipinnum Hayata 



var. subglabratum 

this publication 


2009 

207 

Ohwi, J. Flora of 

Japan. Smithsonian 

Institution, 

USA (1965)/ 


Ohwi, J. Flora of 

Japan. Smithsonian 

Institution, 

USA (1965)/ 


?=Athyrium arisanense (MAN) Knapp, R. 

2011 

=Athyrium 


cryptogrammoides 2009 

=Athyrium delavayi Liu, Y. C. et al. 

2009 

? (MAN) 

=Athyrium arisanense Liu, Y. C. et al. 

2009 

?=Athyrium arisanense (MAN) Knapp, R. 

2011 

?=Athyrium niponicum (MAN) Knapp, R. 

2011 

? (MAN) 

=Diplazium pullingeri 

MAN 


in: J. Geobot. 14: 

4 (1965) 

=Deparia allantodioides MAN 

=Athyrium 


pubescens 


2009

208 

Athyrium reflexipinnum auct. 

non Hayata 

=Athyrium palustre Chang, Y. H. et 

al. 2014 

Athyrium roseum auct. non. 

Christ: Acta Phytotax. Sin. 


2009 

34(2): 190 

Athyrium silvicola auct. non 

Tagawa: Taiwania 50:150 

=Athyrium auriculatum Liu, Y. C. et al. 

2009 

(2005) 

Athyrium sp. 1 in Knapp, R. =Athyrium sp. 


2011 

Athyrium sp. 2 in Knapp, R. =Athyrium palustre this publication 

2011 

Athyrium squamigerum (Mett.) =Diplazium 

MAN 

Ohwi 

squamigerum 

Athyrium subfluviale (Hayata) =Deparia subfluvialis MAN 

Tagawa 

Athyrium subrigescens 

(Hayata) Hayata ex H. Ito 

=Athyrium delavayi var. 

subrigescens 


2009 

Athyrium sungkangense C. M. 


=Athyrium 

erythropodum 


2009 

Athyrium taiwanense Tagawa 


=Athyrium 



2009 

Athyrium taiwanense var. 

pubescens Tagawa (type: 

=Athyrium 



2009 


pubescens 

Athyrium takeoi (Hayata) 

Tagawa 

=Athyrium mupinense Liu, Y. C. et al. 

2009 

Athyrium tenuicaule (Hayata) 

Tagawa 

=Athyrium leiopodum Liu, Y. C. et al. 

2009 

Athyrium tenuisectum (Blume) =Acystopteris 

MAN 

T. Moore 

tenuisecta 

Athyrium tenuissimum (Hayata) 

Merr. 

=Athyrium nakanoi Liu, Y. C. et al. 

2009 

Athyrium thysanocarpum 

Hayata 

=Athyrium anisopterum Liu, Y. C. et al. 

2009 

Athyrium unifurcatum (Baker) =Deparia unifurcata FOC 

C. Chr. 

Athyrium unifurcatum var. =Deparia subfluvialis MAN 

subfluviale (Hayata) H. Ito 

Athyrium virescens (Kunze) =Diplazium virescens MAN 

Ohwi 

Athyrium wichurae (Mett.) =Diplazium wichurae MAN 

Merr. 

Athyrium woodsioides Christ =Athyrium mupinense Liu, Y. C. et al. 

2009

Athyrium xichouense Y. T. 

Hsieh & Z. R. Wang 

Athyrium yakusimense Tagawa 

=Athyrium silvicola Liu, Y. C. et al. 

2009 

=Athyrium 

FOT 2003 

erythropodum 

=Athyrium sp. Knapp, R. 2011 

Athyrium yui auct. non Ching: 

Liu 2008 

Azolla caroliniana Willd. =Azolla filiculoides Knapp, R. 2011 

Azolla imbricata (Roxb.) Nakai =Azolla pinnata FOC 

Azolla japonica Franch. & Sav. ?=Azolla filiculoides Knapp, R. 2011 

Azolla pinnata var. imbricata =Azolla pinnata FOC 

(Roxb.) Bonaparte 

Belvisia formosana (M. Ogata) =Belvisia mucronata MAN 

Ching 

Blechnidium melanopus =Blechnum melanopus MAN 


Blechnidium plagiogyriifrons =Blechnum melanopus MAN 


Blechnopsis orientalis (L.) J. =Blechnum melanopus MAN 

Presl 

Blechnum integripinnulum =Blechnum fraseri MAN 


Blechnum insignis (Hook.) C. =Brainea insignis MAN 

M. Kuo 

Blechnum japonicum L. =Woodwardia japonica MAN 

Blechnum plagiogyriifrons =Blechnum melanopus MAN 


Blechnum stenopterum Hance =Plagiogyria stenoptera MAN 


Bolbitis contaminans auct. non =Bolbitis angustipinna MAN 

(Wall.) Ching: FOT 1975 

Bolbitis formosana Tagawa =Bolbitis subcordata MAN 


Bolbitis hainanensis auct. non =Bolbitis virens var. Knapp, R. 2011 

Ching & C. H. Wang 

compacta 

Bolbitis koidzumiana Tagawa =Bolbitis heteroclita FOT 2003 

Botrychium arisanense =Botrychium 

MAN 

Masam. 

lanuginosum 

Botrychium daucifolium auct. 

non Wall. ex Hook. & Grev.: 

FOT 2003 

=Botrychium 

formosanum 

MAN 

Botrychium japonicum auct. 

non Underw.: FOT 1975 

Botrychium lanuginosum var. 

leptostachyum (Hayata) 

Nakai 

=Botrychium 

formosanum 

=Botrychium 

lanuginosum 

MAN 

MAN 

209

210 

Botrychium leptostachyum =Botrychium 

MAN 


lanuginosum 

Botrychium virginianum var. =Botrychium 

MAN 

lanuginosum (Wall. ex Hook. 

& Grev.) T. Moore 

lanuginosum 

Botrychium zeylanicum (L.) =Helminthostachys MAN 

Sw. 

zeylanica 

Botrypus lanuginosus (Wall. ex =Botrychium 

MAN 

Hook. & Grev.) Holub 

lanuginosum 

Botrypus lanuginosus subsp. =Botrychium 

MAN 

leptostachys (Hayata) Holub lanuginosum 

Botrypus lunaria (L.) Rich. =Botrychium lunaria MAN 

Bowringia insignis Hook. =Brainea insignis MAN 

Brainea formosana Hayata =Brainea insignis MAN 


Brainea insignis var. 

=Brainea insignis MAN 

formosana (Hayata) Tagawa 

Callipteris esculenta (Retz.) J. =Diplazium esculentum MAN 

Sm. ex T. Moore & Houlston 

Callipteris prolifera (Lam.) Bory =Diplazium proliferum FOC 

Callistopteris apiifolia (C. Presl) =Cephalomanes MAN 

Copel. 

apiifolium 

Calymmodon asiaticus auct. =Calymmodon 


non Copel.: Kuo, C. M. 2010 cucullatus 

Calymmodon glabrescens =Calymmodon 

FOT 2003/MAN 

auct. non Copel. 

cucullatus 

Calymmodon gracillimus =Calymmodon 

MAN 

(Copel.) Nakai ex H. Ito cucullatus 

Calymmodon hyalinus auct. =Calymmodon gracilis MAN 

non Copel. 

Campium subcordatum Copel. =Bolbitis subcordata MAN 

Cephalomanes auriculatum =Crepidomanes MAN 

(Blume) Bosch 

auriculatum 

Cephalomanes laciniatum 

(Roxb.) DeVol 

=Cephalomanes 

javanicum var. 

MAN 

Cephalomanes oblongifolium 

C. Presl 

CERATOPTERIDACEAE 

asplenioides 

=Cephalomanes 


asplenioides 

=PTERIDACEAE 

(Subfamily 

Ceratopteridoideae) 

MAN 

Cheilanthes alpina Blume =Hypolepis alpina MAN/Knapp, R. 

2011 

Cheilanthes contiguum Wall. 

(nom. nud.) 

=Onychium lucidum FOT 2003/MAN 

--

Cheilanthes dealbata D. Don =Cheilanthes 

MAN 

formosana 

Cheilanthes farinosa auct. non =Cheilanthes agetae in MAN 

(Forssk.) Kaulf.: FOT 1975 part 

Cheilanthes farinosa auct. non =Cheilanthes 

Saiki 1984/MAN 

(Forssk.) Kaulf.: FOT 2003 formosana in part 

Cheilanthes henryi Christ =Cheilanthes nitidula MAN 

Cheilanthes hirsuta (Poir.) =Cheilanthes 

MAN 

Mett. 

nudiuscula 

Cheilanthes mexicana Davenp. =Cheilanthes krameri Knapp, R. 2011 

Cheilanthes mysurensis auct. =Cheilanthes chusana MAN 

non Wall. ex Hook.: FOT 

1975 

Cheilantes pallida Blume =Hypolepis pallida MAN/Knapp, R. 

2013 

Cheilanthes polypodioides 

Blume 

=Hypolepis 

polypodioides 


2011 

Cheilosoria chusana (Hook.) =Cheilanthes chusana MAN 

Ching 

Cheiropleuria bicuspis auct. 

non. (Blume) C. Presl: FOT 

1975 


integrifolia 

MAN 

Cheiropleuria bicuspis f. 

integrifolia (D. C. Eaton ex 

Hook.) D. C. Eaton ex 

Luerss. 

Cheiropleuria bicuspis var. 

integrifolia (D. C. Eaton ex 

Hook.) D. C. Eaton ex 

Matsum. & Hayata 

Chieniopteris harlandii (Hook.) 

Ching 

Chieniopteris kempii (Copel.) 

Ching 

Chingia ferox (Blume) Holtt. 


integrifolia 


integrifolia 

=Woodwardia harlandii 

=Woodwardia kempii 

=Cyclosorus ferox, 

unconfirmed status 


=Cyclosorus 

acuminatus 

=Cyclosorus ensifer 

MAN 

MAN 

MAN 

MAN 

Christella acuminata (Houtt.) H. 

MAN 

Lev. 

Christella acuminata var. 

MAN 

kuliangensis auct. non 

(Ching) C. M. Kuo: MAN 

Christella arida (D. Don) Holtt. =Cyclosorus aridus MAN 

Christella dentata (Forssk.) =Cyclosorus dentatus MAN 

Brownsey & Jermy 


211

212 

Christella ensifera (Tagawa) 

Holtt. ex C. M. Kuo 

Christella jaculosa (Christ) 

Holtt. 

Christella latipinna (Benth.) H. 

Lev. 

Christella papilio (C. Hope) 

Holtt. 

Christella parasitica (L.) H. Lev. 

Christella subarida (Tatew. & 

Tagawa) Holtt. ex C. M. Kuo 

CHRYSOGRAMMITIS 

Chrysogrammitis glandulosa 

(J. Sm.) Parris 

Cibotium cumingii auct. non 

Kunze: FOT 1975 

Clementea palmiformis Cav. 

Colysis decurrens (Blume) 

Manickam & Irudayaraj 

Colysis elliptica var. pothifolia 

auct. non (Buch.-Ham. ex D. 

Don) Ching: FOT 2003 

Colysis elliptica var. flexiloba 

auct. non (Christ) L. Shi & X. 

C. Zhang 

Colysis flexiloba auct. non 


Colysis longissima (Blume) J. 

Sm. 

Colysis membranacea (D. Don) 

J. Sm. 

Colysis pothifolia auct. non 

(Don) C. Presl: FOT 2003 

Colysis pothifolia var. 

multijugata H. Ito (type: 



=Cyclosorus jaculosus 

=Cyclosorus latipinnus 

=Cyclosorus papilio 

=Cyclosorus 

parasiticus 


=Grammitis, kept. for 

C. glandulosa 

=Grammitis glandulosa 

=Cibotium taiwanense 

=Angiopteris 

palmiformis 

=Leptochilus decurrens 

MAN 

MAN 

MAN 

MAN 

MAN 

MAN 

-- 


MAN 

MAN 

MAN 

=Colysis elliptica MAN/Knapp, R. 

2011 

?=Colysis elliptica (FOC) unresolved 

taxonomic 

situation of 

Colysis elliptica in 


?=Colysis elliptica (FOC) unresolved 

taxonomic 

situation of 

Colysis elliptica in 


=Phymatosorus MAN 

longissimus 

=Microsorum 

MAN 

membranaceum 

=Colysis elliptica Knapp, R. 2011 

? published in: 

Iconogr. Pl. As. 

Orient. 4(1): 330, 

tab. 110 (1941)

213 

Colysis pteropus (Blume) 

Bosman 

Colysis wrightii var. heteroclita 

Tagawa (type: Taiwan) 

Coniogramme parvipinnula 


Coniogramme taipeiensis 

Ching ex K. H. Shing (type: 


Coniogramme taiwanensis 



Cornopteris banahaoensis (C. 

Chr.) K. Iwats. & M. G. Price 

(misspelt species name) 

Cornopteris fluvialis (Hayata) 

Tagawa 

Cornopteris tenuisecta (Blume) 

Tardieu 

Craspedoneuron pallidum 

(Blume) Bosch 

Crepidomanes bilabiatum 

(Nees & Blume) Copel. 

Crepidomanes bipunctatum 

var. latealatum (Bosch) C. B. 

Clarke 

Crepidomanes birmanicum 

(Bedd.) K. Iwats. 

Crepidomanes digitatum (Sw.) 

K. Iwats. 

Crepidomanes grande (Copel.) 

Ebihara & K. Iwats. 

Crepidomanes insigne var. 

makinoi (C. Chr.) Sa. Kurata 

Crepidomanes makinoi (C. 

Chr.) Copel. 

Crepidomanes nanophyllum 


Crepidomanes pervenulosum 

auct. non (Alderw.) Copel. 

=Microsorum pteropus 

MAN 

? published in: Acta 


11: 308 (1942) 

=Coniogramme procera MAN 


in: Acta Bot. 

Yunnan. 3: 229 

(1981) 


in: Acta Bot. 

Yunnan. 3: 236 

=Cornopteris 

banajaoensis 

=Cornopteris 

banajaoensis 

=Acystopteris 

tenuisecta 

=Crepidomanes 

pallidum 

=Crepidomanes 

bipunctatum 

=Crepidomanes 

latealatum 

=Crepidomanes 

birmanicum (but 

partly misapplied to 

C. kalamocarpum) 

=Sphaerocionium 

digitatum 

=Cephalomanes 

grande 

=Crepidomanes 

palmifolium 

=Crepidomanes 

palmifolium 

=Crepidomanes kurzii 

=Crepidomanes 

parvifolium 

(1981) 


MAN 

MAN 

MAN 

MAN 

FOC 


Geobot. 60(1): 

26-40 (2009) 


2011 


2013 

MAN 

MAN 

MAN 

Knapp, R. 2011

214 

Crepidomanes radicans var. 

naseanum (H. Christ) K. 

Iwats. 

Crepidomanes schmidtianum 

var. latifrons (Bosch) K. 

Iwats. 

Crepidophyllum humile (G. 

Forst.) C. F. Reed 

Crepidopteris humilis (G. 

Forst.) Copel. 

Crypsinus echinosporus 

(Tagawa) Tagawa 

Crypsinus engleri (Luerss.) 

Copel. 

Crypsinus hastatus (Thunb.) 

Copel. 

Crypsinus okamotoi (Tagawa) 

Tagawa 

Crypsinus rhynchophyllus auct. 

non (Hook.) Copel.: MAN 

Crypsinus quasidivaricatus 


Crypsinus taeniatus var. 

palmatus auct. non (Blume) 

Tagawa: MAN 

Crypsinus taiwanensis 


Crypsinus veitchii auct. non 

(Baker) Copel.: FOT 1975 

Crypsinus yakushimensis 

(Makino) Tagawa 

Cryptosorus obliquata (Blume) 

J. Sm. 

Ctenitis apiciflora (Wall. ex 

Mett.) Ching 

Ctenitis aureovestita (Rosenst.) 

Ching 

Ctenitis boryana auct. non 

(Willd.) Copel.: FOT 2003 

Ctenitis dissecta (G. Forst.) H. 

Ito 

Ctenitis hendersonii (Bedd.) H. 

Ito 

Ctenitis kawakamii (Hayata) 

Ching 

=Crepidomanes 

birmanicum 

=Crepidomanes 

latifrons 

=Crepidomanes humile 




26-40 (2009) 

MAN 

MAN 

MAN 

=Selliguea 


echinosporus 2011 

=Selliguea engleri MAN/Knapp, R. 

2011 

=Selliguea hastatus MAN/Knapp, R. 

2011 

=Selliguea okamotoi MAN/Knapp, R. 

2011 

=Selliguea okamotoi Knapp, R. 2011 

=Selliguea 

quasidivaricata 

=Selliguea 

falcatopinnata 


2011 


=Selliguea taiwanensis MAN/Knapp, R. 

2011 

=Selliguea 

FOT 2003 


=Selliguea 


yakushimensis 2011 

=Grammitis obliquata FOT 2003 


=Dryopsis 


=Deparia subfluvialis 

=Tectaria dissecta 

=Nothoperanema 

hendersonii 

=Dryopsis kawakamii 

MAN 

MAN 

FOT 2003/MAN 

MAN 

MAN 

MAN

215 

Ctenitis kusukusensis (Hayata) =Tectaria kusukusensis MAN 

H. Ito 

Ctenitis leucostipes (Baker) =Ctenitis eatonii FOT 2003 

Tardieu 

Ctenitis mariformis (Rosenst.) =Dryopsis kawakamii FOT 2003 

Ching 

Ctenitis matsumurae (Makino) =Dryopsis 

FOT 2003 

Koidz. 


Ctenitis maximowicziana (Miq.) =Dryopis 

MAN 

Ching 


Ctenitis rhodolepis (C. B. =Ctenitis 

FOT 2003 

Clarke) Ching 

subglandulosa 

Ctenitis sacholepis H. Ito =Dryopsis kawakamii FOT 2003 

Ctenitis sasaki (Hayata) Ching =Tectaria dissecta MAN 

Ctenitis simozawae (Tagawa) =Lastreopsis tenera FOT 2003 

Ching 

Ctenitis subglandulosa var. =Ctenitis eatonii FOT 2003 

leucostipes (Baker) Shieh 

Ctenitis subtripinnata (Miq.) H. =Ctenitis 

FOT 2003 

Ito 

subglandulosa 

Ctenitis tenuifrons (Hayata) =Tectaria dissecta MAN 

Ching 

Ctenitis transmorrisonensis =Dryopsis 

MAN 


transmorrisonensis 

Ctenitis trichorachis (Hayata) =Nothoperanema MAN 

H. Ito 

squamiseta 

Ctenitopsis angustodissecta =Tectaria dissecta MAN 


Ctenitopsis devexa (Kunze ex =Tectaria devexa MAN 

Mett.) Ching & C. H. Wang 

Ctenitopsis dissecta (G. Forst.) =Tectaria dissecta MAN 

Ching 

Ctenitopsis fuscipes (Wall. ex =Tectaria fuscipes MAN 

Bedd.) Tardieu & C. Chr. 

Ctenitopsis kusukusensis =Tectaria kusukusensis MAN 

(Hayata) C. Chr. ex Tardieu & 

C. Chr. 

Ctenitopsis kusukusensis var. =Tectaria kusukusensis FOC 

crenato-lobata Tagawa (type: 


Ctenitopsis sasaki (Hayata) =Tectaria dissecta MAN 

Ching & C. H. Wang 

Ctenitopsis subfuscipes =Tectaria subfuscipes MAN 


CTENOPTERIS =Grammitis --

216 

Ctenopteris brevivenosa auct. 

non (Alderw.) Holtt. 

=Grammitis mollicoma Kuo, C. M. 

1985/Knapp, R. 

2011 

=Grammitis celebica FOC/Knapp, R. 

2011 

=Grammitis contigua FOC/Knapp, R. 

2011 

=Grammitis curtisii Knapp, R. 2011 

Ctenopteris celebica (Blume) 

Copel. 

Ctenopteris contigua (G. 

Forster) Holtt. 

Ctenopteris curtisii (Baker) 

Copel. 

Ctenopteris glandulosa J. Sm. =Grammitis glandulosa Kew Bull. 53(4): 

912-914 (1998) 

Ctenopteris hayatae (Masam.) =Grammitis doniana Gard. Bull. Sing. 

H. Ito 

58: 248 (2007)/ 


Ctenopteris merrittii (Copel.) 

Tagawa 

=Grammitis glandulosa Kew Bull. 53(4): 

912-914 (1998) 

Ctenopteris mollicoma (Nees & =Grammitis mollicoma Knapp, R. 2011 

Blume) Kunze 

Ctenopteris obliquata (G. =Grammitis obliquata MAN 

Forst.) Holtt. 

Ctenopteris subcorticola 



912-914 (1998) 

Ctenopteris subfalcata auct. =Grammitis doniana Knapp, R. 2011 

non (Blume) Kunze 

Ctenopteris tenuisecta (Blume) =Grammitis tenuisecta MAN 

J. Sm. 

Currania oyamensis (Baker) =Gymnocarpium MAN 

Copel. 

oyamense 

Cyathea formosana (Baker) =Cyathea metteniana MAN 

Copel. 

Cyathea fujiana (Nakai) =Cyathea fenicis MAN 

Tagawa 

Cyathea pustulosa (Christ) =Cyathea lepifera MAN 

Copel. 

Cyathea taiwaniana Nakai =Cyathea spinulosa MAN 

Cyclogramma auriculata (J. =Cyclosorus 

MAN 

Sm.) Ching 

auriculatus 

Cyclogramma leveillei (Christ) =Cyclosorus omeiensis FOC/FOT 2003 

Ching 

Cyclogramma omeiensis =Cyclosorus omeiensis MAN 

(Baker) Tagawa 

Cyclophorus adnascens (Sw.) =Pyrrosia lanceolata MAN 

Desv. 

Cyclophorus grandissimus 


=Pyrrosia sheareri MAN

Cyclophorus gralla (Giesenh.) =Pyrrosia porosa MAN 

C. Chr. 

Cyclophorus lanceolatus (L.) =Pyrrosia lanceolata MAN 

Alston 

Cyclophorus lingua (Thunb.) =Pyrrosia lingua MAN 

Desv. 

Cyclophorus lingua var. =Pyrrosia lingua MAN 

angustifrons Hayata (type: 


Cyclophorus lingua var. =Pyrrosia lingua MAN 

attenuata Rosenst. (type: 


Cyclophorus linearifolius Hook. =Pyrrosia linearifolia FOC 

Cyclophorus matsudai Hayata =Pyrrosia matsudae MAN 


Cyclophorus polydactylus =Pyrrosia polydactyla MAN 

(Hance) C. Chr. 

Cyclophorus porosus C. Presl =Pyrrosia porosa MAN 

Cyclophorus sasakii Hayata =Pyrrosia angustissima FOC 


Cyclophorus sheareri (Baker) =Pyrrosia angustissima MAN 

C. Chr. 

Cyclophorus subfissus Hayata =Pyrrosia porosa MAN 


Cyclophorus taiwanense =Pyrrosia lingua MAN 

(Christ) C. Chr. 

Cyclophorus 

transmorrisonensis Hayata 

=Pyrrosia porosa MAN/Knapp, R. 

2011 


Cyclosorus acuminatus f. =Cyclosorus ensifer MAN 

ensipinnus (Hayata) H. Ito 

Cyclosorus acuminatus f. 

pilosus H. Ito (type: Taiwan) 

Cyclosorus acuminatus var. 

kuliangensis auct. non Ching: 

MAN 

Cyclosorus caoshanensis 



Cyclosorus cuspidatus (Blume) 

C. M. Kuo 

=Cyclosorus 

acuminatus 


217 

(FOC) published 

in: Bot. Mag. 

(Tokyo) 51: 712 

(1937) 

MAN 



Popularis Sin. 

4(1): 321 (1999) 

=Cyclosorus liukiuensis Knapp, R. 2011

218 

Cyclosorus gaoxiongensis 



Cyclosorus gongylodes 

(Schkuhr) Link 

Cyclosorus kotoensis (Hayata) 

W. C. Shieh 

Cyclosorus kuliangensis auct. 

non (Ching) K. H. Shing: 

MAN 

Cyclosorus lanyuensis C. M. 

Kuo (nom. nud.) ("type": 


Cyclosorus oblancifolius 

Tagawa 

Cyclosorus orientalis C. M. Kuo 

(nom. nud.) 

Cyclosorus subaridus Tatewaki 

& Tagawa (type: Taiwan) 

Cyclosorus subnamburensis 



Cyclosorus subpubescens 

auct. non (Blume) Ching: 

FOT 2003 

Cyclosorus truncatus f. 

kwashotensis (Hayata) H. Ito 

Cyclosorus truncatus f. 

laevifrons (Hayata) H. Ito 

Cyclosorus truncatus var. 

kotoensis (Hayata) H. Ito 



=Cyclosorus interruptus FOT 2003 

=Cyclosorus productus 


?=Cyclosorus 

productus 

MAN 

MAN 

(MAN) 

=Cyclosorus dentatus FOT 2003 

? (MAN) 


MAN 



Popularis Sin. 

4(1): 341 (1999) 

=Cyclosorus jaculosus MAN 

=Cyclosorus truncatus 



MAN 

MAN 

MAN 

Cyrtogonellum caducum Ching =Polystichum tenuius FOC 

Cyrtogonellum falcilobum =Polystichum tenuius FOC 

Ching ex Y. T. Hsieh 

Cyrtogonellum fraxinellum =Polystichum 

MAN 


fraxinellum 

Cyrtogonellum salicifolium =Polystichum tenuius FOC 

Ching ex Y. T. Hsieh 

Cyrtogonellum tenuius Ching =Polystichum tenuius FOC 

Cyrtomidictyum lepidocaulon =Polystichum 

MAN 

(Hook.) Ching 

lepidocaulon 

CYRTOMIUM 

=Polystichum (kept for 

C. macrophyllum var. 

simadae) 

--

Cyrtomium caryotideum (Wall. 

ex Hook. & Grev.) C. Presl 

Cyrtomium devexiscapulae 

(Koidz.) Koidz. & Ching 

=Polystichum 

caryotideum 

=Polystichum 

devexiscapulae 

MAN 

219 

alternative 

combination in: 

Bull. Chin. Bot. 

Soc. 2: 96 (1936) 

MAN 

Cyrtomium falcatum (L. f.) C. =Polystichum falcatum 

Presl 

Cyrtomium falcatum subsp. =Polystichum falcatum Knapp, R. 2011 

australe Matsumoto (nom. 

nud.) 

Cyrtomium fortunei J. Sm. =Polystichum fortunei MAN 

Cyrtomium fraxinellum (Christ) =Polystichum 

MAN 

Christ 

fraxinellum 

Cyrtomium hookerianum (C. =Polystichum 

MAN 


integripinnum 

Cyrtomium integripinnum =Polystichum 

MAN 


integripinnum 

Cyrtomium longipes Ching & K. ?=Polystichum 


H. Shing 

taiwanianum 

Cyrtomium macrophyllum =Polystichum 

MAN 


macrophyllum 

Cyrtomium macrophyllum var. =Polystichum simadae MAN/Knapp, R. 

acuminatum (Diels) Tagawa: 

2013 

MAN, Knapp, R. 2011 

Cyrtomium macrophyllum var. 

simadae Tagawa (type: 


Cyrtomium microindusium Sa. 

Kurata 

Cyrtomium nervosum Ching & 

K. H. Shing 

Cyrtomium taiwanense 

Tagawa (mis-spelling of 


Cyrtomium taiwanianum 


Cystopteris formosana Hayata 


Cystopteris japonica auct. non 

Luerss.: FOT 1975 

Cystopteris japonica var. 

taiwaniana Tagawa (type: 


Cystopteris setosa Bedd. 

=Polystichum simadae Knapp, R. 2013 

=Polystichum 

anomophyllum 

=Polystichum 

anomophyllum 

=Polystichum 

taiwanianum 

=Polystichum 

taiwanianum 

=Acystopteris 

tenuisecta 

=Acystopteris 

taiwaniana 

=Acystopteris 

taiwaniana 

=Acystopteris 

tenuisecta 



2013 

MAN 

MAN 

MAN 

MAN 

MAN 

FOT 2003

220 

Cystopteris sphaerocarpa =Cystopteris 

FOT 2003 


moupinensis 

Cystopteris sudetica var. =Cystopteris 

FOT 2003 

moupinensis (Franch.) C. 

Chr. 

moupinensis 

Cystopteris tenuisecta (Blume) =Acystopteris 

MAN 

Mett. 

tenuisecta 

DASYGRAMMITIS 

=Grammitis, kept. for - 

D. mollicoma 

Dasygrammitis mollicoma =Grammitis mollicoma this publication 

(Nees & Blume) Parris 

Davallia biflora Kaulf. =Odontosoria biflora MAN 

Davallia biserrata Blume =Tapeinidium 

MAN 

biserratum 

Davallia bullata auct. non Wall. =Davallia 

FOT 2003/MAN 

trichomanoides 

Davallia calvescens Wall. ex =Microlepia calvescens MAN 

Hook. 

Davallia chinensis (L.) Sm. =Odontosoria chinensis MAN 

Davallia clarkei auct. non =Araiostegia 


Baker: MAN 

parvipinnula 

Davallia divaricata auct. non =Davallia formosana MAN 

Blume: FOT 1975 

Davallia dryopteridifrons 

Hayata 

=Leucostegia truncata MAN/Fraser- 


Davallia gracilis auct. non 

Blume 

=Tapeinidium 

biserratum 

2008 

Kuo, C. M. 


2011 

Davallia hirsuta Sw. =Dennstaedtia hirsuta MAN 

Davallia hookeriana Wall. ex =Microlepia hookeriana FOT 2003 

Hook. 

Davallia immersa Wall. (nom. 

nud.) 


MAN/Fraser- 


2008 

Davallia mariesii T. Moore ex =Davallia 

MAN 

Baker 


Davallia mariesii var. 

=Davallia 

MAN 

stenolepis (Hayata) Hoshiz. trichomanoides 

Davallia multiflora Roxb. =Nephrolepis brownii MAN/Knapp, R. 

2011 

Davallia orientalis C. Chr. apud =Davallia formosana FOT 2003 

Wu 

Davallia parallela Wall. ex 

Hook. 

=Davallia pectinata MAN

Davallia parvipinnula Hayata =Araiostegia 

MAN 


parvipinnula 

Davallia pedata Sm. =Davallia repens MAN 

Davallia perdurans Christ =Araiostegia 

MAN 

parvipinnula 

Davallia pinnata Cav. =Tapeinidium pinnatum MAN 

Davallia platyphylla D. Don =Microlepia obtusiloba MAN 

Davallia pulcherrima Baker =Asplenium 

MAN 

pulcherrimum 

Davallia rhomboidea Wall. ex =Microlepia 

MAN 

Kunze 

rhomboidea 

Davallia speluncae (L.) Baker =Microlepia speluncae this publication 

Davallia stenolepis Hayata =Davallia 

MAN 



Davallia stipellata Wall. (nom. =Acrophorus paleolatus MAN 

nud.) 

Davallia strigosa var. 

rhomboidea (Wall. ex Kunze) 

Hook. & Baker 

=Microlepia 

rhomboidea 

MAN 

Davallia subalpina Hayata 


221 

? published in: J. 

Coll. Sci. Imp. 

Univ. Tokyo 30. 

432 (1911) 

Davallia trapeziformis 

=Microlepia speluncae Moore, S. J. 2010 

(Roxburgh) Kuhn 

Davallia truncata D. Don =Leucostegia truncata Fraser-Jenkins, 

C. R. 2008 

Dennstaedtia leptophylla =Dennstaedtia smithii MAN 


Dennstaedtia strigosa (Thunb.) =Microlepia strigosa MAN 

J. Sm. 

Deparia boryana auct. non 

(Willd.) M. Kato: Kuo, C. M. 

2010 

=Deparia subfluvialis this publication 

Deparia formosana (Rosenst.) 

R. Sano 

Deparia orientalis auct. non (?) 

(Z. R. Wang & J. J. Chien) 

Naikaike 

=Dictyodroma 


formosana 

2013 

=Deparia sp. 1 Knapp, R. 2011 

Diacalpe aspidioides Blume =Peranema aspidioides MAN 

Dicksonia scabra Wall. ex =Dennstaedtia scabra MAN 

Hook. 

Dicksonia scandens Blume =Dennstaedtia 

MAN 

scandens 

Dicksonia smithii Hook. =Dennstaedtia smithii MAN

222 

Dicranopteris blotiana (C. Chr.) 

Tagawa 

Dicranopteris chinensis 

(Rosenst.) Tagawa 

Dicranopteris dichotoma 

(Thunb.) Bernh. 

Dicranopteris glauca (Thunb. 

ex Houtt.) Underw. 

Dicranopteris laevissima 

(Christ) Nakai 

Dicranopteris linearis var. 

montana Holtt. 


subpectinata (Christ) Holtt. 


tetraphylla (Rosenst.) Nakai 

Dicranopteris splendida auct. 

non Hand.-Mazz. 

Dicranopteris warburgii auct. 

non Christ 

DICTYOCLINE 

Dictyocline griffithii T. Moore 

var. griffithii 

Dictyocline griffithii var. 

pinnatifida (Hook.) Bedd. 

Dictyocline griffithii var. wilfordii 


Dictyocline wilfordii (Hook.) J. 

Sm. 

Didymoglossum affine Bosch 

Didymoglossum bipunctatum 

(Poir.) E. Fourn. 

Didymoglossum denticulatum 

(Sw.) Hassk. 

Didymoglossum humile (G. 

Forst.) C. Presl 

Didymoglossum latealatum 

Bosch 

Didymoglossum longisetum C. 

Presl 

Didymoglossum simonsianum 

(Hook.) Bosch 

Didyocionium holochilum 

Bosch 

=Diplopterygium MAN 

blotianum 

=Diplopterygium MAN 

chinensis 

=Dicranopteris linearis FOT 2003 

=Diplopterygium 

glaucum 


laevissimum 

=Dicranopteris 

taiwanensis 


subpectinata 


tetraphylla 


tetraphylla 


subpectinata 

=Cyclosorus, kept for 

D. wilfordii 

=Cyclosorus griffithii 

MAN 

MAN 

Kuo, C. M. 1985 

MAN 

MAN 

FOT 2003 

FOT 2003 

-- 

MAN 

=Cyclosorus griffithii FOT 2003 

=Cyclosorus wilfordii MAN/Knapp, R. 

2011 

=Cyclosorus wilfordii MAN/Knapp, R. 

2011 

=Hymenophyllum FOT 2003 

holochilum 

=Crepidomanes MAN 

bipunctatum 

=Hymenophyllum MAN 

denticulatum 

=Crepidomanes humile MAN 

=Crepidomanes 

latealatum 

=Cephalomanes 

obscurum 

=Hymenophyllum 

simonsianum 


holochilum 

MAN 

FOT 2003 

MAN 

MAN

223 

Diphasiastrum veitchii (Christ) =Lycopodium veitchii MAN 

Holub 

Diphasium multispicatum (J. H. =Lycopodium 

MAN 

Wilce) Rothm. 

multispicatum 

Diplaziopsis brunoniana (Wall. 

ex Bedd.) W. M. Chu 

=Diplaziopsis javanica Christenhusz et 

al. 2011 

Diplaziopsis formosana 

(Rosenst.) M. G. Price 

=Dictyodroma 

formosana 


2011 

Diplazium agyokuense Tagawa =Diplazium latifrons this publication 


Diplazium allantodioides Ching =Diplazium virescens FOT 2003 

Diplazium amamianum ?=Diplazium laxifrons Knapp, R. 2011 

Tagawa 

Diplazium arisanense Hayata 



2009 

Diplazium asperum Blume =Diplazium sikkimense Knapp, R. 2013 

Diplazium bantamense f. 

serratum Tagawa (type: 


? published in: Acta 


2: 199 (1933) 

Diplazium bicuspe Hayata =Asplenium ensiforme MAN 


Diplazium costalisorum Hayata =Diplazium doederleinii MAN 


Diplazium crenato-serratum =Diplazium pullingeri MAN 

var. hirta Rosenst. (type: 


Diplazium decurrenti-alatum =Cornopteris 

MAN 

(Hook.) C. Chr. 

Diplazium donianum var. 

aphanoneuron (Ohwi) 

Tagawa 


=Diplazium 

aphanoneuron 

alternative 



Geobot. 20: 215 

(1962) 

MAN 

Diplazium donianum var. =Diplazium lobatum 

lobatum Tagawa 

Diplazium epirachis Christ =Athyrium epirachis Liu, Y. C. et al. 

2009 

Diplazium fauriei Christ =Diplazium 

MAN 

mettenianum 

Diplazium formosanum =Dictyodroma 


Rosenst. (type: Taiwan) formosana 

2011 

Diplazium fraxineum D. Don =Coniogramme MAN 

fraxinea 

Diplazium fraxinifolium auct. =Diplazium donianum FOT 2003 

non C. Presl.

224 

Diplazium griffithii auct. non T. 

Moore 

Diplazium heterophlebium 

auct. non (Mett.) Diels. 

Diplazium hookerianum Koidz. 

Diplazium inflatisorum Hayata 


Diplazium iridophyllum 


Diplazium japonicum auct. non 

(Thunb.) Bedd. 

Diplazium kappanensis Hayata 


Diplazium lanceum (Thunb.) C. 

Presl 

Diplazium lanceum var. 

crenatum Makino 

Diplazium latifolium var. 

cyclolobum Christ (type: 


Diplazium leiopodum Hayata 


Diplazium macrophyllum Ching 

Diplazium makinoi Yabe ex 

Masam. & Hayata 

Diplazium makinoi var. 

karapinense Hayata (type: 


Diplazium makinoi var. 

multicaudatum M. Ogata 


Diplazium maximum var. 

formosanum Rosenst. (type: 


Diplazium morii Hayata (type: 


Diplazium muricatum auct. non 

(Mett.) Alderw.: MAN 

=Diplazium latifrons (Kuo, C. M. 2010) 


=Dictyodroma 

FOT 2003 

formosana 

=Cornopteris 

FOT 2003 


=Diplazium doederleinii MAN 

=Asplenium 

MAN 

griffithianum 

=Deparia petersenii Knapp, R. 2011 

?=Diplazium 

taiwanense 

=Deparia lancea 

FOC (if both taxa 

are identical, D. 

kappanensis 

replaces D. 

taiwanense as 

being older) 

MAN 

=Deparia lancea MAN/Knapp, R. 

2011 

=Diplazium dilatatum FOC/Knapp, R. 

2011 

=Athyrium leiopodum 

=Diplazium 

megaphyllum 

=Asplenium formosae 

=Asplenium formosae 


2009 

FOT 2003 

MAN 

MAN 


Journ. Jap. Bot. 

11: 32, fig. 3 

(1935) 

=Diplazium latifrons MAN 

=Diplazium doederleinii 

MAN 

=Diplazium kawakamii Knapp, R. 2011

Diplazium odoratissimum 


=Dictyodroma 

formosana 


2013 

Diplazium opacum (D. Don) =Cornopteris opaca MAN 

Christ 

Diplazium petersenii (Kunze) =Deparia petersenii MAN 

Christ 

Diplazium petrii Tardieu =Diplazium latifrons MAN 

Diplazium phaeolepis Tagawa =Diplazium 

Kuo, C. M. 1985 



Diplazium polypodioides auct. =Diplazium sikkimense Knapp, R. 2011 

non Blume: MAN 

Diplazium pulchrum Tagawa =Athyrium pulchrum MAN 


Diplazium squamigerum (Mett.) 

Christ 

=Diplazium 

squamigerum (Mett.) 


Diplazium squamigerum (Wall. 

ex Mett.) C. Hope 

Matsum. 

=Diplazium 

squamigerum (Mett.) 

Matsum. 


Diplazium subrigescens 


=Athyrium delavayi var. 

subrigescens 


2009 

Diplazium subsinuatum (Wall. =Deparia lancea MAN 

ex Hook. & Grev.) Tagawa 

Diplazium tenuicaule Hayata 


=Athyrium leiopodum Liu, Y. C. et al. 

2009 

Diplazium textorii Miq. 

=Diplazium 

FOC 

mettenianum 

Diplazium triangulare Tagawa =Diplazium latifrons FOC/MAN 

Diplazium uraiense Rosenst. =Diplazium dilatatum Knapp, R. 2011 


Diplazium virescens var. 

okinawaense (Tagawa) Sa. 

Kurata 

=Diplazium 

okinawaense 

MAN 

Diploblechnum fraseri (A. 

Cunn.) DeVol 

Diploblechnum integripinnulum 


Doryopteris concolor (Langsd. 

& Fisch.) Kuhn 

Drymoglossum carnosum auct. 

non J. Sm. ex Hook.: MAN 

Drymoglossum carnosum var. 

obovatum Harr. (type: 


Drymoglossum microphyllum 

(C. Presl) C. Chr. 

=Blechnum fraseri 

=Blechnum fraseri 

=Cheilanthes concolor 

=Lemmaphyllum 

microphyllum 

=Lemmaphyllum 

microphyllum 

=Lemmaphyllum 

microphyllum 

MAN 

MAN 

MAN 


2011 


2011 

FOC 

225

226 

Drymoglossum nobukoanum 

Makino (type: Taiwan) 

=Lemmaphyllum 

microphyllum 


2011 

Drymoglossum obovatum 

(Harr.) Christ 

=Lemmaphyllum 

microphyllum 


2011 

Drymotaenium nakaii Hayata =Drymotaenium MAN 


miyoshianum 

Drynaria coronans (Wall. ex =Aglaomorpha 

MAN 

Mett.) J. Sm. ex T. Moore coronans 

Drynaria roosii T. Nakaike =Drynaria fortunei (Kuo 2001) 


(based on FOC is 

D. fortunei a later 

homonym of D. 

roosii) 

Dryoathyrium boryanum auct. =Deparia subfluvialis MAN 

non (Willd.) Ching: FOT 1975 

Dryoathyrium unifurcatum =Deparia unifurcata MAN 

(Baker) Ching 

Dryomenis kwarenkoensis =Tectaria simonsii FOT 2003 

(Hayata) Nakai 

Dryopteris abricana auct. non =Cyclosorus tottoides FOT 2003 

(Desv.) C. Chr. 

Dryopteris adaucta Rosenst. ? (FOC) published 


341 (1915) 

Dryopteris adiantoides T. =Dryopteris hasseltii MAN 

Suzuki (type: Taiwan) 

Dryopteris angustodissecta =Tectaria dissecta MAN 


Dryopteris anastomosans 


=Tectaria griffithii FOT 2003/this 

publication 

Dryopteris angustodissecta =Tectaria dissecta FOT 2003 


Dryopteris apiciflora (Wall. ex =Dryopsis apiciflora MAN 

Mett.) 

Dryopteris arida (Don) O. =Cyclosorus aridus FOT 2003 

Kuntze 

Dryopteris arisanenses 




340 (1915) 

Dryopteris aristata (G. Forst.) =Arachniodes aristata MAN/Knapp, R. 

Kuntze 

Dryopteris athyriiformis 


=Cornopteris 

banajaoensis 

2011 

MAN

Dryopteris atrata (Kunze) 

Ching 

Dryopteris atrosetosa Rosenst. 


Dryopteris aureovestita 


Dryopteris auriculata (L.) 

Kuntze 

Dryopteris austriaca auct. non 

(Jacq.) Woyn. ex Schinz & 

Thell.: MAN 

Dryopteris banajaoensis C. 

Chr. 

Dryopteris bankinsinensis 


Dryopteris barbigera auct. non 

(T. Moore ex Hook.) Kuntze: 

MAN 

Dryopteris barbigera subsp. 

komarovii (Kossinsky) Fraser- 

Jenk. 

Dryopteris beddomei (Baker) 

O. Kuntze 

=Dryopteris atrata, but 

partly misapplied for: 

Dryopteris cycadina 

(e. g. in FOT 1975), 

Dryopteris stenolepis 

(e. g. in MAN) 


squamiseta 

=Dryopsis 


=Nephrolepis cordifolia 


MAN 

MAN 

MAN 

=Dryopteris expansa Knapp, R. 2011 

=Cornopteris 

banajaoensis 

?=Cornopteris opaca 

MAN 

227 

published in: Icon. 


146, f. 73-74 

(1919) 

=Dryopteris komarovii Knapp, R. 2011 

=Dryopteris komarovii 

MAN 

=Thelypteris beddomei FOT 2003 

Dryopteris bipinnata C. Chr. =Dryopteris fuscipes FOC 

Dryopteris castanea Tagawa =Thelypteris japonica MAN 


Dryopteris chapaensis C. Chr. =Dryopteris polita FOT 2003 

& Ching 

Dryopteris chrysocoma auct. 

non (Christ) C. Chr.: FOT 

2003 

=Dryopteris 

woodsiisora 

MAN 

Dryopteris cnemidaria Christ =Pteridrys cnemidaria FOT 2003 

Dryopteris crenata (Forssk.) =Hypodematium MAN 

Kuntze 

crenatum 

Dryopteris cyrtolepis Hayata =Dryopteris wallichiana MAN 



Dryopteris decursive-pinnata =Phegopteris 

FOT 2003 

(van Hall) O. Kuntze 

decursivepinnata 

Dryopteris dilatata var. 

deltoidea (Milde) Takeda 

=Dryopteris expansa FOT 2003

228 

Dryopteris dissecta (Forst.) O. =Tectaria dissecta FOT 2003 

Kuntze 

Dryopteris doiana Ching (nom. =Dryopteris wallichiana FOC 

illeg.) 

Dryopteris eatonii (Baker) =Ctenitis eatonii MAN 

Kuntze 

Dryopteris enneaphylla var. 

pseudosieboldii (Hayata) 

=Dryopteris 

pseudosieboldii 


R. 2011 

Tagawa & Iwats. 

Dryopteris ensifera Tagawa =Cyclosorus ensifer MAN 


Dryopteris erubescens (Wall. =Cyclosorus 

FOT 2003 

ex Hook.) C. Chr. 

erubescens 

Dryopteris erythrosora var. =Dryopteris tenuipes MAN 

tenuipes Rosenst. (type: 


Dryopteris esquirolii Christ =Cyclosorus esquirolii MAN 

Dryopteris fibrillosa (C. B. =Dryopteris edwardsii Knapp, R. 2011 

Clarke) Hand.-Mazz. 

in part, =Dryopteris 

zayuensis in part, 

=Dryopteris 

xanthomelas in part 

Dryopteris filix-mas var. =Dryopteris 

MAN 

serrato-dentata Bedd. 

serratodentata 

Dryopteris fluvialis Hayata =Cornopteris 

MAN 


banajaoensis 

Dryopteris glanduligera =Thelypteris 

FOT 2003 

(Kunze) Chist 

glanduligera 

Dryopteris gongylodes 


(Schkuhr) O. Kuntze 

Dryopteris gracilescens =Thelypteris 

FOT 2003 

(Blume) O. Kuntze 

Dryopteris gracilescens auct. 

non (Blume) O. Kuntze: 

Makino 

Dryopteris gracilescens var. 

glanduligena (Kunze) Makino 

& Nemoto 

Dryopteris grandissima 


Dryopteris gymnopteridifrons 


Dryopteris gymnosora var. 

indusiata (Makino) Makino ex 

Bonap. 

gracilescens 

=Thelypteris 

angustifrons 

=Thelypteris 

glanduligera 

FOT 2003 

FOT 2003 

=Dryopteris marginata FOT 2003 

=Cyclosorus 

MAN 


=Dryopteris tenuicula FOT 2003

Dryopteris hancockii (Copel.) =Cyathea hancockii MAN 

Nakai 

Dryopteris hayatae Tagawa =Dryopteris subexaltata FOC 

Dryopteris hendersonii (Bedd.) =Nothoperanema MAN 

C. Chr. 

hendersonii 

Dryopteris hirsutisquama =Thelypteris uraiensis FOT 2003 


Dryopteris hirtipes auct. non 

Kuntze 

=Dryopteris atrata; 

partly misapplied for: 

Dryopteris cycadina, 

D. stenolepis 

229 


R. 2011 

Dryopteris hirtirachis C. Chr. =Pseudophegopteris MAN 

hirtirachis 

Dryopteris hypophlebia Hayata =Dryopteris fructuosa MAN 


Dryopteris inaensis Tagawa =Polystichum capillipes MAN/Knapp, R. 

2011 

Dryopteris indusiata (Makino) =Dryopteris tenuicula MAN 

Makino & Yamam. 

Dryopteris indusiata var. 

yoshimatsuana Suzuki (type: 




11: 647, fig. 3 

Dryopteris jaculosa (Christ) C. 

Chr. 

Dryopteris kawakamii Hayata 


Dryopteris kodamai Hayata 


Dryopteris kotoensis Hayata 


Dryopteris kusukusensis 


Dryopteris kwashotensis 


Dryopteris labordei auct. non 

(Christ) C. Chr.: FOT 2003 

Dryopteris lachoongensis 

(Bedd.) B. K. Nayar & S. Kaur 

Dryopteris laevifrons Hayata 


Dryopteris laevifrons var. 

kwashotensis (Hayata) 

Tagawa 

Dryopteris lasiocarpa Hayata 




=Dryopteris formosana 


=Tectaria kusukusensis 


=Dryopteris tenuicula 

=Dryopteris fructuosa 


(1935) 

FOT 2003/MAN 

MAN 

FOC 

MAN 

MAN 

MAN 

MAN 


C. R. 2008 

MAN 

=Cyclosorus truncatus FOT 2003 

=Macrothelypteris 

torresiana 

FOT 2003

230 

Dryopteris latipinna (Bentham) =Cyclosorus latipinnus FOC 

Kuntze 

Dryopteris laurisilvicola T. =Dryopteris polita MAN 

Suzuki 

Dryopteris laxa (Franch. & =Thelypteris laxa FOT 2003 

Sav.) C. Chr. 

Dryopteris lepidocaulis (Hook.) =Polystichum 

MAN 

Kuntze 

lepidocaulon 

Dryopteris lepigera auct. non =Ctenitis 

FOT 2003 

(Baker) O. Kuntze 

subglandulosa 

Dryopteris leptorhachia Hayata 



in: Icon. Pl. 

Formosan. 4: 

162, f. 102 (1914) 

Dryopteris leucostipes (Baker) =Ctenitis eatonii MAN 

C. Chr. 

Dryopteris leveillei Christ =Cyclosorus omeiensis FOT 2003 

Dryopteris liukiuensis (Christ) 

C. Chr. 


2011 

Dryopteris mariformis Rosenst. =Dryopsis kawakamii FOT 2003 

Dryopteris matsumurae =Dryopsis 

FOT 2003 

(Makino) C. Chr. 


Dryopteris maximowicziana =Ctenitis 

FOT 2003 

Koidz. 

subglandulosa 

Dryopteris maximowicziana =Dryopsis 

MAN 

(Miq.) C. Chr. 


Dryopteris membranoides =Tectaria kusukusensis FOC 


Dryopteris mingetsuensis =Dryopteris 

FOT 2003 


reflexosquamata 

Dryopteris morrisonensis =Dryopteris 

FOT 2003 



Dryopteris nigra Ching =Dryopteris lepidopoda FOT 2003 

Dryopteris nigrisquama Hayata =Dryopteris scottii MAN 


Dryopteris nigrisquama f. =Dryopteris scottii MAN 

subdecipiens (Hayata) H. Ito 

Dryopteris nokoensis Tagawa =Dryopsis 

FOT 2003 



Dryopteris oblancifolia Tagawa =Cyclosorus dentatus FOT 2003 


Dryopteris ochthodes auct. non =Cyclosorus esquirolii FOT 2003 

(Kunze) C. Chr. 

Dryopteris oligophlebia (Baker) 

C. Chr. 


torresiana 

FOT 2003

Dryopteris opaca (D. Don) C. =Cornopteris opaca MAN 

Chr. 

Dryopteris oppositipenna 


=Athyrium 



2009 

Dryopteris oyamensis (Baker) =Gymnocarpium MAN 

C. Chr. 

oyamense 

Dryopteris pachyphylla Hayata =Dryopteris wallichiana MAN 



Dryopteris paleacea auct. non =Dryopteris wallichiana FOT 2003 

(Sw.) C. Chr. 

Dryopteris paleacea var. =Dryopteris lepidopoda FOT 2003 

khasiana (C. B. Clarke) C. 

Chr. 

Dryopteris parasitica (L.) =Cyclosorus 

MAN 

Kuntze 

parasiticus 

Dryopteris parasitica var. =Cyclosorus latipinnus FOC 

latipinna (Bentham) C. Chr. 

Dryopteris patens (Sw.) O. =Cyclosorus 

FOT 2003 

Kuntze 

parasiticus 

Dryopteris peninsulae 

=Dryopteris lacera Knapp, R. 2011 

Kitagawa 

Dryopteris phaeolepis Hayata =Dryopteris formosana FOC 


Dryopteris phegopteris (L.) C. =Phegopteris 

FOT 2003 

Chr. 

connectilis 

Dryopteris polymorpha var. =Tectaria polymorpha FOT 2003 

pentaphylla (Hook.) Nakai 

Dryopteris prescottianum =Polystichum 

MAN 

Kuntze 

prescottianum 

Dryopteris prolifera (Retz.) C. =Cyclosorus proliferus this publication 

Chr. 

Dryopteris pseudo-africana =Cyclosorus omeiensis FOT 2003 

Makino & Ogata 

Dryopteris pseudosabaei =Dryopteris fructuosa MAN 


Dryopteris psilosora Tagawa =Dryopteris hasseltii MAN 

Dryopteris quadripinnata =Arachniodes 

MAN 


quadripinnata 

Dryopteris reflexipinna Hayata =Dryopteris diffracta MAN 


Dryopteris remota Hayata =Gymnocarpium MAN 


remote-pinnatum 

Dryopteris remotipinnata =Gymnocarpium MAN 


remote-pinnatum 

Dryopteris rufinervis Hayata 


=Pleocnemia rufinervis MAN 

231

232 

Dryopteris sacholepis Hayata 


Dryopteris sasaki Hayata (type: 


Dryopteris sieboldii var. 

heteroneura Tagawa (type: 


Dryopteris simozawae Tagawa 


Dryopteris sinofibrillosa Ching 

Dryopteris somai Hayata (type: 


Dryopteris sophoroides f. 

ensipinna Hayata (type: 


Dryopteris spinulosa var. 

morrisonensis Hayata (type: 


Dryopteris splendens var. 

formosana Rosenst. (type: 


Dryopteris squamaestipes 

auct. non C. Chr. 

Dryopteris squamiseta (Hook.) 

Kuntze 

Dryopteris subaurita Tagawa 


Dryopteris subdecipiens 


Dryopteris subfluvialis Hayata 


Dryopteris subglandulosa 

(Hance) Hayata 

Dryopteris subhispidula 



=Tectaria dissecta 

=Dryopteris 

enneaphylla 

MAN 

MAN 

MAN 

=Lastreopsis tenera FOT 2003 

=Dryopteris edwardsii Knapp, R. 2011 



=Dryopteris 


? published in: Icon. 


287, fig. 112 

=Cyclosorus 

acuminatus 

=Dryopteris expansa 

(1915) 

MAN 

FOC 


in: Hedwigia 

56(5): 343 (1915) 

=Cyclosorus 

FOT 2003/MAN 

auriculatus 

=Nothoperanema MAN 

squamiseta 

=Pseudophegopteris MAN 

subaurita 

=Dryopteris scottii MAN 

=Deparia subfluvialis 

=Ctenitis 

subglandulosa 

?=Cyclosorus 

taiwanensis 

MAN 

MAN 

Dryopteris subintegriloba Seriz. =Dryopteris integriloba MAN 

Dryopteris sublaevifrons ?=Cyclosorus truncatus 


(MAN) published 


342 (1915) 



5: 192 (1936)

Dryopteris sublaxa Hayata 


233 


in: Icon. Pl. 

Formosan. 4: 

183, f. 121 (1914) 

=Ctenitis 


Dryopteris subtripinnata (Miq.) 

Kuntze 

subglandulosa 

Dryopteris subtripinnata var. ? (MAN) published 

bunkikiyensis Rosenst. (type: 



342 (1915) 

Dryopteris subtripinnata var. =Ctenitis eatonii MAN 

leucostipes (Baker) H. Ito 

Dryopteris succulentipes =Cornopteris opaca FOC 


Dryopteris taitoensis Tagawa =Dryopteris polita FOT 2003 


Dryopteris taitungensis Koidz. =Cyathea hancockii MAN 


Dryopteris taiwanensis C. Chr. =Cyclosorus 

MAN 


taiwanensis 

Dryopteris taiwanicola Tagawa =Dryopteris lepidopoda MAN 


Dryopteris takeoi Hayata (type: =Athyrium mupinense Liu, Y. C. et al. 


2009 

Dryopteris tenuifrons Hayata =Tectaria dissecta MAN 


Dryopteris thomsonii (Hook. f.) =Polystichum thomsonii MAN 

Kuntze 

Dryopteris thrichorhachis =Nothoperanema MAN 


squamiseta 

Dryopteris thysanocarpa =Athyrium anisopterum Liu, Y. C. et al. 


2009 

Dryopteris todayensis auct. =Cyclosorus 

FOT 2003 

non Christ 

taiwanensis 

Dryopteris tosensis Kodama =Deparia unifurcata FOT 2003 

Dryopteris transmorrisonensis =Dryopsis 

MAN 



Dryopteris triphylla (Sw.) C. =Cyclosorus triphyllus FOT 2003 

Chr. 

Dryopteris truncata (Poir.) O. =Cyclosorus truncatus FOT 2003 

Kuntze 

Dryopteris unifurcata (Baker) =Deparia unifurcata MAN 

C. Chr. 

Dryopteris uraiensis Rosenst. =Thelypteris uraiensis MAN 


Dryopteris urophylla auct. non =Cyclosorus 

FOT 2003/MAN 

(Wall.) C. Chr. 

gymnopteridifrons

234 

Dryopteris uropinna Price 

Dryopteris ursipes Hayata 


Dryopteris yabei Hayata (type: 


Egenolfia appendiculata 

(Willd.) J. Sm. 

Egenolfia laxireticulata (K. 

Iwats.) C. M. Kuo 

Egenolfia rhizophylla (Kaulf.) 

Fee 

Egenolfia serrulata (J. Sm.) 

Fee 

Elaphoglossum austro-sinicum 

Matthew & Christ 

Elaphoglossum conforme auct. 

non (Sw.) Schott: FOT 1975 

Elaphoglossum 

fuscopunctatum Christ 

Elaphoglossum latifolium auct. 

non (?) (Sw.) J. Sm. 

Elaphoglossum laurifolium 

auct. non (Thouars) T. Moore 

Elaphoglossum lepidopodum 

C. Chr. ex Ogata (Type: 


Elaphoglossum ogatae C. Chr. 

ex M. Ogata (type: Taiwan) 

Elaphoglossum pendulifolium 


Elaphoglossum subellipticum 


Equisetum debile Roxb. ex 

Vaucher 

Equisetum ramosissimum var. 

taikankoense Yamamoto 

=Dryopteris 

FOC 

subtriangularis 

=Dryopteris wallichiana MAN 


=Dryopteris varia FOT 2003 

=Bolbitis appendiculata 

=Bolbitis x laxireticulata 

=Bolbitis rhizophylla 

MAN 

MAN 

MAN 

=Bolbitis rhizophylla FOT 2003 


yoshinagae 


marginatum 


marginatum 


callifolium 


angulatum 


luzonicum 


angulatum 


marginatum 


commutatum 

=Equisetum 

ramosissimum 

subsp. debile 

=Equisetum 

ramosissimum 

subsp. 

ramosissimum 

FOT 2003 

MAN 

FOC 

Knapp, R. 2011; 

follow-up is 

needed as FOC is 

providing a 

reference to 

Elaphoglossum 

luzonicum 

FOT 2003 

MAN 

MAN 

MAN 

MAN 

MAN 

MAN

235 

Glaphyropteridopsis 

erubescens (Wall. ex Hook.) 

Ching 

Glaphyropteris falciloba 

(Hook.) H. Ito 

Glaphyropteris omeiensis 

(Baker) H. Ito 

Gleichenia blotiana C. Chr. 

Gleichenia chinensis Rosenst. 

Gleichenia glauca (Thunb. ex 

Houtt.) Hook. 

Gleichenia kiusiana Makino 

Gleichenia laevissima Christ 

Gleichenia linearis (Burm. f.) C. 

B. Clarke 

Gleichenia linearis var. 

alternans Mett. 

Gleichenia linearis var. 

tetraphylla Rosenst. 

Gleichenia subpectinata Christ 

Goniophlebium amoenum 

(Wall. ex Mett.) Bedd. 

Goniophlebium amoenum var. 

arisanense (Hayata) Rodl- 

Linder 

Goniophlebium argutum auct. 

non (Wall. ex Hook.) J. Sm.: 

FOT 1975 

Goniophlebium caudiceps T. 

Moore (type: Taiwan) 

Goniophlebium formosanum 

(Baker) Rodl-Linder 

Goniophlebium hendersonii 

auct. non Bedd. 

Goniophlebium mengtzeense 

(Christ) Rodl-Linder 

Goniophlebium microrhizoma 

(C. B. Clarke ex Baker) C. B. 

Clarke ex Bedd. 

=Cyclosorus 

erubescens 

MAN 

=Cyclosorus esquirolii FOT 2003 

=Cyclosorus omeiensis 


blotianum 


chinensis 


glaucum 


laevissimum 


laevissimum 

=Dicranopteris linearis 


subpectinata 


tetraphylla 


subpectinata 

=Polypodium amoenum 

=Polypodium amoenum 

=Polypodium 

mengtzeense 

=Lepisorus 

obscurevenulosus 

=Polypodium 

formosanum 

=Polypodium 

microrhizoma 

=Polypodium 

mengtzeense 

=Polypodium 

microrhizoma 

MAN 

MAN 

MAN 

MAN 

MAN 

FOC 

MAN 

FOT 2003/MAN 

MAN 

MAN 

alternative 


Gen. Fil. t. 51 

(1840) 

MAN 

MAN 

MAN 

MAN 

MAN 

MAN 

MAN

236 

Goniophlebium niponicum 

(Mett.) Bedd. 

=Polypodium 

raishaense in part, 

=Polypodium 

transpianense in part 

=Polypodium 


and Knapp, R. 

2013 

Goniophlebium persicifolium 


(Desv.) Bedd. 

persicifolium 

Goniophlebium raishaense =Polypodium 

alternative 

(Rosenst.) C. M. Kuo (?nom. raishaense 

combination, ?not 

nud.): Kuo, C. M. 2010 

validly published 

Goniophlebium transpianense =Polypodium 

alternative 

(Yamamoto) C. M. Kuo 

transpianense combination, ?not 

(?nom. nud.): Kuo, C. M. 

validly published 

2010 

Goniopteris prolifera (Retz.) C. =Cyclosorus proliferus MAN 

Presl 

Gonocormus diffusus (Blume) =Crepidomanes MAN 

Bosch 

minutum 

Gonocormus minutus (Blume) =Crepidomanes MAN 

Bosch 

minutum 

Gonocormus nitidulus (Bosch) =Sphaerocionium MAN 

Prantl 

nitidulum 

Gonocormus prolifer (Blume) =Crepidomanes MAN 

Prantl 

minutum 

Gonocormus saxifragoides (C. =Crepidomanes MAN 

Presl) Bosch 

minutum 

Grammitis dorsipila auct. non =Grammitis fenicis MAN 

(Christ) C. Chr. & Tardieu: 

FOT 1975 

Grammitis fasciculata Blume =Grammitis intromissa MAN 

Grammitis jagoriana auct. non =Grammitis fenicis MAN 

(Mett. ex Kuhn) Tagawa: FOT 

1975 

Grammitis latifolia DeVol =Grammitis intromissa MAN 

Grammitis leptophylla (L.) Sw. =Anogramma 

MAN 

leptophylla 

Grammitis malaica (Alderw.) =Grammitis adspera FOT 2003 

Tagawa 

Grammitis procera Wall. (nom. =Coniogramme procera MAN 

nud.) 

Grammitis setigerum (Blume) =Grammitis intromissa MAN 

Ching 

Grammitis setosa auct. non =Grammitis congener MAN 

(Kaulf.) C. Presl: FOT 1975 

Grammitis subfalcata (Blume) 

Ching 

=Grammitis doniana MAN

Grammitis vestita Wall. (nom. 

nud.) 

=Paraceterach vestita MAN/Knapp, R. 

2011 

Gymnocarpium remotum =Gymnocarpium FOT 2003 


remotepinnatum 

Gymnogramma arifolia (Burm. 

f.) Kuhn 

=Paraceterach cordata MAN/Knapp, R. 

2013 

Gymnogramma aurita auct. 

non Hook.: MAN 


levingei 


2013 

Gymnogramma aurita var. =Pseudophegopteris Knapp, R. 2013 

levingei C. B. Clarke 

levingei 

Gymnogramma calomelanos =Pityrogramma MAN 

(L.) Kaulf. 

calomelanos 

Gymnogramma decurrentialata 

=Cornopteris 

MAN 

Hook. 


Gymnogramma fauriei Christ =Asplenium tenuicaule FOT 2003 

Gymnogramma fraxinea (D. =Coniogramme MAN 

Don) Bedd. 

fraxinea 

Gymnogramma grammitoides =Loxogramme 

MAN 

Baker 

grammitoides 

Gymnogramma japonica =Coniogramme MAN 

(Thunb.) Desv. 

japonica 

Gymnogramma leptophylla (L.) =Anogramma 

MAN 

Desv. 

leptophylla 

Gymnogramma opaca (D. Don) =Cornopteris opaca MAN 

Spreng. 

Gymnogramma rhizophylla =Bolbitis rhizophylla MAN 

Kaulf. 

Gymnogramma salicifolia =Loxogramme 

MAN 

Makino 

salicifolia 

Gymnogramma vestita C. Presl 

(nom. nud.) 


2011 

Gymnogramma vestita Hook. =Paraceterach vestita MAN/Knapp, R. 

2011 

Gymnogramma wrightii Hook. =Colysis wrightii MAN 

Gymnopteris bonii Christ =Tectaria harlandii FOT 2003/MAN 

Gymnopteris contaminans =Bolbitis angustipinna FOT 2003/MAN 

(Wall.) Bedd. 

Gymnopteris decurrens Hook. =Tectaria harlandii MAN 

Gymnopteris lanceolata (L.) T. =Pyrrosia lanceolata MAN 

Moore 

Gymnopteris repanda auct. =Bolbitis subcordata FOT 2003 

non (Blume) Christ 

Gymnopteris vestita (Hook.) =Paraceterach vestita MAN/Knapp, R. 

Underw. 

Gymnosphaera denticulata 

(Baker) Copel. 

=Cyathea hancockii 

2011 

MAN 

237

238 

Gymnosphaera metteniana =Cyathea metteniana MAN 

(Hance) Tagawa 

Gymnosphaera podophylla =Cyathea podophylla MAN 


Haplodictyum cumingii (C. =Cephalomanes MAN 

Presl) Bosch 

cumingii 

HAPLOPTERIS =Vittaria (kept for H. -- 

heterophylla) 

Haplopteris anguste-elongata =Vittaria angusteelongata 

MAN 

(Hayata) E. H. Crane 

Haplopteris flexuosa (Fee) E. =Vittaria flexuosa MAN 

H. Crane 

Haplopteris heterophylla C. W. =Vittaria heterophylla Knapp, R. 2013 

Chen, Y. H. Chang & Y. C. 

Liu 

Haplopteris taeniophylla =Vittaria taeniophylla MAN 

(Copel.) E. H. Crane 

Haplopteris zosterifolia (Willd.) =Vittaria zosterifolia MAN 

E. H. Crane 

Hemigramma decurrens =Tectaria harlandii MAN 


Hemionitis arifolia (Burm. f.) T. 

Moore 


2013 

Hemionitis esculenta Retz. =Diplazium esculentum FOT 2003 

Hemionitis griffithii (T. Moore) =Cyclosorus griffithii FOC/MAN 

Hook. f. & Thomson 

Hemionitis japonica Thunb. =Coniogramme MAN 

japonica 

Hemionitis opaca D. Don =Cornopteris opaca MAN 

Hemionitis parvula (Blume) C. =Antrophyum parvulum MAN 

Presl 

Hemionitis pothifolia auct. non 

Buch.-Ham. ex D. Don: FOT 

=Colysis pothifolia MAN/Knapp, R. 

2011 

2003 

Hemionitis pozoi Lag. =Cyclosorus pozoi MAN 

Hemionitis prolifera Retz. =Cyclosorus proliferus MAN 

Hemionitis sessilifolia Cav. =Antrophyum 

MAN 

sessilifolium 

Hemionitis wilfordii Hook. (type: =Cyclosorus wilfordii Knapp, R. 2011 


Hemionitis vestita (Hook.) J. 

Sm. 


2011 

Hemiphlebium bimarginatum =Trichomanes 

MAN 

(Bosch) Luerss. 

Heteroneuron scalpturatum 

Fee 

bimarginatum 

=Bolbitis scalpturata 

MAN

Hicriopteris blotiana (C. Chr.) 

Ching 

Hicriopteris chinensis 


Hicriopteris glauca (Thunb. ex 

Houtt.) Ching 

Hicriopteris laevissima (Christ) 

Ching 

Hippochaete debilis (Roxb. ex 

Vaucher) Holub 

Hippochaete ramosissima 

subsp. debilis (Roxb. ex 

Vaucher) A. Love & D. Love 

Hippochaete ramosissima 

(Desf.) Borner subsp. 

ramosissima 

Humata chrysanthemifolia 

(Hayata) C. Chr. 

Humata cumingii (Hook.) 

Brack. 

Humata dryopteridifrons 


Humata gaimardiana (Gaud.) 

J. Sm. 

Humata griffithiana (Hook.) C. 

Chr. 

Humata griffithiana var. 

tyermanni (T. Moore) Tagawa 

Humata hookeri (T. Moore) 

Diels 

Humata immersa (C. Presl) 

Mett. 


blotianum 


chinensis 


glaucum 


laevissima 

=Equisetum 

ramosissimum 

subsp. debile 

=Equisetum 

ramosissimum 

subsp. debile 

=Equisetum 

ramosissimum 

subsp. 

ramosissimum 

=Davallia 

chrysanthemifolia 

=Davallia cumingii 


=Davallia pectinata 

=Davallia griffithiana 

=Davallia tyermanni 

=Araiostegia 

parvipinnula 


MAN 

MAN 

MAN 

MAN 

MAN 

MAN 

MAN 

MAN 

MAN 

MAN/Fraser- 


2008 

FOT 2003/MAN 

MAN 

MAN 

MAN 

FOC/Fraser- 


2008 


Humata kinabaluensis auct. =Davallia cumingii 

non Copel.: FOT 2003 

Humata lepida T. Moore =Davallia cumingii FOT 2003/MAN 

Humata macrostegia Tagawa =Davallia 

FOT 2003 



Humata obtusata Alderw. =Davallia 

Kuo, C. M. 1997 


Humata parallela (Wall. ex =Davallia pectinata MAN 

Hook.) Brack. 

Humata pectinata (Sm.) Desv. =Davallia pectinata MAN 

239

240 

Humata pedata (Sm.) J. Sm. =Davallia repens MAN 

Humata perdurans (Christ) =Araiostegia 

MAN 

Hieron. 

parvipinnula 

Humata repens (L. f.) J. Small =Davallia repens MAN 

ex Diels 

Humata solida (G. Forst.) =Davallia solida MAN 

Desv. 

Humata trifoliata Cav. =Davallia cumingii MAN 

Humata tyermanni T. Moore =Davallia tyermanni MAN 

Humata vestita auct. non =Davallia cumingii Knapp, R. 2011 

Blume 

Huperzia cryptomeriana =Huperzia 


(Maxim.) R. D. Dixit (misspelt 

species name) 

cryptomerina 

Huperzia formosana Holub 

(nom. illeg.) ("type": Taiwan) 

=Huperzia 

cryptomerina 


2011 

Huperzia juniperistachys =Huperzia fordii MAN 

(Hayata) Holub 

Huperzia laxa (C. Presl) T. Sen =Huperzia carinata MAN 

& U. Sen 

Huperzia myriophyllifolia =Huperzia serrata MAN 

(Hayata) Holub 

Huperzia selago var. appressa =Huperzia appressa MAN/Knapp, R. 

(Bach. Pyl. ex Desv.) Ching 

Huperzia selago subsp. 

appressa (Bach. Pyl. ex 

Desv.) D. Love 

Huperzia serrata f. 

longipetiolata (Spring) Ching 

Huperzia serrata var. 

longipetiolata (Spring) H. M. 

Chang 

Huperzia serrata f. 

myriophyllifolia (Hayata) C. 

M. Kuo (nom. non rite pub.) 

Huperzia serrata var. 

myriophyllifolia (Hayata) C. 

M. Kuo 

Huperzia taiwanense (C. M. 

Kuo) C. M. Kuo 

Huperzia taiwanensis (Ching) 

Holub (nom. nud.) 

Hydroglossum japonicum 

(Thunb.) Willd. 

Hymenasplenium cheilosorum 

Kunze ex. Mett. 

2011 

=Huperzia appressa MAN/Knapp, R. 

2011 

=Huperzia serrata 




=Huperzia 

cryptomerina 

=Huperzia 

cryptomerina 

=Lygodium japonicum 

=Asplenium 

cheilosorum 

MAN 

FOC 

MAN 

Kuo 



2011 


MAN

Hymenasplenium obscurum =Asplenium obscurum MAN 

(Blume) Tagawa 

Hymensplenium rahaoense (Y. =Asplenium excisum MAN 

Yabe ex Hayata) H. Ito ex 

Tuyama 

Hymensplenium unilaterale =Asplenium excisum MAN 

var. rahaoense (Y. Yabe ex 

Hayata) Nemoto 

Hymenolepis formosana M. =Belvisia mucronata MAN 

Ogata (type: Taiwan) 

Hymenolepis mucronata Fee =Belvisia mucronata MAN 

Hymenophyllum australe auct. =Hymenophyllum FOT 2003 

non Willd. 

javanicum 

Hymenophyllum blumeanum =Hymenophyllum FOT 2003 

Spreng. 

polyanthos 

Hymenophyllum constrictum ?=Hymenophyllum MAN 


polyanthos 

Hymenophyllum crispatoalatum 

?=Hymenophyllum FOT 2003 

Hayata (type: Taiwan) badium 

Hymenophyllum crispatoalatum 


f. remotipinnum 


badium 

Hymenophyllum crispatum =Hymenophyllum MAN 

Wall. ex Hook. & Grev. 

badium 

Hymenophyllum digitatum =Sphaerocionium this publication 

(Sw.) Fosberg 

digitatum 

Hymenophyllum flexile Makino =Hymenophyllum MAN 

badium 

Hymenophyllum hayatae ?=Hymenophyllum MAN 

Masam. 

polyanthos 

Hymenophyllum humile Nees =Hymenophyllum FOT 2003 

ex Blume 

denticulatum 

Hymenophyllum japonicum =Hymenophyllum MAN 

Miq. 

Hymenophyllum javanicum var. 

badium (Hook. & Grev.) C. B. 

Clarke 

Hymenophyllum micranthum 

Bosch 

Hymenophyllum microsorum 

Bosch 

Hymenophyllum nitidulum 

(Bosch) Ebihara & K. Iwats. 

Hymenophyllum okadai 

Masam. 

barbatum 


badium 


javanicum 

?=Hymenophyllum 

polyanthos 


nitidulum 


simonsianum 

MAN 

FOT 2003 

MAN 


MAN 

241

242 

Hymenophyllum 

parallelocarpum Hayata 


Hymenophyllum pilosissimum 

C. Chr. 

Hymenophyllum punctisorum 

Rosenst. 

Hymenophyllum retusilobum 


Hymenophyllum taiwanense 

DeVol 

Hymenophyllum wrightii non 

auct. Bosch: FOT 2003 

Hypodematium taiwanensis 

Ching ex K. H. Shing 

Hypolepis alte-gracillima 


Isoetes kinmenensis (nom. 

nud.) 

Isoetes sp. in Knapp, R. 2011 

Japanobotrychiun lanuginosum 

(Wall. ex Hook. & Grev.) 

Nishida ex Tagawa 

Kaulinia hancockii (Baker) B. 

K. Nayar 

Lacostea auriculata (Blume) 

Prantl 

Lacosteopsis auriculata 

(Blume) T. Nakaike 

Lacosteopsis maxima (Blume) 

T. Nakaike 

Lacosteopsis orientalis auct. 

non (C. Chr.) T. Nakaike: 

MAN 

Lacosteopsis orientalis var. 

naseana (Christ) T. Nakaike 

Lastrea apiciflora (Wall. ex 

Mett.) Bedd. 

Lastrea barbigera auct. non (T. 

Moore ex Hook.) Bedd.: FOT 

2003 

Lastrea beddomei (Baker) 

Bedd. 


polyanthos 

MAN 

=Sphaerocionium this publication 

pilosissimum 

?=Hymenophyllum MAN 

polyanthos 

=Mecodium badium FOT 2003 


devolii 

=Hymenophyllum Knapp, R. 2011 

polyanthos 

?=Hypodematium FOC 

crenatum 

=Hypolepis alpina Knapp, R. 2011 

=Isoetes taiwanensis 

var. kinmenensis 

=Isoetes taiwanensis 

var. kinmenensis 

=Botrychium 

lanuginosum 

=Microsorum insigne 

=Crepidomanes 

auriculatum 

=Crepidomanes 

auriculatum 

=Crepidomanes 

maximum 

=Crepidomanes 

birmanicum 

=Crepidomanes 

birmanicum 



=Thelypteris beddomei 



MAN 

MAN 

MAN 

MAN 

MAN 



26-40 (2009) 



26-40 (2009) 

MAN 


R. 2011 

FOT 2003/MAN

243 

Lastrea crenata (Forssk.) 

Bedd. 

Lastrea decursive-pinnata (van 

Hall) J. Sm. 

Lastrea erubescens (Wall. ex 

Hook.) Copel. 

Lastrea filix-mas var. 

parallelogramma subvar. 

fibrillosa Bedd. 

Lastrea filix-mas var. serratodentata 

Bedd. (nom. nud.) 

Lastrea glanduligera (Kunze) 

T. Moore 

Lastrea gracilescens (Blume) 

T. Moore 

Lastrea hendersonii Bedd. 

Lastrea hookeriana C. Presl 

(nom. nud.) 

Lastrea laxa (Franch. & Sav.) 

Copel. 

Lastrea miqueliana Tagawa 

Lastrea oligophlebia var. 

lasiocarpa (Hayata) H. Ito ex 

Mizushima 

Lastrea omeiensis (Baker) 

Copel. 

Lastrea ornata (Wall. ex Bedd.) 

Copel. 

Lastrea simozawae (Tagawa) 

Tagawa 

Lastrea subochthodes (Ching) 

Tagawa 

Lastrea tenericaulis (Wall. ex 

Hook.) T. Moore 

Lastrea torresiana (Gaud.) T. 

Moore 

Lastrea uraiensis (Rosenst.) 

Copel. 

Lastreopsis recedens (J. Sm. 

ex T. Moore) Ching 

Lastreopsis simozawae 


=Hypodematium 

crenatum 

=Phegopteris 


=Cyclosorus 

erubescens 

=Dryopteris edwardsii 



=Dryopteris 


=Dryopteris 


=Thelypteris 

glanduligera 

=Thelypteris 

gracilescens 


hendersonii 

=Polystichum 

integripinnum 

=Thelypteris laxa 

=Thelypteris 

angustifrons 


torresiana 



polypodioides 

=Thelypteris 

angulariloba 

=Cyclosorus esquirolii 


torresiana 


torresiana 

=Thelypteris uraiensis 

MAN 

FOT 2003 

FOT 2003/MAN 


R. 2011 

MAN 

FOT 2003/MAN 

FOT 2003/MAN 

MAN 

MAN 

FOT 2003/MAN 

FOT 2003/MAN 

FOT 2003/MAN 

FOT 2003/MAN 

FOT 2003/MAN 

FOT 2003/MAN 

FOT 2003/MAN 

FOT 2003/MAN 

FOT 2003/MAN 

FOT 2003/MAN 

=Lastreopsis tenera FOT 2003 

=Lastreopsis tenera FOT 2003

244 

Lecanopteris formosana ?=Grammitis contigua MAN 


Lemmaphyllum carnosum auct. =Lemmaphyllum Knapp, R. 2011 

non (J. Sm. ex Hook.) C. 

Presl: MAN 

microphyllum 

Lemmaphyllum christensenii =Lemmaphyllum FOT 2003/MAN 

Ching 

rostratum 

Lemmaphyllum diversum =Lemmaphyllum MAN 

(Rosenst.) Tagawa 

rostratum 

Lemmaphyllum microphyllum 

var. obovatum (Harr.) C. Chr. 

=Lemmaphyllum 

microphyllum 


2011 

Lemmaphyllum subrostratum =Lemmaphyllum FOT 2003/MAN 

auct. non (C. Chr.) Ching rostratum 

Lepidogrammitis rostrata =Lemmaphyllum MAN 

(Bedd.) Ching 

rostratum 

Lepidotis annotina (L.) P. =Lycopodium 

MAN 

Beauv. 

annotinum 

Lepidotis casuarinoides =Lycopodium 

FOC 

(Spring) Rothm. 

casuarinoides 

Lepidotis cernua (L.) P. Beauv. =Lycopodiella cernua MAN 

Lepidotis phlegmaria (L.) P. =Huperzia phlegmaria MAN 

Beauv. 

Lepisorus angustifrons Tagawa =Lepisorus 

FOT 2003 


pseudoussuriensis 

Lepisorus bicolor (Takeda) =Lepisorus 

FOT 2003 

Ching 

morrisonensis 

Lepisorus heterolepis auct. non =Lepisorus monilisorus FOT 2003 


Lepisorus infraplanicostalis =Lepisorus tosaensis FOT 2003 


Lepisorus megasorus auct. non =Lemmaphyllum FOT 2003/MAN 

(C. Chr.) Ching 

rostratum 

Lepisorus oligolepidus auct. =Lepisorus kawakamii Knapp, R. 2011 

non (Baker) Ching 

Leptochilus angustipinnus =Bolbitis angustipinna MAN 


Leptochilus cuspidatus auct. =Bolbitis heteroclita FOT 2003 

non (Pr.) C. Chr. 

Leptochilus cuspidatus var. 

crenatus Rosenst. (type: 




348 (1915) 

Leptochilus harlandii (Hook.) C. =Tectaria harlandii FOT 2003/MAN 

Chr. 

Leptochilus hemionitideus (C. 

Presl) Noot. 

=Colysis hemionitidea MAN

Leptochilus heteroclitus (C. =Bolbitis heteroclita MAN 


Leptochilus kanashiroi Hayata =Tectaria harlandii MAN 


Leptochilus macrophyllus var. =Colysis wrightii MAN 

wrightii (Hook.) Noot. 

Leptochilus x shintenensis =Colysis x shintenensis this publication 

(Hayata) T. Nakaike 

Leptochilus x shintenensis =Colysis x shintenensis this publication 

(Hayata) X. C. Zhang & Noot. 

Leptochilus virens auct. non =Bolbitis subcordata FOT 2003 

(Wall.) C. Chr. 

Leptochilus zeylanicus (Houtt.) =Tectaria zeilanica FOT 2003 

C. Chr. 

Leptocionium affine (Bosch) =Hymenophyllum FOT 2003 

Bosch 

holochilum 

Leptocionium barbatum Bosch =Hymenophyllum MAN 

barbatum 

Leptocionium denticulatum =Hymenophyllum MAN 

(Sw.) Bosch 

denticulatum 

Leptocionium holochilum =Hymenophyllum MAN 

(Bosch) Bosch 

holochilum 

Leptogramma aurita auct. non 

(Hook.) Bedd.: MAN 


levingei 


2013 

Leptogramma caudata Ching =Cyclosorus tottoides MAN 


Leptogramma decursivepinnata 

=Phegopteris 

MAN 

(H. C. Hall) J. Sm. decursivepinnata 

Leptogramma omeiensis =Cyclosorus omeiensis MAN 


Leptogramma pozoi (Lag.) =Cyclosorus pozoi MAN 

Ching 

Leptogramma totta auct. non =Cyclosorus tottoides FOT 2003/MAN 


Leptogramma tottoides H. Ito =Cyclosorus tottoides MAN 


Leptorumohra quadripinnata =Arachniodes 

MAN 

(Hayata) H. Ito 

quadripinnata 

Leptostegia lucida D. Don =Onychium lucidum MAN 

Leucostegia clarkei (Baker) C. =Araiostegia 

MAN 

Chr. 

parvipinnula 

Leucostegia griffithiana (Hook.) =Davallia griffithiana MAN 

J. Sm. 

Leucostegia hookeri (T. Moore) 

Bedd. 

=Araiostegia 

parvipinnula 

MAN 

245

246 

Leucostegia immersa (Wall. ex 

Hook.) C. Presl 

=Leucostegia truncata Fraser-Jenkins, 

C. R. 2008 

Leucostegia parvipinnnula =Araiostegia 

MAN 


parvipinnula 

Lindsaea chinensis (L.) Mett. =Odontosoria chinensis MAN 

ex Kuhn 

Lindsaea chienii var. deltoidea =Lindsaea javanensis MAN 

(Y. C. Wu) Tagawa 

Lindsaea concinna auct. non J. =Lindsaea lucida FOT 2003/MAN 

Sm. 

Lindsaea cultrata var. japonica =Lindsaea japonica MAN 

Baker 

Lindsaea davallioides auct. non =Lindsaea cultrata FOT 2003 

Blume 

Lindsaea gracilis Blume =Lindsaea lucida Knapp, R. 2011 

Lindsaea kusukusensis Hayata =Lindsaea lucida MAN 


Lindsaea macraeana auct. non =Lindsaea 

FOT 2003/MAN 

Copel. 

yaeyamensis 

Lindsaea merrillii subsp. 

yaeyamensis (Tagawa) K. U. 

Kramer 

=Lindsaea 

yaeyamensis 


Lindsaea merrillii var. 

yaeyamensis (Tagawa) W. C. 

Shieh 

Lindsaea odorata var. japonica 

(Baker) K. U. Kramer 

Lindsaea orbiculata var. chienii 

(Ching) Ohwi 

Lindsaea orbiculata var. 

commixta (Tagawa) K. U. 

Kramer 


deltoidea Y. C. Wu 


recedens (Ching) W. C. 

Shieh 

=Lindsaea 

yaeyamensis 

=Lindsaea japonica 

=Lindsaea chienii 

=Lindsaea commixta 

=Lindsaea javanensis 

=Lindsaea chienii 


MAN 

MAN 

MAN 

MAN 

MAN 

Lindsaea recedens Ching =Lindsaea chienii MAN 

Lindsaea repens auct. non =Lindsaea 

FOT 2003 

(Bory) Bedd. 

yaeyamensis 

Lindsaea securifolia var. 

kusukusensis (Hayata) W. C. 

Shieh 

=Lindsaea lucida MAN 

Lindsaea taiwaniana Ching =Lindsaea orbiculata FOT 2003 

Lindsaea tenera var. chienii =Lindsaea chienii MAN 

(Ching) Tardieu & C. Chr.

Lindsaea tenera var. commixta 

(Tagawa) K. Iwats. 

LINDSAEACEAE 

=Lindsaea commixta 

=DENNSTAEDTIA- 

CEAE (Subfamily 

Lindsaeoideae) 

MAN 

Lomaria adnata Blume =Plagiogyria adnata MAN 

Lomaria euphlebia Kunze =Plagiogyria euphlebia MAN 

Lomaria fraseri A. Cunn. =Blechnum fraseri MAN 

Lomaria glauca Blume =Plagiogyria formosana MAN 

-- 

247 

Lomaria hancockii (Hance) =Blechnum hancockii MAN 

Baker 

Lomaria matsumureana ?=Plagiogyria koidzumii (MAN) 

Makino (nom. nud.?) 

Lomaria spectabilis Kunze =Lomariopsis 

MAN 

spectabilis 

Lomaria stenoptera (Hance) =Plagiogyria stenoptera MAN 

Baker 

Lomariopsis leptocarpa auct. =Lomariopsis 

FOT 2003 

non Fee 

spectabilis 

Lonchitis tenuifolia G. Forst. =Hypolepis tenuifolia MAN 

Lorinseria harlandii (Hook.) J. =Woodwardia harlandii FOT 2003 

Sm. 

Loxogramme confertifolia =Loxogramme 

MAN 


chinensis 

Loxogramme duclouxii auct. =Loxogramme 


non Christ: Kuo, C. M. 2010 remotefrondigera 

Loxogramme ensiformis Ching =Loxogramme 

FOT 2003 

formosana 

Loxogramme fauriei Copel. =Loxogramme 

MAN 


salicifolia 

Loxogramme linearis Copel. 


=Loxogramme 

remotefrondigera 


Geobot. 8(4): 233 

Lunathyrium allantodioides 

(Bedd.) Ching 

Lunathyium orientale auct. non 

(?) Z. 

R. Wang & J. J. Chien 

Lunathyrium petersenii (Kunze) 

H. Ohba 

Lunathyium pycnosorum auct. 

non (Christ) Koidz.: FOT 

1975 

Lunathyrium unifurcatum 

(Baker) Sa. Kurata 

=Deparia allantodioides 

(1939) 

MAN 

=Deparia sp. 1 Knapp, R. 2011 


=Deparia allantodioides 

=Deparia unifurcata 

MAN 

MAN 

MAN

248 

Lycopodiastrum casuarinoides =Lycopodium 

MAN 

(Spring) Holub ex R. D. Dixit casuarinoides 

Lycopodioides delicatula =Selaginella delicatula MAN 

(Desv. ex Poir.) H. S. Ku 

Lycopodioides doederleinii =Selaginella 

MAN 

(Hieron.) H. S. Kung 

doederleinii 

Lycopodioides labordei =Selaginella labordei MAN 

(Hieron. ex Christ) H. S. Kung 

Lycopodioides remotifolia =Selaginella remotifolia MAN 

(Spring) H. S. Kung 

Lycopodium alpinum var. =Lycopodium veitchii MAN 

transmorrisonense Hayata 


Lycopodium appressum 

(Desv.) Petrov 


2011 

Lycopodium carinatum Desv. =Huperzia carinata MAN 

ex Poir. 

Lycopodium casuarinoides var. =Lycopodium 

MAN 

japonicum Nakai 

casuarinoides 

Lycopodium caulescens Wall. =Selaginella involvens FOT 2003 


Lycopodium cernuum L. =Lycopodiella cernua MAN 

Lycopodium chinense var. =Huperzia somae MAN 

somae (Hayata) Masam. 

Lycopodium christensenianum =Huperzia sieboldii MAN 

Christ & Herter 

Lycopodium ciliare Retz. =Selaginella ciliaris MAN 

Lycopodium clavatum auct. =Lycopodium 

MAN 

non. L.: FOT 1975 

pseudoclavatum 

Lycopodium clavatum var. =Lycopodium 

MAN 

nipponicum Nakai 

japonicum 

Lycopodium complanatum =Lycopodium 

MAN 

auct. non. L.: FOT 1975 multispicatum in part, 

=Lycopodium 

yueshanense in part 

Lycopodium cryptomeri(a)num =Huperzia 

MAN 

Maxim. 

cryptomerina 

Lycopodium cunninghamioides =Huperzia 

MAN 


cunninghamioides 

Lycopodium delicatulum Desv. =Selaginella delicatula MAN 

ex Poir. 

Lycopodium fargesii Herter =Huperzia fargesii MAN 

Lycopodium fargesii var. 

gracile Tagawa (type: 


=Huperzia fargesii FOC/MAN

Lycopodium fauriei Rosenst. =Huperzia fargesii MAN 


Lycopodium fordii Baker =Huperzia fordii MAN 

Lycopodium formosanus Herter =Huperzia salvinioides MAN 

ex Hayata 

Lycopodium hamiltonii auct. 

non. Sprengel ex Grev. & 

=Huperzia fordii Kuo, C. M. 

1997/MAN 

Hook. 

Lycopodium involvens Sw. =Selaginella involvens MAN 

Lycopodium juniperstachyum =Huperzia fordii FOT 2003 


Lycopodium laxum C. Presl =Huperzia carinata MAN 

Lycopodium nudum L. =Psilotum nudum MAN 

Lycopodium obscurum auct. =Lycopodium 

MAN 

non. L.: FOT 1975 

Lycopodium obscurum f. 

juniperoideum (Sw.) H. 

Takeda 

juniperoideum 

=Lycopodium 

juniperoideum 

MAN 

249 

Lycopodium phlegmaria L. =Huperzia phlegmaria MAN 

Lycopodium pseudoclavatum 

Ching 

=Lycopodium 

japonicum 


R. 2011 

Lycopodium pulcherrimum 

auct. non. Wall. ex Hook. & 

Grev.: FOT 1975 

=Huperzia 

cryptomerina 

Kuo/Knapp, R. 

2011 

Lycopodium 

quasipolytrichoides Hayata 


Lycopodium reflexo-integrum 


Lycopodium remoganense 


Lycopodium repandum Desv. 

ex Poir. 

Lycopodium salvinioides 

(Herter) Tagawa 

Lycopodium selago var. 

appressum Bach. Pyl. ex 

Desv. 

=Huperzia 

quasipolytrichoides 

=Huperzia 

quasipolytrichoides 

=Huperzia squarrosa 

=Selaginella repanda 

=Huperzia salvinioides 

MAN 

MAN 

MAN 

MAN 

MAN 


2011 

Lycopodium selago var. somae =Huperzia somae MAN 

(Hayata) Masam. 

Lycopodium serratum Thunb. =Huperzia serrata MAN 

Lycopodium serratum var. =Huperzia serrata MAN 

longipetiolatum Spring 

Lycopodium serratum var. 

myriophyllifolium Hayata 


=Huperzia serrata MAN

250 

Lycopodium sieboldii Miq. =Huperzia sieboldii MAN 

Lycopodium sitchense var. =Lycopodium veitchii MAN 

veitchii (Christ) Takeda 

Lycopodium somae Hayata =Huperzia somae MAN 


Lycopodium subdistichum =Huperzia fordii MAN 

Makino 

Lycopodium squarrosum G. =Huperzia squarrosa MAN 

Forst. 

Lycopodium taiwanense C. M. =Huperzia 


Kuo 

cryptomerina 

Lycopodium tereticaule Hayata =Huperzia fargesii MAN 


Lycopodium wightianum auct. =Lycopodium 

Kuo 

non Hook. & Grev. 

yueshanense 

Lygodium japonicum var. =Lygodium japonicum MAN 

microstachyum (Desv.) C. 

Chr. & Tardieu 

Lygodium microstachyum =Lygodium japonicum MAN 

Desv. 

Lygodium microstachyum var. =Lygodium japonicum FOC 

glabrescens Nakai (type: 


Lygodium scandens var. =Lygodium 

MAN 

microphyllum (Cav.) Luerss. microphyllum 

Macroplethus mucronatus =Belvisia mucronata MAN 

(Fee) Tagawa 

Macrothelypteris ornata (Wall. =Macrothelypteris FOT 2003/MAN 

ex Bedd.) Ching 

polypodioides 

Macrothelypteris setigera auct. =Macrothelypteris Knapp, R. 2011 

non (Blume) Ching: CHK torresiana 

Macrothelypteris uraiensis =Thelypteris uraiensis MAN 

(Rosenst.) A. Love & D. Love 

Marattia fraxinea Sm. ex Christ =Marattia pellucida MAN 

Marginaria arisanensis 

=Polypodium amoenum MAN 

(Hayata) Nakai ex H. Ito 

Marginaria formosana (Baker) =Polypodium 

MAN 

Nakai ex H. Ito 

formosanum 

Marginaria pseudoformosana ?=Polypodium 

(MAN) 


raishaense 

Marginaria niponica (Mett.) 


=Polypodium 


=Polypodium 



and Knapp, R. 

2013

Marginaria raishanense 

(Rosenst.) Nakai ex H. Ito 

Marginaria taiwaniana (Hayata) 


Marginaria transpianensis 

(Yamam.) H. Ito 

=Polypodium 

raishaense 

=Polypodium 

mengtzeense 

=Polypodium 

transpianense 

251 


2011 and Knapp, 

R. 2013 

MAN 



R. 2013 

Marsilea crenata C. Presl =Marsilea minuta MAN 

Marsilea natans L. =Salvinia natans MAN 

Matteuccia orientalis (Hook.) =Onoclea orientalis MAN 

Trevis. 

Mecodium badium (Hook. & =Hymenophyllum MAN 

Grev.) Copel. 

badium 

Mecodium crispato-alatum ?=Hymenophyllum FOT 2003 


badium 

Mecodium crispatum (Wall. ex =Hymenophyllum MAN 

Hook. & Grev.) Copel. 

badium 

Mecodium flexile (Makino) =Hymenophyllum MAN 

Copel. 

badium 

Mecodium fimbriatum (J. Sm.) =Hymenophyllum MAN 

Copel. 

fimbriatum 

Mecodium javanicum (Spreng.) =Hymenophyllum MAN 

Copel. 

javanicum 

Mecodium microsorum (Bosch) =Hymenophyllum MAN 

Ching 

polyanthos 

Mecodium okadai (Masam.) ?=Hymenophyllum MAN 

Shieh 

simonsianum 

Mecodium oligosorum (Makino) =Hymenophyllum MAN 

H. Ito 

oligosorum 

Mecodium paniculiflorum (C. =Hymenophyllum Kuo, C. M. 1985 

Presl) Copel. 

paniculiflorum 

Mecodium polyanthos (Sw.) =Hymenophyllum MAN 

Copel. 

polyanthos 

Mecodium productum (Kunze) =Hymenophyllum MAN 

Copel. 

productum 

Mecodium taiwanense Tagawa =Hymenophyllum MAN 


taiwanense 

Mecodium wrightii auct. non =Hymenophyllum Knapp, R. 2011 

Bosch: FOT 2003 

polyanthos 

Meniscium cuspidatum Blume =Cyclosorus liukiuensis MAN/Knapp, R. 

2011 

Meniscium liukiuense Christ 

ex. Matsum. 


2011 

Meniscium parishii Bedd. =Cyclosorus parishii Knapp, R. 2011

252 

Meniscium proliferum (Retz.) =Cyclosorus proliferus MAN 

Sw. 

Meniscium simplex Hook. =Cyclosorus simplex MAN 

Meniscium triphyllum Sw. =Cyclosorus triphyllus MAN 

Meringium acanthoides auct. =Hymenophyllum FOT 2003/MAN 

non (Bosch) Copel. 

denticulatum 

Meringium blandum (Racib.) =Hymenophyllum MAN 

Copel. 

blandum 

Meringium denticulatum (Sw.) =Hymenophyllum MAN 

Copel. 

denticulatum 

Meringium holochilum (Bosch) =Hymenophyllum MAN 

Copel. 

holochilum 

Mertensia laevissima (Christ) =Diplopterygium MAN 

Nakai 

laevissimum 

Metathelypteris adscendens =Thelypteris 

MAN 

(Ching) Ching 

adscendens 

Metathelypteris gracilescens =Thelypteris 

MAN 

(Blume) Ching 

gracilescens 

Metathelypteris laxa (Franch. & =Thelypteris laxa MAN 

Sav.) Ching 

Metathelypteris uraiensis =Thelypteris uraiensis MAN 


Microgonium beccarianum =Trichomanes motleyi FOT 2003/MAN 

(Cesati) Copel. 

Microgonium bimarginatum =Trichomanes 

FOC 

Bosch 

bimarginatum 

Microgonium motleyi (Bosch) =Trichomanes motleyi MAN 

H. Ito 

Microgonium omphalodes =Trichomanes 

MAN 

Vieill. ex Fourn. 

tahitense 

Microgonium tahitense 

=Trichomanes 

MAN 

(Nadeaud) Tindale 

tahitense 

Microlepia biflora (Kaulf.) Mett. =Odontosoria biflora MAN 

Microlepia calvescens (Wall. ex =Microlepia calvescens MAN 

Hook.) Bedd. var. calvescens 

Microlepia calvescens var. =Microlepia 

MAN 

intramarginalis (Tagawa) W. 

C. Shieh 

intramarginalis 

Microlepia chinensis (L.) Mett. =Odontosoria chinensis MAN 

Microlepia gracilis auct. non 

(Blume) J. Sm. 

=Tapeinidium 

biserratum 

Kuo/Knapp, R. 

2011 

Microlepia grandissima Hayata =Microlepia platyphylla FOT 2003 


Microlepia hancei auct. non 

Prantl 

=Microlepia speluncae Knapp, R. 2011

253 

Microlepia herbacea var. 

trichosora (Ching) Seriz. 

Microlepia hirsutissima Hayata 


Microlepia mollifolia Tagawa 


Microlepia pinnata (Cav.) 

Bedd. 

Microlepia quadripinnata 


Microlepia rhomboidea C. 

Presl. (nom. nud.) 

Microlepia speluncae var. 

hancei auct. non (Prantl) C. 

Chr. & Tardieu 

Microlepia strigosa var. 

intramarginalis Tagawa (type: 


Microlepia subpinnata Hayata 


Microlepia taiwaniana Tagawa 


Microlepia trapeziformis auct. 

non (Roxb.) Kuhn: FOT 1975 

Micropolypodium okuboi 

(Yatabe) Hayata 

Micropolypodium 

pseudotrichomanoides 


Microsor(i)um brachylepis 

(Baker) T. Nakaike 

Microsor(i)um buergerianum 

auct. non (Miq.) Ching: FOT 

1975 

Microsorium buergerianum var. 

ohwianum (Tagawa) Tagawa 

Microsor(i)um carinatum auct. 

non (?) (W. M. Chu & Z. R. 

He) S. G. Lu 

=Microlepia trichosora 

=Microlepia obtusiloba 

? (similar to Microlepia 

speluncae with 

differences in 

dissection pattern of 

lamina, hairiness of 

axes and lamina and 

size of segments, 

probably a distinct 

taxon) 

=Tapeinidium pinnatum 

MAN 

MAN 


MAN 

=Arachniodes 

MAN 

quadripinnata 

=Microlepia 

MAN 

rhomboidea 

=Microlepia speluncae Knapp, R. 2011 

=Microlepia 

intramarginalis 

=Microlepia obtusiloba 

=Microlepia 

rhomboidea 

=Microlepia 

rhomboidea 

=Grammitis okuboi 


=Microsorum 

superficiale 

=Microsorum 

superficiale 

=Microsorum 

superficiale 

=Microsorum 

membranaceum 

MAN 

MAN 

MAN 

MAN 

MAN 

MAN 



2013 


R. 2013 


254 

Microsor(i)um dilatatum =Microsorum insigne MAN 

(Bedd.) Sledge 

Microsorium ensatum (Thunb.) =Neocheiropteris MAN 

H. Ito 

ensata 

Microsorium hancockii Baker =Microsorum insigne MAN 

Microsor(i)um fortunei auct. =Microsorum henryi MAN 

non (T. Moore) Ching: FOT 

1975 

Microsorium henryi (Christ) C. =Microsorum henryi Knapp, R. 2011 

M. Kuo, typo in genus 

spelling 

Microsorium insigne (Blume) =Microsorum insigne Knapp, R. 2011 

Copel., typo in genus spelling 

Microsorium membranaceum =Microsorum 


(D. Don) Ching, typo in genus 

spelling 

membranaceum 

Microsorium membranifolium =Phymatosorus MAN 

(R. Br.) Ching 

membranifolius 

Microsor(i)um nigrescens =Phymatosorus MAN 

(Blume) Copel. 


Microsorium ohwianum 


=Microsorum 

superficiale 


R. 2013 

Microsorium pteropus (Blume) =Microsorum pteropus Knapp, R. 2011 

Copel., typo in genus spelling 

Microsorium punctatum (L.) =Microsorum 


Copel., typo in genus spelling punctatum 

Microsorium rubidum (Kunze) =Phymatosorus MAN 

Copel. 

longissimus 

Microsorium steerei (Harr.) =Microsorum steerei Knapp, R. 2011 

Ching, typo in genus spelling 

Microsorium takedae (Nakai) =Microsorum henryi FOT 2003/MAN 

H. Ito 

Microsorum rubidum (Kunze) =Phymatosorus FOC 

Copel. 

longissimus 

Microsorum scolopendrium =Phymatosorus MAN 

(Burm. f.) Copel. 

scolopendria 

Microstegia aspera (Blume) C. 

Presl 

=Diplazium sikkimense MAN/Knapp, R. 

2013 

Microstegia esculenta (Retz.) =Diplazium esculentum MAN 

C. Presl 

Microtrichomanes digitatum 

(Sw.) Copel. 


digitatum 


2011 

Microtrichomanes nitidulum =Sphaerocionium MAN 

(Bosch) Copel. 

Mildella henryi (Christ) C. C. 

Hall & Lellinger 

nitidulum 

=Cheilanthes nitidula 

MAN

Monachosorum maximowiczii 

var. melanocaulon Hayata 


Monachosorum subdigitatum 

var. henryi (Christ) Tagawa 

Monogramma trochoidea J. 

Sm. (nom. nud.) 

Monomelangium pullingeri 


Neocheiropteris phyllomanes 


Neoniphopsis linearifolia 

(Hook.) Nakai 

Neottopteris antiqua (Makino) 

Masam. 

Neottopteris australasica J. 

Sm. 

=Monachosorum 

maximowiczii 

=Monachosorum henryi 

MAN 

MAN 

=Monogramma MAN 

trichoidea 

=Diplazium pullingeri FOT 2003 

=Neocheiropteris 

ensata 

=Pyrrosia linearifolia 

=Asplenium antiquum 

FOT 2003 

MAN 

MAN 

255 

=Asplenium 

MAN 

australasicum 

Neottopteris nidus (L.) J. Sm. =Asplenium nidus MAN 

Nephrodium apiciflorum (Wall. =Dryopsis apiciflora MAN 

ex Mett.) Hook. 

Nephrodium auriculatum (L.) A. =Nephrolepis cordifolia MAN 

Rich. 

Nephrodium banksiifolium C. =Osmunda banksiifolia MAN 

Presl 

Nephrodium barbigerum auct. 

non T. Moore ex Hook.: MAN 

=Dryopteris komarovii MAN/Knapp, R. 

2011 

Nephrodium beddomei Baker =Thelypteris beddomei MAN 

Nephrodium biserratum (Sw.) =Nephrolepis biserrata MAN 

C. Presl 

Nephrodium chrysocoma =Dryopteris 

FOT 2003/MAN 

(Christ) Hand.-Mzt. 

woodsiisora 

Nephrodium cicutarium var. =Tectaria dubia Knapp, R. 2013 

dubium C. B. Clarke & Baker 

Nephrodium clavivenum Yabe 

ex Matsum. & Hayata 

=Cyclosorus liukiuensis FOT 2003/Knapp, 

R. 2011 

Nephrodium decurrens (C. =Tectaria decurrens FOC 

Presl) Baker 

Nephrodium decursivepinnatum 

=Phegopteris 

FOT 2003 

(van Hall) Baker decursivepinnata 

Nephrodium dickinsii (Franch. =Dryopteris dickinsii MAN 

& Sav.) Baker 

Nephrodium diffractum Baker =Dryopteris diffracta MAN 

Nephrodium eatonii Baker =Ctenitis eatonii MAN 

Nephrodium eatonii var. 

formosanum Harrington 

=Ctenitis eatonii Knapp, R. 2011

256 

Nephrodium enneaphyllum 

Baker 

Nephrodium falconeri auct. non 

Hook. 

Nephrodium filix-mas var. 

fibrillosum C. B. Clarke 

=Dryopteris 

enneaphylla 


=Dryopteris edwardsii 



=Dryopteris 


=Dryopteris marginata 

MAN 


R. 2011 


Nephrodium filix-mas var. 

MAN 

marginatum C. B. Clarke 

Nephrodium fuscipes (Wall.) =Tectaria fuscipes FOT 2003/MAN 

Christ 

Nephrodium glanduligerum =Thelypteris 

FOT 2003/MAN 

(Kunze) Makino 

glanduligera 

Nephrodium griffithii auct. non =Tectaria multicaudata FOC 

Baker: Knapp, R. 2011 

Nephrodium gymnosorum var. =Dryopteris tenuicula MAN 

indusiatum Makino 

Nephrodium jaculosum (Christ) =Cyclosorus jaculosus FOT 2003/MAN 

Matsum. & Hayata 

Nephrodium japonicum Baker =Thelypteris japonica MAN 

Nephrodium latipinnum =Cyclosorus latipinnus FOC 

(Bentham) Hook. ex Baker 

Nephrodium leucostipes Baker =Ctenitis eatonii MAN 


Nephrodium matsumurae =Dryopsis 

FOT 2003/MAN 

(Makino) Makino 


Nephrodium membranifolium =Tectaria fuscipes FOT 2003/MAN 

C. Presl 

Nephrodium multicaudatum C. =Tectaria multicaudata this publication 

B. Clarke 

Nephrodium odoratum Baker =Hypodematium FOT 2003/MAN 

crenatum 

Nephrodium papilio C. Hope =Cyclosorus papilio MAN 

Nephrodium parasiticum (L.) =Cyclosorus 

MAN 

Desv. 

parasiticus 

Nephrodium polymorphum =Tectaria polymorpha this publication 

Baker 

Nephrodium serratodentatum =Dryopteris 

MAN 

C. Hope 


Nephrodium sheareri Baker =Athyrium sheareri MAN 

Nephrodium simonsii Baker =Tectaria simonsii MAN 

Nephrodium sparsum Buch.- =Dryopteris sparsa MAN 

Ham. ex D. Don

257 

Nephrodium spectabile C. B. =Nothoperanema FOT 2003 

Clarke 

hendersonii 

Nephrodium squamisetum =Nothoperanema MAN 

Hook. 

squamiseta 

Nephrodium subpedatum Harr. 


=Tectaria polymorpha Fl. Males., Ser. 2, 

2: 87 (1991) 

Nephrodium subtriangulare C. =Dryopteris 

MAN 

Hope 

subtriangularis 

Nephrodium tenerum R. Br. =Lastreopsis tenera MAN 

Nephrodium triphyllum (Sw.) =Cyclosorus triphyllus FOT 2003/MAN 

Diels 

Nephrodium truncatum (Poir.) =Cyclosorus truncatus FOT 2003 

Pr. 

Nephrodium unifurcatum Baker =Deparia unifurcata MAN 

Nephrodium urophyllum auct. =Cyclosorus 

FOT 2003/MAN 

non (Wall.) Keys. 


Nephrodium variolosum (Wall. 

ex Hook.) Baker 

=Tectaria impressa FOT 2003/ 


C. R. 2008 

Nephrodium yunnanense 

Baker 

=Tectaria griffithii MAN/this 

publication 

Nephrolepis arida D. L. Jones ?=Nephrolepis x 

hippocrepicis 

Blumea 50: 312 

(2005) 

Nephrolepis auriculata (L.) =Nephrolepis cordifolia Knapp, R. 2011 

Trimen 

Nephrolepis exaltata var. =Nephrolepis biserrata MAN 

biserrata (Sw.) Baker 

Nephrolepis hirsutula auct. non 

(Forst.) C. Presl. 

=Nephrolepis brownii FOT 2003/Knapp, 

R. 2011 

Nephrolepis hirsutula var. =Nephrolepis biserrata FOC 

biserrata (Sw.) Kuntze 

Nephrolepis multiflora 

=Nephrolepis brownii Knapp, R. 2011 

(Roxburgh) Jarrett & Morton 

Nephrolepis tenuissima Hayata 


=Athyrium nakanoi Liu, Y. C. et al. 

2009 

Nesopteris blepharistoma =Cephalomanes FOT 2003/MAN 

(Copel.) Tagawa 

thysanostomum 

Nesopteris grandis (Copel.) =Cephalomanes Knapp, R. 2013 

Copel. 

grande 

Nesopteris pseudoblepharistoma 

=Cephalomanes MAN 

(Tagawa) Masam. thysanostomum 

Nesopteris thysanostoma =Cephalomanes MAN 

(Makino) Copel. 

Neurogramme aurita (D. Don) 

Christ 

thysanostomum 

=Coniogramme 

fraxinea 

this publication

258 

Niphobolus adnascens (Sw.) =Pyrrosia lanceolata MAN 

Kaulf. 

Niphobolus angustissimus =Pyrrosia angustissima MAN 

Giesenh. ex Diels 

Niphobolus gralla Giesenh. =Pyrrosia porosa MAN 

Niphobolus linearifolius Hook. =Pyrrosia linearifolia MAN 

Niphobolus lingua (Thunb.) =Pyrrosia lingua MAN 

Spreng. 

Niphobolus polydactylon =Pyrrosia polydactyla MAN 

(Hance) Giesenh. ex Diels 

Niphobolus porosus C. Presl =Pyrrosia porosa MAN 

(nom. nud.) 

ONOCLEACEAE 

=DRYTERIDACEAE -- 

(Subfamily 

Athyrioideae / Tribe 

Onocleeae) 

Onychium contiguum auct. non =Onychium lucidum MAN 

Hope: FOT 1975 

Onychium japonicum var. =Onychium lucidum MAN 

lucidum (D. Don) Christ 

Ophiala zeylanica (L.) Desv. =Helminthostachys MAN 

zeylanica 

Ophioderma pendula (L.) C. =Ophioglossum MAN 

Presl 

pendulum 

Ophioderma pendula f. ramosa =Ophioglossum FOC/MAN 

Nakai (type: Taiwan) 

pendulum 

Ophioderma pendula (L.) C. =Ophioglossum FOC 

Presl 

pendulum 

Ophioglossum japonicum =Lygodium japonicum MAN 

Thunb. 

Ophioglossum vulgatum auct. =Ophioglossum MAN 

non L.: FOT 1975 

austroasiaticum 

Ophioglossum vulgatum var. =Ophioglossum MAN 

thermale (Komarov) C. Chr. thermale 

Ophioglossum zeilanicum =Tectaria zeilanica MAN 

Houtt. 

Osmunda bromeliaefolia auct. =Osmunda banksiifolia FOT 2003 

non Copel. 

Osmunda cinnamomea var. =Osmunda 

MAN 

fokiense Copel. 

cinnamomea 

Osmunda claytoniana var. =Osmunda claytoniana MAN 

pilosa (Wall. ex Grev. & 

Hook.) Ching 

Osmunda lunaria L. =Botrychium lunaria MAN 

Osmunda pilosa Wall. ex Grev. 

& Hook. 

=Osmunda claytoniana MAN

Osmunda ternata Thunb. =Botrychium ternatum MAN 

Osmunda zeylanica L. 

=Helminthostachys MAN 

zeylanica 

Osmundastrum cinnamomeum =Osmunda 

MAN 

(L.) C. Presl 

cinnamomea 

Osmundastrum cinnamomeum =Osmunda 

MAN 

var. fokiense (Copel.) 

Tagawa 

cinnamomea 

Osmundastrum claytonianum =Osmunda claytoniana MAN 

(L.) Tagawa 

Osmundastrum japonicum =Osmunda japonica MAN 

(Thunb.) C. Presl 

Osmundopteris lanuginosa 


Nishida 

Pachypleuria repens (L. f.) M. 

Kato 

Pachypleuria trifoliata (Cav.) M. 

Kato 

Paesia taiwanensis W. C. 

Shieh 

Palhinhaea cernua (L.) Franco 

& Vasc. 

Paraceterach arifolia (nom. 

nud.) 

Paragymnopteris vestita 

(Hook.) K. H. Shing 

PARAHEMIONITIS 

Parahemionitis arifolia (Burm. 

f.) Panigrahi 

Paraleptochilus decurrens 


Parathelypteris angulariloba 

(Ching) Ching 

Parathelypteris angustifrons 

(Miq.) Ching 

Parathelypteris beddomei 

(Baker) Ching 

Parathelypteris castanea 


Parathelypteris glanduligera 

(Kunze) Ching 

Parathelypteris hirsutipes auct. 

non (C. B. Clarke) Ching 

=Botrychium 

lanuginosum 

=Davallia repens 

=Davallia cumingii 

=Paesia radula 

=Lycopodiella cernua 

MAN 

MAN 

MAN 

MAN 

MAN 

=Paraceterach cordata Knapp, R. 2011 

=Paraceterach vestita Knapp, R. 2011 

=Paraceterach, kept for -- 

P. cordata 


2013 

=Leptochilus decurrens MAN 

=Thelypteris 

angulariloba 

=Thelypteris 

angustifrons 

=Thelypteris beddomei 

=Thelypteris japonica 

=Thelypteris 

glanduligera 

=Thelypteris 

angulariloba 

MAN 

MAN 

MAN 

MAN 

MAN 

259 

(Kuo, C. M. 2010) 


260 

Parathelypteris japonica =Thelypteris japonica MAN 

(Baker) Ching 

Parathyrium unifurcatum =Deparia unifurcata MAN 

(Baker) Holtt. 

PARKERIACEAE =PTERIDACEAE -- 

Patania scabra (Wall. ex =Dennstaedtia scabra MAN 

Hook.) Bedd. 

Pellaea concolor (Langsd. & =Cheilanthes concolor MAN 

Fisch.) Baker 

Pellaea fauriei Christ =Histiopteris incisa MAN 

Pellaea henryi Christ =Cheilanthes nitidula MAN 

Pellaea nitidula (Hook.) Baker =Cheilanthes nitidula MAN 

Pentarhizidium orientalis =Onoclea orientalis MAN 

(Hook.) Hayata 

Peranema formosana Hayata =Peranema 

MAN 


cyatheoides 

Phegopteris auriculata J. Sm. =Cyclosorus 

FOT 2003/MAN 

auriculatus 

Phegopteris aurita auct. non 

(Hook.) J. Sm.: MAN 


levingei 


2013 

Phegopteris cuspidata (Blume) 

Mett. 


2011 

Phegopteris erubescens (Wall. =Cyclosorus 

MAN 

ex Hook.) J. Sm. 

erubescens 

Phegopteris eximia auct. non 

Mett. ex Kuhn: MAN 

=Polystichum 

scariosum 


2011 

Phegopteris maximowiczii =Monachosorum MAN 

(Baker) Christ 

maximowiczii 

Phegopteris ornata J. Sm. =Macrothelypteris MAN 

polypodioides 

Phegopteris paludosa auct. =Pseudophegopteris FOT 2003/MAN 

non (Blume) J. Sm. 

hirtirachis 

Phegopteris polypodioides Fee =Phegopteris 

FOC 

connectilis 

Phegopteris prolifera (Retz.) =Cyclosorus proliferus MAN 

Kuhn 

Phegopteris punctata (Thunb.) =Hypolepis punctata MAN 

Mett. 

Phegopteris pyrrhorachis var. =Pseudophegopteris MAN 

hirtirachis (C. Chr.) Tagawa hirtirachis 

Phegopteris simplex (Hook.) =Cyclosorus simplex MAN 

Mett. 

Phegopteris subaurita 

=Pseudophegopteris FOT 2003 


subaurita 

Phegopteris tenuifolia (G. 

Forst.) Keyserl. 

=Hypolepis tenuifolia MAN

261 

Phegopteris triphylla (Sw.) 

Mett. 

Phegopteris vulgaris Mett. 

Phlegmariurus carinatus (Desv. 

ex Poir.) Ching 

Phlegmariurus changii T. Y. 

Hsieh 

Phlegmariurus 

cryptomeri(a)nus (Maxim.) 

Ching 

Phlegmariurus 

cunninghamioides (Hayata) 

Ching 

Phlegmariurus fargesii (Herter) 

Ching 

Phlegmariurus fordii (Baker) 

Ching 

Phlegmariurus phlegmaria (L.) 

Holub 

Phlegmariurus salvinioides 

(Herter) Ching 

Phlegmariurus sieboldii (Miq.) 

Ching 

Phlegmariurus squarrosus (G. 

Forst.) A. Love & D. Love 

Phlegmariurus taiwanensis 

Ching (nom. nud.) ("type": 


Phyllitis scolopendrium (L.) 

Newman 

Phymatodes echinospora 


Phymatodes engleri (Luerss.) 

Ching 

Phymatodes engleri var. 

coriacea Tagawa (type: 


Phymatodes hastata (Thunb.) 

Ching 

Phymatodes longissima 


Phymatodes nigrescens 


Phymatodes okamotoi Tagawa 


=Cyclosorus triphyllus 

=Phegopteris 

connectilis 

=Huperzia carinata 

=Huperzia changii 

=Huperzia 

cryptomerina 

=Huperzia 


=Huperzia fargesii 

=Huperzia fordii 

=Huperzia phlegmaria 

=Huperzia salvinioides 

=Huperzia sieboldii 

=Huperzia squarrosa 

=Huperzia 

cryptomerina 

MAN 

FOT 2003 

MAN 


MAN 

MAN 

MAN 

MAN 

MAN 

MAN 

MAN 

MAN 


2011 

=Asplenium 

MAN 

scolopendrium 

=Selliguea echinospora MAN/Knapp, R. 

2011 


2011 

=Selliguea engleri FOC 

=Selliguea hastata MAN/Knapp, R. 

2011 


longissimus 



=Selliguea okamotoi MAN/Knapp, R. 

2011

262 

Phymatodes quasidivaricata 


Phymatodes rhynchophylla 

auct. non (Hook.) Ching: 

MAN 

Phymatodes scolopendria 

(Burm. f.) Ching 

Phymatodes taeniata var. 

palmata auct. non (Blume) 

Ching 

Phymatodes taiwanensis 


Phymatodes takedae Nakai 


Phymatodes yakushimensis 


Phymatopsis echinospora 

(Tagawa) H. Ito 

Phymatopsis engleri (Luerss.) 

H. Ito 

Phymatopsis hastata (Thunb.) 

Kitag. ex H. Ito 

Phymatopsis quasidivaricata 


Phymatopsis rhynchophylla 

auct. non (Hook.) J. Sm. 

Phymatopsis taeniata var. 

palmata auct. non (Blume) 

Ching: MAN 

Phymatopsis taiwanensis 


Phymatopsis yakushimensis 

(Makino) H. Ito 

Phymatopteris echinospora 

(Tagawa) Pic. Serm. 

Phymatopteris engleri (Luerss.) 

Pic. Serm. 

Phymatopteris hastata 

(Thunb.) Pic. Serm. 

Phymatopteris palmata auct. 

non (Blume) Pic. Serm.: MAN 

Phymatopteris quasidivaricata 

(Hayata) Pic. Serm. 

Phymatopteris rhynchophylla 

auct. non (Hook.) Pic. Serm.: 

MAN 

=Selliguea 


quasidivaricata 2011 


=Phymatosorus 

scolopendria 

=Selliguea 


MAN 

Kuo, C. M. 


2013 


2011 

=Microsorum henryi FOT 2003/MAN 

=Selliguea 

yakushimensis 


2011 


2011 


2011 


2011 

=Selliguea 




=Selliguea 




2011 

=Selliguea 




2011 


2011 


2011 

=Selliguea 



=Selliguea 



=Selliguea okamotoi Knapp, R. 2011

Phymatopteris taeniata auct. 

non (Sw.) Pic. Serm.: Kuo, C. 

M. 2010 

Phymatopteris taiwanensis 

(Tagawa) Pic. Serm. 

Phymatopteris yakushimensis 

(Makino) Pic. Serm. 

Phymatosorus nigrescens 

(Blume) Pic. Serm. 

=Selliguea 




2011 

=Selliguea 





Plagiogyria dunnii Copel. =Plagiogyria falcata MAN 

Plagiogyria euphlebia var. =Plagiogyria euphlebia MAN 

grandis (Copel.) De Vol 

Plagiogyria formosana Nakai =Plagiogyria glauca MAN 


Plagiogyria formosana var. =Plagiogyria glauca MAN 

angustata Nakai (type: 


Plagiogyria glauca subsp. 

formosana (Nakai) T. 

Nakaike 

=Plagiogyria glauca 

MAN 

Plagiogyria glauca var. =Plagiogyria glauca MAN 

philippinensis Christ 

Plagiogyria grandis Copel. =Plagiogyria euphlebia MAN 

Plagiogyria hayatana Makino =Plagiogyria falcata MAN 


Plagiogyria henryi Christ =Plagiogyria stenoptera MAN 

Plagiogyria intermedia Copel. =Plagiogyria japonica FOT 2003 

Plagiogyria matsumureana ?=Plagiogyria koidzumii (MAN) 

Makino 

Plagiogyria parva DeVol ex F. 

S. Liew (nom. nud.) ("type": 


=Plagiogyria adnata MAN 

Plagiogyria rankanensis 


Plagiogyria semicordata subsp. 

matsumureana (Makino) T. 

Nakaike 

=Plagiogyria adnata 

?=Plagiogyria koidzumii 

MAN 

(MAN) 

Plagiogyria tenuifolia Copel. =Plagiogyria falcata MAN 

Plananthus squarrosa (G. =Huperzia squarrosa MAN 

Forst.) P. Beauv. 

Plectopteris gracilis Fee =Calymmodon gracilis FOT 2003 

Plenasium banksiifolium (C. =Osmunda banksiifolia MAN 

Presl) C. Presl 

Plenasium claytonianum (L.) C. 

Presl 

=Osmunda claytoniana MAN 

263

264 

Pleocnemia aspidiodes 

(Blume) Mett. 

=Pleocnemia 


Kuo, C. M. 


2011 

MAN 

Pleocnemia cumingiana auct. =Pleocnemia rufinervis 

non C. Presl: FOT 1975 

Pleocnemia devexa (Kunze) =Tectaria devexa FOT 2003 

Alderw. 

Pleocnemia winitii Holtt. =Pleocnemia 

MAN 


Pleopeltis dilatata Bedd. =Microsorum insigne MAN 

Pleopeltis insignis (Blume) =Microsorum insigne MAN 

Bedd. 

Pleopeltis rostratum Bedd. =Lemmaphyllum MAN 

rostratum 

Pleopeltis thunbergianus Kaulf. =Lepisorus 

MAN 

thunbergianus 

Pleurogramme paradoxa Fee =Monogramma MAN 

paradoxa 

Pleuromanes pallidum (Blume) =Crepidomanes MAN 

C. Presl 

pallidum 

Pneumatopteris truncata (Poir.) =Cyclosorus truncatus MAN 

Holtt. 

Polybotrya appendiculata =Bolbitis appendiculata FOT 2003 


Polybotrya duplicato-serrata =Bolbitis rhizophylla MAN 


Polybotrya marginata Blume =Bolbitis appendiculata FOT 2003 

Polybotrya serrulata J. Sm. =Bolbitis rhizophylla FOT 2003 

Polypodiastrum mengtzeense =Polypodium 

MAN 


mengtzeense 

Polypodiastrum taiwanianum =Polypodium 

MAN 


mengtzeense 

Polypodiodes formosana =Polypodium 

MAN 

(Baker) Ching 

formosanum 

Polypodiodes microrhizoma (C. =Polypodium 

MAN 

B. Clarke ex Baker) Ching microrhizoma 

Polypodioides niponica (Mett.) 

Ching 

Polypodiodes raishanensis 

(Rosenst.) S. G. Lu 

Polypodioides transpianensis 

(Yamam.) Saiki 

=Polypodium 


=Polypodium 


=Polypodium 

raishaense 

=Polypodium 

transpianense 



R. 2013 

alternative 

combination 

published in CHK 

MAN

265 

Polypodium acuminatum Houtt. =Cyclosorus 

MAN 

acuminatus 

Polypodium adnascens Sw. =Pyrrosia lanceolata MAN 

Polypodium adspersum Blume =Grammitis adspera MAN 

Polypodium angustissimum =Pyrrosia angustissima MAN 

Baker 

Polypodium argutum auct. non =Polypodium 

MAN 

Wall. ex Hook.: FOT 2003 mengtzeense 

Polypodium argutum var. =Polypodium 

MAN 

mengtzeense (Christ) Christ mengtzeense 

Polypodium arisanense Hayata =Polypodium amoenum MAN 


Polypodium arisanense 

Rosenst. (type Taiwan) 



347 (1915) 

Polypodium aristatum G. Forst. =Arachniodes aristata MAN/Knapp, R. 

2011 

Polypodium asperum auct. non =Cyclosorus 

FOT 2003/MAN 

C. Presl 


Polypodium aspidistrifrons =Microsorum steerei FOT 2003 


Polypodium atkinsonii auct. =Polypodium 

MAN 

non C. Chr.: FOT 1975 

microrhizoma 

Polypodium auriculatum L. =Nephrolepis cordifolia MAN 

Polypodium austriacum auct. 

non Jacq.: MAN 

=Dryopteris expansa MAN/Knapp, R. 

2011 

Polypodium barometz L. =Cibotium barometz MAN 

Polypodium beddomei Baker =Thelypteris beddomei this publication 

Polypodium brachylepis Baker =Microsorum 

superficiale 


2013 

Polypodium buergerianum 

auct. non Miq.: FOT 2003 

=Microsorum 

superficiale 


R. 2013 

Polypodium caudiceps (T. =Lepisorus 

MAN 

Moore) Baker 


Polypodium christii Copel. =Grammitis jagorianum FOT 2003 

Polypodium clathratum C. B. =Lepisorus clathratus MAN 

Clarke 

Polypodium congenerum =Grammitis congener MAN 

(Blume) C. Presl 

Polypodium conjugatum Kaulf. =Dipteris conjugata MAN 

Polypodium connectile Michx. =Phegopteris 

MAN 

connectilis 

Polypodium contiguum (G. =Grammitis contigua MAN 

Forst.) J. Sm. 

Polypodium cordifolium L. =Nephrolepis cordifolia MAN

266 

Polypodium coronans Wall. ex =Aglaomorpha 

MAN 

Mett. 

coronans 

Polypodium crenatum Forssk. =Hypodematium MAN 

crenatum 

Polypodium cucullatum Nees & =Calymmodon 

MAN 

Blume 

cucullatus 

Polypodium curtisii Baker =Grammitis curtisii MAN 

Polypodium decrescens var. 

blechnifrons Hayata (syntype: 

=Grammitis curtisii FOT 2003/Knapp, 

R. 2011 


Polypodium decursivepinnatum =Phegopteris 

MAN 

H. C. Hall 


Polypodium dentatum Forssk. =Cyclosorus dentatus MAN 

Polypodium deorsipinnatum =Polypodium 

MAN 

Copel. (type: Taiwan) 

microrhizoma 

Polypodium dilatatum Wall. ex =Microsorum insigne MAN 

Hook. 

Polypodium dimorphum Baker ?=Tectaria harlandii FOC 


Polypodium dipteris Blume =Dipteris conjugata this publication 

Polypodium dissectum G. =Tectaria dissecta MAN 

Forst. 

Polypodium divaricatum 


=Selliguea 



2011 

Polypodium diversum Rosenst. =Lemmaphyllum MAN 


rostratum 

Polypodium drepanopterum =Athyrium mupinense MAN/Liu, Y. C. et 

auct. non Kunze: MAN 

Polypodium ellipticum var. 

pothifolium auct. non (Buch.- 

Ham. ex D. Don) Makino: 

FOT 2003 

al. 2009 


2011 

Polypodium engleri Luerss. =Selliguea engleri MAN/Knapp, R. 

2011 

Polypodium engleri var. 

hypoleucum Hayata (type: 


=Selliguea engleri FOC 

Polypodium engleri var. 

yakushimense Makino 

Polypodium ensato-sessilifrons 


Polypodium ensatum Thunb. 

Polypodium erubescens Wall. 

ex Hook. 

=Selliguea 

yakushimensis 

=Colysis hemionitidea 

=Neocheiropteris 

ensata 

=Cyclosorus 

erubescens 


2011 

MAN 

MAN 

MAN

Polypodium excavatum var. =Lepisorus 

FOT 2003 

bicolor Takeda 

morrisonensis 

Polypodium falcatopinnatum =Selliguea 




Polypodium falcatum L. f. =Polystichum falcatum MAN 

Polypodium fortunei Kunze ex =Drynaria fortunei MAN 

Mett. 

Polypodium fragile L. =Cystopteris fragilis MAN 

Polypodium glandulosum 

Hook. (1863) non Desv. 


912-914 (1998) 

(1811) 

Polypodium glaucum Thunb. =Diplopterygium MAN 

ex Houtt. 

glaucum 

Polypodium gracillimum Copel. =Calymmodon 

MAN 

cucullatus 

Polypodium hancockii Baker =Microsorum insigne MAN 


Polypodium hainanense C. 

Chr. (syntype: ?Taiwan) 

267 


in: Index Filic. 531 

(1906) 

Polypodium hasseltii Blume =Dryopteris hasseltii MAN 

Polypodium hastatum Thunb. =Selliguea hastata MAN/Knapp, R. 

2011 

Polypodium hastatum var. 

engleri (Luerss.) Christ 


2011 

Polypodium hayatai Masam. =Grammitis doniana FOT 2003/Knapp, 

R. 2011 

Polypodium henryi Christ =Microsorum henryi MAN 

Polypodium hoozanense =Lepisorus tosaensis FOC 


Polypodium hypochrysum =Lepisorus megasorus FOT 2003 


Polypodium infraplanicostale =Lepisorus tosaensis FOT 2003 


Polypodium insigne Blume =Microsorum insigne MAN 

Polypodium intromissum Christ =Grammitis intromissa MAN 

Polypodium jagorianum Mett. =Grammitis jagoriana FOT 2003 

ex Kuhn 

Polypodium kanashiroi Hayata ?=Grammitis contigua MAN 

Polypodium kawakamii Hayata =Lepisorus kawakamii MAN 


Polypodium kuchenense Y. C. =Lepisorus 

MAN 

Wu 

kuchenensis 

Polypodium kusukusense =Colysis wrightii MAN 


Polypodium lehmanni Mett. =Arthromeris lehmannii FOT 2003

268 

Polypodium leptophyllum L. =Anogramma 

MAN 

leptophylla 

Polypodium lineare Burm. f. =Dicranopteris linearis MAN 

Polypodium lineare var. =Lepisorus tosaensis FOT 2003 

caudatum Makino 

Polypodium lineare var. =Lepisorus monilisorus MAN 

monilisorum Hayata (type: 


Polypodium linearifolium =Pyrrosia linearifolia MAN 

(Hook.) Hook. 

Polypodium lingua (Thunb.) =Pyrrosia lingua MAN 

Sw. 

Polypodium longissimum =Phymatosorus MAN 

Blume 

longissimus 

Polypodium loxogramme var. =Loxogramme 

MAN 

lamprocaulon Rosenst. (type: 


grammitoides 

Polypodium marginatum Houtt. =Microlepia marginata MAN 

Polypodium maximowiczii =Monachosorum MAN 

Baker 

maximowiczii 

Polypodium megasorum C. =Lepisorus 

MAN 

Chr. 

megaphyllus 

Polypodium membranaceum =Microsorum 

MAN 

D. Don 

membranaceum 

Polypodium membranifolium R. =Phymatosorus MAN 

Br. 


Polypodium merrittii Copel. =Grammitis glandulosa Kew Bull. 53(4): 

912-914 (1998) 

Polypodium meyenianum =Aglaomorpha 

MAN 

(Schott) Hook. 

meyeniana 

Polypodium mollicomum Nees =Grammitis mollicoma MAN 

& Blume 

Polypodium morii Hayata (type: =Lepisorus tosaensis FOT 2003 


Polypodium morrisonense =Lepisorus 

MAN 


morrisonensis 

Polypodium nigrescens Blume =Phymatosorus MAN 

Polypodium niponicum Mett. 


=Polypodium 


=Polypodium 




R. 2013 

Polypodium normale auct. non =Microsorum henryi FOT 2003/MAN 

D. Don 

Polypodium obliquatum Blume =Grammitis obliquata MAN

269 

Polypodium obscurevenulosum =Lepisorus 

MAN 



Polypodium obtusifrons Hayata 


? published in: Icon. 


250 (1914) 

Polypodium okuboi Yatabe =Grammitis okuboi MAN 

Polypodium omeiense Baker =Cyclosorus omeiensis MAN 

Polypodium ornatum Wall. ex =Macrothelypteris MAN 

Bedd. 

polypodioides 

Polypodium oyamense Baker =Gymnocarpium MAN 

oyamense 

Polypodium palmatum auct. =Selliguea 


non Blume: MAN 


Polypodium papakense =Lepisorus clathratus FOT 2003 

Masam. 

Polypodium parasiticum L. =Cyclosorus 

MAN 

parasiticus 

Polypodium pellucidifolium 


?=Selliguea engleri published in: Icon. 


250 (1914) 

Polypodium phegopteris L. =Phegopteris 

FOT 2003 

connectilis 

Polypodium phymatodes L. =Phymatosorus FOT 2003/MAN 

scolopendria 

Polypodium pinnatum Hayata 


=Arthromeris lehmannii MAN/Knapp, R. 

2011 

Polypodium polydactylon =Pyrrosia polydactyla MAN 

Hance (type: Taiwan) 

Polypodium pseudocucullatum =Grammitis okuboi FOC/MAN 


Polypodium 

pseudoformosanum Tagawa 


Polypodium 

pseudotrichomanoides 


?=Polypodium 

raishaense 


MAN 

MAN 

Polypodium pteropus Blume =Microsorum pteropus MAN 

Polypodium punctatum Thunb. =Hypolepis punctata MAN 

Polypodium pyrrhorhachis 

Kunze 


hirtirachis 


R. 2011 

Polypodium quasidivaricatum 

Hayata 

=Selliguea 



2011 

Polypodium reinwardtii (Blume) 

C. Presl 

=Grammitis reinwardtii MAN

270 

Polypodium remotefrondigerum 

Hayata (type: 


Polypodium rhynchophyllum 

auct. non Hook. 

Polypodium rostratum Hook. 

=Loxogramme 

remotefrondigera 

MAN 


=Lemmaphyllum MAN 

rostratum 

Polypodium rubidum Kunze =Phymatosorus MAN 

longissimus 

Polypodium scolopendria =Phymatosorus MAN 

Burm. f. 

scolopendria 

Polypodium scottii Bedd. =Dryopteris scottii MAN 

Polypodium setigerum Blume =Grammitis intromissa MAN 

Polypodium sheareri Baker =Pyrrosia sheareri MAN 

Polypodium shintenense =Colysis x shintenensis MAN 


Polypodium speluncae L. =Microlepia speluncae MAN 

Polypodium stenolepis Baker =Dryopteris stenolepis MAN 

Polypodium steerei Harr. =Microsorum steerei MAN 

Polypodium subevenosum =Grammitis adspera FOT 2003 

Baker 

Polypodium subfalcatum =Grammitis doniana MAN 

Blume 

Polypodium subtriphyllum =Tectaria subtriphylla MAN 

Hook. & Arn. 

Polypodium suishastagnale =Phymatosorus MAN 


longissimus 

Polypodium superficiale Hook. =Microsorum 

FOT 2003/MAN 

superficiale 

Polypodium taiwanense Christ =Pyrrosia lingua MAN 


Polypodium taiwanianum =Polypodium 

MAN 


mengtzeense 

Polypodium tenuisectum =Grammitis tenuisecta MAN 

Blume 

Polypodium tenuissimum 


=Grammitis doniana FOT 2003/Knapp, 

R. 2011 

(non Copel.) 

Polypodium thunbergianum =Lepisorus 

MAN 

(Kaulf.) C. Chr. 

thunbergianus 

Polypodium tosaense Makino =Lepisorus tosaensis MAN 

Polypodium truncatum Poir. =Cyclosorus truncatus MAN 

Polypodium urceolare Hayata =Grammitis urceolaris MAN 


Polypodium varium L. =Dryopteris varia MAN

271 

Polypodium varium var. 

eurylepidota Rosenst. (type: 


Polypodium wrightii (Hook.) 

Mett. ex Diels 

Polypodium wrightii var. lobata 


Polypodium yakushimense 

(Makino) Makino 

Polystichum aculeatum var. 

durissima Rosenst. (type: 



formosanum Kodama (type: 



nigropaleaceum non Christ: 

MAN 


variiforme Hayata (type: 


Polystichum acuminatum auct. 

non (Diels) C. M. Kuo: MAN, 


Polystichum amabile var. 

chinense Rosenst. 

Polystichum arisanicum 


Polystichum aristatum (G. 

Forst.) C. Presl 

Polystichum atroviridissimum 


Polystichum attenuatum var. 

subattenuatum (Ching et W. 

M. Chu) W. M. Chu et Z. R. 

He 

Polystichum auriculatum (L.) C. 

Presl 

Polystichum auriculatum var. 

stenophyllum (Baker) 

Matthew 

Polystichum constantissimum 


Polystichum deltodon var. 

pseudodeltodon (Tagawa) 

Tagawa 

=Dryopteris varia Kuo, C. M. 1997 

=Colysis wrightii 

=Colysis wrightii 

=Selliguea 

yakushimensis 

=Polystichum 

biaristatum 

=Polystichum 

mucronifolium 

=Polystichum 

piceopaleaceum 

=Polystichum 

scariosum 

MAN 

MAN 


2011 

MAN 


2011 


C. R. 2008 


R. 2011 

=Polystichum simadae Knapp, R. 2013 

=Arachniodes 

chinensis 

=Arachniodes globisora 

=Arachniodes aristata 

=Polystichum 

nepalense 

=Polystichum 

attenuatum 

=Nephrolepis cordifolia 

=Polystichum 

hecatopterum 


MAN 


FOT 2003 

Chang, Y. H. et 


MAN 

FOT 2003 

=Dryopteris formosana FOT 2003 

=Polystichum deltodon FOT 2003

272 

Polystichum diffractum (Baker) =Dryopteris diffracta FOC 

Masam. 

Polystichum dimorphophyllum =Polystichum deltodon MAN 

Hayata 

Polystichum discretum auct. 

non (Buch.-Ham. ex D. Don) 

=Polystichum 



C. R. 2008 

J. Sm.: MAN 

Polystichum duthiei auct. non =Polystichum glaciale Knapp, R. 2011 

(C. Hope) C. Chr.: MAN 

Polystichum duthiei var. 

subbipinnatum Tagawa (type: 

=Polystichum glaciale MAN/Knapp, R. 

2011 


Polystichum eximium auct. non =Polystichum 


(Mett. ex Kuhn) C. Chr.: MAN scariosum 

Polystichum eximium var. 

minus Tagawa 

=Polystichum 

scariosum 


R. 2011 

Polystichum falcatipinnum =Polystichum 

MAN 


manmeiense 

Polystichum falcatum f. 

acuminatum auct. non Diels: 

=Polystichum simadae MAN/Knapp, R. 

2013 

MAN, Knapp, R. 2011 

Polystichum falcatum var. 

acuminatum auct. non Diels 

=Polystichum simadae FOT 2003/Knapp, 

R. 2013 

Polystichum falcatum var. =Polystichum fortunei MAN 

fortunei (J. Sm.) Matsum. 

Polystichum falcatum var. =Polystichum 

MAN 

macrophyllum (Makino) 

Matsum. 

macrophyllum 

Polystichum fukuyamae =Polystichum sinense FOT 2003/MAN 


Polystichum gemmiferum 

Tagawa 

=Polystichum 

scariosum 


R. 2011 

Polystichum gladiipinnum =Polystichum 

FOT 2003 


xiphophyllum 

Polystichum globisorum =Arachniodes globisora MAN 


Polystichum grande Ching =Polystichum 

MAN 

grandifrons 

Polystichum hasseltii Blume =Dryopteris hasseltii FOT 2003/MAN 

Polystichum hookerianum (C. =Polystichum 

MAN 


integripinnum 

Polystichum hololepis Hayata =Dryopteris varia FOT 2003 


Polystichum horidipinnum 


=Polystichum 


FOT 2003

273 

Polystichum ilicifolium auct. 

non (D. Don) T. Moore 

Polystichum inaense (Tagawa) 

Tagawa 

Polystichum kiusiuense 

Tagawa 

Polystichum kodamae Tagawa 

Polystichum lentum var. gelida 


Polystichum lepidocaulon var. 

appendiculatum Nakai 

Polystichum leptopteron 


Polystichum lobatum var. 

chinense Christ 

Polystichum longistipes Hayata 


Polystichum maximowiczii 

(Baker) Diels 

Polystichum maximowiczii var. 

melanocaulon Hayata (type: 


Polystichum mayebarae 

Tagawa 

Polystichum microindusium 

(nom. nud.) 

Polystichum morii Hayata 


Polystichum neolobatum var. 

brevipinnum Tagawa (type: 


Polystichum nervosum (Ching 

& K. H. Shing) C. M. Kuo 

Polystichum nigropaleaceum 

auct. non (Christ) Diels: MAN 

Polystichum nigrospinosum 

Ching 

Polystichum niitakayamense 


Polystichum obtusoauriculatum 

Hayata (type: 


Polystichum pseudoaristatum 

Tagawa 

=Polystichum 

FOT 2003 


=Polystichum capillipes Knapp, R. 2011 

=Polystichum 

grandifrons 

=Polystichum 

mucronifolium 

=Polystichum 

prionolepis 

=Polystichum 

lepidocaulon 

=Polystichum hancockii 

=Polystichum 

neolobatum 


=Monachosorum 

maximowiczii 

=Monachosorum 

maximowiczii 

MAN 


2011 

MAN 

FOT 2003 

MAN 

FOT 2003 

MAN 

MAN 

MAN 

=Polystichum 

FOT 2003 

tsussimense 

=Polystichum 


anomophyllum 

=Polystichum atkinsonii Knapp, R. 2011 

=Polystichum 

neolobatum 

=Polystichum 

anomophyllum 

=Polystichum 


=Arachniodes 

nigrospinosa 

=Polystichum 

pseudostenophyllum 

=Polystichum 

formosanum 

=Arachniodes 


FOT 2003 



C. R. 2008 

MAN 


MAN 

MAN

274 

Polystichum pseudoaristatum 

var. acutum Tagawa 

Polystichum pseudodeltodon 


Polystichum 

pseudomaximowiczii Hayata 


Polystichum rectipinnum 


Polystichum rhomboideum 

Schott 

Polystichum simplicipinnum 


Polystichum simplicius var. 

majus Tagawa 

Polystichum sozanense Ching 

ex H. S. Kung & L. B. Zhang 

Polystichum sp. 2 in Knapp, R. 

2013 

Polystichum spinescens 


Polystichum stenophyllum var. 

abbreviatum Tagawa (type: 


Polystichum subapiciflorum 


Polystichum subattenuatum 

Ching et W. M. Chu 

Polystichum subauriculatum 


Polystichum subobliquum 


Polystichum tacticopterum 

(Kunze) T. Moore 

Polystichum taiwanense 

(Tagawa) C. M. Kuo (mistyping 

of species epithet) 

Polystichium torresianum 

Gaudich. 

Polystichum tosaense (Makino) 

Makino 

Polystichum transmorrisonense 


Polystichum tsus-simense var. 

mayebarae (Tagawa) Kurata 

=Arachniodes 

FOT 1994 


=Polystichum deltodon FOT 2003 

=Polystichum 

hecatopterum 

=Polystichum 

prionolepis 

=Arachniodes 

rhomboidea 


MAN 

MAN 

MAN 

MAN 

=Arachniodes caudata Knapp, R. 2013 

?=Polystichum 

parvipinnulum 

=Polystichum tenuius 

=Polystichum 


?=Polystichum 

pseudostenophyllum 



FOT 2003 


R. 2011 

=Polystichum 

MAN 

biaristatum 

=Polystichum 

Chang, Y. H. et 

attenuatum 


=Polystichum acutidens FOT 2003 

=Polystichum obliquum FOT 2003 

=Polystichum 

mucronifolium 

=Polystichum 

taiwanianum 



=Macrothelypteris MAN 

torresiana 

=Polystichum deltodon FOT 2003 

=Dryopsis 


=Polystichum 

tsussimense 

MAN 

FOT 2003

275 

Polystichum variiforme 


=Polystichum 

scariosum 


R. 2011 

Polystichum varium var. =Dryopteris varia MAN 

eurylepidota Rosenst. (type: 


Polystichum wilsonii Christ =Polystichum sinense MAN 

Pronephium aspera (C. Presl) =Cyclosorus 

MAN 

W. C. Shieh & J. L. Tsai 

(nom. superfl.) 


Pronephium asperum auct. non =Cyclosorus 

MAN 

(C. Presl) Holtt. 


Pronephrium cuspidatum 

(Blume) Holtt. 


2011 

Pronephrium gymnopteridifrons =Cyclosorus 

MAN 

(Hayata) Holtt. 


Pronephium insularis (K. =Cyclosorus insularis MAN 

Iwats.) Holtt. 

Pronephium longipetiolatum (K. =Cyclosorus 

MAN 

Iwats.) Holtt. 


Pronephrium parishii (Bedd.) 

Holtt. 


2011 

Pronephium simplex (Hook.) =Cyclosorus simplex MAN 

Holtt. 

Pronephrium triphyllum var. 

parishii (Bedd.) T. Nakaike 


2011 

Pronephrium triphyllum (Sw.) =Cyclosorus triphyllus MAN 

Holtt. var. triphyllum 

PROSAPTIA =Grammitis, kept for P. -- 

celebica, P contigua, 

P. nutans, P. 

urceolaris 

Prosaptia celebica (Blume) =Grammitis celebica this publication 

Tagawa & K. Iwats. 

Prosaptia contigua (G. Forst.) =Grammitis contigua Knapp, R. 2011 

C. Presl 

Prosaptia kanashiroi (Hayata) ?=Grammitis contigua MAN 

Nakai ex Yamamoto 

Prosaptia khasiana auct. non =Grammitis urceolaris this publication 

(Hook.) C. Chr.: Kuo, C. M. 

2010 

Prosaptia obliquata (Blume) =Grammitis obliquata MAN 

Mett. 

Prosaptia urceolaris (Hayata) =Grammitis urceolaris Knapp, R. 2011 

Copel. 

Protangiopteris somai (Hayata) 

Hayata 

=Angiopteris somae MAN

276 

Pseudocyclosorus esquirolii =Cyclosorus esquirolii MAN 


Pseudocyclosorus tylodes 

(Kunze) Ching 

=Cyclosorus tylodes FOC/Knapp, R. 

2011 

Pseudocyclosorus xylodes =Cyclosorus tylodes Knapp, R. 2011 

(Kunze) Ching (mis-spelling 

of species epithet) 

Pseudocystopteris atkinsonii 

(Bedd.) Ching 

=Athyrium atkinsonii Liu, Y. C. et al. 

2009 

Pseudodrynaria coronans =Aglaomorpha 

MAN 

(Wall. ex Mett.) Ching 

coronans 

Pseudophegopteris aurita auct. =Pseudophegopteris Knapp, R. 2013 

non (Hook.) Ching: MAN levingei 

Pseudophegopteris paludosa =Pseudophegopteris MAN 

auct. non (Blume) Ching: 

FOT 2003 

hirtirachis 

Pteridium aquilinum var. =Pteridium revolutum FOT 2003/MAN 

lanuginosum auct. non Bory. 

Pteridium aquilinum subsp. =Pteridium latiusculum MAN 

latiusculum (Desv.) W. C. 

Shieh 

Pteridium aquilinum subsp. =Pteridium latiusculum MAN 

aquilinum var. latiusculum 

(Desv.) Under ex A. Heller 

Pteridium aquilinum subsp. =Pteridium revolutum MAN 

wightianum (J. Agardh) W. C. 

Shieh 

Pteridium aquilinum subsp. =Pteridium revolutum MAN 

aquilinum var. wightianum (J. 

Agardh) R. M. Tryon 

Pteris aquilina L. =Pteridium sp. FOT 2003 

Pteris argentea S. G. Gmel =Cheilanthes argentea MAN 

Pteris confertinervia Ching ex 

Ching & S. H. Wu (type: 

=Pteris cretica subsp. 

cretica 


C. R. 2008 


Pteris concolor Langsd. & =Cheilanthes concolor MAN 

Fisch. 

Pteris cretica var. albo-lineata =Pteris nipponica Knapp, R. 2011 

T. Moore 

Pteris cretica var. laeta (Wall. 

ex Ettingsh.) C. Chr. & 

Tardieu 


laeta 


2011 

Pteris decurrenti-pinnulata =Pteris formosana FOT 2003 

Bonap. 

Pteris dimidiata Willd. =Pteris semipinnata Fraser-Jenkins, 

C. R. 2008

Pteris dissitifolia auct. non (?) =Pteris semipinnata Knapp, R. 2011 

Baker: CHK 

Pteris ensiformis var. 

=Pteris grevilleana MAN 

grevilleana C. B. Clarke ex 

Bedd. 

Pteris esquirolii auct. non =Pteris pellucidifolia Knapp, R. 2013 

Christ 

Pteris excelsa Gaudich. =Pteris terminalis Knapp, R. 2013 

Pteris excelsissima Hayata 


=Pteris terminalis MAN/Knapp, R. 

2013 

Pteris farinosa auct. non. 

Forssk. 

=Cheilanthes agetae in 

part 

FOT 1975/Saiki 

1984/MAN 

Pteris fauriei var. minor Hieron. =Pteris fauriei 

MAN 


Pteris flavicaulis Hayata (type: =Pteris biaurita FOT 2003 


Pteris guangdongensis auct. =Pteris plumbea Knapp, R. 2011 

non Ching 

Pteris hirsuta Poir. 

=Cheilanthes 

MAN 

nudiuscula 

Pteris incisa Thunb. =Histiopteris incisa MAN 

Pteris interrupta Willd. =Cyclosorus interruptus MAN 

Pteris laeta Wall. ex Ettingsh. =Pteris cretica subsp. 

laeta 


2011 

Pteris latiuscula Desv. =Pteridium latiusculum MAN 

Pteris linearis auct. non Poir.: =Pteris arisanensis Knapp, R. 2011 

MAN 

Pteris linearis var. fauriei =Pteris fauriei 

MAN 

(Hieron.) C. Chr. & Tardieu 

Pteris longifolia auct. non L. =Pteris vittata FOT 2003 

Pteris longifolia var. vittata (L.) =Pteris vittata 


Nic. 

Pteris matsudai Masam. =Pteris pellucida MAN 

Pteris morrisonicola Hayata =Pteris wallichiana MAN 


Pteris nervosa Thunb. 


cretica 


2011 

Pteris nudiuscula R. Br. =Cheilanthes 

MAN 

nudiuscula 

Pteris parkeri hort. (of garden =Pteris nipponica Knapp, R. 2011 

origin) 

Pteris pellucens Ag. =Pteris longipes FOT 2003 

Pteris plumbea var. sintenensis 

Masam. 

=Pteris plumbea Knapp, R. 2011 

277

278 

Pteris quadriaurita var. 

=Pteris fauriei 

MAN 

abbreviata Rosenst. (type: 


Pteris radula Baker =Paesia radula MAN 

Pteris recurvata var. wightiana =Pteridium revolutum MAN 

J. Agardh 

Pteris revoluta Blume =Pteridium revolutum MAN 

Pteris scabristipes Tagawa =Pteris aspericaulis Knapp, R. 2011 


Pteris semipinnata auct. non 

L.: FOT 1975 

=Pteris dispar MAN/Knapp, R. 

2011 

Pteris semipinnata var. dispar 

(Kunze) Hook. & Baker 

=Pteris dispar MAN/Knapp, R. 

2011 

Pteris serrulata var. obtusata =Pteris ensiformis MAN 

Christ (type: Taiwan) 

Pteris siliculosa Desv. =Onychium siliculosum MAN 

Pteris stelleri S. G. Gmel. =Cryptogramma stelleri MAN 

Pteris taiwanensis Ching in ?=Pteris semipinnata 

Ching & S. H. Wu 

Pteris taiwanensis Masam. & 

Suzuki 

Pteris takeoi Hayata (type: 


Pteris thalictroides (L.) Sw. 

(MAN, FOC) 


Bot. Austro Sin. 1: 

16 (1983) 

=Pteris dispar FOT 2003 

=Pteris formosana 

MAN 

=Ceratopteris 

MAN 

thalictroides 

Pteris tokioi Masam. =Pteris amoena Knapp, R. 2013 

Pteris trifoliata Christ =Pteris deltodon FOT 2003 

Pteris venusta Kunze =Pteris pellucida Knapp, R. 2011 

Pteris wightiana Wall. (nom. =Pteridium revolutum FOT 2003/MAN 

nud.) 

Pteris x wulaiensis C. M. Kuo =Pteris wulaiensis this publication 

Ptilopteris hancockii Hance =Polystichum hancockii MAN 

Ptilopteris maximowiczii =Monachosorum MAN 

(Baker) Hance 

maximowiczii 

Pyrrosia adnascens (Sw.) =Pyrrosia lanceolata MAN 

Ching 

Pyrrosia gralla (Giesenh.) =Pyrrosia porosa MAN 

Ching 

Pyrrosia grandissima (Hayata) =Pyrrosia sheareri MAN 

Ching 

Pyrrosia linearifolia var. 

heterolepis Tagawa (type: 


=Pyrrosia linearifolia MAN

Pyrrosia mollis auct. non =Pyrrosia matsudae MAN 

(Kunze) Ching: FOT 1975 

Pyrrosia pseudopolydactylis =Pyrrosia polydactyla MAN 

Seriz. (type: Taiwan) 

Pyrrosia sasakii Hayata (type: =Pyrrosia angustissima MAN 


Pyrrosia subfissa (Hayata) =Pyrrosia porosa MAN 

Ching 

Pyrrosia transmorrisonensis =Pyrrosia porosa Knapp, R. 2011 


Quercifilix zeilanica (Houtt.) 

Copel. 

=Tectaria zeilanica MAN/Knapp, R. 

2011 

Quercifilix zeylanica (Houtt.) =Tectaria zeilanica MAN 

Copel. (mis-spelling in 

species name) 

Reediella humile (G. Forst.) =Crepidomanes humile MAN 

Pic. Serm. 

RHACHIDOSORUS =Athyrium, kept for R. -- 

pulcher 

Rhachidosorus mesosorus 

auct. non (Makino) Ching 

=Athyrium pulchrum Knapp, R. 2011; 

probably a misidentification 

of a 

specimen from S 

Taiwan (J. L. 

Tseng 6602 in 

TAIF) 

Rhachidosorus pulcher =Athyrium pulchrum Knapp, R. 2011 


Rumohra amabilis var. 

=Arachniodes 

MAN 

yakusimensis H. Ito 

yakusimensis 

Rumohra arisanica (Rosenst.) =Arachniodes globisora MAN 

Ching 

Rumohra aristata (G. Forst.) 

Ching 

=Arachniodes aristata MAN/Knapp, R. 

2011 

Rumohra dimorphophylla =Polystichum deltodon MAN 


Rumohra diffracta (Baker) =Dryopteris diffracta FOC 

Ching 

Rumohra festina (Hance) =Arachniodes festina MAN 

Ching 

Rumohra globisora (Hayata) H. =Arachniodes globisora MAN 

Ito 

Rumohra hasseltii (Blume) =Dryopteris hasseltii MAN 

Ching 

Rumohra miyakei H. Ito (type: 


=Arachniodes 


FOT 2003 

279

280 

Rumohra nigrospinosa (Ching) =Arachniodes 

MAN 

Ching 

nigrospinosa 

Rumohra pseudoaristata =Arachniodes 

MAN 

(Tagawa) H. Ito 


Rumohra pseudoaristata var. =Arachniodes 

FOT 2003 

magnipinna H. Ito 


Rumohra quadripinnata =Arachniodes 

MAN 


quadripinnata 

Sagenia coadunata J. Sm. =Tectaria coadunata MAN 

Sagenia fauriei Tagawa =Tectaria fauriei FOT 2003 

Sagenia macrodonta Fee =Tectaria coadunata FOT 2003 

Sagenia pteropus T. Moore =Tectaria devexa MAN 

Sagenia subtriphylla (Hook. & =Tectaria subtriphylla FOT 2003 

Arn.) Bedd. 

Sagenia variolata auct. non =Tectaria phaeocaulis FOT 2003 

(Wall.) T. Moore 

Saxiglossum angustissimum =Pyrrosia angustissima MAN 

(Giesenh. ex Diels) Ching 

Saxiglossum sasakii (Hayata) =Pyrrosia angustissima MAN 

Tagawa 

Saxiglossum taenidoes var. =Pyrrosia angustissima MAN 

sasakii (Hayata) Masam. ex 

Satomi 

Sceptridium daucifolium auct. =Botrychium 

FOC 

non. (Wall. ex Hook. & Grev.) 

Lyon 

formosanum 

Sceptridium formosanum =Botrychium 

MAN 

(Tagawa) Holub 

formosanum 

Sceptridium ternatum (Thunb.) =Botrychium ternatum MAN 

Lyon 

Schellolepis persicifolia (Desv.) =Polypodium 


Pic. Serm. 

persicifolium 

Schizaea biroi auct. non (?) =Schizaea dichotoma Knapp, R. 2011 

Richter 

Schizaea kikuzatonis Ogata =Schizaea dichotoma FOT 2003 

Schizoloma ensifolia (Sw.) J. =Lindsaea ensifolia this publication 

Sm. 

Scleroglossum pusillum =Scleroglossum Knapp, R. 2013 

(Blume) Alderw. 

Selaginella atroviridis auct. non 


Spring 

Selaginella canaliculata (L.) 

Baker 

Selaginella caulescens (Wall.) 

Spring 

sulcatum 

=Selaginella 

doederleinii 

FOT 2003 

=Selaginella delicatula FOT 2003 

=Selaginella involvens FOT 2003

Selaginella doederleinii subsp. present in Taiwan? Knapp, R. 2013 

trachyphylla (Warb.) X. C. 

Zhang 

Selaginella flabellata (L.) =Selaginella delicatula FOT 2003 

Spring. 

Selaginella hachijoense Nakai =Selaginella nipponica FOT 2003 

Selaginella hayatana 

=Selaginella 

MAN 

Kümmerle (type: Taiwan) moellendorffii 

Selaginella hayatana Satake =Selaginella aristata Knapp, R. 2013 


Selaginella henryi Koidz. (type: =Selaginella repanda MAN 


Selaginella involvens auct. non =Selaginella 

FOT 2003 

Spring.: Henry 

tamariscina 

Selaginella japonica Miq. =Selaginella remotifolia FOT 2003 

Selaginella kelungensis Hayata =Selaginella remotifolia MAN 


Selaginella kraussiana auct. =Selaginella remotifolia FOT 2003 

non (Kunze) A. Br. 

Selaginella leptophylla Baker =Selaginella aristata Knapp, R. 2013 

Selaginella leptophylla var. 

wichurae (Warb.) Tagawa 

=Selaginella aristata MAN/Knapp, R. 

2013 

Selaginella longicauda Warb. =Selaginella boninensis FOT 2003 

Selaginella mongholica auct. =Selaginella repanda FOT 2003 

non Rupr. 

Selaginella morrisonensis =Selaginella labordei MAN 


Selaginella plumosa auct. non =Selaginella 


(L.) C. Presl 

doederleinii 

Selaginella proniflora auct. non =Selaginella ciliaris FOT 2003 

Baker 

Selaginella pseudo-involvens =Selaginella 

MAN 


stauntoniana 

Selaginella pseudonipponica =Selaginella helvetica Knapp, R. 2013 

Tagawa 

subsp. pseudonipponica 

Selaginella recurvifolia Warb. =Selaginella 

FOT 2003 

heterostachys 

Selaginella satakeana Koidz. =Selaginella aristata Knapp, R. 2013 

Selaginella savatieri Baker =Selaginella nipponica FOT 2003 

Selaginella somae Hayata =Selaginella boninensis MAN 


Selaginella springiana auct. =Selaginella 

FOT 2003 

non Rosenst. 

Selaginella stenostachya 


281 

doederleinii 

=Selaginella aristata Knapp, R. 2013

282 

Selaginella subcaulescens =Selaginella 

Chang et al. 2012 


moellendorffii 

Selaginella tarokoensis =Selaginella 

MAN 

Yamamoto (type: Taiwan) heterostachys 

Selaginella trachyphylla A. =Selaginella 


Braun ex Hieron. 

doederleinii subsp. 

trachyphylla; present 

in Taiwan? 

Selaginella wichurae Warb. =Selaginella aristata MAN/Knapp, R. 

2013 

Selenodesmium obscurum =Cephalomanes MAN 


obscurum 

Selliguea decurrens (Blume) C. =Leptochilus decurrens this publication 

Presl 

Selliguea echinospora 

(Tagawa) H. Ohashi & K. 

Ohashi 

=Selliguea echinospora 

(Tagawa) Fraser- 

Jenk. 


85(1): 46 (2010) 

Selliguea engleri (Luerss.) H. 

Ohashi & K. Ohashi 

Selliguea hastata (Thunb.) H. 

Ohashi & K. Ohashi 

=Selliguea engleri 

(Luerss.) Fraser- 

Jenk. 

=Selliguea hastata 

(Thunb.) Fraser- 

Jenk. 

Selliguea hemionitidea C. Presl =Colysis hemionitidea MAN 

Selliguea pothifolia auct. non 

(Buch.-Ham. ex D. Don) J. 

Sm.: FOT 2003 

Selliguea rhynchophylla auct. 

non (Hook.) Fraser-Jenk.: 

MAN 

Selliguea rhynchophylla auct. 

non (Hook.) H. Ohashi & K. 

Ohashi 

Selliguea yakushimensis 

(Makino) H. Ohashi & K. 

Ohashi 


85(1): 46 (2010) 


85(1): 46 (2010) 


2011 


=Selliguea okamotoi 

=Selliguea 

yakushimensis 

(Makino) Fraser- 

Jenk. 



85(1): 46 (2010) 

Sorolepidium duthiei auct. non =Polystichum glaciale Knapp, R. 2011 

(Hope) Ching 

Sorolepidium glaciale (Christ) =Polystichum glaciale Knapp, R. 2011 

Christ 

Sorolepidium ovale Y. T. Hsieh =Polystichum glaciale FOC 

Sphaerocionium badium =Hymenophyllum MAN 

(Hook. & Grev.) 

badium 

Sphaeropteris lepifera (J. Sm. 

ex Hook.) R. M. Tryon 

=Cyathea lepifera MAN

Sphaerostephanos kotoensis =Cyclosorus productus MAN 

(Hayata) Holtt. ex C. M. Kuo 

Sphaerostephanos lanyuensis 


("type": Taiwan) 

?=Cyclosorus 

productus 

(MAN) name as 

written on "type" 

specimen label 

Sphaerostephanos productus =Cyclosorus productus MAN 

(Kaulf.) Holtt. 

Sphaerostephanos taiwanensis =Cyclosorus 

MAN 

(C. Chr.) Holtt. ex C. M. Kuo taiwanensis 

Sphenomeris biflora (Kaulf.) =Odontosoria biflora MAN 

Akas. 

Sphenomeris chinensis (L.) =Odontosoria chinensis MAN 

Maxon 

Sphenomeris chusana (L.) =Odontosoria chinensis MAN 

Copel. 

Sphenomeris gracilis (Tagawa) =Odontosoria gracilis Knapp, R. 2011 

Sa. Kurata 

Spicanta melanopus (Hook.) O. =Blechnum melanopus FOT 2003 

Kunze 

Spicantopsis eburnea (Christ) =Blechnum eburneum MAN 

Tagawa 

Spicantopsis eburnea var. =Blechnum eburneum MAN 

obtusa Tagawa (type: 


Spicantopsis hancockii (Hance) =Blechnum hancockii MAN 

Massamune 

Spicantopsis niponica var. =Blechnum hancockii MAN 

hancockii (Hance) Nakai 

Stachygynandrum japonicum =Lycopodium 

MAN 

(Thunb.) P. Beauv. 

japonicum 

Stachygynandrum 

=Selaginella 

MAN 

tamariscinum P. Beauv. tamariscina 

Stegnogramma dictyoclinoides =Cyclosorus 

MAN 

Ching 

dictyoclinoides 

Stegnogramma griffithii (T. =Cyclosorus griffithii MAN 

Moore) K. Iwats. 

Stegnogramma pozoi (Lag.) K. =Cyclosorus pozoi MAN 

Iwats. 

Stegnogramma tottoides =Cyclosorus tottoides MAN 

(Hayata ex H. Ito) C. V. 

Morton 

Stenochlaena sorbifolia auct. =Lomariopsis 

FOT 2003 

non J. Sm. 

spectabilis 

Stenoloma biflorum (Kaulf.) =Odontosoria biflora MAN 

Ching 

Stenoloma chusana (L.) Ching =Odontosoria chinensis MAN 

283

284 

Stenoloma gracile Tagawa =Odontosoria gracilis Knapp, R. 2011 

Struthiopteris eburnea (Christ) =Blechnum eburneum MAN 

Ching 

Struthiopteris eburnea var. =Blechnum eburneum MAN 

obtusa (Tagawa) Tagawa 

Struthiopteris fraseri (A. Cunn.) =Blechnum fraseri FOT 2003 

Ching 

Struthiopteris hancockii =Blechnum hancockii MAN 

(Hance) Tagawa 

Struthiopteris integripinnulum =Blechnum fraseri MAN 


Struthiopteris orientalis Hook. =Onoclea orientalis MAN 

Taenitis miyoshiana Makino =Drymotaenium MAN 

miyoshianum 

Tapeinidium gracile auct. non =Tapeinidium 


(Blume) Alderw.: Kuo 

biserratum 

Tapeinidium pinnatum var. 

biserratum (Blume) W. C. 

Shieh 

=Tapeinidium 

biserratum 

MAN 

Tectaria brachiata auct. non 

(Zoll. & Mor.) Morton: MAN 

=Tectaria impressa 


C. R. 2008/ 


Tectaria cuspidatopinnata 

(Hayata) C. Chr. (nom. nud.?) 

=Tectaria impressa FOT 2003/Knapp, 

R. 2011 

Tectaria grandifolia auct. non 

(C. Presl) Copel: Fern Fl. 

Philipp. (1960) 

=Tectaria sulitii Flora Malesiana 

Ser. II Vol. 2 part 

1: 91 (1991) 

Tectaria jinpingensis Ching & =Tectaria dubia FOC 

Chu H. Wang 

Tectaria griffithii auct. non =Tectaria multicaudata this publication 

(Baker) C. Chr.: Knapp, R. 

2011 

Tectaria kwarenkoensis =Tectaria simonsii MAN 

(Hayata) C. Chr. 

Tectaria laciniata Ching =Tectaria phaeocaulis FOT 2003 

Tectaria leuzeana (Gaud.) =Pleocnemia rufinervis FOT 2003 

Copel. 

Tectaria linlaoensis Ching & C. 

H. Wang 

mis-typing forTectaria 

linloensis; present in 


Tectaria linloensis Ching & C. 

H. Wang 

Tectaria macrodonta (Fee) C. 

Chr. 

Taiwan? 

=Tectaria rockii; Knapp, R. 2013 

present in Taiwan? 

=Tectaria coadunata FOT 2003

Tectaria pachinense Hayata: 

Kuo, C. M. 1997 (? nom. 

nud.) 

285 

=Tectaria simonsii Kuo, C. M. 1997 

(conflicting data: 

Aspidium 

pachinense 

Hayata =T. 

simonsii; Tectaria 

pachinense 

Hayata =T. 

subtriphylla) 

/MAN 

=Tectaria fuscipes FOT 2003/MAN 

Tectaria paradoxa (Fee) 

Sledge 

Tectaria pteropus (T. Moore) =Tectaria devexa MAN 

Shieh 

Tectaria rockii C. Chr. present in Taiwan? Knapp, R. 2013 

Tectaria subpedata (Harr.) =Tectaria polymorpha Fl. Males., Ser. 2, 

Ching 

2: 87 (1991) 

Tectaria subtriphylla var. =Tectaria simonsii MAN 

ebenosa (Nakai) Nemoto 

Tectaria trifolia (Alderw.) C. =Tectaria polymorpha MAN 

Chr. 

Tectaria variolosa (Wall. ex =Tectaria impressa Fraser-Jenkins, 

Hook.) C. Chr. 

C. R. 2008 

Tectaria vasta (Blume) Copel. =Tectaria fauriei FOT 2003 

Tectaria yunnanensis (Baker) =Tectaria griffithii this publication 

Ching 

Thelypteris acuminata (Houtt.) =Cyclosorus 

MAN 

C. V. Morton 

acuminatus 

Thelypteris acuminata var. =Cyclosorus ensifer MAN 

kuliangensis auct. non 

(Ching) K. Iwats.: MAN 

Thelypteris arida (D. Don) C. V. 

Morton 

Thelypteris aspera auct. non 

(C. Presl) K. Iwats. 

Thelypteris auriculata (J. Sm.) 

K. Iwats. 

Thelypteris aurita auct. non 

(Hook.) Ching: MAN 

Thelypteris brunnea var. 

hiritrachis (C. Chr.) Ching 

Thelypteris castanea (Tagawa) 

Ching 

Thelypteris caudata (Ching) C. 

F. Reed 

=Cyclosorus aridus 

=Cyclosorus 


=Cyclosorus 

auriculatus 


levingei 


hirtirachis 

=Thelypteris japonica 

=Cyclosorus tottoides 

MAN 

FOT 2003/MAN 

FOC/MAN 


2013 

MAN 

MAN 

MAN

286 

Thelypteris decursive-pinnata 

(H. C. Hall) Ching 

Thelypteris dentata (Forssk.) E. 

P. St. John 

Thelypteris dictyoclinoides 

(Ching) C. M. Kuo 

Thelypteris ensifer(a) (Tagawa) 

K. Iwats. 

Thelypteris erubescens (Wall. 

ex Hook.) Ching 

Thelypteris esquirolii (Christ) 

Ching 

Thelypteris glanduligera var. 

hyalostegia (Copel.) H. Ito 

Thelypteris glanduligera var. 

typica H. Ito 

Thelypteris gongylodes 

(Schkuhr) K. Iwats. 

Thelypteris griffithii (var. 

griffithii) (T. Moore) C. F. 

Reed 

Thelypteris griffithii var. wilfordii 

(Hook.) C. M. Kuo 

Thelypteris gymnopteridifrons 

(Hayata) C. M. Kuo 

Thelypteris hirsutipes auct. non 

(C. B. Clarke) Ching 

Thelypteris insularis (K. Iwats.) 

K. Iwats. 

Thelypteris interrupta (Willd.) 

K. Iwats. 

Thelypteris jaculosa (Christ) 

Panigrahi 

Thelypteris kotoensis (Hayata) 

K. Iwats. 

Thelypteris latipinna (Benth.) K. 

Iwats. 

Thelypteris leveillei (Christ) C. 

M. Kuo 

Thelypteris levingei (C. B. 

Clarke) Ching 

Thelypteris liukiuensis (Christ) 

Iwats. 

Thelypteris longipetiolata (K. 

Iwats.) K. Iwats. 

=Phegopteris 


=Cyclosorus dentatus 

=Cyclosorus 

dictyoclinoides 


=Cyclosorus 

erubescens 

=Cyclosorus esquirolii 

MAN 

MAN 

MAN 

MAN 

MAN 

MAN 

=Thelypteris 

FOT 2003/MAN 

angustifrons 

=Thelypteris 

FOT 2003/MAN 

glanduligera 


=Cyclosorus griffithii 

=Cyclosorus wilfordii 

=Cyclosorus 


=Thelypteris 

angulariloba 

=Cyclosorus insularis 

=Cyclosorus interruptus 



=Cyclosorus latipinnus 

MAN 


R. 2011 

MAN 

FOT 2003/MAN 

MAN 

MAN 

MAN 

MAN 

MAN 

=Cyclosorus omeiensis FOT 2003 

=Pseudophegopteris FOC 

levingei 

=Cyclosorus liukiuensis Knapp, R. 2011 

=Cyclosorus 


MAN

Thelypteris oligophlebia var. 

lasiocarpa (Hayata) H. Ito 

Thelypteris omeiensis (Baker) 

Ching 

Thelypteris ornata (J. Sm.) 

Ching 

Thelypteris paludosa auct. non 

(Blume) K. Iwats.: Kuo, C. M. 

1985 

Thelypteris papilio (C. Hope) K. 

Iwats. 

Thelypteris parasitica (L.) 

Fosberg 

Thelypteris phegopteris (L.) 

Slosson ex Rydb. 

Thelypteris pozoi (Lag.) C. V. 

Morton 

Thelypteris producta (Kaulf.) C. 

F. Reed 

Thelypteris prolifera (Retz.) C. 

F. Reed 

Thelypteris pyrrhorhachis 

(Kunze) C. M. Kuo 

Thelypteris remotipinnata 

(Hayata) Alston 

Thelypteris setigera auct. non 

(Wall. ex Bedd.) Ching 

Thelypteris simozawae 


Thelypteris simplex (Hook.) K. 

Iwats. 

Thelypteris subarida (Tatew. & 

Tagawa) C. F. Reed 

Thelypteris subaurita (Tagawa) 

Ching 

Thelypteris supubescens auct 

non (Blume) K. Iwats. 

Thelypteris taiwanensis (C. 

Chr.) K. Iwats. 

Thelypteris torresiana 

(Gaudich.) Alston 

Thelypteris tottoides (Hayata 

ex H. Ito) C. V. Morton 

Thelypteris triphylla (Sw.) K. 

Iwats. (var. triphylla) 


torresiana 



polypodioides 


hirtirachis 

=Cyclosorus papilio 

=Cyclosorus 

parasiticus 

=Phegopteris 

connectilis 

=Cyclosorus pozoi 


=Cyclosorus proliferus 


hirtirachis 

=Gymnocarpium 

remotepinnatum 


polypodioides 

=Thelypteris 

angulariloba 

=Cyclosorus simplex 



subaurita 


=Cyclosorus 

taiwanensis 


torresiana 

=Cyclosorus tottoides 

=Cyclosorus triphyllus 

287 

FOT 2003/MAN 

MAN 

MAN 

MAN 

MAN 

MAN 

FOT 2003 

MAN 

MAN 

MAN 


R. 2011 

MAN 

FOT 2003/MAN 

FOT 2003/MAN 

MAN 

MAN 

MAN 

FOT 2003/MAN 

MAN 

MAN 

MAN 

MAN

288 

Thelypteris triphylla var. 

parishii (Bedd.) K. Iwats. 


2011 

Thelypteris truncata (Poir.) K. =Cyclosorus truncatus MAN 

Iwats. 

Thelypteris truncata f. 

=Cyclosorus truncatus MAN 

kwashotensis (Hayata) C. F. 

Reed 

Thelypteris truncata f. 

=Cyclosorus truncatus MAN 

laevifrons (Hayata) C. F. 

Reed 

Thelypteris tylodes (Kunze) 

Ching 

=Cyclosorus tylodes FOC/Knapp, R. 

2011 

Thelypteris xylodes (Kunze) =Cyclosorus tylodes Knapp, R. 2011 

Ching (mis-spelling of 


THEMELIUM =Grammitis, kept for T. -- 

curtisii 

Themelium curtisii (Baker) =Grammitis curtisii this publication 

Parris 

Themelium tenuisectum =Grammitis tenuisecta MAN 

(Blume) Parris 

TOMOPHYLLUM =Grammitis, kept. for T. -- 

donianum 

Trichomanes acutum C. Presl =Crepidomanes Knapp, R. 2011 

acutum 

Trichomanes acutum Makino =Crepidomanes FOC 

ex Christ 

latealatum 

Trichomanes adiantoides L. =Asplenium falcatum FOT 2003/Knapp, 

R. 2013 

Trichomanes anceps Hook. =Crepidomanes FOT 2003/MAN 

maximum 

Trichomanes apiifolium C. =Cephalomanes MAN 

Presl 

apiifolium 

Trichomanes asplenioides C. =Cephalomanes MAN 

Presl 


aspleniodes 

Trichomanes auriculatum =Crepidomanes MAN 

Blume 

auriculatum 

Trichomanes bauerianum auct. =Cephalomanes FOT 2003/MAN 

non Endl. 

apiifolium 

Trichomanes beccarianum 

Cesati 

=Trichomanes motleyi FOC/Knapp, R. 

2011 

Trichomanes belangeri Bory =Crepidomanes FOT 2003/MAN 

Trichomanes bilabiatum Nees 

& Blume 

auriculatum 

=Crepidomanes 

bipunctatum 

MAN

Trichomanes bipunctatum Poir. =Crepidomanes MAN 

bipunctatum 

Trichomanes bipunctatum var. =Crepidomanes MAN 

latealatum (Bosch) Alderw. latealatum 

Trichomanes birmanicum 

Bedd. 

=Crepidomanes 

birmanicum in part, 

=Crepidomanes 



26-40 (2009) 

kalamocarpum in part 

Trichomanes blepharistomum =Cephalomanes FOT 2003/MAN 

Copel. 

thysanostomum 

Trichomanes chinense L. =Odontosoria chinensis MAN 

Trichomanes clathratum =Cephalomanes MAN 


clathratum 

Trichomanes concinum Mett. =Crepidomanes humile FOT 2003/MAN 

Trichomanes contiguum G. =Grammitis contigua MAN 

Forst. 

Trichomanes cumingii (C. =Cephalomanes MAN 


cumingii 

Trichomanes cupressifolium =Crepidomanes MAN 


latifrons 

Trichomanes cupressoides =Cephalomanes FOT 2003/MAN 

auct. non Desv. 

obscurum 

Trichomanes denticulatum =Hymenophyllum MAN 

(Sw.) Poir. 

denticulatum 

Trichomanes diffusum Blume =Crepidomanes MAN 

minutum 

Trichomanes digitatum Sw. =Sphaerocionium 

digitatum 


2011 

Trichomanes dimidiatum C. =Crepidomanes FOT 2003/MAN 

Presl 

auriculatum 

Trichomanes dissectum J. Sm. =Crepidomanes FOT 2003/MAN 

auriculatum 

Trichomanes eminens C. Presl =Cephalomanes FOT 2003/MAN 

apiifolium 

Trichomanes filiculum Bory =Crepidomanes FOT 2003 

latealatum 

Trichomanes formosanum Y. =Crepidomanes Knapp, R. 2011 

Yabe (type: Taiwan) 

rupicola 

Trichomanes grande Copel. =Cephalomanes Knapp, R. 2013 

grande 

Trichomanes humile G. Forst. =Crepidomanes humile MAN 

Trichomanes japonicum =Crepidomanes MAN 

Thunb. 

birmanicum 

Trichomanes japonicum var. 

formosanum Christ ex 

Matsum. (type: Taiwan) 

=Crepidomanes 

maximum 

MAN 

289

290 

Trichomanes javanicum auct. 

non Blume 

Trichomanes javanicum var. 

asplenioides (C. Presl) C. 

Chr. 

Trichomanes kalamocarpum 


=Cephalomanes 


asplenioides 

=Cephalomanes 


asplenioides 

=Crepidomanes 

kalamocarpum 

FOT 2003/MAN 

MAN 

Trichomanes kurzii Bedd. =Crepidomanes kurzii MAN 

Trichomanes laciniatum Roxb. =Cephalomanes 


asplenioides 

MAN 

Trichomanes latealatum 

(Bosch) Christ 

Trichomanes latemarginale 

Eaton 

Trichomanes latifrons Bosch 

Trichomanes latipinnum Copel. 

Trichomanes liukiuense Christ 

ex Matsum. 

Trichomanes liukiuensis Yabe 

Trichomanes longifrons Nakai 


Trichomanes luzonicum C. 

Presl 

Trichomanes makinoi C. Chr. 

Trichomanes maximum (auct. 

non Blume) Ogata 

Trichomanes maximum Blume 

Trichomanes minutissimum 

Alderw. 

Trichomanes minutum Blume 

Trichomanes miyakei Y. Yabe 


Trichomanes nanophyllum 

(Tagawa) C. V. Morton 

=Crepidomanes 

latealatum 

=Crepidomanes 

latemarginale 

=Crepidomanes 

latifrons 

=Cephalomanes 

obscurum 

=Crepidomanes 

maximum 

=Crepidomanes 

maximum 

=Crepidomanes 

latifrons 


=Crepidomanes 

latealatum 

=Cephalomanes 

thysanostomum 

=Crepidomanes 

maximum 

=Trichomanes motleyi 

=Crepidomanes 

minutum 

=Crepidomanes 

maximum 

=Crepidomanes kurzii 



26-40 (2009) 

MAN 

MAN 

MAN 

FOT 2003/MAN 

FOT 2003/MAN 

FOT 2003/MAN 

MAN 

FOT 2003/MAN 

MAN 

FOT 2003/MAN 

MAN 

Copeland 

1933/FOT 


2011 

MAN 

MAN 

MAN

Trichomanes nanum Hook. & 

Baker 

Trichomanes nitidulum Bosch 

Trichomanes naseanum Christ 

Trichomanes obscurum Blume 

Trichomanes omphalodes 

(Vieill. ex E. Fourn.) C. Chr. 

Trichomanes orbiculatum 

Ching ex Ogata 

Trichomanes orientalis auct. 

non C. Chr.: MAN 

Trichomanes pallidum Blume 

Trichomanes palmifolium 


Trichomanes papillatum K. 

Mueller 

Trichomanes parvum Copel. 

Trichomanes peltatum auct. 

non Poir. 

Trichomanes pervenulosum 

auct. non Alderw. 

Trichomanes polyanthos Sw. 

Trichomanes preslii C. V. 

Morton 

Trichomanes proliferum Blume 

Trichomanes 

pseudoblepharistomum 

Tagawa 

Trichomanes pyxidiferum L. 

Trichomanes racemulosum 

Bosch 

Trichomanes radicans var. 

birmanicum (Bedd.) C. Chr. 

=Crepidomanes 

latemarginale 


nitidulum 

=Crepidomanes 

birmanicum 

=Cephalomanes 

obscurum 

=Trichomanes 

tahitense 

=Crepidomanes 

minutum 

=Crepidomanes 

birmanicum 

=Crepidomanes 

pallidum 

=Crepidomanes 

palmifolium 

=Cephalomanes 

obscurum 

=Crepidomanes 

kalamocarpum 

=Trichomanes 

tahitense 

=Crepidomanes 

parvifolium 


polyanthos 

=Cephalomanes 


asplenioides 

=Crepidomanes 

minutum 

=Cephalomanes 

thysanostomum 

=Crepidomanes 

latifrons 

=Cephalomanes 

obscurum 

=Crepidomanes 

birmanicum 

FOT 2003 

MAN 



26-40 (2009) 

MAN 

MAN 

FOT 2003/MAN 



26-40 (2009) 

MAN 


2011 

FOT 2003/MAN 



26-40 (2009) 

FOT 2003/MAN 

291 

Knapp, R. 


2013 

FOT 2003/MAN 

MAN 

MAN 

MAN 

Kuo 

FOT 2003/MAN 


2011

292 

Trichomanes radicans f. 

naseanum (Christ) C. Chr. 


naseanum (Christ) Lellinger 


orientalis auct. non (C. Chr.) 

Lellinger: MAN 

Trichomanes rigidum auct. non 

Sw. 

Trichomanes rupicola Racib. 

=Crepidomanes 

birmanicum 

=Crepidomanes 

birmanicum 

=Crepidomanes 

birmanicum 

Kuo, C. M. 


2011 



26-40 (2009) 



26-40 (2009) 

FOT 2003/MAN 

=Cephalomanes 

obscurum 

=Crepidomanes FOC 

rupicola 

Trichomanes saxatile T. Moore =Cephalomanes FOT 2003/MAN 

obscurum 

Trichomanes saxifragoides C. =Crepidomanes MAN 

Presl 

minutum 

Trichomanes smithii Hook. =Cephalomanes FOT 2003/MAN 

cumingii 

Trichomanes solidum G. Forst. =Davallia solida MAN 

Trichomanes somae Nakai =Crepidomanes MAN 


birmanicum 

Trichomanes striatum D. Don =Crepidomanes Knapp, R. 2011 

birmanicum 

Trichomanes strigosum Thunb. =Microlepia strigosa MAN 

Trichomanes subclathratum (K. ?=Crepidomanes sp. 1 Knapp, R. 2011 

Iwats.) C. V. Morton 

Trichomanes tagawanum K. =Crepidomanes MAN 

Iwats. 

latealatum 

Trichomanes tenuifolium Burm. =Cheilanthes tenuifolia MAN 

f. 

Trichomanes thysanostomum =Cephalomanes MAN 

Makino 

thysanostomum 

Trichomanes yandinense =Trichomanes 

FOT 2003/MAN 

Bailey 

bimarginatum 

Ugena microphylla Cav. =Lygodium 

MAN 

microphyllum 

Urostachys carinatus (Descv. =Huperzia carinata MAN 

ex Poir.) Herter ex Nessel 

Urostachys christensenianus =Huperzia sieboldii MAN 

(Christ & Herter) Herter ex 

Nessel 

Urostachys cryptomeri(a)nus 

(Maxim.) Herter ex Nessel 

Urostachys cunninghamioides 

(Hayata) Herter ex Nessel 

=Huperzia 

cryptomerina 

=Huperzia 


MAN 

MAN

Urostachys fargesii (Herter) =Huperzia fargesii MAN 

Herter ex Nessel 

Urostachys fauriei (Rosenst.) =Huperzia fargesii MAN 

Herter 

Urostachys fordii (Baker) =Huperzia fordii MAN 


Urostachys formosanus Herter =Huperzia salvinioides MAN 

ex Nessel 

Urostachys juniperistachys =Huperzia fordii MAN 


Urostachys phlegmaria (L.) =Huperzia phlegmaria MAN 


Urostachys quasipolytrichoides =Huperzia 

MAN 

(Hayata) Herter ex Nessel quasipolytrichoides 

Urostachys salvinioides Herter =Huperzia salvinioides MAN 

Urostachys selago var. 

appressus (Bach. Pyl. ex 


2011 

Desv.) Herter ex Nessel 

Urostachys serratus (Thunb.) =Huperzia serrata MAN 

Herter 

Urostachys sieboldii (Miq.) =Huperzia sieboldii FOC 


Urostachys squarrosus (G. =Huperzia squarrosa MAN 

Forst.) Herter 

Urostachys subdistichus =Huperzia fordii MAN 

(Makino) Herter ex Nessel 

Urostachys tereticaulis 

=Huperzia fargesii MAN 


Vaginularia paradoxa (Fee ) =Monogramma MAN 

Mett. ex Miq. 

paradoxa 

Vaginularia trichoidea Fee =Monogramma MAN 

trichoidea 

Vandenboschia auriculata =Crepidomanes MAN 


Vandenboschia birmanica 

(Bedd.) Ching 

Vandenboschia latifrons 

(Bosch) Copel. 

Vandenboschia longifrons 

(Nakai) Copel. 

Vandenboschia maxima 


Vandenboschia naseana 

(Christ) Ching & Wang 

auriculatum 

=Crepidomanes 

birmanicum 

=Crepidomanes 

latifrons 

=Crepidomanes 

latifrons 

=Crepidomanes 

maximum 

=Crepidomanes 

birmanicum 



26-40 (2009) 

MAN 

FOT 2003/MAN 

MAN 



26-40 (2009) 

293

294 

Vandenboschia orientalis auct. 

non (C. Chr.) Ching: MAN 

Vandenboschia parva (Copel.) 

Copel. 

Vandenboschia pyxidifera auct. 

non (L.) Copel.: FOT 1975 

Vandenboschia radicans var. 

naseana (Christ) H. Ito 

Vandenboschia radicans var. 

orientalis auct. non (C. Chr.) 

H. Ito: MAN 

Vandenboschia somai auct. 

non ? (Nakai) Copel. 

Vandenboschia subclathrata K. 

Iwats. 

Vittaria agasawarensis 

Kodama 

Vittaria arisanensis Hayata 


Vittaria formosana Nakai (type: 


=Crepidomanes 

birmanicum 

=Crepidomanes 

kalamocarpum 

=Crepidomanes 

latifrons 

=Crepidomanes 

birmanicum 

=Crepidomanes 

birmanicum 



26-40 (2009) 



26-40 (2009) 

MAN 



26-40 (2009) 



26-40 (2009) 

FOT 2003/ 


=Crepidomanes 

birmanicum 

?=Crepidomanes sp. 1 Knapp, R. 2011 

=Vittaria zosterifolia FOT 2003 

=Vittaria taeniophylla 

=Vittaria zosterifolia 

MAN 

MAN 

Vittaria japonica Miq. =Vittaria flexuosa FOT 2003 

Vittaria pusilla Blume 

=Scleroglossum MAN 

sulcatum 

Vittaria stenophylla Copel. =Vittaria angusteelongata 

FOT 2003 

Vittaria suberecta Hayata =Vittaria flexuosa MAN 


Vittaria tortifrons Hayata (type: =Vittaria zosterifolia MAN 


Woodsia polystichoides var. =Woodsia 

FOT 2003 

veitchii Hance 

polystichoides 

WOODSIACEAE =DRYOPTERIDACEAE -- 

Woodwardia orientalis auct. =Woodwardia prolifera MAN 

non Sw.: FOT 1975 

Woodwardia orientalis var. =Woodwardia prolifera MAN 

formosana Rosenst. 

Woodwardia orientalis var. =Woodwardia prolifera MAN 

prolifera (Hook. & Arn.) Ching 

Woodwardia prolifera var. 

formosana (Rosenst.) Ching 

ex Chiu 

=Woodwardia prolifera MAN

295 

Woodwardia radicans var. 

unigemmata Makino 

Woodwardia takeoi Hayata 


Xiphopteris okuboi (Yatabe) 

Copel. 

=Woodwardia 

unigemmata 

=Woodwardia kempii 


MAN 

MAN 

MAN

296

297 

5 R E F E R E N C E S 

Backer, C. A. 1936; reprint 2000, L. J. Veen. Verklarend woordenboek van 

wetenschappelijke plantennamen. Amsterdam/Antwerpen. 

Baker, J. G. 1885. Ferns collected in North Formosa by Mr. William Hancock. 

J. Bot. 23: 102-107. 

Chang, H. M., Chiou, W. L. & Wang, J. C. 2012. Flora of Taiwan, 

Selaginellaceae. Endemic Species Research Institute, Nantou, Taiwan. 

Chang, Y. H. 2013. Review of the taxa new to Taiwan in recent years: 

lycophytes and ferns. Forestry Research Newsletter 113: 29-40 (in Chinese; 

original title: 臺灣植物誌新分類群之增補 — 石松類植物及蕨蕨類植物篇 ). 

Chang, Y. H., Chiou, W. L., Huang, Y. M., Shen, B. N., Lin, C. Y. & Hsu, T. C. 

2014. New additions to the fern flora of Taiwan (2). TW J. of Biodivers. 

16(3): 263-272. 

Chang, Y. H., Wu, W. H., Chen, C. W., Lu, P. F. & Huang, Y. M. 2012. Two 

taxa of ferns newly recorded to Taiwan. TW J. of Biodivers. 14(3-4): 81-90. 

Chao, Y. S., Ebihara, A., Chang, Y. H., Jiang, R. H., Ngan, L. T. & Chiou, W. 

L. 2013. New distribution of Pteris kawabatae Sa. Kurata and recircumscription 

of Pteris arisanensis Tagawa. Taiwania 58(4): 305-310. 

Chen, C. W., Huang, Y. M., Kuo, L. Y., Chang, Y. H., Liu, Y. C. & Chiou, W. L. 

2013. A new vittarioid fern species, Haplopteris heterophylla (Pteridaceae). 

Syst. Bot. 38(4): 901-909. 

Christenhusz, M. J. M. & Chase, M. W. 2014. Trends and concepts in fern 

classification. Ann. Bot. 113(4): 571-594. 

Christenhusz, M. J. M. & Schneider, H. Corrections to Phytotaxa 19: Linear 

sequence of lycophytes and ferns. Phytotaxa 28: 50-52 (2011) 

Christenhusz, M. J. M., Zhang, X. C. & Schneider, H. 2011. A linear sequence 

of extant families and genera of lycophytes and ferns. Phytotaxa 19: 7-54. 

Copeland, E. B. 1933. Trichomanes. Philipp. J. Sci. 51: 119-280, pls. 1-61. 

Copeland, E. B. 1960. Fern flora of the Philippines 3. Bureau of Printing, 

Manila. 

DeVol, C. E. 1979. A check list of the vascular plants of Taiwan 1. 

Pteridophyta, in Li, H. L. et al. Taipei, Taiwan, Epoch Publishing. 

Field, A. R. & Bostock, P. D. 2013. New and existing combinations in 

palaeotropical Phlegmariurus (Lycopodiaceae) and lectotypification of the 

type species Phlegmariurus phlegmaria (L.) T. Sen & U. Sen. PhytoKeys 20: 

33-51. 

Fraser-Jenkins, C. R. 1989. A monograph of the genus Dryopteris 

(Pteridophyta: Dryopteridaceae) in the Indian subcontinent. Bull. Brit. Mus. 

Nat. Hist., Botany series Vol. 18(5): 323-477. 

Fraser-Jenkins, C. R. 1997. An outline monographic study of the genus 

Polystichum in the Indian subcontinent. International Book Distributors, 

Dehra Dun, India.

298 

Fraser-Jenkins, C. R. 2008. Taxonomic revision of three hundred Indian 

subcontinental pteridophytes - a new picture of fern-taxonomy and nomenclature 

in the Indian subcontinent. Bishen Singh Mahendra Pal Singh, Dehra 

Dun, India. 

Fraser-Jenkins, C. R. 2009. A brief comparison of modern pteridophyte 

classifications (families and genera in India). Indian Fern J. 26: 107-126. 

Genaust, H. 1996. Etymologisches Wörterbuch der botanischen Pflanzennamen 

(3. Auflage). Birkhäuser Verlag, Switzerland. 

Gledhill, D. 2008. The names of plants (4 th ed.). Cambridge University Press. 

Hawksworth, D. L. 2010. Terms used in bionomenclature. Global Biodiversity 

Information Facility, Copenhagen. 

Hayata, B. 1908. Flora montana formosae. University of Tokyo. 

Hayata, B. 1909. Some ferns from the mountain regions of Formosa. Bot. 

Mag. (Tokyo) 23: 1-4 & 24-34 & 76-80. 

Hayata, B. 1911. Materials for a flora of Formosa. University of Tokyo. 

Hayata, B. 1914-21. Iconum plantarum formosanarum 4-8 & 10. Government 

of Formosa, Taihoku. 

Hayata, B. 1917. General index to the flora of Formosa, supplement to icones 

plantarum formosanarum 6 (1916). Bureau of Productive Industries, 

Government of Formosa, Taihoku, Taiwan. 

Henry, A. 1896. A list of plants from Formosa. Transactions of the Asiatic 

Society of Japan Vol. 24 - Supplement. 

Hsu, T. C., Lin, C. Y., Chen, C. W., Chiou, W. L. & Chang, Y. H. 2014. 

Confirmation of the occurrence and distribution of three fern species in 

Taiwan. Taiwan J For Sci 29(2): 157-167. 

Huang, T. C. (ed.-in-chief). 1994. Flora of Taiwan 1 (2 nd ed.). Editorial 

Committee of the Flora of Taiwan. 

Huang, T. C. (ed.-in-chief). 2003. Flora of Taiwan 6 (2 nd ed.). Editorial 

Committee of the Flora of Taiwan. 

Huang, T. C., Huang, S. F. & Yang, K. C. 1994. The flora of Taipingtao (Aba 

Itu Island) (part 1). Taiwania 39. 

Ito, H. 1944. Filices japonenses illustratae. Tokyo. 

Ito, H. 1981. Icones filicum japoniae 9. (in Japanese) 

Ito, T. 1928. Illustrations of Taiwan ferns. Kyoto. (in Japanese) 

Iwatsuki, K., Yamazaki, T., Boufford, D. E. & Ohba, H. (eds.). 1995. Flora of 

Japan 1. Kodansha, Tokyo. 

Jaeger, E. C. 1955. A source book of biological names and terms (3 rd ed). 

Riverside College, Riverside, USA. 

Kadoya, T., Takenaka, A., Ishihama, F., Fujita, T., Ogawa, M. et al. 2014. 

Crisis of Japanese vascular flora shown by quantifying extinction risks for 

1618 taxa. PLoS ONE 9(6): e98954. Doi:10.1371/journal.pone.0098954. 

Kawakami, T. 1910. A list of plants of Formosa. Taihoku, Bureau of 

Productive Industry, Govt. of Formosa. 

Knapp, R. 2011. Ferns and fern allies of Taiwan. KBCC Press & Yuan-Liou 

Publishing, Taipei, Taiwan.

299 

Knapp, R. 2013. Ferns and fern allies of Taiwan - Supplement. KBCC Press, 

Taipei, Taiwan. 

Kramer, K. U. & Green, P. S. (eds.) in Kubitzki, K. (ed.). 1990. The families 

and genera of vascular plants 1 - Pteridophytes and Gymnosperms. 

Springer Verlag, Berlin, Heidelberg. 

Kuo, C. M. 1985. Taxonomy and phytogeography of Taiwanese pteridophytes. 

Taiwania 30: 5-99. 

Kuo, C. M. 1997. Manual of Taiwan vascular plants 1. Taipei, Taiwan Council 

of Agriculture (in Chinese). 

Kuo, C. M. 2001. Ferns of Taiwan 1. Yuan-Liou Publishing, Taipei (in 

Chinese). 

Kuo, C. M. 2010. Ferns of Taiwan 2. Yuan-Liou Publishing, Taipei (in 

Chinese). 

Lellinger, D. B. 2007. A modern English - Chinese glossary for taxonomic 

pteridology. Missouri Botanical Garden, St. Louis, Missouri. 

Li, H. L. et al. (eds.). 1975. Flora of Taiwan 1 (1 st ed.). Taipei, Taiwan, Epoch 

Publishing. 

Liew, F. S. 1976. A list of ferns and fern allies found in Orchid Island, Taiwan. 

Taiwania 21. 

Liu, Y. C. 2008. A taxonomic study of Athyrium in Taiwan. Ph. D Thesis, 

National Sun Yat-sen University, Taiwan. 

Liu, Y. C., Chiou, W. L. & Liu, H. Y. 2009. Fern Flora of Taiwan: Athyrium. 

TFRI Extension Series No. 198, Taipei, Taiwan. 

Lu, F. Y. 2011. Flora of Kinmen. Kinmen National Park Headquarters. 

Lu, S. G. & Yang, T. Y. A. 2005. The checklist of Taiwanese pteridophytes 

following Ching's system. Taiwania 50(2): 137-165. 

Masamune, G. 1936. A short flora of Formosa. Taihoku: Kudoa. 

Matsumura, J., and Hayata, B. 1906. Enumeratio Plantarum: in insula 

formosa sponte crescentium hucusque rite cognitarum adjectis descriptionibus 

et figuris specierum pro regione novarum. Journ. Coll. Sci. Tokyo 22. 

McNeill, J., Barrie, F. R., Buck, W. R., Demoulin, V., Greuter, W., 

Hawksworth, D. L., Herendeen, P. S., Knapp, S., Marhold, K., Prado, J., 

Prud’homme van Reine, W. F., Smith, G. F., Wiersema, J. H. & Turland, N. 

J. (eds.). 2012. International Code of Nomenclature for algae, fungi, and 

plants (Melbourne Code). Regnum Vegetabile 154. Koeltz Scientific Books. 

Moore, S. J. 2010. Monographic study on the fern genus Microlepia (Dennstaedtiaceae). 

Ph. D. thesis, National Taiwan Normal University, Taipei, 

Taiwan (in Chinese with English abstract, key and taxa data). 

Moore, S. J., Hsu, T. W., Chiang, C. Y. & Chen, C. C. 1999. An outstanding 

Taiwan fern collector - Bi-Jiao Wang (1953-1992). Endemic Species 

Research Institute, Nantou, Taiwan and Institute of Botany, Academia 

Sinica, Taipei, Taiwan. 

Ogata, M. 1928-40. Icones filicum japoniae 1-8. Sanshusha, Tokyo (in 

Japanese).

300 

Parris, B. S., Kiew, R., Chung, R. C. K., Saw, L. G. & Soepadmo, E. 2010. 

Flora of Peninsular Malaysia: Series I: Ferns and lycophytes 1. Forest 

Research Institute Malaysia, Malayan Forest Records No. 48. 

Rothfels, C. J., Sundue, M. A., Kuo, L. Y., Larsson, A. Kato, M. Schuettpelz, 

E. & Pryer, K. M. 2012. A revised family-level classification for eupolypod II 

ferns (Polypodiidae: Polypodiales). Taxon 61(3): 515-533. 

Saiki, Y. 1994. Note on Ferns (2) - Asiatic species of the Aleuritopteris 

farinosa Complex. J. Phytogeog. Taxon. 32(1): 1-13. 

Sasaki, S. 1928. List of plants of Formosa. Government Research Institute, 

Taihoku, Formosa. 

Stearn, W. T. 2007. Botanical Latin (4 th ed., paperback, reprinted). Timber 

Press, USA. 

Stewart, R. R., Johnson, D. M. & Mickel, T. 1983. Pteridophyte genera - 

Meaning of their names. Fiddlehead Forum 10(4&5): 21-36. 

Swinhoe, R. 1863. Lists of plants from the island of Formosa, or Taiwan. 

Chelsea. 

Tagawa, M. 1940-49. Studies on Formosan ferns 1-7. Acta Phytotax. Geobot. 

9, 10, 11 & 14. 

Wang, J. C., Chiou, W. L. & Chang, H. M. (eds.). 2012. A preliminary Red List 

of Taiwanese vascular plants. Endemic Species Research Institute & 

Taiwan Society of Plant Systematics, Nantou, Taiwan. 

Wu, Z. Y., Raven, P. H. & Hong, D. Y. (eds.). 2013. Flora of China 2-3 

(Pteridophytes). Beijing: Science Press; St. Louis: Missouri Botanical 

Garden Press. 

Yabe, Y. 1902. A note of ferns from the island of Koto (Botel-Tobago). Bot. 

Mag. (Tokyo) 16. 

Yamamoto, Y. 1928. Supplementa iconum plantarum formosanarum 4. 

Government Research Institute, Taihoku, Formosa. 

Yamamoto, Y. 1932. Supplementa iconum plantarum formosanarum 5. 

Government Research Institute, Taihoku, Formosa. 

Yang, Y. P. & Liu, H. Y. 2002. Manual of Taiwan vascular plants 6. The 

Council of Agriculture Press, Taipei, Taiwan (in Chinese).

301 

6 A C K N O W L E D G E M E N T 

My particular thanks go to T. C. Hsu ( 許天銓 , TaiBei, Taiwan) and C. W. Chen 

( 陳正為 , TaiBei, Taiwan) for the numerous and invaluable discussions for new 

and critical taxa. 

I am very grateful to the following experts for the partly extensive 

support regarding etymological data for scientific names, and as well other 

input: P. H. Hovenkamp (Leiden, Netherlands), C. R. Fraser-Jenkins (Kathmandu, 

Nepal), B. S. Parris (Bay of Islands, New Zealand), G. Rouhan (Paris, 

France), M. Sonnberger (Heiligkreuzsteinach, Germany), A. Ebihara (Tsukuba, 

Japan), L. B. Zhang ( 张丽兵 , Missouri, USA), M. Hassler (Bruchsal, Germany), 

X. C. Zhang ( 张宪宪春 , BeiJing, China) and F. Heiker (Markkleeberg, Germany). 

I thank D. A. Dowling (TaiBei, Taiwan) for improving the English of 

this document. Regarding editing I am indebted to D. Chung ( 鍾彥儒 , Marina, 

USA). 

I sincerely thank the Dr. Cecilia Koo Botanic Conservation and 

Environmental Protection Foundation (KBCC) for their continued engagement 

and interest in this project. In particular, I wish to thank C. W. Li ( 李家家維 , 

XinZhu, Taiwan) for his continuous support to get this index realized. 

I feel privileged at having received help to such a great extent from 

curators and their teams, and I would like to particularly mention the following: 

HAST (Biodiversity Research Center, Academia Sinica; TaiBei, Taiwan); TAIF 

(Taiwan Forestry Research Institute; TaiBei, Taiwan); TAI (National Taiwan 

University; TaiBei, Taiwan); B (Botanischer Garten und Botanisches Museum 

Berlin-Dahlem; Berlin, Germany); BM (The Natural History Museum; London, 

UK); BR (National Botanic Garden of Belgium; Meise, Belgium); E (Royal 

Botanic Garden Edinburgh, Edinburg, UK); FI (Natural History Museum; 

Florence, Italy); GH (Harvard University; Cambridge, USA); K (Royal Botanic 

Gardens; Kew, UK); KYO (Kyoto University; Kyoto, Japan); L (Nationaal 

Herbarium Nederland, Leiden University branch; Leiden, Netherlands); MICH 

(University of Michigan; Ann Arbor, USA); NY (New York Botanical Garden; 

New York, USA); P (Muséum National d'Histoire Naturelle; Paris, France); PE 

(Institute of Botany, Chinese Academy of Sciences; BeiJing, China); PR 

(National Museum in Prague; Prague, Czech Republic); PRC (Charles

302 

University in Prague; Prague, Czech Republic); TI (University of Tokyo; Tokyo, 

Japan); US (Smithsonian Institution; Washington, USA); UPS (Uppsala 

University; Uppsala, Sweden); Z (Universität Zürich; Zurich, Switzerland). 

I thank all those who have provided feedback to the content of my 

books, my web albums and other publications, feedback which is reflected in 

this publication. 

The most important help that I have received was from my wife Jing 

Zeng ( 曾競 ) and daughter Ann-Ev Knapp who supported my time intensive 

studies in a multitude of ways.

303 

A P P E N D I X A 

- Taxa endemic to Taiwan 

Based on the information we currently have available, a significant number of 

taxa can be considered unique to Taiwan and its surrounding smaller islands. 

Some of them are rather widely distributed and exist in large numbers. 

However, in most cases, populations are sparse and small. In extreme cases, 

only single populations have been confirmed, and the total number of 

individuals could be less than one hundred. 

For the survival of the following, presumed endemic, taxa, Taiwan is solely 

responsible: 

Adiantum capillus-veneris f. 

lanyuanum 

Adiantum formosanum 

Adiantum meishanianum 

Adiantum taiwanianum 

Angiopteris itoi 

Angiopteris somae 

Antrophyum castaneum 

Araiostegia parvipinnula 

Asplenium cuneatiforme 

Asplenium x wangii 

Athyrium auriculatum 

Athyrium leiopodum 

Athyrium tripinnatum 

Bolbitis x nanjenensis 

Cheilanthes agetae 

Cibotium taiwanense 

Cyclosorus ensifer 

Cyclosorus longipetiolatus 

Deparia dawuense 

Diplazium pseudodoederleinii 

Dryopsis x fauriei 

Dryopteris cacaina 

Dryopteris kwanzanensis 

Very rare on two islands of SE Taiwan 

Rare, but widely distributed in Taiwan 

Very rare, only in one location of SC 



Very rare, only in two very small 

subpopulations 

Very rare, only in two locations 


Common, widely distributed in Taiwan 

Rather common, widely distributed in 


Very rare, only in two locations 

Rare in S Taiwan 

Rare, very scattered in Taiwan 

Occasionally, widely distributed in Taiwan 

Very rare, only known from type location 

(no image available) 

Probably rare, very scattered in Taiwan 

Locally common, widely distributed in 


Locally common in S Taiwan 

Very rare, only known from very few 

locations 

Probably rare, very scattered in Taiwan 




Rare, very scattered in Taiwan

304 

Dryopteris pseudolunanensis 

Dryopteris pseudosieboldii 

Dryopteris subatrata 

Dryopteris subreflexipinna 


Huperzia changii 

Huperzia cunninghamioides 

Hymenophyllum alishanense 

Hymenophyllum devolii 


Isoetes taiwanensis var. 

kinmenensis 

Isoetes taiwanensis var. 

taiwanensis 

Lepisorus kawakamii 

Lepisorus megasorus 

Lepisorus monilisorus 

Lepisorus pseudoussuriensis 

Loxogramme biformis 

Loxogramme remotefrondigera 

Lycopodium yueshanense 

Microlepia intramarginalis 

Microlepia krameri 

Microlepia trichocarpa 

Pleocnemia rufinervis 

Polypodium raishaense 

Polypodium transpianense 

Polystichum parvipinnulum 

Polystichum pseudostenophyllum 

Polystichum simadae 

Polystichum taiwanianum 

Polystichum taizhongense 

Pteris angustipinna 

Pteris longipinna 

Pteris pellucidifolia 

Pteris wulaiensis 

Pyrrosia polydactyla 

Selaginella devolii 




Very rare, very scattered in Taiwan 

Very rare, only known from two locations 

Very rare, only known from three locations 

(no image available) 

Very rare, only scattered in S, E and N 


Very rare, only known from three locations 

Rare in S and SE Taiwan 


Very rare, only known from one location in 

JinMen 

Very rare, only known from one location in 

N Taiwan 





Rare, very scattered in S, SW and C 




Locally not rare, probably widely distributed 

in Taiwan 

Common and widely distributed in N 


Rare, mostly in S and W Taiwan 

Probably common in N Taiwan 




Rare, scattered in high-mountain regions 


Rare and scattered in C and SC Taiwan 

Rare, but rather widely distributed in 


Very rare, only in C Taiwan 

Quite common, widely distributed in Taiwan 

Rare in C Taiwan 

Rare, very scattered in N and E Taiwan 


Very rare, only known from two locations

305 

Selaginella helvetica subsp. 

pseudonipponica 

Selaginella matsuensis 

Selliguea echinospora 

Selliguea falcatopinnata 

Selliguea okamotoi 

Selliguea taiwanensis 

Woodsia okamotoi 

Not uncommon on limestone in E Taiwan, 

also reported from few locations in S 


Not rare in MaZu and JinMen 


Very rare, only known from two small 

locations 

Probably very rare in SC Taiwan 



Undetermined taxa or unpublished hybrids of chapter 2 List were not 

considered here.

306 

Adiantum cap.-veneris f. lanyuanum 

Adiantum formosanum 

Adiantum meishanianum 

Adiantum taiwanianum

307 

Angiopteris itoi 

Angiopteris somae 

Antrophyum castaneum 

Araiostegia parvipinnula

308 

Asplenium cuneatiforme 

Asplenium x wangii 

Athyrium auriculatum 

Athyrium leiopodum

309 

Athyrium tripinnatum 

Cheilanthes agetae 

Cibotium taiwanense 

Cyclosorus ensifer

310 

Cyclosorus longipetiolatus 

Deparia dawuense 

Diplazium pseudodoederleinii 

Dryopsis x fauriei

311 

Dryopteris cacaina 

Dryopteris kwanzanensis 

Dryopteris pseudolunanensis 

Dryopteris pseudosieboldii

312 

Dryopteris subatrata 

Dryopteris subreflexipinna 


Huperzia cunninghamioides

313 

Hymenophyllum alishanense 

Hymenophyllum devolii 


Isoetes taiwanensis var. kinmenensis

314 

Isoetes taiwanensis var. taiwanensis 

Lepisorus kawakamii 

Lepisorus megasorus 

Lepisorus monilisorus

315 

Lepisorus pseudoussuriensis 

Loxogramme biformis 

Loxogramme remotefrondigera 

Lycopodium yueshanense

316 

Microlepia intramarginalis 

Microlepia krameri 

Microlepia trichocarpa 

Pleocnemia rufinervis

317 

Polypodium raishaense 

Polypodium transpianense 

Polystichum parvipinnulum 

Polystichum pseudostenophyllum

318 

Polystichum simadae 

Polystichum taiwanianum 

Polystichum taizhongense 

Pteris angustipinna

319 

Pteris longipinna 

Pteris pellucidifolia 

Pteris wulaiensis 

Pyrrosia polydactyla

320 

Selaginella devolii 

Selaginella helv. subsp. pseudonipp. 

Selaginella matsuensis 

Selliguea echinospora

321 

Selliguea falcatopinnata 

Selliguea okamotoi 

Selliguea taiwanensis 

Woodsia okamotoi

322

323 

A P P E N D I X B 

- Taxonomic systems 

The system of Kramer and Green (1990) 

For Taiwan's taxa their system is arranged as following: 

Class: Psilotatae 

Order: Psilotales 

Family: Psilotaceae 

Psilotum 

Class: Lycopodiatae 

Order: Isoetales 

Family: Isoetaceae 

Isoetes 

Order: Lycopodiales 

Family: Lycopodiaceae 

Huperzia 

Lycopodiella 

Lycopodium 

Order: Selaginellales 

Family: Selaginellaceae 

Selaginella 

Class: Equisetatae 

Order: Equisetales 

Family: Equisetaceae 

Equisetum 

Class: Filicatae 

Family: Aspleniaceae 

Asplenium 

Family: Azollaceae 

Azolla 

Family: Blechnaceae 

Blechnum 

Brainea 

Woodwardia 

Family: Cheiropleuriaceae 

Cheiropleuria 

Family: Cyatheaceae 

Cyathea 

Family: Davalliaceae 

Araiostegia 

Davallia 

Leucostegia

324 

Family: Dennstaedtiaceae 

Subfamily: Dennstaedtioideae 

Dennstaedtia 

Histiopteris 

Hypolepis 

Microlepia 

Paesia 

Pteridium 

Subfamily: Lindsaeoideae 

Lindsaea 

Odontosoria 

Tapeinidium 

Family: Dicksoniaceae 

Cibotium 

Family: Dipteridaceae 

Dipteris 

Family: Dryopteridaceae 

Subfamily: Athyrioideae 

Tribe: Onocleeae 

Onoclea 

Tribe: Physematieae 

Acystopteris 

Athyrium 

Cornopteris 

Cystopteris 

Deparia 

Dictyodroma 

Diplaziopsis 

Diplazium 

Gymnocarpium 

Hypodematium 

Woodsia 

Subfamily: Dryopteridoideae 

Tribe: Dryopterideae 

Acrophorus 

Arachniodes 

Dryopteris 

Nothoperanema 

Peranema 

Polystichum 

Tribe: Tectarieae 

Ctenitis 

Dryopsis 

Lastreopsis 

Pleocnemia 

Pteridrys

Tectaria 

Family: Gleicheniaceae 

Dicranopteris 

Diplopterygium 

Family: Grammitidaceae 

Calymmodon 

Grammitis 

Scleroglossum 

Family: Hymenophyllaceae 

Cephalomanes 

Crepidomanes 

Hymenophyllum 

Sphaerocionium 

Trichomanes 

Family: Lomariopsidaceae 

Bolbitis 

Elaphoglossum 

Lomariopsis 

Family: Marattiaceae 

Angiopteris 

Marattia 

Family: Marsileaceae 

Marsilea 

Family: Monachosoraceae 

Monachosorum 

Family: Nephrolepidaceae 

Nephrolepis 

Family: Oleandraceae 

Arthropteris 

Oleandra 

Family: Ophioglossaceae 

Botrychium 

Helminthostachys 

Ophioglossum 

Family: Osmundaceae 

Osmunda 

Family: Plagiogyriaceae 

Plagiogyria 

Family: Polypodiaceae 

Subfamily: Platycerioideae 

Pyrrosia 

Subfamily: Polypodioideae 

Tribe: Drynarieae 

Aglaomorpha 

Drynaria 

325

326 

Tribe: Lepisoreae 

Belvisia 

Drymotaenium 

Lemmaphyllum 

Lepisorus 

Tribe: Loxogrammeae 

Loxogramme 

Tribe: Microsoreae 

Colysis 

Leptochilus 

Microsorum 

Neocheiropteris 

Phymatosorus 

Tribe: Polypodieae 

Polypodium 

Tribe: Selligueeae 

Arthromeris 

Selliguea 

Family: Pteridaceae 

Subfamily: Adiantoideae 

Adiantum 

Subfamily: Ceratopteridoideae 

Ceratopteris 

Subfamily: Cheilanthoideae 

Cheilanthes 

Coniogramme 

Cryptogramma 

Paraceterach 

Subfamily: Pteridoideae 

Acrostichum 

Pteris 

Subfamily: Taenitidoideae 

Anogramma 

Onychium 

Pityrogramma 

Family: Salviniaceae 

Salvinia 

Family: Schizaeaceae 

Lygodium 

Schizaea 

Family: Thelypteridaceae 

Cyclosorus 


Phegopteris 


Thelypteris

327 

Family: Vittariaceae 

Antrophyum 

Monogramma 

Vittaria 

The system of Christenhusz et al. (2011) 

Taiwan's taxa can be allocated into the following units, consequent 

alternative combinations are shown in chapter 2 List: 

Subclass: Lycopodiidae 



Isoetes 



Huperzia 

Lycopodiella 

Lycopodium 



Selaginella 

Subclass: Equisetidae 



Equisetum 

Subclass: Ophioglossidae 

Order: Ophioglossales 


Botrychium 


Ophioglossum 



Psilotum 

Subclass: Marattiidae 

Order: Marattiales 


Angiopteris 

Ptisana

328 

Subclass: Polypodiidae 

Order: Cyatheales 

Family: Cibotiaceae 

Cibotium 


Alsophila 

Sphaeropteris 

Family: Plagiogyriaceae 

Plagiogyria 

Order: Gleicheniales 


Cheiropleuriaceae 

Dipteris 


Dicranopteris 


Order: Hymenophyllales 


Abrodictyum 

Callistopteris 

Cephalomanes 

Crepidomanes 

Didymoglossum 

Hymenophyllum 

Vandenboschia 

Order: Osmundales 


Osmunda 

Osmundastrum 

Order: Polypodiales 


Asplenium 


Family: Athyriaceae 

Anisocampium 

Athyrium 

Cornopteris 

Deparia 

Diplazium 

Family: Blechnaceae 

Blechnum 

Brainea 

Diploblechnum 

Woodwardia 

Family: Cystopteridaceae 

Acystopteris

Cystopteris 

Gymnocarpium 

Family: Davalliaceae 

Davallia 

Davallodes 


Dennstaedtia 

Histiopteris 

Hypolepis 

Microlepia 

Monachosorum 

Paesia 

Pteridium 

Family: Diplaziopsidaceae 

Diplaziopsis 

Family: Dryopteridaceae 


Acrophorus 

Acrorumohra 

Arachniodes 

Cyrtogonellum 

Cyrtomidictyum 

Cyrtomium 

Diacalpe 

Dryopteris 

Nothoperanema 

Peranema 

Polystichum 

Subfamily: Elaphoglossoideae 

Bolbitis 

Elaphoglossum 

Lastreopsis 

Family: Hypodematiaceae 

Hypodematium 

Leucostegia 

Family: Lindsaeaceae 

Lindsaea 

Odontosoria 

Tapeinidium 

Family: Lomariopsidaceae 

Lomariopsis 


Nephrolepis 

Family: Oleandraceae 

Oleandra 

329

330 

Family: Onocleaceae 

Onoclea 


Subfamily: Drynarioideae 

Aglaomorpha 

Arthromeris 

Drynaria 

Selliguea 

Subfamily: Loxogrammoideae 

Loxogramme 

Subfamily: Microsoroideae 

Goniophlebium 

Lemmaphyllum 

Lepisorus 

Leptochilus 

Microsorum 

Neocheiropteris 

Phymatosorus 

Subfamily: Platycerioideae 

Pyrrosia 


Calymmodon 

Chrysogrammitis 

Dasygrammitis 

Micropolypodium 

Oreogrammitis 

Prosaptia 


Scleroglossum 

Themelium 

Xiphopterella 



Ceratopteris 


Aleuritopteris 

Cheilanthes 

Coniogramme 

Doryopteris 

Mildella 

Paraceterach 

Paragymnopteris 

Subfamily: Cryptogrammoideae 

Cryptogramma 


Acrostichum

331 

Pteris 

Subfamily: Taenitidoideae 

Anogramma 

Onychium 

Pityrogramma 

Subfamily: Vittarioideae 

Adiantum 

Antrophyum 

Haplopteris 

Monogramma 

Family: Rhachidosoraceae 

Rhachidosorus 

Family: Tectariaceae 

Arthropteris 

Pleocnemia 

Pteridrys 

Tectaria 

Family: Thelypteridaceae 

Cyclosorus 


Phegopteris 


Thelypteris 

Family: Woodsiaceae 

Woodsia 

Order: Salviniales 


Marsilea 


Azolla 

Salvinia 

Order: Schizaeales 

Family: Lygodiaceae 

Lygodium 


Schizaea 

The system of Christenhusz and Chase (2014) 

With reference to Taiwan's taxa it is organized as follows (with changes to 

the previous system shown in red fonts): 

Subclass: Lycopodiidae 



Isoetes

332 



Huperzia 

Lycopodiella 

Lycopodium 



Selaginella 

Subclass: Equisetidae 



Equisetum 

Subclass: Ophioglossidae 

Order: Ophioglossales 


Botrychium 


Ophioglossum 



Psilotum 

Subclass: Marattiidae 

Order: Marattiales 


Angiopteris 

Ptisana 

Subclass: Polypodiidae 

Order: Cyatheales 


Subfamily: Plagiogyrioideae 

Plagiogyria 

Subfamily: Cyatheoideae 

Alsophila 

Gymnosphaera 

Sphaeropteris 

Subfamily: Cibotioideae 

Cibotium 

Order: Gleicheniales 


Cheiropleuriaceae 

Dipteris


Dicranopteris 


Order: Hymenophyllales 


Hymenophyllum 

Trichomanes 

Order: Osmundales 


Osmunda 

Osmundastrum 

Order: Polypodiales 


Subfamily: Asplenioideae 

Asplenium 


Subfamily: Blechnoideae 

Blechnum 

Onoclea 

Woodwardia 

Subfamily: Cystopteridoideae 

Acystopteris 

Cystopteris 

Gymnocarpium 

Subfamily: Rhachidosoroideae 

Rhachidosorus 

Subfamily: Diplaziopsidoideae 

Diplaziopsis 

Subfamily: Thelypteridoideae 


Phegopteris 

Thelypteris 

Subfamily: Woodsioideae 

Woodsia 

Subfamily: Athyrioideae 

Athyrium 

Cornopteris 

Deparia 

Diplazium 


Dennstaedtia 

Histiopteris 

Hypolepis 

Microlepia 

Monachosorum 

Paesia 

333

334 

Pteridium 

Family: Lindsaeaceae 

Lindsaea 

Odontosoria 

Tapeinidium 


Nephrolepis 


Subfamily: Davalliaceae 

Davallia 

Davallodes 


Arachniodes 

Bolbitis 

Ctenitis 

Cyrtomidictyum 

Dryopteris 

Elaphoglossum 

Lastreopsis 

Pleocnemia 

Polystichum 

Subfamily: Hypodematiaceae 

Hypodematium 

Leucostegia 

Subfamily: Lomariopsidoideae 

Lomariopsis 

Subfamily: Oleandroideae 

Oleandra 


Tribe: Loxogrammeae 

Loxogramme 

Tribe: Drynarieae 

Drynaria 

Selliguea 

Tribe: Platycerieae 

Pyrrosia 

Tribe: Microsoreae 

Goniophlebium 

Lemmaphyllum 

Lepidomicrosorium 

Lepisorus 

Leptochilus 

Microsorum 

Neolepisorus 

Phymatosorus

Tribe: Polypodieae 

Grammitis 

Subfamily: Tectariaceae 

Arthropteris 

Pteridrys 

Tectaria 



Acrostichum 

Ceratopteris 


Aleuritopteris 

Cheilanthes 

Doryopteris 

Hemionitis (for Hemionitis cordata) 

Mildella 

Pellaea (for Paragymnopteris) 

Subfamily: Cryptogrammoideae 

Coniogramme 

Cryptogramma 


Anogramma 

Onychium 

Pityrogramma 

Pteris 

Subfamily: Vittarioideae 

Adiantum 

Antrophyum 

Haplopteris 

Monogramma 

Order: Salviniales 


Marsilea 


Azolla 

Salvinia 

Order: Schizaeales 


Subfamily: Lygodioideae 

Lygodium 

Subfamily: Schizaeoideae 

Schizaea 

335

336

This publication is closely linked to the following books which contain a 

detailed set of information on the pteridophyte flora of Taiwan: 

Ferns and Fern Allies of Taiwan (2011) 

1064 pages, 4700 colour photographs, hardcover 

English, with Latin and Chinese species index 

Ferns and Fern Allies of Taiwan - Supplement (2013) 

212 pages, 560 colour photographs, paperback 

English, with Latin and Chinese species index 

Selection of book stores carrying these titles: 

B. K. Norton (Taiwan) http://bknortonbknorton.blogspot.tw/ 

KBCC Press (Taiwan) 

http://www.kbcc.org.tw/tw/ 

Koeltz (Germany) 

http://www.koeltz.com/ 

MBG Press (USA) 

http://www.mbgpress.info/ 

NHBS (UK) 

http://www.nhbs.com/ 

Almost all taxa introduced in the above books and in this publication are 

illustrated in the following Internet based photo collection of the author: 

https://picasaweb.google.com/116136418529949606360?noredirect=1 

(or simply use a web browser with search terms "picasa ralf knapp")

KBCC had a fern colection with 1686 taxa in August 2014, including 173 in 

Polypodiaceae,75 in Aspidiaceae,59 in Pteridaceae,40 in Maratiaceae,and 25 in 

Cyatheaceae. 

Dr.ChiouandAmericanplant 

physiologist,Dr.CraigMartin, 

conductedresearchon 

Platycerium 

Dr.Pei-ChunLiaofrom National 

PingtungUniversityofScience 

andTechnologycolected 

Thelypterissamples 

Ferncolectionmanager,Chen 

ChunMing,givingaworkshop 

onfernsinKBCC 

Treeferncolection 

DavaliaceaeColection 

MaratiaceaeColection

• 

Taiwanisoneoftheglobaldiversitycentersofpteridophytes(fern, 

andplantsaliedtothem)withmorethan700speciesdistributedover 

anareaof35800km 2 . 

ThisindexprovidesacondensedupdatetoknowntaxaofTaiwanandits 

neighboringsmalerislandsandislets. 

•referencepublicationforeachacceptedscientificname 

•localthreatclasification(folowingIUCN) 

•local(vernacular)nameinChineseandpinyintranscription 

•basionym andprotologuedata 

•etymologicaldata 

•alternativecombination(forfitingintocurentphylogeneticsystems) 

•typeinformation 

•firstpublicationofnameforTaiwan 

•comprehensivebackgroundinformation 

•extensivelistofsynonyms,misappliednamesandnamesinalternative 

combinations 

•bibliography 

•ilustratedlistofendemictaxa

R. Knapp_2014_Index Ferns Fern Allies Taiwan_KBCC

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