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Ver. 19.2
April 17th, 2024

044.0000 Family Thelypteridaceae Ching ex Pic.Serm.; Webbia 24: 711 (1970)
5-34 genera (depending on genus concept), 1100-1200 spp. New genus concept adopted here from Fawcett & Smith (2021), based on molecular results.
Species listed in database: 1180; hybrids: 38
044.2038 Genus Pneumatopteris Nakai; Bot. Mag. (Tokyo) 47: 179 (1933)
Distribution: Malesia, Pacific. Many species need transfer into other genera.
Species listed in database: 31
= Thelypteris subgen.Pneumatopteris K.Iwats.; Mem. Coll. Sci. Univ. Kyoto, B, Biol. 31: 33 (1964), excl. sect. Macrocyclosorus
= Cyclosorus subgen.Pneumatopteris (Nakai) Panigrahi; Abstr. & Souv. Nation. Symp. Curr. Trends Pterid. 4-6 October 1991: 13 (1991)
species :
Pneumatopteris aberrans Holttum; Blumea 33(2): 401 (1988)
Distribution: Sulawesi
42 SUL
Remarks: generic placement uncertain
Synonyms: hidden
= Thelypteris aberrans (Holttum) Christenh.; Global Fl. 1(4): 25 (2018)
species :
Pneumatopteris auctipinna Holttum; Blumea 21: 310 (1973)
Distribution: C-Sulawesi; N-Sulawesi; E-Java; Lesser Sunda Isl. (Flores); Moluccas (Buru, Ambon)
42 JAW LSI MOL SUL
Remarks: generic placement uncertain, probably in Reholttumia
Synonyms: hidden
= Aspidium truncatum var. celebicum Miq.; Ann. Mus. Bot. Lugduno-Batavi 4: 160 (1869)
= Thelypteris auctipinna (Holttum) Christenh.; Global Fl. 1(4): 27 (2018)
species :
Pneumatopteris brooksii (Copel.) Holttum; Blumea 21: 308 (1973)
Distribution: Borneo (Sarawak)
42 BOR
Remarks: generic placement uncertain
Synonyms: hidden
= Cyclosorus brooksii (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 142 (1947)
= Dryopteris brooksii Copel.; Philipp. J. Sci., Bot. 3: 345 (1908)
= Thelypteris brooksii (Copel.) C.F.Reed; Phytologia 17: 265 (1968)
species :
Pneumatopteris callosa Nakai; Bot. Mag. (Tokyo) 47: 179 (1933)
Distribution: peninsular Malaysia (Cameron Highlands, Pahang); Sumatra; Borneo (Mt. Kinabalu, Sabah); Sulawesi; ?Java; Lesser Sunda Isl. (Bali, Flores, Timor); Moluccas (Seram)
42 BOR LSI MLY MOL SUL SUM
Remarks: Pneumatopteris s. str.
Synonyms: hidden
= Aspidium callosum Blume; Enum. Pl. Javae 2: 152 (1828)
= Aspidium multijugum Christ; Ann. Jard. Bot. Buitenz. 15: 135 (1898)
= Cyclosorus callosus (Blume) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 205 (1938)
= Dryopteris callosa (Blume) C.Chr.; Index Filic. 256 (1905)
= Goniopteris lobbiana Fée; Mém. Foug. 5 [Gen. Filic.] 251 (1852)
= Nephrodium callosum (Blume) Keyserl.; Polyp. Herb. Bunge 48 (1873)
= Thelypteris callosa (Blume) K.Iwats.; Mem. Coll. Sci. Univ. Kyoto, B, Biol. 31: 34 (1964)
species :
Pneumatopteris cheesmaniae Holttum; Blumea 21: 307 (1973)
Distribution: New Guinea (Yapen Isl.)
42 NWG
Remarks: generic placement uncertain
Conservation Status: only known from type
Synonyms: hidden
= Thelypteris cheesmaniae (Holttum) Christenh.; Global Fl. 1(4): 28 (2018)
species :
Pneumatopteris comorensis Holttum; Blumea 21: 318 (1973)
Distribution: Comores (Anjouan)
29 COM
Remarks: probably in Reholttumia
Synonyms: hidden
= Thelypteris comorensis (Holttum) Christenh.; Global Fl. 1(4): 28 (2018)
species :
Pneumatopteris dicranogramma (Alderw.) Holttum; Blumea 21: 301 (1973)
Distribution: Sumatra (G. Kerinci, G. Dempo)
42 SUM
Remarks: Pneumatopteris s. str.
Synonyms: hidden
= Cyclosorus dicranogramma (Alderw.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 244 (1941)
= Dryopteris dicranogramma Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 3, 5: 202 (1922)
= Thelypteris dicranogramma (Alderw.) C.F.Reed; Phytologia 17: 272 (1968)
species :
Pneumatopteris dilatata Holttum; Blumea 33(2): 401 (1988)
Distribution: Sulawesi
42 SUL
Remarks: Pneumatopteris s. str.
Synonyms: hidden
= Thelypteris shelob Christenh.; Global Fl. 1(4): 37 (2018), nom. nov.
species :
Pneumatopteris eburnea Holttum; Fl. Males. Ser. 2,1(5): 434 (1982)
Distribution: Borneo (Sarawak, Mt. Mulu)
42 BOR
Conservation Status: only known from type
Synonyms: hidden
= Thelypteris eburnea (Holttum) Christenh.; Global Fl. 1(4): 29 (2018)
species :
Pneumatopteris egenolfioides Holttum; Blumea 21: 306 (1973)
Distribution: New Guinea (Irian Jaya: Habbema)
42 NWG
Remarks: generic placement uncertain
Synonyms: hidden
= Thelypteris egenolfioides (Holttum) Christenh.; Global Fl. 1(4): 29 (2018)
species :
Pneumatopteris florencei (A.R.Sm. & Lorence) A.R.Sm. & Lorence; Fl. Marquesas Isl. 1: 210 (2019)
Distribution: Marquesas Isl. (Hiva Oa)
61 MRQ
Remarks: Pneumatopteris s. str.
Synonyms: hidden
= Cyclosorus florencei A.R.Sm. & Lorence; PhytoKeys 4: 40 (-42, fig. 12) (2011)
= Thelypteris florencei (A.R.Sm. & Lorence) Christenh.; Global Fl. 1(4): 30 (2018)
species :
Pneumatopteris glabra (Copel.) Holttum; Blumea 21: 301 (1973)
Distribution: Philippines (Luzon, Samar, Bohol, Mindanao)
42 PHI
Remarks: Pneumatopteris s. str.
Synonyms: hidden
= Cyclosorus glaber Copel.; Philipp. J. Sci. 81: 34 (1952) (non Thelypteris glabra (Brack.) Ching)
= Cyclosorus viridis Copel.; Philipp. J. Sci. 81: 35, pl. 24 (1952)
= Thelypteris sevillana C.F.Reed; Phytologia 17: 313 (1968), nom. nov.
= Thelypteris viridis (Copel.) C.F.Reed; Phytologia 17: 324 (1968)
species :
Pneumatopteris glandulifera (Brack.) Holttum; Blumea 21: 302 (1973)
Distribution: Solomon Isl. (Choiseul, Guadalcanal, Santa Ysabel, Kolombangara, Malaita); Vanuatu; Tonga (Tafahi); American Samoa (Tutuila, Olosega, Ta´u); Cook Isl. (Rarotonga)
43 SOL 60 ASA TON VAN 61 COO
Remarks: Pneumatopteris s. str.
Synonyms: hidden
= Aspidium glanduliferum (Brack.) Mett.; Ann. Mus. Bot. Lugduno-Batavi 1: 232 (1863)
= Aspidium glanduliferum (Brack.) Kuhn; Verh. K. K. Zool.-Bot. Ges. Wien 19: 577 (1869), isonym
= Aspidium pteroides Mett.; (1874) non Sw. 1801
= Cyclosorus glanduliferus (Brack.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 142 (1947)
= Cyclosorus obstructus (Copel.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 247 (1941)
= Dryopteris obstructa Copel.; Univ. Calif. Publ. Bot. 12: 378 (1931)
= Dryopteris subspinosa C.Chr.; Index Filic. 296 (1905)
= Dryopteris tannensis C.Chr.; Index Filic. 297 (1905)
= Goniopteris glandulifera Brack.; Expl. exp. 16: 29 (1854)
= Goniopteris tannensis (C.Chr.) Ching; Sunyatsenia 5: 239 (1940)
= Nephrodium muricatum Diels; Engl. & Prantl, Nat. Pflanzenfam. 1(4): 179 (1899)
= Pneumatopteris obstructa (Copel.) Holttum; Blumea 21: 302 (1973)
= Polypodium brackenridgei Hook.; Sp. Fil. 5: 9 (1863)
= Polypodium excelsum Baker; Hook. & Baker, Syn. Fil. ed. 2: 505 (1874) (non Desv. (1827))
= Polypodium muricatum Powell; Baker, Syn. 506 (1874) (non L. (1753))
= Thelypteris glandulifera (Brack.) C.F.Reed; Phytologia 17: 278 (1968)
= Thelypteris obstructa (Copel.) C.F.Reed; Phytologia 17: 297 (1968)
= Thelypteris tannensis (C.Chr.) C.F.Reed; Phytologia 17: 319 (1968)
variant :
var. angusta (C.Chr.) Holttum; Allertonia 1(3): 230 (1977)
Distribution: [Western] Samoa (´Upolu)
60 SAM
Synonyms: hidden
= Dryopteris subspinosa var. angusta C.Chr.; Bull. Bernice P. Bishop Mus. 177: 93 (1943)
= Nephrodium pteroides J.Sm.; Cat. Cult. Ferns 54 (1857)
species :
Pneumatopteris humbertii Holttum; Blumea 21: 313 (1973)
Distribution: Madagascar
29 MDG
Remarks: genus wrong, probably in Pseudocyclosorus-Trigonospora clade
Synonyms: hidden
= Thelypteris humbertii (Holttum) Christenh.; Global Fl. 1(4): 32 (2018)
species :
Pneumatopteris japenensis Holttum; Fl. Males. Ser. 2,1(5): 426 (1982)
Distribution: New Guinea (Yapen Isl.)
42 NWG
Remarks: probably in Reholttumia, description and material lacking
Conservation Status: only known from type
Synonyms: hidden
= Thelypteris japenensis (Holttum) Christenh.; Global Fl. 1(4): 32 (2018)
species :
Pneumatopteris latisquamata Holttum; Blumea 21: 305 (1973)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Remarks: generic placement uncertain
Conservation Status: only known from type
Synonyms: hidden
= Thelypteris latisquamata (Holttum) Christenh.; Global Fl. 1(4): 32 (2018)
species :
Pneumatopteris lithophila Holttum; Fl. Males. Ser. 2,1(5): 422 (1982)
Distribution: Philippines (W-Samar)
42 PHI
Remarks: generic placement uncertain
Conservation Status: only known from 2 collections
Synonyms: hidden
= Thelypteris lithophila (Holttum) Christenh.; Global Fl. 1(4): 33 (2018)
species :
Pneumatopteris mesocarpa (Copel.) Holttum; Blumea 21: 303 (1973)
Distribution: Society Isl. (Huahine)
61 SCI
Remarks: Pneumatopteris s. str.
Synonyms: hidden
= Cyclosorus mesocarpus (Copel.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 246 (1941)
= Dryopteris mesocarpa Copel.; Bull. Bernice P. Bishop Mus. 93: 9, t. 7 A (1932)
= Thelypteris mesocarpa (Copel.) C.F.Reed; Phytologia 17: 293 (1968)
species :
Pneumatopteris microauriculata Holttum; Blumea 21: 311 (1973)
Distribution: Borneo (Mt. Kinabalu, Sabah)
42 BOR
Remarks: genus and status uncertain, description and material lacking
Synonyms: hidden
= Thelypteris microauriculata (Holttum) K.Iwats. & M.Kato; Acta Phytotax. Geobot. 34(4-6): 137 (1983)
species :
Pneumatopteris nephrolepioides (C.Chr.) Holttum; Blumea 21: 320 (1973)
Distribution: Moluccas (Seram); New Guinea (Papua New Guinea)
42 MOL NWG
Remarks: generic placement uncertain
Synonyms: hidden
= Dryopteris nephrolepidioides C.Chr.; Brittonia 2: 295 (1937)
= Lastrea nephrolepidioides (C.Chr.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 139 (1947)
= Thelypteris nephrolepioides (C.Chr.) C.F.Reed; Phytologia 17: 295 (1968)
species :
Pneumatopteris parksii (F.Ballard) Holttum; Blumea 21: 302 (1973)
Distribution: Vanuatu; Fiji (Viti Levu, Vanua Levu, Taveuni)
60 FIJ VAN
Remarks: Pneumatopteris s. str.
Synonyms: hidden
= Cyclosorus microsorus (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 143 (1947), nom. nov. illeg.
= Dryopteris microsora Copel.; Bull. Bernice P. Bishop Mus. 59: 12 (1929), nom. nov. illeg., non Kuntze 1891
= Dryopteris parksii F.Ballard; Bull. Misc. Inform. Kew 1937: 347 (1937), nom. nov.
= Thelypteris microsora (Copel.) C.F.Reed; Phytologia 17: 293 (1968), nom. illeg.
= Thelypteris parksii (F.Ballard) Christenh.; Global Fl. 1(4): 35 (2018)
species :
Pneumatopteris patentipinna Holttum; Blumea 21: 307 (1973)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Remarks: generic placement uncertain
Synonyms: hidden
= Thelypteris patentipinna (Holttum) Christenh.; Global Fl. 1(4): 35 (2018)
species :
Pneumatopteris prismatica (Desv.) Holttum; Blumea 21: 316 (1973)
Distribution: Malawi; Mauritius; La Réunion
26 MLW 29 MAU REU
Remarks: probably in Reholttumia
Synonyms: hidden
= Aspidium caudiculatum Sieber ex Kunze; Linnaea 24: 280 (1851)
= Aspidium mascarenense Kaulf.; Spreng., Syst. 4: 101 (1827)
= Cyclosorus prismaticus (Desv.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 248 (1941)
= Cyclosorus prismaticus (Desv.) Tardieu; (1958), isonym
= Dryopteris caudiculata (Sieber ex Kunze) C.Chr.; Cat. Pl. Madagasc., Pterid.: 25 (1932)
= Dryopteris prismatica (Desv.) C.Chr.; Dansk Bot. Ark. 7: 202 (1932)
= Nephrodium caudiculatum (Sieber ex Kunze) C.Presl; Abh. Königl. Böhm. Ges. Wiss., 5, 6: 46 (1851), non J. Sm.
= Nephrodium caudiculatum Sieber; C. Presl, Tent. Pterid. 81 (1836)
= Nephrodium prismaticum Desv.; Mem. Soc. Linn. Paris 6, 2: 256 (1827)
= Thelypteris caudiculata (Sieber ex Kunze) C.F.Reed; Phytologia 17: 266 (1968)
= Thelypteris prismatica (Desv.) Schelpe; Bol. Soc. Broter. Ser. 2, 41: 217 (1967)
species :
Pneumatopteris sibelana Holttum; Blumea 21: 301 (1973)
Distribution: Moluccas (Bacan)
42 MOL
Remarks: Pneumatopteris s. str.
Conservation Status: only known from type
Synonyms: hidden
= Thelypteris sibeliana (Holttum) Christenh.; Global Fl. 1(4): 37 (2018)
species :
Pneumatopteris stokesii (E.D.Br. ex E.D.Br. & F.Br.) Holttum; Blumea 21: 302 (1973)
Distribution: Austral Isl. (Rapa Iti)
61 AUI
Remarks: Pneumatopteris s. str.
Synonyms: hidden
= Cyclosorus stokesii (E.D.Br.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 249 (1941)
= Dryopteris stokesii E.D.Br.; Bull. Bernice P. Bishop Mus. 89 [Fl. S. E. Polynesia II]: 20, f. 6 (1931)
= Thelypteris stokesii (E.D.Br. ex E. & F.Br.) C.F.Reed; Phytologia 17: 316 (1968)
species :
Pneumatopteris sumbawensis (C.Chr.) Holttum; Blumea 21: 323 (1973)
Distribution: Lesser Sunda Isl. (Sumbawa)
42 LSI
Remarks: generic placement uncertain
Conservation Status: only known from type
Synonyms: hidden
= Dryopteris sumbawensis C.Chr. ex Rensch; Hedwigia 74: 231 (1934)
= Thelypteris sumbawensis (C.Chr.) Christenh.; Global Fl. 1(4): 39 (2018)
species :
Pneumatopteris tobaica Holttum; Blumea 21: 316 (1973)
Distribution: N-Sumatra
42 SUM
Remarks: probably in Reholttumia
Conservation Status: only known from type
Synonyms: hidden
= Thelypteris tobaica (Holttum) Christenh.; Global Fl. 1(4): 39 (2018)
species :
Pneumatopteris transversaria (Brack.) Holttum; Blumea 21: 319 (1973)
Distribution: Niue; American Samoa (Tutuila?); [Western] Samoa (Savai´i, ´Upolu)
60 ASA NUE SAM
Remarks: genus and status uncertain, description and material lacking
Synonyms: hidden
= Cyclosorus transversarius (Brack.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 250 (1941)
= Dryopteris transversaria (Brack.) Brause; Bot. Jahrb. Syst. 56: 104 (1920)
= Nephrodium transversarium Brack.; U.S. Expl. Exped., Filic. 16: 187 (1864)
= Thelypteris transversaria (Brack.) C.F.Reed; Phytologia 17: 321 (1968)
species :
Pneumatopteris usambarensis Holttum; Blumea 21: 312 (1973)
Distribution: Kenya; Tanzania
25 KEN TAN
Remarks: probably in Reholttumia
Synonyms: hidden
= Thelypteris usambarensis (Holttum) Christenh.; Global Fl. 1(4): 39 (2018)
species :
Pneumatopteris venulosa (Hook.) Holttum; Blumea 21: 315 (1973)
Distribution: Bioko Isl. [Fernando Poo]; Principe Isl.; São Tomé; Annobon Isl.; Equatorial Guinea (Rio Muni); ?Chad
23 ANN BIO EQG PRI SAO
Remarks: probably in Reholttumia
Conservation Status: IUCN R
Synonyms: hidden
= Aspidium elatum Mett.; Kuhn, Filic. Afr. 130 (1868) (non Bojer (1837))
= Cyclosorus elatus (Mett.) Alston; Bol. Soc. Broter., ser. 2, 30: 13 (1956)
= Cyclosorus venulosus (Hook.) Tardieu; Notul. Syst. (Paris) 14: 345 (1953)
= Dryopteris elata (Mett.) C.Chr.; Index Filic. 263 (1905) (non Kuntze (1891))
= Dryopteris venulosa Kuntze; Revis. Gen. Pl. 2: 814 (1891), nom. nov. for Nephrodium venulosum Hook., non Desv. (1827)
= Nephrodium venulosum Hook.; Sp. Fil. 4: 71 (1862) (non Desv. (1827))
= Thelypteris elata (Mett.) Schelpe; J. S. Afr. Bot. 31: 265. 1965
= Thelypteris venulosa (Kuntze) C.F.Reed; Phytologia 17: 323 (1968)
species :
Pneumatopteris versteeghii Holttum; Blumea 21: 321 (1973)
Distribution: New Guinea (Irian Jaya)
42 NWG
Remarks: generic placement uncertain
Conservation Status: only known from type
Synonyms: hidden
= Thelypteris versteeghii (Holttum) Christenh.; Global Fl. 1(4): 39 (2018)
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A complete, synonymic checklist of the Higher Plants of the World

Version 19.2 (April 17th, 2024)

Covering 354,696 accepted species,
8,644 hybrids, 47,758 accepted infraspecific taxa and 996,379 synonyms,
in total 1,407,477* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,290 pictures of 15,532 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life