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Chlaenius

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Description

Chlaenius is a large and diverse genus of ground beetle. It is native to the Palearctic (including Europe), the Near East, North Africa, Afrotropical region, and the Nearctic. Worldwide, roughly 1,000 species are currently recognized with the majority of known species occurring in the Oriental and Afrotropical regions. The genus is divided into many subgenera.

From Wikipedia article at https://en.wikipedia.org/wiki/Chlaenius, which is released under the Creative Commons Attribution-Share-Alike License 3.0.

Source Data

Source	ID	Link
Global Biodiversity Information Facility ID (GBIF)	4988973	https://www.gbif.org/species/4988973
PaleoBioDB ID (PBDB)	71818	https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=71818

Rank

genus

Taxonomy (GBIF,PBDB)

Life : Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : Chlaenius

Taxonomic Status (GBIF)

accepted

Classification
(PBDB,GBIF)

Rank	Name	Author
-	Eukaryota
-	Opisthokonta	Cavalier-Smith 1987
kingdom	Animalia
-	Bilateria
-	Eubilateria	Ax 1987
-	Protostomia	Grobben 1908
-	Ecdysozoa
-	Panarthropoda
phylum	Arthropoda	Latreille 1829
subphylum	Mandibulata
-	Pancrustacea
superclass	Hexapoda	Latreille 1825
-	Cercofilata	Kukalová-Peck 1987
class	Insecta	Linnaeus 1758
-	Dicondylia
-	Paranotalia	Sroka et al. 2014
subclass	Pterygota	Brauer 1885
-	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
-	Holometabola
-	Aparaglossata
-	Coleopterida	Kukalová-Peck and Lawrence 2004
order	Coleoptera	Linnaeus 1758
suborder	Adephaga	Clairville 1806
family	Carabidae	Latreille 1802
subfamily	Harpalinae
tribe	Chlaeniini
subtribe	Chlaeniina	Brullé 1834
genus	Chlaenius	Bonelli 1810

Scientific Name

Chlaenius Bonelli, 1810

Name Published In

Bonelli, Franco A. 1810. Observations entomologiques. Première partie (cicindélètes et portion des carabiques). Félix Galletti, Turin.: 1-58; Tabula synoptica.

Opinions (PBDB)

Name	Rank	Opinion	Evidence	Author
Chlaenius	genus	belongs to Carabiques	stated with evidence	Dejean, 1821
Chlaenius	genus	belongs to Chlaenii	stated with evidence	LeConte, 1861
Chlaenius	genus	belongs to Carabidae	stated with evidence	Scudder, 1890
Chlaenius	genus	belongs to Chlaeniina	stated with evidence	Bouchard et al., 2011
Chlaenius	genus	belongs to Chlaeniini	stated without evidence	Tamutis et al., 2011

Status (PBDB)

extant

Taxon Size (PBDB)

Extant Size (PBDB)

7 (58%)

First Recorded Appearance

28.4 - 23.0 Ma
Cenozoic

Environment

terrestrial (based on Insecta)

Motility

actively mobile (based on Arthropoda)

Taphonomy

chitin (based on Insecta)

Primary Reference (PBDB)

F. A. Bonelli. 1810. Observations Entomologiques. Première Partie (Cicindélètes et Portion des Carabiques)

Wikipedia

https://en.wikipedia.org/wiki/Chlaenius

External Images

Fossil Distribution

Subtaxa

Name	Status	Common Name(s)	Fossil Occurrences	Oldest	Youngest
Chlaenius (Agostenus) subgenus	listed (PBDB)		4	2.59 Ma Pleistocene	0 Ma Extant
Chlaenius (Chlaeniellus) subgenus	listed (PBDB)		1	0.13 Ma Pleistocene	0 Ma Extant
Chlaenius abacoides species	accepted (GBIF)	No associated record in PBDB
Chlaenius aberanus species	accepted (GBIF)	No associated record in PBDB
Chlaenius aberrans species	accepted (GBIF)	No associated record in PBDB
Chlaenius abjectus species	accepted (GBIF)	No associated record in PBDB
Chlaenius abstersus species	accepted (GBIF)	No associated record in PBDB
Chlaenius accedens species	accepted (GBIF)	No associated record in PBDB
Chlaenius acroxanthus species	accepted (GBIF)	No associated record in PBDB
Chlaenius acutecostatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius adagidensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius adametzi species	accepted (GBIF)	No associated record in PBDB
Chlaenius adonis species	accepted (GBIF)	No associated record in PBDB
Chlaenius aeneocephalus species	accepted (GBIF)	No associated record in PBDB
Chlaenius aeratus species	accepted (GBIF)	No associated record in PBDB
Chlaenius aestivus species	accepted (GBIF)	No associated record in PBDB
Chlaenius aethiopicus species	accepted (GBIF)	No associated record in PBDB
Chlaenius afganus species	accepted (GBIF)	No associated record in PBDB
Chlaenius africanicus species	accepted (GBIF)	No associated record in PBDB
Chlaenius agilis species	accepted (GBIF)	No associated record in PBDB
Chlaenius agiloides species	accepted (GBIF)	No associated record in PBDB
Chlaenius akanensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius alaotraensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius albissoni species	accepted (GBIF)	No associated record in PBDB
Chlaenius alesi species	accepted (GBIF)	No associated record in PBDB
Chlaenius alexanderdostali species	accepted (GBIF)	No associated record in PBDB
Chlaenius alfredpuchneri species	accepted (GBIF)	No associated record in PBDB
Chlaenius allacteus species	accepted (GBIF)	No associated record in PBDB
Chlaenius allardi species	accepted (GBIF)	No associated record in PBDB
Chlaenius allardianus species	accepted (GBIF)	No associated record in PBDB
Chlaenius alluaudi species	accepted (GBIF)	No associated record in PBDB
Chlaenius alluaudianus species	accepted (GBIF)	No associated record in PBDB
Chlaenius alternans species	accepted (GBIF)	No associated record in PBDB
Chlaenius alternatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius alutaceus species	accepted (GBIF)	No associated record in PBDB
Chlaenius amabilis species	accepted (GBIF)	No associated record in PBDB
Chlaenius amarae species	accepted (GBIF)	No associated record in PBDB
Chlaenius amauropterus species	accepted (GBIF)	No associated record in PBDB
Chlaenius amazonicus species	accepted (GBIF)	No associated record in PBDB
Chlaenius ammon species	accepted (GBIF)	No associated record in PBDB
Chlaenius amoenus species	accepted (GBIF)	No associated record in PBDB
Chlaenius ampanihyensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius amplians species	accepted (GBIF)	No associated record in PBDB
Chlaenius amplicollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius amplipennis species	accepted (GBIF)	No associated record in PBDB
Chlaenius amydrus species	accepted (GBIF)	No associated record in PBDB
Chlaenius analis species	accepted (GBIF)	No associated record in PBDB
Chlaenius analisimilis species	accepted (GBIF)	No associated record in PBDB
Chlaenius andamanensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius androyanus species	accepted (GBIF)	No associated record in PBDB
Chlaenius angolanus species	accepted (GBIF)	No associated record in PBDB
Chlaenius angustatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius anischenkoi species	accepted (GBIF)	No associated record in PBDB
Chlaenius annulipes species	accepted (GBIF)	No associated record in PBDB
Chlaenius antennatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius anthraceus species	accepted (GBIF)	No associated record in PBDB
Chlaenius anthracoderus species	accepted (GBIF)	No associated record in PBDB
Chlaenius aodai species	accepted (GBIF)	No associated record in PBDB
Chlaenius apollo species	accepted (GBIF)	No associated record in PBDB
Chlaenius arakanensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius argentinicus species	accepted (GBIF)	No associated record in PBDB
Chlaenius armenus species	accepted (GBIF)	No associated record in PBDB
Chlaenius arnoldi species	accepted (GBIF)	No associated record in PBDB
Chlaenius aruwimius species	accepted (GBIF)	No associated record in PBDB
Chlaenius aspericollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius assamensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius assecla species	accepted (GBIF)	No associated record in PBDB
Chlaenius aterrimus species	accepted (GBIF)	No associated record in PBDB
Chlaenius athleta species	accepted (GBIF)	No associated record in PBDB
Chlaenius atratus species	accepted (GBIF)	No associated record in PBDB
Chlaenius atripes species	accepted (GBIF)	No associated record in PBDB
Chlaenius atropos species	accepted (GBIF)	No associated record in PBDB
Chlaenius attenuatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius augustus species	accepted (GBIF)	No associated record in PBDB
Chlaenius aurifex species	accepted (GBIF)	No associated record in PBDB
Chlaenius auripilis species	accepted (GBIF)	No associated record in PBDB
Chlaenius aurolimbatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius australis species	accepted (GBIF)	No associated record in PBDB
Chlaenius azurescens species	accepted (GBIF)	No associated record in PBDB
Chlaenius backanensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius baehri species	accepted (GBIF)	No associated record in PBDB
Chlaenius baeticus species	accepted (GBIF)	No associated record in PBDB
Chlaenius bakwuensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius baliensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius balthasari species	accepted (GBIF)	No associated record in PBDB
Chlaenius bandjermasinensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius bangkokensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius baoxingensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius barkeri species	accepted (GBIF)	No associated record in PBDB
Chlaenius basilimbatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius baxi species	accepted (GBIF)	No associated record in PBDB
Chlaenius beatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius bengalensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius benyovszkyi species	accepted (GBIF)	No associated record in PBDB
Chlaenius bergerdostali species	accepted (GBIF)	No associated record in PBDB
Chlaenius berndjaegeri species	accepted (GBIF)	No associated record in PBDB
Chlaenius betioky species	accepted (GBIF)	No associated record in PBDB
Chlaenius betrokaensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius bicolor species	accepted (GBIF)	No associated record in PBDB
Chlaenius bifenestratus species	accepted (GBIF)	No associated record in PBDB
Chlaenius bilyi species	accepted (GBIF)	No associated record in PBDB
Chlaenius binghami species	accepted (GBIF)	No associated record in PBDB
Chlaenius bioculatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius bipustulatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius birmanicus species	accepted (GBIF)	No associated record in PBDB
Chlaenius bisignatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius bivulnerus species	accepted (GBIF)	No associated record in PBDB
Chlaenius blanditus species	accepted (GBIF)	No associated record in PBDB
Chlaenius boali species	accepted (GBIF)	No associated record in PBDB
Chlaenius bocandei species	accepted (GBIF)	No associated record in PBDB
Chlaenius bodhidharma species	accepted (GBIF)	No associated record in PBDB
Chlaenius bohemani species	accepted (GBIF)	No associated record in PBDB
Chlaenius boisduvalii species	accepted (GBIF)	No associated record in PBDB
Chlaenius bonelli species	accepted (GBIF)	No associated record in PBDB
Chlaenius borellyi species	accepted (GBIF)	No associated record in PBDB
Chlaenius borgouensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius bottegoi species	accepted (GBIF)	No associated record in PBDB
Chlaenius boueti species	accepted (GBIF)	No associated record in PBDB
Chlaenius boukali species	accepted (GBIF)	No associated record in PBDB
Chlaenius braminus species	accepted (GBIF)	No associated record in PBDB
Chlaenius brasiliensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius breuningi species	accepted (GBIF)	No associated record in PBDB
Chlaenius brevilabris species	accepted (GBIF)	No associated record in PBDB
Chlaenius brevior species	accepted (GBIF)	No associated record in PBDB
Chlaenius breviusculus species	accepted (GBIF)	No associated record in PBDB
Chlaenius brunnescens species	accepted (GBIF)	No associated record in PBDB
Chlaenius bulirschi species	accepted (GBIF)	No associated record in PBDB
Chlaenius buquetii species	accepted (GBIF)	No associated record in PBDB
Chlaenius caecus species	accepted (GBIF)	No associated record in PBDB
Chlaenius caeruleiceps species	accepted (GBIF)	No associated record in PBDB
Chlaenius caeruleicollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius caesitius species	accepted (GBIF)	No associated record in PBDB
Chlaenius caffer species	accepted (GBIF)	No associated record in PBDB
Chlaenius cambodiensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius camerunensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius canariensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius carbonatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius caurinus species	accepted (GBIF)	No associated record in PBDB
Chlaenius cavilabrum species	accepted (GBIF)	No associated record in PBDB
Chlaenius cebuensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius cecrops species	accepted (GBIF)	No associated record in PBDB
Chlaenius celer species	accepted (GBIF)	No associated record in PBDB
Chlaenius ceramensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius chagga species	accepted (GBIF)	No associated record in PBDB
Chlaenius chainatensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius chalcoderus species	accepted (GBIF)	No associated record in PBDB
Chlaenius chalcothorax species	accepted (GBIF)	No associated record in PBDB
Chlaenius cham species	accepted (GBIF)	No associated record in PBDB
Chlaenius championi species	accepted (GBIF)	No associated record in PBDB
Chlaenius changwatensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius chapanus species	accepted (GBIF)	No associated record in PBDB
Chlaenius chaudoiri species	accepted (GBIF)	No associated record in PBDB
Chlaenius chengduensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius cheni species	accepted (GBIF)	No associated record in PBDB
Chlaenius cherensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius chiangmaiensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius chlorochrous species	accepted (GBIF)	No associated record in PBDB
Chlaenius chlorodius species	accepted (GBIF)	No associated record in PBDB
Chlaenius chomthongensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius chrysocephalus species	accepted (GBIF)	No associated record in PBDB
Chlaenius chrysoderus species	accepted (GBIF)	No associated record in PBDB
Chlaenius chuanqianensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius chuji species	accepted (GBIF)	No associated record in PBDB
Chlaenius cinctus species	accepted (GBIF)	No associated record in PBDB
Chlaenius circumcinctus species	accepted (GBIF)	No associated record in PBDB
Chlaenius circumdatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius circumductus species	accepted (GBIF)	No associated record in PBDB
Chlaenius circumscriptus species	accepted (GBIF)	No associated record in PBDB
Chlaenius clarksoni species	accepted (GBIF)	No associated record in PBDB
Chlaenius claviger species	accepted (GBIF)	No associated record in PBDB
Chlaenius clypeopatens species	accepted (GBIF)	No associated record in PBDB
Chlaenius coeruleipennis species	accepted (GBIF)	No associated record in PBDB
Chlaenius coeruleus species	accepted (GBIF)	No associated record in PBDB
Chlaenius colasi species	accepted (GBIF)	No associated record in PBDB
Chlaenius collarti species	accepted (GBIF)	No associated record in PBDB
Chlaenius colmanti species	accepted (GBIF)	No associated record in PBDB
Chlaenius columbinus species	accepted (GBIF)	No associated record in PBDB
Chlaenius comans species	accepted (GBIF)	No associated record in PBDB
Chlaenius comes species	accepted (GBIF)	No associated record in PBDB
Chlaenius communimacula species	accepted (GBIF)	No associated record in PBDB
Chlaenius conformis species	accepted (GBIF)	No associated record in PBDB
Chlaenius congoanus species	accepted (GBIF)	No associated record in PBDB
Chlaenius congoensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius conradsi species	accepted (GBIF)	No associated record in PBDB
Chlaenius consobrinus species	accepted (GBIF)	No associated record in PBDB
Chlaenius consors species	accepted (GBIF)	No associated record in PBDB
Chlaenius contractus species	accepted (GBIF)	No associated record in PBDB
Chlaenius controversus species	accepted (GBIF)	No associated record in PBDB
Chlaenius convexus species	accepted (GBIF)	No associated record in PBDB
Chlaenius cookei species	accepted (GBIF)	No associated record in PBDB
Chlaenius coquerelii species	accepted (GBIF)	No associated record in PBDB
Chlaenius corbetti species	accepted (GBIF)	No associated record in PBDB
Chlaenius cordicollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius cordifer species	accepted (GBIF)	No associated record in PBDB
Chlaenius corrosulus species	accepted (GBIF)	No associated record in PBDB
Chlaenius coscinioderus species	accepted (GBIF)	No associated record in PBDB
Chlaenius cosciniophorus species	accepted (GBIF)	No associated record in PBDB
Chlaenius costiger species	accepted (GBIF)	No associated record in PBDB
Chlaenius costipennis species	accepted (GBIF)	No associated record in PBDB
Chlaenius costulatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius coxalis species	accepted (GBIF)	No associated record in PBDB
Chlaenius crebrepunctatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius crenistriatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius cribellatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius cribellicollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius cribricollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius croesus species	accepted (GBIF)	No associated record in PBDB
Chlaenius croyi species	accepted (GBIF)	No associated record in PBDB
Chlaenius cruciatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius crudelis species	accepted (GBIF)	No associated record in PBDB
Chlaenius cruralis species	accepted (GBIF)	No associated record in PBDB
Chlaenius cubanus species	accepted (GBIF)	No associated record in PBDB
Chlaenius cumatilis species	accepted (GBIF)	No associated record in PBDB
Chlaenius cupreocinctus species	accepted (GBIF)	No associated record in PBDB
Chlaenius cupreolineatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius cupreolus species	accepted (GBIF)	No associated record in PBDB
Chlaenius cupreopurpureus species	accepted (GBIF)	No associated record in PBDB
Chlaenius cupripennis species	accepted (GBIF)	No associated record in PBDB
Chlaenius cuprithorax species	accepted (GBIF)	No associated record in PBDB
Chlaenius cursor species	accepted (GBIF)	No associated record in PBDB
Chlaenius curtii species	accepted (GBIF)	No associated record in PBDB
Chlaenius curtulus species	accepted (GBIF)	No associated record in PBDB
Chlaenius cyaneus species	accepted (GBIF)	No associated record in PBDB
Chlaenius cyanipennis species	accepted (GBIF)	No associated record in PBDB
Chlaenius cyanostolus species	accepted (GBIF)	No associated record in PBDB
Chlaenius cylindricollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius daer species	accepted (GBIF)	No associated record in PBDB
Chlaenius dajuensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius dalibaiensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius daressalaami species	accepted (GBIF)	No associated record in PBDB
Chlaenius darfurensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius darlingensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius darlingtoni species	accepted (GBIF)	No associated record in PBDB
Chlaenius davidsoni species	accepted (GBIF)	No associated record in PBDB
Chlaenius decellei species	accepted (GBIF)	No associated record in PBDB
Chlaenius decipiens species	accepted (GBIF)	No associated record in PBDB
Chlaenius decorsei species	accepted (GBIF)	No associated record in PBDB
Chlaenius dejeanii species	accepted (GBIF)	No associated record in PBDB
Chlaenius delicatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius deliciolus species	accepted (GBIF)	No associated record in PBDB
Chlaenius densaticollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius denticulatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius deplanatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius deserticola species	accepted (GBIF)	No associated record in PBDB
Chlaenius deuvei species	accepted (GBIF)	No associated record in PBDB
Chlaenius deyrollei species	accepted (GBIF)	No associated record in PBDB
Chlaenius dhawalagiriensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius diabolicus species	accepted (GBIF)	No associated record in PBDB
Chlaenius dianus species	accepted (GBIF)	No associated record in PBDB
Chlaenius dichrous species	accepted (GBIF)	No associated record in PBDB
Chlaenius diephelus species	accepted (GBIF)	No associated record in PBDB
Chlaenius differens species	accepted (GBIF)	No associated record in PBDB
Chlaenius difficilis species	accepted (GBIF)	No associated record in PBDB
Chlaenius dilatatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius dilutipes species	accepted (GBIF)	No associated record in PBDB
Chlaenius dimidiatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius dinodoides species	accepted (GBIF)	No associated record in PBDB
Chlaenius discicollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius discopictus species	accepted (GBIF)	No associated record in PBDB
Chlaenius dispreticus species	accepted (GBIF)	No associated record in PBDB
Chlaenius dissidens species	accepted (GBIF)	No associated record in PBDB
Chlaenius dissimilis species	accepted (GBIF)	No associated record in PBDB
Chlaenius distigma species	accepted (GBIF)	No associated record in PBDB
Chlaenius ditulus species	accepted (GBIF)	No associated record in PBDB
Chlaenius diversus species	accepted (GBIF)	No associated record in PBDB
Chlaenius dives species	accepted (GBIF)	No associated record in PBDB
Chlaenius djaina species	accepted (GBIF)	No associated record in PBDB
Chlaenius dohrnii species	accepted (GBIF)	No associated record in PBDB
Chlaenius dondoensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius doriae species	accepted (GBIF)	No associated record in PBDB
Chlaenius dorsalis species	accepted (GBIF)	No associated record in PBDB
Chlaenius dostojevskii species	accepted (GBIF)	No associated record in PBDB
Chlaenius douei species	accepted (GBIF)	No associated record in PBDB
Chlaenius drescheri species	accepted (GBIF)	No associated record in PBDB
Chlaenius dureli species	accepted (GBIF)	No associated record in PBDB
Chlaenius dusaultii species	accepted (GBIF)	No associated record in PBDB
Chlaenius duvaucelii species	accepted (GBIF)	No associated record in PBDB
Chlaenius ecuadoricus species	accepted (GBIF)	No associated record in PBDB
Chlaenius effugiens species	accepted (GBIF)	No associated record in PBDB
Chlaenius eggeri species	accepted (GBIF)	No associated record in PBDB
Chlaenius electrinus ✝ species	accepted (GBIF) listed (PBDB)		1	0.01 Ma Pleistocene	0.01 Ma Pleistocene
Chlaenius elegans species	accepted (GBIF)	No associated record in PBDB
Chlaenius elgonensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius elisabethanus species	accepted (GBIF)	No associated record in PBDB
Chlaenius elongatulus species	accepted (GBIF)	No associated record in PBDB
Chlaenius elongatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius emarginatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius eneides species	accepted (GBIF)	No associated record in PBDB
Chlaenius epicosmoides species	accepted (GBIF)	No associated record in PBDB
Chlaenius episcopalis species	accepted (GBIF)	No associated record in PBDB
Chlaenius epistrophus species	accepted (GBIF)	No associated record in PBDB
Chlaenius eritreaensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius ernesti species	accepted (GBIF)	No associated record in PBDB
Chlaenius eruditus species	accepted (GBIF)	No associated record in PBDB
Chlaenius erythrocnemis species	accepted (GBIF)	No associated record in PBDB
Chlaenius erythropus species	accepted (GBIF)	No associated record in PBDB
Chlaenius eryx species	accepted (GBIF)	No associated record in PBDB
Chlaenius etoshaensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius eugrammus species	accepted (GBIF)	No associated record in PBDB
Chlaenius eurybates species	accepted (GBIF)	No associated record in PBDB
Chlaenius euryscopus species	accepted (GBIF)	No associated record in PBDB
Chlaenius evae species	accepted (GBIF)	No associated record in PBDB
Chlaenius exaratus species	accepted (GBIF)	No associated record in PBDB
Chlaenius exilis species	accepted (GBIF)	No associated record in PBDB
Chlaenius extensus species	accepted (GBIF)	No associated record in PBDB
Chlaenius eyeni species	accepted (GBIF)	No associated record in PBDB
Chlaenius fairmairei species	accepted (GBIF)	No associated record in PBDB
Chlaenius fallax species	accepted (GBIF)	No associated record in PBDB
Chlaenius farai species	accepted (GBIF)	No associated record in PBDB
Chlaenius fasciger species	accepted (GBIF)	No associated record in PBDB
Chlaenius fastigatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius feanus species	accepted (GBIF)	No associated record in PBDB
Chlaenius femoratus species	accepted (GBIF)	No associated record in PBDB
Chlaenius fenestratus species	accepted (GBIF)	No associated record in PBDB
Chlaenius feronoides species	accepted (GBIF)	No associated record in PBDB
Chlaenius festivus species	accepted (GBIF)	No associated record in PBDB
Chlaenius fimbriatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius fizianus species	accepted (GBIF)	No associated record in PBDB
Chlaenius flaccidus species	accepted (GBIF)	No associated record in PBDB
Chlaenius flavicornis species	accepted (GBIF)	No associated record in PBDB
Chlaenius flaviguttatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius flavipes species	accepted (GBIF)	No associated record in PBDB
Chlaenius flavofemoratus species	accepted (GBIF)	No associated record in PBDB
Chlaenius flavomaculatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius flavoscriptus species	accepted (GBIF)	No associated record in PBDB
Chlaenius fletcheri species	accepted (GBIF)	No associated record in PBDB
Chlaenius floridanus species	accepted (GBIF)	No associated record in PBDB
Chlaenius formosensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius forreri species	accepted (GBIF)	No associated record in PBDB
Chlaenius fouquei species	accepted (GBIF)	No associated record in PBDB
Chlaenius frater species	accepted (GBIF)	No associated record in PBDB
Chlaenius fraterculus species	accepted (GBIF)	No associated record in PBDB
Chlaenius fraternus species	accepted (GBIF)	No associated record in PBDB
Chlaenius freyellus species	accepted (GBIF)	No associated record in PBDB
Chlaenius freyi species	accepted (GBIF)	No associated record in PBDB
Chlaenius freynei species	accepted (GBIF)	No associated record in PBDB
Chlaenius fugax species	accepted (GBIF)	No associated record in PBDB
Chlaenius fulgidicollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius fulvicollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius fulvipes species	accepted (GBIF)	No associated record in PBDB
Chlaenius furvus ✝ species	listed (PBDB)		5	20.4 Ma Miocene	20.4 Ma Miocene
Chlaenius fuscicornis species	accepted (GBIF)	No associated record in PBDB
Chlaenius gabonensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius gabonicus species	accepted (GBIF)	No associated record in PBDB
Chlaenius gambelaensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius gansuensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius gaoligongensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius gardoensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius garnerae species	accepted (GBIF)	No associated record in PBDB
Chlaenius geayi species	accepted (GBIF)	No associated record in PBDB
Chlaenius gebleri species	listed (PBDB)		1	1.81 Ma Pleistocene	0 Ma Extant
Chlaenius geisthardti species	accepted (GBIF)	No associated record in PBDB
Chlaenius gemmingeri species	accepted (GBIF)	No associated record in PBDB
Chlaenius geniculatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius germanus species	accepted (GBIF)	No associated record in PBDB
Chlaenius gestroi species	accepted (GBIF)	No associated record in PBDB
Chlaenius ghesquierei species	accepted (GBIF)	No associated record in PBDB
Chlaenius gikondo species	accepted (GBIF)	No associated record in PBDB
Chlaenius glabratus species	accepted (GBIF)	No associated record in PBDB
Chlaenius glabricollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius glaucinus species	accepted (GBIF)	No associated record in PBDB
Chlaenius glaucus species	accepted (GBIF)	No associated record in PBDB
Chlaenius gonioderus species	accepted (GBIF)	No associated record in PBDB
Chlaenius gonospilus species	accepted (GBIF)	No associated record in PBDB
Chlaenius goossensi species	accepted (GBIF)	No associated record in PBDB
Chlaenius goryi species	accepted (GBIF)	No associated record in PBDB
Chlaenius gotschii species	accepted (GBIF)	No associated record in PBDB
Chlaenius greensladei species	accepted (GBIF)	No associated record in PBDB
Chlaenius greyanus species	accepted (GBIF)	No associated record in PBDB
Chlaenius grootfonteinensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius grundmanni species	accepted (GBIF)	No associated record in PBDB
Chlaenius guatemalenus species	accepted (GBIF)	No associated record in PBDB
Chlaenius guineensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius gundlachi species	accepted (GBIF)	No associated record in PBDB
Chlaenius gunganagarensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius guttula species	accepted (GBIF)	No associated record in PBDB
Chlaenius haeckeli species	accepted (GBIF)	No associated record in PBDB
Chlaenius hainanensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius hamatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius hamifer species	accepted (GBIF)	No associated record in PBDB
Chlaenius harpalinus species	accepted (GBIF)	No associated record in PBDB
Chlaenius heidenfelderi species	accepted (GBIF)	No associated record in PBDB
Chlaenius helvetorum species	accepted (GBIF)	No associated record in PBDB
Chlaenius hemichlorus species	accepted (GBIF)	No associated record in PBDB
Chlaenius henryi species	accepted (GBIF)	No associated record in PBDB
Chlaenius herbaceus species	accepted (GBIF)	No associated record in PBDB
Chlaenius heyrovskyi species	accepted (GBIF)	No associated record in PBDB
Chlaenius hildebrandti species	accepted (GBIF)	No associated record in PBDB
Chlaenius himalayicus species	accepted (GBIF)	No associated record in PBDB
Chlaenius holomelas species	accepted (GBIF)	No associated record in PBDB
Chlaenius hornburgi species	accepted (GBIF)	No associated record in PBDB
Chlaenius horni species	accepted (GBIF)	No associated record in PBDB
Chlaenius hostilis species	accepted (GBIF)	No associated record in PBDB
Chlaenius howa species	accepted (GBIF)	No associated record in PBDB
Chlaenius huedepohli species	accepted (GBIF)	No associated record in PBDB
Chlaenius humeralis species	accepted (GBIF)	No associated record in PBDB
Chlaenius humphreyi species	accepted (GBIF)	No associated record in PBDB
Chlaenius hunanensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius hutiaoxiaensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius hypocrita species	accepted (GBIF)	No associated record in PBDB
Chlaenius imitatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius immaculatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius immunitus species	accepted (GBIF)	No associated record in PBDB
Chlaenius imperialis species	accepted (GBIF)	No associated record in PBDB
Chlaenius impictus species	accepted (GBIF)	No associated record in PBDB
Chlaenius impressicollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius impunctifrons species	accepted (GBIF)	No associated record in PBDB
Chlaenius inaequalis species	accepted (GBIF)	No associated record in PBDB
Chlaenius incognitus species	accepted (GBIF)	No associated record in PBDB
Chlaenius inderiensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius indicus species	accepted (GBIF)	No associated record in PBDB
Chlaenius indutus species	accepted (GBIF)	No associated record in PBDB
Chlaenius infantulus species	accepted (GBIF)	No associated record in PBDB
Chlaenius infersus species	accepted (GBIF)	No associated record in PBDB
Chlaenius inops species	accepted (GBIF)	No associated record in PBDB
Chlaenius insignis species	accepted (GBIF)	No associated record in PBDB
Chlaenius instabilis species	accepted (GBIF)	No associated record in PBDB
Chlaenius insulanus species	accepted (GBIF)	No associated record in PBDB
Chlaenius insularis species	accepted (GBIF)	No associated record in PBDB
Chlaenius intermedius species	accepted (GBIF)	No associated record in PBDB
Chlaenius interruptus species	accepted (GBIF)	No associated record in PBDB
Chlaenius iranensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius isaloensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius itombwanus species	accepted (GBIF)	No associated record in PBDB
Chlaenius ivorensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius jacobsoni species	accepted (GBIF)	No associated record in PBDB
Chlaenius jactus species	accepted (GBIF)	No associated record in PBDB
Chlaenius jaechi species	accepted (GBIF)	No associated record in PBDB
Chlaenius jaegeri species	accepted (GBIF)	No associated record in PBDB
Chlaenius jakeschi species	accepted (GBIF)	No associated record in PBDB
Chlaenius jaleswarensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius jamaicae species	accepted (GBIF)	No associated record in PBDB
Chlaenius janaki species	accepted (GBIF)	No associated record in PBDB
Chlaenius janthinus species	accepted (GBIF)	No associated record in PBDB
Chlaenius janus species	accepted (GBIF)	No associated record in PBDB
Chlaenius jeanneli species	accepted (GBIF)	No associated record in PBDB
Chlaenius jordani species	accepted (GBIF)	No associated record in PBDB
Chlaenius jos species	accepted (GBIF)	No associated record in PBDB
Chlaenius jucundulus species	accepted (GBIF)	No associated record in PBDB
Chlaenius judianensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius junceus species	accepted (GBIF)	No associated record in PBDB
Chlaenius jureceki species	accepted (GBIF)	No associated record in PBDB
Chlaenius juvencus species	accepted (GBIF)	No associated record in PBDB
Chlaenius kabompo species	accepted (GBIF)	No associated record in PBDB
Chlaenius kafakumbae species	accepted (GBIF)	No associated record in PBDB
Chlaenius kanarae species	accepted (GBIF)	No associated record in PBDB
Chlaenius kapangae species	accepted (GBIF)	No associated record in PBDB
Chlaenius kashituensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius kashmiricus species	accepted (GBIF)	No associated record in PBDB
Chlaenius kaszabi species	accepted (GBIF)	No associated record in PBDB
Chlaenius katanga species	accepted (GBIF)	No associated record in PBDB
Chlaenius katanganus species	accepted (GBIF)	No associated record in PBDB
Chlaenius kathmanduensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius kathrynae species	accepted (GBIF)	No associated record in PBDB
Chlaenius keniaensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius kenyerii species	accepted (GBIF)	No associated record in PBDB
Chlaenius kerkvoordeae species	accepted (GBIF)	No associated record in PBDB
Chlaenius kilimanus species	accepted (GBIF)	No associated record in PBDB
Chlaenius kindermanni species	accepted (GBIF)	No associated record in PBDB
Chlaenius kira species	accepted (GBIF)	No associated record in PBDB
Chlaenius kirki species	accepted (GBIF)	No associated record in PBDB
Chlaenius kirschenhoferi species	accepted (GBIF)	No associated record in PBDB
Chlaenius kivuanus species	accepted (GBIF)	No associated record in PBDB
Chlaenius kivuensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius klapperichi species	accepted (GBIF)	No associated record in PBDB
Chlaenius koenigi species	accepted (GBIF)	No associated record in PBDB
Chlaenius kolariensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius kolbei species	accepted (GBIF)	No associated record in PBDB
Chlaenius koltzei species	accepted (GBIF)	No associated record in PBDB
Chlaenius kotys species	accepted (GBIF)	No associated record in PBDB
Chlaenius kraatzi species	accepted (GBIF)	No associated record in PBDB
Chlaenius kryzhanovskyi species	accepted (GBIF)	No associated record in PBDB
Chlaenius kulti species	accepted (GBIF)	No associated record in PBDB
Chlaenius kurosawai species	accepted (GBIF)	No associated record in PBDB
Chlaenius kwazuluensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius lacunosus species	accepted (GBIF)	No associated record in PBDB
Chlaenius laeticollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius laetiusculus species	accepted (GBIF)	No associated record in PBDB
Chlaenius laetoides species	accepted (GBIF)	No associated record in PBDB
Chlaenius laetus species	accepted (GBIF)	No associated record in PBDB
Chlaenius laevicollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius laeviplaga species	accepted (GBIF)	No associated record in PBDB
Chlaenius laeviusculus species	accepted (GBIF)	No associated record in PBDB
Chlaenius lafertei species	accepted (GBIF)	No associated record in PBDB
Chlaenius lamottei species	accepted (GBIF)	No associated record in PBDB
Chlaenius langmusiensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius langsonensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius laosensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius laotinus species	accepted (GBIF)	No associated record in PBDB
Chlaenius lapillus species	accepted (GBIF)	No associated record in PBDB
Chlaenius lastii species	accepted (GBIF)	No associated record in PBDB
Chlaenius lateralis species	accepted (GBIF)	No associated record in PBDB
Chlaenius latesternalis species	accepted (GBIF)	No associated record in PBDB
Chlaenius laticollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius latipalpis species	accepted (GBIF)	No associated record in PBDB
Chlaenius latipennis species	accepted (GBIF)	No associated record in PBDB
Chlaenius latithorax species	accepted (GBIF)	No associated record in PBDB
Chlaenius lativittis species	accepted (GBIF)	No associated record in PBDB
Chlaenius latreillei species	accepted (GBIF)	No associated record in PBDB
Chlaenius lederi species	accepted (GBIF)	No associated record in PBDB
Chlaenius leigongshanensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius leishanensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius leleupi species	accepted (GBIF)	No associated record in PBDB
Chlaenius leopoldi species	accepted (GBIF)	No associated record in PBDB
Chlaenius leprieuri species	accepted (GBIF)	No associated record in PBDB
Chlaenius leptopus species	accepted (GBIF)	No associated record in PBDB
Chlaenius leucops species	accepted (GBIF)	No associated record in PBDB
Chlaenius leucoristus species	accepted (GBIF)	No associated record in PBDB
Chlaenius leucoscelis species	accepted (GBIF)	No associated record in PBDB
Chlaenius leytensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius li species	accepted (GBIF)	No associated record in PBDB
Chlaenius limbatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius limbicollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius limbipennis species	accepted (GBIF)	No associated record in PBDB
Chlaenius lineatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius lineellus species	accepted (GBIF)	No associated record in PBDB
Chlaenius lineicinctus species	accepted (GBIF)	No associated record in PBDB
Chlaenius linwensini species	accepted (GBIF)	No associated record in PBDB
Chlaenius liocephalus species	accepted (GBIF)	No associated record in PBDB
Chlaenius lioderus species	accepted (GBIF)	No associated record in PBDB
Chlaenius liothorax species	accepted (GBIF)	No associated record in PBDB
Chlaenius lirifer species	accepted (GBIF)	No associated record in PBDB
Chlaenius lissoderus species	accepted (GBIF)	No associated record in PBDB
Chlaenius lithophilus species	accepted (GBIF)	No associated record in PBDB
Chlaenius litongaensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius loango species	accepted (GBIF)	No associated record in PBDB
Chlaenius lobozianus species	accepted (GBIF)	No associated record in PBDB
Chlaenius loeblianus species	accepted (GBIF)	No associated record in PBDB
Chlaenius lombokensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius lomii species	accepted (GBIF)	No associated record in PBDB
Chlaenius lomsakensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius longeantennatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius longicornis species	accepted (GBIF)	No associated record in PBDB
Chlaenius longulus species	accepted (GBIF)	No associated record in PBDB
Chlaenius louwerensi species	accepted (GBIF)	No associated record in PBDB
Chlaenius loveridgei species	accepted (GBIF)	No associated record in PBDB
Chlaenius loxias species	accepted (GBIF)	No associated record in PBDB
Chlaenius lucasii species	accepted (GBIF)	No associated record in PBDB
Chlaenius lucidicollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius lucidulus species	accepted (GBIF)	No associated record in PBDB
Chlaenius luculentus species	accepted (GBIF)	No associated record in PBDB
Chlaenius lugens species	accepted (GBIF)	No associated record in PBDB
Chlaenius luisae species	accepted (GBIF)	No associated record in PBDB
Chlaenius lujai species	accepted (GBIF)	No associated record in PBDB
Chlaenius lulengae species	accepted (GBIF)	No associated record in PBDB
Chlaenius luluanus species	accepted (GBIF)	No associated record in PBDB
Chlaenius lunatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius luteicauda species	accepted (GBIF)	No associated record in PBDB
Chlaenius luteoapicalis species	accepted (GBIF)	No associated record in PBDB
Chlaenius luzonicus species	accepted (GBIF)	No associated record in PBDB
Chlaenius lyperus species	accepted (GBIF)	No associated record in PBDB
Chlaenius macropus species	accepted (GBIF)	No associated record in PBDB
Chlaenius maculatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius maculiceps species	accepted (GBIF)	No associated record in PBDB
Chlaenius maculiger species	accepted (GBIF)	No associated record in PBDB
Chlaenius madangensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius madecassus species	accepted (GBIF)	No associated record in PBDB
Chlaenius madrasensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius makalolo species	accepted (GBIF)	No associated record in PBDB
Chlaenius makokou species	accepted (GBIF)	No associated record in PBDB
Chlaenius malachinus species	accepted (GBIF)	No associated record in PBDB
Chlaenius malcheri species	accepted (GBIF)	No associated record in PBDB
Chlaenius mandli species	accepted (GBIF)	No associated record in PBDB
Chlaenius manowianus species	accepted (GBIF)	No associated record in PBDB
Chlaenius manus species	accepted (GBIF)	No associated record in PBDB
Chlaenius maowensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius maputoensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius marginellus species	accepted (GBIF)	No associated record in PBDB
Chlaenius marginicollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius marginipennis species	accepted (GBIF)	No associated record in PBDB
Chlaenius marianensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius marleyi species	accepted (GBIF)	No associated record in PBDB
Chlaenius marondera species	accepted (GBIF)	No associated record in PBDB
Chlaenius martinbaehri species	accepted (GBIF)	No associated record in PBDB
Chlaenius masoni species	accepted (GBIF)	No associated record in PBDB
Chlaenius maxi species	accepted (GBIF)	No associated record in PBDB
Chlaenius maxillosus species	accepted (GBIF)	No associated record in PBDB
Chlaenius mederici species	accepted (GBIF)	No associated record in PBDB
Chlaenius medioguttatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius mediornatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius melampus species	accepted (GBIF)	No associated record in PBDB
Chlaenius melancholicus species	accepted (GBIF)	No associated record in PBDB
Chlaenius melanopterus species	accepted (GBIF)	No associated record in PBDB
Chlaenius melanopus species	accepted (GBIF)	No associated record in PBDB
Chlaenius menevillei species	accepted (GBIF)	No associated record in PBDB
Chlaenius merkli species	accepted (GBIF)	No associated record in PBDB
Chlaenius merklianus species	accepted (GBIF)	No associated record in PBDB
Chlaenius meteorus species	accepted (GBIF)	No associated record in PBDB
Chlaenius meticulosus species	accepted (GBIF)	No associated record in PBDB
Chlaenius meyeri species	accepted (GBIF)	No associated record in PBDB
Chlaenius micans species	accepted (GBIF)	No associated record in PBDB
Chlaenius microspilus species	accepted (GBIF)	No associated record in PBDB
Chlaenius milleti species	accepted (GBIF)	No associated record in PBDB
Chlaenius milloti species	accepted (GBIF)	No associated record in PBDB
Chlaenius mkongoensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius moaboensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius modestus species	accepted (GBIF)	No associated record in PBDB
Chlaenius moestus species	accepted (GBIF)	No associated record in PBDB
Chlaenius moheliensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius moluccensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius momboensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius monachus species	accepted (GBIF)	No associated record in PBDB
Chlaenius monardi species	accepted (GBIF)	No associated record in PBDB
Chlaenius montanus species	accepted (GBIF)	No associated record in PBDB
Chlaenius montivagus species	accepted (GBIF)	No associated record in PBDB
Chlaenius moraveci species	accepted (GBIF)	No associated record in PBDB
Chlaenius morettoi species	accepted (GBIF)	No associated record in PBDB
Chlaenius morio species	accepted (GBIF)	No associated record in PBDB
Chlaenius morosus species	accepted (GBIF)	No associated record in PBDB
Chlaenius mpanga species	accepted (GBIF)	No associated record in PBDB
Chlaenius multicolor species	accepted (GBIF)	No associated record in PBDB
Chlaenius mutatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius naeviger species	accepted (GBIF)	No associated record in PBDB
Chlaenius nanlingensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius natalensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius nebraskensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius nebrioides species	accepted (GBIF)	No associated record in PBDB
Chlaenius nemoralis species	accepted (GBIF)	No associated record in PBDB
Chlaenius neocaledonicus species	accepted (GBIF)	No associated record in PBDB
Chlaenius neochloodes species	accepted (GBIF)	No associated record in PBDB
Chlaenius nepalensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius nepos species	accepted (GBIF)	No associated record in PBDB
Chlaenius newdehliensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius niger species	accepted (GBIF)	No associated record in PBDB
Chlaenius nigerrimus species	accepted (GBIF)	No associated record in PBDB
Chlaenius nigrans species	accepted (GBIF)	No associated record in PBDB
Chlaenius nigratus species	accepted (GBIF)	No associated record in PBDB
Chlaenius nigricans species	accepted (GBIF)	No associated record in PBDB
Chlaenius nigricornis species	accepted (GBIF)	No associated record in PBDB
Chlaenius nigripennis species	accepted (GBIF)	No associated record in PBDB
Chlaenius nigrita species	accepted (GBIF)	No associated record in PBDB
Chlaenius nigroscelis species	accepted (GBIF)	No associated record in PBDB
Chlaenius nigrosuturatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius nilgiricus species	accepted (GBIF)	No associated record in PBDB
Chlaenius nimbanus species	accepted (GBIF)	No associated record in PBDB
Chlaenius nitidiceps species	accepted (GBIF)	No associated record in PBDB
Chlaenius nitidicollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius nitidifrons species	accepted (GBIF)	No associated record in PBDB
Chlaenius nitidulus species	accepted (GBIF)	No associated record in PBDB
Chlaenius noeli species	accepted (GBIF)	No associated record in PBDB
Chlaenius noguchii species	accepted (GBIF)	No associated record in PBDB
Chlaenius nossibianus species	accepted (GBIF)	No associated record in PBDB
Chlaenius notabilis species	accepted (GBIF)	No associated record in PBDB
Chlaenius notula species	accepted (GBIF)	No associated record in PBDB
Chlaenius nubicus species	accepted (GBIF)	No associated record in PBDB
Chlaenius nyassaensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius nyika species	accepted (GBIF)	No associated record in PBDB
Chlaenius obenbergeri species	accepted (GBIF)	No associated record in PBDB
Chlaenius oberthueri species	accepted (GBIF)	No associated record in PBDB
Chlaenius obesus species	accepted (GBIF)	No associated record in PBDB
Chlaenius obliquatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius oblongus species	accepted (GBIF)	No associated record in PBDB
Chlaenius obscuripennis species	accepted (GBIF)	No associated record in PBDB
Chlaenius obscurus species	accepted (GBIF)	No associated record in PBDB
Chlaenius obsidianoides species	accepted (GBIF)	No associated record in PBDB
Chlaenius obsidianus species	accepted (GBIF)	No associated record in PBDB
Chlaenius obsoletus species	accepted (GBIF)	No associated record in PBDB
Chlaenius obtusus species	accepted (GBIF)	No associated record in PBDB
Chlaenius occultus species	accepted (GBIF)	No associated record in PBDB
Chlaenius ochroperas species	accepted (GBIF)	No associated record in PBDB
Chlaenius ocreatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius oculatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius oligochrysus species	accepted (GBIF)	No associated record in PBDB
Chlaenius olivieri species	accepted (GBIF)	No associated record in PBDB
Chlaenius olthofi species	accepted (GBIF)	No associated record in PBDB
Chlaenius omochlorus species	accepted (GBIF)	No associated record in PBDB
Chlaenius oneili species	accepted (GBIF)	No associated record in PBDB
Chlaenius oodioides species	accepted (GBIF)	No associated record in PBDB
Chlaenius opacipennis species	accepted (GBIF)	No associated record in PBDB
Chlaenius ophonoides species	accepted (GBIF)	No associated record in PBDB
Chlaenius opisthographus species	accepted (GBIF)	No associated record in PBDB
Chlaenius orbicollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius orbiculicollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius orbus species	accepted (GBIF)	No associated record in PBDB
Chlaenius orphanus species	accepted (GBIF)	No associated record in PBDB
Chlaenius ostrinus species	accepted (GBIF)	No associated record in PBDB
Chlaenius ovalipennis species	accepted (GBIF)	No associated record in PBDB
Chlaenius ovampo species	accepted (GBIF)	No associated record in PBDB
Chlaenius overlaeti species	accepted (GBIF)	No associated record in PBDB
Chlaenius oxygonus species	accepted (GBIF)	No associated record in PBDB
Chlaenius pacholatkoi species	accepted (GBIF)	No associated record in PBDB
Chlaenius pachys species	accepted (GBIF)	No associated record in PBDB
Chlaenius paenulatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius pailin species	accepted (GBIF)	No associated record in PBDB
Chlaenius pailinensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius pallidecornis species	accepted (GBIF)	No associated record in PBDB
Chlaenius pallipes species	accepted (GBIF)	No associated record in PBDB
Chlaenius palpalis species	accepted (GBIF)	No associated record in PBDB
Chlaenius paluensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius pan species	accepted (GBIF)	No associated record in PBDB
Chlaenius panagaeoides species	accepted (GBIF)	No associated record in PBDB
Chlaenius pandagensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius pantarensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius parallelus species	accepted (GBIF)	No associated record in PBDB
Chlaenius parepus species	accepted (GBIF)	No associated record in PBDB
Chlaenius paromius species	accepted (GBIF)	No associated record in PBDB
Chlaenius patricius species	accepted (GBIF)	No associated record in PBDB
Chlaenius patrizii species	accepted (GBIF)	No associated record in PBDB
Chlaenius patruelis species	accepted (GBIF)	No associated record in PBDB
Chlaenius paulyi species	accepted (GBIF)	No associated record in PBDB
Chlaenius pauxillus species	accepted (GBIF)	No associated record in PBDB
Chlaenius paykulli species	accepted (GBIF)	No associated record in PBDB
Chlaenius pecirkai species	accepted (GBIF)	No associated record in PBDB
Chlaenius pectinipes species	accepted (GBIF)	No associated record in PBDB
Chlaenius peltastes species	accepted (GBIF)	No associated record in PBDB
Chlaenius pendjariensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius pennsylvanicus species	accepted (GBIF)	No associated record in PBDB
Chlaenius pericallus species	accepted (GBIF)	No associated record in PBDB
Chlaenius perinetanus species	accepted (GBIF)	No associated record in PBDB
Chlaenius peringueyi species	accepted (GBIF)	No associated record in PBDB
Chlaenius perpunctatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius perrieri species	accepted (GBIF)	No associated record in PBDB
Chlaenius persimilis species	accepted (GBIF)	No associated record in PBDB
Chlaenius perspicillaris species	accepted (GBIF)	No associated record in PBDB
Chlaenius pertinax species	accepted (GBIF)	No associated record in PBDB
Chlaenius peruanus species	accepted (GBIF)	No associated record in PBDB
Chlaenius peterseni species	accepted (GBIF)	No associated record in PBDB
Chlaenius phaenoderus species	accepted (GBIF)	No associated record in PBDB
Chlaenius phenax species	accepted (GBIF)	No associated record in PBDB
Chlaenius philemon species	accepted (GBIF)	No associated record in PBDB
Chlaenius piceus species	accepted (GBIF)	No associated record in PBDB
Chlaenius pictus species	accepted (GBIF)	No associated record in PBDB
Chlaenius piligenys species	accepted (GBIF)	No associated record in PBDB
Chlaenius pilosicoerulans species	accepted (GBIF)	No associated record in PBDB
Chlaenius pimalicus species	accepted (GBIF)	No associated record in PBDB
Chlaenius planipennis species	accepted (GBIF)	No associated record in PBDB
Chlaenius planulatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius platensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius platyderus species	accepted (GBIF)	No associated record in PBDB
Chlaenius platynoides species	accepted (GBIF)	No associated record in PBDB
Chlaenius plausibilis species	accepted (GBIF)	No associated record in PBDB
Chlaenius pleuroderus species	accepted (GBIF)	No associated record in PBDB
Chlaenius plicatipennis ✝ species	accepted (GBIF) listed (PBDB)		1	0.13 Ma Pleistocene	0.13 Ma Pleistocene
Chlaenius poecilinus species	accepted (GBIF)	No associated record in PBDB
Chlaenius pongraczi species	accepted (GBIF)	No associated record in PBDB
Chlaenius porinus species	accepted (GBIF)	No associated record in PBDB
Chlaenius porphyrius species	accepted (GBIF)	No associated record in PBDB
Chlaenius porrectus species	accepted (GBIF)	No associated record in PBDB
Chlaenius posticalis species	accepted (GBIF)	No associated record in PBDB
Chlaenius posticemaculatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius posticus species	accepted (GBIF)	No associated record in PBDB
Chlaenius postmaculatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius postscriptus species	accepted (GBIF)	No associated record in PBDB
Chlaenius prabangensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius pradieri species	accepted (GBIF)	No associated record in PBDB
Chlaenius praefectus species	accepted (GBIF)	No associated record in PBDB
Chlaenius prasinus species	accepted (GBIF)	No associated record in PBDB
Chlaenius pratensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius pretiosus species	accepted (GBIF)	No associated record in PBDB
Chlaenius primaevus ✝ species	accepted (GBIF) listed (PBDB)		1	28.4 Ma Oligocene	28.4 Ma Oligocene
Chlaenius principalis species	accepted (GBIF)	No associated record in PBDB
Chlaenius privatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius probsti species	accepted (GBIF)	No associated record in PBDB
Chlaenius procerulus species	accepted (GBIF)	No associated record in PBDB
Chlaenius prolixus species	accepted (GBIF)	No associated record in PBDB
Chlaenius propeagilis species	accepted (GBIF)	No associated record in PBDB
Chlaenius propinquus species	accepted (GBIF)	No associated record in PBDB
Chlaenius prostenus species	accepted (GBIF)	No associated record in PBDB
Chlaenius protensus species	accepted (GBIF)	No associated record in PBDB
Chlaenius pseudobipustulatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius pseudocruciatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius pseudocupreolus species	accepted (GBIF)	No associated record in PBDB
Chlaenius pseudoglabratus species	accepted (GBIF)	No associated record in PBDB
Chlaenius pseudolulengae species	accepted (GBIF)	No associated record in PBDB
Chlaenius pseudopalpalis species	accepted (GBIF)	No associated record in PBDB
Chlaenius pseudopoeciloides species	accepted (GBIF)	No associated record in PBDB
Chlaenius pseudoraffrayi species	accepted (GBIF)	No associated record in PBDB
Chlaenius pseudotristis species	accepted (GBIF)	No associated record in PBDB
Chlaenius pterostichoides species	accepted (GBIF)	No associated record in PBDB
Chlaenius ptuchodes species	accepted (GBIF)	No associated record in PBDB
Chlaenius puberulus species	accepted (GBIF)	No associated record in PBDB
Chlaenius pubifer species	accepted (GBIF)	No associated record in PBDB
Chlaenius pubipennis species	accepted (GBIF)	No associated record in PBDB
Chlaenius puchneri species	accepted (GBIF)	No associated record in PBDB
Chlaenius pudicus species	accepted (GBIF)	No associated record in PBDB
Chlaenius pugni species	accepted (GBIF)	No associated record in PBDB
Chlaenius pulchellus species	accepted (GBIF)	No associated record in PBDB
Chlaenius pulcher species	accepted (GBIF)	No associated record in PBDB
Chlaenius pulchriceps species	accepted (GBIF)	No associated record in PBDB
Chlaenius pumilio species	accepted (GBIF)	No associated record in PBDB
Chlaenius punctatostriatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius puncticephalis species	accepted (GBIF)	No associated record in PBDB
Chlaenius puncticollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius punctulatus ✝ species	accepted (GBIF) listed (PBDB)		1	1.80 Ma Pleistocene	1.80 Ma Pleistocene
Chlaenius purpureus species	accepted (GBIF)	No associated record in PBDB
Chlaenius purpuricollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius pusillus species	accepted (GBIF)	No associated record in PBDB
Chlaenius putzeysi species	accepted (GBIF)	No associated record in PBDB
Chlaenius pyrrhopodus species	accepted (GBIF)	No associated record in PBDB
Chlaenius pyrrhos species	accepted (GBIF)	No associated record in PBDB
Chlaenius qinqchensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius quadricolor species	accepted (GBIF)	No associated record in PBDB
Chlaenius quadrimaculatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius quadrinotatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius quadriornatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius quadripustulatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius quadrisignatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius quadrisulcatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius quadrivittatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius radama species	accepted (GBIF)	No associated record in PBDB
Chlaenius raffrayi species	accepted (GBIF)	No associated record in PBDB
Chlaenius rafiki species	accepted (GBIF)	No associated record in PBDB
Chlaenius rajasthanensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius rambouseki species	accepted (GBIF)	No associated record in PBDB
Chlaenius rampurensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius ranavalonae species	accepted (GBIF)	No associated record in PBDB
Chlaenius rapyllii species	accepted (GBIF)	No associated record in PBDB
Chlaenius rebellis species	accepted (GBIF)	No associated record in PBDB
Chlaenius reichei species	accepted (GBIF)	No associated record in PBDB
Chlaenius reitteri species	accepted (GBIF)	No associated record in PBDB
Chlaenius retropictus species	accepted (GBIF)	No associated record in PBDB
Chlaenius rhodesianus species	accepted (GBIF)	No associated record in PBDB
Chlaenius rhysonotus species	accepted (GBIF)	No associated record in PBDB
Chlaenius richardsi species	accepted (GBIF)	No associated record in PBDB
Chlaenius riparius species	accepted (GBIF)	No associated record in PBDB
Chlaenius ripicola species	accepted (GBIF)	No associated record in PBDB
Chlaenius robertae species	accepted (GBIF)	No associated record in PBDB
Chlaenius robustus species	accepted (GBIF)	No associated record in PBDB
Chlaenius rodriguezi species	accepted (GBIF)	No associated record in PBDB
Chlaenius roeschkei species	accepted (GBIF)	No associated record in PBDB
Chlaenius rotundatulus species	accepted (GBIF)	No associated record in PBDB
Chlaenius rotundithorax species	accepted (GBIF)	No associated record in PBDB
Chlaenius rotundus species	accepted (GBIF)	No associated record in PBDB
Chlaenius rubenticollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius rubricrus species	accepted (GBIF)	No associated record in PBDB
Chlaenius rudesculptus species	accepted (GBIF)	No associated record in PBDB
Chlaenius rudicollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius ruficauda species	accepted (GBIF)	No associated record in PBDB
Chlaenius rufifemoratus species	accepted (GBIF)	No associated record in PBDB
Chlaenius rufithorax species	accepted (GBIF)	No associated record in PBDB
Chlaenius rufomarginatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius rugulosus species	accepted (GBIF)	No associated record in PBDB
Chlaenius rukwaensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius ruthmuellerae species	accepted (GBIF)	No associated record in PBDB
Chlaenius ruvumaensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius sabahensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius sagaingensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius saginatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius sagittarius species	accepted (GBIF)	No associated record in PBDB
Chlaenius sallei species	accepted (GBIF)	No associated record in PBDB
Chlaenius samoensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius sanatonialis species	accepted (GBIF)	No associated record in PBDB
Chlaenius sangaicus species	accepted (GBIF)	No associated record in PBDB
Chlaenius sankuruensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius sassanus species	accepted (GBIF)	No associated record in PBDB
Chlaenius savanicola species	accepted (GBIF)	No associated record in PBDB
Chlaenius scabricollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius scapularis species	accepted (GBIF)	No associated record in PBDB
Chlaenius schatzmayri species	accepted (GBIF)	No associated record in PBDB
Chlaenius scheerpeltzi species	accepted (GBIF)	No associated record in PBDB
Chlaenius schillhammeri species	accepted (GBIF)	No associated record in PBDB
Chlaenius schmidti species	accepted (GBIF)	No associated record in PBDB
Chlaenius schoenherri species	accepted (GBIF)	No associated record in PBDB
Chlaenius schoutedeni species	accepted (GBIF)	No associated record in PBDB
Chlaenius schuelei species	accepted (GBIF)	No associated record in PBDB
Chlaenius sciakyi species	accepted (GBIF)	No associated record in PBDB
Chlaenius scotti species	accepted (GBIF)	No associated record in PBDB
Chlaenius sculptilis species	accepted (GBIF)	No associated record in PBDB
Chlaenius scutellaris species	accepted (GBIF)	No associated record in PBDB
Chlaenius sehnali species	accepted (GBIF)	No associated record in PBDB
Chlaenius seiferti species	accepted (GBIF)	No associated record in PBDB
Chlaenius sellatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius semenowi species	accepted (GBIF)	No associated record in PBDB
Chlaenius semicupreus species	accepted (GBIF)	No associated record in PBDB
Chlaenius semicyaneus species	accepted (GBIF)	No associated record in PBDB
Chlaenius semipurpureus species	accepted (GBIF)	No associated record in PBDB
Chlaenius semiviridis species	accepted (GBIF)	No associated record in PBDB
Chlaenius semperi species	accepted (GBIF)	No associated record in PBDB
Chlaenius senegalensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius sennaariensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius sericeus species	accepted (GBIF)	No associated record in PBDB
Chlaenius sericimicans species	accepted (GBIF)	No associated record in PBDB
Chlaenius sexguttatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius sexmaculatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius seyrigi species	accepted (GBIF)	No associated record in PBDB
Chlaenius shaanxinensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius sibutensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius sibuti species	accepted (GBIF)	No associated record in PBDB
Chlaenius siccus species	accepted (GBIF)	No associated record in PBDB
Chlaenius sichuanensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius sierraleonensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius siffointei species	accepted (GBIF)	No associated record in PBDB
Chlaenius signatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius silvestrii species	accepted (GBIF)	No associated record in PBDB
Chlaenius simba species	accepted (GBIF)	No associated record in PBDB
Chlaenius simbabwensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius similatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius similis species	accepted (GBIF)	No associated record in PBDB
Chlaenius simillimus species	accepted (GBIF)	No associated record in PBDB
Chlaenius simplex species	accepted (GBIF)	No associated record in PBDB
Chlaenius simulatorius species	accepted (GBIF)	No associated record in PBDB
Chlaenius sinensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius sinuatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius sivorii species	accepted (GBIF)	No associated record in PBDB
Chlaenius skukuzaensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius sobrinus species	accepted (GBIF)	No associated record in PBDB
Chlaenius soccatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius soginoides species	accepted (GBIF)	No associated record in PBDB
Chlaenius sokotranus species	accepted (GBIF)	No associated record in PBDB
Chlaenius solitarius species	accepted (GBIF)	No associated record in PBDB
Chlaenius sollicitus species	accepted (GBIF)	No associated record in PBDB
Chlaenius somaliae species	accepted (GBIF)	No associated record in PBDB
Chlaenius somereni species	accepted (GBIF)	No associated record in PBDB
Chlaenius sonomae species	accepted (GBIF)	No associated record in PBDB
Chlaenius soricinus species	accepted (GBIF)	No associated record in PBDB
Chlaenius sparsepunctatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius spathulifer species	accepted (GBIF)	No associated record in PBDB
Chlaenius specularis species	accepted (GBIF)	No associated record in PBDB
Chlaenius splendidus species	accepted (GBIF)	No associated record in PBDB
Chlaenius spoliatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius startellus species	accepted (GBIF)	No associated record in PBDB
Chlaenius stenoristus species	accepted (GBIF)	No associated record in PBDB
Chlaenius sterbai species	accepted (GBIF)	No associated record in PBDB
Chlaenius stevenii species	accepted (GBIF)	No associated record in PBDB
Chlaenius stichai species	accepted (GBIF)	No associated record in PBDB
Chlaenius straneoi species	accepted (GBIF)	No associated record in PBDB
Chlaenius stschukini species	accepted (GBIF)	No associated record in PBDB
Chlaenius stungtrengensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius stygius species	accepted (GBIF)	No associated record in PBDB
Chlaenius suavis species	accepted (GBIF)	No associated record in PBDB
Chlaenius subcostatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius suberbiei species	accepted (GBIF)	No associated record in PBDB
Chlaenius subferrugineus species	accepted (GBIF)	No associated record in PBDB
Chlaenius submarginatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius subovatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius subsulcatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius suensoni species	accepted (GBIF)	No associated record in PBDB
Chlaenius sulcicollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius sulcipennis species	accepted (GBIF)	No associated record in PBDB
Chlaenius sumbawanus species	accepted (GBIF)	No associated record in PBDB
Chlaenius sumptuosus species	accepted (GBIF)	No associated record in PBDB
Chlaenius superbus species	accepted (GBIF)	No associated record in PBDB
Chlaenius superstes species	accepted (GBIF)	No associated record in PBDB
Chlaenius suppletor species	accepted (GBIF)	No associated record in PBDB
Chlaenius surdipennis species	accepted (GBIF)	No associated record in PBDB
Chlaenius suvorovi species	accepted (GBIF)	No associated record in PBDB
Chlaenius swahilius species	accepted (GBIF)	No associated record in PBDB
Chlaenius syangyaensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius sykesi species	accepted (GBIF)	No associated record in PBDB
Chlaenius synaptus species	accepted (GBIF)	No associated record in PBDB
Chlaenius syriacus species	accepted (GBIF)	No associated record in PBDB
Chlaenius systolocranioides species	accepted (GBIF)	No associated record in PBDB
Chlaenius tamdaoensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius tanahrataensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius tansaniensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius tartagalensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius taveuniensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius tchabalensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius teani species	accepted (GBIF)	No associated record in PBDB
Chlaenius tecospilus species	accepted (GBIF)	No associated record in PBDB
Chlaenius tenellus species	accepted (GBIF)	No associated record in PBDB
Chlaenius tenuelimbatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius tenuicollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius tenuis species	accepted (GBIF)	No associated record in PBDB
Chlaenius terminatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius testaceicrus species	accepted (GBIF)	No associated record in PBDB
Chlaenius tetracelis species	accepted (GBIF)	No associated record in PBDB
Chlaenius tetragonoderus species	accepted (GBIF)	No associated record in PBDB
Chlaenius tetraphacus species	accepted (GBIF)	No associated record in PBDB
Chlaenius tetraspilus species	accepted (GBIF)	No associated record in PBDB
Chlaenius texanus species	accepted (GBIF)	No associated record in PBDB
Chlaenius thieleni species	accepted (GBIF)	No associated record in PBDB
Chlaenius thiesensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius tibialis species	accepted (GBIF)	No associated record in PBDB
Chlaenius timorensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius tinantae species	accepted (GBIF)	No associated record in PBDB
Chlaenius tingaudi species	accepted (GBIF)	No associated record in PBDB
Chlaenius tiomanensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius titschacki species	accepted (GBIF)	No associated record in PBDB
Chlaenius togatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius toliaraensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius tomentosus species	accepted (GBIF)	No associated record in PBDB
Chlaenius topali species	accepted (GBIF)	No associated record in PBDB
Chlaenius toubaensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius touzalini species	accepted (GBIF)	No associated record in PBDB
Chlaenius trachys species	accepted (GBIF)	No associated record in PBDB
Chlaenius transfuga species	accepted (GBIF)	No associated record in PBDB
Chlaenius transvaalensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius trapezicollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius treichi species	accepted (GBIF)	No associated record in PBDB
Chlaenius trichrous species	accepted (GBIF)	No associated record in PBDB
Chlaenius tricolor species	accepted (GBIF)	No associated record in PBDB
Chlaenius trigonotomoides species	accepted (GBIF)	No associated record in PBDB
Chlaenius trinotatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius tripustulatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius tristis species	accepted (GBIF)	No associated record in PBDB
Chlaenius tsanerenaensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius tschitscherini species	accepted (GBIF)	No associated record in PBDB
Chlaenius tshibindensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius tudicus species	accepted (GBIF)	No associated record in PBDB
Chlaenius tuky species	accepted (GBIF)	No associated record in PBDB
Chlaenius udaipurensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius uluguruanus species	accepted (GBIF)	No associated record in PBDB
Chlaenius umtalianus species	accepted (GBIF)	No associated record in PBDB
Chlaenius unicolor species	accepted (GBIF)	No associated record in PBDB
Chlaenius uninotatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius unyorensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius uzungwensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius vadoni species	accepted (GBIF)	No associated record in PBDB
Chlaenius vafer species	accepted (GBIF)	No associated record in PBDB
Chlaenius validicornis species	accepted (GBIF)	No associated record in PBDB
Chlaenius validus species	accepted (GBIF)	No associated record in PBDB
Chlaenius variabilipes species	accepted (GBIF)	No associated record in PBDB
Chlaenius varians species	accepted (GBIF)	No associated record in PBDB
Chlaenius variicornis species	accepted (GBIF)	No associated record in PBDB
Chlaenius variipes species	accepted (GBIF)	No associated record in PBDB
Chlaenius variolosus species	accepted (GBIF)	No associated record in PBDB
Chlaenius velocipes species	accepted (GBIF)	No associated record in PBDB
Chlaenius velutinus species	accepted (GBIF)	No associated record in PBDB
Chlaenius venustulus species	accepted (GBIF)	No associated record in PBDB
Chlaenius vertagoides species	accepted (GBIF)	No associated record in PBDB
Chlaenius veselyi species	accepted (GBIF)	No associated record in PBDB
Chlaenius vestitus species	accepted (GBIF)	No associated record in PBDB
Chlaenius vethi species	accepted (GBIF)	No associated record in PBDB
Chlaenius victor species	accepted (GBIF)	No associated record in PBDB
Chlaenius victoriae species	accepted (GBIF)	No associated record in PBDB
Chlaenius viduus species	accepted (GBIF)	No associated record in PBDB
Chlaenius vientianensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius vietnamensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius vietnami species	accepted (GBIF)	No associated record in PBDB
Chlaenius vinhphuensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius violaceipennis species	accepted (GBIF)	No associated record in PBDB
Chlaenius violatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius virens species	accepted (GBIF)	No associated record in PBDB
Chlaenius virescens species	accepted (GBIF)	No associated record in PBDB
Chlaenius virgulatus species	accepted (GBIF)	No associated record in PBDB
Chlaenius virgulifer species	accepted (GBIF)	No associated record in PBDB
Chlaenius viridicollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius viridis species	accepted (GBIF)	No associated record in PBDB
Chlaenius vitalisi species	accepted (GBIF)	No associated record in PBDB
Chlaenius vitticollis species	accepted (GBIF)	No associated record in PBDB
Chlaenius vividus species	accepted (GBIF)	No associated record in PBDB
Chlaenius vulneratus species	accepted (GBIF)	No associated record in PBDB
Chlaenius waddellii species	accepted (GBIF)	No associated record in PBDB
Chlaenius wallacei species	accepted (GBIF)	No associated record in PBDB
Chlaenius watsaensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius wegneri species	accepted (GBIF)	No associated record in PBDB
Chlaenius wenchuanensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius wewalkai species	accepted (GBIF)	No associated record in PBDB
Chlaenius wittei species	accepted (GBIF)	No associated record in PBDB
Chlaenius wittmerianus species	accepted (GBIF)	No associated record in PBDB
Chlaenius wrasei species	accepted (GBIF)	No associated record in PBDB
Chlaenius xanthacrus species	accepted (GBIF)	No associated record in PBDB
Chlaenius xanthomerus species	accepted (GBIF)	No associated record in PBDB
Chlaenius xanthospilus species	accepted (GBIF)	No associated record in PBDB
Chlaenius yamdena species	accepted (GBIF)	No associated record in PBDB
Chlaenius yangonensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius yunnanulus species	accepted (GBIF)	No associated record in PBDB
Chlaenius zanzibaricus species	accepted (GBIF)	No associated record in PBDB
Chlaenius ziloensis species	accepted (GBIF)	No associated record in PBDB
Chlaenius zygogrammus species	accepted (GBIF)	No associated record in PBDB

Obsolete Names

Name	Source	Taxon Rank	Taxonomy
Iberodinodes Basilewsky & Grundmann, 1954	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Iberodinodes*
Agrochlaenius Basilewsky & Grundmann, 1954	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Agrochlaenius*
Chaelinus Basilewsky & Grundmann, 1954	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Chaelinus*
Chinelaus Basilewsky & Grundmann, 1954	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Chinelaus*
Lissauchenus Desmarest, 1851-22	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Lissauchenus*
Chlaenionus Kuntzen, 1913	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Chlaenionus*
Chlaenitidius Jeannel, 1949	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Chlaenitidius*
Epomis Dejean, 1826	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Epomis*
Laenichus Lucnik, 1933	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Laenichus*
Rhyzotrachelus Chaudoir, 1876	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Rhyzotrachelus*
Dibolochilus Lacordaire, 1854	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Dibolochilus*
Aleptocerus de la Ferté-Sénectère, 1851	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Aleptocerus*
Pelasmus Motschoulsky, 1850	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Pelasmus*
Goniodinoides Jeannel, 1949	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Goniodinoides*
Chlaeniostenodes Basilewsky, 1953	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Chlaeniostenodes*
Capsochlaenius Basilewsky, 1950	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Capsochlaenius*
Amomoglossus Chaudoir, 1876	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Amomoglossus*
Pseudanomoglossus Bell, 1960	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Pseudanomoglossus*
Ilaenchus Lucnik, 1933	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Ilaenchus*
Merochlaenius Grundman, 1955	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Merochlaenius*
Chlaeniopus Grundmann, 1955	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Chlaeniopus*
Chaelinus Lucnik, 1933	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Chaelinus*
Nectochlaenius Antoine, 1961	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Nectochlaenius*
Pachydinodes Kuntzen, 1919	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Pachydinodes*
Chlaenius Dejean, 1826	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Chlaenius*
Chlaeniostenus Kuntzen, 1919	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Chlaeniostenus*
Trachychlaenites Kuntzen, 1919	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Trachychlaenites*
Rhizotrachelus Lacordaire, 1854	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Rhizotrachelus*
Anomoglossus de Chaudoir, 1856	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Anomoglossus*
Agrochlaenius Lucnik, 1933	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Agrochlaenius*
Glyptoderus de la Ferté-Sénectère, 1851	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Glyptoderus*
Oochlaenius Alluaud, 1933	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Oochlaenius*
Amblygenius de la Ferté-Sénectère, 1851	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Amblygenius*
Calochlaenius Kuntzen, 1913	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Calochlaenius*
Parachlaenites Jeannel, 1949	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Parachlaenites*
Paradinodes Apfelbeck, 1904	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Paradinodes*
Ocybatoides Jeannel, 1949	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Ocybatoides*
Leptodinodes Jeannel, 1949	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Leptodinodes*
Cyaneodinodes Jeannel, 1949	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Cyaneodinodes*
Paratrachelus Basilewsky, 1950	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Paratrachelus*
Naelichus Lucnik, 1933	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Naelichus*
Chloenius Brullé, 1832	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Chloenius*
Wilmernstus Basilewsky, 1965	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Wilmernstus*
Chlaenites Motschoulsky, 1860	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Chlaenites*
Ilaenchus Basilewsky & Grundmann, 1954	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Ilaenchus*
Barymorphus de la Ferté-Sénectère, 1851	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Barymorphus*
Agostenus Motschoulsky, 1850	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Agostenus*
Platychlaenius Jeannel, 1949	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Platychlaenius*
Pachychlaenius Grundmann, 1955	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Pachychlaenius*
Diaphoropsophus de Chaudoir, 1850	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Diaphoropsophus*
Achlaenius Mandl, 1992	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Achlaenius*
Prochlaeniellus Basilewsky, 1965	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Prochlaeniellus*
Umtalius Péringuey, 1926	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Umtalius*
Syntelestes Gistl, 1857	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Syntelestes*
Baidochlaenius Basilewsky, 1950	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Baidochlaenius*
Stenodinodes Kuntzen, 1919	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Stenodinodes*
Aeacus de la Ferté-Sénectère, 1851	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Aeacus*
Ocybatus Laferté-Sénectère, 1851	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Ocybatus*
Rhysotrachelus Boheman, 1848	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Rhysotrachelus*
Compsochlaenius Alluaud, 1916	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Compsochlaenius*
Eurydactylus Ferté-Sénectère, 1851	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Eurydactylus*
Pseudochlaeniellus Jeannel, 1949	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Pseudochlaeniellus*
Aulacosomus Grundmann, 1955	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Aulacosomus*
Afrochlaenius Basilewsky, 1965	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Afrochlaenius*
Eudinodes Basilewsky, 1965	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Eudinodes*
Iberodinodes Wagner, 1932	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Iberodinodes*
Tomochilus Laferté-Sénéctère, 1851	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Tomochilus*
Brachylobus de Chaudoir, 1876	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Brachylobus*
Basilewskyellus Grundmann, 1956	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Basilewskyellus*
Pelasmomimus Grundmann, 1955	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Pelasmomimus*
Dilobochilus Ferté-Sénectère, 1851	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Dilobochilus*
Lissauchenius Macleay, 1825	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Lissauchenius*
Terraleus Fairmaire, 1899	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Terraleus*
Ocydromus Laferté-Sénectère, 1851	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Ocydromus*
Chlaeniodromus Basilewsky, 1950	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Chlaeniodromus*
Spilochlaenius Jeannel, 1949	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Spilochlaenius*
Callistometus Grundmann, 1956	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Callistometus*
Chinelaus Lucnik, 1933	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Chinelaus*
Sericochlaenius Grundmann, 1955	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Sericochlaenius*
Randallius Bousquet, 2012	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Randallius*

Synonymy List

Year	Name and Author
1810	Chlaenius Bonelli
1821	Chlaenius Dejean p. 8
1861	Chlaenius LeConte p. 28
1890	Chlaenius Scudder p. 517
1900	Chlaenius Scudder p. 33
1983	Shanwangicarabus Hong p. 4
1989	Chlaenius Zhang p. 99
2011	Chlaenius Bouchard et al.
2011	Chlaenius Tamutis et al.
2016	Chlaenius Webster

References

Bonelli F. A. , Observations Entomologiques. Première Partie (Cicindélètes et Portion des Carabiques), 1-58

Dejean P. F. M. , Catalogue de la collection de Coléoptères de M. le Baron Dejean, 1-136

Scudder S. H. The Tertiary Insects of North America, Report of the United States Geological Survey of the Territories XIII, 1-734

Scudder S. H. Adephagous and clavicorn Coleoptera from the Tertiary deposits at Florissant, Colorado with descriptions of a few other forms and a systematic list of the non-rhynchophorus Tertiary Coleoptera of North America, Monographs of the United States Geological Survey 40, 1-148

Hong Y. C. (1983) Fossil insects in the diatoms of Shanwang, Bulletin of the Tianjin Institute of Geology and Mineral Resources 8, 1-15

Zhang J. F. (1989) , Fossil Insects From Shanwang, Shandong, China, 1-459

Bouchard P., Bousquet Y., et al (2011) Family-group names in Coleoptera (Insecta), ZooKeys 88, 1-972

Tamutis V., Tamuté B., et al (2011) A catalogue of Lithuanian beetles (Insecta, Coleoptera), ZooKeys 121, 1-494

Webster R. P. (2016) Checklist of the Coleoptera of New Brunswick, Canada, ZooKeys 573, 387-512

Bell RT (1960) A revision of the genus Chlaenius Bonelli (Coleoptera: Carabidae) in North America. Miscellaneous Publications of the Entomological Society of America 1: 97-166. - via Checklist of Beetles (Coleoptera) of Canada and Alaska. Second Edition.

Gustafsson, B. (Col.) (2004) Catalogus Coleopterorum Sueciae. Naturhistoriska riksmuseet, Stockholm, Excelfil hämtad från http://www.nrm.se/en/catalogus.html.se. - via Dyntaxa. Svensk taxonomisk databas

CoL2006 - via The Interim Register of Marine and Nonmarine Genera

Hallan, J. (2000-current). Biology Catalog. Web compilation accessible at http://bug.tamu.edu/research/collection/hallan/ (accessed June 2012). - via The Interim Register of Marine and Nonmarine Genera

SN2000: Brands, S. J. (compiler) 1989-2005. Systema Naturae 2000. Amsterdam, The Netherlands (2006 version). Available online at http://sn2000.taxonomy.nl/. - via The Interim Register of Marine and Nonmarine Genera

as per family (nonmarine except where stated) - via The Interim Register of Marine and Nonmarine Genera

Bousquet, Yves, and André Larochelle, 1993: Catalogue of the Geadephaga (Coleoptera: Trachypachidae, Rhysodidae, Carabidae including Cicindelini) of America North of Mexico. Memoirs of the Entomological Society of Canada, no. 167. 397. - via Integrated Taxonomic Information System (ITIS)

SN2000, PaleoDB - via The Interim Register of Marine and Nonmarine Genera

Bousquet, Yves, 2012: Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico. ZooKeys, issue 245. 1-1722. - via Integrated Taxonomic Information System (ITIS)

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

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