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Revised checklist of Hawaiian mosses 

Tropical Bryology 25: 35-69, 2004 

A revised checklist of Hawaiian mosses 1 

G. W. Staples A , C. T. Imada A , W. J. Hoe, B and C. W. Smith C 

A – Hawaii Biological Survey, Bishop Museum, 1525 Bernice St., Honolulu, Hawaii 96817 

U.S.A. 

B – deceased; 

C – Department of Botany, The Natural History Museum, Cromwell Road, London, SW7 

5BD, U.K. 

Abstract. A revised and updated literature-based checklist of Hawaiian mosses is presented. 

Geographic coverage includes the eight main Hawaiian Islands; the Northwestern Hawaiian Islands 

are excluded. The checklist is alphabetically ordered by scientific names; the family is noted for each 

genus. Synonyms and misapplied names are cross-referenced to the accepted names. A bibliography 

of supporting references is included. 

Introduction 

The Hawaii Biological Survey (HBS) was 

established as a program of Bishop Museum by 

the Hawaii State Legislature in 1992, specifically 

to create and maintain accurate inventories of all 

species present in the Hawaiian Islands, and to 

document their presence with voucher specimen 

collections and bibliographies of relevant 

scientific literature. In the ensuing eleven years, 

comprehensive taxonomic inventories have been 

compiled for many groups of organisms, both 

native and alien, that comprise the Hawaiian biota. 

Vascular plants and marine algae have been the 

primary focus of botanical inventory efforts; 

research on non-marine algae, terrestrial 

cryptogamic plants, fungi, and lichens has lagged 

behind. 

This checklist is intended to fill one of 

the gaps in our knowledge of Hawaiian terrestrial 

1 Contribution No. 2004–001 to the Hawaii 

Biological Survey. 

TROPICAL BRYOLOGY 25 (2004) 

35 

cryptogamic plants—the mosses. Also in 

preparation by HBS staff members are checklists 

and bibliographies for the Hawaiian anthocerotes 

(hornworts) and hepatics (liverworts). A nonmarine 

algal checklist for the Hawaiian Islands is 

also in an advanced stage of preparation (A. 

Sherwood, pers. comm.). Eventually a checklist 

of Hawaiian lichens is planned. These ongoing 

inventories will help the Hawaii Biological Survey 

to fulfill its mission by systematically compiling 

literature-based inventories for each of these 

groups of organisms. 

Genesis of the current checklist 

This checklist of Hawaiian mosses begins 

where its predecessor—the checklist compiled 

by Hoe (1974)—left off, and updates that 

document by revising the nomenclature, adding 

new taxa and distributional records, and 

providing a bibliography of literature that 

supports the names in current use. The checklist

36 

and bibliography were compiled by a 

collaborative effort between HBS staff members 

(Staples, Imada) working in partnership with two 

bryological specialists (Hoe, Smith), who had 

assembled a cache of literature citations and new 

distribution records pertinent to Hawaiian 

mosses. Funding from the USDA–PLANTS 

database project in 1998 provided the impetus 

for Staples and Imada to undertake conversion 

of Hoe’s 1974 moss checklist to an electronic 

format and then begin to update the taxonomy 

and distribution information it contained. Hoe 

and Smith generously shared their preliminary 

cache of information in order to facilitate the 

update process. Sadly, in May 2000, Hoe died 

suddenly, a great loss to the botanical community 

in Hawai‘i and to the world bryological 

community. After some reorganization the project 

resumed and was completed by Staples and 

Imada. In the final version of the checklist, 

taxonomic decisions were made by Staples, with 

review and critical input from Smith and two 

independent reviewers. 

Coverage 

It must be emphasized that this is a 

literature-based taxonomic checklist, not a 

specimen-based inventory. It is compiled from 

literature published from the mid-1970s to the 

present (2003). The starting point is the checklist 

published by Hoe (1974), who summarized the 

literature and reconciled the scientific names for 

Hawaiian mosses in use up to that time. The 

present checklist attempts to account for all 

scientific names in use for Hawaiian mosses in 

the latter decades of the twentieth century; it is 

not a comprehensive nomenclator for all names 

that have ever been published for, or applied to, 

Hawaiian mosses. No nomenclatural novelties or 

taxonomic changes are initiated in this checklist. 

Geographically, coverage focuses on the 

eight main Hawaiian Islands. There are, as yet, 

no published reports for bryophytes from the 

uninhabited Northwestern Hawaiian Islands, 

although a few bryophyte collections were made 

there in the last decades of the twentieth century. 

It had been Bill Hoe’s intention to identify these 

voucher specimens and write them up for 

publication but the specimens remain unstudied 

as this checklist goes to print. They await 

Staples et al. 

inclusion in a future iteration of the Hawaiian 

moss checklist. 

In the last quarter of the twentieth 

century there was a renaissance of bryological 

research worldwide and as a result many 

revisions, floristic accounts, and several 

monographs have been published that impact 

Hawaiian moss nomenclature. These regional and 

global studies resulted in significant taxonomic 

changes, as well as reporting discovery of several 

taxa new to the Islands, either rare native mosses 

or newly naturalized alien species. This checklist 

aims to summarize all this information in a concise 

format, as follows. 

Format 

Accepted scientific name 

The main body of the checklist is 

alphabetically organized by genus, species, and 

infraspecific taxon. Accepted names appear in 

bold italic type. Authors are included for the 

genera, species, and infraspecific epithets. 

Authors of moss genera have been standardized 

against the classification proposed in Bryophyte 

Biology (Shaw & Goffinet 2000); authors for five 

genera not found in that classification have been 

checked on the MOST website (cited below). All 

author names are abbreviated according to 

Authors of Plant Names style (Brummitt & Powell 

1992). References that explain the taxonomic 

concept or provide geographic distribution data 

are cited after the accepted taxon name. 

The first mention of a genus includes 

the family assignment on the same line. Most 

genera are still placed in the same families 

recognized by Hoe (1974), but a few new family 

placements have been made since then. In such 

cases, we provide first the family assignment 

used by Hoe (1974), and mention alternative 

family placements afterward, with a reference to 

the source. 

Synonyms and misapplied names (in 

italics) follow the accepted name in paragraph 

style. These are limited primarily to names in use 

for Hawaiian mosses in the latter part of the 

twentieth century, although a few names from 

other parts of the world that enter into the 

nomenclature for particular taxa have been 

included to enable tracking names through the 

literature. 

TROPICAL BRYOLOGY 25 (2004)


Status 

Following the taxonomy is a column 

indicating status of the taxon in the Hawaiian 

flora: end indicates an endemic, ind indicates 

indigenous, and nat signifies naturalized. A ? 

following any of these signifies uncertainty about 

the status of the taxon in the Hawaiian flora. While 

some research has been devoted to the mode of 

introduction of pteridophytes and angiosperms 

to Hawai‘i, resulting in a list of species that are 

likely to have been introduced by Polynesian 

voyagers that settled the archipelago and another 

(much larger) list of plants that arrived following 

European contact, little attention has been 

devoted to bryophytes in this regard. The authors 

have used their best judgment in cases where 

the status is not clear and appended a ? to 

whatever status we assigned, to draw attention 

to the species and invite investigation from 

specialists. 

One case worth noting is Racopilum 

cuspidigerum, which is widespread in the 

Southern Hemisphere but is found only in the 

Hawaiian Islands in the Northern Hemisphere. It 

is a prime candidate as a Polynesian introduction 

to Hawai‘i, and indeed to many of the other 

Polynesian islands in the South Pacific (S. Olson, 

pers. comm.). 

Island distribution 

Following the status category is a 

summary of geographic distribution on the eight 

main Hawaiian Islands. The island name 

abbreviations are explained at the foot of each 

page. An island abbreviation appears in the row 

to the right of the taxon name to signify that the 

taxon is reported in the literature from that island. 

Absence of the island name abbreviation 

indicates the moss is not (yet) known from that 

island. Given the inadequate state of moss 

collecting in the Hawaiian Islands, many new 

island distribution records are sure to be found 

in the future. 

Synonym list and Literature Consulted 

Following the main body of the checklist, 

an alphabetical list of all synonyms and 

misapplied names is provided. This list crossreferences 

every synonym or misapplied name 

to the main entry found under the scientific name 


37 

accepted in this checklist. The checklist 

concludes with a bibliography of the references 

cited in the checklist; more than 130 bryological 

titles were consulted in the course of the project, 

but only those providing taxonomic or 

distribution data pertinent to this checklist are 

included here. 

Anomalies and incomplete records 

A few moss names without any island 

distribution are included, based on (mostly) older 

literature that reports the species from the 

Hawaiian (or sometimes Sandwich) Islands 

without stating on which island the species was 

found. Two examples in the list are Dicranoweisia 

cirrata and Plagiothecium denticulatum. Their 

presence in the Hawaiian Islands needs to be 

confirmed and documented with contemporary 

collections. 

Another anomaly is Mniobryoides 

degeneri, an invalid name published without a 

description or indication of a type specimen by 

H. Hörmann. It has been pointed out (S. Olson, 

pers. comm.) that this is probably a 

misidentification for some well-known species. 

However, M. degeneriae has never been assessed 

by a competent bryologist to ascertain its true 

identity. We leave the name in the checklist to 

point out that more study is required. 

A note about family circumscriptions 

The past 25 years has brought an 

explosion in research on bryophyte systematics 

and a great deal of new information has accrued, 

which has led to many changes in moss 

classification. Family classification in particular 

has become increasingly problematic as new data 

provide insights into relationships between 

genera. Several new families have been erected 

and other genera have been reassigned. We have 

indicated the family where Hoe (1974) assigned 

each genus, followed by alternative family 

assignments where these were discovered. Two 

principle reference works were consulted for 

family assignments: first, Bryophyte Biology 

(Shaw & Goffinet 2000), and in a few cases the 

bryological nomenclator included in the 

W 3 MOST database of mosses (http:// 

mobot.mobot.org/W3T/Search/most.html) on the 

TROPICOS website.

38 

The way forward 

It is hoped that by bringing together the literature 

and providing an updated checklist of names in 

current use, one or more competent bryologists 

might be inspired to undertake preparation of a 

moss flora for the Hawaiian Islands. The last 

floristic account of the Hawaiian mosses is that 

by Bartram (1933, 1939). Bill Hoe intended to write 

a modern flora of the Hawaiian mosses, but his 

untimely death prevented that. It is our hope that 

professional bryologists will take up the challenge 

to complete this much needed reference work. 

Those who would study Hawaiian 

mosses will find type material and historic 

specimens widely distributed in herbaria around 

the world. In the last decades of the twentieth 

century, D. Vitt and his students as well as a team 

of cryptogamic botanists from the National 

Museum of Tokyo collected mosses throughout 

the main Hawaiian Islands. But by far the most 

comprehensive late twentieth century collections 

of Hawaiian mosses are those made by Hoe 

between 1965 and 1995. Today the first set of 

Hoe’s collections resides in two herbaria: the 

Herbarium Pacificum, Bishop Museum, Honolulu 

(BISH) and the Claude Philipps Herbarium, 

Delaware State University, Dover (DOV), with 

duplicates widely distributed in other herbaria. 

Hoe donated his personal herbarium to BISH in 

1983. His residual duplicates and some new 

material acquired after 1983 were deposited at 

DOV after the sale of Hoe’s bryological library 

and herbarium. The Hoe collections in BISH and 

DOV provide an excellent resource for bryological 

researchers and afford a solid foundation for any 

floristic account of Hawaiian mosses. 

Hawaiian Moss Flora in Summary 

Endemic Indigenous Alien Total 

Families0 41 1 42 

Genera 2 126 7 135 

Species 75 166 14 255 

Total named taxa 90 169 14 273 


Acknowledgments 

Funding for initial data compilation was 

made available by the USDA-PLANTS program; 

we thank Scott Petersen and Mark Skinner for 

their support and assistance. Our Museum 

colleagues Katie Anderson and Karin Kami 

assisted with data entry and bibliographic 

compilation. Bishop Museum librarian Patrice 

Belcher obtained numerous critical references 

through interlibrary loan services. Reviews of the 

draft checklist by Storrs Olson, U.S. National 

Museum, and James Shevock, U.S. Geological 

Survey, brought to light several bryological 

articles overlooked during the compilation as well 

as many suggestions for its improvement. Bill 

Buck, New York Botanical Garden, added further 

literature citations and name changes when he 

edited the manuscript for Tropical Bryology. The 

authors gratefully acknowledge all assistance 

rendered; any errors of fact or interpretation are 

the sole responsibility of the authors. 

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stenocarpus (Thér.) Zander includes 

Trichostomum spirale Grout. Miscellanea 

Bryologica et Lichenologica 9(4): 72–74. 

Zander, R.H. 1993. Genera of Pottiaceae: mosses 

of harsh environments. Bulletin of the Buffalo 

Society of Natural Sciences 32: i–vi, 1–378. 

Zander, R.H., and P.M. Eckel. 1982. 

Hymenostylium recurvirostrum var. insigne 

and Barbula amplexifolia in British Columbia, 

Canada. Canadian Journal of Botany 60: 1596– 

1600. 

Zander, R.H., and W.J. Hoe. 1979. Geographic 

disjunction and heterophylly in Tortella 

fragilis var. tortelloides (= Sarconeurum 

tortelloides). Bryologist 82(1): 84–87. 

Zander, R.H., and D.H. Vitt. 1979. Gametophytic 

distinctions of Zygodon (Orthotrichaceae) and 

Anoectangium and Leptodontium (Pottiaceae) 

and the status of Anoectangium 

rubrigemmium of Hawaii. Canadian Journal of 

Botany 57(3): 292–296. 

Zanten, B.O. van. 1959. Trachypodaceae—a 

critical revision. Blumea 9(2): 477–575. 

Zanten, B.O. van. 1971. Musci. Pp. 173–227. In 

Zinderen-Bakker, E.M. van, J.M. 

Winterbottom, and R.A. Dyer (eds.), Marion 

and Prince Edward Islands. A. A. Balkema, 

Cape Town. 

Zilstra, G. 1990. Report of the Committee for 

Bryophyta: I. Taxon 39(2): 289–292. 


Island abbreviations: Ni Ni‘ihau; K Kaua‘i; O 

O‘ahu; Mo Moloka‘i; M Maui; Ka Kahoolawe; 

H Hawai‘i. 




FullName STATUS NI K O MO L M KA H 

ACROPORIUM Mitt. (Sematophyllaceae) 

Acroporium fuscoflavum (Paris) Broth. [Hoe 1974, 

Miller et al. 1978] 

Acroporium fuscoflavum (Paris) Broth. var. 

baldwinii (Paris) Broth. 

?Acroporium sigmatodontium misapplied 

AEROBRYOPSIS M.Fleisch. (Meteoriaceae) 

Aerobryopsis subdivergens (Broth.) Broth. ssp. scariosa 

(E.B.Bartram) Nog. [Hoe 1974, Noguchi 1976] 

Aerobryopsis scariosa E.B.Bartram 

Ptychomnion aciculare misapplied 

Aerobryopsis wallichii (Brid.) M.Fleisch. [Hoe 1974, 

Noguchi 1976] 

Aerobryopsis longissima (Dozy & Molk.) M.Fleisch. 

Aerobryopsis longissima (Dozy & Molk.) 

M.Fleisch. var. dozyana M.Fleisch. 

ind K O Mo L M H 

ind K O Mo M 


AMPHIDIUM Schimp. (Dicranaceae, or 

Rhabdoweisiaceae [Shaw & Goffinet 2000]) 

Amphidium lapponicum (Hedw.) Schimp. [Hoe 1979] ind M 

Amphidium tortuosum (Hornsch.) Cufod. [Frahm et al. 

2000, Muñoz & Pando 2000] 

Amphidium cyathicarpum (Mont.) Broth. 

Grimmia haleakalae Reichardt 

Grimmia haliacalae Reichardt 

ind M H 

ANDREAEA Hedw. (Andreaeaceae) 

Andreaea acutifolia Hook. F. & Wilson ssp. acuminata 

(Mitt.) Vitt [Vitt 1980a, Vitt & Hoe 1980] 

Andreaea acuminata Mitt. 

Andreaea rupestris Hedw. [Hoe 1977, Hoe 1979, Vitt 

1980a, Vitt & Hoe 1980] 

Andreaea perpapillosa Broth. 

ANOECTANGIUM Schwägr. (Pottiaceae) 

Anoectangium aestivum (Hedw.) Mitt. [Hoe 1974, 

Zander 1977, Zander & Vitt 1979] 

Anoectangium euchloron (Schwägr.) Mitt. 

Anoectangium haleakalae (Müll.Hal.) Paris var. 

haleakalae 

Anoectangium haleakalae (Müll.Hal.) Paris var. 

laxulum E.B.Bartram 

Anoectangium harttiae E.B.Bartram 

ind M H 

ind M 


ANOMOBRYUM Schimp. (Bryaceae) 

Anomobryum angustirete Broth. [Hoe 1974] end K L M 

43

44 


FullName 

BALDWINIELLA M.Fleisch. (Neckeraceae) 

STATUS NI K O MO L M KA H 

Baldwiniella kealeensis (Reichardt) E.B.Bartram [Hoe 

1974, Hoe 1975] 

end K O Mo L M H 

BARBELLOPSIS Broth. (Meteoriaceae) 

Barbellopsis trichophora (Mont.) W.R.Buck [Hoe 1974, 

Streimann 1993, Buck 1998] 

Barbella trichophora (Mont.) M.Fleisch. 

BARBULA Hedw. (Pottiaceae) 

Barbula indica (Hook.) Spreng. [Hoe 1978, Zander 1993, 

Darigo 1996] 

Barbula cruegeri Sond. ex Müll.Hal. 

Barbula unguiculata misapplied 

ind K O Mo M H 

nat O M 

BARTRAMIA Hedw. (Bartramiaceae) 

Bartramia baldwinii Müll.Hal. [Hoe 1974] end M 

Bartramia halleriana Hedw. [Hoe 1974] ind M 

BRACHYMENIUM Schwägr. (Bryaceae) 

Brachymenium exile (Dozy & Molk.) Bosch & Sande 

Lac. [Hoe 1974] 

Brachymenium indicum (Dozy & Molk.) Bosch & Sande 

Lac. [Hoe 1979] 


ind O H 

BRACHYTHECIUM Schimp. (Brachytheciaceae) 

Brachythecium hawaiicum E.B.Bartram [Hoe 1974] end M H 

Brachythecium lamprocarpum (Müll.Hal.) A.Jaeger 

[Hoe 1974] 

ind K M H 

Brachythecium plumosum (Hedw.) B.S.G. [Hoe 1974] 

Brachythecium oxyrrhynchum (Dozy & Molk.) 

A.Jaeger 

Cirriphyllum oxyrrhynchum (Dozy & Molk.) 

M.Fleisch. 


Brachythecium rutabulum (Hedw.) B.S.G. [Hoe 1974] ind M 

BREUTELIA (Bruch & Schimp.) Schimp. 

(Bartramiaceae) 

Breutelia affinis (Hook.) Mitt. [Hoe 1974, Virtanen 1997] 

Breutelia kilaueae (Müll.Hal.) Broth. 

ind H 

Breutelia arundinifolia (Duby) M.Fleisch. [Hoe 1974, 

Virtanen 1997] 

ind K O M H 

Breutelia crassicaulis (Müll.Hal.) Paris [Virtanen 1997] ind M H 

BRYOERYTHROPHYLLUM P.C.Chen (Pottiaceae) 

Bryoerythrophyllum recurvirostre (Hedw.) P.C.Chen 

[Hoe 1974] 

Didymodon recurvirostris (Hedw.) Jenn. 

ind M 





BRYUM Hedw. (Bryaceae) 

Bryum argenteum Hedw. [Hoe 1974, Miller et al. 1978, 

Tan & Koponen 1989] 

Bryum argenteum Hedw. var. lanatum (P. Beauv.) 

Hampe 

Bryum leucophyllum Dozy & Molk. 

Bryum atrovirens Brid. [Hoe 1974] 

Bryum erythrocarpum Schwägr. 

ind K O M H 

ind O M H 

Bryum baldwinii Broth. [Hoe 1974] end M 

Bryum caespiticium Hedw. [Hoe 1974] ind M H 

Bryum chrysoneuron Müll.Hal. [Miller et al. 1978] ind H 

Bryum conicum Hornsch. [Ochi 1980] 

Bryum ceramiocarpum Müll.Hal. 

ind M 

Bryum flavituber R.Wilczek & Demaret [Wilczek & 

Demaret 1976] 

nat O 

Bryum hawaiicum Hoe [Hoe 1974] 

Bryum crassicostatum Broth. 

end M H 

Bryum klinggraeffii Schimp. [Hoe 1979] nat O 

Bryum lisae De Not. var. cuspidatum (Bruch & Schimp.) nat O 

Margad. [Olson 1999a] 

Bryum mildeanum Jur. [Hoe 1974, Ochi 1974, Hoe 1979, 

Ignatov & Afonina 1992] 

Bryum apiculatum Schwägr. 

Bryum mauiensis Broth. 

Bryum megalostegium Sull. 

Bryum nitens Hook. 

ind K O L M 

Bryum pseudocapillare Besch. [Hoe 1975] nat? O 

Bryum radiculosum Brid. [Olson 1999a] nat K 

BUCKIELLA Ireland (Hypnaceae) 

Buckiella draytonii (Sull.) Ireland [Hoe 1974, Ireland 

2001] 

Hypnum draytoni(i) Sull. 

Hypnum eudorae Sull. 

Plagiothecium draytonii (Sull.) Müll.Hal. 

end K O Mo M H 

CALYMPERES Sw. (Calymperaceae) 

Calymperes fasciculatum Dozy & Molk. [Hoe 1974, 

Reese et al. 1986] 

Calymperes hawaiiense E.B.Bartram 

ind O 

Calymperes tenerum Müll.Hal. [Hoe 1974] ind K O Mo M H 

CAMPTOCHAETE Reichardt (Lembophyllaceae) 

Camptochaete pulvinata (Hook.f. & Wilson) A.Jaeger 

[Hoe 1974, Tangney 1997] 

ind M 

CAMPYLOPODIUM (Müll.Hal.) Besch. (Dicranaceae) 

Campylopodium medium (Duby) Giese & J.-P.Frahm ind K O Mo M H 

45

46 



[Hoe 1974, Giese & Frahm 1985] 

Campylopodium euphorocladum (Müll.Hal.) Besch. 

CAMPYLOPUS Brid. (Dicranaceae) 

Campylopus exasperatus (Nees & Blume) Brid. [Frahm 

1991] 

Campylopus fragilis (Brid.) Bruch. & Schimp. ssp. 

zollingerianus (Müll.Hal.) J.-P.Frahm [Hoe 1974, 

Frahm 1991, Frahm 1992] 

Campylopus boswellii (Müll.Hal.) Paris 

Campylopus boswellii (Müll.Hal.) Paris var. 

capitulatus E.B.Bartram 

Campylopus hawaiicus (Müll.Hal.) A.Jaeger var. 

densifolius (Ångstr.) J.-P.Frahm [Frahm 1978, Frahm 

1991] 

Campylopus densifolius Ångstr. 

Campylopus densiretis Broth. 

Campylopus wawreanus Reichardt 


hawaiicoflexuosus (Müll.Hal.) J.-P.Frahm [Frahm 

1978, Frahm 1991] 

Campylopus densifolius Ångstr. var. falcatus Broth. 

Campylopus densifolius Ångstr. var. 

hawaiicoflexuosus (Müll.Hal.) H.A.Mill. 

Campylopus hawaiicoflexuosus (Müll.Hal.) 

Müll.Hal. 

Campylopus hawaiicoflexuosus (Müll.Hal.) Paris 

Campylopus perangustifolius Müll.Hal. 

Campylopus purpureoflavescens Hampe, in part 

Dicranum hawaiicoflexuosum Müll.Hal. 


hawaiicus [Frahm 1978, Frahm 1991] 

Campylopus densifolius Ångstr. var. 

purpureoflavescens (Hampe ex Müll.Hal.) 

H.A.Mill. 

Campylopus purpureoflavescens Hampe, in part 

Dicranum hawaiicum Müll.Hal. 

Campylopus incurvatus J.-P.Frahm & Hoe [Frahm & Hoe 

1978, Frahm 1991] 

Campylopus laxitextus Sande Lac. [Hoe 1974, Frahm 

1991] 

Campylopus fumarioli Müll.Hal. 

Campylopus praemorsus (Müll.Hal.) A.Jaeger [Frahm 

1976, Hoe 1979, Frahm 1991] 

Campylopus exasperatus (Nees & Blume) Brid. 

Campylopus microcephalus (Müll.Hal.) Paris 

Dicranum microcephalum Müll.Hal. 

Dicranum praemorsum Müll.Hal. 

ind K 

ind K O Mo M 




end M H 


end K O L M H 




Campylopus schmidii (Müll.Hal.) A.Jaeger ssp. 

hemitrichius (Müll.Hal.) J.-P.Frahm [Hoe 1974, Frahm 

1991] 

Campylopus skottsbergii Broth. 

Campylopus tubulosus E.B.Bartram 

Dicranum skottsbergii Broth. 

ind K O M H 

Campylopus schmidii (Müll.Hal.) A.Jaeger ssp. schmidii 

[Hoe 1974, Hoe 1977, Frahm 1991] 

Campylopus aureus Bosch & Sande Lac. 

Campylopus introflexus misapplied 

Campylopus introflexus (Hedw.) Brid. var. 

polytrichoides misapplied 

Campylopus polytrichoides misapplied 

ind K O M H 

Campylopus umbellatus (Arn.) Paris var. pterotoneuron 

(Müll.Hal.) J.-P.Frahm [Hoe 1974, Frahm 1991] 

Campylopus geniculatus Ångstr. 

ind O 

Campylopus umbellatus (Arn.) Paris var. umbellatus 

[Hoe 1974, Frahm 1991] 

?Campylopus didrichsenii (Müll.Hal.) A.Jaeger 

?Campylopus pertristis (Müll.Hal.) Paris 

Campylopus richardii misapplied 

Campylopus sandwicensis Paris 

Campylopus umbellatus (Arn.) E.B.Bartram 

Dicranum didrichsenii Müll.Hal. 

Dicranum pertriste Müll.Hal. 

Thysanomitrion hawaiicum Müll.Hal. 

Thysanomitrion umbellatum Arn. 

ind K O L M H 

Campylopus wheeleri (Müll.Hal.) Paris [Hoe 1974, 

Frahm 1991] 

Dicranoloma wheeleri (Müll.Hal.) Paris 

Dicranum wheeleri Hampe ex Müll.Hal. 

end K M H 

CERATODON Brid. (Ditrichaceae) 

Ceratodon purpureus (Hedw.) Brid. ssp. purpureus [Hoe 

1974, Hoe 1977, Miller et al. 1978, Burley & Pritchard 

1990] 

Ceratodon purpureus (Hedw.) Brid. fma. purpureus 

Ceratodon purpureus (Hedw.) Brid. fma. xanthopus 

(Sull.) E.Britton ex Grout 

CHENIA R.H.Zander (Pottiaceae) 

Chenia leptophylla (Müll.Hal.) R.H.Zander [Hoe 1974, 

Zander 1993] 

Tortula rhizophylla (Sakurai) Z.Iwats. & K.Saito 

Tortula vectensis E.F.Warb. & Crundw. 

CLAOPODIUM (Lesq. & James) Renauld & Cardot 

(Thuidiaceae, or Leskeaceae [Shaw & Goffinet 2000]) 

Claopodium prionophyllum (Müll. Hal.) Broth. [Hoe 

1974] 


ind K O M H 

nat O 

ind O Mo M H 

47

48 



Claopodium amblystegioides Dixon 

Claopodium nervosum M.Fleisch. 


Claopodium whippleanum (Sull.) Renauld & Cardot 

[Hoe 1974] 

Claopodium hawaiiense R.S.Williams 

ind K O M 

CLASTOBRYOPSIS M.Fleisch. (Sematophyllaceae) 

Clastobryopsis robusta (Broth.) M.Fleisch. [Hoe 1974, 

Tan & Jia 1999] 

Aptychella robusta (Broth.) M.Fleisch. 

CTENIDIUM (Schimp.) Mitt. (Hypnaceae) 

Ctenidium elegantulum Broth. [Hoe 1974, Nishimura 

1985] 

ind K M 

end O M H 

CYRTOPUS (Brid.) Hook.f. (Cyrtopodaceae) 

Cyrtopus setosus (Hedw.) Hook.f. [Hoe 1974] ind H 

DALTONIA Hook. & Taylor (Hookeriaceae, or 

Daltoniaceae [Shaw & Goffinet 2000]) 

Daltonia baldwinii Broth. [Hoe 1974] end M H 

Daltonia contorta Müll.Hal. [Hoe 1974, Hoe 1979] ind K O Mo L M H 

Daltonia pseudostenophylla E.B.Bartram [Hoe 1974] ind O M H 

Daltonia rufescens Broth. [Hoe 1974] ind O M H 

DICRANELLA (Müll.Hal.) Schimp. (Dicranaceae) 

Dicranella exilis Sull. [Hoe 1974] end M 

Dicranella hawaiica (Müll.Hal.) Broth. var. hawaiica 

[Hoe 1974] 

Microdus hawaiicus (Müll.Hal.) Paris 

end K Mo M 

Dicranella hawaiica (Müll.Hal.) Broth. var. tomentella 

E.B.Bartram [Hoe 1974] 

end O M 

Dicranella hochreutineri Cardot [Hoe 1974] 

Dicranella hochreuteri misapplied 

ind K O L M H 

Dicranella integrifolia E.B.Bartram [Hoe 1974] end O Mo M H 

Dicranella rigidula E.B.Bartram [Hoe 1974] end K O H 

DICRANODONTIUM Bruch & Schimp. (Dicranaceae) 

Dicranodontium denudatum (Brid.) E.Britton [Hoe 1974] ind M 

Dicranodontium falcatum Broth. var. atrovirens (Broth.) 

E.B.Bartram [Hoe 1974] 

end M H 

Dicranodontium porodictyon Cardot & Thér. [Hoe 1974, 

Hoe 1979, Frahm 1997] 

Dicranodontium falcatum Broth. var. falcatum 

Dicranodontium hawaiicum Broth. 


DICRANOLOMA (Renauld) Renauld (Dicranaceae) 

Dicranoloma gracile Broth. ex E.B.Bartram [Hoe 1974] end M 





DICRANOWEISIA Milde (Dicranaceae, or 


Dicranoweisia cirrata (Hedw.) Lindb. [Hoe 1974, Miller 

et al. 1978] 

DICRANUM Hedw. (Dicranaceae) 

Dicranum speirophyllum Mont. var. breviflagellare 

(Müll.Hal.) E.B.Bartram [Hoe 1974] 

Dicranum speirophyllum Mont. var. condensatum (Sull.) 

Wijk & Margad. [Hoe 1974] 

Dicranum speirophyllum Mont. var. elongatum (Sull.) 

Wijk & Margad. [Hoe 1974] 

Dicranum speirophyllum Mont. var. speirophyllum [Hoe 

1974] 

DIDYMODON Hedw. (Pottiaceae) 

Didymodon insulanus (De Not.) M.O.Hill [Hoe 1974, 

Zander 1978, Corley et al. 1981] 

Barbula vinealis Brid. var. cylindrica (Taylor) 

Boulay 

Barbula vinealis Brid. var. flaccida B.S.G. 

Didymodon vinealis (Brid.) R.H.Zander var. 

flaccidus (Bruch & Schimp.) R.H.Zander 

ind H? 

end O M H 

end H 

end M 


ind O M 

DISTICHIUM Bruch & Schimp. (Ditrichaceae) 

Distichium capillaceum (Hedw.) B.S.G. [Hoe 1974] ind H 

DISTICHOPHYLLUM Dozy & Molk. (Hookeriaceae, 

or Daltoniaceae [Shaw & Goffinet 2000]) 

Distichophyllum freycinetii (Schwägr.) Mitt. var. 

crasseturgescens (Müll.Hal.) E.B.Bartram ex Hoe [Hoe 

1974] 

Mniadelphus freycineti (Schwägr.) Müll.Hal. var. 

crasseturgescens Müll.Hal. 

end 

Distichophyllum freycinetii (Schwägr.) Mitt. var. 

freycinetii [Hoe 1974] 


Distichophyllum paradoxum (Mont.) Mitt. [Hoe 1974] end K O Mo L M H 

ECTROPOTHECIUM Mitt. (Hypnaceae) 

Ectropothecium arcuatum Mitt. [Hoe 1974] end K O M H 

Ectropothecium decurrens (Sull.) Nishimura [Hoe 1974, 

Nishimura 1985] 

Ctenidium decurrens (Sull.) Broth. 

Ctenidium pulcherrimum Broth. 

Ctenidium submalacobolum Broth. 

Cupressina decurrens (Sull.) Müll.Hal. 

Ectropothecium rockii Broth. 

Hypnum decurrens Sull. 


49

50 



Stereodon decurrens (Sull.) Mitt. 

Taxiphyllum gracile Broth. 


Ectropothecium sandwichense (Hook. & Arn.) Mitt. 

[Hoe 1974, Hoe 1979] 


Ectropothecium tutuilum (Sull.) Mitt. [Miller et al. 1978] ind H 

Ectropothecium viridifolium E.B.Bartram [Hoe 1974, end Ni O L M 

Wichman & St. John 1990] 

Ectropothecium zollingeri (Müll. Hal.) A.Jaeger var. 

filicaule (M.Fleisch.) H.A.Mill., H.Whittier & 

B.Whittier [Hoe 1974, Hoe 1975, Miller et al. 1978] 

Glossadelphus zollingeri (Müll.Hal.) M.Fleisch. var. 

filicaulis (M.Fleisch.) M.Fleisch. 


ENCALYPTA Hedw. (Encalyptaceae) 

Encalypta ciliata Hedw. [Horton 1983] 

Encalypta scabrata E.B.Bartram 

ind M H 

Encalypta rhabdocarpa Schwägr. [Hoe 1974] 

Encalypta rhaptocarpa misapplied 

ind H 

Encalypta sandwicensis Sull. [Hoe 1974, Hoe 1977] ind M H 

ENTODON Müll.Hal. (Entodontaceae) 

Entodon plicatus Müll.Hal. [Hoe 1974, Buck 1990] 

Entodon hillebrandii Müll.Hal. 

Entodon solanderi (Ångstr.) A.Jaeger 

Entodon subcuspidatus (Müll.Hal.) E.B.Bartram [Hoe 

1974, Hoe 1979] 

ENTODONTOPSIS Broth. (Stereophyllaceae) 

Entodontopsis anceps (Bosch & Sande Lac.) W.R.Buck 

& Ireland [Hoe 1974, Buck & Ireland 1985] 

Stereophyllum oahuense Broth. 

ENTOSTHODON Schwägr. (Funariaceae) 

Entosthodon subintegrus (Broth.) H.A.Mill., H.Whittier 

& B.Whittier [Miller et al. 1978] 

Funaria subintegra Broth. 

ind K O M H 

ind K Mo M 

ind O 


FABRONIA Raddi (Fabroniaceae) 

Fabronia ciliaris (Brid.) Brid. [Hoe 1977] ind M 

Fabronia degeneri E.B.Bartram [Hoe 1974] end O Mo H 

Fabronia nietneri Müll.Hal. [Hoe 1974, Hoe 1975] ind K O M H 

FISSIDENS Hedw. (Fissidentaceae) 

Fissidens aphelotaxifolius Pursell [Pursell & Hoe 1977] ind M 

Fissidens bryoides Hedw. [Pursell & Hoe 1977] 

Fissidens hawaiicus E.B.Bartram 

Fissidens insularis E.B.Bartram 

Fissidens oahuensis E.B.Bartram 





Fissidens delicatulus Ångstr. [Pursell & Hoe 1977] end K O L M H 

Fissidens elegans Brid. [Pursell & Hoe 1977] 

Fissidens baldwinii Broth. 

Fissidens bishopii Hoe 


Fissidens hoei Pursell [Pursell & Hoe 1977] end O L 

Fissidens kilaueae Hoe & H.A.Crum [Pursell & Hoe 

1977] 

end K O L M H 

Fissidens lancifolius E.B.Bartram [Pursell & Hoe 1977] end K O Mo L M H 

Fissidens lindbergii Mitt. [Pursell & Hoe 1977, Darigo 

1996, Pursell 1997] 

Fissidens crumii Hoe 

nat O M 

Fissidens nothotaxifolius Pursell & Hoe [Pursell & Hoe 

1977] 


Fissidens pacificus Ångstr. [Hoe 1974, Pursell & Hoe 

1977] 

Fissidens mauiensis Müll.Hal. 

end K O M H 

Fissidens submarginatus Bruch [Pursell & Hoe 1977, 

Hoe 1979, Bruggeman-Nannenga & Pursell 1995] 

Fissidens intermedius Müll.Hal. 


Fissidens taxifolius Hedw. [Pursell & Hoe 1977] ind O M 

FLORIBUNDARIA M.Fleisch. (Meteoriaceae) 

Floribundaria floribunda (Dozy & Molk.) M.Fleisch. 

[Hoe 1974, Tan 1984] 

Floribundaria baldwinii Broth. 


ind O M H 

FUNARIA Hedw. (Funariaceae) 

Funaria hygrometrica Hedw. [Hoe 1974, Hoe 1975] ind O H 

GAROVAGLIA Endl. (Pterobryaceae, or Garovagliaceae 

[Shaw & Goffinet 2000]) 

Garovaglia powellii Mitt. var. haleakalae (Broth.) During 

[During 1977] 

Garovaglia haleakalae Broth. 

ind M 

GLOSSADELPHUS M.Fleisch. 1 (Sematophyllaceae) 

Glossadelphus acutifolius E.B.Bartram [Hoe 1974, Tixier 

1988] 

end M 

Glossadelphus limnobioides Broth. [Hoe 1974, Tixier 

1988] 

end O M H 

Glossadelphus mauiensis Broth. [Hoe 1974, Tixier 1988] end M 

Glossadelphus ogatae Broth. & Yasuda [Hoe 1974, Hoe 

1979, Tixier 1988] 

Glossadelphus chrysobasilaris Broth. 

ind M H 

1 Buck (1987b) reduced the type species of Glossadelphus to Phyllodon so the genus name is not usable. 

The correct placement for the 4 species left here remains to be determined. 

51

52 



GRIMMIA Hedw. (Grimmiaceae) 


Grimmia anodon Bruch & Schimp. [Hoe 1977, Hoe 

1979, Muñoz & Pando 2000] 

Grimmia apocarpa Hedw. var. pulvinata (Hedw.) 

G.N.Jones 

Gymnostomum pulvinatum Hedw. 

ind M H 

Grimmia laevigata (Brid.) Brid. [Hoe 1974, Hoe 1979, 

Muñoz & Pando 2000] 

ind M H 

Grimmia longirostris Hook. [Hoe 1977,Muñoz 1998] 

Grimmia pilifera misapplied 

ind M 

Grimmia mauiense Greven [Greven 2003] end M H 

Grimmia maunakeaense Greven [Greven 2003] end M H 

Grimmia pulvinata (Hedw.) Sm. [Muñoz & Pando 2000] 

Leersia pulvinata Hedw. 

ind M 

Grimmia trichophylla Grev. [Hoe 1974, Muñoz & Pando 

2000] 

ind M H 

HAGENIELLA Broth. (Sematophyllaceae) 

Hageniella micans (Mitt.) B.C.Tan & Y.Jia [Hoe 1974, 

Tan & Jia 1999] 

Hageniella pacifica Broth. 

HAPLOCLADIUM Müll.Hal. (Leskeaceae) 

Haplocladium microphyllum (Hedw.) Broth. [Hoe 1974, 

Watanabe & Iwatuski 1981, Zilstra 1990] 

Bryohaplocladium microphyllum (Hedw.) R.Watan. 

& Z.Iwats. 

HAPLOHYMENIUM Dozy & Molk. (Thuidiaceae, or 

Anomodontaceae [Shaw & Goffinet 2000]) 

Haplohymenium triste (Ces.) Kindb. [Hoe 1974, Hoe 

1979] 

HETEROPHYLLIUM (Schimp.) Kindb. 

(Sematophyllaceae) 

Heterophyllium subauriculatum (Müll.Hal.) Broth. [Hoe 

1974] 

HOLOMITRIUM Brid. (Dicranaceae) 

Holomitrium seticalycinum Müll.Hal. [Hoe 1974, Hoe 

1979] 

Holomitrium squarrifolium E.B.Bartram [Hoe 1974] 

Holomitrium ferriei misapplied 

HOMALIODENDRON M.Fleisch. (Neckeraceae) 

Homaliodendron flabellatum (Sm.) M.Fleisch. [Hoe 

1974] 

Homalia densa Bosw. 

Homalia praelonga Reichardt 

ind K M H 

ind O 

ind K O M H 

end M H 


end O H 






HOOKERIA Sm. (Hookeriaceae) 

Hookeria acutifolia Hook. & Grev. [Hoe 1974, Hoe 

1975] 

HYMENOSTYLIUM Brid. (Pottiaceae) 

Hymenostylium recurvirostrum (Hedw.) Dixon var. 

cylindricum (E.B.Bartram) R.H.Zander [Zander & 

Eckel 1982] 

Merceyopsis crassinervis (Broth.) E.B.Bartram 

Molendoa crassinervis (Broth.) E.B.Bartram 

Hymenostylium recurvirostrum (Hedw.) Dixon var. 

recurvirostrum [Zander 1993] 

Hymenostylium firmum (Müll.Hal.) Broth. ex 

E.B.Bartram 

HYOPHILA Brid. (Pottiaceae) 

Hyophila involuta (Hook.) A.Jaeger [Hoe 1974] 

Hyophila dozy-molkenboeri M.Fleisch. 

HYPNUM Hedw. (Hypnaceae) 

Hypnum plumaeforme Wilson [Crum & Mueller- 

Dombois 1968, Hoe 1974] 

HYPOPTERYGIUM Brid. (Hypopterygiaceae) 

Hypopterygium sandwicense Broth. [Hoe 1974, Kruijer 

2002] 

ind O L M H 

ind H 

ind K O M H 


nat M H 

end M H 

ISOPTERYGIUM Mitt. (Hypnaceae) 

Isopterygium albescens (Hook.) A.Jaeger [Hoe 1974] ind K O Mo L M H 

Isopterygium vineale E.B.Bartram [Hoe 1974] end K O Mo M H 

LEPTOBRYUM (Bruch & Schimp.) Wilson (Bryaceae, 

or Meesiaceae [Shaw & Goffinet 2000]) 

Leptobryum pyriforme (Hedw.) Wilson [Hoe 1974] ind M H 

LEPTODONTIUM (Müll.Hal.) Lindb. (Pottiaceae) 

Leptodontium flexifolium (Dicks. ex With.) Hampe [Hoe 

1974, Hoe 1977] 

Leptodontium brevicaule E.B.Bartram 

ind M H 

LEUCOBRYUM Hampe (Leucobryaceae) 

Leucobryum gracile Sull. var. gracile [Hoe 1974] end K O Mo L M H 

Leucobryum gracile Sull. var. hamatum E.B.Bartram 

[Hoe 1974] 

end K O Mo M 

Leucobryum pachyphyllum Müll.Hal. [Hoe 1974] end K O Mo L M 

Leucobryum seemannii Mitt. var. fumarioli (Müll.Hal.) 

H.A.Mill. [Hoe 1974] 

end H 

53

54 



Leucobryum hawaiense (Reichardt) E.B.Bartram var. 

fumarioli (Müll.Hal.) E.B.Bartram 

Leucobryum solfatarae Müll.Hal. var. fumarioli 

(Müll.Hal.) Wijk & Margad. 


Leucobryum seemannii Mitt. var. seemannii [Hoe 1974, 

Hoe 1975] 

Leucobryum hawaiense (Reichardt) E.B.Bartram 

Leucobryum solfatarae Müll.Hal. var. hawaiense 


(Reichardt) Wijk & Marg. 

Leucobryum seemannii Mitt. var. solfatarae (Müll.Hal.) 

H.A.Mill. [Hoe 1974] 

Leucobryum hawaiense (Reichardt) E.B.Bartram var. 

solfatarae (Müll.Hal.) E.B.Bartram 

Leucobryum solfatarae Müll.Hal. 

end K Mo M H 

LEUCOLOMA Brid. (Dicranaceae) 

Leucoloma molle (Müll.Hal.) Mitt. [Hoe 1974] ind O M H 

Leucoloma scaberulum E.B.Bartram [Hoe 1974] end O 

LEUCOPHANES Brid. (Leucobryaceae, or 

Calymperaceae [Shaw & Goffinet 2000]) 

Leucophanes glaucum (Schwägr.) Mitt. [Hoe 1974] ind O 

LIMBELLA (Müll.Hal.) Müll.Hal. (Amblystegiaceae) 

Limbella tricostata (Sull.) Müll.Hal. [Hoe 1974, Christy 

1987] 

Sciaromium tricostatum (Sull.) Mitt. 

MACROCOMA (Müll.Hal.) Grout (Orthotrichaceae) 

Macrocoma tenue (Hook. & Grev.) Vitt ssp. sullivantii 

(Müll.Hal.) Vitt [Hoe 1974, Vitt 1980b, Vitt 1981] 

Macrocoma intricata (Müll.Hal.) H.A.Miller, 

H.Whittier & B.Whittier 

Macromitrium intricatum Müll.Hal. 



MACROMITRIUM Brid. (Orthotrichaceae) 

Macromitrium brevisetum Mitt. [Hoe 1974, Hoe 1979] end K O Mo L M H 

Macromitrium emersulum Müll.Hal. [Hoe 1974, Hoe 

1975] 


Macromitrium incurvifolium (Hook. & Grev.) Schwägr. 

[Hoe 1974] 

Macromitrium cumingii Müll.Hal. 

ind K M H 

Macromitrium microstomum (Hook. & Grev.) Schwägr. 

[Hoe 1974, Vitt 1983] 

Macromitrium owahiense Müll.Hal. 

Macromitrium reinwardtii Schwägr. 


Macromitrium piliferum Schwägr. [Hoe 1974] end K O Mo L M H 




MACROTHAMNIUM M.Fleisch. (Hylocomiaceae) 


Macrothamnium macrocarpum (Reinw. & Hornsch.) 

M.Fleisch. [Hoe 1974] 

ind M 

MICRODUS Schimp. ex Besch. (Dicranaceae) 

Microdus hillebrandii (Müll.Hal.) Paris [Hoe 1974] 

Dicranella hillebrandii (Müll.Hal.) Broth. 


ind O Mo 

MNIOBRYOIDES Hörmann (Bryaceae) 

Mniobryoides degeneriae Hörmann [Hoe 1974] end H 

MNIUM Hedw. (Mniaceae) 

Mnium marginatum (With.) P.Beauv. [Hoe 1974] 

Mnium serratum Schrad. ex Brid. 

ind M 

NECKERA Hedw. (Neckeraceae) 

Neckera hawaiicopennata Müll.Hal. [Hoe 1974] end M 

NECKEROPSIS Reichardt (Neckeraceae) 

Neckeropsis lepineana (Mont.) M.Fleisch. [Hoe 1974, 

Enroth 1989] 

ind K O M 

Neckeropsis obtusata (Mont.) Broth. [Olson 1999b] nat K 

OCTOBLEPHARUM Hedw. (Leucobryaceae, or 

Calymperaceae [Shaw & Goffinet 2000]) 

Octoblepharum albidum Hedw. [Hoe 1974, Hoe 1975] ind K O Mo M H 

ORTHODONTIUM Wilson (Bryaceae, or 

Orthodontiaceae [Shaw & Goffinet 2000]) 

Orthodontium pellucens (Hook.) B.S.G. [Hoe 1974, Hoe 

1977] 

ind K M 

ORTHOTRICHUM Hedw. (Orthotrichaceae) 

Orthotrichum alpestre Hornsch. var. berggrenii 

(E.B.Bartram) Lewinsky [Hoe 1974, Lewinksy 1993] 

Orthotrichum berggrenii E.B.Bartram 

end O? H 

Orthotrichum diaphanum Schrad. ex Brid. [Hoe 1974, 

Hoe 1977, Lewinsky 1993] 

ind M H 

Orthotrichum hawaiicum Müll.Hal. [Hoe 1974] end M H 

Orthotrichum incrassatum Lewinsky [Lewinsky 1993] end H 

Orthotrichum lyellii Hook. & Taylor [Hoe 1974, 

Lewinsky 1993] 

Orthotrichum verrucatum Müll.Hal. 

ind 

Orthotrichum rupestre Schleich. & Schwägr. [Hoe 1974, 

Lewinsky 1993] 

Orthotrichum hillebrandii Müll.Hal. 

ind O M H 

55

56 



OXYRRHYNCHIUM (Schimp.) Warnst. 

(Brachytheciaceae) 


Oxyrrhynchium vagans (A.Jaeger) Ignatova & Huttunen 

[Hoe 1974, Hoe 1975, Ignatov & Huttunen 2002] 

Eurhynchium vagans (A.Jaeger) E.B.Bartram 

Rhynchostegium vagans A.Jaeger 


OXYSTEGUS (Limpr.) Hilp. (Pottiaceae) 

Oxystegus tenuirostris (Hook. & Taylor) A.J.E.Sm. var. 

gemmiparus (Schimp.) R.H.Zander [Hoe 1977, Hoe 

1979, Zander 1982] 

Trichostomum tenuirostre (Hook. & Taylor) Lindb. 

PALAMOCLADIUM Müll.Hal. (Brachytheciaceae) 

Palamocladium wilkesianum (Sull.) Müll.Hal. [Hoe 

1974, Hofmann 1997] 

Hypnum wilkesianum Sull. 

Meteorium pallidovirens Müll.Hal. 

Meteorium sciuroides Müll.Hal. 

Palamocladium aptychoides Müll.Hal. 

Palamocladium sciurellum Müll.Hal. 

Palamocladium wilkesianum (Sull.) Müll.Hal. var. 

altisetum Müll.Hal. 

Palamocladium wilkesianum (Sull.) Müll.Hal. var. 

sciuroides (Müll.Hal.) Wijk & Margad. 

Pleuropus aptychoides (Müll.Hal.) Broth. 

Pleuropus pallidovirens (Müll.Hal.) Broth. 

Pleuropus sciurellus (Müll.Hal.) Broth. 

Pleuropus wilkesianus (Sull.) Broth. 

Pleuropus wilkesianus (Sull.) Broth. var. sciuroides 

(Müll.Hal.) E.B.Bartram 

PELEKIUM Mitt. (Thuidiaceae) 

Pelekium versicolor (Hornsch. ex Müll.Hal.) Touw [Hoe 

1974, Touw 2001] 

Thuidium crenulatum Mitt. 

Thuidium nanophyllum Müll.Hal. 

ind K O M 


ind O M H 

PHILONOTIS Brid. (Bartramiaceae) 

Philonotis falcata (Hook.) Mitt. [Hoe 1974] ind K O Mo M 

Philonotis hastata (Duby) Wijk & Margad. [Hoe 1974] 

Philonotis laxissima Mitt. 

ind O M H 

Philonotis hawaica (Müll.Hal.) Broth. [Hoe 1974] end K O Mo L M H 

Philonotis turneriana (Schwägr.) Mitt. var. sullivantii 

(Müll.Hal.) E.B.Bartram [Hoe 1974] 

end O M H 

Philonotis turneriana (Schwägr.) Mitt. var. turneriana 

[Hoe 1974] 





PILOTRICHELLA (Müll.Hal.) Besch. (Meteoriaceae, 

or Lembophyllaceae [Shaw & Goffinet 2000]) 

Pilotrichella mauiensis (Sull.) A.Jaeger [Hoe 1974] end O M H 

PLAGIOMNIUM T.J.Kop. (Mniaceae) 

Plagiomnium rostratum (Schrad.) T.J.Kop. [Hoe 1974] 

Mnium rostratum Schrad. 

Plagiomnium prorepens (Müll.Hal.) T.J.Kop. 

Plagiomnium rhynchosporum (Hook.) T.J.Kop. 

PLAGIOPUS Brid. (Bartramiaceae) 

Plagiopus oederiana (Sw.) H.A.Crum & L.E.Anderson 

[Hoe 1974] 

Plagiopus longisetus (Brid.) E.B.Bartram 

Plagiopus oederi (Brid.) Limpr. 



ind M 

PLAGIOTHECIUM Bruch & Schimp. 

(Plagiotheciaceae) 

Plagiothecium cavifolium (Brid.) Z.Iwats. [Hoe 1977] ind M 

Plagiothecium denticulatum (Hedw.) Schimp. [Hoe 

1974] 

ind 

Plagiothecium mauiense Broth. [Hoe 1974] end M 

PLATYHYPNIDIUM M.Fleisch. (Amblystegiaceae, or 

Brachytheciaceae [Shaw & Goffinet 2000]) 

Platyhypnidium muelleri (A.Jaeger) M.Fleisch. [Hoe 

1974] 

Eurhynchium mulleri misapplied 

POGONATUM P. Beauv. (Polytrichaceae) 

Pogonatum tahitense Schimp. ex Besch. [Hoe 1974] 

Pogonatum baldwinii (Müll.Hal.) Paris 

POHLIA Hedw. (Bryaceae, or Mniaceae [Shaw & 

Goffinet 2000]) 

Pohlia baldwinii (Broth. ex E.B.Bartram) W.Schultze- 

Motel [Hoe 1974] 

Webera baldwinii Broth. ex E.B.Bartram 

Pohlia cruda (Hedw.) Lindb. [Hoe 1974, Hoe 1979] 

Webera cruda (Hedw.) Fürnr. 

Pohlia flexuosa Hook. [Hoe 1974, Gangulee 1974] 

Pohlia gracilescens (E.B.Bartram) E.B.Bartram 

Pohlia gracillima (Cardot) Horik. & Ochi 

Pohlia leucostoma (Bosch & Sande Lac.) M.Fleisch. 

Pohlia leucostomoides (Broth.) E.B.Bartram 

Webera gracilescens E.B.Bartram 

Webera leucostomoides Broth. 

Pohlia mauiensis (Broth. ex E.B.Bartram) W.Schultze- 

Motel [Hoe 1974] 

ind K O Mo M 


end M 

ind M H 


end M H 

57

58 



POLYTRICHUM Hedw. (Polytrichaceae) 

Polytrichum commune Hedw. [Hoe 1974] ind H 

Polytrichum juniperinum Hedw. [Hoe 1974, Hyvönen 

1986] 

ind H 

Polytrichum piliferum Hedw. [Hoe 1974] ind M H 

PSEUDOSCLEROPODIUM (Limpr.) M.Fleisch. 


Pseudoscleropodium purum (Hedw.) M.Fleisch. [Hoe 

1974] 

PSEUDOSYMBLEPHARIS Broth. (Pottiaceae) 

Pseudosymblepharis angustata (Mitt.) Hilp. [Hoe 1974, 

Hoe 1979, Norris & Koponen 1989] 

Pseudosymblepharis mauiensis (Mull.Hal.) Broth. 

PSEUDOTAXIPHYLLUM Z.Iwats. (Hypnaceae) 

Pseudotaxiphyllum elegans (Brid.) Z.Iwats. [Hoe 1979, 

Iwatsuki 1987] 

Isopterygium elegans (Brid.) Lindb. 

nat H 


ind M 

PTEROGONIDIUM Broth. (Sematophyllaceae) 

Pterogonidium pulchellum (Hook.) Müll.Hal. [Hoe 1979] nat? O 

PTYCHOMITRIUM Fürnr. (Ptychomitriaceae) 

Ptychomitrium lepidomitrium (Müll.Hal.) Schimp. ex 

Besch. [Hoe 1974, Cao et al. 2003] 

Ptychomitrium mauiense Broth. 

PYLAISIADELPHA Broth. (Meteoriaceae) 

Pylaisiadelpha opaeodon (Sull.) W.R. Buck [Hoe 1974, 

Buck 1984] 

Brotherella opaeodon (Sull.) Broth. 

Hypnum opaeodon Sull. 

PYRRHOBRYUM Mitt. (Rhizogoniaceae) 

Pyrrhobryum spiniforme (Hedw.) Mitt. [Hoe 1974, 

Koponen et al. 1986] 

Rhizogonium spiniforme (Hedw.) Bruch 

RACOMITRIUM Brid. (Grimmiaceae) 

Racomitrium crispulum (Hook.f. & Wilson) Hook.f. & 

Wilson [Hoe 1974] 

Racomitrium fasciculare (Schrad. ex Hedw.) Brid. var. 

erosum Broth. [Hoe 1974] 

Racomitrium lanuginosum (Hedw.) Brid. var. 

lanuginosum [Hoe 1974] 

ind M H 



ind K M H 

end M 

ind M 





pruinosum Wilson [Hoe 1974] 


sandvicense Reichardt 


RACOPILUM P.Beauv. (Racopilaceae) 

Racopilum cuspidigerum (Schwägr.) Ångstr. [Hoe 1974] 

Rhacomitrium cuspidigerum misapplied 

RADULINA W.R.Buck & B.C.Tan (Sematophyllaceae) 

Radulina hamata (Dozy & Molk.) W.R.Buck & B.C.Tan 

[Hoe 1974, Buck & Tan 1989] 

Trichosteleum hamatum (Dozy & Molk.) A.Jaeger 

REMYELLA Müll.Hal. (Brachytheciaceae) 

Remyella hawaica Müll.Hal. [Hoe 1974, Ignatov & 

Huttunen 2002] 

Rhynchostegiella hawaica (Müll.Hal.) Broth. 

RHABDOWEISIA Bruch & Schimp. (Dicranaceae, or 


Rhabdoweisia crenulata (Mitt.) H.Jameson [Hoe 1975, 

Frahm et al. 2000] 

Rhabdoweisia crispata (With.) Lindb. [Hoe 1974, Hoe 

1977, Hoe 1979] 

Rhabdoweisia denticulata B.S.G. 


nat? K O Mo L M H 


end O 

ind M 

RHAPHIDOSTICHUM M.Fleisch. (Sematophyllaceae) 

Rhaphidostichum pustulatum Hoe [Hoe 1974] end O 

ind M H 

RHIZOGONIUM Brid. (Rhizogoniaceae) 

Rhizogonium pungens Sull. [Hoe 1974] end K O Mo L M H 

RHODOBRYUM (Schimp.) Limpr. (Bryaceae) 

Rhodobryum giganteum (Schwägr.) Paris [Hoe 1974, 

Crosby et al. 1983, Tan & Iwatsuki 1991] 

Bryum giganteum (Schwägr.) Arn. 

RHYNCHOSTEGIUM Bruch & Schimp. 


Rhynchostegium celebicum (Sande Lac.) A.Jaeger [Hoe 

1974, Hoe 1977] 

Eurhynchium celebicum (Sande Lac.) E.B.Bartram 

Rhynchostegium gaudichaudii (Mont.) A.Jaeger [Hoe 

1974] 

Rhynchostegium selaginellifolium Müll.Hal. var. 

recurvirameum (Müll.Hal.) Wijk & Margad. [Hoe 

1974] 

Eurhynchium selaginellifolium var. recurvirameum 

ind K O M 

ind K O Mo M 

ind 

end K O H 

59

60 



(Müll.Hal.) E.B.Bartram 


Rhynchostegium selaginellifolium Müll.Hal. var. 

selaginellifolium [Hoe 1974] 

Eurhynchium selaginellifolium (Müll.Hal.) 

E.B.Bartram 


ROSULABRYUM J.R.Spence (Bryaceae) 

Rosulabryum billardierei (Schwägr.) J.R.Spence 

[Mohamed 1979, Anderson et al. 1990, Spence 1996] 

Bryum andicola Hook. 

Bryum decaisnei Dozy & Molk. 

Bryum limbatomarginatum Müll.Hal. 

Bryum truncorum misapplied 

Rosulabryum capillare (Hedw.) J.R.Spence [Hoe 1974, 

Hoe 1977, Spence 1996] 

Bryum capillare Hedw. 

Bryum vinoviride E.B.Bartram 

ind O M H 

ind K O M H 

SAELANIA Lindb. (Ditrichaceae) 

Saelania glaucescens (Hedw.) Broth. [Hoe 1974] ind M 

SCHISTIDIUM Bruch & Schimp. (Grimmiaceae) 

Schistidium apocarpum (Hedw.) Bruch & Schimp. 

[Schultze-Motel 1963, Muñoz & Pando 2000] 

Grimmia apocarpa Hedw. 

Schistidium strictum (Turner) Loeske ex Mårtensson 

[Hoe 1974, Cao & Churchill 1995, Muñoz & Pando 

2000] 

Grimmia apocarpa Hedw. var. stricta (Turner) Hook. 

& Taylor 

Grimmia scabrifolia Broth. 

Grimmia stricta Turner 

SCHIZYMENIUM Harv. (Bryaceae, or Mniaceae [Shaw 

& Goffinet 2000]) 

Schizymenium campylocarpum (Hook. & Arn.) A.J.Shaw 

var. campylocarpum [Zanten 1971, Shaw 1985] 

Mielichhoferia campylocarpa (Hook. & Arn.) Mitt. 

Weissia campylocarpa Hook. & Arn. in Hook. 

Schizymenium nealiae (E.B.Bartram) A.J.Shaw [Hoe 

1974, Shaw 1985] 

Mielichhoferia nealiae E.B.Bartram 

Schizymenium pulvinatum (Müll.Hal.) A.J.Shaw [Hoe 

1974, Shaw 1985] 

Mielichhoferia pulvinata Müll.Hal. 

ind M 

ind M 

ind H 

end H 

end M H 

SCOPELOPHILA (Mitt.) Lindb. (Pottiaceae) 

Scopelophila infericola Hoe [Hoe 1974] end H 




SEMATOPHYLLUM Mitt. (Sematophyllaceae) 


Sematophyllum hawaiiense (Broth.) Broth. [Hoe 1974, 

Hoe 1979] 

ind K O L M H 

Sematophyllum subpinnatum (Brid.) E.Britton [Hoe 

1974, Hoe 1979, Buck 1983] 

Sematophyllum caespitosum misapplied 

ind O H 

SPHAGNUM L. (Sphagnaceae) 

Sphagnum palustre L. [Hoe 1978, Karlin & Andrus 

1995] 

Sphagnum henryense misapplied 

Sphagnum lonchocladum Müll.Hal. 

Sphagnum perichaetiale misapplied 

Sphagnum vitjianum misapplied 

Sphagnum wheeleri Müll.Hal. [Hoe 1974, Karlin & 

Andrus 1995, Karlin 2001] 

Sphagnum compactum misapplied 

Sphagnum rigidulum Warnst. 

Sphagnum vulcanicum Warnst. 

SPLACHNOBRYUM Müll.Hal. (Splachnobryaceae, or 

Pottiaceae [Shaw & Goffinet 2000]) 

Splachnobryum obtusum (Brid.) Müll.Hal. [Hoe 1974, 

Arts 2001] 


ind H 

end M 

nat O 

STREPTOPOGON Mitt. (Pottiaceae) 

Streptopogon erythrodontus (Taylor) Wilson [Hoe 1974] ind M 

SYMPHYODON Mont. (Symphyodontaceae) 

Symphyodon pygmaeus (Broth.) S.He & Snider [Hoe 

1974, Hoe 1977, He & Snider 1992] 

Glossadelphus abortivapicus Hoe 

SYNTRICHIA Brid. (Pottiaceae) 

Syntrichia fragilis (Taylor) Ochyra [Hoe 1974, Kürschner 

2000] 

Tortula alpina (Brid.) Arn. var. inermis (Milde) De 

Not. 

Tortula fragilis Taylor 

Syntrichia princeps (De Not.) Mitt. [Hoe 1974, 

Kürschner 2000] 

Tortula princeps De Not. 

SYRRHOPODON Schwägr. (Calymperaceae) 

Syrrhopodon armatus Mitt. [Hoe 1974, Orbán & Reese 

1986] 

Syrrhopodon oahuensis Broth. 

ind O M 

ind M H 

ind M H 

ind O M H 

Syrrhopodon hawaiicus Müll.Hal. [Hoe 1974] end K O Mo M H 

Syrrhopodon prolifer Schwägr. [Hoe 1974, Reese et al. end O Mo M H 

61

62 



1986] 

Syrrhopodon kilaueae Müll.Hal. 

Syrrhopodon semiliber (Mitt.) Besch. [Hoe 1974, Reese 

et al. 1986] 

Calymperes semiliberum Mitt. 

Calymperopsis semiliber (Mitt.) M.Fleisch. 

ind O H 

TAXIPHYLLUM M.Fleisch. (Hypnaceae) 

Taxiphyllum laevifolium (Mitt.) W.R.Buck [Hoe 1974, 

Buck 1987b, Tixier 1988] 

Glossadelphus baldwinii Broth. 

Glossadelphus laevifolius (Mitt.) E.B.Bartram 

Taxiphyllum torrentium (Besch.) W.R.Buck [Hoe 1974, 

Buck 1987b, Tixier 1988] 

Glossadelphus irroratus E.B.Bartram 

Glossadelphus torrentium (Besch.) M.Fleisch. 

TAXITHELIUM Mitt. (Sematophyllaceae) 

Taxithelium mundulum (Sull.) E.B.Bartram [Hoe 1974] 

Taxithelium undulatum misapplied 


ind M 


TAYLORIA Hook. (Splachnaceae) 

Tayloria sandwicensis (Müll.Hal.) Broth. [Hoe 1974] end M 

THAMNIOPSIS (Mitt.) Fleisch. (Hookeriaceae, or 

Callicostaceae [Buck 1987a]) 

Thamniopsis utacamundiana (Mont.) W.R Buck [Hoe 

1974, Buck 1987a] 

Hookeria purpurea Müll.Hal. 

Hookeriopsis purpurea (Müll.Hal.) Broth. 

Hookeriopsis purpurea (Müll.Hal.) Broth. var. 

acuminatula (Müll.Hal.) E.B.Bartram 

Hookeriopsis purpurea (Müll.Hal.) Broth. var. 

ligulacea (Müll.Hal.) E.B.Bartram 

THAMNOBRYUM Nieuwl. (Neckeraceae) 

Thamnobryum speciosum (Broth.) Hoe [Hoe 1974] 

Thamnium speciosum Broth. 

THUIDIUM Bruch & Schimp. (Thuidiaceae) 

Thuidium cymbifolium (Dozy & Molk.) Dozy & Molk. 

[Hoe 1974, Touw 2001] 

Thuidium glabratum Broth. 

?Thuidium havaiense Reichardt 

Thuidium oahuense Broth. 

Thuidium plicatum Mitt. 

Thuidium plicatum Mitt. var. brevifolium 

E.B.Bartram 


end M H 





TIMMIA Hedw. (Timmiaceae) 


Timmia megapolitana Hedw. ssp. bavarica (Hessl.) 

Brassard [Brassard 1984] 

Timmia bavarica Hessl. 

ind H 

TORTELLA (Lindb.) Limpr. (Pottiaceae) 

Tortella fragilis (Hook. & Wilson) Limpr. var. 

tortelloides (S.W.Greene) R.H.Zander & Hoe [Zander 

& Hoe 1979] 

Sarconeurum tortelloides S.W.Greene 

Tortella tortelloides (S.W.Greene) H.Rob. 

ind M 

Tortella humilis (Hedw.) Jenn. [Hoe 1974] 

Tortella caespitosa (Schwägr.) Limpr. 

ind K O L M H 

Tortella tortuosa (Hedw.) Limpr. [Hoe 1979] ind L 

TORTULA Hedw. (Pottiaceae) 

Tortula atrovirens (Sm.) Lindb. [Hoe 1974, Hoe 1979, 

Corley et al. 1981, Zander 1993] 

Desmatodon convolutus (Brid.) Grout 

TRACHYLOMA Brid. (Pterobryaceae, or 

Trachylomataceae [Shaw & Goffinet 2000]) 

Trachyloma indicum Mitt. [Hoe 1974] 

Trachyloma tahitense Besch. 

TRACHYPODOPSIS M.Fleisch. (Trachypodaceae, or 

Meteoriaceae [Shaw & Goffinet 2000]) 

Trachypodopsis auriculata (Mitt.) M.Fleisch. [Hoe 1974] 

Trachypodopsis ornans (Reichardt) M.Fleisch. 

Trachypus ornans (Reichardt) Müll.Hal. fma. 

simplex Broth. 

Trachypodopsis serrulata (P.Beauv.) M.Fleisch. var. 

crispatula (Hook.) Zanten [Hoe 1979] 

TRACHYPUS Reinw. & Hornsch. (Trachypodaceae, or 

Meteoriaceae [Shaw & Goffinet 2000]) 

Trachypus bicolor Reinw. & Hornsch. [Hoe 1974, Hoe 

1979] 

Trachypus humilis Lindb. var. flagellifer (Broth. ex 

Herzog) H.A.Mill. [Zanten 1959, Miller 1967, Hoe 

1974] 

Trachypus humilis Lindb. var. tenerrimus (Herzog) 

Zanten 

Trachypus mauiensis Broth. 

Trachypus humilis Lindb. var. humilis [Hoe 1974, Hoe 

1977] 

TREMATODON Michx. (Dicranaceae, or Bruchiaceae 

[Shaw & Goffinet 2000]) 


ind M H 

ind M 


ind H 

ind K M H 

ind K O M 

ind K M 

63

64 



Trematodon latinervis Müll.Hal. [Hoe 1974] 

Trematodon longicollis misapplied 


TRICHOSTOMUM Bruch (Pottiaceae) 

Trichostomum brachydontium Bruch [Sollman 1984] 

Trichostomum oblongifolium E.B.Bartram 

Trichostomum crispulum Bruch [Hoe 1974, Hoe 1975, 

Norris & Koponen 1989] 

Trichostomum bartramii H.A.Mill. 

Trichostomum mauiense Broth. 



ULOTA D.Mohr (Orthotrichaceae) 

Ulota cervina Hoe & H.A.Crum [Hoe 1974] end M 

VESICULARIA (Müll.Hal.) Müll.Hal. (Hypnaceae) 

Vesicularia inflectens (Brid.) Müll.Hal. [Hoe 1979] ind M 

Vesicularia perviridis (Ångstr.) Müll.Hal. [Hoe 1974] 

Vesicularia graminicolor Müll.Hal. 


WEISSIA Hedw. (Pottiaceae) 

Weissia controversa Hedw. [Hoe 1974, Wichman & St. 

John 1990] 

Weisia viridula Hedw. 

ind Ni K O H 

Weissia ovalis (R.S.Williams) E.B.Bartram [Hoe 1974] end K O Mo L M H 

ZYGODON Hook. & Taylor (Orthotrichaceae) 

Zygodon reinwardtii (Hornsch.) A.Braun [Hoe 1974] ind M 

Zygodon rubrigemmius (Hoe & H.A.Crum) R.H.Zander 

& Vitt [Hoe 1974, Zander & Vitt 1979] 

Anoectangium rubrigemmium Hoe & H.A.Crum 

end M 

Zygodon tetragonostomus A.Braun ex B.S.G. [Hoe 1974] ind K O M H 



CROSS-REFERENCED LIST OF SYNONYMS AND MISAPPLIED NAMES 

Acroporium fuscoflavum var. baldwinii See Acroporium fuscoflavum 

?Acroporium sigmatodontium misapplied See Acroporium fuscoflavum 

Aerobryopsis longissima See Aerobryopsis wallichii 

Aerobryopsis longissima var. dozyana See Aerobryopsis wallichii 

Aerobryopsis scariosa See Aerobryopsis subdivergens ssp. scariosa 

Amphidium cyathicarpum See Amphidium tortuosum 

Andreaea acuminata See Andreaea acutifolia ssp. acuminata 

Andreaea perpapillosa See Andreaea rupestris 

Anoectangium euchloron See Anoectangium aestivum 

Anoectangium haleakalae var. haleakalae See Anoectangium aestivum 

Anoectangium haleakalae var. laxulum See Anoectangium aestivum 

Anoectangium harttiae See Anoectangium aestivum 

Anoectangium rubrigemmium See Zygodon rubrigemmius 

Aptychella robusta See Clastobryopsis robusta 

Barbella trichophora See Barbellopsis trichophora 

Barbula cruegeri See Barbula indica 

Barbula unguiculata misapplied See Barbula indica 

Barbula vinealis var. cylindrica See Didymodon insulanus 

Barbula vinealis var. flaccida See Didymodon insulanus 

Brachythecium oxyrrhynchum See Brachythecium plumosum 

Breutelia kilaueae See Breutelia affinis 

Brotherella opaeodon See Pylaisiadelpha opaeodon 

Bryohaplocladium microphyllum See Haplocladium microphyllum 

Bryum andicola Hook. See Rosulabryum billardierei 

Bryum apiculatum See Bryum mildeanum 

Bryum argenteum var. lanatum See Bryum argenteum 

Bryum capillare See Rosulabryum capillare 

Bryum ceramiocarpum See Bryum conicum 

Bryum crassicostatum See Bryum hawaiicum 

Bryum decaisnei See Rosulabryum billardierei 

Bryum erythrocarpum See Bryum atrovirens 

Bryum giganteum See Rhodobryum giganteum 

Bryum leucophyllum See Bryum argenteum 

Bryum limbatomarginatum See Rosulabryum billardierei 

Bryum mauiensis See Bryum mildeanum 

Bryum megalostegium See Bryum mildeanum 

Bryum nitens See Bryum mildeanum 

Bryum truncorum misapplied See Rosulabryum billardierei 

Bryum vinoviride See Rosulabryum capillare 

Calymperes hawaiiense See Calymperes fasciculatum 

Calymperes semiliberum See Syrrhopodon semiliber 

Calymperopsis semiliber See Syrrhopodon semiliber 

Campylopodium euphorocladum See Campylopodium medium 

Campylopus aureus See Campylopus schmidii ssp. schmidii 

Campylopus boswellii See Campylopus fragilis ssp. zollingerianus 

Campylopus boswellii var. capitulatus See Campylopus fragilis ssp. zollingerianus 

Campylopus densifolius var. densifolius See Campylopus hawaiicus var. densifolius 

Campylopus densifolius var. falcatus See Campylopus hawaiicus var. hawaiicoflexuosus 


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Campylopus densifolius var. hawaiicoflexuosus See Campylopus hawaiicus var. hawaiicoflexuosus 

Campylopus densifolius var. purpureoflavescens See Campylopus hawaiicus var. hawaiicus 

Campylopus densiretis See Campylopus hawaiicus var. densifolius 

?Campylopus didrichsenii See Campylopus umbellatus var. umbellatus 

Campylopus exasperatus See Campylopus praemorsus 

Campylopus fumarioli See Campylopus laxitextus 

Campylopus geniculatus See Campylopus umbellatus var. pterotoneuron 

Campylopus hawaiicoflexuosus See Campylopus hawaiicus var. hawaiicoflexuosus 

Campylopus introflexus misapplied See Campylopus schmidii ssp. schmidii 

Campylopus introflexus var. polytrichoides misapplied See Campylopus schmidii ssp. schmidii 

Campylopus microcephalus See Campylopus praemorsus 

Campylopus perangustifolius See Campylopus hawaiicus var. hawaiicoflexuosus 

?Campylopus pertristis See Campylopus umbellatus var. umbellatus 

Campylopus polytrichoides misapplied See Campylopus schmidii ssp. schmidii 

Campylopus purpureoflavescens, in part See Campylopus hawaiicus var. hawaiicoflexuosus, C. 

hawaiicus var. hawaiicus 

Campylopus richardii misapplied See Campylopus umbellatus var. umbellatus 

Campylopus sandwicensis See Campylopus umbellatus var. umbellatus 

Campylopus skottsbergii See Campylopus schmidii ssp. hemitrichius 

Campylopus tubulosus See Campylopus schmidii ssp. hemitrichius 

Campylopus wawreanus See Campylopus hawaiicus var. densifolius 

Ceratodon purpureus fma. xanthopus See Ceratodon purpureus ssp. purpureus 

Cirriphyllum oxyrrhynchum See Brachythecium plumosum 

Claopodium amblystegioides See Claopodium prionophyllum 

Claopodium hawaiiense See Claopodium whippleanum 

Claopodium nervosum See Claopodium prionophyllum 

Ctenidium decurrens See Ectropothecium decurrens 

Ctenidium pulcherrimum See Ectropothecium decurrens 

Ctenidium submalacobolum See Ectropothecium decurrens 

Cupressina decurrens See Ectropothecium decurrens 

Desmatodon convolutus See Tortula atrovirens 

Dicranella hillebrandii See Microdus hillebrandii 

Dicranella hochreuteri misapplied See Dicranella hochreutineri 

Dicranodontium falcatum var. falcatum See Dicranodontium porodictyon 

Dicranodontium hawaiicum See Dicranodontium porodictyon 

Dicranoloma wheeleri See Campylopus wheeleri 

Dicranum didrichsenii See Campylopus umbellatus var. umbellatus 

Dicranum hawaiicoflexuosum See Campylopus hawaiicus var. hawaiicoflexuosus 

Dicranum hawaiicum See Campylopus hawaiicus var. hawaiicus 

Dicranum microcephalum See Campylopus praemorsus 

Dicranum pertriste See Campylopus umbellatus var. umbellatus 

Dicranum praemorsum See Campylopus praemorsus 

Dicranum skottsbergii See Campylopus schmidii ssp. hemitrichius 

Dicranum wheeleri See Campylopus wheeleri 

Didymodon recurvirostris See Bryoerythrophyllum recurvirostre 

Didymodon vinealis var. flaccidus See Didymodon insulanus 

Ectropothecium rockii See Ectropothecium decurrens 

Encalypta rhaptocarpa misapplied See Encalypta rhabdocarpa 

Encalypta scabrata See Encalypta ciliata 

Entodon hillebrandii See Entodon plicatus 



Entodon solanderi See Entodon plicatus 

Eurhynchium celebicum See Rhynchostegium celebicum 

Eurhynchium muelleri misapplied See Platyhypnidium muelleri 

Eurhynchium selaginellifolium See Rhynchostegium selaginellifolium var. selaginellifolium 

Eurhynchium selaginellifolium var. recurvirameum See Rhynchostegium selaginellifolium var. 

recurvirameum 

Eurhynchium vagans See Oxyrrhynchium vagans 

Fissidens baldwinii See Fissidens elegans 

Fissidens bishopii See Fissidens elegans 

Fissidens crumii See Fissidens lindbergii 

Fissidens hawaiicus See Fissidens bryoides 

Fissidens insularis See Fissidens bryoides 

Fissidens intermedius See Fissidens submarginatus 

Fissidens mauiensis See Fissidens pacificus 

Fissidens oahuensis See Fissidens bryoides 

Floribundaria baldwinii See Floribundaria floribunda 

Funaria subintegra See Entosthodon subintegrus 

Garovaglia haleakalae See Garovaglia powellii var. haleakalae 

Glossadelphus abortivapicus See Symphyodon pygmaeus 

Glossadelphus baldwinii See Taxiphyllum laevifolium 

Glossadelphus chrysobasilaris See Glossadelphus ogatae 

Glossadelphus irroratus See Taxiphyllum torrentium 

Glossadelphus laevifolius See Taxiphyllum laevifolium 

Glossadelphus torrentium See Taxiphyllum torrentium 

Glossadelphus zollingeri var. filicaulis See Ectropothecium zollingeri var. filicaule 

Grimmia apocarpa See Schistidium apocarpum 

Grimmia apocarpa var. pulvinata See Grimmia anodon 

Grimmia apocarpa var. stricta See Schistidium strictum 

Grimmia haleakalae See Amphidium tortuosum 

Grimmia haliacalae See Amphidium tortuosum 

Grimmia pilfera See Grimmia longirostris 

Grimmia scabrifolia See Schistidium strictum 

Grimmia stricta See Schistidium strictum 

Gymnostoma pulvinata See Grimmia anodon 

Hageniella pacifica See Hageniella micans 

Holomitrium ferriei misapplied See Holomitrium squarrifolium 

Homalia densa See Homaliodendron flabellatum 

Homalia praelonga See Homaliodendron flabellatum 

Hookeria purpurea See Thamniopsis utacamundiana 

Hookeriopsis purpurea See Thamniopsis utacamundiana 

Hookeriopsis purpurea var. acuminatula See Thamniopsis utacamundiana 

Hookeriopsis purpurea var. ligulacea See Thamniopsis utacamundiana 

Hymenostylium firmum See Hymenostylium recurvirostrum var. recurvirostrum 

Hyophila dozy-molkenboeri See Hyophila involuta 

Hypnum decurrens See Ectropothecium decurrens 

Hypnum draytoni(i) See Buckiella draytonii 

Hypnum eudorae See Buckiella draytonii 

Hypnum opaeodon See Pylaisiadelpha opaeodon 

Hypnum wilkesianum See Palamocladium wilkesianum 

Isopterygium elegans See Pseudotaxiphyllum elegans 


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68 

Leptodontium brevicaule See Leptodontium flexifolium 

Leucobryum hawaiense See Leucobryum seemannii var. seemannii 

Leucobryum hawaiense var. fumarioli See Leucobryum seemannii var. fumarioli 

Leucobryum hawaiense var. solfatarae See Leucobryum seemannii var. solfatarae 

Leucobryum solfatarae See Leucobryum seemannii var. solfatarae 

Leucobryum solfatarae var. fumarioli See Leucobryum seemannii var. fumarioli 

Leucobryum solfatarae var. hawaiense See Leucobryum seemannii var. seemannii 

Macrocoma intricata See Macrocoma tenue ssp. sullivantii 

Macromitrium cumingii See Macromitirum incurvifolium 

Macromitrium intricatum See Macrocoma tenue ssp. sullivantii 

Macromitrium owahiense See Macromitrium microstomum 

Macromitrium reinwardtii See Macromitrium microstomum 

Merceyopsis crassinervis See Hymenostylium recurvirostrum var. cylindricum 

Meteorium pallidovirens See Palamocladium wilkesianum 

Meteorium sciuroides See Palamocladium wilkesianum 

Microdus hawaiicus See Dicranella hawaiica var. hawaiica 

Mielichhoferia campylocarpa See Schizymenium campylocarpum var. campylocarpum 

Mielichhoferia nealiae See Schizymenium nealiae 

Mielichhoferia pulvinata See Schizymenium pulvinatum 

Mniadelphus freycineti var. crasseturgescens See Distichophyllum freycinetii var. 

crasseturgescens 

Mnium rostratum See Plagiomnium rostratum 

Mnium serratum See Mnium marginatum 

Molendoa crassinervis See Hymenostylium recurvirostrum var. cylindricum 

Orthotrichum berggrenii See Orthotrichum alpestre var. berggrenii 

Orthotrichum hillebrandii See Orthotrichum rupestre 

Orthotrichum verrucatum See Orthotrichum lyellii 

Palamocladium aptychoides See Palamocladium wilkesianum 

Palamocladium sciurellum See Palamocladium wilkesianum 

Palamocladium wilkesianum var. altisetum See Palamocladium wilkesianum 

Palamocladium wilkesianum var. sciuroides See Palamocladium wilkesianum 

Philonotis laxissima See Philonotis hastata 

Plagiomnium prorepens See Plagiomnium rostratum 

Plagiomnium rhynchosporum See Plagiomnium rostratum 

Plagiopus longisetus See Plagiopus oederiana 

Plagiopus oederi See Plagiopus oederiana 

Plagiothecium draytonii See Buckiella draytonii 

Pleuropus aptychoides See Palamocladium wilkesianum 

Pleuropus pallidovirens See Palamocladium wilkesianum 

Pleuropus sciurellus See Palamocladium wilkesianum 

Pleuropus wilkesianus See Palamocladium wilkesianum 

Pleuropus wilkesianus var. sciuroides See Palamocladium wilkesianum 

Pogonatum baldwinii See Pogonatum tahitense 

Pohlia gracilescens See Pohlia flexuosa 

Pohlia gracillima See Pohlia flexuosa 

Pohlia leucostoma See Pohlia flexuosa 

Pohlia leucostomoides See Pohlia flexuosa 

Pseudosymblepharis mauiensis See Pseudosymblepharis angustata 

Ptychomitrium mauiense See Ptychomitrium lepidomitrium 

Ptychomnion aciculare misapplied See Aerobryopsis subdivergens ssp. scariosa 




Racomitrium lanuginosum var. sandvicense See Racomitrium lanuginosum var. pruinosum 

Rhabdoweisia denticulata See Rhabdoweisia crispata 

Rhacomitrium cuspidigerum misapplied See Racopilum cuspidigerum 

Rhizogonium spiniforme See Pyrrhobryum spiniforme 

Rhynchostegiella hawaica See Remyella hawaica 

Rhynchostegium vagans See Oxyrrhynchium vagans 

Sarconeurum tortelloides See Tortella fragilis var. tortelloides 

Sciaromium tricostatum See Limbella tricostata 

Sematophyllum caespitosum See Sematophyllum subpinnatum 

Sphagnum compactum misapplied See Sphagnum wheeleri 

Sphagnum henryense misapplied See Sphagnum palustre 

Sphagnum lonchocladum See Sphagnum palustre 

Sphagnum perichaetiale misapplied See Sphagnum palustre 

Sphagnum rigidulum See Sphagnum wheeleri 

Sphagnum vitjianum misapplied See Sphagnum palustre 

Sphagnum vulcanicum See Sphagnum wheeleri 

Stereodon decurrens See Ectropothecium decurrens 

Stereophyllum oahuense See Entodontopsis anceps 

Syrrhopodon kilaueae See Syrrhopodon prolifer 

Syrrhopodon oahuensis See Syrrhopodon armatus 

Taxiphyllum gracile See Ectropothecium decurrens 

Taxithelium undulatum misapplied See Taxithelium mundulum 

Thamnium speciosum See Thamnobryum speciosum 

Thuidium crenulatum See Pelekium versicolor 

Thuidium glabratum See Thuidium cymbifolium 

?Thuidium havaiense See Thuidium cymbifolium 

Thuidium nanophyllum See Pelekium versicolor 

Thuidium oahuense See Thuidium cymbifolium 

Thuidium plicatum See Thuidium cymbifolium 

Thuidium plicatum var. brevifolium See Thuidium cymbifolium 

Thysanomitrion hawaiicum See Campylopus umbellatus var. umbellatus 

Thysanomitrion umbellatum See Campylopus umbellatus var. umbellatus 

Timmia bavarica See Timmia megapolitana ssp. bavarica 

Tortella caespitosa See Tortella humilis 

Tortella tortelloides See Tortella fragilis var. tortelloides 

Tortula alpina var. inermis See Syntrichia fragilis 

Tortula fragilis See Syntrichia fragilis 

Tortula princeps See Syntrichia princeps 

Tortula rhizophylla See Chenia leptophylla 

Tortula vectensis See Chenia leptophylla 

Trachyloma tahitense See Trachyloma indicum 

Trachypodopsis ornans See Trachypodopsis auriculata 

Trachypus humilis var. tenerrimus See Trachypus humilis var. flagellifer 

Trachypus mauiensis See Trachypus humilis var. flagellifer 

Trachypus ornans fma. simplex See Trachypodopsis auriculata 

Trematodon longicollis misapplied See Trematodon latinervis 

Trichosteleum hamatum See Radulina hamata 

Trichostomum bartramii See Trichostomum crispulum 

Trichostomum mauiense See Trichostomum crispulum 

Trichostomum oblongifolium See Trichostomum brachydontium 


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Trichostomum tenuirostre See Oxystegus tenuirostris var. gemmiparus 

Vesicularia graminicolor See Vesicularia perviridis 

Webera baldwinii See Pohlia baldwinii 

Webera cruda See Pohlia cruda 

Webera gracilescens See Pohlia flexuosa 

Webera leucostomoides See Pohlia flexuosa 

Weisia viridula See Weissia controversa 

Weissia campylocarpa See Schizymenium campylocarpum var. campylocarpum

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