044.0000
Family
Thelypteridaceae Ching ex Pic.Serm.; Webbia 24: 711 (1970)
5-34 genera (depending on genus concept), 1100-1200 spp. New genus concept adopted here from Fawcett & Smith (2021), based on molecular results.
Species listed in database: 1180; hybrids: 38
044.2039
Genus
Sphaerostephanos J.Sm.; Hook. & Baker, Gen. Fil. t. 24 (1839)
Distribution: Tropical Africa, Asia, Pacific Islands.
Species listed in database: 192
= Mesochlaena R.Br.; Benn. & Br. Pl. Jav. Rar. 5 (1838) (nom. illeg.)
= Proferea C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6: 259 (1851)
= Thelypteris subgen.Glaphyropteridopsis sect.Neocyclosorus K.Iwats.; Mem. Coll. Sci. Univ. Kyoto, B, Biol. 31: 30 (1964) (quoad spec. typ.)
= Thelypteris subgen.Pneumatopteris sect.Macrocyclosorus K.Iwats.; (quoad spec. typ.)
= Thelypteris subgen.Sphaerostephanos K.Iwats.; Mem. Coll. Sci. Univ. Kyoto, B, Biol. 31: 32 (1964)
= Cyclosorus subgen.Sphaerostephanos (J.Sm.) Panigrahi; Abstr. & Souv. Nation. Symp. Curr. Trends Pterid. 4-6 October 1991: 13 (1991)
species :
Sphaerostephanos acrostichoides
(Desv.) Holttum; Fl. Males. Ser. 2,1(5): 456 (1982)
Distribution: C-Sulawesi; SE-Sulawesi; Philippines (Bohol, Mindanao, Guimaras); E-Java; Lesser Sunda Isl. (Sumba, Timor); Moluccas (Ambon); New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch. (Admiralty Isl.)
42 JAW LSI MOL NWG PHI SUL 43 BIS
Synonyms: hidden
= Aspidium riedleanum Gaudich.; Voy. Uranie [Freycinet] 326 (1827)
= Cyclodium acrostichoides (Desv.) J.Sm.; Lond. J. Bot. 1: 199 (1842), nomen
= Cyclosorus angustus (Copel.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 242 (1941)
= Cyclosorus boholensis Copel.; Philipp. J. Sci. 81: 31, pl. 23 (1952)
= Cyclosorus perpiliferus (Alderw.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 247 (1941)
= Cyclosorus reederi Copel.; Amer. Fern J. 43: 12 (1953)
= Cyclosorus riparius (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 143 (1947)
= Dryopteris angusta Copel.; Philipp. J. Sci., Bot. 9: 3 (1914)
= Dryopteris canescens subsp. acrostichoides (Desv.) C.Chr.; Index Filic. 256 (1905)
= Dryopteris perpilifera Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 11: 12 (1913)
= Dryopteris pseudoarbuscula Alderw.; Philipp. J. Sci. 11: 106 (1916)
= Dryopteris riedleana (Gaudich.) Alderw.; Handb. 230 (1908)
= Dryopteris riparia Copel.; Univ. Calif. Publ. Bot. 18: 221 (1942)
= Nephrodium acrostichoides (Desv.) J.Sm.; J. Bot. 8: 411 (1841), nomen
= Nephrodium acrostichoides Desv.; Mém. Soc. Linn. Paris 6: 255 (1827) (non Thelypteris acrostichoides (Michx.) Newl.)
= Nephrodium riedleanum (Gaudich.) T.Moore; Index Filic. (T. Moore) 102 (1858)
= Nephrodium smithianum C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6: 46 (1851)
= Sphaerostephanos smithianus (C.Presl) Holttum; Kalikasan 4: 65 (1975)
= Thelypteris angusta (Copel.) C.F.Reed; Phytologia 17: 259 (1968)
= Thelypteris boholensis (Copel.) C.F.Reed; Phytologia 17: 204 (1968)
= Thelypteris draco Christenh.; Global Fl. 1(4): 29 (2018), nom. nov. for Nephrodrium acrostichoides Desv.
= Thelypteris perpilifera (Alderw.) C.F.Reed; Phytologia 17 (1968), 303
= Thelypteris reederi (Copel.) C.F.Reed; Phytologia 17: 308 (1968)
= Thelypteris riparia (Copel.) C.F.Reed; Phytologia 17: 309 (1968)
species :
Sphaerostephanos adenostegius
(Copel.) Holttum; Fl. Males. 1: 466 (1981)
Distribution: Sulawesi; New Guinea (Irian Jaya, Papua New Guinea)
42 NWG SUL
Synonyms: hidden
= Cyclosorus adenostegius (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 142 (1947)
= Dryopteris adenostegia Copel.; Univ. Calif. Publ. Bot. 18: 220 (1942)
= Pronephrium nothofageti Holttum; Blumea 20: 118 (1972)
= Thelypteris adenostegia (Copel.) C.F.Reed; Phytologia 17: 258 (1968)
species :
Sphaerostephanos alatellus
(Christ) Holttum; Fl. Males. Ser. 2,1(5): 451 (1982)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Aspidium alatellum (Christ) Christ; Bull. Herb. Boissier, sér, 2, 1: 454 (1901)
= Cyclosorus alatellus (Christ) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 141 (1947)
= Dryopteris alatella (Christ) C.Chr.; Index Filic. 251 (1905)
= Nephrodium alatellum Christ; K. Schum. & Lauterb., Fl. Schutzgeb. Südsee 112 (1900)
= Thelypteris alatella (Christ) C.F.Reed; Phytologia 17: 259 (1968)
species :
Sphaerostephanos albosetosus
(Copel.) Holttum; Fl. Males. Ser. 2,1(5): 505 (1982)
Distribution: New Guinea (Irian Jaya: near Lake Habbema)
42 NWG
Conservation Status: Only known from type
Synonyms: hidden
= Cyclosorus albosetosus (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 142 (1947)
= Dryopteris albosetosa Copel.; Univ. Calif. Publ. Bot. 18: 221 (1942)
= Thelypteris albosetosa (Copel.) C.F.Reed; Phytologia 17: 259 (1968)
species :
Sphaerostephanos alcasidii
Holttum; Fl. Males. Ser. 2,1(5): 503 (1982)
Distribution: Philippines (Luzon, Mt. Data)
42 PHI
Conservation Status: Only known from type
Synonyms: hidden
= Thelypteris alcasidii (Holttum) Christenh.; Global Fl. 1(4): 25 (2018)
species :
Sphaerostephanos alpinus
Holttum; Fl. Males. Ser. 2,1(5): 488 (1982)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Thelypteris alpina (Holttum) Christenh.; Global Fl. 1(4): 25 (2018)
species :
Sphaerostephanos alticola
Holttum; Fl. Males. Ser. 2,1(5): 484 (1982)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Conservation Status: Only known from 2 collections
Synonyms: hidden
= Thelypteris alticola (Holttum) Christenh.; Global Fl. 1(4): 25 (2018)
species :
Sphaerostephanos angustibasis
Holttum; Fl. Males. Ser. 2,1(5): 476 (1982)
Distribution: Moluccas (Tidore)
42 MOL
Conservation Status: Only known from type
Synonyms: hidden
= Thelypteris angustibasis (Holttum) Christenh.; Global Fl. 1(4): 27 (2018)
species :
Sphaerostephanos angustifolius
(C.Presl) Holttum; Kalikasan 4: 66 (1975)
Distribution: Philippines
42 PHI
Synonyms: hidden
= Aspidium angustifolium (C.Presl) Mett.; Salomon, Nom. 43 (1883)
= Cyclosorus angustifolius (C.Presl) Copel.; Fern Fl. Philip. 2: 349 (1960)
= Nephrodium angustifolium C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 48 (1851) (non Thelypteris angustifolia (Willd.) Proctor)
= Nephrodium mucronatum J.Sm.; J. Bot. 8: 412 (1841) (p. p.)
= Thelypteris isildurii Christenh.; Global Fl. 1(4): 32 (2018), nom. nov. superfl.
= Thelypteris setulosa C.F.Reed; Phytologia 17: 313 (1968), nom. nov.
species :
Sphaerostephanos appendiculatus
(Blume) Holttum; Kalikasan 4: 62 (1975)
Distribution: Sumatra; N-Sulawesi; SW-Sulawesi; Philippines (Luzon); Java; Lesser Sunda Isl. (Bali, Flores)
42 JAW LSI PHI SUL SUM
Synonyms: hidden
= Aspidium pilosiusculum Mett.; Ann. Mus. Bot. Lugduno-Batavi 1: 233 (1863), nom. nov.
= Cyclosorus appendiculatus (Blume) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 242 (1941)
= Cyclosorus brunnescens (C.Chr. ex Kjellb. & C.Chr.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 243 (1941)
= Dryopteris appendiculata (Blume) C.Chr.; Index Filic. 252 (1905)
= Dryopteris brunnescens C.Chr.; Bot. Jahrb. Syst. 66: 44 (1933)
= Dryopteris perakensis var. sumatrensis Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 11: 13 (1913)
= Goniopteris appendiculata (Blume) C.Presl; Tent. Pterid. I: 7, f. 11 (1836)
= Gymnogramma appendiculata Blume; Enum. Pl. Javae 2: addendum to p. 112 (1828)
= Nephrodium appendiculatum (Blume) Racib.; Fl. Buitenz. 1: 186 (1898)
= Phegopteris appendiculata (Blume) Mett. ex Salomon; Nomencl. Gefäßkrypt. 273 (1883)
= Thelypteris appendiculata (Blume) C.F.Reed; Phytologia 17: 260 (1968)
= Thelypteris brunnescens (C.Chr. ex Kjellb. & C.Chr.) C.F.Reed; Phytologia 17: 265 (1968)
species :
Sphaerostephanos aquatilis
(Copel.) Holttum; Fl. Males. Ser. 2,1(5): 466 (1982)
Distribution: New Guinea (SE-Papua New Guinea)
42 NWG
Conservation Status: Only known from 2 collections
Synonyms: hidden
= Cyclosorus aquatilis (Copel.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 242 (1941)
= Dryopteris aquatilis Copel.; Philipp. J. Sci., Bot. 6: 75 (1911)
= Dryopteris caudiculata Rosenst.; Repert. Spec. Nov. Regni Veg. 9: 426 (1911) (non Alderw. (1909))
= Thelypteris aquatilis (Copel.) C.F.Reed; Phytologia 17: 260 (1968)
species :
Sphaerostephanos arbuscula
(Willd.) Holttum; Nayar & Kaur, Comp. to Bedd. Handb. Ferns Brit. India 207 (1974)
Distribution: Kenya; Tanzania; Madagascar; Mauritius; La Réunion; India (Kerala, ?Madhya Pradesh, Tamil Nadu); Sri Lanka; ?New Caledonia
25 KEN TAN 29 MAU MDG REU 40 IND SRL
Synonyms: hidden
= Aspidium arbuscula Willd.; Linnaea Sp. Pl. 5: 233 (1810)
= Aspidium hookeri Wall.; Numer. List no. 64 (1829), nomen
= Aspidium hookeri Wall. ex Hook.; Hooker´s Icon. Pl. 6: 10, pl. 992 (1854), non Sw. 1830
= Cyclosorus arbuscula (Willd.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 194 (1938)
= Dryopteris arbuscula (Willd.) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Nephrodium arbuscula (Willd.) Desv.; Mem. Soc. Linn. Paris 6, 2: 253 (1827)
= Nephrodium auriculatum Fée; Mém. Foug. 10 (Icon. Esp. Nouv.): 42, t. 42, f. 2 (1865)
= Nephrodium falcilobum var. puberum C.B.Clarke; Trans. Linn. Soc. Lond., 2 Bot., 1: 515 (1880)
= Nephrodium hookeri Wall. ex T.Moore & Houlston; Gard. Mag. Bot. 3 (1851)
= Nephrodium puberum J.Sm.; Cat. Cult. Ferns 54 (1857)
= Sphaerostephanos arbuscula subsp. africanus Holttum; J. S. Afr. Bot. 40: 165 (1974)
= Thelypteris arbuscula (Willd.) K.Iwats.; Acta Phytotax. Geobot. 21: 170 (1965)
species :
Sphaerostephanos archboldii
(C.Chr.) Holttum; Fl. Males. Ser. 2,1(5): 485 (1982)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Cyclosorus archboldii (C.Chr.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 142 (1947)
= Cyclosorus protectus (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 143 (1947)
= Dryopteris archboldii C.Chr.; Brittonia 2: 297 (1937)
= Dryopteris protecta Copel.; Univ. Calif. Publ. Bot. 18: 221 (1942)
= Thelypteris archboldii (C.Chr.) C.F.Reed; Phytologia 17: 260 (1968)
= Thelypteris protecta (Copel.) C.F.Reed; Phytologia 17: 306 (1968)
species :
Sphaerostephanos arfakianus
(Baker) Holttum; Fl. Males. 1: 484 (1981)
Distribution: Moluccas (Seram, ?Ambon); New Guinea (Irian Jaya, Papua New Guinea)
42 MOL NWG
Synonyms: hidden
= Cyclosorus arfakianus (Baker) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 142 (1947)
= Dryopteris arborea Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 28: 24 (1918) (non Brause (1914))
= Dryopteris arfakiana (Baker) C.Chr.; Index Filic. 253 (1905)
= Dryopteris rosenburghii C.Chr.; Index Filic., Suppl. Tert.: 96 (1934)
= Dryopteris sepikensis Brause; Bot. Jahrb. Syst. 56: 101 (1920)
= Phegopteris arfakiana (Baker) Alderw.; Handb. 502 (1908)
= Polypodium arfakianum Baker; Beccari, Malesia 3: 45 (1880)
= Thelypteris arfakiana (Baker) C.F.Reed; Phytologia 17: 260 (1968)
species :
Sphaerostephanos atasripii
(Rosenst.) Holttum; Webbia 30: 194 (1976)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Synonyms: hidden
= Cyclosorus atasripii (Rosenst.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 142 (1947)
= Cyclosorus deminuens Holttum; Blumea 13: 135 (1965)
= Dryopteris atasripii Rosenst.; Meded. Rijks Herb. no. 31: 6 (1917)
= Thelypteris atasripii (Rosenst.) C.F.Reed; Phytologia 17: 261 (1968)
species :
Sphaerostephanos austerus
(Brause) Holttum; Fl. Males. Ser. 2,1(5): 502 (1982)
Distribution: New Guinea (Papua New Guinea: Sepik)
42 NWG
Conservation Status: Only known from type
Synonyms: hidden
= Cyclosorus austerus (Brause) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 243 (1941)
= Dryopteris austera Brause; Bot. Jahrb. Syst. 56: 108 (1920)
= Thelypteris austera (Brause) C.F.Reed; Phytologia 17: 262 (1968)
species :
Sphaerostephanos bakeri
(Harr.) S.E.Fawc. & A.R.Sm.; A generic revision of the Thelypteridaceae: 81 (2021)
Distribution: Philippines (Luzon, Panay, Negros Oriental, Mindanao)
42 PHI
Synonyms: hidden
= Cyclosorus bakeri (Harr.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 142 (1947)
= Dryopteris bakeri (Harr.) Copel.; Philipp. J. Sci., Bot. 2: 405 (1907)
= Dryopteris canescens subsp. bakeri (Harr.) C.Chr.; Index Filic. 256 (1906)
= Haplodictyum bakeri (Harr.) Ching; Sunyatsenia 5: 251 (1940)
= Nephrodium bakeri Harr.; J. Linn. Soc., Bot. 16: 29 (1877)
= Pronephrium bakeri (Harr.) Holttum; Fl. Males. Ser. 2,1(5): 516 (1982)
= Thelypteris bakeri (Harr.) C.F.Reed; Phytologia 17: 262 (1968)
species :
Sphaerostephanos baramensis
(C.Chr.) Holttum; Fl. Males. 1: 473 (1981)
Distribution: Borneo (N-Sarawak, E-Kalimantan, Mt. Kinabalu, Sabah)
42 BOR
Synonyms: hidden
= Dryopteris baramensis C.Chr.; Gard. Bull. Straits Settlem. 7: 246 (1934)
= Thelypteris baramensis (C.Chr.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 6: 287 (1936)
species :
Sphaerostephanos batacorum
(Rosenst.) Holttum; Fl. Males. 1: 476 (1981)
Distribution: Sumatra
42 SUM
Synonyms: hidden
= Cyclosorus batacorum (Rosenst.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 243 (1941)
= Dryopteris batacorum Rosenst.; Repert. Spec. Nov. Regni Veg. 13: 217 (1914), excl. var. winkleri
= Dryopteris stipellata var. obtusata Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 3, 2: 151 (1920)
= Thelypteris batacorum (Rosenst.) C.F.Reed; Phytologia 17: 263 (1968)
species :
Sphaerostephanos batjanensis
(Rosenst.) Holttum; Fl. Males. 1: 467 (1981)
Distribution: N-Sulawesi; Moluccas (Bacan, Seram)
42 MOL SUL
Synonyms: hidden
= Aspidium canescens f. gymnogrammoides Christ; Ann. Jard. Bot. Buitenz. I, 5: 131 (1898)
= Cyclosorus batjanensis (Rosenst.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 243 (1941)
= Dryopteris batjanensis Rosenst.; Meded. Rijks Herb. no. 31: 5 (1917)
= Thelypteris batjanensis (Rosenst.) C.F.Reed; Phytologia 17: 263 (1968)
species :
Sphaerostephanos batulantensis
Holttum; Fl. Males. Ser. 2,1(5): 473 (1982)
Distribution: Lesser Sunda Isl. (W-Sumbawa)
42 LSI
Conservation Status: Only known from type
Synonyms: hidden
= Thelypteris batulantensis (Holttum) Christenh.; Global Fl. 1(4): 27 (2018)
species :
Sphaerostephanos beccarianus
(Ces.) S.E.Fawc. & A.R.Sm.; A generic revision of the Thelypteridaceae: 81 (2021)
Distribution: ?Moluccas (?Seram); New Guinea (Irian Jaya, Papua New Guinea); Solomon Isl. (Bougainville, Kolombangara, Makira, Santa Ysabel); Vanuatu; Fiji (Viti Levu, Vanua Levu, Taveuni)
42 NWG 43 SOL 60 FIJ VAN
Synonyms: hidden
= Cyclosorus beccarianus (Ces.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 142 (1947)
= Dryopteris cesatiana C.Chr.; Index Filic. 257 (1905), nom. nov.
= Dryopteris oblanceolata Copel.; Philipp. J. Sci., Bot. 9: 3 (1914)
= Goniopteris simplicifolia var. viliensis Carruth.; Seem., Fl. Vit. 366 (1893)
= Meniscium beccarianum Ces.; Rend. Acad. Napoli 16: 27, 30 (1877) (non Nephrodium beccarianum Ces. (1876))
= Phegopteris beccariana (Ces.) Alderw.; Handb. 509 (1908)
= Phegopteris oblanceolata (Copel.) Alderw.; Malayan Ferns Fern Allies 320 (1917)
= Phegopteris rutteniana Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 28: 36 (1918)
= Pronephrium beccarianum (Ces.) Holttum; Blumea 20: 107 (1972)
= Thelypteris cesatiana (C.Chr.) C.F.Reed; Phytologia 17: 267 (1968)
species :
Sphaerostephanos benoiteanus
(Gaudich.) Holttum; Fl. Males. Ser. 2,1(5): 500 (1982)
Distribution: New Guinea (Irian Jaya: Rawak Isl.)
42 NWG
Conservation Status: Only known from type
Synonyms: hidden
= Aspidium benoiteanum Gaudich.; Voy. Uranie [Freycinet] (331), t. 11 (1827), as ´benoitianum´, nom. nud. pro syn.
= Dryopteris benoiteana (Gaudich.) Alderw.; Malayan Ferns 225 (1909)
= Nephrodium benoiteanum (Gaudich.) C.Presl; Tent. Pterid. 81 (1836)
= Polystichum benoiteanum Gaudich.; Voy. Uranie [Freycinet] 331 (1827)
= Thelypteris benoiteana (Gaudich.) Christenh.; Global Fl. 1(4): 27 (2018)
species :
Sphaerostephanos braithwaitei
Holttum; Allertonia 1(3): 206 (1977)
Distribution: Solomon Isl. (Bougainville, Gizo, Guadalcanal, Santa Ysabel, Kolombangara, Makira, Rendova, Santa Ysabel)
43 SOL
Synonyms: hidden
= Thelypteris braithwaitei (Holttum) Christenh.; Global Fl. 1(4): 28 (2018)
species :
Sphaerostephanos brauseanus
(Holttum) S.E.Fawc. & A.R.Sm.; A generic revision of the Thelypteridaceae: 81 (2021)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Dryopteris canescens var. novoguineensis Brause; Bot. Jahrb. Syst. 49: 22 (1912)
= Pronephrium brauseanum Holttum; Blumea 20: 107 (1972)
= Thelypteris brauseanum (Holttum) Christenh.; Global Fl. 1(4): 28 (2018)
species :
Sphaerostephanos bulusanicus
(Holttum) S.E.Fawc. & A.R.Sm.; A generic revision of the Thelypteridaceae: 81 (2021)
Distribution: Philippines (Luzon)
42 PHI
Conservation Status: IUCN R
Synonyms: hidden
= Haplodictyum bulusanicum Holttum; Kalikasan 2: 61 (1973)
= Pronephrium bulusanicum (Holttum) Holttum; Fl. Males. Ser. 2,1(5): 515 (1982)
= Thelypteris bulusanica (Holttum) Christenh.; Global Fl. 1(4): 28 (2018)
species :
Sphaerostephanos canescens
(Blume) Holttum; Fl. Males. Ser. 2,1(5): 497 (1982)
Distribution: N-Sulawesi; W-Java; Moluccas (Tidore, Halmaheira)
42 JAW MOL SUL
Synonyms: hidden
= Aspidium canescens Christ; Ann. Jard. Bot. Buitenz. 16: 130 (1897)
= Aspidium diversilobum Mett.; Pheg. u. Asp. 100, n. 239 (1858)
= Cyclosorus canescens (Blume) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 141 (1947)
= Dryopteris blumei (J.Sm.) Alderw.; Malayan Ferns 231 (1909)
= Dryopteris canescens (Blume) C.Chr.; Index Filic. 250 (1905) (quoad typus tantum)
= Goniopteris canescens (Blume) C.Presl; Tent. Pterid. 183 (1836)
= Gymnogramma canescens (Blume) Blume; Fl. Javae, Fil. 93, t. 40 (1829)
= Haplodictyum canescens (Blume) Ching; Sunyatsenia 5: 251 (1940)
= Nephrodium blumei J.Sm.; J. Bot. 8: 411 (1841) (non Hook. (1862))
= Nephrodium canescens (Blume) Christ; Verh. Naturf. Ges. Basel 11: 208 (1895)
= Nephrodium subdimorphum Christ; Verh. Naturf. Ges. Basel 11: 16 (1894)
= Phegopteris canescens (Blume) Mett.; Ann. Mus. Bot. Lugduno-Batavi 1: 223 (1863) (non Mett. 1858)
= Pleocnemia blumei (J.Sm.) T.Moore; Index Filic. (T. Moore) 87 (1857)
= Polypodium canescens Blume; Enum. Pl. Javae 2: 133 (1828)
= Thelypteris canescens (Blume) K.Iwats.; Mem. Coll. Sci. Univ. Kyoto, B, Biol. 31: 34 (1964)
species :
Sphaerostephanos carrii
Holttum; Fl. Males. Ser. 2,1(5): 463 (1982)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Thelypteris carrii (Holttum) Christenh.; Global Fl. 1(4): 28 (2018)
species :
Sphaerostephanos cartilagidens
P.M.Zamora & Co; Nat. Appl. Sci. Bull. Univ. Philipp. 32(1-4): 43 (1980)
Distribution: Palawan; Philippines
42 PHI PLW
Synonyms: hidden
= Thelypteris cartilagidens (P.M.Zamora & Co) Christenh.; Global Fl. 1(4): 28 (2018)
species :
Sphaerostephanos castaneus
(A.R.Sm. & Lorence) A.R.Sm. & Lorence; Fl. Marquesas Isl. 1: 213 (2019)
Distribution: Marquesas Isl. (Nuku Hiva)
61 MRQ
Synonyms: hidden
= Cyclosorus castaneus A.R.Sm. & Lorence; PhytoKeys 4: 39 (-40, fig. 11) (2011)
= Thelypteris minasmorgulensis Christenh.; Global Fl. 1(4): 33 (2018), nom. nov.
species :
Sphaerostephanos cataractorum
(W.H.Wagner & Grether) Holttum; Fl. Males. Ser. 2,1(5): 468 (1982)
Distribution: New Guinea (Papua New Guinea); Bismarck Arch. (Admiralty Isl., New Britain); Solomon Isl. (Bougainville, Santa Ysabel)
42 NWG 43 BIS SOL
Synonyms: hidden
= Cyclosorus cataractorum W.H.Wagner & Grether; Univ. Calif. Publ. Bot. 23: 50, t. 16 (1948)
= Thelypteris cataractorum (W.H.Wagner & Grether) C.F.Reed; Phytologia 17: 266 (1968)
species :
Sphaerostephanos caulescens
Holttum; Fl. Males. Ser. 2,1(5): 472 (1982)
Distribution: Borneo (N-Sarawak, Mt. Kinabalu, Sabah); N-Sulawesi; Moluccas (Bacan)
42 BOR MOL SUL
Synonyms: hidden
= Dryopteris porphyricola C.Chr.; Gard. Bull. Straits Settlem. 7: 244 (1934) (non Copel.)
= Thelypteris caulescens (Holttum) Christenh.; Global Fl. 1(4): 28 (2018)
species :
Sphaerostephanos confertus
(Brause) Holttum; Webbia 30: 194 (1976)
Distribution: New Guinea (Irian Jaya, Papua New Guinea, Waigeu Isl.)
42 NWG
Synonyms: hidden
= Cyclosorus confertus (Brause) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 149 (1947)
= Cyclosorus terrestris (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 143 (1947)
= Dryopteris conferta Brause; Bot. Jahrb. Syst. 49: 22 (1912)
= Dryopteris terrestris Copel.; Univ. Calif. Publ. Bot. 18: 221 (1942)
= Thelypteris conferta (Brause) C.F.Reed; Phytologia 17: 268 (1968)
= Thelypteris terrestris (Copel.) C.F.Reed; Phytologia 17: 319 (1968)
species :
Sphaerostephanos convergens
Holttum; Fl. Males. Ser. 2,1(5): 477 (1982)
Distribution: Philippines (W-Samar)
42 PHI
Conservation Status: Only known from type
Synonyms: hidden
= Thelypteris convergens (Holttum) Christenh.; Global Fl. 1(4): 28 (2018)
species :
Sphaerostephanos cyrtocaulos
(Alderw.) Holttum; Fl. Males. Ser. 2,1(5): 473 (1982)
Distribution: Sumatra (near G. Kerinci)
42 SUM
Synonyms: hidden
= Cyclosorus cyrtocaulos (Alderw.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 243 (1941)
= Dryopteris cyrtocaulos Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 3, 5: 201 (1922)
= Thelypteris cyrtocaulos (Alderw.) C.F.Reed; Phytologia 17: 270 (1968)
species :
Sphaerostephanos daymanianus
Holttum; Fl. Males. Ser. 2,1(5): 451 (1982)
Distribution: New Guinea (Papua New Guinea: Milne Bay, Morobe Districts); Solomon Isl. (Bougainville)
42 NWG 43 SOL
Synonyms: hidden
= Thelypteris daymaniana (Holttum) Christenh.; Global Fl. 1(4): 29 (2018)
species :
Sphaerostephanos debilis
(Mett.) Holttum; Fl. Males. Ser. 2,1(5): 448 (1982)
Distribution: Moluccas (Ambon); New Guinea (Irian Jaya)
42 MOL NWG
Synonyms: hidden
= Cyclosorus debilis (Mett.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 142 (1947) (non (Baker) Ching (1941))
= Dryopteris debilis (Mett.) C.Chr.; Index Filic. 260 (1905)
= Nephrodium debile (Mett.) Diels; Engl. & Prantl, Nat. Pflanzenfam. 1(4): 178 (1899)
= Nephrodium warburgii Diels; K. Schum. & Lauterb., Fl. Schutzgeb. Südsee 112 (1901)
= Phegopteris debilis Mett.; Ann. Mus. Bot. Lugduno-Batavi 1: 123, t. 6, f. 1 (1863)
= Polypodium debile (Mett.) Baker; Hook. & Baker, Syn. Fil. 313 (1867)
= Thelypteris debilis (Mett.) C.F.Reed; Phytologia 17: 270 (1968)
species :
Sphaerostephanos decadens
(Baker) Holttum; Allertonia 1(3): 209 (1977)
Distribution: Vanuatu; Tonga (´Eua); Fiji (Viti Levu, Vanua Levu); Wallis & Futuna (Futuna Isl.); ?[Western] Samoa
60 FIJ TON VAN WAL
Synonyms: hidden
= Cyclosorus decadens (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 244 (1941)
= Cyclosorus degeneri Copel.; J. Arnold Arbor. 30: 438 (1949)
= Cyclosorus orthocaulis K.Iwats.; Amer. Fern J. 53: 135 (1963)
= Dryopteris decadens (Baker) C.Chr.; Index Filic. 260 (1905)
= Nephrodium decadens Baker; J. Bot. 24: 183 (1886)
= Thelypteris decadens (Baker) C.F.Reed; Phytologia 17: 270 (1968)
= Thelypteris degeneri (Copel.) C.F.Reed; Phytologia 17: 271 (1968)
= Thelypteris orthocaulis (K.Iwats.) C.F.Reed; Phytologia 17: 299 (1968)
species :
Sphaerostephanos dichrotrichoides
(Alderw.) Holttum; Kalikasan 4: 65 (1975)
Distribution: Philippines (Mindanao, Panay?)
42 PHI
Synonyms: hidden
= Cyclosorus dichrotrichoides (Alderw.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 244 (1941)
= Dryopteris dichrotricha Copel.; Philipp. J. Sci., Bot. 7: 54 (1912) (non Copel. (1911))
= Dryopteris dichrotrichoides Alderw.; Malayan Ferns Fern Allies, Corr. 48 (1917), nom. nov.
= Dryopteris weberi Copel.; Philipp. J. Sci.38: 135 (1929) (nom. nov. superfl.)
= Lastrea dichrotrichoides (Alderw.) Copel.; Fern Fl. Philip. 329 (1960)
= Lastrea weberi (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] [Copeland] 140 (1947)
= Thelypteris dichrotrichoides (Alderw.) C.F.Reed; Phytologia 17: 272 (1968)
species :
Sphaerostephanos dichrotrichus
(Copel.) Holttum; Fl. Males. Ser. 2,1(5): 453 (1982)
Distribution: New Guinea (Papua New Guinea); Bismarck Arch. (Admiralty Isl.)
42 NWG 43 BIS
Synonyms: hidden
= Cyclosorus dichrotrichus (Copel.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 244 (1941)
= Dryopteris dichrotricha Copel.; Philipp. J. Sci., Bot. 6: 74 (1911)
= Thelypteris dichrotricha (Copel.) C.F.Reed; Phytologia 17: 272 (1968)
species :
Sphaerostephanos dimidiolobatus
Holttum; Fl. Males. Ser. 2,1(5): 459 (1982)
Distribution: Bismarck Arch. (S-New Ireland)
43 BIS
Conservation Status: Only known from 2 collections
Synonyms: hidden
= Thelypteris dimidiolobata (Holttum) Christenh.; Global Fl. 1(4): 29 (2018)
species :
Sphaerostephanos diminutus
(Copel.) M.G.Price; Contr. Univ. Michigan Herb. 17: 277 (1990)
Distribution: Philippines (Luzon, Mindanao)
42 PHI
Synonyms: hidden
= Cyclosorus diminutus (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 142 (1947)
= Dryopteris diminuta Copel.; Philipp. J. Sci. 40: 298 (1929)
= Pronephrium diminutum (Copel.) Holttum; Blumea 21: 115 (1972)
= Sphaerostephanos lucbanii Holttum; Fl. Males. Ser. 2,1(5): 463 (1982)
= Thelypteris lucbanii (Holttum) Christenh.; Global Fl. 1(4): 33 (2018)
= Thelypteris diminuta (Copel.) C.F.Reed; Phytologia 17: 273 (1968)
= Thelypteris diminuta (Copel.) Christenh.; Global Fl. 1(4): 29 (2018), isonym
species :
Sphaerostephanos dimorphus
(Brause) Holttum; Fl. Males. Ser. 2,1(5): 501 (1982)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Cyclosorus dimorphus (Brause) Copel.; Philipp. J. Sci. 78: 442 (1951)
= Cyclosorus morobensis (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 143 (1947)
= Dryopteris dimorpha Brause; Bot. Jahrb. Syst. 56: 100 (1920)
= Dryopteris morobensis Copel.; Univ. Calif. Publ. Bot. 18: 221 (1942)
= Thelypteris dimorpha (Brause) C.F.Reed; Phytologia 17: 273 (1968)
= Thelypteris morobensis (Copel.) C.F.Reed; Phytologia 17: 294 (1968)
species :
Sphaerostephanos diversilobus
(C.Presl) Holttum; Kalikasan 4: 57 (1975)
Distribution: Sulawesi; Philippines (Luzon, Panay)
42 PHI SUL
Synonyms: hidden
= Cyclosorus asymetricus (Fée) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 142 (1947)
= Cyclosorus diversilobus (C.Presl) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 244 (1941)
= Dryopteris acromanes Christ; Philipp. J. Sci., Bot. 2: 200 (1907)
= Dryopteris asymetrica (Fée) Christ; Geogr. d. Farne 226 (1910)
= Dryopteris diversiloba (C.Presl) Christ; Philipp. J. Sci., Bot. 2: 199 (1907), nomen
= Goniopteris asymetrica Fée; Mém. Foug. 5 [Gen. Filic.] 253 (1852)
= Nephrodium diversilobum C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6: 47 (1851)
= Phegopteris acromanes (Christ) Alderw.; Handb. 505 (1908)
= Phegopteris diversiloba (C.Presl) Alderw.; Handb. 503 (1908)
= Pronephrium acrostichoides C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 259 (1851)
= Thelypteris asymetrica (Fée) C.F.Reed; Phytologia 17: 261 (1968)
= Thelypteris diversiloba (C.Presl) C.F.Reed; Phytologia 17: 273 (1968)
= Thelypteris diversiloba (C.Presl) Christenh.; Global Fl. 1(4): 29 (2018), isonym
species :
Sphaerostephanos doodioides
(Copel.) Holttum; Allertonia 1(3): 203 (1977)
Distribution: Solomon Isl. (Guadalcanal, Makira); New Caledonia; Vanuatu (Espírito Santo)
43 SOL 60 NWC VAN
Synonyms: hidden
= Cyclosorus doodioides (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 142 (1947)
= Dryopteris doodioides Copel.; Philipp. J. Sci. 60: 107, t. 11 (1936)
= Thelypteris doodioides (Copel.) C.F.Reed; Phytologia 17: 273 (1968)
species :
Sphaerostephanos echinosporus
(Alderw.) Holttum; Fl. Males. Ser. 2,1(5): 495 (1982)
Distribution: S-Sumatra
42 SUM
Conservation Status: Only known from type
Synonyms: hidden
= Cyclosorus echinosporus (Alderw.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 244 (1941)
= Dryopteris echinospora Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 3, 2: 149 (1920)
= Thelypteris echinospora (Alderw.) C.F.Reed; Phytologia 17: 274 (1968)
species :
Sphaerostephanos efogensis
Holttum; Fl. Males. Ser. 2,1(5): 482 (1982)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Thelypteris efogensis (Holttum) Christenh.; Global Fl. 1(4): 29 (2018)
species :
Sphaerostephanos ekutiensis
Holttum; Fl. Males. Ser. 2,1(5): 483 (1982)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Conservation Status: Only known from type
Synonyms: hidden
= Thelypteris ekutiensis (Holttum) Christenh.; Global Fl. 1(4): 29 (2018)
species :
Sphaerostephanos elatus
(Bojer) Holttum; J. S. Afr. Bot. 40: 167 (1974)
subspecies :
subsp. elatus
Distribution: Madagascar; Mauritius; La Réunion; Rodrigues; Seychelles
29 MAU MDG REU ROD SEY
Remarks: probably in a new genus as sister to Abacopteris, might also belong in Pseudocyclosorus
Synonyms: hidden
= Aspidium albolineatum Boivin; Salomon, Nom. 42 (1883)
= Aspidium elatum Bojer; Hortus Maurit. 390 (1837) (nomen), ex Baker, Syn. 502 (1874) (non 298 (1867))
= Aspidium mauritianum (Fée) Kuhn; Filic. Afr. 136 (1868), non Desv.
= Aspidium maximum Bojer; Baker, Syn. 502 (1874)
= Aspidium procerum Bojer; Kuhn, Filic. Afr. 139 (1868) (nomen), ex Decken, Reisen Ost-Afr. 38: Bot. 41 (1879) (non Spreng. (1827))
= Cyclosorus mauritianus (Fée) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 246 (1941)
= Dryopteris mauritiana (Fée) C.Chr.; Index Filic. 277 (1905), nom. nov., non Nephodium mauritianum Fée (1852), nec Desv. (1827)
= Dryopteris procera Kuntze; Revis. Gen. Pl. 2: 811 (1891)
= Nephrodium elatum Baker; Hook. & Baker, Syn. Fil. ed. 2: 502 (1874)
= Nephrodium elatum var. mauritianum (Fée) Baker; Flora of Mauritius and the Seychelles: 499 (1877)
= Nephrodium mauritianum Fée; Mém. Foug. 5 (Gen. Filic.): 308 (1852)
= Nephrodium procerum (Bojer ex Decken) Baker; Hook. & Baker, Syn. Fil. ed. 2: 502 (1874)
= Thelypteris dodo Christenh.; Global Fl. 1(4): 29 (2018), nom. nov.
= Thelypteris mauritiana (Fée) C.F.Reed; Phytologia 17: 291 (1968)
subspecies :
subsp. thomensis
Holttum; J. South Afr. Bot. 40(2): 167 (1974)
Distribution: São Tomé
23 SAO
Synonyms: hidden
= Thelypteris mauritiana subsp. thomensis (Holttum) Viane; Bull. Soc. Roy. Bot. Belg. 118(1): 54 (1985)
species :
Sphaerostephanos ellipticus
(Rosenst.) Holttum; Kalikasan 4: 66 (1975)
Distribution: Philippines (Mindanao, Negros)
42 PHI
Conservation Status: Only known from types
Synonyms: hidden
= Cyclosorus ellipticus (Rosenst.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 142 (1947)
= Dryopteris elliptica Rosenst.; Meded. Rijks Herb. no. 31: 6 (1917)
= Sphaerostephanos ellipticus var. glabrior Holttum; Fl. Males. Ser. 2,1(5): 475 (1982)
= Thelypteris elliptica (Rosenst.) C.F.Reed; Phytologia 17: 274 (1968)
species :
Sphaerostephanos eminens
(Baker) Holttum; Fl. Males. Ser. 2,1(5): 449 (1982)
Distribution: C-Sumatra; N-Sumatra
42 SUM
Synonyms: hidden
= Dryopteris eminens (Baker) C.Chr.; Index Filic. 263 (1905)
= Dryopteris pilososquamata var. obtusata Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 16: 12 (1914)
= Mesochlaena talamauensis Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 28: 34 (1918)
= Nephrodium eminens Baker; J. Bot. 18: 213 (1880)
= Sphaerostephanos talamauensis (Alderw.) C.Chr.; Index Filic., Suppl. Tert.: 172 (1934)
= Thelypteris eminens (Baker) Christenh.; Global Fl. 1(4): 29 (2018)
species :
Sphaerostephanos erectus
(Copel) Holttum; Kalikasan 4: 64 (1975)
Distribution: Philippines (Leyte, Biliran)
42 PHI
Synonyms: hidden
= Cyclosorus erectus Copel.; Philipp. J. Sci. 81: 30, t. 22 (1952)
= Thelypteris erecta (Copel.) C.F.Reed; Phytologia 17: 275 (1968)
species :
Sphaerostephanos erwinii
Holttum; Fl. Males. Ser. 2,1(5): 504 (1982)
Distribution: Moluccas (Seram); New Guinea (Irian Jaya)
42 MOL NWG
Synonyms: hidden
= Thelypteris erwinii (Holttum) Christenh.; Global Fl. 1(4): 30 (2018)
species :
Sphaerostephanos exindusiatus
Holttum; Fl. Males. Ser. 2,1(5): 463 (1982)
Distribution: SW-Sulawesi
42 SUL
Conservation Status: Only known from type
Synonyms: hidden
= Thelypteris peregrinii Christenh.; Global Fl. 1(4): 35 (2018), nom. nov.
species :
Sphaerostephanos flavoviridis
Holttum; Fl. Males. Ser. 2,1(5): 450 (1982)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Conservation Status: Only known from 2 specimens
Synonyms: hidden
= Thelypteris flavoviridis (Holttum) Christenh.; Global Fl. 1(4): 30 (2018)
species :
Sphaerostephanos foliolosus
Holttum; Fl. Males. Ser. 2,1(5): 488 (1982)
Distribution: N-Sulawesi; Moluccas (Talaud)
42 MOL SUL
Synonyms: hidden
= Thelypteris foliolosa (Holttum) Christenh.; Global Fl. 1(4): 30 (2018)
variant :
var. minor
Holttum; Blumea 33(2): 403 (1988)
Distribution: Sulawesi
42 SUL
species :
Sphaerostephanos foxworthyi
Holttum; Fl. Males. Ser. 2,1(5): 475 (1982)
Distribution: Philippines (Luzon)
42 PHI
Conservation Status: Only known from type
Synonyms: hidden
= Thelypteris foxworthyi (Holttum) Christenh.; Global Fl. 1(4): 30 (2018)
species :
Sphaerostephanos gaudichaudii
Holttum; Kew Bull. 34(2): 228 (1979)
Distribution: Vietnam
41 VIE
Synonyms: hidden
= Thelypteris gaudichaudii (Holttum) Christenh.; Global Fl. 1(4): 30 (2018)
species :
Sphaerostephanos glandulosus
(Blume) S.E.Fawc. & A.R.Sm.; A generic revision of the Thelypteridaceae: 81 (2021)
Distribution: peninsular Thailand; peninsular Malaysia (Perak); Sumatra; Sulawesi; Java
41 THA 42 JAW MLY SUL SUM
Synonyms: hidden
= Abacopteris glandulosa (Blume) Fée; Mém. Foug. 5 [Gen. Filic.] 310 (1852)
= Aspidium glandulosum Blume; Enum. Pl. Javae 2: 144 (1828)
= Cyclodium glandulosum (Blume) C.Presl; Tent. Pterid. 85, t. 2, f. 21 (1836)
= Cyclosorus bartlettii (Copel.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 243 (1941)
= Cyclosorus excrescens (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 142 (1947)
= Cyclosorus glandulosus (Blume) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 227 (1938)
= Cyclosorus iridescens (Alderw.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 142 (1947)
= Dryopteris bartlettii Copel.; Univ. Calif. Publ. Bot. 14: 374, t. 58 (1929)
= Dryopteris excrescens Copel.; Univ. Calif. Publ. Bot. 14: 374 (1929)
= Dryopteris glandulosa (Blume) Kuntze; Revis. Gen. Pl. 2: 812 (1891), et C. Chr., Ind. 268 (1905) (non Dryopteris glandulosa (Desv.) C. Chr.)
= Dryopteris iridescens Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 11: 11 (1913)
= Dryopteris malayensis C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afh., ser. 7, 10: 171 (1913) (nom. nov. illegit.)
= Nephrodium glandulosum (Blume) Hook.; Sp. Fil. 4: 76 (1862), isonym
= Nephrodium glandulosum (Blume) J.Sm.; J. Bot. (Hook.) 3: 411 (1841)
= Nephrodium glandulosum (Blume) T.Moore; Index Filic. (T. Moore) 1 (1857), isonym
= Nephrodium meniscioides C.B.Clarke; Trans. Linn. Soc. London, Bot. 1: 532 (1880)
= Pronephrium glandulosum (Blume) Holttum; Blumea 20: 118 (1972)
= Thelypteris bartlettii (Copel.) C.F.Reed; Phytologia 17: 263 (1968)
= Thelypteris glandulosa (Blume) Tagawa & K.Iwats.; Acta Phytotax. Geobot. 23: 54 (1968), non (Desv.) Proctor. 1960
= Thelypteris iridescens (Alderw.) C.F.Reed; Phytologia 17: 285 (1968)
= Thelypteris malayensis (C.Chr.) C.F.Reed; Phytologia 17: 291 (1968)
= Thelypteris my Christenh.; Global Fl. 1(4): 34 (2018), nom. nov. for Aspidium glandulosum Blume
species :
Sphaerostephanos grandescens
Holttum; Fl. Males. Ser. 2,1(5): 465 (1982)
Distribution: Philippines (Mindoro, Mt. Halcon)
42 PHI
Synonyms: hidden
= Thelypteris grandescens (Holttum) Christenh.; Global Fl. 1(4): 31 (2018)
species :
Sphaerostephanos gregarius
(Copel.) Holttum; Fl. Males. Ser. 2,1(5): 480 (1982)
Distribution: Sulawesi; New Guinea (Papua New Guinea); Solomon Isl. (Bougainville, Mono)
42 NWG SUL 43 SOL
Synonyms: hidden
= Cyclosorus gregarius Copel.; J. Arnold Arbor. 24: 440 (1943)
= Thelypteris gregaria (Copel.) C.F.Reed; Phytologia 17: 280 (1968)
species :
Sphaerostephanos gymnorachis
Holttum; Fl. Males. Ser. 2,1(5): 479 (1982)
Distribution: Borneo (Mt. Kinabalu, Sabah)
42 BOR
Synonyms: hidden
= Thelypteris gymnorachis (Holttum) Christenh.; Global Fl. 1(4): 31 (2018)
species :
Sphaerostephanos hamiferus
(Alderw.) Holttum; Fl. Males. Ser. 2,1(5): 496 (1982)
Distribution: C-Sumatra
42 SUM
Synonyms: hidden
= Cyclosorus hamiferus (Alderw.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 245 (1941)
= Dryopteris hamifera Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 16: 12 (1914)
= Thelypteris hamifera (Alderw.) C.F.Reed; Phytologia 17: 281 (1968)
species :
Sphaerostephanos hastatopinnatus
(Brause) Holttum; Fl. Males. Ser. 2,1(5): 470 (1982)
Distribution: C-Sulawesi; Moluccas (Seram); New Guinea (Irian Jaya, Papua New Guinea)
42 MOL NWG SUL
Synonyms: hidden
= Cyclosorus hastatopinnatus (Brause) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 245 (1941)
= Dryopteris hastatopinnata Brause; Bot. Jahrb. Syst. 66: 112 (1920)
= Thelypteris hastatopinnata (Brause) C.F.Reed; Phytologia 17: 281 (1968)
species :
Sphaerostephanos hellwigensis
Holttum; Fl. Males. Ser. 2,1(5): 500 (1982)
Distribution: New Guinea (Irian Jaya: Mt. Hellwig)
42 NWG
Synonyms: hidden
= Thelypteris hellwigensis (Holttum) Christenh.; Global Fl. 1(4): 31 (2018)
species :
Sphaerostephanos hendersonii
Holttum; Fl. Males. Ser. 2,1(5): 493 (1982)
Distribution: peninsular Malaysia (Cameron Highlands, Pahang)
42 MLY
Synonyms: hidden
= Thelypteris hendersonii (Holttum) Christenh.; Global Fl. 1(4): 31 (2018)
species :
Sphaerostephanos hernaezii
Holttum; Fl. Males. Ser. 2,1(5): 459 (1982)
Distribution: Philippines (Samar)
42 PHI
Conservation Status: Only known from type
Synonyms: hidden
= Thelypteris hernaezii (Holttum) Christenh.; Global Fl. 1(4): 31 (2018)
species :
Sphaerostephanos heterocarpos
(Blume) Holttum; Nayar & Kaur, Comp. to Bedd. Handb. Ferns Brit. India 209 (1974)
Distribution: China (Guangdong, Guangxi, Hainan); ?Andaman Isl.; ?Nicobar Isl.; peninsular Thailand; SE-Thailand; Laos; Cambodia; Vietnam; peninsular Malaysia; Singapore; Sumatra; Borneo (Mt. Kinabalu, Sabah, Sarawak); Sulawesi; Palawan; Philippines; Java; Lesser Sunda Isl. (Lombok, Bali, Flores, Sumbawa); Moluccas; New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch. (New Hanover); Solomon Isl. (Gizo, Guadalcanal, Santa Ysabel, Kolombangara, Makira, Malaita, New Georgia, Rendova, Vangunu, Vangunu); Australia (NE-Queensland); Santa Cruz Isl.; Vanuatu; Fiji (Viti Levu, Vanua Levu); American Samoa (Tutuila, Olosega, Ta´u); [Western] Samoa; Palau (Babeldaob, Koror); Northern Marianas (Rota Isl.); Fed. States of Micronesia (Pohnpei, Kosrae)
36 CHI 41 CBD LAO THA VIE 42 BOR JAW LSI MLY MOL NWG PHI PLW SNG SUL SUM 43 BIS SOL 50 AUS 60 ASA FIJ SAM SCZ VAN 62 MIC MRN PLU
Synonyms: hidden
= Aspidium heterocarpon Blume; Enum. Pl. Javae 2: 155 (1828)
= Cyclosorus bordenii (Christ) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 243 (1941)
= Cyclosorus heterocarpos (Blume) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 180 (1938)
= Cyclosorus suprastrigosus (Rosenst.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 143 (1947)
= Dryopteris bordenii Christ; Philipp. J. Sci., Bot. 2: 204 (1907)
= Dryopteris heterocarpos (Blume) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Dryopteris mixta Rosenst.; Repert. Spec. Nov. Regni Veg. 12: 172 (1913)
= Dryopteris suprastrigosa Rosenst.; Repert. Spec. Nov. Regni Veg. 10: 335 (1912)
= Lastrea mixta (Rosenst.) Copel.; Philipp. J. Sci. 78: 426 (1951)
= Nephrodium heterocarpon (Blume) T.Moore; Index Filic. (T. Moore) 93 (1858)
= Nephrodium pubescens Brack.; U.S. Expl. Exped., Filic. 16: 186 (1854) (non Don (1825))
= Polypodium longifolium Roxb.; Calcutta J. Nat. Hist. 4: 462 (1844), non Cav. (1802), nec C. Presl (1822)
= Sphaerostephanos heterocarpos var. bordenii Holttum; Kalikasan 4: 67 (1975)
= Thelypteris bordenii (Christ) C.F.Reed; Phytologia 17: 264 (1968)
= Thelypteris heterocarpos (Blume) C.V.Morton; Amer. Fern J. 49: 113 (1959)
= Thelypteris mixta (Rosenst.) C.F.Reed; Phytologia 17: 293 (1968)
= Thelypteris suprastrigosa (Rosenst.) C.F.Reed; Phytologia 17: 318 (1968)
species :
Sphaerostephanos heterophyllus
(C.Presl) S.E.Fawc. & A.R.Sm.; A generic revision of the Thelypteridaceae: 82 (2021)
Distribution: Philippines (Samar, S-Luzon, Mindanao)
42 PHI
Synonyms: hidden
= Aspidium blumei Kunze ex Mett.; Fil. Hort. Bot. Lips. 94, t. 22, f. 5 (1856)
= Aspidium blumei var. subpinnata Mett.; Farngatt. IV: 98 (1858)
= Aspidium canescens (Blume) Christ; Christ, Farnkr. Erde 244 (1897)
= Aspidium heterophyllum (C.Presl) Hook.; Hooker´s Icon. Pl. 6: 10 (1854), t. 920
= Dryopteris heterophylla (C.Presl) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Haplodictyum heterophyllum C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6: 51 (1851)
= Nephrodium heterophyllum (C.Presl) Hook.; Sp. Fil. 4: 62 (1862)
= Pleocnemia heterophylla (C.Presl) Alderw.; Handb. 171 (1908)
= Pronephrium heterophyllum (C.Presl) Holttum; Fl. Males. Ser. 2,1(5): 515 (1982)
= Thelypteris heterophylla (C.Presl) K.Iwats.; Mem. Coll. Sci. Univ. Kyoto, B, Biol. 31: 32 (1964)
species :
Sphaerostephanos hirsutus
(Kunze ex Mett.) Holttum; Kalikasan 4: 63 (1975)
Distribution: Sulawesi; Philippines
42 PHI SUL
Remarks: not in India
Synonyms: hidden
= Aspidium hirsutum Kunze ex Mett.; Farngatt. IV: 107 (1858), nom. nov.
= Cyclosorus adenophorus (C.Chr.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 242 (1941)
= Dryopteris adenophora C.Chr.; Index Filic. 251 (1905) (nom. nov. superfl. for Nephrodium hirsutum J. Sm. ex C. Presl)
= Dryopteris hirsuta (Kunze ex Mett.) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Nephrodium hirsutum J.Sm.; J. Bot. (Hooker) 3: 412 (1841), nomen, ex C. Presl, Epim. 48 (1849) (non Don (1825) nec Bory (1828))
= Thelypteris adenophora (C.Chr.) C.F.Reed; Phytologia 17: 258 (1968)
= Thelypteris hirsuta (Kunze ex Mett.) Christenh.; Global Fl. 1(4): 31 (2018), isonym
= Thelypteris hirsuta (Kunze ex Mett.) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 1: 507 (2016)
variant :
var. celebicus
Holttum; Fl. Males. Ser. 2,1(5): 456 (1982)
Distribution: C-Sulawesi; SW-Sulawesi; Lesser Sunda Isl.
42 LSI SUL
variant :
var. elatus
Holttum; Blumea 33(2): 403 (1988)
Distribution: Sulawesi
42 SUL
species :
Sphaerostephanos hispidifolius
(Alderw.) Holttum; Fl. Males. 1: 466 (1981)
Distribution: Borneo (Sarawak, Kalimantan)
42 BOR
Synonyms: hidden
= Dryopteris hispidifolia Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 20: 15 (1915)
= Nephrodium hispidulum Christ; Ann. Jard. Bot. Buitenz. 20: 107 (1906) (non (Decne.) Baker)
= Thelypteris hispidifolia (Alderw.) C.F.Reed; Phytologia 17: 283 (1968)
= Cyclosorus hispidifolius (Alderw.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 245 (1941)
species :
Sphaerostephanos hispiduliformis
(C.Chr.) Holttum; Fl. Males. Ser. 2,1(5): 470 (1982)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Cyclosorus hispiduliformis (C.Chr.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 245 (1941)
= Dryopteris hispida Brause; Bot. Jahrb. Syst. 56: 102 (1920) (non Kuntze)
= Dryopteris hispiduliformis C.Chr.; Index Filic., Suppl. Tert.: 88 (1934)
= Thelypteris hispida (Brause) C.F.Reed; Phytologia 17: 283 (1968)
= Thelypteris hispiduliformis (C.Chr.) C.F.Reed; Phytologia 17: 283 (1968)
variant :
var. brassii
Holttum; Fl. Males. Ser. 2,1(5): 470 (1982)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Cyclosorus strigosissimus (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 143 (1947)
= Dryopteris strigosissima Copel.; Univ. Calif. Publ. Bot. 18: 221 (1942)
= Thelypteris strigosissima (Copel.) C.F.Reed; Phytologia 17: 316 (1968)
variant :
var. vinkii
Holttum; Fl. Males. Ser. 2,1(5): 470 (1982)
Distribution: New Guinea (Papua New Guinea: Sepik)
42 NWG
species :
Sphaerostephanos hoalensis
Holttum; Fl. Males. Ser. 2,1(5): 488 (1982)
Distribution: Moluccas (Seram)
42 MOL
Conservation Status: Only known from 2 collections
Synonyms: hidden
= Thelypteris hoalensis (Holttum) Christenh.; Global Fl. 1(4): 31 (2018)
species :
Sphaerostephanos hochreutineri
(C.Chr.) Holttum; Allertonia 1(3): 207 (1977)
Distribution: [Western] Samoa (´Upolu)
60 SAM
Synonyms: hidden
= Dryopteris hochreutineri Christ; Annuaire Conserv. Jard. Bot. Genève 16-16. 188 (1912)
= Thelypteris hochreutineri (C.Chr.) Christenh.; Global Fl. 1(4): 31 (2018)
species :
Sphaerostephanos humilis
Holttum; Fl. Males. Ser. 2,1(5): 468 (1982)
Distribution: C-Sulawesi
42 SUL
Conservation Status: Only known from type
Synonyms: hidden
= Thelypteris humilis (Holttum) Christenh.; Global Fl. 1(4): 32 (2018)
species :
Sphaerostephanos immucosus
Holttum; Blumea 33(2): 404 (1988)
Distribution: Sulawesi
42 SUL
Synonyms: hidden
= Thelypteris immucosa (Holttum) Christenh.; Global Fl. 1(4): 32 (2018)
species :
Sphaerostephanos incisus
(Copel.) S.E.Fawc. & A.R.Sm.; A generic revision of the Thelypteridaceae: 82 (2021)
Distribution: Moluccas (Ternate, Halmaheira)
42 MOL
Synonyms: hidden
= Pneumatopteris incisa Holttum; Blumea 21: 317 (1973)
= Thelypteris incisa (Holttum) Christenh.; Global Fl. 1(4): 32 (2018)
species :
Sphaerostephanos inconspicuus
(Copel.) Holttum; Fl. Males. Ser. 2,1(5): 493 (1982)
Distribution: Borneo (Mt. Kinabalu, Sabah)
42 BOR
Synonyms: hidden
= Dryopteris inconspicua Copel.; Philipp. J. Sci., Bot. 12: 55 (1917)
= Thelypteris inconspicua (Copel.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 252 (1941)
species :
Sphaerostephanos indrapurae
Holttum; Fl. Males. Ser. 2,1(5): 475 (1982)
Distribution: C-Sumatra (Mt. Kerinci)
42 SUM
Conservation Status: Only known from type
Synonyms: hidden
= Thelypteris indrapurae (Holttum) Christenh.; Global Fl. 1(4): 32 (2018)
species :
Sphaerostephanos irayensis
(Copel.) Holttum; Fl. Males. Ser. 2,1(5): 495 (1982)
Distribution: Moluccas (Bacan)
42 MOL
Synonyms: hidden
= Cyclosorus irayensis Copel.; Philipp. J. Sci. 81: 28 (1952)
= Plesioneuron irayense (Copel.) Holttum; Blumea 22: 239 (1975)
= Sphaerostephanos fenixii Holttum; Kalikasan 4: 55 (1975)
= Thelypteris irayensis (Copel.) C.F.Reed; Phytologia 17: 285 (1968)
species :
Sphaerostephanos isomorphus
Holttum; Fl. Males. Ser. 2,1(5): 458 (1982)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Conservation Status: Only known from type
Synonyms: hidden
= Thelypteris isomorpha (Holttum) Christenh.; Global Fl. 1(4): 32 (2018)
species :
Sphaerostephanos kalkmanii
Holttum; Fl. Males. Ser. 2,1(5): 454 (1982)
Distribution: New Guinea (Irian Jaya)
42 NWG
Conservation Status: Only known from type
Synonyms: hidden
= Thelypteris kalkmanii (Holttum) Christenh.; Global Fl. 1(4): 32 (2018)
species :
Sphaerostephanos kurzii
Holttum; Kew Bull. 34(2): 231 (1979)
Distribution: Nicobar Isl.
41 NCB
Remarks: genus uncertain
Conservation Status: IUCN CR or EX for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Cyclosorus kurzii (Holttum) Panigrahi; Pl. Sci. Res. (India) 17(1-2): 52 (1995)
= Thelypteris kurzii (Holttum) Fraser-Jenk.; New Species Syndr. Indian Pteridol. 280 (1997)
species :
Sphaerostephanos lamii
Holttum; Fl. Males. Ser. 2,1(5): 455 (1982)
Distribution: New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch. (Admiralty Isl.)
42 NWG 43 BIS
Synonyms: hidden
= Thelypteris lamii (Holttum) Christenh.; Global Fl. 1(4): 32 (2018)
species :
Sphaerostephanos larutensis
(Bedd.) C.Chr.; Index Filic., Suppl. Tert.: 172 (1934)
Distribution: peninsular Thailand; peninsular Malaysia (Perak); Sumatra; Borneo
41 THA 42 BOR MLY SUM
Synonyms: hidden
= Cyclosorus larutensis (Bedd.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 245 (1941)
= Dryopteris larutensis (Bedd.) C.Chr.; Index Filic. 273 (1905)
= Mesochlaena larutensis (Bedd.) Alderw.; Bull. Dép. Agric. Indes Néerl. 18: 8 (1908)
= Mesochlaena moluccana R.Br. ex J.Sm.; J. Bot. 3: 18 (1840)
= Mesochlaena sumatrensis Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 3, 2: 160 (1920)
= Nephrodium larutense Bedd.; Suppl. Ferns Brit. Ind. 73 (1892)
= Sphaerostephanos sumatrensis (Alderw.) C.Chr.; Index Filic., Suppl. Tert.: 172 (1934)
= Stegnogramma mesochlaena Fée; Mém. Foug. 5 (Gen. Filic.): 204 (1852)
= Stegnogramma moluccana Fée; Mém. Foug. 5 (Gen. Filic.): 204 (1852)
= Thelypteris larutensis (Bedd.) C.F.Reed; Phytologia 17: 286 (1968), isonym
= Thelypteris larutensis (Bedd.) Tagawa & K.Iwats.; Acta Phytotax. Geobot. 23: 53 (1968)
species :
Sphaerostephanos lastreoides
(C.Presl) Holttum; Kalikasan 4: 54 (1975)
Distribution: Palawan; Philippines (Luzon, Negros, Mindanao)
42 PHI PLW
Synonyms: hidden
= Aspidium lastreoides (C.Presl) Mett.; Salomon, Nom. 57 (1883)
= Cyclosorus duplosetosus Copel.; Philipp. J. Sci. 81: 32 (1952)
= Pronephrium lastreoides C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6: 259 (1851)
= Thelypteris lastreoides (C.Presl) Christenh.; Global Fl. 1(4): 32 (2018)
= Thelypteris duplosetosa (Copel.) C.F.Reed; Phytologia 17: 274 (1968)
species :
Sphaerostephanos latebrosus
(Kunze ex Mett.) Holttum; Nayar & Kaur, Comp. to Bedd. Handb. Ferns Brit. India 209 (1974)
Distribution: China (S-Yunnan); Nepal; India (Assam State, West Bengal); Bangladesh; peninsular Malaysia; Borneo (Mt. Kinabalu, Sarawak); Sulawesi; Palawan; Philippines
36 CHI 40 BAN IND NEP 42 BOR MLY PHI PLW SUL
Remarks: the report by Lin & Iwatsuki (Fl. of. China) from Bangladesh, but not India, was due to misplacing Haldibari and Northern Bengal
Synonyms: hidden
= Aspidium dimorphum Kunze; Bot. Zeitung (Berlin) 261 (1848)
= Aspidium latebrosum Kunze; Bot. Zeitung (Berlin) 261 (1848) (nomen), ex Mett., Pheg. u. Asp. 104, n. 248 (1858)
= Cyclosorus angustipes (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 142 (1947)
= Cyclosorus glabellus Ching ex W.M.Chu & S.G.Lu; Fl. Yunnanica 20: 608 (2006)
= Cyclosorus glaucostipes (Bedd.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 245 (1941)
= Cyclosorus halconensis Copel.; Philipp. J. Sci. 81: 29 (1952)
= Cyclosorus heterocarpus var. glaucostipes (Bedd.) Holttum; Rev. Fl. Mal. 2: 271 (1955)
= Cyclosorus latebrosus (Kunze ex Mett.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 245 (1941)
= Cyclosorus polypterus Copel.; Philipp. J. Sci. 84: 161 (1955)
= Cyclosorus sagittifolioides Copel.; Philipp. J. Sci. 81: 29, 30, pl. 21 (1952)
= Cyclosorus todayensis (Christ) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 249 (1941)
= Dryopteris angustipes Copel.; Philipp. J. Sci., Bot. 7: 60 (1912)
= Dryopteris glaucostipes (Bedd.) C.Chr.; Index Filic. 268 (1905)
= Dryopteris latebrosa (Kunze ex Mett.) C.Chr.; Index Filic. 274 (1905)
= Dryopteris todayensis Christ; Philipp. J. Sci., Bot. 2: 193 (1907)
= Nephrodium dimorphum (Kunze) T.Moore; Index Filic. (T. Moore) 90 (1858)
= Nephrodium glaucostipes Bedd.; Suppl. Ferns Brit. Ind. 80 (1892)
= Nephrodium latebrosum (Kunze ex Mett.) T.Moore; Index Filic. (T. Moore) 95 (1858)
= Sphaerostephanos todayensis (Christ) Holttum; Kalikasan 4: 58 (1975)
= Thelypteris angustipes (Copel.) C.F.Reed; Phytologia 17: 260 (1968)
= Thelypteris halconensis (Copel.) C.F.Reed; Phytologia 17: 281 (1968)
= Thelypteris heterocarpus var. glaucostipes (Bedd.) C.F.Reed; Phytologia 17: 282 (1968)
= Thelypteris latebrosa (Kunze ex Mett.) C.F.Reed; Phytologia 17: 287 (1968)
= Thelypteris polyptera (Copel.) C.F.Reed; Phytologia 17: 305 (1968)
= Thelypteris sagittifolioides (Copel.) C.F.Reed; Phytologia 17: 311 (1968)
= Thelypteris todayensis (Christ) C.F.Reed; Phytologia 17: 319 (1968)
species :
Sphaerostephanos lithophyllus
(Copel.) Holttum; Fl. Males. Ser. 2,1(5): 496 (1982)
Distribution: Borneo (Mt. Kinabalu, Sabah)
42 BOR
Synonyms: hidden
= Cyclosorus lithophyllus (Copel.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 246 (1941)
= Dryopteris lithophylla Copel.; Philipp. J. Sci., Bot. 12: 57 (1917)
= Thelypteris lithophylla (Copel.) C.F.Reed; Phytologia 17: 289 (1968)
species :
Sphaerostephanos lobangensis
(C.Chr.) Holttum; Fl. Males. Ser. 2,1(5): 479 (1982)
Distribution: Borneo (Mt. Kinabalu, Sabah)
42 BOR
Conservation Status: Only known from 2 collections
Synonyms: hidden
= Cyclosorus lobangensis (C.Chr. ex C.Chr. & Holttum) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 142 (1947)
= Dryopteris lobangensis C.Chr.; Gard. Bull. Straits Settlem. 7: 246 (1934)
= Thelypteris lobangensis (C.Chr. ex C.Chr. & Holttum) C.F.Reed; Phytologia 17: 289 (1968)
species :
Sphaerostephanos lobatus
(Copel.) Holttum; Kalikasan 4: 64 (1975)
Distribution: Palawan; Philippines (Luzon)
42 PHI PLW
Synonyms: hidden
= Cyclosorus lobatus (Copel.) Copel.; Philipp. J. Sci. 81: 33 (1952)
= Dryopteris canescens var. lobata Christ; Philipp. J. Sci., Bot. 2: 198 (1907)
= Thelypteris lobata (Copel.) C.F.Reed; Phytologia 17: 289 (1968)
species :
Sphaerostephanos loherianus
(Christ) Holttum; Kalikasan 4: 61 (1975)
Distribution: Philippines (N-Luzon)
42 PHI
Synonyms: hidden
= Aspidium loherianum Christ; Bull. Herb. Boissier 6: 191 (1893)
= Cyclosorus loherianus (Christ) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 142 (1947)
= Dryopteris loheriana (Christ) C.Chr.; Index Filic. 275 (1905)
= Nephrodium loherianum Copel.; Polyp. Philipp. 30 (1905)
= Thelypteris loheriana (Christ) C.F.Reed; Phytologia 17: 289 (1968)
species :
Sphaerostephanos longbawanensis
(K.Iwats. & M.Kato) S.E.Fawc. & A.R.Sm.; A generic revision of the Thelypteridaceae: 82 (2021)
Distribution: Borneo (Kalimantan, Sarawak)
42 BOR
Synonyms: hidden
= Thelypteris longbawanensis K.Iwats. & M.Kato; Acta Phytotax. Geobot. 34(4-6): 137 (1983)
species :
Sphaerostephanos maemonensis
(W.H.Wagner & Grether) Holttum; Allertonia 1(3): 209 (1977)
Distribution: Palau (Babeldaob); Northern Marianas (Saipan); Guam; Fed. States of Micronesia (Truk, Moen, Dublon, Pohnpei, Kosrae)
62 GUM MIC MRN PLU
Synonyms: hidden
= Aspidium adenopteris Kunze ex Mett.; Pheg. u. Asp. 107, n. 255 (1858)
= Cyclosorus maemonensis Wagner & Grether; Occas. Pap. Bernice Pauahi Bishop Mus. 19: 54, f. 5 (1948)
= Nephrodium adenopteris Kunze ex T.Moore; Index Filic. (T. Moore) 85 (1858)
= Polypodium adenopteris Kunze; hb.
= Thelypteris maemonensis (W.H.Wagner & Grether) Stone; Micronesica 2: 135 (1967)
species :
Sphaerostephanos magnus
(Copel.) Holttum; Fl. Males. Ser. 2,1(5): 465 (1982)
Distribution: Philippines (Negros)
42 PHI
Conservation Status: Only known from 2 collections
Synonyms: hidden
= Cyclosorus magnus Copel.; Philipp. J. Sci. 81: 30 (1952)
= Thelypteris magna (Copel.) C.F.Reed; Phytologia 17: 291 (1968)
species :
Sphaerostephanos major
(Copel.) Holttum; Fl. Males. Ser. 2,1(5): 464 (1982)
Distribution: Philippines (Mountains of, N-Luzon)
42 PHI
Synonyms: hidden
= Cyclosorus dimorphus Copel.; Philipp. J. Sci. 83: 99, pl. 5 (1954) (non (Brause) Copel. (1960))
= Haplodictyum majus Copel.; Elmer, Leafl. Philipp. Bot. 9 (1920)
= Thelypteris copelandii C.F.Reed; Phytologia 17: 269. 1968, nom. nov.
= Thelypteris major (Copel.) Christenh.; Global Fl. 1(4): 33 (2018)
species :
Sphaerostephanos makassaricus
Holttum; Fl. Males. Ser. 2,1(5): 498 (1982)
Distribution: SW-Sulawesi
42 SUL
Synonyms: hidden
= Thelypteris makassarica (Holttum) Christenh.; Global Fl. 1(4): 33 (2018)
species :
Sphaerostephanos maximus
(K.Iwats. & M.Kato) S.E.Fawc. & A.R.Sm.; A generic revision of the Thelypteridaceae: 82 (2021)
Distribution: Borneo (Kalimantan)
42 BOR
Synonyms: hidden
= Thelypteris maxima K.Iwats. & M.Kato; Acta Phytotax. Geobot. 34(4-6): 138 (1983)
species :
Sphaerostephanos melanophlebius
(Copel.) S.E.Fawc. & A.R.Sm.; A generic revision of the Thelypteridaceae: 82 (2021)
Distribution: Philippines (Negros, Mindanao, ?Luzon); Moluccas (Seram)
42 MOL PHI
Synonyms: hidden
= Cyclosorus melanophlebius (Copel.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 246 (1941)
= Dryopteris canescens var. subsimplicifolia Christ; Philipp. J. Sci., Bot. 2: 199 (1907)
= Dryopteris melanophlebia Copel.; Philipp. J. Sci., Bot. 6: 147 (1911)
= Phegopteris melanophlebia (Copel.) Alderw.; Malayan Ferns Fern Allies 319 (1917)
= Pronephrium melanophlebium (Copel.) Holttum; Blumea 20: 108 (1972)
= Thelypteris melanophlebia (Copel.) C.F.Reed; Phytologia 17: 292 (1968)
species :
Sphaerostephanos melanorachis
Holttum; Fl. Males. Ser. 2,1(5): 457 (1982)
Distribution: Moluccas (Bacan)
42 MOL
Synonyms: hidden
= Thelypteris melanorachis (Holttum) Christenh.; Global Fl. 1(4): 33 (2018)
species :
Sphaerostephanos menadensis
Holttum; Fl. Males. Ser. 2,1(5): 469 (1982)
Distribution: NE-Sulawesi
42 SUL
Conservation Status: Only known from type
Synonyms: hidden
= Thelypteris menadensis (Holttum) Christenh.; Global Fl. 1(4): 33 (2018)
species :
Sphaerostephanos mengenianus
Holttum; Fl. Males. Ser. 2,1(5): 491 (1982)
Distribution: Bismarck Arch. (New Britain)
43 BIS
Conservation Status: Only known from type
Synonyms: hidden
= Thelypteris mengeniana (Holttum) Christenh.; Global Fl. 1(4): 33 (2018)
species :
Sphaerostephanos metcalfei
(Baker) Holttum; Allertonia 1(3): 207 (1977)
Distribution: Santa Cruz Isl.; Vanuatu (Aneityum)
60 SCZ VAN
Synonyms: hidden
= Dryopteris metcalfei (Baker) C.Chr.; Index Filic. 277 (1905)
= Polypodium metcalfei Baker; Ann. Bot. (Oxford) 5: 461 (1891)
= Thelypteris metcalfei (Baker) C.F.Reed; Phytologia 17: 293 (1968)
species :
Sphaerostephanos microlonchus
(Christ) S.E.Fawc. & A.R.Sm.; A generic revision of the Thelypteridaceae: 82 (2021)
Distribution: Philippines (Luzon, Negros)
42 PHI
Synonyms: hidden
= Cyclosorus microlonchus (Christ) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 246 (1941)
= Dryopteris caudiculata Alderw.; Handb. 820 (1908)
= Dryopteris microloncha Christ; Philipp. J. Sci., Bot. 2: 202 (1907)
= Pneumatopteris microloncha (Christ) Holttum; Blumea 21: 308 (1973)
= Thelypteris microloncha (Christ) C.F.Reed; Phytologia 7: 293 (1968)
species :
Sphaerostephanos micropinnatus
(Holttum) S.E.Fawc. & A.R.Sm.; A generic revision of the Thelypteridaceae: 82 (2021)
Distribution: New Guinea (N-Papua New Guinea: Sepik, Madang Districts)
42 NWG
Synonyms: hidden
= Pronephrium micropinnatum Holttum; Blumea 20: 108 (1972)
= Thelypteris micropinnata (Holttum) Christenh.; Global Fl. 1(4): 33 (2018)
species :
Sphaerostephanos mindorensis
Holttum; Kalikasan 4: 54 (1975)
Distribution: Philippines (Mindoro)
42 PHI
Remarks: status uncertain, known only from two collections
Synonyms: hidden
= Thelypteris mindorensis (Holttum) Christenh.; Global Fl. 1(4): 34 (2018)
species :
Sphaerostephanos mjobergii
Holttum; Fl. Males. Ser. 2,1(5): 494 (1982)
Distribution: Borneo (Sarawak)
42 BOR
Conservation Status: Only known from type
Synonyms: hidden
= Thelypteris mjobergii (Holttum) Christenh.; Global Fl. 1(4): 34 (2018)
species :
Sphaerostephanos morotaiensis
Holttum; Fl. Males. Ser. 2,1(5): 453 (1982)
Distribution: Moluccas (Morotai, Obi)
42 MOL
Synonyms: hidden
= Thelypteris morotaiensis (Holttum) Christenh.; Global Fl. 1(4): 34 (2018)
species :
Sphaerostephanos moseleyi
Holttum; Fl. Males. Ser. 2,1(5): 460 (1982)
Distribution: Moluccas (Aru Isl.)
42 MOL
Synonyms: hidden
= Thelypteris moseleyi (Holttum) Christenh.; Global Fl. 1(4): 34 (2018)
species :
Sphaerostephanos multiauriculatus
(Copel.) Holttum; Fl. Males. Ser. 2,1(5): 492 (1982)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Cyclosorus multiauriculatus (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 143 (1947)
= Dryopteris multiauriculata Copel.; Univ. Calif. Publ. Bot. 18: 221 (1942)
= Thelypteris multiauriculata (Copel.) C.F.Reed; Phytologia 17: 294 (1968)
species :
Sphaerostephanos muluensis
Holttum; Fl. Males. Ser. 2,1(5): 499 (1982)
Distribution: Borneo (Sarawak)
42 BOR
Synonyms: hidden
= Thelypteris muluensis (Holttum) Christenh.; Global Fl. 1(4): 34 (2018)
species :
Sphaerostephanos mundus
(Rosenst.) Holttum; Fl. Males. Ser. 2,1(5): 503 (1982)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Synonyms: hidden
= Cyclosorus farinosus (Brause) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 244 (1941)
= Cyclosorus farinosus (Brause) Copel.; Philipp. J. Sci.78: 446 (1951), isonym
= Cyclosorus oblongus (Brause) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 247 (1941)
= Cyclosorus oblongus (Brause) Copel.; Philipp. J. Sci.78: 446 (1951), isonym
= Dryopteris farinosa Brause; Bot. Jahrb. Syst. 56: 111 (1920)
= Dryopteris munda Rosenst.; Meded. Rijks Herb. no. 31: 5 (1917)
= Dryopteris oblonga Brause; Bot. Jahrb. Syst. 56: 109 (1920)
= Thelypteris farinosa (Brause) C.F.Reed; Phytologia 17: 276 (1968)
= Thelypteris munda (Rosenst.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 252 (1941)
= Thelypteris oblonga (Brause) C.F.Reed; Phytologia 17: 297 (1968)
species :
Sphaerostephanos mutabilis
(Brause) Holttum; Fl. Males. Ser. 2,1(5): 494 (1982)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Conservation Status: Only known from 2 collections
Synonyms: hidden
= Cyclosorus mutabilis (Brause) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 143 (1947)
= Dryopteris mutabilis Brause; Bot. Jahrb. Syst. 56: 97 (1920)
= Thelypteris mutabilis (Brause) C.F.Reed; Phytologia 17: 295 (1968)
species :
Sphaerostephanos nakaikei
Holttum; Fl. Males. Ser. 2,1(5): 482 (1982)
Distribution: New Guinea (Papua New Guinea: Mt. Wilhelm, Mt. Giluwe)
42 NWG
Synonyms: hidden
= Thelypteris nakaikei (Holttum) Christenh.; Global Fl. 1(4): 34 (2018), nom. nov. illeg.
= Thelypteris toshiyukii Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 2: 463 (2018), nom. nov.
species :
Sphaerostephanos neotoppingii
Holttum; Fl. Males. Ser. 2,1(5): 500 (1982), nom. nov.
Distribution: ?peninsular Malaysia (Malacca); Sumatra; Borneo (Sabah, Kalimantan)
42 BOR SUM
Synonyms: hidden
= Cyclosorus toppingii (Copel.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 224 (1938)
= Dryopteris toppingii Copel.; Philipp. J. Sci., Bot. 12: 56 (1917)
= Nephrodium indicum Ridl.; J. Mal. Br. R. As. Soc. 4: 73 (1926)
= Thelypteris toppingii (Copel.) K.Iwats.; Acta Phytotax. Geobot. 21: 168 (1965), not Sphaerostephanos toppingii (Copel.) C. Chr. (1934)
species :
Sphaerostephanos norrisii
(Rosenst.) Holttum; Fl. Males. Ser. 2,1(5): 485 (1982)
Distribution: peninsular Malaysia (Cameron Highlands, Pahang); Sumatra; Borneo (Sabah, Sarawak); Philippines (Mindanao)
42 BOR MLY PHI SUM
Synonyms: hidden
= Cyclosorus elmerorum (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 142 (1947)
= Cyclosorus subfalcinellus (Alderw.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 249 (1941)
= Dryopteris elmerorum Copel.; Philipp. J. Sci.40: 295, pl. 2 (1929)
= Dryopteris indica var. malayensis (Bedd.) Alderw.; Handb. 224 (1908)
= Dryopteris norrisii Rosenst.; Meded. Rijks Herb. no. 31: 8 (1917)
= Dryopteris subfalcinella Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 3, 2: 151 (1920)
= Lastrea norrisii Ridl.; J. Malayan Branch Roy. Asiat. Soc. 4: 69 (1926)
= Nephrodium pennigerum var. malayense Bedd.; Suppl. Ferns Brit. Ind. 74 (1892)
= Pronephrium elmerorum (Copel.) Holttum; Blumea 20: 118 (1972)
= Sphaerostephanos elmerorum (Copel.) Holttum; Kalikasan 4: 55 (1975)
= Thelypteris elmerorum (Copel.) C.F.Reed; Phytologia 17: 274 (1968)
= Thelypteris norrisii (Rosenst.) Christenh.; Global Fl. 1(4): 34 (2018)
= Thelypteris subfalcinella (Alderw.) C.F.Reed; Phytologia 17: 317 (1968)
species :
Sphaerostephanos novae-britanniae
Holttum; Fl. Males. Ser. 2,1(5): 477 (1982)
Distribution: Bismarck Arch. (New Britain)
43 BIS
Conservation Status: Only known from type
Synonyms: hidden
= Thelypteris novae-brittaniae (Holttum) Christenh.; Global Fl. 1(4): 35 (2018)
species :
Sphaerostephanos novoguineensis
(Brause) Holttum; Fl. Males. Ser. 2,1(5): 469 (1982)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Dryopteris glaucescens Brause; Bot. Jahrb. Syst. 56: 85 (1920)
= Dryopteris novoguineensis Brause; Bot. Jahrb. Syst. 49: 21 (1912)
= Lastrea novoguineensis (Brause) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 139 (1947)
= Thelypteris glaucescens (Brause) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 251 (1941)
= Thelypteris novoguineensis (Brause) C.F.Reed; Phytologia 17: 297 (1968)
species :
Sphaerostephanos nudisorus
Holttum; Fl. Males. Ser. 2,1(5): 476 (1982)
Distribution: C-Sulawesi
42 SUL
Conservation Status: Only known from type
Synonyms: hidden
= Thelypteris nudisora (Holttum) Christenh.; Global Fl. 1(4): 35 (2018)
species :
Sphaerostephanos obtusifolius
(Rosenst.) Holttum; Fl. Males. Ser. 2,1(5): 503 (1982)
Distribution: New Guinea (Papua New Guinea); Bismarck Arch. (Admiralty Isl.)
42 NWG 43 BIS
Synonyms: hidden
= Cyclosorus obtusifolius (Rosenst.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 143 (1947)
= Dryopteris obtusifolia Rosenst.; Repert. Spec. Nov. Regni Veg. 10: 336 (1912)
= Phegopteris obtusifolia (Rosenst.) Alderw.; Malayan Ferns Fern Allies 315 (1917)
= Thelypteris obtusifolia (Rosenst.) C.F.Reed; Phytologia 17: 297 (1968)
species :
Sphaerostephanos omatianus
Holttum; Fl. Males. Ser. 2,1(5): 483 (1982)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Conservation Status: Only known from type
Synonyms: hidden
= Thelypteris omatiana (Holttum) Christenh.; Global Fl. 1(4): 35 (2018)
species :
Sphaerostephanos oosorus
(Baker) Holttum; Fl. Males. Ser. 2,1(5): 500 (1982)
Distribution: Borneo (Sabah, P. Balambagan)
42 BOR
Synonyms: hidden
= Cornopteris oosora (Baker) Tardieu; Amer. Fern J. 48: 32 (1958)
= Cyclosorus oosorus (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 247 (1941)
= Dryopteris oosora (Baker) C.Chr.; Index Filic. 280 (1905)
= Nephrodium oosorum Baker; Bull. Misc. Inform. Kew 1896: 41 (1896)
= Thelypteris oosora (Baker) C.F.Reed; Phytologia 17: 298 (1968)
species :
Sphaerostephanos paripinnatus
(Copel.) Holttum; Fl. Males. Ser. 2,1(5): 476 (1982)
Distribution: New Guinea (Irian Jaya)
42 NWG
Conservation Status: Only known from type
Synonyms: hidden
= Cyclosorus paripinnatus (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 143 (1947)
= Dryopteris paripinnata Copel.; Univ. Calif. Publ. Bot. 18. 220 (1942)
= Thelypteris paripinnata (Copel.) C.F.Reed; Phytologia 17: 302 (1968)
species :
Sphaerostephanos penniger
(Hook.) Holttum; Nayar & Kaur, Comp. to Bedd. Handb. Ferns Brit. India 209 (1974)
Distribution: peninsular Thailand; SW-Thailand; peninsular Malaysia; Sumatra; Borneo (Mt. Kinabalu, Sabah, Kalimantan); Sulawesi; Java; Lesser Sunda Isl. (Lombok, Bali)
41 THA 42 BOR JAW LSI MLY SUL SUM
Remarks: reports from India refer to T. articulata fide Fraser-Jenkins 2008
Synonyms: hidden
= Aspidium megaphyllum Mett.; Ann. Mus. Bot. Lugduno-Batavi 1: 233 (1863), nom. nov for A. pennigerum Blume.
= Aspidium pennigerum Blume; Enum. Pl. Javae 2: 153 (1828), non (G. Forst.) Sw. 1801, nec al.
= Christella megaphylla H.Lév.; Flore du Kouy-tcheou 474-476 (1915)
= Cyclosorus megaphyllus (Mett.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 225 (1938)
= Dryopteris megaphylla (Mett.) C.Chr.; Index Filic. 277 (1905)
= Dryopteris multilineata (Bedd.) Kuntze; Revis. Gen. Pl. 2: 811 (1891)
= Goniopteris lineata Bedd.; Handb. Ferns Brit. India, t. 3 (1865)
= Nephrodium abruptum Bedd.; Ferns South. India, t. 86 (1863)
= Nephrodium articulatum J.Sm.; Cat. Cult. Ferns 54 (1857)
= Nephrodium multilineatum Bedd.; Suppl. Ferns Brit. Ind. 80 (1892) (non Aspidium multilineatum Mett. 1858)
= Nephrodium pennigerum (Hook.) T.Moore; Index Filic. (T. Moore) 100 (1858) (non C. Presl (1825))
= Nephrodium pennigerum Hook.; Sp. Fil. [W. J. Hooker] 4: 82 (1862), nom. nov. for Aspidium pennigerum Blume, non Nephrodium pennigerum (G. Forst.) C. Presl 1825, nec al., nec Thelypteris pennigera (G. Forst.) Allan, Fl. New Zealand 1: 51. 1961
= Nephrodium pennigerum var. multilineatum C.B.Clarke; Trans. Linn. Soc. London, Bot. 1: 532 (1880)
= Thelypteris megaphylla (Mett.) K.Iwats.; Mem. Coll. Sci. Univ. Kyoto, B, Biol. 31: 34 (1964)
variant :
var. excellens
(Blume) Holttum; Fl. Males. Ser. 2,1(5): 462 (1982)
Distribution: Java
42 JAW
Conservation Status: Only known from type
Synonyms: hidden
= Aspidium excellens Blume; Enum. Pl. Javae 2: 160 (1828)
= Dryopteris excellens (Blume) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Nephrodium excellens (Blume) Hook.; Sp. Fil. 4: 63 (1862)
= Pleocnemia excellens (Blume) Alderw.; Handb. 171 (1908)
= Proferea excellens (Blume) C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6: 259 (1851)
= Tectaria excellens (Blume) C.Chr.; Index Filic., Suppl. Tert.: 179 (1934)
variant :
var. karoensis
Holttum; Fl. Males. Ser. 2,1(5): 462 (1982)
Distribution: N-Sumatra
42 SUM
Conservation Status: Only known from type
species :
Sphaerostephanos pentaphyllus
(Rosenst.) S.E.Fawc. & A.R.Sm.; A generic revision of the Thelypteridaceae: 82 (2021)
Distribution: New Guinea (N-Papua New Guinea); Bismarck Arch. (New Ireland)
42 NWG 43 BIS
Synonyms: hidden
= Cyclosorus pentaphyllus (Rosenst.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 143 (1947)
= Dryopteris pentaphylla Rosenst.; Repert. Spec. Nov. Regni Veg. 12: 529 (1913)
= Phegopteris pentaphylla (Rosenst.) Alderw.; Malayan Ferns Fern Allies 317 (1917)
= Pronephrium pentaphyllum (Rosenst.) Holttum; Blumea 20: 108 (1972)
= Thelypteris pentaphylla (Rosenst.) C.F.Reed; Phytologia 17: 303 (1968)
species :
Sphaerostephanos perglanduliferus
(Alderw.) Holttum; Kalikasan 4: 59 (1975)
Distribution: S-Sumatra; Borneo (Mt. Kinabalu, Sabah); SW-Sulawesi; Philippines (Negros); Palau (Babeldaob)
42 BOR PHI SUL SUM 62 PLU
Synonyms: hidden
= Cyclosorus perglanduliferus (Alderw.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 247 (1941)
= Dryopteris perglandulifera Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 3, 2: 150 (1920)
= Dryopteris perglandulifera var. firmior Alderw.
= Meniscium pseudarfakianum (Hosok.) Hosok.; Trans. Nat. Hist. Soc. Formosa 32 286 (1942)
= Phegopteris pseudarfakiana Hosok.; Trans. Nat. Hist. Soc. Formosa 28: 147 (1938)
= Thelypteris perglandulifera (Alderw.) C.F.Reed; Phytologia 17: 303 (1968)
= Thelypteris pseudarfakiana (Hosok.) C.F.Reed; Phytologia 17: 306 (1968)
species :
Sphaerostephanos petiolatus
(K.Iwats. & M.Kato) S.E.Fawc. & A.R.Sm.; A generic revision of the Thelypteridaceae: 82 (2021)
Distribution: Borneo (Kalimantan)
42 BOR
Synonyms: hidden
= Thelypteris petiolata K.Iwats. & M.Kato; Acta Phytotax. Geobot. 34(4-6): 139 (1983)
species :
Sphaerostephanos pilosissimus
Holttum; Fl. Males. Ser. 2,1(5): 504 (1982)
Distribution: WC-Sumatra (G. Kerinci)
42 SUM
Synonyms: hidden
= Thelypteris haisuli Christenh.; Global Fl. 1(4): 31 (2018), nom. nov.
species :
Sphaerostephanos pilosiusculus
(Racib.) S.E.Fawc. & A.R.Sm.; A generic revision of the Thelypteridaceae: 82 (2021)
Distribution: Sumatra; Java; Lesser Sunda Isl. (Flores)
42 JAW LSI SUM
Synonyms: hidden
= Cyclosorus debilis (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 244 (1941)
= Cyclosorus pilosiusculus (Zippel ex Racib.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 247 (1941)
= Dryopteris pilosiuscula (Zippel ex Racib.) C.Chr.; Index Filic. 284 (1905)
= Dryopteris pseudoreptans C.Chr.; Index Filic. 286 (1905), nom. nov.
= Nephrodium debile Baker; J. Bot. 18: 212 (1880)
= Nephrodium pilosiusculum Zipp. ex Racib.; Fl. Buitenz. 1: 189 (1898)
= Pronephrium debile (Baker) Holttum; Blumea 20: 118 (1972)
= Thelypteris glorfindelii Christenh.; Global Fl. 1(4): 30 (2018), nom. nov. for Nephrodium debile Baker
= Thelypteris pilosiuscula (Zipp. ex Racib.) C.F.Reed; Phytologia 17: 304 (1968)
= Thelypteris pseudoreptans (C.Chr.) C.F.Reed; Phytologia 17: 306 (1968)
species :
Sphaerostephanos pilososquamatus
(Alderw.) Holttum; Fl. Males. 1: 481 (1981)
Distribution: New Guinea (Papua New Guinea); Bismarck Arch. (Admiralty Isl.)
42 NWG 43 BIS
Synonyms: hidden
= Cyclosorus megaphylloides (Rosenst.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 143 (1947)
= Cyclosorus paraphysatus (Copel.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 247 (1941)
= Cyclosorus pilososquamatus (Alderw.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 247 (1941)
= Dryopteris megaphylloides Rosenst.; Repert. Spec. Nov. Regni Veg. 12: 174 (1913)
= Dryopteris paraphysata Copel.; Philipp. J. Sci., Bot. 6: 74 (1911)
= Dryopteris pilososquamata Alderw.; Bull. Dép. Agric. Indes Néerl. 21: 4 (1908)
= Thelypteris megaphylloides (Rosenst.) C.F.Reed; Phytologia 17: 292 (1968)
= Thelypteris paraphysata (Copel.) C.F.Reed; Phytologia 17: 301 (1968)
= Thelypteris pilososquamata (Alderw.) C.F.Reed; Phytologia 17: 304 (1968)
species :
Sphaerostephanos plurifolius
(Alderw.) Holttum; Fl. Males. Ser. 2,1(5): 492 (1982)
Distribution: Sumatra
42 SUM
Synonyms: hidden
= Cyclosorus plurifolius (Alderw.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 248 (1941)
= Dryopteris plurifolia Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 3, 5: 201 (1922)
= Thelypteris plurifolia (Alderw.) C.F.Reed; Phytologia 17: 305 (1968)
species :
Sphaerostephanos plurivenosus
Holttum; Fl. Males. Ser. 2,1(5): 496 (1982)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Thelypteris plurivenosa (Holttum) Christenh.; Global Fl. 1(4): 36 (2018)
species :
Sphaerostephanos polisianus
Holttum; Kalikasan 4: 65 (1975)
Distribution: Philippines (Luzon)
42 PHI
Conservation Status: Only known from type
Synonyms: hidden
= Thelypteris polisiana (Holttum) Christenh.; Global Fl. 1(4): 36 (2018)
species :
Sphaerostephanos polycarpos
(Blume) Copel.; Univ. Calif. Publ. Bot. 16: 60 (1929)
Distribution: Nicobar Isl.; peninsular Thailand; S-Thailand; Vietnam; peninsular Malaysia; Singapore; Sumatra; Borneo (Mt. Kinabalu, Sabah); Sulawesi; Philippines; Java; Lesser Sunda Isl. (Lombok, Bali); New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch.; Solomon Isl. (Bougainville, Gizo, Guadalcanal, Kolombangara, Makira, Rendova, Vella Lavella); Santa Cruz Isl.; [Western] Samoa (´Upolu)
41 NCB THA VIE 42 BOR JAW LSI MLY NWG PHI SNG SUL SUM 43 BIS SOL 60 SAM SCZ
Conservation Status: IUCN EN for India (Fraser-Jenk. 2012). Vulnerable (Singapore)
Synonyms: hidden
= Aspidium heterodon Blume; Enum. Pl. Javae 2: 157 (1828)
= Aspidium javanicum (R.Br. ex Mett.) Mett.; Pheg. u. Asp. 103, n. 245 (1858)
= Aspidium perakense Bedd.; J. Bot. 26: 4 (1888)
= Aspidium polycarpon Blume; Enum. Pl. Javae 2: 156 (1828)
= Cyclosorus perakensis (Bedd.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 247 (1941)
= Cyclosorus perakensis var. sumatrensis Alderw.; Bull. Jard. Bot. Buitenz. II: 13 (1913)
= Cyclosorus polycarpos (Blume) Holttum; Rev. Fl. Mal. 2: 283, f. 164 (1955)
= Didymochlaena polycarpos (Blume) Baker; Hook. & Baker, Syn. Fil. 248 (1867)
= Dryopteris microchlamys (de Vriese) Alderw.; Malayan Ferns 226 (1909)
= Dryopteris perakensis (Bedd.) C.Chr.; Index Filic. 284 (1905)
= Dryopteris polycarpos (Blume) Christ; Philipp. J. Sci., Bot. 2: 202 (1907)
= Lastrea microchlamys de Vriese; Tijdschr. Wis.- Natuurk. Wetensch. Eerste Kl. Kon. Ned. Inst. Wetensch. 1: 155 (1848)
= Mesochlaena asplenioides (J.Sm. ex Hook.) J.Sm.; J. Bot. (Hooker) 3: 18 (1840)
= Mesochlaena javanica (Fée) R.Br. ex Mett.; Fil. Hort. Bot. Lips.: 96 (1856)
= Mesochlaena polycarpos (Blume) Bedd.; Handb. Ferns Brit. India, Suppl. 13 (1876)
= Mesochlaena toppingii Copel.; Philipp. J. Sci., C 12: 57 (1917)
= Nephrodium heterodon (Blume) T.Moore; Index Filic. (T. Moore) 93 (1858)
= Nephrodium javanicum (R.Br.) Hook.; Fil. Exot. t. 61 (1859)
= Nephrodium microchlamys Baker; J. Linn. Soc., Bot. 15: 107 (1876), non de Vriese 1848
= Nephrodium perakense (Bedd.) Baker; Ann. Bot. (Oxford) 5: 319 (1891)
= Nephrodium polycarpon (Blume) Keyserl.; Polyp. Herb. Bunge (1873)
= Sphaerostephanos asplenioides J.Sm. ex Hook.; Gen. Fil.: t. 24 (1839)
= Sphaerostephanos toppingii (Copel.) C.Chr.; Index Filic., Suppl. Tert.: 172 (1934)
= Sphaerostephanos unijugus Copel.; Philipp. J. Sci.60: 109 (1936)
= Stegnogramma asplenioides (J.Sm. ex Hook.) Fée; Mém. Foug. 5. (Gen. Filic.) 204 (1852)
= Stegnogramma javanica (R.Br.) Fée; Mém. Foug. 5 (Gen. Filic.): 204 (1852)
= Thelypteris perakensis (Bedd.) C.F.Reed; Phytologia 17: 303 (1968)
= Thelypteris polycarpos (Blume) K.Iwats.; Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 31: 32 (1964)
species :
Sphaerostephanos polyotis
(C.Chr.) Holttum; Fl. Males. Ser. 2,1(5): 464 (1982)
Distribution: SW-Sulawesi
42 SUL
Synonyms: hidden
= Cyclosorus polyotis (C.Chr. ex Kjellb. & C.Chr.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 248 (1941)
= Dryopteris polyotis C.Chr.; Bot. Jahrb. Syst. 66: 46 (1933)
= Thelypteris polyotis (C.Chr. ex Kjellb. & C.Chr.) C.F.Reed; Phytologia 17: 305 (1968)
species :
Sphaerostephanos porphyricola
(Copel.) Holttum; Kalikasan 4: 59 (1975)
Distribution: Borneo (Sarawak, Mt. Kinabalu); Philippines; Moluccas (Sula Arch.: Tawi-Tawi)
42 BOR MOL PHI
Synonyms: hidden
= Cyclosorus porphyricola (Copel.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 248 (1941)
= Dryopteris porphyricola Copel.; Philipp. J. Sci., Bot. 7: 60 (1912)
= Thelypteris porphyricola (Copel.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 6: 287 (nomen tantum) (1936)
species :
Sphaerostephanos posthumi
Holttum; Fl. Males. Ser. 2,1(5): 493 (1982) (as ´posthumii´)
Distribution: Lesser Sunda Isl. (Flores)
42 LSI
Conservation Status: Only known from type
Synonyms: hidden
= Thelypteris posthumi (Holttum) Christenh.; Global Fl. 1(4): 36 (2018)
species :
Sphaerostephanos potamios
Holttum; Fl. Males. Ser. 2,1(5): 502 (1982)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Conservation Status: Only known from type
Synonyms: hidden
= Thelypteris potamia (Holttum) Christenh.; Global Fl. 1(4): 36 (2018)
species :
Sphaerostephanos productus
(Kaulf.) Holttum; Kalikasan 4: 59 (1975)
Distribution: Taiwan (Lan Yu Isl.); Palawan; Philippines (Luzon, Mindanao)
38 TAI 42 PHI PLW
Synonyms: hidden
= Aspidium productum Kaulf.; Enum. Filic. Chamisso: 237 (1824)
= Cyclosorus basilaris (C.Presl) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 235 (1938)
= Cyclosorus kotoensis (Hayata) W.C.Shieh; J. Sci. Engin. (Natl. Chung-Hsing Univ.) 13: 45 (1976)
= Cyclosorus leucadenius Copel.; Philipp. J. Sci. 81: 27 (1952)
= Cyclosorus philippinensis (Baker) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 143 (1947)
= Cyclosorus productus (Kaulf.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 248 (1941)
= Cyclosorus pseudohirsutus (Rosenst.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 248 (1941)
= Cyclosorus serratus Copel.; Philipp. J. Sci. 81: 36 (1952)
= Cyclosorus truncatus var. kotoensis (Hayata) H.Itô; Bot. Mag. (Tokyo) 51: 729 (1937)
= Cyclosorus weberi Copel.; Philipp. J. Sci. 81: 36, t. 25 (1952)
= Dryopteris basilaris (C.Presl) C.Chr.; Index Filic. 254 (1905) ex Christ, Philipp. J. Sci., Bot. 2. 196 (1907)
= Dryopteris kotoensis Hayata; Icon. Pl. Formosan. 5: 279, f. 107 (1915)
= Dryopteris luzonica var. puberula Christ; Philipp. J. Sci., Bot. 2: 197 (1907)
= Dryopteris philippinensis (Baker) Copel.; Philipp. J. Sci. 56: 100 (1935)
= Dryopteris producta (Kaulf.) C.Chr.; Index Filic. 286 (1905)
= Dryopteris pseudohirsuta Rosenst.; Meded. Rijks Herb. no. 31: 7 (1917)
= Nephrodium basilare C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 258 (1851)
= Nephrodium caudiculatum J.Sm.; J. Bot. (Hooker) 3: 411 (1841)
= Nephrodium philippinense Baker; Ann. Bot. (Oxford) 5: 327 (1891) (non Phegopteris philippinensis (Fée) Mett. (1858))
= Nephrodium productum (Kaulf.) C.Presl; Tent. Pterid. 81 (1836)
= Sphaerostephanos kotoensis (Hayata) Holttum; C. M. Kuo, Fl. Taiwan 1: 436 (1975) ex Kuo, H. L. Li et al., Fl. Taiwan 1: 436 (1975)
= Sphaerostephanos punctatus Holttum; Kalikasan 4: 60 (1975)
= Thelypteris endorica Christenh.; Global Fl. 1(4): 29 (2018), nom. nov. for Cyclosorus serratus Copel.
= Thelypteris kotoensis (Hayata) K.Iwats.; Acta Phytotax. Geobot. 21: 42 (1964)
= Thelypteris leucadenia (Copel.) C.F.Reed; Phytologia 17: 287 (1968)
= Thelypteris producta (Kaulf.) C.F.Reed; Phytologia 17: 306 (1968)
= Thelypteris pseudohirsuta (Rosenst.) C.F.Reed; Phytologia 17: 306 (1968)
= Thelypteris ugoensis C.F.Reed; Phytologia 17: 322 (1968), nom. nov.
= Thelypteris weberi (Copel.) C.F.Reed; Phytologia 17: 324 (1968)
species :
Sphaerostephanos pseudomegaphyllus
(Alderw.) Holttum; Fl. Males. Ser. 2,1(5): 452 (1982)
Distribution: Sumatra (Padang Highlands)
42 SUM
Synonyms: hidden
= Aspidium megaphyllum var. hirsutum Mett.; Ann. Mus. Bot. Lugduno-Batavi 1: 233 (1863)
= Dryopteris megaphylloides Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 20: 16 (1915) (non Rosenst. (1913))
= Dryopteris pseudomegaphylla Alderw.; Malayan Ferns Fern Allies 180 (1917)
= Thelypteris pseudomegaphylla (Alderw.) Christenh.; Global Fl. 1(4): 36 (2018)
species :
Sphaerostephanos pterosporus
(Alderw.) Holttum; Fl. Males. Ser. 2,1(5): 486 (1982)
Distribution: peninsular Malaysia (several localities); S-Sumatra; Borneo (W-Sarawak)
42 BOR MLY SUM
Synonyms: hidden
= Cyclosorus pterosporus (Alderw.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 248 (1941)
= Dryopteris bungoensis C.Chr.; Dansk Bot. Ark. 9, pl. 3: 59 (1937)
= Dryopteris pterospora Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 3, 2: 148 (1920)
= Thelypteris pterospora (Alderw.) C.F.Reed; Phytologia 17: 307 (1968)
variant :
var. altilobus
Holttum; Fl. Males. Ser. 2,1(5): 487 (1982)
Distribution: peninsular Malaysia (Perak)
42 MLY
variant :
var. crenatus
Holttum; Fl. Males. Ser. 2,1(5): 487 (1982)
Distribution: Borneo (Sarawak); Brunei
42 BOR BRN
species :
Sphaerostephanos pullenii
Holttum; Fl. Males. Ser. 2,1(5): 480 (1982)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Thelypteris pullenii (Holttum) Christenh.; Global Fl. 1(4): 36 (2018)
species :
Sphaerostephanos pycnosorus
(C.Chr.) Holttum; Allertonia 1(3): 204 (1977)
Distribution: American Samoa (Ta´u); [Western] Samoa
60 ASA SAM
Synonyms: hidden
= Dryopteris pycnosora C.Chr.; Bull. Bernice P. Bishop Mus. 177: 83 (1943)
= Thelypteris pycnosora (C.Chr.) Christenh.; Global Fl. 1(4): 36 (2018)
species :
Sphaerostephanos reconditus
Holttum; Fl. Males. Ser. 2,1(5): 472 (1982)
Distribution: Borneo (Sarawak, Mt. Mulu)
42 BOR
Conservation Status: Only known from type
Synonyms: hidden
= Thelypteris recondita (Holttum) Christenh.; Global Fl. 1(4): 36 (2018)
species :
Sphaerostephanos reineckei
(C.Chr.) Holttum; Allertonia 1(3): 205 (1977)
Distribution: American Samoa (Tutuila, Ofu?, Ta´u); [Western] Samoa (Savai´i, ´Upolu)
60 ASA SAM
Synonyms: hidden
= Dryopteris reineckei C.Chr.; Bull. Bernice P. Bishop Mus. 177: 90 (1943)
= Aspidium savaiiense Christ; Bot. Jahrb. Syst. 23: 350 (1896)
= Thelypteris reineckei (C.Chr.) Christenh.; Global Fl. 1(4): 36 (2018)
species :
Sphaerostephanos richardsii
(Baker) Holttum; Allertonia 1(3): 205 (1977)
Distribution: Solomon Isl. (Kolombangara, Makira, Malaita, Santa Ysabel); New Caledonia; Vanuatu
43 SOL 60 NWC VAN
Synonyms: hidden
= Aspidium herusianum E.Fourn.; Ann. Sci. Nat., Bot., sér. V, 18: 295 (1873) (nomen))
= Cyclosorus epaleatus (C.Chr.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 244 (1941)
= Cyclosorus richardsii (Baker) Brownlie; Fl. Nouv.-Caléd. 3: 218 (1969)
= Dryopteris epaleata C.Chr.; Index Filic., Suppl. Tert.: 85 (1934)
= Dryopteris francii Copel.; Univ. Calif. Publ. Bot. 14: 357 (1929) (non C. Chr. (1925))
= Dryopteris richardsii (Baker) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Lastrea richardsii (Baker) T.Moore; Gard. Chron., n.s., 15: 104 (1881)
= Lastrea richardsii var. multifida T.Moore; Gard. Chron., n.s., 15: 104 (1881)
= Nephrodium richardsii Baker; Hook. & Baker, Syn. Fil. ed. 2: 494 (1874)
= Nephrodium richardsii var. multifidum (T.Moore) J.H.Veitch; Hort. Veitch.: 325 (1906)
= Thelypteris epaleata (C.Chr.) C.F.Reed; Phytologia 17: 275 (1968)
= Thelypteris herusiana (E.Fourn.) Brownlie; (1969)
= Thelypteris richardsii (Baker) Christenh.; Global Fl. 1(4): 36 (2018)
species :
Sphaerostephanos rigidus
(Ridl.) Holttum; Fl. Males. Ser. 2,1(5): 484 (1982)
Distribution: New Guinea (Irian Jaya)
42 NWG
Conservation Status: Only known from 2 collections
Synonyms: hidden
= Cyclosorus rigidus (Ridl.) Copel.; Philipp. J. Sci. 78: 452 (1951)
= Cyclosorus wollastonii (Alderw.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 143 (1947)
= Dryopteris ridleyana Brause; Bot. Jahrb. Syst. 56: 112 (1920) (non Dryopteris ridleyana Underw.)
= Dryopteris wollastonii (Alderw.) C.Chr.; Index Filic., Suppl. Tert.: 101 (1934)
= Goniopteris rigida Ridl.; Trans. Linn. Soc. London, Bot. 9: 258 (1916)
= Phegopteris wollastonii Alderw.; Malayan Ferns Fern Allies 515 (1917) (nom. nov., non P. rigida (Hook. & Grev.) Mett.)
= Thelypteris rigida (Ridl.) C.F.Reed; Phytologia 17: 309 (1961)
= Thelypteris wollastonii (Alderw.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 255 (1941)
species :
Sphaerostephanos roemerianus
(Rosenst.) Holttum; Fl. Males. Ser. 2,1(5): 499 (1982)
Distribution: New Guinea (Irian Jaya)
42 NWG
Conservation Status: Only known from type
Synonyms: hidden
= Cyclosorus roemerianus (Rosenst.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 143 (1947)
= Dryopteris roemeriana Rosenst.; Nova Guinea 8: 723 (1912)
= Thelypteris roemeriana (Rosenst.) C.F.Reed; Phytologia 17: 310 (1968)
species :
Sphaerostephanos rudis
(Ridl.) Holttum; Fl. Males. Ser. 2,1(5): 501 (1982)
Distribution: New Guinea (Irian Jaya: incl. Yapen Isl.)
42 NWG
Synonyms: hidden
= Dryopteris rufopilosa Brause; Bot. Jahrb. Syst. 56: 106 (1920), nom. nov.
= Goniopteris rudis Ridl.; Trans. Linn. Soc., Bot. 9: 259 (1916) (non Dryopteris rudis C. Chr.)
= Phegopteris ridleyana Alderw.; Malayan Ferns Fern Allies 515 (1917) (non Dryopteris ridleyana Brause (1920))
= Thelypteris ridleyana (Alderw.) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 2: 465 (2018)
= Thelypteris valinorica Christenh.; Global Fl. 1(4): 39 (2018), nom. nov. superfl. for Goniopteris rudis Ridl.
variant :
var. micans
(Brause) Holttum; Fl. Males. Ser. 2,1(5): 501 (1982)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Synonyms: hidden
= Cyclosorus micans (Brause) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 143 (1947)
= Dryopteris micans Brause; Bot. Jahrb. Syst. 56: 98 (1920)
= Thelypteris micans (Brause) C.F.Reed; Phytologia 17: 293 (1968)
species :
Sphaerostephanos sagittifolius
(Blume) Holttum; Fl. Males. Ser. 2,1(5): 487 (1982)
Distribution: Sumatra; Java; Lesser Sunda Isl. (Lombok, Flores)
42 JAW LSI SUM
Synonyms: hidden
= Aspidium sagittifolium Blume; Enum. Pl. Javae 2: 153 (1828)
= Cyclosorus sagittifolius (Blume) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 248 (1941)
= Dryopteris sagittifolia (Blume) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Nephrodium sagittifolium (Blume) T.Moore; Index Filic. (T. Moore) 103 (1858)
= Thelypteris sagittifolia (Blume) C.F.Reed; Phytologia 17: 311 (1968)
species :
Sphaerostephanos santomasii
Holttum; Kalikasan 4: 62 (1975)
Distribution: Philippines (N-Luzon, Mt. San to Tomas, Mt. Nangaoto &, Mt. Data)
42 PHI
Synonyms: hidden
= Thelypteris santomasii (Holttum) Christenh.; Global Fl. 1(4): 37 (2018)
species :
Sphaerostephanos sarasinorum
Holttum; Fl. Males. Ser. 2,1(5): 498 (1982)
Distribution: C-Sulawesi
42 SUL
Synonyms: hidden
= Thelypteris sarasinorum (Holttum) Christenh.; Global Fl. 1(4): 37 (2018)
species :
Sphaerostephanos scandens
Holttum; Allertonia 1(3): 211 (1977)
Distribution: Solomon Isl. (Guadalcanal, Kolombangara, Rendova, Santa Ysabel)
43 SOL
Synonyms: hidden
= Thelypteris scandens (Holttum) Christenh.; Global Fl. 1(4): 37 (2018)
species :
Sphaerostephanos scopulorum
(Holttum) S.E.Fawc. & A.R.Sm.; A generic revision of the Thelypteridaceae: 82 (2021)
Distribution: Moluccas (Aru Isl.); New Guinea (Irian Jaya)
42 MOL NWG
Synonyms: hidden
= Pronephrium scopulorum Holttum; Fl. Males. Ser. 2,1(5): 532 (1982)
= Thelypteris scopulorum (Holttum) Christenh.; Global Fl. 1(4): 37 (2018)
species :
Sphaerostephanos semicordatus
Holttum; Fl. Males. Ser. 2,1(5): 486 (1982)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Thelypteris semicordata (Holttum) Christenh.; Global Fl. 1(4): 37 (2018)
species :
Sphaerostephanos semimetralis
Holttum; Fl. Males. Ser. 2,1(5): 486 (1982)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Synonyms: hidden
= Thelypteris semimetralis (Holttum) Christenh.; Global Fl. 1(4): 37 (2018)
species :
Sphaerostephanos seramensis
(M.Kato) comb.ined.
Distribution: Moluccas (Seram)
42 MOL
Synonyms: hidden
= Nannothelypteris seramensis M.Kato; Acta Phytotax. Geobot. 48(1): 52 (1997)
species :
Sphaerostephanos sessilipinna
(Copel.) Holttum; Kalikasan 4: 53 (1975)
Distribution: Philippines (Negros, Mindanao)
42 PHI
Synonyms: hidden
= Cyclosorus austrophilippinus (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 142 (1947)
= Cyclosorus canlaonensis (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 142 (1947)
= Cyclosorus sessilipinnus (Copel.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 249 (1941)
= Dryopteris austrophilippina Copel.; Philipp. J. Sci.40: 300 (1929)
= Dryopteris canlaonensis Copel.; Philipp. J. Sci.40: 300 (1929)
= Dryopteris sessilipinna Copel.; Philipp. J. Sci., Bot. 6: 145 (1911)
= Thelypteris austrophilippina (Copel.) C.F.Reed; Phytologia 17: 262 (1968)
= Thelypteris canlaonensis (Copel.) C.F.Reed; Phytologia 17: 266 (1968)
= Thelypteris sessilipinna C.F.Reed; Phytologia 17: 313 (1968)
species :
Sphaerostephanos simplicifolius
(J.Sm. ex Hook.) Holttum; Kalikasan 4: 61 (1975)
Distribution: Philippines (Samar, Leyte, Biliran)
42 PHI
Synonyms: hidden
= Abacopteris simplicifolia (J.Sm.) Fée; Mém. Foug. 5. (Gen. Filic.) 310 (1852), nom. inval.
= Aspidium simplicifolium J.Sm. ex Hook.; Hooker´s Icon. Pl. 6: 10 (1854), t. 919
= Cyclosorus simplicifolius (J.Sm. ex Hook.) Copel.; J. Arnold Arbor. 30: 438 (1949)
= Dryopteris simplicifolia (J.Sm. ex Hook.) Christ; Philipp. J. Sci., Bot. 2: 206 (1907)
= Goniopteris simplicifolia (J.Sm. ex Hook.) Carruth.; Seem., Fl. Vit. 366 (1873)
= Nephrodium simplicifolium J.Sm.; J. Bot. 8: 411 (1841), nomen
= Phegopteris simplicifolia (J.Sm. ex Hook.) Alderw.; Handb. 500 (1908)
= Polypodium simplicifolium (J.Sm. ex Hook.) Hook.; Sp. Fil. 5: 2 (1863)
= Thelypteris simplicifolia (J.Sm. ex Hook.) C.F.Reed; Phytologia 17: 314 (1968)
species :
Sphaerostephanos solutus
Holttum; Fl. Males. Ser. 2,1(5): 459 (1982)
Distribution: Lesser Sunda Isl. (Timor)
42 LSI
Conservation Status: Only known from type
Synonyms: hidden
= Aspidium hispidulum var. solutum Miq.; Ann. Mus. Bot. Lugduno-Batavi 4: 160 (1868)
= Thelypteris soluta (Holttum) Christenh.; Global Fl. 1(4): 38 (2018)
species :
Sphaerostephanos spenceri
(Copel. ex Christ) Holttum; Kalikasan 4: 58 (1975)
Distribution: Philippines (E- & SE-Mindanao)
42 PHI
Synonyms: hidden
= Cyclosorus spenceri (Copel. ex Christ) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 143 (1947)
= Dryopteris spenceri Copel. ex Christ; Philipp. J. Sci., Bot. 2: 290 (1907)
= Nephrodium spenceri Copel.; ms.
= Phegopteris spenceri (Copel. ex Christ) Alderw.; Handb. 508 (1908)
= Thelypteris spenceri (Copel. ex Christ) C.F.Reed; Phytologia 17: 314 (1968)
species :
Sphaerostephanos squamatellus
Holttum; Fl. Males. Ser. 2,1(5): 497 (1982)
Distribution: Borneo (Mt. Kinabalu, Sabah)
42 BOR
Synonyms: hidden
= Thelypteris squamatella (Holttum) Christenh.; Global Fl. 1(4): 38 (2018)
species :
Sphaerostephanos stenodontus
(Copel.) Holttum; Kalikasan 4: 56 (1975)
Distribution: Philippines (Panay)
42 PHI
Conservation Status: Only known from type
Synonyms: hidden
= Cyclosorus stenodontus Copel.; Philipp. J. Sci. 81: 28, t. 20 (1952)
= Thelypteris stenodonta (Copel.) C.F.Reed; Phytologia 17: 315 (1968)
species :
Sphaerostephanos stipellatus
(Blume) Holttum; Fl. Males. Ser. 2,1(5): 492 (1982)
Distribution: peninsular Malaysia (Johore, Pahang, Trengganu, Perak); Sumatra; Java
42 JAW MLY SUM
Synonyms: hidden
= Aspidium stipellatum Blume; Enum. Pl. Javae 2: 152 (1828)
= Cyclosorus stipellatus (Blume) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 228 (1938)
= Dryopteris stipellata (Blume) Kuntze; Revis. Gen. Pl. 2: 811 (1891)
= Nephrodium stipellatum (Blume) T.Moore; Index Filic. (T. Moore) 105 (1858)
= Thelypteris stipellata (Blume) K.Iwats.; Acta Phytotax. Geobot. 21: 168 (1965)
species :
Sphaerostephanos stresemannii
Holttum; Fl. Males. Ser. 2,1(5): 490 (1982)
Distribution: Moluccas (Buru)
42 MOL
Synonyms: hidden
= Thelypteris stresemannii (Holttum) Christenh.; Global Fl. 1(4): 38 (2018)
species :
Sphaerostephanos subalpinus
(Alderw.) Holttum; Fl. Males. Ser. 2,1(5): 474 (1982)
Distribution: Moluccas (Ternate, Halmaheira, Seram?)
42 MOL
Synonyms: hidden
= Aspidium hispidulum var. ternatense Miq.; Ann. Mus. Bot. Lugduno-Batavi 4: 160 (1869)
= Cyclosorus subalpinus (Alderw.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10 249 (1941)
= Dryopteris subalpina Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 3,5: 200 (1922)
= Polypodium acutum Roxb.; Calcutta J. Nat. Hist. 4: 492 (1844)
= Thelypteris subalpina (Alderw.) C.F.Reed; Phytologia 17: 317 (1968)
species :
Sphaerostephanos subappendiculatus
(Copel.) S.E.Fawc. & A.R.Sm.; A generic revision of the Thelypteridaceae: 82 (2021)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Cyclosorus subappendiculatus (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 143 (1947)
= Dryopteris subappendiculata Copel.; Univ. Calif. Publ. Bot. 18: 220 (1942)
= Pneumatopteris subappendiculata (Copel.) Holttum; Blumea 21: 303 (1973)
= Thelypteris subappendicuata (Copel.) C.F.Reed; Phytologia 17: 317 (1968)
species :
Sphaerostephanos subcanescens
Holttum; Blumea 33(2): 404 (1988)
Distribution: Sulawesi
42 SUL
Synonyms: hidden
= Thelypteris subcanescens (Holttum) Christenh.; Global Fl. 1(4): 38 (2018)
species :
Sphaerostephanos subcordatus
Holttum; Fl. Males. Ser. 2,1(5): 469 (1982)
Distribution: Philippines (W-Samar)
42 PHI
Conservation Status: Only known from type
Synonyms: hidden
= Thelypteris subcordata (Holttum) Christenh.; Global Fl. 1(4): 38 (2018)
species :
Sphaerostephanos suboppositus
Holttum; Fl. Males. Ser. 2,1(5): 498 (1982)
Distribution: N-Sulawesi
42 SUL
Synonyms: hidden
= Thelypteris subopposita (Holttum) Christenh.; Global Fl. 1(4): 38 (2018)
species :
Sphaerostephanos subpectinatus
(Copel.) Holttum; Allertonia 1(3): 207 (1977)
Distribution: Cook Isl. (Rarotonga); Marquesas Isl. (Ua Huka, Fatu Hiva, Ua Pou, Hiva Oa, Tahuata); Society Isl. (Tahiti, Raiatea, Moorea)
61 COO MRQ SCI
Synonyms: hidden
= Cyclosorus subpectinatus (Copel.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 249 (1941)
= Dryopteris subpectinata Copel.; Bull. Bernice P. Bishop Mus. 93, t. 7 D (1932)
= Thelypteris subpectinata (Copel.) C.F.Reed; Phytologia 17: 317 (1968)
species :
Sphaerostephanos subtruncatus
(Bory) Holttum; Holttum & P. Chandra, Kew Bull. 26: 80 (1971)
Distribution: Seychelles (Silhouette Isl.); India (Goa, Karnataka, Kerala, Tamil Nadu); Sri Lanka
29 SEY 40 IND SRL
Synonyms: hidden
= Cyclosorus wynadensis (B.K.Nayar & Geev.) Mazumdar & R.Mukhop.; Bionature (Bhopal) 33(1): 34 (2013)
= Dryopteris mauritiana var. gardineri C.Chr.; Trans. Linn. Soc. London, Bot. 7(19): 413 (1912)
= Polypodium subtruncatum Bory; Bél., Voy. Indes Or., Bot. 2: 32 (1833)
= Sphaerostephanos wynadensis B.K.Nayar & Geev.; Bull. Bot. Surv. India 28(1-4): 137 (1986 publ. 1988) (fide Fraser-Jenkins 1997)
= Thelypteris subtruncata (Bory) Christenh.; Global Fl. 1(4): 38 (2018), isonym
= Thelypteris subtruncata (Bory) Panigrahi; Phytologia 31: 371 (1975)
species :
Sphaerostephanos subulifolius
(Alderw.) Holttum; Fl. Males. Ser. 2,1(5): 494 (1982)
Distribution: S-Sumatra
42 SUM
Synonyms: hidden
= Dryopteris subulifolia Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 28: 22 (1918)
= Thelypteris subulifolia (Alderw.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 254 (1941)
species :
Sphaerostephanos sudesticus
Holttum; Fl. Males. Ser. 2,1(5): 448 (1982)
Distribution: New Guinea (Louisiade Arch.: Sudest Isl.)
42 NWG
Conservation Status: Only known from type
Synonyms: hidden
= Thelypteris sudestica (Holttum) Christenh.; Global Fl. 1(4): 38 (2018)
species :
Sphaerostephanos superbus
(Brause) S.E.Fawc. & A.R.Sm.; A generic revision of the Thelypteridaceae: 82 (2021)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Synonyms: hidden
= Cyclosorus deltipterus (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 142 (1947)
= Cyclosorus superbus (Brause) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 249 (1941)
= Dryopteris deltiptera Copel.; Univ. Calif. Publ. Bot. 18 (1942)
= Dryopteris superba Brause; Bot. Jahrb. Syst. 56: 105 (1920)
= Pneumatopteris superba (Brause) Holttum; Blumea 21: 303 (1973)
= Thelypteris deltiptera (Copel.) C.F.Reed; Phytologia 17: 271 (1968)
= Thelypteris superba (Brause) C.F.Reed; Phytologia 17: 318 (1968)
species :
Sphaerostephanos taiwanensis
(C.Chr.) Holttum ex Kuo; H. L. Li et al., Fl. Taiwan 1: 436 (1975)
Distribution: China (Fujian, Guangdong, Guangxi, Jiangxi); Ryukyu Isl.; Taiwan; Palau
36 CHI 38 NNS TAI 62 PLU
Synonyms: hidden
= Aspidium lobulatum Christ; Bull. Herb. Boissier, sér, 2, 4: 614 (1904) (non Blume (1828))
= Christella taiwanensis (C.Chr.) Holttum; (1975)
= Cyclosorus subheterocarpus Ching; (1976), nom. nud.
= Cyclosorus taiwanensis (C.Chr.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 181. (1938), isonym
= Cyclosorus taiwanensis (C.Chr.) H.Itô; Bot. Mag. (Tokyo) 51: 728, f. 2 (1937)
= Cyclosorus tungkooshanensis Ching; (1976), nom. nud.
= Dryopteris subhispidula Rosenst.; Hedwigia 66: 343 (1915)
= Dryopteris taiwanensis C.Chr.; Index Filic. 297 (1905)
= Thelypteris taiwanensis (C.Chr.) K.Iwats.; Mem. Coll. Sci. Univ. Kyoto, B, Biol. 31: 183 (1965)
species :
Sphaerostephanos tamiensis
(Brause) comb.ined.
Distribution: New Guinea
42 NWG
Remarks: status uncertain, type incomplete. Holttum never published the combination in Sphaerostephanos but mentioned that it belongs there.
Synonyms: hidden
= Dryopteris tamiensis Brause; Bot. Jahrb. Syst. 49: 23 (1912)
species :
Sphaerostephanos tandikatensis
(Alderw.) Holttum; Fl. Males. Ser. 2,1(5): 463 (1982)
Distribution: C-Sumatra; S-Sumatra
42 SUM
Synonyms: hidden
= Dryopteris tandikatensis Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 11: 11 (1913)
= Thelypteris tandikatensis (Alderw.) Christenh.; Global Fl. 1(4): 39 (2018)
species :
Sphaerostephanos telefominicus
Holttum; Fl. Males. Ser. 2,1(5): 499 (1982)
Distribution: New Guinea (Papua New Guinea: Telefomin)
42 NWG
Conservation Status: Only known from type
Synonyms: hidden
= Thelypteris telefominica (Holttum) Christenh.; Global Fl. 1(4): 39 (2018)
species :
Sphaerostephanos tephrophyllus
(Copel.) Holttum; Kalikasan 4: 58 (1975)
Distribution: Philippines (Mindanao)
42 PHI
Synonyms: hidden
= Cyclosorus tephrophyllus (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 143 (1947)
= Dryopteris tephrophylla Copel.; Philipp. J. Sci. 40: 296 (1929)
= Thelypteris tephrophylla (Copel.) C.F.Reed; Phytologia 17: 319 (1968)
species :
Sphaerostephanos tibangensis
(C.Chr.) Holttum; Fl. Males. Ser. 2,1(5): 485 (1982)
Distribution: E-Borneo (Mt. Tibang)
42 BOR
Synonyms: hidden
= Dryopteris tibangensis C.Chr.; Dansk Bot. Ark. 9(3): 66 (1937)
= Pronephrium tibangense (C.Chr.) Holttum; Blumea 20: 118 (1972)
= Thelypteris tibangensis (C.Chr.) Christenh.; Global Fl. 1(4): 39 (2018)
species :
Sphaerostephanos trichochlamys
Holttum; Fl. Males. Ser. 2,1(5): 465 (1982)
Distribution: Borneo (Sabah, Mt. Kinabalu, Sarawak, G. Mulu)
42 BOR
Synonyms: hidden
= Thelypteris trichochlamys (Holttum) Christenh.; Global Fl. 1(4): 39 (2018)
species :
Sphaerostephanos trimetralis
Holttum; Fl. Males. Ser. 2,1(5): 451 (1982)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Conservation Status: Only known from type
Synonyms: hidden
= Thelypteris trimetralis (Holttum) Christenh.; Global Fl. 1(4): 39 (2018)
species :
Sphaerostephanos uaniensis
Holttum; Fl. Males. Ser. 2,1(5): 494 (1982)
Distribution: New Guinea (New Britain)
42 NWG
Conservation Status: Only known from type
Synonyms: hidden
= Thelypteris uaniensis (Holttum) Christenh.; Global Fl. 1(4): 39 (2018)
species :
Sphaerostephanos uniauriculatus
(Copel.) Holttum; Fl. Males. Ser. 2,1(5): 467 (1982)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Conservation Status: Only known from type
Synonyms: hidden
= Cyclosorus uniauriculatus (Copel.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 250 (1941)
= Dryopteris uniauriculata Copel.; Philipp. J. Sci., Bot. 9: 3 (1914)
= Thelypteris uniauriculata (Copel.) C.F.Reed; Phytologia 17: 322 (1968)
species :
Sphaerostephanos urdanetensis
(Copel.) Holttum; Fl. Males. Ser. 2,1(5): 467 (1982)
Distribution: Philippines (Mindanao)
42 PHI
Synonyms: hidden
= Cyclosorus matutumensis (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 143 (1947)
= Cyclosorus urdanetensis (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 143 (1947)
= Dryopteris matutumensis Copel.; Philipp. J. Sci. 40: 299, t. 3 (1929)
= Dryopteris urdanetensis Copel.; Leafl. Philipp. Bot. 6: 1682 (1913)
= Thelypteris matutumensis (Copel.) C.F.Reed; Phytologia 17: 291 (1968)
= Thelypteris urdanetensis (Copel.) C.F.Reed; Phytologia 17: 322 (1968)
species :
Sphaerostephanos veitchii
Holttum; Allertonia 1: 204 (1977)
Distribution: New Guinea (Papua New Guinea); Bismarck Arch. (New Britain); Solomon Isl. (Bougainville, Gizo, Guadalcanal, Santa Ysabel, Kolombangara, Makira, Malaita, Mono, Nggelas, Rendova, Russells, Ulawa, Makira, Nggatokae, Ranongga, Tetepare, Vangunu, Vella Lavella)
42 NWG 43 BIS SOL
Synonyms: hidden
= Nephrodium amboinense var. subglandulosum Baker; J. Linn. Soc., Bot. 19: 295 (1882)
= Thelypteris veitchii (Holttum) Christenh.; Global Fl. 1(4): 39 (2018)
species :
Sphaerostephanos vestigiatus
(Copel.) Holttum; Fl. Males. Ser. 2,1(5): 453 (1982)
Distribution: New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch. (Admiralty Isl.)
42 NWG 43 BIS
Synonyms: hidden
= Cyclosorus vestigiatus (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 143 (1947)
= Dryopteris vestigiata Copel.; Univ. Calif. Publ. Bot. 18: 220 (1942)
= Thelypteris vestigiata (Copel.) C.F.Reed; Phytologia 17: 323 (1968)
species :
Sphaerostephanos warburgii
(Kuhn & C.Chr.) Holttum; Allertonia 1(3): 202 (1977)
Distribution: C-Sulawesi; SE-Sulawesi; Moluccas (Tanimbar); New Guinea (Papua New Guinea); Bismarck Arch. (Admiralty Isl., Admiralty Isl.); Guam
42 MOL NWG SUL 43 BIS 62 GUM
Synonyms: hidden
= Aspidium warburgii Kuhn & Christ; Warb., Monsunia 1: 81 (1900)
= Cyclosorus warburgii (Kuhn & Christ ex Christ) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 250 (1941)
= Dryopteris depauperata Copel.
= Dryopteris calcicola C.Chr.; Bot. Jahrb. Syst. 66: 44 (1933)
= Dryopteris warburgii (Kuhn & Christ) C.Chr.; Index Filic. 98 (1905)
= Thelypteris warburgii (Kuhn & Christ) B.C.Stone; Micronesica 2: 3 (1966)
species :
Sphaerostephanos wauensis
Holttum; Fl. Males. Ser. 2,1(5): 505 (1982)
Distribution: New Guinea (Papua New Guinea: Morobe District)
42 NWG
Synonyms: hidden
= Thelypteris wauensis (Holttum) Christenh.; Global Fl. 1(4): 39 (2018)
species :
Sphaerostephanos williamsii
(Copel.) Holttum; Kalikasan 4: 53 (1975)
Distribution: Philippines (Mindanao)
42 PHI
Conservation Status: Only known from type
Synonyms: hidden
= Dryopteris williamsii Copel.; Brittonia 1: 67, pl. 1 (1931)
= Lastrea williamsii (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 140 (1947)
= Thelypteris williamsii (Copel.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 255 (1941)
species :
Sphaerostephanos woitapensis
Holttum; Fl. Males. Ser. 2,1(5): 483 (1982)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Thelypteris woitapensis (Holttum) Christenh.; Global Fl. 1(4): 40 (2018)
species :
Sphaerostephanos womersleyi
(Holttum) S.E.Fawc. & A.R.Sm.; A generic revision of the Thelypteridaceae: 82 (2021)
Distribution: Sulawesi; Moluccas (Seram); New Guinea (Papua New Guinea); Bismarck Arch. (New Ireland); Solomon Isl. (Guadalcanal); ?Fiji
42 MOL NWG SUL 43 BIS SOL
Synonyms: hidden
= Pronephrium womersleyi Holttum; Blumea 20: 108 (1972)
= Thelypteris womersleyi (Holttum) Christenh.; Global Fl. 1(4): 40 (2018)
