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Revised checklist of Hawaiian mosses<br />
<strong>Tropical</strong> <strong>Bryology</strong> 25: 35-69, 2004<br />
A revised checklist of Hawaiian mosses 1<br />
G. W. Staples A , C. T. Imada A , W. J. Hoe, B and C. W. Smith C<br />
A – Hawaii Biological Survey, Bishop Museum, 1525 Bernice St., Honolulu, Hawaii 96817<br />
U.S.A.<br />
B – deceased;<br />
C – Department of Botany, The Natural History Museum, Cromwell Road, London, SW7<br />
5BD, U.K.<br />
Abstract. A revised and updated literature-based checklist of Hawaiian mosses is presented.<br />
Geographic coverage includes the eight main Hawaiian Islands; the Northwestern Hawaiian Islands<br />
are excluded. The checklist is alphabetically ordered by scientific names; the family is noted for each<br />
genus. Synonyms and misapplied names are cross-referenced to the accepted names. A bibliography<br />
of supporting references is included.<br />
Introduction<br />
The Hawaii Biological Survey (HBS) was<br />
established as a program of Bishop Museum by<br />
the Hawaii State Legislature in 1992, specifically<br />
to create and maintain accurate inventories of all<br />
species present in the Hawaiian Islands, and to<br />
document their presence with voucher specimen<br />
collections and bibliographies of relevant<br />
scientific literature. In the ensuing eleven years,<br />
comprehensive taxonomic inventories have been<br />
compiled for many groups of organisms, both<br />
native and alien, that comprise the Hawaiian biota.<br />
Vascular plants and marine algae have been the<br />
primary focus of botanical inventory efforts;<br />
research on non-marine algae, terrestrial<br />
cryptogamic plants, fungi, and lichens has lagged<br />
behind.<br />
This checklist is intended to fill one of<br />
the gaps in our knowledge of Hawaiian terrestrial<br />
1 Contribution No. 2004–001 to the Hawaii<br />
Biological Survey.<br />
TROPICAL BRYOLOGY 25 (2004)<br />
35<br />
cryptogamic plants—the mosses. Also in<br />
preparation by HBS staff members are checklists<br />
and bibliographies for the Hawaiian anthocerotes<br />
(hornworts) and hepatics (liverworts). A nonmarine<br />
algal checklist for the Hawaiian Islands is<br />
also in an advanced stage of preparation (A.<br />
Sherwood, pers. comm.). Eventually a checklist<br />
of Hawaiian lichens is planned. These ongoing<br />
inventories will help the Hawaii Biological Survey<br />
to fulfill its mission by systematically compiling<br />
literature-based inventories for each of these<br />
groups of organisms.<br />
Genesis of the current checklist<br />
This checklist of Hawaiian mosses begins<br />
where its predecessor—the checklist compiled<br />
by Hoe (1974)—left off, and updates that<br />
document by revising the nomenclature, adding<br />
new taxa and distributional records, and<br />
providing a bibliography of literature that<br />
supports the names in current use. The checklist
36<br />
and bibliography were compiled by a<br />
collaborative effort between HBS staff members<br />
(Staples, Imada) working in partnership with two<br />
bryological specialists (Hoe, Smith), who had<br />
assembled a cache of literature citations and new<br />
distribution records pertinent to Hawaiian<br />
mosses. Funding from the USDA–PLANTS<br />
database project in 1998 provided the impetus<br />
for Staples and Imada to undertake conversion<br />
of Hoe’s 1974 moss checklist to an electronic<br />
format and then begin to update the taxonomy<br />
and distribution information it contained. Hoe<br />
and Smith generously shared their preliminary<br />
cache of information in order to facilitate the<br />
update process. Sadly, in May 2000, Hoe died<br />
suddenly, a great loss to the botanical community<br />
in Hawai‘i and to the world bryological<br />
community. After some reorganization the project<br />
resumed and was completed by Staples and<br />
Imada. In the final version of the checklist,<br />
taxonomic decisions were made by Staples, with<br />
review and critical input from Smith and two<br />
independent reviewers.<br />
Coverage<br />
It must be emphasized that this is a<br />
literature-based taxonomic checklist, not a<br />
specimen-based inventory. It is compiled from<br />
literature published from the mid-1970s to the<br />
present (2003). The starting point is the checklist<br />
published by Hoe (1974), who summarized the<br />
literature and reconciled the scientific names for<br />
Hawaiian mosses in use up to that time. The<br />
present checklist attempts to account for all<br />
scientific names in use for Hawaiian mosses in<br />
the latter decades of the twentieth century; it is<br />
not a comprehensive nomenclator for all names<br />
that have ever been published for, or applied to,<br />
Hawaiian mosses. No nomenclatural novelties or<br />
taxonomic changes are initiated in this checklist.<br />
Geographically, coverage focuses on the<br />
eight main Hawaiian Islands. There are, as yet,<br />
no published reports for bryophytes from the<br />
uninhabited Northwestern Hawaiian Islands,<br />
although a few bryophyte collections were made<br />
there in the last decades of the twentieth century.<br />
It had been Bill Hoe’s intention to identify these<br />
voucher specimens and write them up for<br />
publication but the specimens remain unstudied<br />
as this checklist goes to print. They await<br />
Staples et al.<br />
inclusion in a future iteration of the Hawaiian<br />
moss checklist.<br />
In the last quarter of the twentieth<br />
century there was a renaissance of bryological<br />
research worldwide and as a result many<br />
revisions, floristic accounts, and several<br />
monographs have been published that impact<br />
Hawaiian moss nomenclature. These regional and<br />
global studies resulted in significant taxonomic<br />
changes, as well as reporting discovery of several<br />
taxa new to the Islands, either rare native mosses<br />
or newly naturalized alien species. This checklist<br />
aims to summarize all this information in a concise<br />
format, as follows.<br />
Format<br />
Accepted scientific name<br />
The main body of the checklist is<br />
alphabetically organized by genus, species, and<br />
infraspecific taxon. Accepted names appear in<br />
bold italic type. Authors are included for the<br />
genera, species, and infraspecific epithets.<br />
Authors of moss genera have been standardized<br />
against the classification proposed in Bryophyte<br />
Biology (Shaw & Goffinet 2000); authors for five<br />
genera not found in that classification have been<br />
checked on the MOST website (cited below). All<br />
author names are abbreviated according to<br />
Authors of Plant Names style (Brummitt & Powell<br />
1992). References that explain the taxonomic<br />
concept or provide geographic distribution data<br />
are cited after the accepted taxon name.<br />
The first mention of a genus includes<br />
the family assignment on the same line. Most<br />
genera are still placed in the same families<br />
recognized by Hoe (1974), but a few new family<br />
placements have been made since then. In such<br />
cases, we provide first the family assignment<br />
used by Hoe (1974), and mention alternative<br />
family placements afterward, with a reference to<br />
the source.<br />
Synonyms and misapplied names (in<br />
italics) follow the accepted name in paragraph<br />
style. These are limited primarily to names in use<br />
for Hawaiian mosses in the latter part of the<br />
twentieth century, although a few names from<br />
other parts of the world that enter into the<br />
nomenclature for particular taxa have been<br />
included to enable tracking names through the<br />
literature.<br />
TROPICAL BRYOLOGY 25 (2004)
Revised checklist of Hawaiian mosses<br />
Status<br />
Following the taxonomy is a column<br />
indicating status of the taxon in the Hawaiian<br />
flora: end indicates an endemic, ind indicates<br />
indigenous, and nat signifies naturalized. A ?<br />
following any of these signifies uncertainty about<br />
the status of the taxon in the Hawaiian flora. While<br />
some research has been devoted to the mode of<br />
introduction of pteridophytes and angiosperms<br />
to Hawai‘i, resulting in a list of species that are<br />
likely to have been introduced by Polynesian<br />
voyagers that settled the archipelago and another<br />
(much larger) list of plants that arrived following<br />
European contact, little attention has been<br />
devoted to bryophytes in this regard. The authors<br />
have used their best judgment in cases where<br />
the status is not clear and appended a ? to<br />
whatever status we assigned, to draw attention<br />
to the species and invite investigation from<br />
specialists.<br />
One case worth noting is Racopilum<br />
cuspidigerum, which is widespread in the<br />
Southern Hemisphere but is found only in the<br />
Hawaiian Islands in the Northern Hemisphere. It<br />
is a prime candidate as a Polynesian introduction<br />
to Hawai‘i, and indeed to many of the other<br />
Polynesian islands in the South Pacific (S. Olson,<br />
pers. comm.).<br />
Island distribution<br />
Following the status category is a<br />
summary of geographic distribution on the eight<br />
main Hawaiian Islands. The island name<br />
abbreviations are explained at the foot of each<br />
page. An island abbreviation appears in the row<br />
to the right of the taxon name to signify that the<br />
taxon is reported in the literature from that island.<br />
Absence of the island name abbreviation<br />
indicates the moss is not (yet) known from that<br />
island. Given the inadequate state of moss<br />
collecting in the Hawaiian Islands, many new<br />
island distribution records are sure to be found<br />
in the future.<br />
Synonym list and Literature Consulted<br />
Following the main body of the checklist,<br />
an alphabetical list of all synonyms and<br />
misapplied names is provided. This list crossreferences<br />
every synonym or misapplied name<br />
to the main entry found under the scientific name<br />
TROPICAL BRYOLOGY 25 (2004)<br />
37<br />
accepted in this checklist. The checklist<br />
concludes with a bibliography of the references<br />
cited in the checklist; more than 130 bryological<br />
titles were consulted in the course of the project,<br />
but only those providing taxonomic or<br />
distribution data pertinent to this checklist are<br />
included here.<br />
Anomalies and incomplete records<br />
A few moss names without any island<br />
distribution are included, based on (mostly) older<br />
literature that reports the species from the<br />
Hawaiian (or sometimes Sandwich) Islands<br />
without stating on which island the species was<br />
found. Two examples in the list are Dicranoweisia<br />
cirrata and Plagiothecium denticulatum. Their<br />
presence in the Hawaiian Islands needs to be<br />
confirmed and documented with contemporary<br />
collections.<br />
Another anomaly is Mniobryoides<br />
degeneri, an invalid name published without a<br />
description or indication of a type specimen by<br />
H. Hörmann. It has been pointed out (S. Olson,<br />
pers. comm.) that this is probably a<br />
misidentification for some well-known species.<br />
However, M. degeneriae has never been assessed<br />
by a competent bryologist to ascertain its true<br />
identity. We leave the name in the checklist to<br />
point out that more study is required.<br />
A note about family circumscriptions<br />
The past 25 years has brought an<br />
explosion in research on bryophyte systematics<br />
and a great deal of new information has accrued,<br />
which has led to many changes in moss<br />
classification. Family classification in particular<br />
has become increasingly problematic as new data<br />
provide insights into relationships between<br />
genera. Several new families have been erected<br />
and other genera have been reassigned. We have<br />
indicated the family where Hoe (1974) assigned<br />
each genus, followed by alternative family<br />
assignments where these were discovered. Two<br />
principle reference works were consulted for<br />
family assignments: first, Bryophyte Biology<br />
(Shaw & Goffinet 2000), and in a few cases the<br />
bryological nomenclator included in the<br />
W 3 MOST database of mosses (http://<br />
mobot.mobot.org/W3T/Search/most.html) on the<br />
TROPICOS website.
38<br />
The way forward<br />
It is hoped that by bringing together the literature<br />
and providing an updated checklist of names in<br />
current use, one or more competent bryologists<br />
might be inspired to undertake preparation of a<br />
moss flora for the Hawaiian Islands. The last<br />
floristic account of the Hawaiian mosses is that<br />
by Bartram (1933, 1939). Bill Hoe intended to write<br />
a modern flora of the Hawaiian mosses, but his<br />
untimely death prevented that. It is our hope that<br />
professional bryologists will take up the challenge<br />
to complete this much needed reference work.<br />
Those who would study Hawaiian<br />
mosses will find type material and historic<br />
specimens widely distributed in herbaria around<br />
the world. In the last decades of the twentieth<br />
century, D. Vitt and his students as well as a team<br />
of cryptogamic botanists from the National<br />
Museum of Tokyo collected mosses throughout<br />
the main Hawaiian Islands. But by far the most<br />
comprehensive late twentieth century collections<br />
of Hawaiian mosses are those made by Hoe<br />
between 1965 and 1995. Today the first set of<br />
Hoe’s collections resides in two herbaria: the<br />
Herbarium Pacificum, Bishop Museum, Honolulu<br />
(BISH) and the Claude Philipps Herbarium,<br />
Delaware State University, Dover (DOV), with<br />
duplicates widely distributed in other herbaria.<br />
Hoe donated his personal herbarium to BISH in<br />
1983. His residual duplicates and some new<br />
material acquired after 1983 were deposited at<br />
DOV after the sale of Hoe’s bryological library<br />
and herbarium. The Hoe collections in BISH and<br />
DOV provide an excellent resource for bryological<br />
researchers and afford a solid foundation for any<br />
floristic account of Hawaiian mosses.<br />
Hawaiian Moss Flora in Summary<br />
Endemic Indigenous Alien Total<br />
Families0 41 1 42<br />
Genera 2 126 7 135<br />
Species 75 166 14 255<br />
Total named taxa 90 169 14 273<br />
Staples et al.<br />
Acknowledgments<br />
Funding for initial data compilation was<br />
made available by the USDA-PLANTS program;<br />
we thank Scott Petersen and Mark Skinner for<br />
their support and assistance. Our Museum<br />
colleagues Katie Anderson and Karin Kami<br />
assisted with data entry and bibliographic<br />
compilation. Bishop Museum librarian Patrice<br />
Belcher obtained numerous critical references<br />
through interlibrary loan services. Reviews of the<br />
draft checklist by Storrs Olson, U.S. National<br />
Museum, and James Shevock, U.S. Geological<br />
Survey, brought to light several bryological<br />
articles overlooked during the compilation as well<br />
as many suggestions for its improvement. Bill<br />
Buck, New York Botanical Garden, added further<br />
literature citations and name changes when he<br />
edited the manuscript for <strong>Tropical</strong> <strong>Bryology</strong>. The<br />
authors gratefully acknowledge all assistance<br />
rendered; any errors of fact or interpretation are<br />
the sole responsibility of the authors.<br />
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Norris, D.H., and T. Koponen. 1989. Bryophyte<br />
flora of the Huon Peninsula, Papua New<br />
Guinea. XXVIII. Acta Botanica Fennica 137:<br />
81–138.<br />
Ochi, H. 1974. Some Bryaceous „Old World“<br />
mosses, also distributed in the New World.<br />
Journal of the Faculty of Education, Tottori<br />
University, Natural Sciences 25(1–2): 35–41.<br />
TROPICAL BRYOLOGY 25 (2004)
Revised checklist of Hawaiian mosses<br />
Ochi, H. 1980. A revision of the Neotropical<br />
Bryoideae, Musci (first part). Journal of the<br />
Faculty of Education, Tottori University,<br />
Natural Sciences 29(2): 49–154 (see p. 100).<br />
Olson, S.L. 1999a. Muscis Hawaiiensis<br />
Mantissae I: Two weedy species of Bryum new<br />
to the archipelago. <strong>Tropical</strong> <strong>Bryology</strong> 17: 129–<br />
130.<br />
Olson, S.L. 1999b. Muscis Hawaiiensis<br />
Mantissae II. Neckeropsis obtusata (Mont.)<br />
Fleisch. restored to the flora of the archipelago.<br />
<strong>Tropical</strong> <strong>Bryology</strong> 17: 131–132.<br />
Orbán, S., and W.D. Reese. 1986. Notes on the<br />
taxonomy of African Syrrhopodon. Abstracta<br />
Botanica 10: 349–355.<br />
Pursell, R.A. 1997. Taxonomic notes on<br />
neotropical Fissidens. II. An addendum.<br />
Bryologist 100(2): 193–197.<br />
Pursell, R.A., and W.J. Hoe. 1977. Fissidens in<br />
Hawaii. Journal of the Hattori Botanical<br />
Laboratory 43: 81–106.<br />
Reese, W.D., T. Koponen, and D.H. Norris. 1986.<br />
Bryophyte flora of the Huon Peninsula, Papua<br />
New Guinea. XIX. Acta Botanica Fennica 133:<br />
151–202.<br />
Schultze-Motel, W. 1963. Beitrag zur Kenntnis<br />
der Laubmoose der Hawaii-Inseln.<br />
Willdenowia 3: 97–107.<br />
Shaw, A.J., and B. Goffinet (eds.). 2000.<br />
Bryophyte Biology. Cambridge University Press,<br />
New York, x + 476 pp.<br />
Shaw, J. 1985. Nomenclatural changes in the<br />
Bryaceae subfamily Mielichhoferioideae.<br />
Bryologist 88(1): 28–30.<br />
Sollman, P. 1984. Notes on pottiaceous mosses<br />
II. Lindbergia 10(1): 53–56.<br />
Spence, J.R. 1996. Rosulabryum genus novum<br />
(Bryaceae). Bryologist 99(2): 221–225.<br />
Streimann, H. 1993. Barbella trichophora, an<br />
older name for B. cubensis (Musci:<br />
Meteroriaceae). Bryologist 96: 223–225.<br />
Tan, B.C. 1984. Additional notes on Asiatic and<br />
Pacific taxa of Meteoriaceae. Yushania 1(4):<br />
7–12.<br />
Tan, B.C., and Z. Iwatsuki. 1991. A new annotated<br />
Philippine moss checklist. Harvard Papers in<br />
Botany 3: 1–64.<br />
Tan, B.C., and Y. Jia. 1999. A preliminary revision<br />
of Chinese Sematophyllaceae. Journal of the<br />
Hattori Botanical Laboratory 86: 1–70.<br />
TROPICAL BRYOLOGY 25 (2004)<br />
41<br />
Tan, B.C., and T. Koponen. 1989. Additions and<br />
corrections for Philippine moss flora.<br />
Cryptogamie, Bryologie-Lichénologie 10(3):<br />
235–245.<br />
Tangney, R.S. 1997. A taxonomic revision of the<br />
genus Camptochaete Reichdt.,<br />
Lembophyllaceae (Musci). Journal of the<br />
Hattori Botanical Laboratory 81: 53–121.<br />
Tixier, P. 1988. Le genre Glossadelphus Fleisch.<br />
(Sematophyllaceae, Musci) et sa valeur. Nova<br />
Hedwigia 46(3–4): 319–356.<br />
Touw, A. 2001. A taxonomic revision of the<br />
Thuidiaceae (Musci) of tropical Asia, the<br />
western Pacific, and Hawaii. Journal of the<br />
Hattori Botanical Laboratory 91: 1–136.<br />
Virtanen, V. 1997. The genus Breutelia<br />
(Bartramiaceae, Musci) in Southeast Asia and<br />
Oceania. Bryologist 100(3): 324–348.<br />
Vitt, D.H. 1980a. A comparative study of<br />
Andreaea acutifolia, A. mutabilis, and A.<br />
rupestris. New Zealand Journal of Botany 18:<br />
367–377.<br />
Vitt, D.H. 1980b. The genus Macrocoma I.<br />
Typification of names and taxonomy of the<br />
species. Bryologist 83(4): 405–436.<br />
Vitt, D.H. 1981. The genus Macrocoma II.<br />
Geographical variation in the Macrocoma<br />
tenue—M. sullivantii species complex.<br />
Bryologist 83(4): 437–450.<br />
Vitt, D.H. 1983. The New Zealand species of the<br />
pantropical genus Macromitrium<br />
(Orthotrichaceae: Musci): taxonomy,<br />
phylogeny and phytogeography. Journal of<br />
the Hattori Botanical Laboratory 54: 1–94.<br />
Vitt, D.H., and W.J. Hoe. 1980. Andreaea in<br />
Hawaii. Bryologist 83(2): 212–214.<br />
W 3 MOST data base. http://mobot.mobot.org/<br />
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Watanabe, R., and Z. Iwatsuki. 1981. New name<br />
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Wichman, J.R., and H. St. John. 1990. A chronicle<br />
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Wilczek, R., and F. Demaret. 1976. Bryum<br />
flavituber Wilczek & Demaret, espèce nouvelle<br />
des Iles Hawaii. Bulletin de la Jardin Botanique<br />
National de Belgique 46: 407–408.
42<br />
Zander, R.H. 1977. The tribe Pleuroweisieae<br />
(Pottiaceae, Musci) in Middle America.<br />
Bryologist 80(2): 233–269.<br />
Zander, R.H. 1978. New combinations in<br />
Didymodon (Musci) and a key to the taxa in<br />
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Zander, R.H. 1982. Oxystegus tenuirostris var.<br />
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Bryologica et Lichenologica 9(4): 72–74.<br />
Zander, R.H. 1993. Genera of Pottiaceae: mosses<br />
of harsh environments. Bulletin of the Buffalo<br />
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Zander, R.H., and P.M. Eckel. 1982.<br />
Hymenostylium recurvirostrum var. insigne<br />
and Barbula amplexifolia in British Columbia,<br />
Canada. Canadian Journal of Botany 60: 1596–<br />
1600.<br />
Zander, R.H., and W.J. Hoe. 1979. Geographic<br />
disjunction and heterophylly in Tortella<br />
fragilis var. tortelloides (= Sarconeurum<br />
tortelloides). Bryologist 82(1): 84–87.<br />
Zander, R.H., and D.H. Vitt. 1979. Gametophytic<br />
distinctions of Zygodon (Orthotrichaceae) and<br />
Anoectangium and Leptodontium (Pottiaceae)<br />
and the status of Anoectangium<br />
rubrigemmium of Hawaii. Canadian Journal of<br />
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Zanten, B.O. van. 1959. Trachypodaceae—a<br />
critical revision. Blumea 9(2): 477–575.<br />
Zanten, B.O. van. 1971. Musci. Pp. 173–227. In<br />
Zinderen-Bakker, E.M. van, J.M.<br />
Winterbottom, and R.A. Dyer (eds.), Marion<br />
and Prince Edward Islands. A. A. Balkema,<br />
Cape Town.<br />
Zilstra, G. 1990. Report of the Committee for<br />
Bryophyta: I. Taxon 39(2): 289–292.<br />
Staples et al.<br />
Island abbreviations: Ni Ni‘ihau; K Kaua‘i; O<br />
O‘ahu; Mo Moloka‘i; M Maui; Ka Kahoolawe;<br />
H Hawai‘i.<br />
TROPICAL BRYOLOGY 25 (2004)
Revised checklist of Hawaiian mosses<br />
TROPICAL BRYOLOGY 25 (2004)<br />
<strong>Full</strong>Name STATUS NI K O MO L M KA H<br />
ACROPORIUM Mitt. (Sematophyllaceae)<br />
Acroporium fuscoflavum (Paris) Broth. [Hoe 1974,<br />
Miller et al. 1978]<br />
Acroporium fuscoflavum (Paris) Broth. var.<br />
baldwinii (Paris) Broth.<br />
?Acroporium sigmatodontium misapplied<br />
AEROBRYOPSIS M.Fleisch. (Meteoriaceae)<br />
Aerobryopsis subdivergens (Broth.) Broth. ssp. scariosa<br />
(E.B.Bartram) Nog. [Hoe 1974, Noguchi 1976]<br />
Aerobryopsis scariosa E.B.Bartram<br />
Ptychomnion aciculare misapplied<br />
Aerobryopsis wallichii (Brid.) M.Fleisch. [Hoe 1974,<br />
Noguchi 1976]<br />
Aerobryopsis longissima (Dozy & Molk.) M.Fleisch.<br />
Aerobryopsis longissima (Dozy & Molk.)<br />
M.Fleisch. var. dozyana M.Fleisch.<br />
ind K O Mo L M H<br />
ind K O Mo M<br />
ind K O Mo L M H<br />
AMPHIDIUM Schimp. (Dicranaceae, or<br />
Rhabdoweisiaceae [Shaw & Goffinet 2000])<br />
Amphidium lapponicum (Hedw.) Schimp. [Hoe 1979] ind M<br />
Amphidium tortuosum (Hornsch.) Cufod. [Frahm et al.<br />
2000, Muñoz & Pando 2000]<br />
Amphidium cyathicarpum (Mont.) Broth.<br />
Grimmia haleakalae Reichardt<br />
Grimmia haliacalae Reichardt<br />
ind M H<br />
ANDREAEA Hedw. (Andreaeaceae)<br />
Andreaea acutifolia Hook. F. & Wilson ssp. acuminata<br />
(Mitt.) Vitt [Vitt 1980a, Vitt & Hoe 1980]<br />
Andreaea acuminata Mitt.<br />
Andreaea rupestris Hedw. [Hoe 1977, Hoe 1979, Vitt<br />
1980a, Vitt & Hoe 1980]<br />
Andreaea perpapillosa Broth.<br />
ANOECTANGIUM Schwägr. (Pottiaceae)<br />
Anoectangium aestivum (Hedw.) Mitt. [Hoe 1974,<br />
Zander 1977, Zander & Vitt 1979]<br />
Anoectangium euchloron (Schwägr.) Mitt.<br />
Anoectangium haleakalae (Müll.Hal.) Paris var.<br />
haleakalae<br />
Anoectangium haleakalae (Müll.Hal.) Paris var.<br />
laxulum E.B.Bartram<br />
Anoectangium harttiae E.B.Bartram<br />
ind M H<br />
ind M<br />
ind K O Mo L M H<br />
ANOMOBRYUM Schimp. (Bryaceae)<br />
Anomobryum angustirete Broth. [Hoe 1974] end K L M<br />
43
44<br />
Staples et al.<br />
<strong>Full</strong>Name<br />
BALDWINIELLA M.Fleisch. (Neckeraceae)<br />
STATUS NI K O MO L M KA H<br />
Baldwiniella kealeensis (Reichardt) E.B.Bartram [Hoe<br />
1974, Hoe 1975]<br />
end K O Mo L M H<br />
BARBELLOPSIS Broth. (Meteoriaceae)<br />
Barbellopsis trichophora (Mont.) W.R.Buck [Hoe 1974,<br />
Streimann 1993, Buck 1998]<br />
Barbella trichophora (Mont.) M.Fleisch.<br />
BARBULA Hedw. (Pottiaceae)<br />
Barbula indica (Hook.) Spreng. [Hoe 1978, Zander 1993,<br />
Darigo 1996]<br />
Barbula cruegeri Sond. ex Müll.Hal.<br />
Barbula unguiculata misapplied<br />
ind K O Mo M H<br />
nat O M<br />
BARTRAMIA Hedw. (Bartramiaceae)<br />
Bartramia baldwinii Müll.Hal. [Hoe 1974] end M<br />
Bartramia halleriana Hedw. [Hoe 1974] ind M<br />
BRACHYMENIUM Schwägr. (Bryaceae)<br />
Brachymenium exile (Dozy & Molk.) Bosch & Sande<br />
Lac. [Hoe 1974]<br />
Brachymenium indicum (Dozy & Molk.) Bosch & Sande<br />
Lac. [Hoe 1979]<br />
ind K O Mo M H<br />
ind O H<br />
BRACHYTHECIUM Schimp. (Brachytheciaceae)<br />
Brachythecium hawaiicum E.B.Bartram [Hoe 1974] end M H<br />
Brachythecium lamprocarpum (Müll.Hal.) A.Jaeger<br />
[Hoe 1974]<br />
ind K M H<br />
Brachythecium plumosum (Hedw.) B.S.G. [Hoe 1974]<br />
Brachythecium oxyrrhynchum (Dozy & Molk.)<br />
A.Jaeger<br />
Cirriphyllum oxyrrhynchum (Dozy & Molk.)<br />
M.Fleisch.<br />
ind K O Mo M H<br />
Brachythecium rutabulum (Hedw.) B.S.G. [Hoe 1974] ind M<br />
BREUTELIA (Bruch & Schimp.) Schimp.<br />
(Bartramiaceae)<br />
Breutelia affinis (Hook.) Mitt. [Hoe 1974, Virtanen 1997]<br />
Breutelia kilaueae (Müll.Hal.) Broth.<br />
ind H<br />
Breutelia arundinifolia (Duby) M.Fleisch. [Hoe 1974,<br />
Virtanen 1997]<br />
ind K O M H<br />
Breutelia crassicaulis (Müll.Hal.) Paris [Virtanen 1997] ind M H<br />
BRYOERYTHROPHYLLUM P.C.Chen (Pottiaceae)<br />
Bryoerythrophyllum recurvirostre (Hedw.) P.C.Chen<br />
[Hoe 1974]<br />
Didymodon recurvirostris (Hedw.) Jenn.<br />
ind M<br />
TROPICAL BRYOLOGY 25 (2004)
Revised checklist of Hawaiian mosses<br />
TROPICAL BRYOLOGY 25 (2004)<br />
<strong>Full</strong>Name STATUS NI K O MO L M KA H<br />
BRYUM Hedw. (Bryaceae)<br />
Bryum argenteum Hedw. [Hoe 1974, Miller et al. 1978,<br />
Tan & Koponen 1989]<br />
Bryum argenteum Hedw. var. lanatum (P. Beauv.)<br />
Hampe<br />
Bryum leucophyllum Dozy & Molk.<br />
Bryum atrovirens Brid. [Hoe 1974]<br />
Bryum erythrocarpum Schwägr.<br />
ind K O M H<br />
ind O M H<br />
Bryum baldwinii Broth. [Hoe 1974] end M<br />
Bryum caespiticium Hedw. [Hoe 1974] ind M H<br />
Bryum chrysoneuron Müll.Hal. [Miller et al. 1978] ind H<br />
Bryum conicum Hornsch. [Ochi 1980]<br />
Bryum ceramiocarpum Müll.Hal.<br />
ind M<br />
Bryum flavituber R.Wilczek & Demaret [Wilczek &<br />
Demaret 1976]<br />
nat O<br />
Bryum hawaiicum Hoe [Hoe 1974]<br />
Bryum crassicostatum Broth.<br />
end M H<br />
Bryum klinggraeffii Schimp. [Hoe 1979] nat O<br />
Bryum lisae De Not. var. cuspidatum (Bruch & Schimp.) nat O<br />
Margad. [Olson 1999a]<br />
Bryum mildeanum Jur. [Hoe 1974, Ochi 1974, Hoe 1979,<br />
Ignatov & Afonina 1992]<br />
Bryum apiculatum Schwägr.<br />
Bryum mauiensis Broth.<br />
Bryum megalostegium Sull.<br />
Bryum nitens Hook.<br />
ind K O L M<br />
Bryum pseudocapillare Besch. [Hoe 1975] nat? O<br />
Bryum radiculosum Brid. [Olson 1999a] nat K<br />
BUCKIELLA Ireland (Hypnaceae)<br />
Buckiella draytonii (Sull.) Ireland [Hoe 1974, Ireland<br />
2001]<br />
Hypnum draytoni(i) Sull.<br />
Hypnum eudorae Sull.<br />
Plagiothecium draytonii (Sull.) Müll.Hal.<br />
end K O Mo M H<br />
CALYMPERES Sw. (Calymperaceae)<br />
Calymperes fasciculatum Dozy & Molk. [Hoe 1974,<br />
Reese et al. 1986]<br />
Calymperes hawaiiense E.B.Bartram<br />
ind O<br />
Calymperes tenerum Müll.Hal. [Hoe 1974] ind K O Mo M H<br />
CAMPTOCHAETE Reichardt (Lembophyllaceae)<br />
Camptochaete pulvinata (Hook.f. & Wilson) A.Jaeger<br />
[Hoe 1974, Tangney 1997]<br />
ind M<br />
CAMPYLOPODIUM (Müll.Hal.) Besch. (Dicranaceae)<br />
Campylopodium medium (Duby) Giese & J.-P.Frahm ind K O Mo M H<br />
45
46<br />
Staples et al.<br />
<strong>Full</strong>Name STATUS NI K O MO L M KA H<br />
[Hoe 1974, Giese & Frahm 1985]<br />
Campylopodium euphorocladum (Müll.Hal.) Besch.<br />
CAMPYLOPUS Brid. (Dicranaceae)<br />
Campylopus exasperatus (Nees & Blume) Brid. [Frahm<br />
1991]<br />
Campylopus fragilis (Brid.) Bruch. & Schimp. ssp.<br />
zollingerianus (Müll.Hal.) J.-P.Frahm [Hoe 1974,<br />
Frahm 1991, Frahm 1992]<br />
Campylopus boswellii (Müll.Hal.) Paris<br />
Campylopus boswellii (Müll.Hal.) Paris var.<br />
capitulatus E.B.Bartram<br />
Campylopus hawaiicus (Müll.Hal.) A.Jaeger var.<br />
densifolius (Ångstr.) J.-P.Frahm [Frahm 1978, Frahm<br />
1991]<br />
Campylopus densifolius Ångstr.<br />
Campylopus densiretis Broth.<br />
Campylopus wawreanus Reichardt<br />
Campylopus hawaiicus (Müll.Hal.) A.Jaeger var.<br />
hawaiicoflexuosus (Müll.Hal.) J.-P.Frahm [Frahm<br />
1978, Frahm 1991]<br />
Campylopus densifolius Ångstr. var. falcatus Broth.<br />
Campylopus densifolius Ångstr. var.<br />
hawaiicoflexuosus (Müll.Hal.) H.A.Mill.<br />
Campylopus hawaiicoflexuosus (Müll.Hal.)<br />
Müll.Hal.<br />
Campylopus hawaiicoflexuosus (Müll.Hal.) Paris<br />
Campylopus perangustifolius Müll.Hal.<br />
Campylopus purpureoflavescens Hampe, in part<br />
Dicranum hawaiicoflexuosum Müll.Hal.<br />
Campylopus hawaiicus (Müll.Hal.) A.Jaeger var.<br />
hawaiicus [Frahm 1978, Frahm 1991]<br />
Campylopus densifolius Ångstr. var.<br />
purpureoflavescens (Hampe ex Müll.Hal.)<br />
H.A.Mill.<br />
Campylopus purpureoflavescens Hampe, in part<br />
Dicranum hawaiicum Müll.Hal.<br />
Campylopus incurvatus J.-P.Frahm & Hoe [Frahm & Hoe<br />
1978, Frahm 1991]<br />
Campylopus laxi<strong>text</strong>us Sande Lac. [Hoe 1974, Frahm<br />
1991]<br />
Campylopus fumarioli Müll.Hal.<br />
Campylopus praemorsus (Müll.Hal.) A.Jaeger [Frahm<br />
1976, Hoe 1979, Frahm 1991]<br />
Campylopus exasperatus (Nees & Blume) Brid.<br />
Campylopus microcephalus (Müll.Hal.) Paris<br />
Dicranum microcephalum Müll.Hal.<br />
Dicranum praemorsum Müll.Hal.<br />
ind K<br />
ind K O Mo M<br />
end K O Mo M H<br />
end K O Mo L M H<br />
ind K O Mo L M H<br />
end M H<br />
ind K O Mo L M H<br />
end K O L M H<br />
TROPICAL BRYOLOGY 25 (2004)
Revised checklist of Hawaiian mosses<br />
<strong>Full</strong>Name STATUS NI K O MO L M KA H<br />
Campylopus schmidii (Müll.Hal.) A.Jaeger ssp.<br />
hemitrichius (Müll.Hal.) J.-P.Frahm [Hoe 1974, Frahm<br />
1991]<br />
Campylopus skottsbergii Broth.<br />
Campylopus tubulosus E.B.Bartram<br />
Dicranum skottsbergii Broth.<br />
ind K O M H<br />
Campylopus schmidii (Müll.Hal.) A.Jaeger ssp. schmidii<br />
[Hoe 1974, Hoe 1977, Frahm 1991]<br />
Campylopus aureus Bosch & Sande Lac.<br />
Campylopus introflexus misapplied<br />
Campylopus introflexus (Hedw.) Brid. var.<br />
polytrichoides misapplied<br />
Campylopus polytrichoides misapplied<br />
ind K O M H<br />
Campylopus umbellatus (Arn.) Paris var. pterotoneuron<br />
(Müll.Hal.) J.-P.Frahm [Hoe 1974, Frahm 1991]<br />
Campylopus geniculatus Ångstr.<br />
ind O<br />
Campylopus umbellatus (Arn.) Paris var. umbellatus<br />
[Hoe 1974, Frahm 1991]<br />
?Campylopus didrichsenii (Müll.Hal.) A.Jaeger<br />
?Campylopus pertristis (Müll.Hal.) Paris<br />
Campylopus richardii misapplied<br />
Campylopus sandwicensis Paris<br />
Campylopus umbellatus (Arn.) E.B.Bartram<br />
Dicranum didrichsenii Müll.Hal.<br />
Dicranum pertriste Müll.Hal.<br />
Thysanomitrion hawaiicum Müll.Hal.<br />
Thysanomitrion umbellatum Arn.<br />
ind K O L M H<br />
Campylopus wheeleri (Müll.Hal.) Paris [Hoe 1974,<br />
Frahm 1991]<br />
Dicranoloma wheeleri (Müll.Hal.) Paris<br />
Dicranum wheeleri Hampe ex Müll.Hal.<br />
end K M H<br />
CERATODON Brid. (Ditrichaceae)<br />
Ceratodon purpureus (Hedw.) Brid. ssp. purpureus [Hoe<br />
1974, Hoe 1977, Miller et al. 1978, Burley & Pritchard<br />
1990]<br />
Ceratodon purpureus (Hedw.) Brid. fma. purpureus<br />
Ceratodon purpureus (Hedw.) Brid. fma. xanthopus<br />
(Sull.) E.Britton ex Grout<br />
CHENIA R.H.Zander (Pottiaceae)<br />
Chenia leptophylla (Müll.Hal.) R.H.Zander [Hoe 1974,<br />
Zander 1993]<br />
Tortula rhizophylla (Sakurai) Z.Iwats. & K.Saito<br />
Tortula vectensis E.F.Warb. & Crundw.<br />
CLAOPODIUM (Lesq. & James) Renauld & Cardot<br />
(Thuidiaceae, or Leskeaceae [Shaw & Goffinet 2000])<br />
Claopodium prionophyllum (Müll. Hal.) Broth. [Hoe<br />
1974]<br />
TROPICAL BRYOLOGY 25 (2004)<br />
ind K O M H<br />
nat O<br />
ind O Mo M H<br />
47
48<br />
Staples et al.<br />
<strong>Full</strong>Name<br />
Claopodium amblystegioides Dixon<br />
Claopodium nervosum M.Fleisch.<br />
STATUS NI K O MO L M KA H<br />
Claopodium whippleanum (Sull.) Renauld & Cardot<br />
[Hoe 1974]<br />
Claopodium hawaiiense R.S.Williams<br />
ind K O M<br />
CLASTOBRYOPSIS M.Fleisch. (Sematophyllaceae)<br />
Clastobryopsis robusta (Broth.) M.Fleisch. [Hoe 1974,<br />
Tan & Jia 1999]<br />
Aptychella robusta (Broth.) M.Fleisch.<br />
CTENIDIUM (Schimp.) Mitt. (Hypnaceae)<br />
Ctenidium elegantulum Broth. [Hoe 1974, Nishimura<br />
1985]<br />
ind K M<br />
end O M H<br />
CYRTOPUS (Brid.) Hook.f. (Cyrtopodaceae)<br />
Cyrtopus setosus (Hedw.) Hook.f. [Hoe 1974] ind H<br />
DALTONIA Hook. & Taylor (Hookeriaceae, or<br />
Daltoniaceae [Shaw & Goffinet 2000])<br />
Daltonia baldwinii Broth. [Hoe 1974] end M H<br />
Daltonia contorta Müll.Hal. [Hoe 1974, Hoe 1979] ind K O Mo L M H<br />
Daltonia pseudostenophylla E.B.Bartram [Hoe 1974] ind O M H<br />
Daltonia rufescens Broth. [Hoe 1974] ind O M H<br />
DICRANELLA (Müll.Hal.) Schimp. (Dicranaceae)<br />
Dicranella exilis Sull. [Hoe 1974] end M<br />
Dicranella hawaiica (Müll.Hal.) Broth. var. hawaiica<br />
[Hoe 1974]<br />
Microdus hawaiicus (Müll.Hal.) Paris<br />
end K Mo M<br />
Dicranella hawaiica (Müll.Hal.) Broth. var. tomentella<br />
E.B.Bartram [Hoe 1974]<br />
end O M<br />
Dicranella hochreutineri Cardot [Hoe 1974]<br />
Dicranella hochreuteri misapplied<br />
ind K O L M H<br />
Dicranella integrifolia E.B.Bartram [Hoe 1974] end O Mo M H<br />
Dicranella rigidula E.B.Bartram [Hoe 1974] end K O H<br />
DICRANODONTIUM Bruch & Schimp. (Dicranaceae)<br />
Dicranodontium denudatum (Brid.) E.Britton [Hoe 1974] ind M<br />
Dicranodontium falcatum Broth. var. atrovirens (Broth.)<br />
E.B.Bartram [Hoe 1974]<br />
end M H<br />
Dicranodontium porodictyon Cardot & Thér. [Hoe 1974,<br />
Hoe 1979, Frahm 1997]<br />
Dicranodontium falcatum Broth. var. falcatum<br />
Dicranodontium hawaiicum Broth.<br />
ind K O Mo M H<br />
DICRANOLOMA (Renauld) Renauld (Dicranaceae)<br />
Dicranoloma gracile Broth. ex E.B.Bartram [Hoe 1974] end M<br />
TROPICAL BRYOLOGY 25 (2004)
Revised checklist of Hawaiian mosses<br />
TROPICAL BRYOLOGY 25 (2004)<br />
<strong>Full</strong>Name STATUS NI K O MO L M KA H<br />
DICRANOWEISIA Milde (Dicranaceae, or<br />
Rhabdoweisiaceae [Shaw & Goffinet 2000])<br />
Dicranoweisia cirrata (Hedw.) Lindb. [Hoe 1974, Miller<br />
et al. 1978]<br />
DICRANUM Hedw. (Dicranaceae)<br />
Dicranum speirophyllum Mont. var. breviflagellare<br />
(Müll.Hal.) E.B.Bartram [Hoe 1974]<br />
Dicranum speirophyllum Mont. var. condensatum (Sull.)<br />
Wijk & Margad. [Hoe 1974]<br />
Dicranum speirophyllum Mont. var. elongatum (Sull.)<br />
Wijk & Margad. [Hoe 1974]<br />
Dicranum speirophyllum Mont. var. speirophyllum [Hoe<br />
1974]<br />
DIDYMODON Hedw. (Pottiaceae)<br />
Didymodon insulanus (De Not.) M.O.Hill [Hoe 1974,<br />
Zander 1978, Corley et al. 1981]<br />
Barbula vinealis Brid. var. cylindrica (Taylor)<br />
Boulay<br />
Barbula vinealis Brid. var. flaccida B.S.G.<br />
Didymodon vinealis (Brid.) R.H.Zander var.<br />
flaccidus (Bruch & Schimp.) R.H.Zander<br />
ind H?<br />
end O M H<br />
end H<br />
end M<br />
end K O Mo M H<br />
ind O M<br />
DISTICHIUM Bruch & Schimp. (Ditrichaceae)<br />
Distichium capillaceum (Hedw.) B.S.G. [Hoe 1974] ind H<br />
DISTICHOPHYLLUM Dozy & Molk. (Hookeriaceae,<br />
or Daltoniaceae [Shaw & Goffinet 2000])<br />
Distichophyllum freycinetii (Schwägr.) Mitt. var.<br />
crasseturgescens (Müll.Hal.) E.B.Bartram ex Hoe [Hoe<br />
1974]<br />
Mniadelphus freycineti (Schwägr.) Müll.Hal. var.<br />
crasseturgescens Müll.Hal.<br />
end<br />
Distichophyllum freycinetii (Schwägr.) Mitt. var.<br />
freycinetii [Hoe 1974]<br />
end K O Mo L M H<br />
Distichophyllum paradoxum (Mont.) Mitt. [Hoe 1974] end K O Mo L M H<br />
ECTROPOTHECIUM Mitt. (Hypnaceae)<br />
Ectropothecium arcuatum Mitt. [Hoe 1974] end K O M H<br />
Ectropothecium decurrens (Sull.) Nishimura [Hoe 1974,<br />
Nishimura 1985]<br />
Ctenidium decurrens (Sull.) Broth.<br />
Ctenidium pulcherrimum Broth.<br />
Ctenidium submalacobolum Broth.<br />
Cupressina decurrens (Sull.) Müll.Hal.<br />
Ectropothecium rockii Broth.<br />
Hypnum decurrens Sull.<br />
end K O Mo M H<br />
49
50<br />
Staples et al.<br />
<strong>Full</strong>Name<br />
Stereodon decurrens (Sull.) Mitt.<br />
Taxiphyllum gracile Broth.<br />
STATUS NI K O MO L M KA H<br />
Ectropothecium sandwichense (Hook. & Arn.) Mitt.<br />
[Hoe 1974, Hoe 1979]<br />
ind K O Mo L M H<br />
Ectropothecium tutuilum (Sull.) Mitt. [Miller et al. 1978] ind H<br />
Ectropothecium viridifolium E.B.Bartram [Hoe 1974, end Ni O L M<br />
Wichman & St. John 1990]<br />
Ectropothecium zollingeri (Müll. Hal.) A.Jaeger var.<br />
filicaule (M.Fleisch.) H.A.Mill., H.Whittier &<br />
B.Whittier [Hoe 1974, Hoe 1975, Miller et al. 1978]<br />
Glossadelphus zollingeri (Müll.Hal.) M.Fleisch. var.<br />
filicaulis (M.Fleisch.) M.Fleisch.<br />
ind K O Mo L M H<br />
ENCALYPTA Hedw. (Encalyptaceae)<br />
Encalypta ciliata Hedw. [Horton 1983]<br />
Encalypta scabrata E.B.Bartram<br />
ind M H<br />
Encalypta rhabdocarpa Schwägr. [Hoe 1974]<br />
Encalypta rhaptocarpa misapplied<br />
ind H<br />
Encalypta sandwicensis Sull. [Hoe 1974, Hoe 1977] ind M H<br />
ENTODON Müll.Hal. (Entodontaceae)<br />
Entodon plicatus Müll.Hal. [Hoe 1974, Buck 1990]<br />
Entodon hillebrandii Müll.Hal.<br />
Entodon solanderi (Ångstr.) A.Jaeger<br />
Entodon subcuspidatus (Müll.Hal.) E.B.Bartram [Hoe<br />
1974, Hoe 1979]<br />
ENTODONTOPSIS Broth. (Stereophyllaceae)<br />
Entodontopsis anceps (Bosch & Sande Lac.) W.R.Buck<br />
& Ireland [Hoe 1974, Buck & Ireland 1985]<br />
Stereophyllum oahuense Broth.<br />
ENTOSTHODON Schwägr. (Funariaceae)<br />
Entosthodon subintegrus (Broth.) H.A.Mill., H.Whittier<br />
& B.Whittier [Miller et al. 1978]<br />
Funaria subintegra Broth.<br />
ind K O M H<br />
ind K Mo M<br />
ind O<br />
end K O Mo L M H<br />
FABRONIA Raddi (Fabroniaceae)<br />
Fabronia ciliaris (Brid.) Brid. [Hoe 1977] ind M<br />
Fabronia degeneri E.B.Bartram [Hoe 1974] end O Mo H<br />
Fabronia nietneri Müll.Hal. [Hoe 1974, Hoe 1975] ind K O M H<br />
FISSIDENS Hedw. (Fissidentaceae)<br />
Fissidens aphelotaxifolius Pursell [Pursell & Hoe 1977] ind M<br />
Fissidens bryoides Hedw. [Pursell & Hoe 1977]<br />
Fissidens hawaiicus E.B.Bartram<br />
Fissidens insularis E.B.Bartram<br />
Fissidens oahuensis E.B.Bartram<br />
ind K O Mo L M H<br />
TROPICAL BRYOLOGY 25 (2004)
Revised checklist of Hawaiian mosses<br />
<strong>Full</strong>Name STATUS NI K O MO L M KA H<br />
Fissidens delicatulus Ångstr. [Pursell & Hoe 1977] end K O L M H<br />
Fissidens elegans Brid. [Pursell & Hoe 1977]<br />
Fissidens baldwinii Broth.<br />
Fissidens bishopii Hoe<br />
ind K O Mo L M H<br />
Fissidens hoei Pursell [Pursell & Hoe 1977] end O L<br />
Fissidens kilaueae Hoe & H.A.Crum [Pursell & Hoe<br />
1977]<br />
end K O L M H<br />
Fissidens lancifolius E.B.Bartram [Pursell & Hoe 1977] end K O Mo L M H<br />
Fissidens lindbergii Mitt. [Pursell & Hoe 1977, Darigo<br />
1996, Pursell 1997]<br />
Fissidens crumii Hoe<br />
nat O M<br />
Fissidens nothotaxifolius Pursell & Hoe [Pursell & Hoe<br />
1977]<br />
end K O Mo L M H<br />
Fissidens pacificus Ångstr. [Hoe 1974, Pursell & Hoe<br />
1977]<br />
Fissidens mauiensis Müll.Hal.<br />
end K O M H<br />
Fissidens submarginatus Bruch [Pursell & Hoe 1977,<br />
Hoe 1979, Bruggeman-Nannenga & Pursell 1995]<br />
Fissidens intermedius Müll.Hal.<br />
ind K O Mo L M H<br />
Fissidens taxifolius Hedw. [Pursell & Hoe 1977] ind O M<br />
FLORIBUNDARIA M.Fleisch. (Meteoriaceae)<br />
Floribundaria floribunda (Dozy & Molk.) M.Fleisch.<br />
[Hoe 1974, Tan 1984]<br />
Floribundaria baldwinii Broth.<br />
TROPICAL BRYOLOGY 25 (2004)<br />
ind O M H<br />
FUNARIA Hedw. (Funariaceae)<br />
Funaria hygrometrica Hedw. [Hoe 1974, Hoe 1975] ind O H<br />
GAROVAGLIA Endl. (Pterobryaceae, or Garovagliaceae<br />
[Shaw & Goffinet 2000])<br />
Garovaglia powellii Mitt. var. haleakalae (Broth.) During<br />
[During 1977]<br />
Garovaglia haleakalae Broth.<br />
ind M<br />
GLOSSADELPHUS M.Fleisch. 1 (Sematophyllaceae)<br />
Glossadelphus acutifolius E.B.Bartram [Hoe 1974, Tixier<br />
1988]<br />
end M<br />
Glossadelphus limnobioides Broth. [Hoe 1974, Tixier<br />
1988]<br />
end O M H<br />
Glossadelphus mauiensis Broth. [Hoe 1974, Tixier 1988] end M<br />
Glossadelphus ogatae Broth. & Yasuda [Hoe 1974, Hoe<br />
1979, Tixier 1988]<br />
Glossadelphus chrysobasilaris Broth.<br />
ind M H<br />
1 Buck (1987b) reduced the type species of Glossadelphus to Phyllodon so the genus name is not usable.<br />
The correct placement for the 4 species left here remains to be determined.<br />
51
52<br />
Staples et al.<br />
<strong>Full</strong>Name<br />
GRIMMIA Hedw. (Grimmiaceae)<br />
STATUS NI K O MO L M KA H<br />
Grimmia anodon Bruch & Schimp. [Hoe 1977, Hoe<br />
1979, Muñoz & Pando 2000]<br />
Grimmia apocarpa Hedw. var. pulvinata (Hedw.)<br />
G.N.Jones<br />
Gymnostomum pulvinatum Hedw.<br />
ind M H<br />
Grimmia laevigata (Brid.) Brid. [Hoe 1974, Hoe 1979,<br />
Muñoz & Pando 2000]<br />
ind M H<br />
Grimmia longirostris Hook. [Hoe 1977,Muñoz 1998]<br />
Grimmia pilifera misapplied<br />
ind M<br />
Grimmia mauiense Greven [Greven 2003] end M H<br />
Grimmia maunakeaense Greven [Greven 2003] end M H<br />
Grimmia pulvinata (Hedw.) Sm. [Muñoz & Pando 2000]<br />
Leersia pulvinata Hedw.<br />
ind M<br />
Grimmia trichophylla Grev. [Hoe 1974, Muñoz & Pando<br />
2000]<br />
ind M H<br />
HAGENIELLA Broth. (Sematophyllaceae)<br />
Hageniella micans (Mitt.) B.C.Tan & Y.Jia [Hoe 1974,<br />
Tan & Jia 1999]<br />
Hageniella pacifica Broth.<br />
HAPLOCLADIUM Müll.Hal. (Leskeaceae)<br />
Haplocladium microphyllum (Hedw.) Broth. [Hoe 1974,<br />
Watanabe & Iwatuski 1981, Zilstra 1990]<br />
Bryohaplocladium microphyllum (Hedw.) R.Watan.<br />
& Z.Iwats.<br />
HAPLOHYMENIUM Dozy & Molk. (Thuidiaceae, or<br />
Anomodontaceae [Shaw & Goffinet 2000])<br />
Haplohymenium triste (Ces.) Kindb. [Hoe 1974, Hoe<br />
1979]<br />
HETEROPHYLLIUM (Schimp.) Kindb.<br />
(Sematophyllaceae)<br />
Heterophyllium subauriculatum (Müll.Hal.) Broth. [Hoe<br />
1974]<br />
HOLOMITRIUM Brid. (Dicranaceae)<br />
Holomitrium seticalycinum Müll.Hal. [Hoe 1974, Hoe<br />
1979]<br />
Holomitrium squarrifolium E.B.Bartram [Hoe 1974]<br />
Holomitrium ferriei misapplied<br />
HOMALIODENDRON M.Fleisch. (Neckeraceae)<br />
Homaliodendron flabellatum (Sm.) M.Fleisch. [Hoe<br />
1974]<br />
Homalia densa Bosw.<br />
Homalia praelonga Reichardt<br />
ind K M H<br />
ind O<br />
ind K O M H<br />
end M H<br />
end K O Mo L M H<br />
end O H<br />
ind K O Mo L M H<br />
TROPICAL BRYOLOGY 25 (2004)
Revised checklist of Hawaiian mosses<br />
TROPICAL BRYOLOGY 25 (2004)<br />
<strong>Full</strong>Name STATUS NI K O MO L M KA H<br />
HOOKERIA Sm. (Hookeriaceae)<br />
Hookeria acutifolia Hook. & Grev. [Hoe 1974, Hoe<br />
1975]<br />
HYMENOSTYLIUM Brid. (Pottiaceae)<br />
Hymenostylium recurvirostrum (Hedw.) Dixon var.<br />
cylindricum (E.B.Bartram) R.H.Zander [Zander &<br />
Eckel 1982]<br />
Merceyopsis crassinervis (Broth.) E.B.Bartram<br />
Molendoa crassinervis (Broth.) E.B.Bartram<br />
Hymenostylium recurvirostrum (Hedw.) Dixon var.<br />
recurvirostrum [Zander 1993]<br />
Hymenostylium firmum (Müll.Hal.) Broth. ex<br />
E.B.Bartram<br />
HYOPHILA Brid. (Pottiaceae)<br />
Hyophila involuta (Hook.) A.Jaeger [Hoe 1974]<br />
Hyophila dozy-molkenboeri M.Fleisch.<br />
HYPNUM Hedw. (Hypnaceae)<br />
Hypnum plumaeforme Wilson [Crum & Mueller-<br />
Dombois 1968, Hoe 1974]<br />
HYPOPTERYGIUM Brid. (Hypopterygiaceae)<br />
Hypopterygium sandwicense Broth. [Hoe 1974, Kruijer<br />
2002]<br />
ind O L M H<br />
ind H<br />
ind K O M H<br />
ind K O Mo M H<br />
nat M H<br />
end M H<br />
ISOPTERYGIUM Mitt. (Hypnaceae)<br />
Isopterygium albescens (Hook.) A.Jaeger [Hoe 1974] ind K O Mo L M H<br />
Isopterygium vineale E.B.Bartram [Hoe 1974] end K O Mo M H<br />
LEPTOBRYUM (Bruch & Schimp.) Wilson (Bryaceae,<br />
or Meesiaceae [Shaw & Goffinet 2000])<br />
Leptobryum pyriforme (Hedw.) Wilson [Hoe 1974] ind M H<br />
LEPTODONTIUM (Müll.Hal.) Lindb. (Pottiaceae)<br />
Leptodontium flexifolium (Dicks. ex With.) Hampe [Hoe<br />
1974, Hoe 1977]<br />
Leptodontium brevicaule E.B.Bartram<br />
ind M H<br />
LEUCOBRYUM Hampe (Leucobryaceae)<br />
Leucobryum gracile Sull. var. gracile [Hoe 1974] end K O Mo L M H<br />
Leucobryum gracile Sull. var. hamatum E.B.Bartram<br />
[Hoe 1974]<br />
end K O Mo M<br />
Leucobryum pachyphyllum Müll.Hal. [Hoe 1974] end K O Mo L M<br />
Leucobryum seemannii Mitt. var. fumarioli (Müll.Hal.)<br />
H.A.Mill. [Hoe 1974]<br />
end H<br />
53
54<br />
Staples et al.<br />
<strong>Full</strong>Name<br />
Leucobryum hawaiense (Reichardt) E.B.Bartram var.<br />
fumarioli (Müll.Hal.) E.B.Bartram<br />
Leucobryum solfatarae Müll.Hal. var. fumarioli<br />
(Müll.Hal.) Wijk & Margad.<br />
STATUS NI K O MO L M KA H<br />
Leucobryum seemannii Mitt. var. seemannii [Hoe 1974,<br />
Hoe 1975]<br />
Leucobryum hawaiense (Reichardt) E.B.Bartram<br />
Leucobryum solfatarae Müll.Hal. var. hawaiense<br />
end K O Mo L M H<br />
(Reichardt) Wijk & Marg.<br />
Leucobryum seemannii Mitt. var. solfatarae (Müll.Hal.)<br />
H.A.Mill. [Hoe 1974]<br />
Leucobryum hawaiense (Reichardt) E.B.Bartram var.<br />
solfatarae (Müll.Hal.) E.B.Bartram<br />
Leucobryum solfatarae Müll.Hal.<br />
end K Mo M H<br />
LEUCOLOMA Brid. (Dicranaceae)<br />
Leucoloma molle (Müll.Hal.) Mitt. [Hoe 1974] ind O M H<br />
Leucoloma scaberulum E.B.Bartram [Hoe 1974] end O<br />
LEUCOPHANES Brid. (Leucobryaceae, or<br />
Calymperaceae [Shaw & Goffinet 2000])<br />
Leucophanes glaucum (Schwägr.) Mitt. [Hoe 1974] ind O<br />
LIMBELLA (Müll.Hal.) Müll.Hal. (Amblystegiaceae)<br />
Limbella tricostata (Sull.) Müll.Hal. [Hoe 1974, Christy<br />
1987]<br />
Sciaromium tricostatum (Sull.) Mitt.<br />
MACROCOMA (Müll.Hal.) Grout (Orthotrichaceae)<br />
Macrocoma tenue (Hook. & Grev.) Vitt ssp. sullivantii<br />
(Müll.Hal.) Vitt [Hoe 1974, Vitt 1980b, Vitt 1981]<br />
Macrocoma intricata (Müll.Hal.) H.A.Miller,<br />
H.Whittier & B.Whittier<br />
Macromitrium intricatum Müll.Hal.<br />
end K O Mo M H<br />
ind K O Mo M H<br />
MACROMITRIUM Brid. (Orthotrichaceae)<br />
Macromitrium brevisetum Mitt. [Hoe 1974, Hoe 1979] end K O Mo L M H<br />
Macromitrium emersulum Müll.Hal. [Hoe 1974, Hoe<br />
1975]<br />
end K O Mo L M H<br />
Macromitrium incurvifolium (Hook. & Grev.) Schwägr.<br />
[Hoe 1974]<br />
Macromitrium cumingii Müll.Hal.<br />
ind K M H<br />
Macromitrium microstomum (Hook. & Grev.) Schwägr.<br />
[Hoe 1974, Vitt 1983]<br />
Macromitrium owahiense Müll.Hal.<br />
Macromitrium reinwardtii Schwägr.<br />
ind K O Mo L M H<br />
Macromitrium piliferum Schwägr. [Hoe 1974] end K O Mo L M H<br />
TROPICAL BRYOLOGY 25 (2004)
Revised checklist of Hawaiian mosses<br />
<strong>Full</strong>Name<br />
MACROTHAMNIUM M.Fleisch. (Hylocomiaceae)<br />
STATUS NI K O MO L M KA H<br />
Macrothamnium macrocarpum (Reinw. & Hornsch.)<br />
M.Fleisch. [Hoe 1974]<br />
ind M<br />
MICRODUS Schimp. ex Besch. (Dicranaceae)<br />
Microdus hillebrandii (Müll.Hal.) Paris [Hoe 1974]<br />
Dicranella hillebrandii (Müll.Hal.) Broth.<br />
TROPICAL BRYOLOGY 25 (2004)<br />
ind O Mo<br />
MNIOBRYOIDES Hörmann (Bryaceae)<br />
Mniobryoides degeneriae Hörmann [Hoe 1974] end H<br />
MNIUM Hedw. (Mniaceae)<br />
Mnium marginatum (With.) P.Beauv. [Hoe 1974]<br />
Mnium serratum Schrad. ex Brid.<br />
ind M<br />
NECKERA Hedw. (Neckeraceae)<br />
Neckera hawaiicopennata Müll.Hal. [Hoe 1974] end M<br />
NECKEROPSIS Reichardt (Neckeraceae)<br />
Neckeropsis lepineana (Mont.) M.Fleisch. [Hoe 1974,<br />
Enroth 1989]<br />
ind K O M<br />
Neckeropsis obtusata (Mont.) Broth. [Olson 1999b] nat K<br />
OCTOBLEPHARUM Hedw. (Leucobryaceae, or<br />
Calymperaceae [Shaw & Goffinet 2000])<br />
Octoblepharum albidum Hedw. [Hoe 1974, Hoe 1975] ind K O Mo M H<br />
ORTHODONTIUM Wilson (Bryaceae, or<br />
Orthodontiaceae [Shaw & Goffinet 2000])<br />
Orthodontium pellucens (Hook.) B.S.G. [Hoe 1974, Hoe<br />
1977]<br />
ind K M<br />
ORTHOTRICHUM Hedw. (Orthotrichaceae)<br />
Orthotrichum alpestre Hornsch. var. berggrenii<br />
(E.B.Bartram) Lewinsky [Hoe 1974, Lewinksy 1993]<br />
Orthotrichum berggrenii E.B.Bartram<br />
end O? H<br />
Orthotrichum diaphanum Schrad. ex Brid. [Hoe 1974,<br />
Hoe 1977, Lewinsky 1993]<br />
ind M H<br />
Orthotrichum hawaiicum Müll.Hal. [Hoe 1974] end M H<br />
Orthotrichum incrassatum Lewinsky [Lewinsky 1993] end H<br />
Orthotrichum lyellii Hook. & Taylor [Hoe 1974,<br />
Lewinsky 1993]<br />
Orthotrichum verrucatum Müll.Hal.<br />
ind<br />
Orthotrichum rupestre Schleich. & Schwägr. [Hoe 1974,<br />
Lewinsky 1993]<br />
Orthotrichum hillebrandii Müll.Hal.<br />
ind O M H<br />
55
56<br />
Staples et al.<br />
<strong>Full</strong>Name<br />
OXYRRHYNCHIUM (Schimp.) Warnst.<br />
(Brachytheciaceae)<br />
STATUS NI K O MO L M KA H<br />
Oxyrrhynchium vagans (A.Jaeger) Ignatova & Huttunen<br />
[Hoe 1974, Hoe 1975, Ignatov & Huttunen 2002]<br />
Eurhynchium vagans (A.Jaeger) E.B.Bartram<br />
Rhynchostegium vagans A.Jaeger<br />
ind K O Mo L M H<br />
OXYSTEGUS (Limpr.) Hilp. (Pottiaceae)<br />
Oxystegus tenuirostris (Hook. & Taylor) A.J.E.Sm. var.<br />
gemmiparus (Schimp.) R.H.Zander [Hoe 1977, Hoe<br />
1979, Zander 1982]<br />
Trichostomum tenuirostre (Hook. & Taylor) Lindb.<br />
PALAMOCLADIUM Müll.Hal. (Brachytheciaceae)<br />
Palamocladium wilkesianum (Sull.) Müll.Hal. [Hoe<br />
1974, Hofmann 1997]<br />
Hypnum wilkesianum Sull.<br />
Meteorium pallidovirens Müll.Hal.<br />
Meteorium sciuroides Müll.Hal.<br />
Palamocladium aptychoides Müll.Hal.<br />
Palamocladium sciurellum Müll.Hal.<br />
Palamocladium wilkesianum (Sull.) Müll.Hal. var.<br />
altisetum Müll.Hal.<br />
Palamocladium wilkesianum (Sull.) Müll.Hal. var.<br />
sciuroides (Müll.Hal.) Wijk & Margad.<br />
Pleuropus aptychoides (Müll.Hal.) Broth.<br />
Pleuropus pallidovirens (Müll.Hal.) Broth.<br />
Pleuropus sciurellus (Müll.Hal.) Broth.<br />
Pleuropus wilkesianus (Sull.) Broth.<br />
Pleuropus wilkesianus (Sull.) Broth. var. sciuroides<br />
(Müll.Hal.) E.B.Bartram<br />
PELEKIUM Mitt. (Thuidiaceae)<br />
Pelekium versicolor (Hornsch. ex Müll.Hal.) Touw [Hoe<br />
1974, Touw 2001]<br />
Thuidium crenulatum Mitt.<br />
Thuidium nanophyllum Müll.Hal.<br />
ind K O M<br />
end K O Mo L M H<br />
ind O M H<br />
PHILONOTIS Brid. (Bartramiaceae)<br />
Philonotis falcata (Hook.) Mitt. [Hoe 1974] ind K O Mo M<br />
Philonotis hastata (Duby) Wijk & Margad. [Hoe 1974]<br />
Philonotis laxissima Mitt.<br />
ind O M H<br />
Philonotis hawaica (Müll.Hal.) Broth. [Hoe 1974] end K O Mo L M H<br />
Philonotis turneriana (Schwägr.) Mitt. var. sullivantii<br />
(Müll.Hal.) E.B.Bartram [Hoe 1974]<br />
end O M H<br />
Philonotis turneriana (Schwägr.) Mitt. var. turneriana<br />
[Hoe 1974]<br />
ind K O Mo L M H<br />
TROPICAL BRYOLOGY 25 (2004)
Revised checklist of Hawaiian mosses<br />
<strong>Full</strong>Name STATUS NI K O MO L M KA H<br />
PILOTRICHELLA (Müll.Hal.) Besch. (Meteoriaceae,<br />
or Lembophyllaceae [Shaw & Goffinet 2000])<br />
Pilotrichella mauiensis (Sull.) A.Jaeger [Hoe 1974] end O M H<br />
PLAGIOMNIUM T.J.Kop. (Mniaceae)<br />
Plagiomnium rostratum (Schrad.) T.J.Kop. [Hoe 1974]<br />
Mnium rostratum Schrad.<br />
Plagiomnium prorepens (Müll.Hal.) T.J.Kop.<br />
Plagiomnium rhynchosporum (Hook.) T.J.Kop.<br />
PLAGIOPUS Brid. (Bartramiaceae)<br />
Plagiopus oederiana (Sw.) H.A.Crum & L.E.Anderson<br />
[Hoe 1974]<br />
Plagiopus longisetus (Brid.) E.B.Bartram<br />
Plagiopus oederi (Brid.) Limpr.<br />
TROPICAL BRYOLOGY 25 (2004)<br />
ind K O Mo M H<br />
ind M<br />
PLAGIOTHECIUM Bruch & Schimp.<br />
(Plagiotheciaceae)<br />
Plagiothecium cavifolium (Brid.) Z.Iwats. [Hoe 1977] ind M<br />
Plagiothecium denticulatum (Hedw.) Schimp. [Hoe<br />
1974]<br />
ind<br />
Plagiothecium mauiense Broth. [Hoe 1974] end M<br />
PLATYHYPNIDIUM M.Fleisch. (Amblystegiaceae, or<br />
Brachytheciaceae [Shaw & Goffinet 2000])<br />
Platyhypnidium muelleri (A.Jaeger) M.Fleisch. [Hoe<br />
1974]<br />
Eurhynchium mulleri misapplied<br />
POGONATUM P. Beauv. (Polytrichaceae)<br />
Pogonatum tahitense Schimp. ex Besch. [Hoe 1974]<br />
Pogonatum baldwinii (Müll.Hal.) Paris<br />
POHLIA Hedw. (Bryaceae, or Mniaceae [Shaw &<br />
Goffinet 2000])<br />
Pohlia baldwinii (Broth. ex E.B.Bartram) W.Schultze-<br />
Motel [Hoe 1974]<br />
Webera baldwinii Broth. ex E.B.Bartram<br />
Pohlia cruda (Hedw.) Lindb. [Hoe 1974, Hoe 1979]<br />
Webera cruda (Hedw.) Fürnr.<br />
Pohlia flexuosa Hook. [Hoe 1974, Gangulee 1974]<br />
Pohlia gracilescens (E.B.Bartram) E.B.Bartram<br />
Pohlia gracillima (Cardot) Horik. & Ochi<br />
Pohlia leucostoma (Bosch & Sande Lac.) M.Fleisch.<br />
Pohlia leucostomoides (Broth.) E.B.Bartram<br />
Webera gracilescens E.B.Bartram<br />
Webera leucostomoides Broth.<br />
Pohlia mauiensis (Broth. ex E.B.Bartram) W.Schultze-<br />
Motel [Hoe 1974]<br />
ind K O Mo M<br />
ind K O Mo L M H<br />
end M<br />
ind M H<br />
ind K O Mo M H<br />
end M H<br />
57
58<br />
Staples et al.<br />
<strong>Full</strong>Name STATUS NI K O MO L M KA H<br />
POLYTRICHUM Hedw. (Polytrichaceae)<br />
Polytrichum commune Hedw. [Hoe 1974] ind H<br />
Polytrichum juniperinum Hedw. [Hoe 1974, Hyvönen<br />
1986]<br />
ind H<br />
Polytrichum piliferum Hedw. [Hoe 1974] ind M H<br />
PSEUDOSCLEROPODIUM (Limpr.) M.Fleisch.<br />
(Brachytheciaceae)<br />
Pseudoscleropodium purum (Hedw.) M.Fleisch. [Hoe<br />
1974]<br />
PSEUDOSYMBLEPHARIS Broth. (Pottiaceae)<br />
Pseudosymblepharis angustata (Mitt.) Hilp. [Hoe 1974,<br />
Hoe 1979, Norris & Koponen 1989]<br />
Pseudosymblepharis mauiensis (Mull.Hal.) Broth.<br />
PSEUDOTAXIPHYLLUM Z.Iwats. (Hypnaceae)<br />
Pseudotaxiphyllum elegans (Brid.) Z.Iwats. [Hoe 1979,<br />
Iwatsuki 1987]<br />
Isopterygium elegans (Brid.) Lindb.<br />
nat H<br />
ind K O Mo M H<br />
ind M<br />
PTEROGONIDIUM Broth. (Sematophyllaceae)<br />
Pterogonidium pulchellum (Hook.) Müll.Hal. [Hoe 1979] nat? O<br />
PTYCHOMITRIUM Fürnr. (Ptychomitriaceae)<br />
Ptychomitrium lepidomitrium (Müll.Hal.) Schimp. ex<br />
Besch. [Hoe 1974, Cao et al. 2003]<br />
Ptychomitrium mauiense Broth.<br />
PYLAISIADELPHA Broth. (Meteoriaceae)<br />
Pylaisiadelpha opaeodon (Sull.) W.R. Buck [Hoe 1974,<br />
Buck 1984]<br />
Brotherella opaeodon (Sull.) Broth.<br />
Hypnum opaeodon Sull.<br />
PYRRHOBRYUM Mitt. (Rhizogoniaceae)<br />
Pyrrhobryum spiniforme (Hedw.) Mitt. [Hoe 1974,<br />
Koponen et al. 1986]<br />
Rhizogonium spiniforme (Hedw.) Bruch<br />
RACOMITRIUM Brid. (Grimmiaceae)<br />
Racomitrium crispulum (Hook.f. & Wilson) Hook.f. &<br />
Wilson [Hoe 1974]<br />
Racomitrium fasciculare (Schrad. ex Hedw.) Brid. var.<br />
erosum Broth. [Hoe 1974]<br />
Racomitrium lanuginosum (Hedw.) Brid. var.<br />
lanuginosum [Hoe 1974]<br />
ind M H<br />
ind K O Mo M H<br />
ind K O Mo L M H<br />
ind K M H<br />
end M<br />
ind M<br />
TROPICAL BRYOLOGY 25 (2004)
Revised checklist of Hawaiian mosses<br />
<strong>Full</strong>Name STATUS NI K O MO L M KA H<br />
Racomitrium lanuginosum (Hedw.) Brid. var.<br />
pruinosum Wilson [Hoe 1974]<br />
Racomitrium lanuginosum (Hedw.) Brid. var.<br />
sandvicense Reichardt<br />
ind K O Mo M H<br />
RACOPILUM P.Beauv. (Racopilaceae)<br />
Racopilum cuspidigerum (Schwägr.) Ångstr. [Hoe 1974]<br />
Rhacomitrium cuspidigerum misapplied<br />
RADULINA W.R.Buck & B.C.Tan (Sematophyllaceae)<br />
Radulina hamata (Dozy & Molk.) W.R.Buck & B.C.Tan<br />
[Hoe 1974, Buck & Tan 1989]<br />
Trichosteleum hamatum (Dozy & Molk.) A.Jaeger<br />
REMYELLA Müll.Hal. (Brachytheciaceae)<br />
Remyella hawaica Müll.Hal. [Hoe 1974, Ignatov &<br />
Huttunen 2002]<br />
Rhynchostegiella hawaica (Müll.Hal.) Broth.<br />
RHABDOWEISIA Bruch & Schimp. (Dicranaceae, or<br />
Rhabdoweisiaceae [Shaw & Goffinet 2000])<br />
Rhabdoweisia crenulata (Mitt.) H.Jameson [Hoe 1975,<br />
Frahm et al. 2000]<br />
Rhabdoweisia crispata (With.) Lindb. [Hoe 1974, Hoe<br />
1977, Hoe 1979]<br />
Rhabdoweisia denticulata B.S.G.<br />
TROPICAL BRYOLOGY 25 (2004)<br />
nat? K O Mo L M H<br />
ind K O Mo L M H<br />
end O<br />
ind M<br />
RHAPHIDOSTICHUM M.Fleisch. (Sematophyllaceae)<br />
Rhaphidostichum pustulatum Hoe [Hoe 1974] end O<br />
ind M H<br />
RHIZOGONIUM Brid. (Rhizogoniaceae)<br />
Rhizogonium pungens Sull. [Hoe 1974] end K O Mo L M H<br />
RHODOBRYUM (Schimp.) Limpr. (Bryaceae)<br />
Rhodobryum giganteum (Schwägr.) Paris [Hoe 1974,<br />
Crosby et al. 1983, Tan & Iwatsuki 1991]<br />
Bryum giganteum (Schwägr.) Arn.<br />
RHYNCHOSTEGIUM Bruch & Schimp.<br />
(Brachytheciaceae)<br />
Rhynchostegium celebicum (Sande Lac.) A.Jaeger [Hoe<br />
1974, Hoe 1977]<br />
Eurhynchium celebicum (Sande Lac.) E.B.Bartram<br />
Rhynchostegium gaudichaudii (Mont.) A.Jaeger [Hoe<br />
1974]<br />
Rhynchostegium selaginellifolium Müll.Hal. var.<br />
recurvirameum (Müll.Hal.) Wijk & Margad. [Hoe<br />
1974]<br />
Eurhynchium selaginellifolium var. recurvirameum<br />
ind K O M<br />
ind K O Mo M<br />
ind<br />
end K O H<br />
59
60<br />
Staples et al.<br />
<strong>Full</strong>Name<br />
(Müll.Hal.) E.B.Bartram<br />
STATUS NI K O MO L M KA H<br />
Rhynchostegium selaginellifolium Müll.Hal. var.<br />
selaginellifolium [Hoe 1974]<br />
Eurhynchium selaginellifolium (Müll.Hal.)<br />
E.B.Bartram<br />
end K O Mo M H<br />
ROSULABRYUM J.R.Spence (Bryaceae)<br />
Rosulabryum billardierei (Schwägr.) J.R.Spence<br />
[Mohamed 1979, Anderson et al. 1990, Spence 1996]<br />
Bryum andicola Hook.<br />
Bryum decaisnei Dozy & Molk.<br />
Bryum limbatomarginatum Müll.Hal.<br />
Bryum truncorum misapplied<br />
Rosulabryum capillare (Hedw.) J.R.Spence [Hoe 1974,<br />
Hoe 1977, Spence 1996]<br />
Bryum capillare Hedw.<br />
Bryum vinoviride E.B.Bartram<br />
ind O M H<br />
ind K O M H<br />
SAELANIA Lindb. (Ditrichaceae)<br />
Saelania glaucescens (Hedw.) Broth. [Hoe 1974] ind M<br />
SCHISTIDIUM Bruch & Schimp. (Grimmiaceae)<br />
Schistidium apocarpum (Hedw.) Bruch & Schimp.<br />
[Schultze-Motel 1963, Muñoz & Pando 2000]<br />
Grimmia apocarpa Hedw.<br />
Schistidium strictum (Turner) Loeske ex Mårtensson<br />
[Hoe 1974, Cao & Churchill 1995, Muñoz & Pando<br />
2000]<br />
Grimmia apocarpa Hedw. var. stricta (Turner) Hook.<br />
& Taylor<br />
Grimmia scabrifolia Broth.<br />
Grimmia stricta Turner<br />
SCHIZYMENIUM Harv. (Bryaceae, or Mniaceae [Shaw<br />
& Goffinet 2000])<br />
Schizymenium campylocarpum (Hook. & Arn.) A.J.Shaw<br />
var. campylocarpum [Zanten 1971, Shaw 1985]<br />
Mielichhoferia campylocarpa (Hook. & Arn.) Mitt.<br />
Weissia campylocarpa Hook. & Arn. in Hook.<br />
Schizymenium nealiae (E.B.Bartram) A.J.Shaw [Hoe<br />
1974, Shaw 1985]<br />
Mielichhoferia nealiae E.B.Bartram<br />
Schizymenium pulvinatum (Müll.Hal.) A.J.Shaw [Hoe<br />
1974, Shaw 1985]<br />
Mielichhoferia pulvinata Müll.Hal.<br />
ind M<br />
ind M<br />
ind H<br />
end H<br />
end M H<br />
SCOPELOPHILA (Mitt.) Lindb. (Pottiaceae)<br />
Scopelophila infericola Hoe [Hoe 1974] end H<br />
TROPICAL BRYOLOGY 25 (2004)
Revised checklist of Hawaiian mosses<br />
<strong>Full</strong>Name<br />
SEMATOPHYLLUM Mitt. (Sematophyllaceae)<br />
STATUS NI K O MO L M KA H<br />
Sematophyllum hawaiiense (Broth.) Broth. [Hoe 1974,<br />
Hoe 1979]<br />
ind K O L M H<br />
Sematophyllum subpinnatum (Brid.) E.Britton [Hoe<br />
1974, Hoe 1979, Buck 1983]<br />
Sematophyllum caespitosum misapplied<br />
ind O H<br />
SPHAGNUM L. (Sphagnaceae)<br />
Sphagnum palustre L. [Hoe 1978, Karlin & Andrus<br />
1995]<br />
Sphagnum henryense misapplied<br />
Sphagnum lonchocladum Müll.Hal.<br />
Sphagnum perichaetiale misapplied<br />
Sphagnum vitjianum misapplied<br />
Sphagnum wheeleri Müll.Hal. [Hoe 1974, Karlin &<br />
Andrus 1995, Karlin 2001]<br />
Sphagnum compactum misapplied<br />
Sphagnum rigidulum Warnst.<br />
Sphagnum vulcanicum Warnst.<br />
SPLACHNOBRYUM Müll.Hal. (Splachnobryaceae, or<br />
Pottiaceae [Shaw & Goffinet 2000])<br />
Splachnobryum obtusum (Brid.) Müll.Hal. [Hoe 1974,<br />
Arts 2001]<br />
TROPICAL BRYOLOGY 25 (2004)<br />
ind H<br />
end M<br />
nat O<br />
STREPTOPOGON Mitt. (Pottiaceae)<br />
Streptopogon erythrodontus (Taylor) Wilson [Hoe 1974] ind M<br />
SYMPHYODON Mont. (Symphyodontaceae)<br />
Symphyodon pygmaeus (Broth.) S.He & Snider [Hoe<br />
1974, Hoe 1977, He & Snider 1992]<br />
Glossadelphus abortivapicus Hoe<br />
SYNTRICHIA Brid. (Pottiaceae)<br />
Syntrichia fragilis (Taylor) Ochyra [Hoe 1974, Kürschner<br />
2000]<br />
Tortula alpina (Brid.) Arn. var. inermis (Milde) De<br />
Not.<br />
Tortula fragilis Taylor<br />
Syntrichia princeps (De Not.) Mitt. [Hoe 1974,<br />
Kürschner 2000]<br />
Tortula princeps De Not.<br />
SYRRHOPODON Schwägr. (Calymperaceae)<br />
Syrrhopodon armatus Mitt. [Hoe 1974, Orbán & Reese<br />
1986]<br />
Syrrhopodon oahuensis Broth.<br />
ind O M<br />
ind M H<br />
ind M H<br />
ind O M H<br />
Syrrhopodon hawaiicus Müll.Hal. [Hoe 1974] end K O Mo M H<br />
Syrrhopodon prolifer Schwägr. [Hoe 1974, Reese et al. end O Mo M H<br />
61
62<br />
Staples et al.<br />
<strong>Full</strong>Name STATUS NI K O MO L M KA H<br />
1986]<br />
Syrrhopodon kilaueae Müll.Hal.<br />
Syrrhopodon semiliber (Mitt.) Besch. [Hoe 1974, Reese<br />
et al. 1986]<br />
Calymperes semiliberum Mitt.<br />
Calymperopsis semiliber (Mitt.) M.Fleisch.<br />
ind O H<br />
TAXIPHYLLUM M.Fleisch. (Hypnaceae)<br />
Taxiphyllum laevifolium (Mitt.) W.R.Buck [Hoe 1974,<br />
Buck 1987b, Tixier 1988]<br />
Glossadelphus baldwinii Broth.<br />
Glossadelphus laevifolius (Mitt.) E.B.Bartram<br />
Taxiphyllum torrentium (Besch.) W.R.Buck [Hoe 1974,<br />
Buck 1987b, Tixier 1988]<br />
Glossadelphus irroratus E.B.Bartram<br />
Glossadelphus torrentium (Besch.) M.Fleisch.<br />
TAXITHELIUM Mitt. (Sematophyllaceae)<br />
Taxithelium mundulum (Sull.) E.B.Bartram [Hoe 1974]<br />
Taxithelium undulatum misapplied<br />
ind K O Mo M H<br />
ind M<br />
ind K O Mo L M H<br />
TAYLORIA Hook. (Splachnaceae)<br />
Tayloria sandwicensis (Müll.Hal.) Broth. [Hoe 1974] end M<br />
THAMNIOPSIS (Mitt.) Fleisch. (Hookeriaceae, or<br />
Callicostaceae [Buck 1987a])<br />
Thamniopsis utacamundiana (Mont.) W.R Buck [Hoe<br />
1974, Buck 1987a]<br />
Hookeria purpurea Müll.Hal.<br />
Hookeriopsis purpurea (Müll.Hal.) Broth.<br />
Hookeriopsis purpurea (Müll.Hal.) Broth. var.<br />
acuminatula (Müll.Hal.) E.B.Bartram<br />
Hookeriopsis purpurea (Müll.Hal.) Broth. var.<br />
ligulacea (Müll.Hal.) E.B.Bartram<br />
THAMNOBRYUM Nieuwl. (Neckeraceae)<br />
Thamnobryum speciosum (Broth.) Hoe [Hoe 1974]<br />
Thamnium speciosum Broth.<br />
THUIDIUM Bruch & Schimp. (Thuidiaceae)<br />
Thuidium cymbifolium (Dozy & Molk.) Dozy & Molk.<br />
[Hoe 1974, Touw 2001]<br />
Thuidium glabratum Broth.<br />
?Thuidium havaiense Reichardt<br />
Thuidium oahuense Broth.<br />
Thuidium plicatum Mitt.<br />
Thuidium plicatum Mitt. var. brevifolium<br />
E.B.Bartram<br />
ind K O Mo M H<br />
end M H<br />
ind K O Mo L M H<br />
TROPICAL BRYOLOGY 25 (2004)
Revised checklist of Hawaiian mosses<br />
<strong>Full</strong>Name<br />
TIMMIA Hedw. (Timmiaceae)<br />
STATUS NI K O MO L M KA H<br />
Timmia megapolitana Hedw. ssp. bavarica (Hessl.)<br />
Brassard [Brassard 1984]<br />
Timmia bavarica Hessl.<br />
ind H<br />
TORTELLA (Lindb.) Limpr. (Pottiaceae)<br />
Tortella fragilis (Hook. & Wilson) Limpr. var.<br />
tortelloides (S.W.Greene) R.H.Zander & Hoe [Zander<br />
& Hoe 1979]<br />
Sarconeurum tortelloides S.W.Greene<br />
Tortella tortelloides (S.W.Greene) H.Rob.<br />
ind M<br />
Tortella humilis (Hedw.) Jenn. [Hoe 1974]<br />
Tortella caespitosa (Schwägr.) Limpr.<br />
ind K O L M H<br />
Tortella tortuosa (Hedw.) Limpr. [Hoe 1979] ind L<br />
TORTULA Hedw. (Pottiaceae)<br />
Tortula atrovirens (Sm.) Lindb. [Hoe 1974, Hoe 1979,<br />
Corley et al. 1981, Zander 1993]<br />
Desmatodon convolutus (Brid.) Grout<br />
TRACHYLOMA Brid. (Pterobryaceae, or<br />
Trachylomataceae [Shaw & Goffinet 2000])<br />
Trachyloma indicum Mitt. [Hoe 1974]<br />
Trachyloma tahitense Besch.<br />
TRACHYPODOPSIS M.Fleisch. (Trachypodaceae, or<br />
Meteoriaceae [Shaw & Goffinet 2000])<br />
Trachypodopsis auriculata (Mitt.) M.Fleisch. [Hoe 1974]<br />
Trachypodopsis ornans (Reichardt) M.Fleisch.<br />
Trachypus ornans (Reichardt) Müll.Hal. fma.<br />
simplex Broth.<br />
Trachypodopsis serrulata (P.Beauv.) M.Fleisch. var.<br />
crispatula (Hook.) Zanten [Hoe 1979]<br />
TRACHYPUS Reinw. & Hornsch. (Trachypodaceae, or<br />
Meteoriaceae [Shaw & Goffinet 2000])<br />
Trachypus bicolor Reinw. & Hornsch. [Hoe 1974, Hoe<br />
1979]<br />
Trachypus humilis Lindb. var. flagellifer (Broth. ex<br />
Herzog) H.A.Mill. [Zanten 1959, Miller 1967, Hoe<br />
1974]<br />
Trachypus humilis Lindb. var. tenerrimus (Herzog)<br />
Zanten<br />
Trachypus mauiensis Broth.<br />
Trachypus humilis Lindb. var. humilis [Hoe 1974, Hoe<br />
1977]<br />
TREMATODON Michx. (Dicranaceae, or Bruchiaceae<br />
[Shaw & Goffinet 2000])<br />
TROPICAL BRYOLOGY 25 (2004)<br />
ind M H<br />
ind M<br />
ind K O Mo M H<br />
ind H<br />
ind K M H<br />
ind K O M<br />
ind K M<br />
63
64<br />
Staples et al.<br />
<strong>Full</strong>Name STATUS NI K O MO L M KA H<br />
Trematodon latinervis Müll.Hal. [Hoe 1974]<br />
Trematodon longicollis misapplied<br />
ind K O Mo M H<br />
TRICHOSTOMUM Bruch (Pottiaceae)<br />
Trichostomum brachydontium Bruch [Sollman 1984]<br />
Trichostomum oblongifolium E.B.Bartram<br />
Trichostomum crispulum Bruch [Hoe 1974, Hoe 1975,<br />
Norris & Koponen 1989]<br />
Trichostomum bartramii H.A.Mill.<br />
Trichostomum mauiense Broth.<br />
ind K O Mo M H<br />
ind K O Mo L M H<br />
ULOTA D.Mohr (Orthotrichaceae)<br />
Ulota cervina Hoe & H.A.Crum [Hoe 1974] end M<br />
VESICULARIA (Müll.Hal.) Müll.Hal. (Hypnaceae)<br />
Vesicularia inflectens (Brid.) Müll.Hal. [Hoe 1979] ind M<br />
Vesicularia perviridis (Ångstr.) Müll.Hal. [Hoe 1974]<br />
Vesicularia graminicolor Müll.Hal.<br />
end K O Mo L M H<br />
WEISSIA Hedw. (Pottiaceae)<br />
Weissia controversa Hedw. [Hoe 1974, Wichman & St.<br />
John 1990]<br />
Weisia viridula Hedw.<br />
ind Ni K O H<br />
Weissia ovalis (R.S.Williams) E.B.Bartram [Hoe 1974] end K O Mo L M H<br />
ZYGODON Hook. & Taylor (Orthotrichaceae)<br />
Zygodon reinwardtii (Hornsch.) A.Braun [Hoe 1974] ind M<br />
Zygodon rubrigemmius (Hoe & H.A.Crum) R.H.Zander<br />
& Vitt [Hoe 1974, Zander & Vitt 1979]<br />
Anoectangium rubrigemmium Hoe & H.A.Crum<br />
end M<br />
Zygodon tetragonostomus A.Braun ex B.S.G. [Hoe 1974] ind K O M H<br />
TROPICAL BRYOLOGY 25 (2004)
Revised checklist of Hawaiian mosses<br />
CROSS-REFERENCED LIST OF SYNONYMS AND MISAPPLIED NAMES<br />
Acroporium fuscoflavum var. baldwinii See Acroporium fuscoflavum<br />
?Acroporium sigmatodontium misapplied See Acroporium fuscoflavum<br />
Aerobryopsis longissima See Aerobryopsis wallichii<br />
Aerobryopsis longissima var. dozyana See Aerobryopsis wallichii<br />
Aerobryopsis scariosa See Aerobryopsis subdivergens ssp. scariosa<br />
Amphidium cyathicarpum See Amphidium tortuosum<br />
Andreaea acuminata See Andreaea acutifolia ssp. acuminata<br />
Andreaea perpapillosa See Andreaea rupestris<br />
Anoectangium euchloron See Anoectangium aestivum<br />
Anoectangium haleakalae var. haleakalae See Anoectangium aestivum<br />
Anoectangium haleakalae var. laxulum See Anoectangium aestivum<br />
Anoectangium harttiae See Anoectangium aestivum<br />
Anoectangium rubrigemmium See Zygodon rubrigemmius<br />
Aptychella robusta See Clastobryopsis robusta<br />
Barbella trichophora See Barbellopsis trichophora<br />
Barbula cruegeri See Barbula indica<br />
Barbula unguiculata misapplied See Barbula indica<br />
Barbula vinealis var. cylindrica See Didymodon insulanus<br />
Barbula vinealis var. flaccida See Didymodon insulanus<br />
Brachythecium oxyrrhynchum See Brachythecium plumosum<br />
Breutelia kilaueae See Breutelia affinis<br />
Brotherella opaeodon See Pylaisiadelpha opaeodon<br />
Bryohaplocladium microphyllum See Haplocladium microphyllum<br />
Bryum andicola Hook. See Rosulabryum billardierei<br />
Bryum apiculatum See Bryum mildeanum<br />
Bryum argenteum var. lanatum See Bryum argenteum<br />
Bryum capillare See Rosulabryum capillare<br />
Bryum ceramiocarpum See Bryum conicum<br />
Bryum crassicostatum See Bryum hawaiicum<br />
Bryum decaisnei See Rosulabryum billardierei<br />
Bryum erythrocarpum See Bryum atrovirens<br />
Bryum giganteum See Rhodobryum giganteum<br />
Bryum leucophyllum See Bryum argenteum<br />
Bryum limbatomarginatum See Rosulabryum billardierei<br />
Bryum mauiensis See Bryum mildeanum<br />
Bryum megalostegium See Bryum mildeanum<br />
Bryum nitens See Bryum mildeanum<br />
Bryum truncorum misapplied See Rosulabryum billardierei<br />
Bryum vinoviride See Rosulabryum capillare<br />
Calymperes hawaiiense See Calymperes fasciculatum<br />
Calymperes semiliberum See Syrrhopodon semiliber<br />
Calymperopsis semiliber See Syrrhopodon semiliber<br />
Campylopodium euphorocladum See Campylopodium medium<br />
Campylopus aureus See Campylopus schmidii ssp. schmidii<br />
Campylopus boswellii See Campylopus fragilis ssp. zollingerianus<br />
Campylopus boswellii var. capitulatus See Campylopus fragilis ssp. zollingerianus<br />
Campylopus densifolius var. densifolius See Campylopus hawaiicus var. densifolius<br />
Campylopus densifolius var. falcatus See Campylopus hawaiicus var. hawaiicoflexuosus<br />
TROPICAL BRYOLOGY 25 (2004)<br />
65
66<br />
Staples et al.<br />
Campylopus densifolius var. hawaiicoflexuosus See Campylopus hawaiicus var. hawaiicoflexuosus<br />
Campylopus densifolius var. purpureoflavescens See Campylopus hawaiicus var. hawaiicus<br />
Campylopus densiretis See Campylopus hawaiicus var. densifolius<br />
?Campylopus didrichsenii See Campylopus umbellatus var. umbellatus<br />
Campylopus exasperatus See Campylopus praemorsus<br />
Campylopus fumarioli See Campylopus laxi<strong>text</strong>us<br />
Campylopus geniculatus See Campylopus umbellatus var. pterotoneuron<br />
Campylopus hawaiicoflexuosus See Campylopus hawaiicus var. hawaiicoflexuosus<br />
Campylopus introflexus misapplied See Campylopus schmidii ssp. schmidii<br />
Campylopus introflexus var. polytrichoides misapplied See Campylopus schmidii ssp. schmidii<br />
Campylopus microcephalus See Campylopus praemorsus<br />
Campylopus perangustifolius See Campylopus hawaiicus var. hawaiicoflexuosus<br />
?Campylopus pertristis See Campylopus umbellatus var. umbellatus<br />
Campylopus polytrichoides misapplied See Campylopus schmidii ssp. schmidii<br />
Campylopus purpureoflavescens, in part See Campylopus hawaiicus var. hawaiicoflexuosus, C.<br />
hawaiicus var. hawaiicus<br />
Campylopus richardii misapplied See Campylopus umbellatus var. umbellatus<br />
Campylopus sandwicensis See Campylopus umbellatus var. umbellatus<br />
Campylopus skottsbergii See Campylopus schmidii ssp. hemitrichius<br />
Campylopus tubulosus See Campylopus schmidii ssp. hemitrichius<br />
Campylopus wawreanus See Campylopus hawaiicus var. densifolius<br />
Ceratodon purpureus fma. xanthopus See Ceratodon purpureus ssp. purpureus<br />
Cirriphyllum oxyrrhynchum See Brachythecium plumosum<br />
Claopodium amblystegioides See Claopodium prionophyllum<br />
Claopodium hawaiiense See Claopodium whippleanum<br />
Claopodium nervosum See Claopodium prionophyllum<br />
Ctenidium decurrens See Ectropothecium decurrens<br />
Ctenidium pulcherrimum See Ectropothecium decurrens<br />
Ctenidium submalacobolum See Ectropothecium decurrens<br />
Cupressina decurrens See Ectropothecium decurrens<br />
Desmatodon convolutus See Tortula atrovirens<br />
Dicranella hillebrandii See Microdus hillebrandii<br />
Dicranella hochreuteri misapplied See Dicranella hochreutineri<br />
Dicranodontium falcatum var. falcatum See Dicranodontium porodictyon<br />
Dicranodontium hawaiicum See Dicranodontium porodictyon<br />
Dicranoloma wheeleri See Campylopus wheeleri<br />
Dicranum didrichsenii See Campylopus umbellatus var. umbellatus<br />
Dicranum hawaiicoflexuosum See Campylopus hawaiicus var. hawaiicoflexuosus<br />
Dicranum hawaiicum See Campylopus hawaiicus var. hawaiicus<br />
Dicranum microcephalum See Campylopus praemorsus<br />
Dicranum pertriste See Campylopus umbellatus var. umbellatus<br />
Dicranum praemorsum See Campylopus praemorsus<br />
Dicranum skottsbergii See Campylopus schmidii ssp. hemitrichius<br />
Dicranum wheeleri See Campylopus wheeleri<br />
Didymodon recurvirostris See Bryoerythrophyllum recurvirostre<br />
Didymodon vinealis var. flaccidus See Didymodon insulanus<br />
Ectropothecium rockii See Ectropothecium decurrens<br />
Encalypta rhaptocarpa misapplied See Encalypta rhabdocarpa<br />
Encalypta scabrata See Encalypta ciliata<br />
Entodon hillebrandii See Entodon plicatus<br />
TROPICAL BRYOLOGY 25 (2004)
Revised checklist of Hawaiian mosses<br />
Entodon solanderi See Entodon plicatus<br />
Eurhynchium celebicum See Rhynchostegium celebicum<br />
Eurhynchium muelleri misapplied See Platyhypnidium muelleri<br />
Eurhynchium selaginellifolium See Rhynchostegium selaginellifolium var. selaginellifolium<br />
Eurhynchium selaginellifolium var. recurvirameum See Rhynchostegium selaginellifolium var.<br />
recurvirameum<br />
Eurhynchium vagans See Oxyrrhynchium vagans<br />
Fissidens baldwinii See Fissidens elegans<br />
Fissidens bishopii See Fissidens elegans<br />
Fissidens crumii See Fissidens lindbergii<br />
Fissidens hawaiicus See Fissidens bryoides<br />
Fissidens insularis See Fissidens bryoides<br />
Fissidens intermedius See Fissidens submarginatus<br />
Fissidens mauiensis See Fissidens pacificus<br />
Fissidens oahuensis See Fissidens bryoides<br />
Floribundaria baldwinii See Floribundaria floribunda<br />
Funaria subintegra See Entosthodon subintegrus<br />
Garovaglia haleakalae See Garovaglia powellii var. haleakalae<br />
Glossadelphus abortivapicus See Symphyodon pygmaeus<br />
Glossadelphus baldwinii See Taxiphyllum laevifolium<br />
Glossadelphus chrysobasilaris See Glossadelphus ogatae<br />
Glossadelphus irroratus See Taxiphyllum torrentium<br />
Glossadelphus laevifolius See Taxiphyllum laevifolium<br />
Glossadelphus torrentium See Taxiphyllum torrentium<br />
Glossadelphus zollingeri var. filicaulis See Ectropothecium zollingeri var. filicaule<br />
Grimmia apocarpa See Schistidium apocarpum<br />
Grimmia apocarpa var. pulvinata See Grimmia anodon<br />
Grimmia apocarpa var. stricta See Schistidium strictum<br />
Grimmia haleakalae See Amphidium tortuosum<br />
Grimmia haliacalae See Amphidium tortuosum<br />
Grimmia pilfera See Grimmia longirostris<br />
Grimmia scabrifolia See Schistidium strictum<br />
Grimmia stricta See Schistidium strictum<br />
Gymnostoma pulvinata See Grimmia anodon<br />
Hageniella pacifica See Hageniella micans<br />
Holomitrium ferriei misapplied See Holomitrium squarrifolium<br />
Homalia densa See Homaliodendron flabellatum<br />
Homalia praelonga See Homaliodendron flabellatum<br />
Hookeria purpurea See Thamniopsis utacamundiana<br />
Hookeriopsis purpurea See Thamniopsis utacamundiana<br />
Hookeriopsis purpurea var. acuminatula See Thamniopsis utacamundiana<br />
Hookeriopsis purpurea var. ligulacea See Thamniopsis utacamundiana<br />
Hymenostylium firmum See Hymenostylium recurvirostrum var. recurvirostrum<br />
Hyophila dozy-molkenboeri See Hyophila involuta<br />
Hypnum decurrens See Ectropothecium decurrens<br />
Hypnum draytoni(i) See Buckiella draytonii<br />
Hypnum eudorae See Buckiella draytonii<br />
Hypnum opaeodon See Pylaisiadelpha opaeodon<br />
Hypnum wilkesianum See Palamocladium wilkesianum<br />
Isopterygium elegans See Pseudotaxiphyllum elegans<br />
TROPICAL BRYOLOGY 25 (2004)<br />
67
68<br />
Leptodontium brevicaule See Leptodontium flexifolium<br />
Leucobryum hawaiense See Leucobryum seemannii var. seemannii<br />
Leucobryum hawaiense var. fumarioli See Leucobryum seemannii var. fumarioli<br />
Leucobryum hawaiense var. solfatarae See Leucobryum seemannii var. solfatarae<br />
Leucobryum solfatarae See Leucobryum seemannii var. solfatarae<br />
Leucobryum solfatarae var. fumarioli See Leucobryum seemannii var. fumarioli<br />
Leucobryum solfatarae var. hawaiense See Leucobryum seemannii var. seemannii<br />
Macrocoma intricata See Macrocoma tenue ssp. sullivantii<br />
Macromitrium cumingii See Macromitirum incurvifolium<br />
Macromitrium intricatum See Macrocoma tenue ssp. sullivantii<br />
Macromitrium owahiense See Macromitrium microstomum<br />
Macromitrium reinwardtii See Macromitrium microstomum<br />
Merceyopsis crassinervis See Hymenostylium recurvirostrum var. cylindricum<br />
Meteorium pallidovirens See Palamocladium wilkesianum<br />
Meteorium sciuroides See Palamocladium wilkesianum<br />
Microdus hawaiicus See Dicranella hawaiica var. hawaiica<br />
Mielichhoferia campylocarpa See Schizymenium campylocarpum var. campylocarpum<br />
Mielichhoferia nealiae See Schizymenium nealiae<br />
Mielichhoferia pulvinata See Schizymenium pulvinatum<br />
Mniadelphus freycineti var. crasseturgescens See Distichophyllum freycinetii var.<br />
crasseturgescens<br />
Mnium rostratum See Plagiomnium rostratum<br />
Mnium serratum See Mnium marginatum<br />
Molendoa crassinervis See Hymenostylium recurvirostrum var. cylindricum<br />
Orthotrichum berggrenii See Orthotrichum alpestre var. berggrenii<br />
Orthotrichum hillebrandii See Orthotrichum rupestre<br />
Orthotrichum verrucatum See Orthotrichum lyellii<br />
Palamocladium aptychoides See Palamocladium wilkesianum<br />
Palamocladium sciurellum See Palamocladium wilkesianum<br />
Palamocladium wilkesianum var. altisetum See Palamocladium wilkesianum<br />
Palamocladium wilkesianum var. sciuroides See Palamocladium wilkesianum<br />
Philonotis laxissima See Philonotis hastata<br />
Plagiomnium prorepens See Plagiomnium rostratum<br />
Plagiomnium rhynchosporum See Plagiomnium rostratum<br />
Plagiopus longisetus See Plagiopus oederiana<br />
Plagiopus oederi See Plagiopus oederiana<br />
Plagiothecium draytonii See Buckiella draytonii<br />
Pleuropus aptychoides See Palamocladium wilkesianum<br />
Pleuropus pallidovirens See Palamocladium wilkesianum<br />
Pleuropus sciurellus See Palamocladium wilkesianum<br />
Pleuropus wilkesianus See Palamocladium wilkesianum<br />
Pleuropus wilkesianus var. sciuroides See Palamocladium wilkesianum<br />
Pogonatum baldwinii See Pogonatum tahitense<br />
Pohlia gracilescens See Pohlia flexuosa<br />
Pohlia gracillima See Pohlia flexuosa<br />
Pohlia leucostoma See Pohlia flexuosa<br />
Pohlia leucostomoides See Pohlia flexuosa<br />
Pseudosymblepharis mauiensis See Pseudosymblepharis angustata<br />
Ptychomitrium mauiense See Ptychomitrium lepidomitrium<br />
Ptychomnion aciculare misapplied See Aerobryopsis subdivergens ssp. scariosa<br />
Staples et al.<br />
TROPICAL BRYOLOGY 25 (2004)
Revised checklist of Hawaiian mosses<br />
Racomitrium lanuginosum var. sandvicense See Racomitrium lanuginosum var. pruinosum<br />
Rhabdoweisia denticulata See Rhabdoweisia crispata<br />
Rhacomitrium cuspidigerum misapplied See Racopilum cuspidigerum<br />
Rhizogonium spiniforme See Pyrrhobryum spiniforme<br />
Rhynchostegiella hawaica See Remyella hawaica<br />
Rhynchostegium vagans See Oxyrrhynchium vagans<br />
Sarconeurum tortelloides See Tortella fragilis var. tortelloides<br />
Sciaromium tricostatum See Limbella tricostata<br />
Sematophyllum caespitosum See Sematophyllum subpinnatum<br />
Sphagnum compactum misapplied See Sphagnum wheeleri<br />
Sphagnum henryense misapplied See Sphagnum palustre<br />
Sphagnum lonchocladum See Sphagnum palustre<br />
Sphagnum perichaetiale misapplied See Sphagnum palustre<br />
Sphagnum rigidulum See Sphagnum wheeleri<br />
Sphagnum vitjianum misapplied See Sphagnum palustre<br />
Sphagnum vulcanicum See Sphagnum wheeleri<br />
Stereodon decurrens See Ectropothecium decurrens<br />
Stereophyllum oahuense See Entodontopsis anceps<br />
Syrrhopodon kilaueae See Syrrhopodon prolifer<br />
Syrrhopodon oahuensis See Syrrhopodon armatus<br />
Taxiphyllum gracile See Ectropothecium decurrens<br />
Taxithelium undulatum misapplied See Taxithelium mundulum<br />
Thamnium speciosum See Thamnobryum speciosum<br />
Thuidium crenulatum See Pelekium versicolor<br />
Thuidium glabratum See Thuidium cymbifolium<br />
?Thuidium havaiense See Thuidium cymbifolium<br />
Thuidium nanophyllum See Pelekium versicolor<br />
Thuidium oahuense See Thuidium cymbifolium<br />
Thuidium plicatum See Thuidium cymbifolium<br />
Thuidium plicatum var. brevifolium See Thuidium cymbifolium<br />
Thysanomitrion hawaiicum See Campylopus umbellatus var. umbellatus<br />
Thysanomitrion umbellatum See Campylopus umbellatus var. umbellatus<br />
Timmia bavarica See Timmia megapolitana ssp. bavarica<br />
Tortella caespitosa See Tortella humilis<br />
Tortella tortelloides See Tortella fragilis var. tortelloides<br />
Tortula alpina var. inermis See Syntrichia fragilis<br />
Tortula fragilis See Syntrichia fragilis<br />
Tortula princeps See Syntrichia princeps<br />
Tortula rhizophylla See Chenia leptophylla<br />
Tortula vectensis See Chenia leptophylla<br />
Trachyloma tahitense See Trachyloma indicum<br />
Trachypodopsis ornans See Trachypodopsis auriculata<br />
Trachypus humilis var. tenerrimus See Trachypus humilis var. flagellifer<br />
Trachypus mauiensis See Trachypus humilis var. flagellifer<br />
Trachypus ornans fma. simplex See Trachypodopsis auriculata<br />
Trematodon longicollis misapplied See Trematodon latinervis<br />
Trichosteleum hamatum See Radulina hamata<br />
Trichostomum bartramii See Trichostomum crispulum<br />
Trichostomum mauiense See Trichostomum crispulum<br />
Trichostomum oblongifolium See Trichostomum brachydontium<br />
TROPICAL BRYOLOGY 25 (2004)<br />
69
70<br />
Trichostomum tenuirostre See Oxystegus tenuirostris var. gemmiparus<br />
Vesicularia graminicolor See Vesicularia perviridis<br />
Webera baldwinii See Pohlia baldwinii<br />
Webera cruda See Pohlia cruda<br />
Webera gracilescens See Pohlia flexuosa<br />
Webera leucostomoides See Pohlia flexuosa<br />
Weisia viridula See Weissia controversa<br />
Weissia campylocarpa See Schizymenium campylocarpum var. campylocarpum<br />
Staples et al.<br />
TROPICAL BRYOLOGY 25 (2004)