ACARORUM
CATALOGUS
IX
ACARIFORMES
Acaridida
Petar Beron
ACARORUM CATALOGUS
Petar Beron
ACARORUM
CATALOGUS
IX
Acariformes
Acaridida
Schizoglyphoidea (Schizoglyphidae)
Histiostomatoidea (Histiostomatidae,
Guanolichidae)
Canestrinioidea (Canestriniidae, Chetochelacaridae, Lophonotacaridae, Heterocoptidae)
Hemisarcoptoidea (Chaetodactylidae,
Hyadesiidae, Algophagidae, Hemisarcoptidae,
Carpoglyphidae, Winterschmidtiidae)
Petar Beron
Pensoft
National Museum of Natural History, Sofia
Bulgarian Academy of Sciences
Sofia–Moscow
2021
ACARORUM CATALOGUS IX
Acariformes
Acaridida
Schizoglyphoidea (Schizoglyphidae)
Histiostomatoidea (Histiostomatidae, Guanolichidae)
Canestrinioidea (Canestriniidae, Chetochelacaridae, Lophonotacaridae,
Heterocoptidae)
Hemisarcoptoidea (Chaetodactylidae, Hyadesiidae, Algophagidae, Hemisarcoptidae,
Carpoglyphidae, Winterschmidtiidae
Petar Beron
First published 2021
ISBN 978-619-248-057-8 (paperback)
ISBN 978-619-248-058-5 (e-book)
Pensoft Series Faunistica No 224
Beron P. 2021. Acarorum Catalogus IX. Schizoglyphoidea (Schizoglyphidae). Histiostomatoidea
(Histiostomatidae, Guanolichidae). Canestrinioidea (Canestriniidae, Chetochelacaridae,
Lophonotacaridae, Heterocoptidae). Hemisarcoptoidea (Chaetodactylidae, Hyadesiidae,
Algophagidae, Hemisarcoptidae, Carpoglyphidae, Winterschmidtiidae. Pensoft & National
Museum of Natural History, Sofia, 396 pp.
© PENSOFT Publishers
© National Museum of Natural History, Sofia
Pensoft Publishers
Prof. Georgi Zlatarski 12, Sofia 1700, Bulgaria
info@pensoft.net
www.pensoft.net
Printed in Bulgaria, December 2021
Contents
Introduction ............................................................................................................................... 9
Acariformes, Acaridida...........................................................................................................10
The Mite Superfamily Schizoglyphoidea ..............................................................................10
The Mite Family Schizoglyphidae .........................................................................................10
Catalogue of Schizoglyphidae ........................................................................................10
Index of Schyzoglyphidae ...............................................................................................10
Distribution of Schizoglyphidae ....................................................................................11
Hosts of Schizoglyphidae ...............................................................................................11
Bibliography of Schizoglyphidae ...................................................................................11
The Mite Superfamily Histiostomatoidea (Anoetoidea) ....................................................11
The Mite Family Histiostomatidae (Anoetidae)..................................................................11
Composition of Histiostomatidae (Anoetidae) ...........................................................12
Catalogue of Histiostomatidae ......................................................................................14
Index of Histiostomatidae ..............................................................................................98
Distribution of Histiostomatidae ................................................................................108
Hosts of Histiostomatidae ............................................................................................115
Bibliography of Histiostomatidae ................................................................................122
The Mite Family Guanolichidae Fain..................................................................................141
Composition of Guanolichidae ...................................................................................141
Catalogue of Guanolichidae .........................................................................................142
Index of Guanolichidae ................................................................................................143
Distribution of Guanolichidae .....................................................................................143
Hosts of Guanolichidae.................................................................................................143
Bibliography of Guanolichidae ....................................................................................143
The Mite Superfamily Canestrinioidea ...............................................................................144
The Mite Family Canestriniidae ..........................................................................................144
Composition of Canestriniidae ...................................................................................145
Catalogue of the Family Canestriniidae .....................................................................147
Index of Canestriniidae ................................................................................................213
Distribution of Canestriniidae .....................................................................................219
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Acarorum Catalogus IX
Hosts of Canestriniidae ................................................................................................225
Bibliography of Canestriniidae ....................................................................................236
The Mite Family Chetochelacaridae Fain ...........................................................................243
Index of Chetochelacaridae..........................................................................................244
Distribution of Chetochelacaridae ..............................................................................244
Hosts of Chetochelacaridae..........................................................................................244
Bibliography of Chetochelacaridae .............................................................................244
The Mite Family Lophonotacaridae Fain ...........................................................................244
Index of Lophonotacaridae ..........................................................................................245
Distribution of Lophonotacaridae...............................................................................245
Hosts of Lophonotacaridae ..........................................................................................245
Bibliography of Lophonotacaridae ..............................................................................245
The Mite Family Heterocoptidae Fain ................................................................................246
Composition of Heterocoptidae ..................................................................................246
Catalogue of Heterocoptidae .......................................................................................246
Index of Heterocoptidae ...............................................................................................253
Distribution of Heterocoptidae ...................................................................................254
Hosts of Heterocoptidae ...............................................................................................255
Bibliography of Heterocoptidae...................................................................................256
The Mite Superfamily Hemisarcoptoidea...........................................................................257
The Mite Family Chaetodactylidae .....................................................................................258
Composition of Chaetodactylidae ..............................................................................258
Catalogue of Chaetodactylidae ....................................................................................258
Index of Chaetodactylidae............................................................................................284
Distribution of Chaetodactylidae ................................................................................286
Hosts of Chaetodactylidae............................................................................................290
Bibliography of Chaetodactylidae ...............................................................................293
The Mite Family Hyadesiidae ..............................................................................................303
Composition of Hyadesiidae........................................................................................303
Catalogue of Hyadesiidae .............................................................................................304
Index of Hyadesiidae .....................................................................................................310
Distribution of Hyadesiidae .........................................................................................311
Bibliography of Hyadesiidae ........................................................................................312
The Mite Family Algophagidae Fain ...................................................................................315
Acarorum Catalogus IX 7
Composition of Algophagidae .....................................................................................316
Catalogue of Algophagidae ..........................................................................................316
Index of Algophagidae ..................................................................................................321
Distribution of Algophagidae ......................................................................................322
Hosts of Algophagidae ..................................................................................................323
Bibliography of Algophagidae .....................................................................................323
The Mite Family Hemisarcoptidae ......................................................................................326
Composition of Hemisarcoptidae ...............................................................................326
Catalogue of Hemisarcoptidae.....................................................................................326
Index of Hemisarcoptidae ............................................................................................332
Distribution of Hemisarcoptidae.................................................................................332
Hosts of Hemisarcoptidae ............................................................................................334
Bibliography of Hemisarcoptidae ................................................................................335
The Mite Family Carpoglyphidae........................................................................................339
Composition of Carpoglyphidae .................................................................................339
Catalogue of Carpoglyphidae ......................................................................................340
Index of Carpoglyphidae ..............................................................................................342
Distribution of Carpoglyphidae ..................................................................................342
Hosts of Carpoglyphidae ..............................................................................................343
Bibliography of Carpoglyphidae..................................................................................343
The Mite Family Winterschmidtiidae (Saproglyphidae) ..................................................344
Composition of Winterschmidtiidae ..........................................................................345
Catalogue of Winterschmidtiidae ...............................................................................346
Index of Winterschmidtiidae .......................................................................................377
Distribution of Winterschmidtiidae ...........................................................................380
Hosts of Winterschmidtiidae .......................................................................................384
Bibliography of Winterschmidtiidae...........................................................................387
Acknowledgements ...............................................................................................................396
Acarorum Catalogus IX 9
Introduction
The present 9th volume of the series Acarorum Catalogus contains lists of mites of 13
families, 225 genera and 1268 species of the superfamilies Schizoglyphoidea, Histiostomatoidea, Canestrinioidea and Hemisarcoptoidea. Most of these mites live on insects
or other animals (as parasites, phoretic or commensals), some inhabit rotten plant material, dung or fungi. Mites of the families Chetochelacaridae and Lophonotacaridae are
specialised to live with Myriapods (Diplopoda). The peculiar aquatic or intertidal mites
of the families Hyadesidae and Algophagidae are also included.
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Acarorum Catalogus IX
Acariformes, Acaridida
The Mite Superfamily Schizoglyphoidea
Contains so far only one family of mites, found in New Guinea on Coleoptera. The
postnymphal stage remains unknown.
The Mite Family Schizoglyphidae
Schizoglyphidae Mahunka, 1978: 115
Monotypic genus of mites found on Tenebrionidae beetles.
Catalogue of Schizoglyphidae
Fam. Schizoglyphidae
Schizoglyphidae Mahunka, 1978: 115
Type genus: Schizoglyphus Mahunka, 1978
Genus Schyzoglyphus Mahunka
Schyzoglyphus Mahunka, 1978: 115
Type species: Schizoglyphus biroi Mahunka, 1978
Schizoglyphus biroi Mahunka
Schizoglyphus biroi Mahunka, 1978: 115
Type locality: Geelvink [Bay]
Distribution: Indonesia (Western New Guinea)
Host: Tagalus tibialis (Coleoptera: Tenebrionidae)
Index of Schyzoglyphidae
(Names in bold are accepted as valid)
biroi, Schizoglyphus 10
Schizoglyphidae 10
Schizoglyphus 10
Schizoglyphoidea 10
Acarorum Catalogus IX 11
Distribution of Schizoglyphidae
Indonesia (Mahunka, 1978) – Schizoglyphus biroi [Western New Guinea]
Hosts of Schizoglyphidae
Insecta
Coleoptera
Fam. Tenebrionidae
Tagalus tibialis – Schizoglyphus biroi
Bibliography of Schizoglyphidae
Mahunka S. 1978. Schizoglyphidae fam. n. and new taxa of Acaridae and Anoetidae (Acari:
Acarida). – Acta Zoologica Academiae Scientiarum Hungaricae, 24(1–2): 115.
The Mite Superfamily Histiostomatoidea
(Anoetoidea)
The Superfamily contains only one family - Histiostomatidae (Anoetidae).
The Mite Family Histiostomatidae
(Anoetidae)
Histiostomatidae Berlese, 1897
Anoetidae Oudemans, 1904; Hughes & Jackson, 1958
Small or middle-sized mites (250–900 µm), living in rotten plant material, in the dung
and fungi, some of them on animals (Hirudinea, Crustacea, Myriapoda, Insecta, Mammalia). Their study starts with the description of Hypopus feroniarum Dufour, 1839,
Hypopus alicola Dujardin, 1842 and the first genera Anoetus Dujardin, 1842 and Histiostoma Kramer, 1876. Follow the many publications of Oudemans, who described in
1904 the family Anoetidae. However, OConnor (1984) insisted that the family name
Histiostomatidae Berlese, 1897 antedates Anoetidae Oudemans, 1904, and since than
the widely known name Anoetidae has been replaced. Vitzthum (1931) composed
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Acarorum Catalogus IX
the review of 10 genera of Anoetidae. Two revisions of the family took place almost
simultaneously: of Scheucher (1957) and of Hughes & Jackson (1958). Until the revision of Scheucher (1957) only 12 valid genera have been described, seven of them by
Oudemans: Anoetus Dujardin, 1842, Histiostoma Kramer, 1876, Zwickia Oudemans,
1924, Anoetoglyphus Vitzthum, 1927, Glyphanoetus Oudemans, 1929, Myianoetus
Oudemans, 1929, Seliea Oudemans, 1929, Munduytia Oudemans, 1929, Cederhjelmia
Oudemans, 1931, Creutzeria Oudemans, 1932, Chiropteranoetus Womersley, 1942,
Prowichmannia Radford, 1950. From the 45 genera described after 1950 16 are due to
A. Fain and co-authors and 20 to S. Mahunka and co-authors (most of them monospecific) . The other major contributors are Sevastyanov, OConnor, Southcott, Fashing,
Woodring, Samšinák, Nesbitt, Ashfaq & Sarwar, Ashfaq, Chaudhri & Aheer, Eraky, and
others. From the 58 genera 34 are monotypic. Only six genera include more than 10 sp.
– Histiostoma (236, Anoetus (93), Myianoetus (48), Anoetoglyphus (15), Glyphanoetus
(14), and Rhopalanoetus (11).
Still exists the Anoetus – Histiostoma problem. According to Woodring & Moser
(1970),”…there is sufficient distinction between the concept of Anoetus and Histiostoma
to warrant the maintenance of the separate groups at the generic level. In this case the
older name Anoetus has priority over Histiostoma, and Histiostoma Kramer, 1876 must
be synonymized under Anoetus Dujardin, 1842, which is the type and name-bearing
genus of the family Anoetidaе”. However, for Sevastianov (1975) Histiostoma is junior
homonyme of Anoetus.
Composition of Histiostomatidae (Anoetidae)
Fam. Histiostomatidae (Anoetidae) - 58 gen., 522 sp.
Subfam. Ceylanoetinae Mahunka, 1974–6 gen., 15 sp.
Ceylanoetus Mahunka, 1974–1 sp.
Chiropteranoetus Womersley, 1942–1 sp.
Conglanoetus Mahunka, 1978 (= Comyianoetus Fain et Philips, 1979) – 3 sp.
Capronomoia Mahunka, 1976–7 sp.
Creutzeria Oudemans, 1932–2 sp.
Curculanoetus Fain, 1974–1 sp.
Subfam. Fibulanoetinae Fain, Camerik, Lukoschus et Kniest, 1980–3 gen., 18 sp.
Fibulanoetus Mahunka, 1973–3 sp.
Ghanoetus Mahunka, 1973–1 sp.
Glyphanoetus Oudemans, 1929–14 sp.
Subfam. Histiostomatinae Berlese, 1897 (Anoetinae Oudemans, 1904) - 46 gen.,
489 sp.
Ameranoetus Ide et Mahunka, 1978–1 sp.
Acarorum Catalogus IX 13
Amyzanoetus Fain, 1976–1 sp.
Ancyranoetus Fain et Santiago-Blay, 1993–1 sp.
Anoetoglyphus Vitzthum, 1927–15 sp.
Anoetus Dujardin, 1842–93 sp. (?)(= Nodipalpus Karpelles, 1893)
Aphodanoetus Bongers, OConnor et Lukoschus, 1985–1 sp.
Auricanoetus Fain et Zumpt, 1974–1 sp.
Austranoetus Fain, 1976–1 sp.
Bonomoia Oudemans, 1911–9 sp.
Bothyanoetus Fain et Camerik, 1978–1 sp.
Cederhjelmia Oudemans, 1931–1 sp.
Chiloanoetus Fain, 1974–1 sp.
Hauserania Mahunka, 1972–2 sp.
Hexanoetus Fain et Philips, 1979–1 sp.
Histiostoma Kramer, 1876–236 sp.
Cavanoetus Mahunka, 1982
Histiostoma Kramer, 1876
Ichtanoetus Fain et Belpaire, 1985
Hormosianoetus Fain, 1980–1 sp.
Hymenanoetus Mahunka, 1963–1 sp.
Insulanoetus Sevastyanov, 1973–1 sp.
Kanoetus Samšinák, 1966–3 sp.
Kaszabanoetus Mahunka, 1976–1 sp.
Loxanoetus Fain, 1970–2 sp.
Munduytia Oudemans, 1929–1 sp.
Nepenthacarus Fashing, 2002–1 sp.
Otanoetus Fain et Zumpt, 1974–1 sp.
Ovanoetus Fain et van Goethem, 1985–1 sp.
Peripatetes Mahunka, 1976–1 sp.
Porrhanoetus Mahunka, 1963–1 sp.
Probonomoia Fain et Rack, 1987–9 sp.
Prowichmannia Radford, 1950–3 sp.
Psyllanoetus Fain et Beaucournu, 1973–3 sp.
Pteranoetus Mahunka, 1978–4 sp.
Rhaphidothrix Mahunka, 1967–3 sp.
Rhopalanoetus Scheucher, 1957–11 sp.
Richardanoetus Samšinák, 1989–1 sp.
Sarraceniopus Fashing et OConnor, 1984–3 sp.
Scheucheria Mahunka, 1969–1 sp.
Scutanoetus Mahunka, 1969–1 sp.
Seliea Oudemans, 1929–1 sp.
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Acarorum Catalogus IX
Semianoetus Mahunka, 1976–2 sp.
Stercoranoetus Mahunka et Mahunka-Papp, 1991–2 sp.
Synanoetus Mahunka, 1972–1 sp.
Syringanoetus Fain, 1980–2 sp.
Teinokyra Mahunka, 1973–1 sp.
Traskorchestianoetus Fain et Colloff, 1990–2 sp.
Xenanoetus Mahunka, 1969–5 sp.
Zwickia Oudemans, 1924–3 sp.
Subfam. Myianoetinae Scheucher, 1957–48 sp.
Myianoetus Oudemans, 1929–48 sp.
Subfam. Scolianoetinae Fain, 1974–5 sp.
Scolianoetus Fain, 1974–5 sp.
Subfam. Spinanoetinae Scheucher, 1957–4 sp
Spinanoetus Scheucher, 1957–4 sp
Catalogue of Histiostomatidae
Family Histiostomatidae Berlese
Histiostomatidae Berlese, 1897
Anoetidae Oudemans, 1904: 191. Type: Anoetus Dujardin, 1842
Nodipalpidae Oudemans, 1923: 206
Kaszabanoetinae Mahunka, 1976: 318
Type genus: Histiostoma Kramer, 1876
Subfamily Ceylanoetinae Mahunka
Ceylanoetinae Mahunka, 1974c: 507
Type genus: Ceylanoetus Mahunka, 1974
Glyphanoetinae Mahunka, 1972: 371
Type genus: Glyphanoetus Oudemans, 1929
Genus Ceylanoetus Mahunka
Ceylanoetus Mahunka, 1974c: 506
Type species: Ceylanoetus excavatus Mahunka, 1974
Ceylanoetus excavatus Mahunka
Ceylanoetus excavatus Mahunka, 1974c: 507
Type locality: Central, Mahaweli Ganga, “7 Meilen flussabwärts von Kandy”
Distribution: Sri Lanka
Acarorum Catalogus IX 15
Genus Chiropteranoetus Womersley
Chiropteranoetus Womersley, 1942: 92
Type species: Chiropteranoetus chalinolobus Womersley, 1942
Chiropteranoetus chalinolobus Womersley
Chiropteranoetus chalinolobus Womersley, 1942: 92
Type locality: Australia
Distribution: Australia
Genus Conglanoetus Mahunka
Conglanoetus Mahunka, 1978: 143
Comyianoetus Fain & Philips 1979: 147 – Type: Comyianoetus denticulatus Fain et Philips,
1979
Type species: Conglanoetus ornatofoveolatus Mahunka, 1978
Conglanoetus denticulatus (Fain et Philips)
Comyianoetus denticulatus Fain & Philips, 1979: 147; Fain et al., 1993: 17
Conglanoetus denticulatus (Fain et Philips):
Type locality: Syracuse, N.Y.
Distribution: Norway, USA (N.Y.)
Host: nest of Bubo bubo and on Trox scaber (Coleoptera: Trocidae)
Conglanoetus ornatofoveolatus Mahunka
Conglanoetus ornatofoveolatus Mahunka, 1978: 143
Type locality: Oriente 00°24’S, 76°36’W, Limoncocha
Distribution: Ecuador
Host: Labidus praedator (Hymenoptera: Formicidae, Dorylinae)
Conglanoetus puteanus (Fain)
Comyianoetus puteanus Fain, in Fain, Vangeluwe, Degreef & Wauthy,1993: 17
Conglanoetus puteanus (Fain):
Type locality: Belgium
Distribution: Belgium
Genus Capronomoia Mahunka
Capronomoia Mahunka, 1976b: 321
Type species: Bonomoia sphaerocerae Vitzthum, 1922
Capronomoia asper Ashfaq et Chaudhri
Capronomoia asper Ashfaq & Chaudhri, 1988: 355
16
Acarorum Catalogus IX
Copronomoia asper Ashfaq & Sarwar, 2001: 129
Type locality: 5 km N of Faisalabad
Distribution: Pakistan
Capronomoia bellulus Ashfaq et Chaudhri
Capronomoia bellulus Ashfaq & Chaudhri, 1988: 352
Copronomoia bellulus: Ashfaq & Sarwar, 2001: 129
Type locality: 3 km W Murree
Distribution: Pakistan
Capronomoia gradatus Ashfaq et Sarwar
Copronomoia gradatus Ashfaq & Sarwar, 2001: 127
Type locality: Quetta
Distribution: Pakistan
Capronomoia mahunkai Eraky
Copronomoia mahunkai Eraky, 1999: 55; Abdelgayed et al., 2019: 32
Type locality: Assiut
Distribution: Egypt
Capronomoia pango Ashfaq et Chaudhri
Capronomoia pango Ashfaq & Chaudhri, 1988: 350
Copronomoia pango Ashfaq & Sarwar, 2001: 129
Type locality: 5 km east of Faisalabad
Distribution: Pakistan
Capronomoia sphaerocerae (Vitzthum)
Bonomoia sphaerocerae Vitzthum, 1922: 545; Scheucher, 1957: 365; Mahunka, 1970: 70
Capronomoia sphaerocerae (Vitzthum): Mahunka, 1976: 321
Type locality: Germany
Distribution: Germany, Hungary, Saudi Arabia, Syria
Hosts: Sphaerocera subsultans (Diptera: Sphaeroceridae [Syrphidae]); also on Borboridae,
Scatophagidae and other Diptera.
Capronomoia trossis Ashfaq et Sarwar
Capronomoia trossis Ashfaq & Sarwar, 2001: 125
Type locality: Khanewal
Distribution: Pakistan
Genus Creutzeria Oudemans
Creutzeria Oudemans, 1932: 207
Type species: Creutzeria tobaica Oudemans, 1932
Acarorum Catalogus IX 17
Creutzeria seychellensis Nesbitt
Creutzeria seychellensis Nesbitt, 1979: 1202
Type locality: Seychelles
Distribution: Seychelles
Creutzeria tobaica Oudemans
Creutzeria tobaica Oudemans, 1932: 207; Buitendijk, 1945: 363
Type locality: Haranggaol
Distribution: Indonesia
Genus Curculanoetus Fain
Curculanoetus Fain, 1974: 66
Type species: Curculanoetus rhynchophorus Fain, 1974
Curculanoetus rhynchophorus Fain
Curculanoetus rhynchophorus Fain, 1974: 66
Type locality: Lemba nr. Kinshasa
Distribution: DR Congo
Host: Rhynchophorus phoenicis (Coleoptera: Curculionidae)
Subfamily Fibulanoetinae Fain, Camerik, Lukoschus et Kniest
Fibulanoetinae Fain, Camerik, Lukoschus & Kniest, 1980: 39
Type genus: Fibulanoetus Mahunka, 1973
Genus Fibulanoetus Mahunka
Fibulanoetus Mahunka, 1973a: 102
Type species: Fibulanoetus labiatus Mahunka, 1973
Fibulanoetus labiatus Mahunka
Fibulanoetus labiatus Mahunka, 1973a: 102
Type locality: Meru
Distribution: Tanzania
Fibulanoetus longitarsus Fain, Camerik, Lukoschus et Kniest
Fibulanoetus longitarsus Fain, Camerik, Lukoschus & Kniest, 1980: 42
Type locality: Thamaga
Distribution: Botswana
Host: Scarabaeus galenus (Coleoptera: Scarabaeidae)
Fibulanoetus mahunkai Fain, Camerik, Lukoschus et Kniest
Fibulanoetus mahunkai Fain, Camerik, Lukoschus & Kniest, 1980: 41
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Acarorum Catalogus IX
Type locality: Mahalapye
Distribution: Botswana
Host: Scarabaeus galenus (Coleoptera: Scarabaeidae)
Genus Ghanoetus Mahunka
Ghanoetus Mahunka, 1973a: 104
Type species: Ghanoetus marmoreus Mahunka, 1973
Ghanoetus marmoreus Mahunka
Ghanoetus marmoreus Mahunka, 1973a: 104; 1975: 60
Type locality: Kumasi (Ghana)
Distribution: Gambia, Ghana
Genus Glyphanoetus Oudemans
Glyphanoetus Oudemans, 1929: 450
Type species: Glyphanoetus fulmeki Oudemans, 1929 (= Hypopus phyllotrichus Berlese,
1881)
Glyphanoetus africanus Mahunka
Glyphanoetus africanus Mahunka, 1967a: 172
Type locality: Kindamba, Meya, Bangu Forest
Distribution: Rep. of the Congo (Brazzaville)
Glyphanoetus brevisetus Mahunka
Glyphanoetus brevisetus Mahunka, 1963a: 38
Type locality: Cazombo (Alto Zambeze)
Distribution: Angola
Glyphanoetus eboris Fain
Glyphanoetus eboris Fain, 1974: 88
Type locality: La Savane de Lamto
Distribution: Ivory Coast
Glyphanoetus flagellatus Mahunka
Glyphanoetus flagellatus Mahunka, 1963d: 370
Type locality: Olkokola, nest of Tachyoryctes sp.
Distribution: Tanzania
Glyphanoetus incanus Ashfaq, Sarwar et Akbar
Glyphanoetus incanus Ashfaq, Sarwar & Akbar, 1999: 21
Type locality: Lahore, Shimla Hill
Acarorum Catalogus IX 19
Distribution: Pakistan
Glyphanoetus leptos Ashfaq, Sarwar Akbar
Glyphanoetus leptos Ashfaq, Sarwar & Akbar, 1999: 23
Type locality: Azad Kashmir, Kotli
Distribution: Pakistan
Glyphanoetus longus Mahunka
Glyphanoetus longus Mahunka, 1969b: 371
Type locality: Mt. Meru, eastern slope
Distribution: Tanzania
Glyphanoetus mahunkai Eraky
Glyphanoetus mahunkai Eraky, 1994: 218
Type locality: Egypt
Distribution: Egypt
Glyphanoetus micidus Chaudhri
Glyphanoetus micidus Chaudhri, 1986: 212
Type locality: Pakistan
Distribution: Pakistan
Glyphanoetus nomiensis Cross
Glyphanoetus nomiensis Cross, 1968: 329
Type locality: Wyoming
Distribution: USA (Wyoming, Louisiana)
Hosts: Nomia melanderi, N. nortoni (Hymenoptera: Halictidae)
Glyphanoetus pappi Mahunka
Glyphanoetus pappi Mahunka, 1982: 63
Type locality: Juan Fernandez Islands
Distribution: Juan Fernandez Islands
Glyphanoetus phyllotrichus (Berlese)
Hypopus phyllotrichus Berlese, 1881: AMS Fasc.9, No 2
Anoetus phyllotrichus Oudemans, 1905b: 238 – from “Synchitodes crenatus” - Velp
Glyphanoetus fulmeki Oudemans, 1929: 451
Glyphanoetus phyllotrichus (Berlese): Scheucher, 1957: 351; Samšinák, 1984: 62; Mahunka,
1970: 59; Mahunka & Mahunka-Papp: 1991: 286
Type locality: Padua (Italy)
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Acarorum Catalogus IX
Distribution: Austria, Germany, Holland, Hungary, Italy; Cabo Verde
Hosts: Ips laricis, I. proximus (Coleoptera: Curculionidae, Scolytinae), Limosina talparum
(Diptera: Sphaeroceridae)
Glyphanoetus processum Eraky
Glyphanoetus processum Eraky, 1994: 217
Type locality: Egypt
Distribution: Egypt
Glyphanoetus szunyoghyi Mahunka
Glyphanoetus szunyoghyi Mahunka, 1961: 528 (as “Gylphanoetus”)
Type locality: Arusha
Distribution: Tanzania
Glyphanoetus venustissimus (Berlese)
Anoetus venustissimus Berlese, 1918: 115
Glyphanoetus venustissimus (Berlese): Scheucher, 1957: 354
Type locality: Florence
Distribution: Germany, Italy
Subfamily Histiostomatinae Scheucher
Anoetinae Oudemans, 1904: 191 - Type genus: Anoetus Dujardin, 1842
Histiostomatini Scheucher, 1957: 287; Fain, 1974: 58
Histiostominae Scheucher: Fain, 1974: 63
Histiostomatinae Berlese, 1897; Fain, 1974: 58
Type genus: Histiostoma Kramer, 1876: 105
Genus Ameranoetus Ide et Mahunka
Ameranoetus Ide & Mahunka, 1978: 47
Type species: Ameranoetus hematobii Ide et Mahunka, 1978
Ameranoetus hematobii Ide et Mahunka
Ameranoetus hematobii Ide & Mahunka, 1978: 48
Type locality: USA
Distribution: USA
Host: Haematobia irritans (Diptera: Muscidae)
Genus Amyzanoetus Fain
Amyzanoetus Fain, 1976: 318
Type species: Amyzanoetus halophilus Fain, 1976
Acarorum Catalogus IX 21
Amyzanoetus halophilus Fain
Amyzanoetus halophilus Fain, 1976: 318
Type locality: Kerguelen Island
Distribution: Kerguelen Island
Genus Ancyranoetus Fain et Santiago-Blay
Ancyranoetus Fain & Santiago-Blay, 1993: 337
Type species: Ancyranoetus virkkii Fain et Santiago-Blay, 1993
Ancyranoetus virkkii Fain et Santiago-Blay
Ancyranoetus virkkii Fain & Santiago-Blay, 1993: 339
Type locality: Puerto Rico
Distribution: Puerto Rico
Host: Lema nigripes (Coleoptera: Chrysomelidae, Criocerinae)
Genus Anoetoglyphus Oudemans ]
Anoetoglyphus Oudemans [Vitzthum in litt.], 1927: 75
Type species: Anoetoglyphus ateuchi Oudemans, 1927
Anoetoglyphus africanus Mahunka
Anoetoglyphus africanus Mahunka, 1965: 448
Type locality: Lefinie Reservation, Nambouli River
Distribution: Congo - Brazaville
Anoetoglyphus antennatus Mahunka
Anoetoglyphus antennatus Mahunka, 1975: 58
Type locality: Kumasi
Distribution: Ghana
Anoetoglyphus ateuchi Vitzthum
Anoetoglyphus ateuchi Oudemans, 1927: 75 (Vitzthum, in litt.)
Type locality: South Italy
Distribution: Italy, Sicily
Host: Scarabaeus semipunctatus (Coleoptera: Scarabaeidae)
Anoetoglyphus bifastigatus Mahunka
Anoetoglyphus (?) bifastigatus Mahunka, 1967: 9
Type locality: Dundo
Distribution: Angola
Host: Rhynchophorus phoenicis (Coleoptera: Curculionidae)
22
Acarorum Catalogus IX
Anoetoglyphus copridis Samšinák
Anoetoglyphus copridis Samšinák, 1962: 92
Type locality: Neusiedler See
Distribution: Austria, Hungary
Host: Copris lunaris (Coleoptera: Scarabaeidae, Coprinae)
Anoetoglyphus ellipticus Mahunka
Anoetoglyphus ellipticus Mahunka, 1973a: 114
Type locality: Nakpanduri
Distribution: Ghana
Anoetoglyphus globulus Mahunka
Anoetoglyphus globulus Mahunka, 1975: 55
Type locality: Kumasi
Distribution: Ghana
Anoetoglyphus immaturus Mahunka
Anoetoglyphus (?) immaturus Mahunka, 1965: 448
Type locality: Lefinie Reservation, Nambouli River
Distribution: Republic of Congo - Brazaville
Anoetoglyphus langi Mahunka
Anoetoglyphus langi Mahunka, 1978: 131
Type locality: Lai Khe
Distribution: Vietnam
Hosts: Coleoptera (Scarabaeidae)
Anoetoglyphus latus Mahunka
Anoetoglyphus latus Mahunka, 1978: 56
Type locality: Ghana
Distribution: Ghana
Anoetoglyphus levis Mahunka
Anoetoglyphus levis Mahunka, 1967d: 161
Type locality: Cuc-phuong, Bong
Distribution: Vietnam
Anoetoglyphus lootsi Mahunka
Anoetoglyphus lootsi Mahunka, 1979: 140
Type locality: Transvaal, Nylsvley Res., 35 km NE from Pretoria
Acarorum Catalogus IX 23
Distribution: South Africa
Anoetoglyphus polygonatus Mahunka
Anoetoglyphus polygonatus Mahunka, 1973a: 302
Type locality: Nakpanduri
Distribution: Ghana
Anoetoglyphus szunyoghyi Mahunka
Anoetoglyphus szunyoghyi Mahunka, 1973a: 115; 1975: 59
Type locality: Banda-Kwanta (Ghana)
Distribution: Gambia, Ghana
Anoetoglyphus tunesiensis Mahunka
Anoetoglyphus tunesiensis Mahunka, 1981: 135
Type locality: Tunisia
Distribution: Tunisia
Genus Anoetus Dujardin
Anoetus Dujardin, 1842: 316
Nodipalpus Karpelles, 1893: 128
Type species: Hypopus alicola Dujardin, 1842
Anoetus aelleni Mahunka
Anoetus aelleni Mahunka, 1974d: 508
Type locality: Central, Mahaweli Ganga, “7 Meilen flussabwärts von Kandy”
Distribution: Sri Lanka
Anoetus akodoni Mahunka
Anoetus akodoni Mahunka, 1963c: 499
Type locality: El Bolsόn
Distribution: Argentina
Anoetus alicola (Dujardin)
Hypopus alicola Dujardin, 1842: 316
Anoetus discrepans Oudemans, 1903 : 103. Type: Bremen, on Halictus 4-notatus; Oudemans, 1903: 103
Anoetus alicola (Dujardin): Buitendijk, 1945: 361; Scheucher, 1957: 374
Type locality: Hülden an de Aller
Distribution: Germany
Hosts: Halictus 4-notatus (Hymenoptera: Halictidae)
24
Acarorum Catalogus IX
Anoetus angularis Mahunka
Anoetus angularis Mahunka, 1974d : 511
Type locality: Uva, Diyaluma Waterfall
Distribution: Sri Lanka
Anoetus anoetopsis Mahunka
Anoetus anoetopsis Mahunka, 1973a : 89
Type locality: Bui Camp
Distribution: Ghana
Host: Coleoptera
Anoetus bradymeri Mahunka
Anoetus bradymeri Mahunka, 1972c: 349
Type locality: Tulagi
Distribution: Salomon Islands
Hosts: Bradymerus leveri, Androsus greensladei (Coleoptera: Tenebrionidae)
Anoetus bushlandi Hughes et Jackson
Anoetus bushlandi Hughes & Jackson, 1958: 133
Type locality: Kerrville, Texas
Distribution: USA (Texas)
Hosts: on stable fly
Anoetus cacuminatum Mahunka
Anoetus cacuminatum Mahunka, 1969: 180
Type locality: Mongolia
Distribution: Mongolia
Anoetus caljubiensis Gad Hamada Hassan et Sevastianov
Anoetus caljubiensis Gad Hamada Hassan & Sevastianov, in Radi & Sevastyanov, 1991: 48
Type locality: Egypt
Distribution: Egypt
Host: Cataglyphis bicolor (Hymenoptera: Formicidae)
Anoetus capillatus Mahunka
Anoetus capillatus Mahunka, 1967d: 151
Type locality: Giang Seo
Distribution: Vietnam
Anoetus castrii Mahunka
Anoetus castrii Mahunka, 1972d: 306
Acarorum Catalogus IX 25
Type locality: Maipu, Quebrada, La Plata
Distribution: Chile
Anoetus ceylanensis Mahunka
Anoetus ceylanensis Mahunka, 1974d : 511
Type locality: Central, Mahaweli Ganga, “7 Meilen flussabwärts von Kandy”
Distribution: Sri Lanka
Anoetus chilensis Mahunka
Anoetus chilensis Mahunka, 1972d: 309
Type locality: Curacavi, Los Cerillos
Distribution: Chile
Anoetus cirrosus Mahunka
Anoetus cirrosus Mahunka, 1972d: 308
Type locality: Maipu, Quebrada, La Plata
Distribution: Chile
Anoetus collis Mahunka
Anoetus collis Mahunka, 1965: 445
Type locality: Lefinie Reservation, Nambouli River
Distribution: Congo (Brazzaville)
Anoetus congoensis Mahunka
Anoetus congoensis Mahunka, 1967a: 166
Type locality: Kindamba, Meya, Bangu forest
Distribution: Rep. of the Congo (Brazzaville)
Anoetus coprophilus Mahunka
Anoetus coprophilus Mahunka, 1965: 446
Type locality: Lefinie Reservation, Nambouli River
Distribution: Rep. of Congo (Brazzaville)
Anoetus debilis Woodring
Anoetus debilis Woodring, 1973: 319
Type locality: Turrialba, Cartago Prov.
Distribution: Costa Rica
Anoetus decorus Mahunka
Anoetus decorus Mahunka, 1967d: 152
Type locality: Cuc-phuong, Bong
Distribution: Vietnam
26
Acarorum Catalogus IX
Anoetus dolifer Mahunka
Anoetus dolifer Mahunka, 1977: 94
Type locality: Switzerland, Wallis, Bagnes
Distribution: France, Switzerland
Hosts: Myrmica ruginodis, M. sabuleti (Hymenoptera: Formicidae - anthills)
Anoetus domrowi (Fain)
Anoetostoma domrowi Fain, 1968: 246
Type locality: Butemu Village, Finisterre Range, Madang, Central District
Distribution: Papua New Guinea
Host: Scoliophthalmus sp. near micans (Diptera: Chloropidae)
Anoetus dudichi Mahunka
Anoetus dudichi Mahunka, 1962: 431
Type locality: Jόsva, Nagyoldal
Distribution: Hungary
Anoetus endroedyyoungai Mahunka
Anoetus endroedyyoungai Mahunka, 1973a : 92
Type locality: Bui Camp
Distribution: Ghana
Anoetus exornatus Sevastianov
Anoetus exornatus Sevastianov, 1973: 849
Type locality: Roshtino Village, Leningradskaya Obl.
Distribution: Russia
Host: Formica rufa (Hymenoptera: Formicidae)
Anoetus fetters Samšinák
Anoetus fetters Samšinák, 1989: 97
Type locality: DR Congo
Distribution: DR Congo
Host: Metaborborus calcaratus (Diptera: Sphaeroceridae)
Anoetus fibulatus Mahunka
Anoetus fibulatus Mahunka, 1973a : 94
Type locality: Meru
Distribution: Tanzania
Anoetus ghanensis Mahunka
Anoetus ghanensis Mahunka, 1973: 94
Acarorum Catalogus IX 27
Type locality: Ghana
Distribution: Ghana
Anoetus grapefruit Samšinák
Anoetus grapefruit Samšinák, 1989: 95
Type locality: Centre Co. Fillmore Cap
Distribution: USA
Host: Diptera indet.
Anoetus hauseri Mahunka
Anoetus hauseri Mahunka, 1972: 1196; 1972e: 951
Type locality: Kephallinia, Aenos
Distribution: Greece (Kephallinia)
Anoetus halictonida (Woodring)
Histiosoma halictonida Woodring, 1973: 312
Anoetus halictonida (Woodring)
Type locality: Canada
Distribution: Canada
Anoetus helena Samšinak
Anoetus helena Samšinak, 1989: 95
Type locality: Ecuador
Distribution: Ecuador
Host: Richardia (plant)
Anoetus heliocopridis Vitzthum
Anoetus heliocopridis Vitzthum, 1926: 182; Mahunka, 1974a : 141
Type locality: Meru (Tanzania)
Distribution: Ghana, Tanzania
Anoetus helobius Sevastianov
Anoetus helobius Sevastianov, 1973: 851
Type locality: Hadjibeyskiy Liman nr. Odessa
Distribution: Ukraine
Anoetus horridus Mahunka
Anoetus horidus Mahunka, 1972c: 352
Type locality: San Cristobal
Distribution: Salomon Islands
28
Acarorum Catalogus IX
Hosts: Leiochrodes suturalis, Scyophagus piceus (Coleoptera: Tenebrionidae)
Anoetus hospitus Sevastianov
Anoetus hospitus Sevastianov, 1973: 851
Type locality: Tchemerovtsi Village, Hmelnitskaya Obl.
Distribution: Ukraine
Host: anthip of Myrmica ruginodis (Hymenoptera: Formicidae)
Anoetus humilis Fain
Anoetus humilis Fain, 1974b: 86
Type locality: La Savane de Lamto
Distribution: Ivory Coast
Anoetus impressus Mahunka
Histiostoma impressum Mahunka, 1963a : 31; 1963b: 59
Anoetus impressus (Mahunka): Mahunka, 1967a: 167; 1973a: 96
Type locality: Angola, Lumeje II, N of Santando Station of C.F.B., Poste de Bussaco (80
km E of Luso)
Distribution: Angola, Rep. of the Congo (Brazaville), DR Congo, Ghana
Anoetus indetonsus Sevastianov
Anoetus indetonsus Sevastianov, 1973: 852
Type locality: Odessa area
Distribution: Ukraine
Host: Coleoptera (Carabidae)
Anoetus indicus Oudemans
Anoetus indicus Oudemans, 1911: 173; Hughes & Jackson, 1958: 80; Mahunka, 1963a:
32; 1973a : 96
Histiostoma indicus (Oudemans): Buitendijl, 1945; Hughes & Jackson, 1958: 80
Histiostoma indica (Oudemans): Mahunka, 1963b: 60
Type locality: Java, Banjoewangi
Distribution: Angola, Ghana, Indonesia (Java)
Host: Blattida fam., gen. sp.
Anoetus insolita Woodring et Moser
Anoetus insolita Woodring & Moser, 1970: 1237
Type locality: Elizabeth, La.
Distribution: USA
Host: from the boring dust of Dendroctonus frontalis and P. taeda (Coleoptera: Curculionidae)
Acarorum Catalogus IX 29
Anoetus irinus Hassan et Sevastianov
Anoetus irinus Hassan & Sevastianov, in Radi & Sevastyanov 1991: 45
Type locality: Egypt
Distribution: Egypt
Host: Iris tingitana (plant)
Anoetus kaszabi Mahunka
Anoetus kaszabi Mahunka, 1969: 181
Type locality: Mongolia
Distribution: Mongolia
Anoetus komarpictoris Samšinák
Anoetus komarpictoris Samšinák, 1970: 33
Type locality: Kozákov
Distribution: Czech Rep.
Host: Formica lemani (Hymenoptera: Formicidae)
Anoetus longipes (Oudemans)
Anoetus longipes Oudemans, 1911: 171; Mahunka, 1967d: 154
Histiostoma longipes (Oudemans): Buitendijk, 1945: 362
Type locality: Banjoewangi
Distribution: Indonesia (Java), Vietnam
Host: Blattida fam. gen. sp.
Anoetus longisetus Mahunka
Anoetus longisetus Mahunka, 1967a: 167; 1973a : 96
Type locality: Kindamba, Meya, Louolo river
Distribution: Republic of the Congo, Ghana
Anoetus longitubus Mahunka
Anoetus longitubus Mahunka, 1969b: 365
Type locality: Mt. Meru, eastern slope
Distribution: Tanzania
Anoetus lunulatus Mahunka
Anoetus lunulatus Mahunka, 1971: 1198
Type locality: Kephallinia
Distribution: Greece (Kephallinia)
Anoetus machadoi Mahunka
Anoetus machadoi Mahunka, 1963b: 63; 1967a: 168
30
Acarorum Catalogus IX
Type locality: Dundo, Cundueji River (Angola)
Distribution: Angola, Rep. of the Congo
Anoetus macrosuctus Fain
Anoetus macrosuctus Fain, 1974b: 86
Type locality: La Savane de Lamto
Distribution: Ivory Coast
Anoetus microti Mahunka
Anoetus microti Mahunka, 1969: 182
Type locality: Mongolia
Distribution: Mongolia
Anoetus moshtohorensis Gad Hamada, Hassan et Sevastianov
Anoetus moshtohorensis Gad Hamada Hassan & Sevastianov, in Radi & Sevastyanov
1991: 47
Type locality: Egypt
Distribution: Egypt
Host: Labidura riparia (Dermaptera: Labiduridae)
Anoetus myrmicarum (Scheucher)
Histiostoma myrmicarum Scheucher, 1957: 300; Mahunka, 1961: 445
Anoetus myrmicarum (Scheucher): Samšinak, 1961: 204; Kurosa, 1976: 62
Type locality: Erlangen, Germany
Distribution: China, Czech Rep., Germany, Hungary, Malaysia
Hosts: Coptotermes formosanus, Macrotermes malaccensis (Isoptera: Rhinotermitidae,
Termitidae); ants (Hymenoptera, Formicidae: Myrmica, Lasius, Camponotus, Formica)
Anoetus neglectus Oudemans
Anoetus neglectus Oudemans, 1902: 46
Type locality: Bremen
Distribution: Germany
Host: Nicrophorus humator (Coleoptera: Silphidae)
Anoetus notabilis Sellnick
Anoetus notabilis Sellnick, 1940: 127
Type locality: Island
Distribution: Island
Anoetus opertus Sevastianov
Anoetus opertus Sevastianov, 1973: 852
Type locality: Tchemerovtsi Village, Hmelnitskaya Obl.
Acarorum Catalogus IX 31
Distribution: Ukraine
Host: Myrmica rubra (Hymenoptera: Formicidae)
Anoetus ornatissimus Mahunka
Anoetus ornatissimus Mahunka, 1973a: 96
Type locality: Meru
Distribution: Tanzania
Anoetus ornatus Mahunka
Anoetus ornatus Mahunka, 1967d: 154
Type locality: Tu-li
Distribution: Vietnam
Anoetus pannonicus Mahunka
Anoetus pannonicus Mahunka, 1968: 251
Type locality: Hungary
Distribution: Hungary
Host: phoretic on Dryocoetes villosus (Coleoptera: Curculionidae)
Anoetus parmatus Sevastianov
Anoetus parmatus Sevastianov, 1973: 853
Type locality: Tchemerovtsi Village, Hmelnitskaya Obl.
Distribution: Ukraine
Anoetus pauliani Mahunka
Anoetus pauliani Mahunka, 1967a: 168
Type locality: Mount Foari Reservation
Distribution: Rep. of the Congo (Brazzaville)
Anoetus pini (Scheucher)
Histiostoma pini Scheucher, 1957: 317
Anoetus pini (Scheucher): Woodring & Mosaer, 1970: 1255
Type locality: Erlangen
Distribution: Belgium, Hungary, Germany
Host: phoretic on Hylastes ater and H. cunicularius (Coleoptera: Curculionidae)
Anoetus pocsi Mahunka
Anoetus pocsi Mahunka, 1967d: 156
Type locality: Giang Seo
Distribution: Vietnam
32
Acarorum Catalogus IX
Anoetus polynesiacus Vitzthum
Anoetus polynesiacus Vitzthum, 1920: 64
Type locality: Fiji
Distribution: Fiji
Host: Eubactrus semiaeneus (Coleoptera: Brenthidae)
Anoetus porosus Mahunka
Anoetus porosus Mahunka, 1969: 183
Type locality: Mongolia
Distribution: Mongolia
Anoetus privorai Samšinák
Anoetus přivorai Samšinák, 1962: 89
Type locality: Roccamena, Palermo (Sicily)
Distribution: Sicily
Host: Aphaenogaster testaceopilosa (Hymenoptera: Formicidae)
Anoetus punctulatus Mahunka
Anoetus punctulatus Mahunka, 1967a: 169
Type locality: Brazzaville, Orstrom Parc
Distribution: Rep. of the Congo (Brazzaville)
Anoetus reticulatus Mahunka
Anoetus reticulatus Mahunka, 1969b: 366
Type locality: Mt. Meru, eastern slope
Distribution: Tanzania
Anoetus retractus Sevastianov
Anoetus retractus Sevastianov, 1973: 853
Type locality: Jaralgach Island (Black Sea)
Distribution: Jaralgach Isl., Crimea
Hosts: Bledius bicornis, B. sarmaticus (Coleoptera: Staphylinidae, Oxythelinae)
Anoetus samiensis Mahunka
Anoetus samiensis Mahunka, 1972: 952
Type locality: Kephallinia: Sami
Distribution: Greece (Kephalinia)
Anoetus serratus (Mahunka)
Histiostoma serrata Mahunka, 1962b: 429
Anoetus serratus (Mahunka), 1970: 26; Sevastianov, 1975: 400
Acarorum Catalogus IX 33
Type locality: Balatonfenyves
Distribution: Hungary
Host: Demetrias imperialis (Coleoptera: Carabidae)
Anoetus shamyi Samšinák
Anoetus shamyi Samšinák, 1970: 36
Type locality: Tzasweri
Distribution: Russia, Caucasus
Host: Formica lemani (Hymenoptera: Formicidae)
Anoetus shanensis Mahunka
Anoetus shanensis Mahunka, 1973a: 96
Type locality: Borbiri Mt Reserve
Distribution: Ghana
Anoetus sharae Gad Hamada Hassan et Sevastianov
Anoetus sharae Gad Hamada Hassan & Sevastianov, in Radi & Sevastyanov 1991: 49
Type locality: Egypt
Distribution: Egypt
Anoetus sinuosus Mahunka
Anoetus sinuosus Mahunka, 1973a : 299
Type locality: Sese
Distribution: Ghana
Anoetus solomoniensis Mahunka
Anoetus solomoniensis Mahunka, 1972c: 355
Type locality: San Cristobal
Distribution: Salomon Islands
Host: Leiochrodes suturalis (Coleoptera: Tenebrionidae)
Anoetus sordida Woodring et Moser
Anoetus sordida Woodring & Moser, 1970: 1238
Type locality: Elizabeth, Louisiana
Distribution: Canada, USA (Texas, Georgia, Louisiana)
Host: boring dust in galleries of Dendroctonus frontalis, Ips avulsus and I. calligraphus
(Col.: Curculionidae, Scolytinae), phoretic on Carcinops sp. (Histeridae) and Corticeus
coynei (Tenebrionidae)
Anoetus spinipes Mahunka
Anoetus spinipes Mahunka, 1973a: 99
34
Acarorum Catalogus IX
Type locality: Kumasi
Distribution: Ghana
Anoetus spinosus Mahunka
Anoetus spinosus Mahunka, 1967d: 158
Type locality: Cuc-phuong, Bong
Distribution: Vietnam
Anoetus striatus Mahunka
Histiostoma striatum Mahunka, 1963a: 36
Histiostoma striata Mahunka, 1963b: 62
Anoetus striatus (Mahunka): Mahunka, 1967a: 170
Type locality: Angola, Lumeje II, N of Santando Station of C.F.B., Poste de Bussaco (80
km E of Luso)
Distribution: Angola, Rep. of the Congo (Brazzaville)
Hosts: Dichaetodrilus incrassatus (Coleoptera: Carabidae), Rhynchophorus phoenicis (Col.:
Curculionidae)
Anoetus sumatrensis Oudemans
Anoetus sumatrensis Oudemans, 1903: 103
Type locality: Sumatra
Distribution: Indonesia
Anoetus tauffliebi Mahunka
Anoetus tauffliebi Mahunka, 1967a: 170
Type locality: Mount Foari Reservation
Distribution: Republic of the Congo (Brazzaville)
Anoetus topali Mahunka
Anoetus ? topali Mahunka, 1963c: 497; 1972 : 311
Type locality: Argentina, Chubut
Distribution: Argentina, Chile
Anoetus toxici Mahunka
Anoetus toxici Mahunka, 1972: 497; 1972 : 357
Type locality: Guadalcanal
Distribution: Salomon Islands
Host: Toxicum quadricorne rufipes (Coleoptera: Tenebrionidae)
Anoetus turcastanae Oudemans
Anoetus turcastanae Oudemans, 1917: 392
Acarorum Catalogus IX 35
Anoetus turcestanae Oudemans: Buitendijk, 1945: 361
Type locality: “Turkestan”
Distribution: “Turkestan”
Host: Homalocopris tmolus (Coleoptera: Scarabaeidae)
Anoetus urcerosus Sevastianov
Anoetus urcerosus Sevastianov, 1973: 855
Type locality: Tchemerovtsi Village, Hmelnitskaya Obl.
Distribution: Ukraine
Host: anthip of Formica rufa (Hymenoptera: Formicidae)
Anoetus variolatus Mahunka
Anoetus variolatus Mahunka, 1973a: 100
Type locality: Borbiri Mt. Res.
Distribution: Ghana
Anoetus vilhenaorum Mahunka
Anoetus vilhenaorum Mahunka, 1963b: 65
Type locality: Dundo, Cundueji River
Distribution: Angola
Anoetus virginiana Woodring et Moser
Anoetus virginiana Woodring & Moser, 1970: 1237; Kurosa, 1976: 62
Type locality: Virginia (USA)
Distribution: Malaysia, USA
Anoetus vitzthumi (Scheucher)
Histiostoma vitzthumi Scheucher, 1957: 313
Anoetus vitzthumi (Scheucher): Woodring & Moser, 1970: 1256; Sevastianov, 1975: 400
Type locality: Schleswig-Holstein
Distribution: Austria, Germany
Host: Dendroctonus micans (Coleoptera: Curculionidae, Scolitinae)
Anoetus wastis Gad Hamada Hassan et Sevastianov
Anoetus wastis Gad Hamada Hassan & Sevastianov, in Radi & Sevastyanov 1991: 49
Type locality: Egypt
Distribution: Egypt
Host: Labidura riparia (Dermaptera)
Genus Aphodanoetus Bongers, OConnor et Lukoschus
Aphodanoetus Bongers, OConnor & Lukoschus, 1985: 33
36
Acarorum Catalogus IX
Type species: Aphodanoetus teinophallus Bongers, OConnor et Lukoschus, 1985
Aphodanoetus teinophallus Bongers, OConnor et Lukoschus
Aphodanoetus teinophallus Bongers, OConnor & Lukoschus, 1985: 36
Type locality: Nijmegen
Distribution: Holland
Host: Aphodius frater (Coleoptera: Scarabaeidae, Aphodiinae)
Genus Auricanoetus Fain et Zumpt
Auricanoetus Fain & Zumpt, 1974: 99 [as subgenus of Loxanoetus)
Auricanoetus Fain & Zumpt: Fain 1983: 849 [as genus]
Type species: Loxanoetus (Auricanoetus) longitarsus Fain et Zumpt, 1974
Auricanoetus longitarsus (Fain et Zumpt)
Loxanoetus (Auricanoetus) longitarsus Fain et Zumpt, 1974: 99; Domrow & Ladds, 1984:
759
Auricanoetus longitarsus (Fain et Zumpt): Fain 1983: 849
Type locality: Karumu Game Reserve
Distribution: Uganda
Host: Syncerus caffer (Artiodactyla: Bovidae)
Genus Austranoetus Fain
Austranoetus Fain, 1976: 314
Type species: Austranoetus kerguelenensis Fain, 1976
Austranoetus kerguelenensis Fain
Austranoetus kerguelenensis Fain, 1976: 314
Type locality: Kerguelen
Distribution: Kerguelen
Genus Bonomoia Oudemans
Bonomoia Oudemans, 1911: 174
Type species: Bonomoia primitiva Oudemans, 1911
Bonomoia africana Mahunka
Bonomoia africana Mahunka, 1967a: 171
Type locality: Mount Foari Reservation
Distribution: Republic of the Congo
Bonomoia certa Woodring et Moser
Bonomoia certa Woodring & Moser, 1970: 1252
Type locality: Moskow Mtn., Moskow, Idaho
Acarorum Catalogus IX 37
Distribution: USA
Hosts: Dendroctonus brevicomis, D. pseudotsugae, Scolytus ventralis (Coleoptera: Curculionidae, Scolytinae)
Bonomoia picturata Sevastianov
Bonomoia picturata Sevastianov, 1973: 81
Type locality: Khmelnitskij Rayon
Distribution: Ukraine
Bonomoia primitiva Oudemans
Bonomoia primitiva Oudemans, 1911: 174
Type locality: Banjoewangi, Java
Distribution: Indonesia (Java)
Host: Blattida fam., gen., sp.
Bonomoia probata Sevastianov
Bonomoia probata Sevastianov, 1973: 82
Type locality: Khmelnitskij Rayon
Distribution: Ukraine
Bonomoia recondita Sevastianov
Bonomoia recondita Sevastianov, 1973: 83
Type locality: Mariyskaya ASSR, Orshanskiy Rayon
Distribution: Russia
Host: Myrmica ruginodis (Hymenoptera: Formicidae)
Bonomoia reticulata Mahunka
Bonomoia reticulata Mahunka, 1967d: 162
Type locality: Cuc-puong, Bong
Distribution: Vietnam
Bonomoia rugosa Mahunka
Bonomoia rugosa Mahunka, 1973a: 301
Type locality: Kwandaso
Distribution: Ghana
Bonomoia zoheirii Metwali et Ahmad
Bonomoia zoheirii Metwali & Ahmad, 1987: 85
Type locality: Sharkya Governorate
Distribution: Egypt
Host: ants (Formicidae)
38
Acarorum Catalogus IX
Bonomoia sp.
Hosts: Hybomitra affinis, H. epistates, H. illota, H. lasiophthalmus, H. nuda, Tabanus
quinquestriatus (Canada)(Mullen et al., 1979)
Genus Bothyanoetus Fain et Camerik
Bothyanoetus Fain & Camerik, 1978: 170
Type species: Bothyanoetus pereirai Fain et Camerik, 1978
Bothyanoetus pereirai Fain et Camerik
Bothyanoetus pereirai Fain & Camerik, 1978: 170
Type locality: Cotia, Sao Paulo
Distribution: Brazil
Host: Bothynus deiphobus (Coleoptera: Scarabaeidae)
Genus Cederhjelmia Oudemans
Cederhjelmia Oudemans, 1931: 261
Type species: Cederhjelmia quadriuncinata Oudemans, 1931
Cederhjelmia quadriuncinata Oudemans
Cederhjelmia quadriuncinata Oudemans, 1931: 179
Type locality: South Brazil
Distribution: Brazil
Genus Chiloanoetus Fain
Chiloanoetus Fain, 1974: 63
Type species: Chiloanoetus mayumbensis Fain, 1974
Chiloanoetus mayumbensis Fain
Chiloanoetus mayumbensis Fain, 1974: 65
Type locality: Chiloango, Mayumbe
Distribution: DR Congo
Host: Prioscelis serrata (Coleoptera: Tenebrionidae)
Genus Hauserania Mahunka
Hauserania Mahunka, 1972e: 954
Type species: Hauserania ornata Mahunka, 1972
Hauserania graeca Mahunka
Hauserania graeca Mahunka, 1972e: 956
Type locality: Kephalinia, Sami
Distribution: Greece (Kephalinia)
Acarorum Catalogus IX 39
Hauserania ornata Mahunka
Hauserania ornata Mahunka, 1972e: 954
Type locality: Kephalinia, on the road Livadion - Argostolion
Distribution: Greece (Kephalinia)
Genus Hexanoetus Fain et Philips
Hexanoetus Fain & Philips, 1979: 150
Type species: Hexanoetus conoidalis Fain et Philips, 1979
Hexanoetus conoidalis Fain et Philips
Hexanoetus conoidalis Fain & Philips, 1979: 150
Type locality: Syracuse, N.Y.
Distribution: USA (N.Y.)
Genus Histiostoma Kramer
Histiostoma Kramer, 1876: 105
Zschachia Oudemans, 1929: 451. Type: Zschachia laevia Oudemans, 1929
Type species: Hypopus julorum C.L. Koch, 1843 (= Histiostoma pectineum Kramer, 1876)
Subgenus Histiostoma Kramer
Histiostoma Kramer, 1876: 105
Type species: Histiostoma pectineum Kramer, 1876 (= Hypopus feroniarum Dufour, 1839)
Histiostoma (H.) abietis Scheucher
Histiostoma abietis Scheucher, 1957: 335
Anoetus abietis (Scheucher): Woodring & Moser, 1970: 1255
Type locality: Regensburg
Distribution: Germany
Host: Ips curvidens (Coleoptera: Curculionidae, Scolytinae)
Histiostoma (H.) acrocini Mahunka
Histiostoma acrocini Mahunka, 1982: 83
Type locality: Brazil
Distribution: Brazil
Host: phoretic on Acrocinus longimanus (Coleoptera: Cerambycidae)
Histiostoma (H.) adarcuale Mahunka
Histiostoma adarcuale Mahunka, 1979: 334
Type locality: Puerto Viejo
Distribution: Costa Rica
40
Acarorum Catalogus IX
Histiostoma (H.) adsimile Fain
Histiostoma adsimile Fain, 1974b: 85
Type locality: La Savane de Lamto
Distribution: Ivory Coast
Histiostoma (H.) alphaticum Fain
Histiostoma alphaticum Fain, 1974b: 82
Type locality: La Savane de Lamto
Distribution: Ivory Coast
Histiostoma (H.) alii Eraky
Histiostoma alii Eraky, 2000: 8
Type locality: Sohag
Distribution: Egypt
Histiostoma (H.) amargynus Mahunka
Histiostoma amargynus Mahunka, 1976b: 309
Type locality: Nondugl, 2200–2700 m
Distribution: Papua New Guinea
Host: Amarygmus [as “Amargynus”] cfr. capillatus (Coleoptera: Tenebrionidae)
Histiostoma (H.) americanum Banks
Histiostoma americanum Banks, 1906: 11
Type locality: USA
Distribution: USA
Histiostoma (H.) amieti Mahunka
Histiostoma amieti Mahunka, 1979: 236
Type locality: Cameroon
Distribution: Cameroon
Histiostoma (H.) andrei Mahunka
Histiostoma andrei Mahunka, 1963b: 57
Type locality: Nordeste: forest n. Cassai
Distribution: Angola
Histiostoma (H.) angolensis Mahunka
Histiostoma angolensis Mahunka, 1963a: 27; 1963b: 58
Type locality: Cavuemba River, Alto Cuílo, Poste de Cacolo
Distribution: Angola
Acarorum Catalogus IX 41
Histiostoma (H.) antillense Mahunka
Histiostoma antillense Mahunka, 1981: 350
Type locality: Castries, Vigie
Distribution: St. Lucia, Windward Islands
Histiostoma arcuata Mahunka
Histiostoma arcuata Mahunka, 1963a: 28
Histiostoma arcuatus Mahunka:
Type locality: Lumeje II, N of Santando Station of C.F.B., Poste de Bussaco (80 km E of
Luso)
Distribution: Angola, Egypt
Histiostoma (H.) arenatus Negm
Histiostoma arenatus Negm, 2007; Abdelgayed et al., 2019: 32
Type locality: Egypt
Distribution: Egypt
Histiostoma (H.) australiense Mahunka
Histiostoma australiense Mahunka, 1975: 317; Tagami & Halliday, 2013: 218
Type locality: Australia
Distribution: Australia
Host: Titanolabis colossea (Dermaptera: Anisolabididae)
Histiostoma (H.) bacilligerum Mahunka
Histiostoma bacilligerum Mahunka, 1963a: 30
Type locality: Cazombo (Alto Zambeze)
Distribution: Angola
Histiostoma (H.) bakeri Hughes et Jackson
Histiostoma bakeri Hughes & Jackson, 1958: 45
Type locality: Norris Dam, Tenn.
Distribution: USA (Tenn.)
Histiostoma (H.) baloghi Mahunka
Histiostoma baloghi Mahunka, 1962: 517
Type locality: Bükk – Gebirge, Belko
Distribution: Hungary
Histiostoma (H.) banjuwangicus (Oudemans)
Anoetus banjuwangicus Oudemans, 1911: 172
42
Acarorum Catalogus IX
Histiostoma banjuwangicus (Oudemans): Buitendijk, 1945: 361; Hughes & Jackson, 1958:
49
Type locality: Banjoewangi
Distribution: Indonesia (Java)
Histiostoma (H.) bananai Eraky et Shoker
Histiostoma bananai Eraky & Shoker, 1993: 53
Type locality: Egypt
Distribution: Egypt
Histiostoma (H.) berghii Jensen
Histiostoma berghii Jensen, 1896: 72
Histiostoma berghi (Jensen): Buitendijk, 1945: 361
Type locality: Copenhagen
Distribution: Denmark
Host: in cocoons of Aulastoma gulo (Hirudinea)
Histiostoma (H.) berndhauseri Mahunka
Histiostoma berndhauseri Mahunka, 1992: 396
Type locality: Cascade de Faarumai
Distribution: Tahiti
Host: Diplopoda
Histiostoma (H.) bernoni Mahunka
Histiostoma bernoni Mahunka, 1979: 134
Type locality: Kenya
Distribution: Kenya
Host: Coleoptera (Scarabaeidae)
Histiostoma (H.) bifoveolatum Mahunka
Histiostoma bifoveolatum Mahunka, 1979: 135
Type locality: Poroto Mt.
Distribution: Tanzania
Histiostoma (H.) blomquisti Wirth et Moser
Histiostoma blomquisti Wirth & Moser, 2010: 357
Type locality: Pineille and New Orleans
Distribution: USA
Host: Solenopsis invicta (Hymenoptera: Formicidae)
Histiostoma (H.) brevimanum (Oudemans)
Lipstorpia brevimana Oudemans, 1914: 73
Acarorum Catalogus IX 43
Anoetus brevimanus: Oudemans, 1917: 392
Histiostoma brevimanus (Oudemans): Buitendijk, 1945: 361; Hughes & Jackson, 1958: 52;
Bongers, OConnor & Lukoschus, 1985: 6
Type locality: Sittard
Distribution: Holland
Histiostoma (H.) brevipodum Berlese
Histiostoma brevipodum Berlese, 1910: 270
Type locality: Palermo
Distribution: Sicily
Habitat: “inter folia putria”
Histiostoma (H.) brevipes Banks
Histiostoma brevipes Banks, 1906: 11
Type locality: USA
Distribution: USA
Histiostoma (H.) calicifera Mahunka
Histiostoma calicifera Mahunka, 1974b: 375; Eraky 2000: 15
Type locality: Banda-Nkwanta
Distribution: Ghana
Histiostoma (H.) camerunensis Mahunka
Histiostoma camerunensis Mahunka, 1979: 238
Type locality: Cameroon
Distribution: Cameroon
Histiostoma (H.) campanula (Oudemans)
Anoetus campanula Oudemans, 1914: 71
Histiostoma campanulae (Oudemans): Buitendijk, 1945: 361; Hughes & Jackson, 1958: 54
Type locality: Banjoewangi
Distribution: Indonesia (Java)
Histiostoma (H.) camphori Eraky
Histiostoma camphori Eraky, 1999: 72; Abdelgayed et al., 2019: 32
Type locality: Upper Egypt (Assiut)
Distribution: Egypt
Histiostoma (H.) capillatum Mahunka
Histiostoma capillata Mahunka, 1978: 143
Histiostoma capillatum Mahunka: Klimov & Khaustov, 2018
Type locality: Oriente, 00o24S, 76o36’W, Limoncocha
44
Acarorum Catalogus IX
Distribution: Ecuador
Host: Labidus praedator (Hymenoptera: Formicidae, Dorylinae)
Histiostoma (H.) caro Ashfaq, Chaudhri et Aheer
Histiostoma caro Ashfaq, Chaudhri & Aheer, 1985: 42
Type locality: Pakistan
Distribution: Pakistan
Histiostoma (H.) cataglyphi Yousef et Metwally
Histiostoma cataglyphi Yousef & Metwally, 1974: 724
Type locality: Zagazig. Sharkia governorate
Distribution: Egypt
Histiostoma (H.) chilopophila Mahunka
Histiostoma chilopophila Mahunka, 1979: 241
Type locality: Cameroon
Distribution: Cameroon
Histiostoma (H.) cirratum (Oudemans)
Anoetus cirratus Oudemans, 1911: 173
Histiostoma cirratus (Oudemans): Buitendijk, 1945: 361
Type locality: Banjoewangi
Distribution: Indonesia (Java)
Host: Blattodea fam., gen. sp.
Histiostoma (H.) clavatum Mahunka
Histiostoma clavatum Mahunka, 1963b: 58
Type locality: Dundo
Distribution: Angola
Host: Brachytripes membranaceus (Orthoptera: Grylloidea)
Histiostoma (H.) clavifer Mahunka
Histiostoma clavifer Mahunka, 1978a: 146
Type locality: 4 km W of Villa Neily, 8o40’N, 82o56’W
Distribution: Costa Rica
Host: Eciton hamatum (Hymenoptera: Formicidae, Dorylinae)
Histiostoma (H.) conclavicola (Oudemans)
Anoetus conclavicola Oudemans, 1929: 461
Histiostoma (H.) conclavicola (Oud.)
Type locality: Stanleyville (Kisangani)
Acarorum Catalogus IX 45
Distribution: DR Congo
Histiostoma (H.) conjunctum (Woodring et Moser)
Anoetus conjuncta Woodring & Moser, 1970: 1250
Histiostoma conjuncta (Woodring et Moser): Woodring & Moser, 1975: 83
Histiostoma conjunctum (Woodring et Moser): Klimov & Khaustov, 2018: 2412
Type locality: Tegusigalpa, Honduras
Distribution:Guatemala, Honduras, USA
Hosts: galleries of Dendroctonus frontalis, Ips avulsus, I. calligraphus, I. cribricollis, I.
grandicollis
Histiostoma (H.) continuatum Mahunka
Histiostoma continuata Mahunka, 1978a: 146
Type locality: Oriente, 00o24S, 76o36’W, Limoncocha
Distribution: Ecuador
Host: Labidus sp. (Hymenoptera: Formicidae, Dorylinae)
Histiostoma (H.) coprae Chinniah et Mohanasundaram
Histiostoma coprae Chinniah & Mohanasundaram, 1995: 73
Type locality: Tamil Nadu
Distribution: India
Host: Copris (Coleoptera)
Histiostoma (H.) coronatum Mahunka
Histiostoma coronatum Mahunka, 1979: 335
Type locality: Puerto Viejo
Distribution: Costa Rica
Histiostoma (H.) cossi Scheucher
Histiostoma cossi Scheucher, 1957: 329
Type locality: Erlangen
Distribution: Germany
Hosts: Cossus cossus, C. ligniperda (Lepidoptera: Cossidae)
Histiostoma (H.) crantor Ashfaq, Chaudhri et Aheer
Histiostoma crantor Ashfaq, Chaudhri & Aheer, 1985: 22
Type locality: Pakistan
Distribution: Pakistan
Histiostoma (H.) crassipes (Oudemans)
Anoetus crassipes Oudemans, 1912: 234; ? Vitzthum, 1923
46
Acarorum Catalogus IX
Histiostoma crassipes (Oudemans): Klimov & Khaustov, 2018: 2413
Type locality: Tanzania
Distribution: Tanzania, (?) Nepal
Hosts: Platypus dispar, Ips stebbingi (Coleoptera: Curculionidae), Hololepta africanae
(Col.: Histeridae)
Histiostoma (H.) crypturgi Scheucher
Histiostoma crypturgi Scheucher , 1957: 310; Klimov & Khaustov, 2018: 2413
Anoetus crypturgi (Scheucher): Woodring & Moser, 1970: 1255; Sevastianov, 1975: 400
Type locality: Erlangen and Regensburg
Distribution: Germany, Poland, Slovakia
Hosts: Crypturgus pusillus (Coleoptera: Curculionidae, Scolytinae), Rhagium sp. (Cerambycidae), Aphodius prodromus (Scarabaeidae)
Histiostoma (H.) cubanum Mahunka
Histiostoma cubanum Mahunka, 1978: 111
Type locality: La Habana
Distribution: Cuba
Host: Acanthoderes circumflexa (Coleoptera: Cerambycidae)
Histiostoma (H.) cyrtandrae (Vitzthum)
Anoetus cyrtandrae Vitzthum, 1931: 110
Histiostoma cyrtandrae (Vitzthum)
Type locality: at Ngebel Lake, Middle Java
Distribution: Indonesia
Histiostoma (H.) dactyloscopicum Mahunka et Eraky
Histiostoma dactyloscopicum Mahunka & Eraky, 1987: 130
Type locality: Hungary
Distribution: Hungary
Histiostoma (H.) darwishii Eraky
Histiostoma darwishii Eraky, 1994: 220; Abdelgayed et al., 2019: 32
Type locality: Egypt
Distribution: Egypt
Histiostoma (H.) decemvirgae Jacot
Histiostoma decemvirgae Jacot, 1939: 199
Type locality: Western North Carolina
Distribution: USA
Acarorum Catalogus IX 47
Histiostoma (H.) densus Ashfaq, Chaudhri et Aheer
Histiostoma densus Ashfaq, Chaudhri & Aheer, 1985 18
Type locality: Pakistan
Distribution: Pakistan
Histiostoma (H.) diadematum Mahunka
Histiostoma diademata Mahunka, 1978a: 148
Type locality: Oriente, 00o24S, 76o36’W, Limoncocha
Distribution: Ecuador
Host: Neivamyrmex gibbatus (Hymenoptera: Formicidae, Dorylinae)
Histiostoma (H.) disiunctum Mahunka
Histiostoma disiuncta Mahunka, 1962b: 428
Anoetus disiunctus
Type locality: Balatonfenyves
Distribution: Hungary
Host: Demetrias imperialis (Coleoptera: Carabidae)
Histiostoma (H.) distentosetum Mahunka
Histiostoma distentoseta Mahunka, 1974b: 375
Type locality: Banda-Nkwanta
Distribution: Ghana
Histiostoma (H.) dryocoeti Scheucher
Histiostoma dryocoeti Scheucher, 1957: 315
Anoetus dryocoeti (Scheucher): Woodring & Moser, 1970: 1255; Sevastianov, 1975: 397
Type locality: “Erlanger Umgebung und Oberpfalz”
Distribution: Germany
Hosts: Dryocoetus autographus (Coleoptera: Curculionidae, Scolytinae), Dorcus parallelopipedus (Coleoptera: Lucanidae)
Histiostoma (H.) dugesi (Claparède)
Hypopus dugesii Claparède, 1868: 506
Histiostoma dugesi (Clap.): Buitendijk, 1945: 361
Type locality: Giseh
Distribution: Egypt
Histiostoma (H.) duplex Mahunka
Histiostoma duplex Mahunka, 1979: 337
Type locality: Puerto Viejo
Distribution: Costa Rica
48
Acarorum Catalogus IX
Histiostoma (H.) edax Ashfaq, Chaudhri et Aheer
Histiostoma edax Ashfaq, Chaudhri & Aheer, 1985: 33
Type locality: Pakistan
Distribution: Pakistan
Histiostoma (H.) editum Mahunka
Histiostoma edita Mahunka, 1974b: 378
Type locality: Kwandaso
Distribution: Ghana
Histiostoma (H.) egyptiacum Eraky et Shoker
Histiostoma egyptiacum Eraky & Shoker, 1993: 51
Type locality: Egypt
Distribution: Egypt
Histiostoma (H.) eickworti Woodring
Histiostoma eickworti Woodring, 1973: 316
Type locality: Costa Rica
Distribution: Costa Rica
Host: Augochlora spp. (Hymenoptera: Halictidae)
Histiostoma (H.) elzingai Mahunka
Histiostoma elzingai Mahunka, 1978a: 148
Type locality: Oriente, 00o24S, 76o36’W, Limoncocha
Distribution: Ecuador
Host: Eciton lucanoides (Hymenoptera: Formicidae, Dorylinae)
Histiostoma (H.) endroedii (Mahunka)
Anoetus endroedii Mahunka, 1973a: 90
Histiostoma endroedii: Mahunka, 1979: 128
Type locality: Abofour (Ghana)
Distribution: Ghana, Gambia
Host: Coleoptera
Histiostoma (H.) ensifer (Oudemans)
Anoetus ensifer Oudemans, 1912: 236
Histiostoma ensifer (Oudemans): Buitendijk, 1945: 361
Type locality: Tanganyika, Amani
Distribution: Tanzania
Histiostoma (H.) erugo Ashfaq, Chaudhri et Aheer
Histiostoma erugo Ashfaq, Chaudhri & Aheer, 1985: 27
Acarorum Catalogus IX 49
Type locality: Pakistan
Distribution: Pakistan
Histiostoma (H.) farghali Eraky
Histiostoma farghali Eraky, 2000: 13
Type locality: Egypt
Distribution: Egypt
Histiostoma (H.) feroniarum (Dufour)
Hypopus feroniarum Dufour, 1839: 278
Histiostoma pectineum Kramer, 1876
Acarus mamillaris Canestrini, 1878: 140
Tyroglyphus rostro-serratus Mégnin, 1873: 372
Rhyzoglyphus rostro-serratus:
Histiostoma rostro-serratum: Michael, 1901:
Anoetus feroniarum (Dufour): Mahunka, 1971: 1195; 1972e: 951
Histiostoma feroniarum (Dufour): Scheucher, 1957: 290; Costa, 1967: 107; Caceres & Fain,
1975: 146; Chen et al., 1982; Shen et al. 1993; Qin & Rohitha 1996: 65; Fain, 1991:
79; Rebollero & Arroyo, 1996; Sanchez-Covisa et al., 1999
Type locality: France
Distribution: Australia, Austria, China, France, Germany, Greece (Kephallinia, Zakynthos), Israel, Italy, New Zealand, Peru, Spain, Canary Is. (Tenerife)
Hosts: Cormocephalus aurantiipes (Chilopoda: Scolopendridae), Hister merdarius, H.
stercorarius, H. unicolor (Coleoptera: Histeridae); Spalax ehrenbergi (Mammalia:
Rodentia)
Histiostoma (H.) flabelliforme Leonardi
Histiostoma flabelliforme Leonardi, 1899: 922
Type locality: Portici
Distribution: Italy
Histiostoma (H.) flaccus Sarwar
Histiostoma flaccus Sarwar, 1998: 2
Type locality: Pakistan
Distribution: Pakistan
Histiostoma (H.) flagellifer Leonardi
Histiostoma flagellifer Leonardi, 1897
Type locality: Italy
Distribution: Italy
50
Acarorum Catalogus IX
Histiostoma (H.) formosanum Philipsen et Coppel
Histiostoma formosana Philipsen & Coppel, 1977: 497; Wang et al., 2002
Type locality: Taiwan
Distribution: Taiwan, China
Host: Coptotermes formosanus (Isoptera: Rhinotermitidae)
Histiostoma (H.) formicarum Vitzthum
Histiostoma formicarum Vitzthum, 1941: 315
Type locality: Tirol
Distribution: Austria
Host: Myrmica sulcinodis (Hymenoptera: Formicidae)
Histiostoma (H.) fortis Ashfaq, Sarwar et Amiad
Histiostoma fortis Ashfaq, Sarwar & Amiad, 2000: 37
Type locality: Rawalakot
Distribution: Pakistan
Host: from decomposing vegetable matter
Histiostoma (H.) foveosimile Mahunka
Histiostoma foveosimile Mahunka, 1975: 320
Type locality: Kiunga
Distribution: Papua New Guinea
Histiostoma (H.) fragosus Ashfaq, Chaudhri et Aheer
Histiostoma fragosus Ashfaq, Chaudhri & Aheer, 1985: 30
Type locality: Pakistan
Distribution: Pakistan
Histiostoma (H.) fucatifer Mahunka
Histiostoma fucatifer Mahunka, 1975b: 47
Type locality: Gambia
Distribution: Gambia
Host: Ploceus-nest
Histiostoma (H.) fuliginosi Scheucher
Histiostoma fuliginosi Scheucher, 1957: 336
Type locality: Erlangen
Distribution: Germany
Histiostoma (H.) furcata Mahunka
Histiostoma furcata Mahunka, 1975b: 50
Type locality: Ghana
Acarorum Catalogus IX 51
Distribution: Ghana
Histiostoma (H.) galerus Sarwar, Ashfaq et Islam,
Histiostoma galerus Sarwar, Ashfaq & Islam, 1998: 4
Type locality: Pakistan
Distribution: Pakistan
Histiostoma (H.) gambiense Mahunka
Histiostoma gambiense Mahunka, 1975b: 53
Type locality: Gambia
Distribution: Gambia
Histiostoma (H.) gampsoceri Mahunka
Histiostoma gampsoceri Mahunka, 1976b: 308
Type locality: “Formosa” (Taiwan)
Distribution: Taiwan
Host: Gampsocera mutata (Diptera: Chloropidae)
Histiostoma (H.) genavensium Mahunka
Histiostoma genavensium Mahunka, 1992: 396
Type locality: Viti Levu: entre Yanuca et Nadi
Distribution: Fiji
Host: Diplopoda
Histiostoma (H.) gervaisi Oudemans
Histiostoma gervaisi Oudemans, 1937: 2145 – nomen novum pro Hypopus ovalis Gervais,
1844
Type locality: Paris
Distribution: France
Host: Lithobius forficatus (Chilopoda: Lithobiidae)
Histiostoma (H.) gladiger (Vitzthum)
Anoetus gladiger Vitzthum, 1926a: 489; Woodring & Moser, 1970: 1255
Histiostoma gladiger Scheucher, 1957: 334; Cooreman, 1963: 39
Type locality: “Nikolajewsk an der Mundung des Amur”
Distribution: Austria, Belgium, Bulgaria, France (incl. Corsica), Germany, Russia (Far East)
Hosts: Ips cembrae, I. sexdentatus, I. subelongatus (Coleoptera: Curculionidae, Scolytinae)
Histiostoma (H.) globotuberum Takami et Kuwahara
Histiostoma globotuberum Takami & Kuwahara, 2005: 86
Type locality: Kyoto and Tsukuba
Distribution: Japan
52
Acarorum Catalogus IX
Histiostoma (H.) gordius (Vitzthum)
Anoetus phyllotrichus Vitzthum, 1921a:
Anoetus gordius Vitzthum, 1923a: 164; Mahunka, 1970: 27; Woodring & Moser, 1970:
1255; Sevastianov, 1975: 400
Anoetus julorum Vitzthum, 1926:
Histiostoma gordius (Vitzthum): Scheucher, 1957: 311; Cooreman, 1963: 37; Hunter &
Davis, 1963: 5;
Distribution: Austria, Belgium, Bosnia & Herzegovina, Germany, Hungary, Romania
Hosts: Ips laricis (Ipidae), Orthomicus [Ips] laricis, O. proximus (Coleoptera: Curculionidae, Scolytinae)
Histiostoma (H.) gorgo Mahunka
Histiostoma gorgo Mahunka, 1974b: 380
Type locality: Banda-Nkwanta
Distribution: Ghana
Histiostoma (H.) gracilipes Banks
Histiostoma gracilipes Banks, 1906: 10
Type locality: USA
Distribution: USA
Histiostoma (H.) gracilis Ashfaq, Sarwar et Amiad
Histiostoma gracilis Ashfaq, Sarwar & Amiad, 2000: 34
Type locality: Haripur
Distribution: Pakistan
Host: from gram (Cicer arietinum)
Histiostoma (H.) granulatum Vitzthum
Histiostoma granulatum Vitzthum, 1935: 93
Type locality: Hivaoa: Kaava Mt.
Distribution: Marquesan Islands
Host: Brachypeplus sp. (Coleoptera: Nitidulidae)
Histiostoma (H.) guanophilum Mahunka
Histiostoma guanophilum Mahunka, 1982: 293
Type locality: New Mexico
Distribution: USA
Histiostoma (H.) halicticolum Fain et Erteld
Histiostoma halicticola Fain & Erteld, 1998: 48
Histiostoma halicticolum Fain & Erteld:
Type locality: Berlin, Doberitzer Heide natural reserve
Acarorum Catalogus IX 53
Distribution: Germany
Host: Halictus sexcinctus (Hymenoptera: Halictidae)
Histiostoma (H.) halictonida Woodring
Histiostoma halictonida Woodring, 1973: 312; Engel & Fain, 2003: 649
Anoetus halictonida: Achtymichuk, 2017
Type locality: USA
Distribution: Canada, Costa Rica, Panama, USA
Hosts: Halictus rubicundus, Megalopta spp. (Hymenoptera: Halictidae)
Histiostoma (H.) heinemanni Hill et Deahl
Histiostoma heinemanni Hill & Deahl, 1978: 318
Type locality: Pennsylvania
Distribution: USA
Histiostoma (H.) herbali Eraky et al.
Anoetus herbali Eraky et al., 2017: 183
Type locality: Assiut, Abu Teeg
Distribution: Egypt
Host: from Citrus orchards
Histiostoma (H.) heriades Fain et Pauly
Histiostoma heriades Fain & Pauly, 2001: 136
Type locality: Sainte Marie Island, Kalalao Forest
Distribution: Madagascar
Host: Heriades erythrosoma (Hymenoptera: Megachilidae)
Histiostoma (H.) himalayae (Vitzthum)
Anoetus himalayae Vitzthum, 1923: 162
Histiostoma himalayae (Vitzthum): Fain, Athias-Binche & Carton, 1993: 73;
Type locality: Himalaya
Distribution: Himalaya
Host: Polygraphus minor (Coleoptera: Curculionidae, Scolytinae [Ipidae])
Histiostoma (H.) histeri Mahunka
Histiostoma histeri Mahunka, 1976: 97
Type locality: Bükk-Gebirge, Czalajka völgy
Distribution: Hungary, Romania
Host: Platysoma compressum (Coleoptera: Histeridae)
Histiostoma (H.) horridum Berlese
Histiostoma horridum Berlese, 1913: 104
54
Acarorum Catalogus IX
Type locality: Java
Distribution: Indonesia (Java)
Histiostoma (H.) huettleri Mahunka
Histiostoma huettleri Mahunka, 1975: 45
Type locality: Gambia
Distribution: Gambia
Histiostoma (H.) humiditatis (Vitzthum)
Anoetus humiditatis Vitzthum, 1927: 98
Histiostoma humiditatis (Vitzthum): Womersley, 1942; Hughes & Jackson, 1958: 78;
Tseng, 1984
Type locality: Thaya, Lower Austria
Distribution: Australia, Austria, Taiwan
Host: Titanolabis colossea (Dermaptera: Anisolabididae)
Histiostoma (H.) hungaricum Mahunka
Histiostoma hungarica Mahunka, 1976: 100
Histiostoma hungaricum Mahunka: Klimov & Khaustov, 2018
Type locality: Slofok
Distribution: Hungary
Host: Carcinopus 14-striata (Coleoptera: Histeridae)
Histiostoma (H.) imbecillus Ashfaq, Chaudhri et Aheer
Histiostoma imbecillus Ashfaq, Chaudhri & Aheer, 1985: 39
Type locality: Pakistan
Distribution: Pakistan
Histiostoma (H.) incurvatum Mahunka
Histiostoma incurvata Mahunka, 1978: 152
Type locality: Canal Zone, Barro Colorado
Distribution: Panama
Host: Eciton burchelli (Hymenoptera: Formicidae, Dorylinae)
Histiostoma (H.) indetonsum (Sevastianov)
Anoetus indetonsus Sevastianov, 1973: 852
Histiostoma indetonsum: Klimov & Khaustov, 2018:
Type locality: Odesa
Distribution: Ukraine
Histiostoma (H.) inquilinum Cross
Histiostoma inquilinus Cross, 1968: 333
Acarorum Catalogus IX 55
Type locality: Maryland
Distribution: USA
Host: Xenoglossa strenua (Hymenoptera: Apidae)
Histiostoma (H.) insulare (Oudemans)
Anoetus insularis Oudemans, 1914: 70; Mahunka, 1967: 15; 1970: 23; 1971: 1198
Histiostoma insulare: Scheucher, 1957: 292
Histiostoma insularis (Oudemans): Hughes & Jackson, 1958: 80
Type locality: Kephallinia
Distribution: Germany, Greece (Kephalinia), Holland, Hungary, Mongolia
Histiostoma (H.) ionicum Mahunka
Histiostoma ionicum Mahunka, 1977: 123
Type locality: Greece
Distribution: Greece
Histiostoma irinum (Hassan et Sevastianov)
Anoetus irinus Hassan & Sevastianov, 1991: 45
Histiostoma irinum (Hassan et Sevastianov): Klimov & Khaustov, 2018:
Type locality: Low Egypt
Distribution: Egypt
Histiostoma (H.) janosi Mahunka
Histiostoma janosi Mahunka, 1975: 319
Type locality: New Guinea
Distribution: New Guinea
Histiostoma (H.) julorum (C.L. Koch)
Hypopus julorum C.L.Koch, 1843: H. 38, No20; Berlese, 1886: AMC, 29, 4
Phyllostoma pectineum Kramer, 1876: 36
Histiostoma pectineum Kramer, 1876: 105
Anoetus julorum (C.L. Koch): Vitzthum, 1926a: 480; Sevastianov, 1975: 398
Histiostoma julorum (C.L. Koch): Canestrini, 1888: 17; Canestrini & Kramer, 1899: 135;
Haarløv, 1957:32
Type locality: Germany
Distribution: Austria, Denmark, Germany, Italy
Hosts: Julus mediterraneus, J. unilineatus (Diplopoda: Julida, Julidae)
Histiostoma (H.) junctus Ashfaq, Chaudhri et Aheer
Histiostoma junctus Ashfaq, Chaudhri & Aheer, 1985: 25
Type locality: Pakistan
56
Acarorum Catalogus IX
Distribution: Pakistan
Histiostoma (H.) karachiensis Khan
Histiostoma karachiensis Khan, 1969: 207
Type locality: Karachi
Distribution: Pakistan
Histiostoma (H.) kerguelenense Fain
Histiostoma kerguelenense Fain, 1976: 312
Type locality: Kerguelen Island
Distribution: Kerguelen Island
Histiostoma laevis (Oudemans)
Zschachia laevis Oudemans, 1929: 451
Type locality: Valkeveen
Distribution: Holland
Histiostoma (H.) lekiensis Mahunka
Histiostoma lekiensis Mahunka, 1979: 241
Type locality: Cameroon
Distribution: Cameroon
Histiostoma (H.) lineatula Mahunka
Histiostoma lineatula Mahunka, 1976: 101
Type locality: Sopron (Hungary)
Distribution: Hungary, Romania
Host: Platysoma compressum (Coleoptera: Histeridae)
Histiostoma (H.) lineosculpturatum Mahunka
Histiostoma lineosculpturatum Mahunka, 1978: 121
Type locality: New Britain, Gazelle Pen., Mt. Sinewit
Distribution: Papua New Guinea [New Britain]
Host: Platolenes sp. (Coleoptera: Tenebrionidae)
Histiostoma (H.) litorale (Oudemans)
Anoetus litoralis Oudemans, 1914: 71; 1914 : 116
Histiostoma litorale (Oudemans): Scheucher, 1957: 333; Mahunka, 1976b: 294; Lotfollahi
et al., 2010
Type locality: Holland (Terschelling I.)
Distribution: Germany, Holland, Iran
Host: Diplopoda, Coleoptera (Carabidae)
Acarorum Catalogus IX 57
Histiostoma (H.) loebli Mahunka
Histiostoma loebli Mahunka, 1979: 423
Type locality: Israel
Distribution: Israel
Histiostoma (H.) loksai Mahunka
Histiostoma loksai Mahunka, 1961: 14
Type locality: Hungary
Distribution: Hungary
Histiostoma longisetum (Mahunka)
Anoetus longisetus Mahunka, 1967: 167
Histiostoma longisetum: Klimov & Khaustov, 2018:
Type locality: Congo
Distribution: Congo
Histiostoma (H.) lorentzi (Oudemans)
Anoetus lorentzi Oudemans, 1905a: 223
Histiostoma lorentzi (Oudemans): Buitendijk, 1945: 362
Type locality: Perboeangan
Distribution: Indonesia (Western New Guinea)
Host: Coleoptera: Coccinellidae
Histiostoma (H.) louisianae Woodring
Histiostoma louisianae Woodring, 1963: 49
Type locality: Louisiana
Distribution: USA
Histiostoma (H.) luxus Ashfaq, Chaudhri et Aheer
Histiostoma luxus Ashfaq, Chaudhri & Aheer, 1985: 36
Type locality: Pakistan
Distribution: Pakistan
Histiostoma (H.) magdolnae Mahuna et Eraky
Histiostoma magdolnae Mahunka & Eraky, 1987: 130
Type locality: Hungary
Distribution: Hungary
Histiostoma (H.) mahunkai Fain
Histiostoma mahunkai Fain, 1974b: 80; Tagami, 2013; 2016: 19
Type locality: Ivory Coast, la Savane de Lamto
58
Acarorum Catalogus IX
Distribution: Ivory Coast, Japan
Host: Anisolabella (Gonolabis) marginalis (Dermaptera: Anisolabididae)
Histiostoma (H.) manicae Samšinák
Histiostoma manicae Samšiṅák, 1958: 291
Type locality: Radhošt (Beskyden)
Distribution: Czech Rep.
Host: Manica rubida (Hymenoptera: Formicidae)
Histiostoma (H.) mannai Eraky
Histiostoma mannai Eraky, 1999: 53
Type locality: Aswan
Distribution: Egypt
Host: Associated with Amitermes desertorum (Insecta)
Histiostoma (H.) mantelli Clark
Histiostoma mantelli Clark, 2010: 10
Type locality: Tongaporutu, North Taranaki
Distribution: New Zealand
Host: Eudyptula minor (Aves: Spheniscidae)
Histiostoma (H.) manuri Eraky
Histiostoma manuri Eraky, 2000: 10
Type locality: Egypt
Distribution: Egypt
Histiostoma (H.) margaritata Mahunka
Histiostoma margaritata Mahunka, 1976: 102
Histiostoma margaritatum Mahunka: Klimov & Khaustov, 2018
Type locality: Mecsek – Gebirge (Hungary)
Distribution: Hungary, Romania
Host: Platysoma compressum (Coleoptera: Histeridae)
Histiostoma (H.) marianae Mahunka
Histiostoma marianae Mahunka, 1979: 338
Type locality: Osa Pen., 2.5 mi SW of Rincon
Distribution: Costa Rica
Histiostoma (H.) maritimum (Oudemans)
Anoetus maritimus Oudemans, 1914: 70; 1914: 110; Mahunka, 1970: 24
Histiostoma maritimum (Oudemans): Scheucher, 1957: 296; Mahunka, 1976b: 326
Type locality: Holland (Terschelling I.)
Acarorum Catalogus IX 59
Distribution: Germany, Holland , Hungary
Hosts: Julus unilineatus (Diplopoda: Julidae), Bembidium assimile, B. doris, B. tenellum,
Elaphrus riparius, Platynus obscurus Coleoptera: Carabidae), Heterocerus marginatus
(Col.: Heteroceridae), Hydrophilus caraboides (Col.: Hydrophilidae)
Histiostoma (H.) matsakisi Mahunka
Histiostoma matsakisi Mahunka, 1977: 125
Type locality: Greece
Distribution: Greece
Histiostoma medium (Woodring et Moser)
Anoetus media Woodring & Moser, 1970: 1238
Histiostoma medium Klimov & Khaustov, 2018:
Type locality: Pineville, Louisiana
Distribution: USA (Louisiana, Florida)
Host: galleries of Dendroctonus terebrans (Col.: Curculionidae, Scolytinae)
Histiostoma (H.) melolonthae Scheucher
Histiostoma melolonthae Scheucher, 1957: 335
Type locality: Erlangen
Distribution: Germany
Hosts: Melolontha hippocastani, Polyphylla fullo (Coleoptera: Scarabaeidae, Melolonthinae)
Histiostoma (H.) meszarosorum Mahunka
Histiostoma meszarosorum Mahunka, 1978: 113
Type locality: La Habana
Distribution: Cuba
Host: Acanthoderes circumflexa (Coleoptera: Cerambycidae)
Histiostoma (H.) metwallii Fawzy et Mtewallii
Histiostoma metwalliii Fawzy & Mtewallii, 2007: 125 (also metwllii)
Type locality: Kafr El-Sheikh
Distribution: Egypt
Histiostoma (H.) minimum Mahunka
Histiostoma minimum Mahunka, 1963a: 33
Type locality: Cazombo (Alto Zambeze)
Distribution: Angola
Histiostoma (H.) mirabile Mahunka
Histiostoma mirabile Mahunka, 1981a: 353
Type locality: Micoud, Troumasée Estate
60
Acarorum Catalogus IX
Distribution: Windward-Islands
Histiostoma (H.) montanum Clark
Histiostoma montanum Clark, 2010b: 193
Type locality: North Egmont, Mt Egmont, Taranaki
Distribution: New Zealand
Host: from shrub Brachyglottis elaeagnifolia litter at 1100–1300 m
Histiostoma (H.) moralesi Mahunka
Histiostoma moralesi Mahunka, 1978a: 152
Type locality: 4 km W of Villa Neily, 8o40’N, 82o56’W
Distribution: Costa Rica
Host: Eciton hamatum (Hymenoptera: Formicidae, Dorylinae)
Histiostoma (H.) murchiei Hughes et Jackson
Histiostoma murchiei Hughes & Jackson, 1958: 88
Type locality: Carp River, Emmet County, Michigan
Distribution: USA
Host: in cocoons of Allolobophora (Lumbricidae)
Histiostoma (H.) muscae Chinniah et Mohanasundaram
Histiostoma muscae Chinniah & Mohanasundaram, 1995: 74
Type locality: Tamil Nadu
Distribution: India
Host: Musca domestica (Diptera: Muscidae)
Histiostoma (H.) nasseri Eraky
Histiostoma nasseri Eraky, 1994: 159; Abdelgayed et al., 2019: 32
Type locality: Egypt
Distribution: Egypt
Histiostoma (H.) necopinus (Sevastianov)
Anoetus necopinus Sevastianov, 1973: 852
Histiostoma necopinus (Sevastianov): Mahunka, 1976
Type locality: Hajibeyskiy Liman nr. Odessa
Distribution: Ukraine
Host: Galeruca sp. (Coleoptera: Chrysomelidae)
Histiostoma (H.) negmi Eraky
Histiostoma negmi Eraky, 2000: 5
Type locality: Sohag
Distribution: Egypt
Acarorum Catalogus IX 61
Histiostoma (H.) novamsterdamense Fain
Histiostoma novamsterdamense Fain, 1976: 565
Type locality: Nouvelle Amsterdam, Pointe d’Entrecasteaux
Distribution: Nouvelle Amsterdam
Histiostoma onioni Eraky et Shoker
Histiostoma onioni Eraky & Shoker, 1994; Abdelgayed et al., 2019: 32
Type locality: Sohag
Distribution: Egypt
Histiostoma (H.) onthophagae Chinniah et Mohanasundaram
Histiostoma onthophagae Chinniah & Mohanasundaram, 1995: 74
Type locality: Tamil Nadu
Distribution: India
Host: Onthophagus pactolus (Coleoptera: Scarabaeidae)
Histiostoma (H.) orientalis Mahunka
Histiostoma orientalis Mahunka, 1974: 104
Type locality: unknown
Distribution: unknown
Host: Halictus tatrazonianellus (Hymenoptera: Halictidae)
Histiostoma (H.) oudemansi Scheucher
Histiostoma oudemansi Scheucher, 1957: 315
Anoetus oudemansi (Scheucher): Sevastianov, 1975: 400
Type locality: Erlangen
Distribution: Germany
Histiostoma (H.) ovale (J. Müller)
Uropoda ovalis J. Müller, 1860
Histiostoma ovalis (J.Müller): Samšiṅák, 1958: 293; Mahunka, 1977b: 104; Moraza, Fernandez & Jurc, 2013: 597; Vissa et al., 2019
Histiostoma ovale (J. Müller): Klimov & Khaustov, 2018: 53
Type locality: Moravia (Slovakia)
Distribution: Austria, Belgium, Bulgaria, Croatia, Czech Rep., France, Germany, Corsica,
Hungary, Italy, Poland, Portugal, Romania, Slovakia, Slovenia, Ukraine
Hosts: Ips sexdentatus (Coleoptera: Curculionidae - Scolytinae), Platysoma compressum
(Col.: Histeridae)
Histiostoma (H.) ovalis marani Samšiṅák
Histiostoma ovalis f. marani Samšiṅák, 1958: 294
62
Acarorum Catalogus IX
Type locality: Vizzavona
Distribution: Corsica
Host: Ips sexdentatus (Coleoptera: Curculionidae - Scolytinae)
Histiostoma (H.) palustre Wirth
Histiostoma palustre Wirth, 2003: 262
Type locality: nr. Teufelsee, Berlin/Grunewald
Distribution: Germany
Histiostoma (H.) papillatum Halliday et Collins
Histiostoma papillata Halliday & Collins, 2002: 155
Type locality: Victoria, Deakin University, Warrnambool
Distribution: Australia
Host: Maccullochella peelii peelii (Pisces: Percichthyidae)
Histiostoma (H.) pauciputeolum Fain et Philips
Histiostoma pauciputeolum Fain & Philips, 1979: 147
Type locality: Syracuse, N.Y.
Distribution: USA
Host: In the nest of Otus asio (Aves, Strigiformes)
Histiostoma (H.) perruptum Mahunka
Histiostoma perruptum Mahunka, 1979: 339
Type locality: Osa Pen., 2.5 mi SW of Rincon
Distribution: Costa Rica
Histiostoma (H.) phyllophorum (Oudemans)
Anoetus phyllophorus Oudemans, 1905b: 238; 1912: 236
Anoetus phyllophorum Oudemans: Mahunka, 1973a: 99
Histiostoma phyllophorum (Oudemans): Scheucher, 1957: 318
Histiostoma nichollsi Womersley, 1941: 484 - Australia
Histiostoma phyllophorus (Oudemans): Mahunka & Mahunka-Papp: 1991: 286
Type locality: Leiden
Distribution: Australia, Germany, Ghana, Holland, Tanzania
Hosts: Oniscus asellus (Isopoda: Onyscidae), Oxythyrea funesta (Coleoptera: Scarabaeidae), Passalidae
Histiostoma (H.) phytophaga Ewing
Histiostoma phytophaga Ewing, 1911: 41
Type locality: USA
Distribution: USA
Acarorum Catalogus IX 63
Histiostoma (H.) piceae Scheucher
Histiostoma piceae Scheucher, 1957: 304; Beron, 1973; Takov
et al. 2009; Walter & Latonas, 2012
Anoetus piceae (Scheucher): Woodring & Moser, 1970: 1255
Type locality: Erlangen, Nurnberg, Regensburg, Harz
Distribution: Bulgaria, Canada, Croatia, France, Germany, Japan, Slovakia, Sweden, Russia
Host: Ips typographus (Coleoptera: Scolytidae)
Histiostoma (H.) pickaxei Eraky et Shoker
Histiostoma pickaxei Eraky & Shoker, 1995: 23; Negm & Alatawi, 2011:
Type locality: Egypt
Distribution: Egypt, Saudi Arabia
Histiostoma (H.) piloseta Hughes et Jackson
Histiostoma piloseta Hughes & Jackson, 1958: 100
Type locality: Ellerson, Va.
Distribution: USA
Host: from compost
Histiostoma (H.) polaki (Oudemans)
Anoetus polaki Oudemans, 1914: 72
Histiostoma polaki (Oudemans):
Type locality: Tanzania (“Deutsch Oost Afrika”)
Distribution: Spirostreptes sp. (Diplopoda: Spirostreptidae)
Histiostoma (H.) polypori (Oudemans)
Anoetus polypori Oudemans, 1914: 72
Histiostoma polypori (Oudemans): Buitendijk, 1945: 362; Behura, 1950: 1025; 1956; 1957:
51; Scheucher, 1957: 298; Chmielewski, 1984: 87; Lotfollahi et al., 2010; Tagami,
2016: 19
Type locality: Holland (Oosterbreek)
Distribution: Germany, Great Britain, Holland, Hungary, Iran, Poland
Hosts: Eugamasus cornutus (Acari: Gamasidae), Forficula auricularia (Dermaptera:
Forficulidae)
Histiostoma (H.) pontigerum Mahunka
Histiostoma pontigerum Mahunka, 1963b: 60
Type locality: Dundo, nr. Cundueji River
Distribution: Angola
Histiosoma (H.) prodectoris (Vitzthum)
Anoetus prodectoris Vitzthum, 1920: 62
64
Acarorum Catalogus IX
Histiosoma prodectoris (Vitzthum):
Type locality: “Bua-Kraeng in Süd-Celebes und Luzon Philippinen”
Distribution: Indonesia (Sulawesi), the Philippines (Luzon)
Hosts: Prodector fruhstorferi, Ectocemus cinnamomeus (Coleoptera: Brenthidae)
Histiosoma (H.) prophthalmi (Vitzthum)
Anoetus prophthalmi Vitzthum, 1920: 59
Histiosoma prophthalmi (Vitzthum): Hughes & Jackson, 1958: 107
Type locality: “Vorderindien, einschl. Assam”
Distribution: India
Host: Prophthalmus potens (Coleoptera: Brenthidae)
Histiostoma (H.) protuberans Hughes et Jackson
Histiostoma protuberans Hughes & Jackson, 1958: 109
Type locality: 8 mi west of Richmond, Va.
Distribution: USA
Histiostoma (H.) pseudolongipes Mahunka
Histiostoma pseudolongipes Mahunka, 1992: 398
Type locality: Viti Levu: entre Yanuca et Nadi
Distribution: Fiji
Host: Diplopoda
Histiostoma (H.) pulchrum Kramer
Histiostoma pulchrum Kramer, 1886: 259; Scheucher, 1957: 304
Type locality: Germany
Distribution: England, Germany
Histiostoma (H.) punctaticeps Mahunka
Histiostoma punctaticeps Mahunka, 1963b: 61
Type locality: Dundo, nr. Cundueji River
Distribution: Angola
Histiostoma (H.) pyriforme Michael
Histiostoma pyriforme Michael, 1901: 221
Type locality: Great Britain
Distribution: Great Britain, Italy
Histiostoma (H.) radiferum Scheucher
Histiostoma radiferum Scheucher, 1957: 324
Anoetus radiferum (Scheucher): Sevastianov, 1975: 386
Type locality: Erlangen
Acarorum Catalogus IX 65
Distribution: Germany
Histiostoma (H.) rakha Fawzy et Mtewallii
Histiostoma rakha Fawzy & Mtewallii, 2007: 131
Type locality: Kafr El-Sheikh
Distribution: Egypt
Histiostoma (H.) reticulatissimum Mahunka
Histiostoma reticulatissimum Mahunka, 1978: 123
Type locality: NE Wau, Morobe Distr. 1200 m
Distribution: Papua New Guinea
Histiostoma (H.) reticulofoveolatum Mahunka
Histiostoma reticulofoveolatum Mahunka, 1978: 124
Type locality: Karubana, 1500 m
Distribution: Papua New Guinea
Host: Amarygmus [as “Amargynus”] cf. deformipes (Coleoptera: Tenebrionidae)
Histiostoma (H.) rettenmeyerorum Mahunka
Histiostoma rettenmeyerorum Mahunka, 1978a: 156
Type locality: Oriente, 00o24S, 76o36’W, Limoncocha
Distribution: Ecuador
Host: Labidus praedator (Hymenoptera: Formicidae, Dorylinae)
Histiostoma (H.) rimosum Mahunka
Histiostoma rimosa Mahunka, 1978: 156
Type locality: Oriente, 00o24S, 76o36’W, Limoncocha
Distribution: Ecuador
Host: Labidum sp. (Hymenoptera: Formicidae, Dorylinae)
Histiostoma (H.) rizkii Eraky
Histiostoma rizkii Eraky, 1994: 157; Abdelgayed et al., 2019: 32
Type locality: Egypt
Distribution: Egypt
Histiostoma (H.) rostratum Mahunka
Histiostoma rostrata Mahunka, 1978a: 156
Type locality: 4 km W of Villa Neily, 8o40’N, 82o56’W
Distribution: Costa Rica
Host: Eciton hamatum (Hymenoptera: Formicidae, Dorylinae)
66
Acarorum Catalogus IX
Histiostoma (H.) ruehmi Scheucher
Histiostoma ruehmi Scheucher, 1957: 331
Anoetus ruehmi (Scheucher): Sevastianov, 1975: 400
Type locality: Nürnberg
Distribution: Germany
Host: larve of Melanotus rufipes (Coleoptera: Elateridae)
Histiostoma (H.) rugosum Mahunka
Histiostoma rugosa Mahunka, 1974b: 382
Type locality: Kumasi
Distribution: Ghana
Histiostoma (H.) sachsi Scheucher
Histiostoma sachsi Scheucher, 1957: 319
Anoetus sachsi (Scheucher): Sevastianov, 1975: 400
Type locality: Erlangen
Distribution: Germany
Histiostoma (H.) sapromyzarum (Dufour)
Hypopus sapromyzarum Dufour, 1839: 278
Hypopus filicum Dujardin, 1849: 261
Anoetus sapromyzarum: Oudemans, 1914: 107; Vitzthum, 1926a: 488; Mahunka, 1963c:
497; 1972e: 952
Histiostoma sapromyzarum (Dufour): Vizthum, 1932: 700; Theodorides, 1955; Scheucher,
1957: 302; Mahunka, 1963e: 369
Type locality: France
Distribution: Argentina, Australia, Bolivia, Brazil, France, Germany, Greece, Holland,
Hungary, Italy, Philippines, Syria
Hosts: Melanotus rufipes (Coleoptera: Elateridae); Schizotrachelus bakeri (Brenthidae),
Bubas bubalus (Scarabaeidae); Myodes [Clethrionomys] glareolus (Mammalia: Rodentia)
Histiostoma (H.) sarrai Eraky et Shoker
Histiostoma sarrai Eraky & Shoker, 1994: ?
Type locality: Egypt
Distribution: Egypt
Histiostoma (H.) schauenbergi Mahunka
Histiostoma schauenbergi Mahunka, 1978b: 274
Type locality: Indian Ocean Islands
Distribution: Indian Ocean Islands
Acarorum Catalogus IX 67
Histiostoma (H.) schoutedeni Fain et Elsen
Histiostoma schoutedeni Fain & Elsen, 1972: 86
Type locality: Mangai on Kasai River
Distribution: DR Congo
Host: Glossina fuscipes quanzensis (Diptera: Glossinidae)
Histiostoma (H.) scinditum Mahunka
Histiostoma scindita Mahunka, 1978: 160; Eraky, 2000: 11
Type locality: Oriente, 00o24S, 76o36’W, Limoncocha
Distribution: Ecuador
Host: Eciton lucanoides (Hymenoptera: Formicidae, Dorylinae)
Histiostoma (H.) sextoni Hughes et Jackson
Histiostoma sextoni Hughes & Jackson, 1958: 116
Type locality: Chickahominy Swamp, near Ellerson, Va.
Distribution: USA
Host: in duck pen
Histiostoma shiramba Klimov et Khaustov
Histiostoma shiramba Klimov & Khaustov, 2018: 2375
Type locality: Sakhalin Island
Distribution: Russia (Sakhalin, Primorskyi Kray)
Host: in the galleries of Polygraphus proximus (Coleoptera: Curculionidae, Scolytinae)
Histiostoma (H.) simile Mahunka
Histiostoma simile Mahunka, 1963a: 34
Type locality: Cavuemba River, Alto Cuílo, Poste de Cacolo
Distribution: Angola
Histiostoma (H.) siphunculinae Chinniah et Mohanasundaram
Histiostoma siphunculinae Chinniah & Mohanasundaram, 1995: 77
Type locality: Tamil-Nadu
Distribution: India
Host: Siphunculina sp. (Diptera: Chloropidae)
Histiostoma (H.) solimani Eraky
Histiostoma solimani Eraky, 1997: 107
Type locality: Assiut
Distribution: Egypt
Histiostoma (H.) spinitarsus (Oudemans)
Anoetus spinitarsus Oudemans, 1917: 392
68
Acarorum Catalogus IX
Histiostoma spinitarsus (Oudemans): Buitendijk, 1945: 363
Type locality: South America
Distribution: South America
Host: Phanaeus sp. (Coleoptera: Scarabaeidae)
Histiostoma (H.) stammeri Scheucher
Histiostoma stammeri Scheucher, 1957: 321
Anoetus stammeri (Scheucher): Sevastianov, 1975: 398
Type locality: Germany
Distribution: Germany
Hosts: Philonus, Ontolestes (Coleoptera: Staphylinidae)
Histiostoma (H.) strenzkei Scheucher
Histiostoma strenzkei Scheucher, 1957: 326
Type locality: Plön
Distribution: Germany
Histiostoma (H.) striatosimile Mahunka
Histiostoma striatosimile Mahunka, 1975: 48
Type locality: Ghana
Distribution: Ghana
Histiostoma (H.) subsimile Mahunka
Histiostoma subsimile Mahunka, 1963b: 62
Type locality: forest near Mussalomuca River
Distribution: Angola
Host: Col.: Carabidae
Hirstiosoma (H.) sylvestre (Woodring et Moser)
Anoetus gordius of Hughes & Jackson (1958), not A. gordius Vitzthum, 1923
Anoetus sylvestris Woodring & Moser, 1970: 1248
Hirstiosoma sylvestre (Woodring et Moser):
Type locality: (Tirol)( Vitzthum, 1923), USA (Ellerson, Va., Hughes & Jackson,1958)
Distribution: (Austria, Bosna & Herzegovina) (Vitzthum, 1923) USA (Hughes & Jackson, 1958: 70)
Histiostoma (H.) szelenyii Mahunka
Anoetus szelenyii Mahunka, 1974 : 101
Histiostoma (H.) szelenyii Mahunka: Fain & Erteld, 1998: 52
Type locality: Afghanistan
Distribution: Afghanistan
Acarorum Catalogus IX 69
Host: Halictus holtzi (Diptera: Halictidae)
Histiostoma (H.) szentivanyi Mahunka
Histiostoma szentivanyi Mahunka, 1978: 126
Type locality: NE Nondugl, 2200–2700 m
Distribution: Papua New Guinea
Host: Amarygmus [as “Amargymus”] cf. capillatus (Coleoptera: Tenebrionidae)
Histiostoma (H.) tanypus Fain et Camerik
Histiostoma tanypus Fain & Camerik, 1978: 171
Type locality: Cotia, Sao Paulu
Distribution: Brazil
Host: Hymenoptera: Vespidae gen. sp.
Histiostoma (H.) tarsalis Banks (nomen dubium)
Histiostoma tarsalis Banks, 1910: 12
Type locality: USA
Distribution: USA
Histiostoma (H.) telatum Mahunka
Histiostoma telatum Mahunka, 1979: 137
Type locality: Tanzania
Distribution: Tanzania
Host: Coleoptera
Histiostoma (H.) tesselata Jacot
Histiostoma tesselata Jacot, 1939: 199
Type locality: Western North Carolina
Distribution: USA
Histiostoma (H.) timidum Fain
Histiostoma timidum Fain, 1974b: 82
Type locality: La Savane de Lamto
Distribution: Ivory Coast
Histiostoma (H.) tinydorsalis Eraky
Histiostoma tinydorsalis Eraky, 1999: 70
Type locality: Upper Egypt (Assiut)
Distribution: Egypt
Histiostoma (H.) titanolabi Tagami et Halliday
Histiostoma titanolabi Tagami & Halliday, 2013: 222
70
Acarorum Catalogus IX
Type locality: Australia
Distribution: Australia
Host: Titanolabis colossea (Dermaptera: Anisolabididae)
Histiostoma (H.) toxopei (Oudemans)
Anoetus toxopei Oudemans, 1927: 230
Histiostoma toxopei (Oudemans): Buitendijk, 1945: 363
Type locality: Wai Eno – Wai Temoen, Boeroe
Distribution: Indonesia
Histiostoma (H.) trichophorum (Oudemans)
Anoetus trichophorus Oudemans, 1912: 235
Histiostoma trichophorus (Oudemans): Buitendijk, 1945: 363
Histiostoma trichophorum (Oudemans): Scheucher, 1957: 306
Type locality: Mkulumusi
Distribution: Tanzania
Hosts: Helops quisquilius, Hypophloeus bicolor (Coleoptera: Tenebrionidae); Platypus
dispar (Col.: Platypodidae)
Histiostoma (H.) ulmi Scheucher
Histiostoma ulmi Scheucher, 1957: 308;
Anoetus ulmi (Scheucher): Woodring & Moser, 1970: 1256; Sevastianov, 1975: 400
Type locality: Erlangen
Distribution: Germany
Host: Hypophloeus bicolor (Coleoptera: Tenebrionidae)
Histiostoma (H.) umbrosa Mahunka
Histiostoma umbrosa Mahunka, 1975b: 51
Type locality: Gambia
Distribution: Gambia
Histiostoma (H.) unidentatum Fain, Athias-Binche et Carton
Histiostoma unidentatum Fain, Athias-Binche & Carton, 1993: 71
Type locality: Thailand
Distribution: Australia, Thailand
Host: Drosophila bipectinata (Diptera) imported from Thailand
Histiostoma (H.) uvafer Mahunka
Histiostoma uvafer Mahunka, 1974b: 383
Type locality: Kwandaso
Distribution: Ghana
Acarorum Catalogus IX 71
Histiostoma (H.) uxoria Mahunka
Histiostoma uxoria Mahunka, 1974b: 385
Type locality: Tamale
Distribution: Ghana
Histiostoma (H.) validum Banks
Histiostoma valida Banks, 1906: 141
Type locality: Canada
Distribution: Canada
Histiostoma (H.) varium (Woodring et Moser)
Anoetus varia Woodring & Moser, 1970: 1243; Moser & Roton, 1971; Moser, 1976: 811;
Histiostoma varia (Woodring et Moser): Stone & Simpson, 1991;
Type locality: Elizabeth, La., USA
Distribution: Australia, USA, Honduras, Mexico, Canada
Hosts: boring dust of Ips avulsus, I. calligraphus, I. grandicollis, Dendroctonus frontalis
(Coleoptera: Curculionidae, Scolytinae), phoretic on Carcinops sp. (Histeridae) and
Corticeus coynei (Tenebrionidae)
Histiostoma (H.) variabilis Mahunka
Histiostoma variabilis Mahunka, 1979: 244
Type locality: Cameroon
Distribution: Cameroon
Histiostoma (H.) verruca Jacot
Histiostoma verruca Jacot, 1939: 201
Type locality: Western North Carolina
Distribution: USA
Histiostoma (H.) verrucogenicula Jacot
Histiostoma verrucogenicula Jacot, 1939: 200
Type locality: Western North Carolina
Distribution: USA
Hirstiosoma virginianum Woodring et Moser
Hirstiosoma virginiana Woodring & Moser, 1970: 1255
Hirstiosoma virginianum Woodring & Moser:
Type locality: Virginia, Florida, Tennessee
Distribution: USA
Histiostoma (H.) volgini Mahunka
Histiostoma volgini Mahunka, 1979: 138
72
Acarorum Catalogus IX
Type locality: Transvaal, Nylsvley Reservation
Distribution: South Africa
Histiostoma (H.) vulpis (Dujardin)
Hypopus vulpis Dujardin, 1849: 263
Histiostoma vulpis (Dujardin):
Type locality: Transvaal, Nylsvley Reservation
Distribution: South Africa
Histiostoma (H.) watanabei Kurosa
Histiostoma watanabei Kurosa, 2002: 28
Type locality: Japan
Distribution: Japan
Host: Phoretic on Lathrobium (Lathrobium) (Coleoptera: Staphylinidae)
Histiostoma (H.) wauense Mahunka
Histiostoma wauense Mahunka, 1976b: 311
Type locality: Torricelli Mts.
Distribution: Papua New Guinea
Host: Platolenes femoratus (Coleoptera: Tenebrionidae)
Histiostoma (H.) zaheri Fawzy et Mtewallii
Histiostoma zaheri Fawzy & Mtewallii, 2007: 124
Type locality: Kafr El-Sheikh
Distribution: Egypt
Subgenus Cavanoetus Mahunka
Cavanoetus Mahunka, 1982: 290
Type species: Histiostoma pierrestrinatii Mahunka, 1982
Histiostoma (Cavanoetus) pierrestrinatii Mahunka
Histiostoma (Cavanoetus) pierrestrinatii Mahunka, 1982: 290
Type locality: New Mexico
Distribution: USA
Subgenus Ichtanoetus Fain et Belpaire
Ichtanoetus Fain & Belpaire, 1985: 286
Type species: Histiostoma anguillarum Fain et Belpaire, 1985
Histiostoma (Ichtanoetus) anguillarum Fain et Belpaire
Histiostoma (Ichtanoetus) anguillarum Fain & Belpaire, 1985: 287
Acarorum Catalogus IX 73
Type locality: Loire
Distribution: France
Host: from the gills of young eels (Anguilla Anguilla, Anguillidae, Anguilliformes)
Histiostoma (Ichtanoetus) cyrtandrae (Vitzthum)
Anoetus cyrtandrae Vitzthum, 1931: 110
Histiostoma (Ichtanoetus) cyrtandrae (Vitzthum): Fain & Belpaire, 1985: 287
Type locality: Java
Distribution: Indonesia
Host: in the axill fluid of Colocasia indica
Histiostoma (Ichtanoetus) nigrellii Hughes et Jackson
Histiostoma nigrellii Hughes & Jackson, 1958: 95
Histiostoma (Ichtanoetus) nigrellii: Fain & Belpaire, 1985: 287
Type locality: aquarium in New York
Distribution: USA
Host: goldfish tank
Histiostoma (Ichtanoetus) ocellatum Fain et Lambrechts
Histiostoma (Ichtanoetus) ocellatum Fain & Lambrechts, 1987: 95
Type locality: aquarium containing Discus fish, Antwerp
Distribution: Holland
Histiostoma (Ichtanoetus) piscium Fain et Lambrechts
Histiostoma piscium Fain & Lambrechts, 1985: 121; Tagami, 2016:
Histiostoma (Ichtanoetus) piscium: Fain & Belpaire, 1985: 287
Type locality: aquarium in Antwerpen (imported from Thailand)
Distribution: Japan, Thailand
Hosts: Pangasius sutchi (Pisces: Siluriformes, Pangasiidae); Periplaneta fuliginosa (Dermaptera: Blattidae)
Genus Hormosianoetus Fain
Hormosianoetus Fain, 1980: 753
Type species: Hormosianoetus aeschlimanni Fain, 1980
Hormosianoetus aeschlimanni Fain
Hormosianoetus aeschlimanni Fain, 1980: 756; Eraky & Shoker 1993: 56
Type locality: Zurich
Distribution: Switzerland
Host: Drosophila sp. (Diptera: Drosophilidae)
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Acarorum Catalogus IX
Hormosianoetus laboratorium (Hughes)
Histiostoma laboratorium Hughes, 1950: 177; Scheucher, 1957: 333
? Histiostoma genetica (Stolpe, 1938)
Hormosianoetus laboratorium (Hughes): Fain, Athias-Binche & Carton, 1993: 75
Type locality: Switzerland
Distribution: Guadeloupe, Morocco, Switzerland
Hosts: Drosophyla willstoni, D. subobscura (Diptera: Drosophylidae)
Hormosianoetus mahunkai Eraky et Shoker
Hormosianoetus mahunkai Eraky & Shoker, 1993: 53; Eraky & Mohamed, 2019: 2
Type locality: from Sohag governorate
Distribution: Egypt
Host: from Fusarium culture
Hormosianoetus mallotae (Fashing)
Mauduytia mallotae Fashing, 1973: 455
Hormosianoetus mallotae: Fain, 1980: 755
Type locality: Johnson County (Texas)
Distribution: USA (Texas, Pennsylvania, Illinois)
Hosts: Mallota posticata, M. bautius (Diptera: Syrphidae)
Genus Hymenanoetus Mahunka
Hymenanoetus Mahunka, 1963a: 39
Type species: Hymenanoetus nodosus Mahunka, 1963
Hymenanoetus nodosus Mahunka
Hymenanoetus nodosus Mahunka, 1963a: 39
Type locality: Angola, Lumeje II, N of Santando Station of C.F.B., Poste de Bussaco (80
km E of Luso)
Distribution: Angola
Genus Insulanoetus Sevastyanov
Insulanoetus Sevastyanov, 1973: 855
Type species: Insulanoetus brachypodus Sevastyanov, 1973
Insulanoetus brachypodus Sevastyanov
Insulanoetus brachypodus Sevastyanov, 1973: 856
Type locality: Jaralgach Island (Black Sea)
Distribution: Crimea
Hosts: Bledius bicornis, B. sarmaticus (Coleoptera: Staphylinidae, Oxythelinae)
Acarorum Catalogus IX 75
Genus Kanoetus Samšinák
Kanoetus Samšinák, 1966: 126
Type species: Kanoetus cubanus Samšinák, 1966
Kanoetus australis Mahunka
Kanoetus australis Mahunka, 1975a: 60
Type locality: Kpington
Distribution: South Africa
Kanoetus contrasetosus Mahunka
Kanoetus contrasetosus Mahunka, 1974 : 388
Type locality: Banda-Nkwanta
Distribution: Ghana
Kanoetus cubanus Samšinák
Kanoetus cubanus Samšinák, 1966: 126
Type locality: Cuba
Distribution: Cuba
Kanoetus samsinaki Mahunka
Kanoetus samsinaki Mahunka, 1974: 386
Type locality: Kumasi
Distribution: Ghana
Genus Kaszabanoetus Mahunka
Kaszabanoetus Mahunka, 1976b: 317
Type species: Kaszabanoetus ulomae Mahunka, 1976
Kaszabanoetus ulomae Mahunka
Kaszabanoetus ulomae Mahunka, 1976b: 318
Type locality: Akcagoga
Distribution: Turkey
Host: Uloma culinaria (Coleoptera: Tenebrionidae)
Genus Loxanoetus Fain
Loxanoetus Fain, 1970: 173
Type species: Loxanoetus bassoni Fain, 1970 [Redescription, Fain 1983: 847]
Loxanoetus bassoni Fain
Loxanoetus bassoni Fain, 1970: 173; Fain & Zumpt, 1974: 99; Fain 1983: 847
Type locality: Kruger Park (South Africa)
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Acarorum Catalogus IX
Distribution: South Africa, Namibia, Uganda
Hosts: Loxodonta africana (Mammalia, Proboscidea: Elephantidae), Syncerus caffer (Artiodactyla: Bovidae)
Loxanoetus lenae Domrow et Ladds
Loxanoetus lenae Domrow & Ladds, 1984: 759
Type locality: Townsville, Qeensland
Distribution: from zoo in Australia
Host: Elephas maximus (Mammalia, Proboscidea: Elephantidae)
Genus Munduytia Oudemans
Munduytia Oudemans, 1929: 450 [also “Mauduytia” in Scheucher, 1957]
Type species: Anoetus tropicus Oudemans, 1911
Munduytia tropicus (Oudemans)
Anoetus tropicus Oudemans, 1911a: 190
Munduytia tropicus (Oudemans): Oudemans, 1929: 450
Mauduytia tropicus: Scheucher, 1957: 379
Type locality: Java, Banjoewangi
Distribution: Indonesia (Java)
Host: Blattodea fam., gen. sp.
Genus Nepenthacarus Fashing
Nepenthacarus Fashing, 2002: 8
Type species: Nepenthacarus warreni Fashing, 2002
Nepenthacarus warreni Fashing
Nepenthacarus warreni Fashing, 2002: 8
Type locality: Far North Queensland
Distribution: Australia
Genus Otanoetus Fain et Zumpt
Otanoetus Fain & Zumpt, 1974: 100 [placement?]
Type species: Otanoetus wetzeli Fain et Zumpt, 1974
Otanoetus wetzeli Fain et Zumpt
Otanoetus wetzeli Fain & Zumpt, 1974: 100
Type locality: Karumu Game Reserve (Uganda)
Distribution: Namibia, Uganda
Hosts: Loxodontha africana (Proboscidea: Elephantidae), Syncerus caffer (Artiodactyla:
Bovidae)
Acarorum Catalogus IX 77
Genus Ovanoetus Fain et van Goethem
Ovanoetus Fain & van Goethem, 1985: 227
Type species: Ovanoetus wautersi Fain et van Goethem, 1985
Ovanoetus wautersi Fain et van Goethem
Ovanoetus wautersi Fain & van Goethem, 1985: 228
Type locality: St. Vincent Republic, Grenadines Isl., Bequia Id. (Windward Group)
Distribution: St. Vincent Republic, Grenadines Isl., Bequia Id. (Windward Group)
Host: Coenobita clypeata (Crustacea: Paguridae)
Genus Peripatetes Mahunka
Peripatetes Mahunka, 1976b: 315
Type species: Peripatetes acanthotarsus Mahnunka, 1976
Peripatetes acanthotarsus Mahunka
Peripatetes acanthotarsus Mahunka, 1976b: 315
Type locality: Budafok
Distribution: Hungary
Host: Staphylinidae (Coleoptera)
Genus Porrhanoetus Mahunka
Porrhanoetus Mahunka, 1963a: 41
Type species: Porrhanoetus machadoi Mahunka, 1963
Porrhanoetus machadoi (Mahunka)
Porrhanoetus machadoi (Mahunka): Mahunka, 1963a: 41; 1967: 168
Type locality: Angola, Cavuemba River, Alto Cuílo, Poste de Cacolo
Distribution: Angola, Congo - Brazzaville
Genus Probonomoia Fain et Rack
Probonomoia Fain & Rack, 1987: 70
Type species: Probonomoia columbiana Fain et Rack, 1987
Probonomoia africana (Mahunka)
Bonomoia africana Mahunka, 1967: 171
Probonomoia africana (Mahunka): Fain & Rack 1987: 71
Type locality: Mount Foari Reservation
Distribution: Congo (Brazzaville)
Probonomoia chrysis Dimitrieff et Fain
Probonomoia chrysis Dimitrieff & Fain, 1994: 105
78
Acarorum Catalogus IX
Type locality: Spain
Distribution: Spain
Host: Chrysis sp. (Hymenoptera: Chrysididae)
Probonomoia columbiana Fain et Rack
Probonomoia columbiana Fain & Rack, 1987: 71
Type locality: Paramo de Chisaca, 40 km south of Bogota
Distribution: Columbia
Host: from the flower of Espeletia grandiflora (Asteraceae)
Probonomoia congoensis (Fain et Elsen)
Bonomoia congoensis Fain & Elsen, 1972: 89
Probonomoia congoensis (Fain et Elsen): Fain & Rack, 1987: 71
Type locality: Mongati
Distribution: DR Congo
Host: Glossina fuscipes quanzensis (Diptera: Glossinidae)
Probonomoia humprechti (Samšinák)
Bonomoia humprechti Samšinák, 1956: 354
Probonomoia humprechti (Samšinák): Klimov & Khausov, 2018: 2411
Type locality: Czech Republic
Distribution: Czech Republic, Hungary, Romania
Hosts: Laphria flava (Diptera: Asilidae), Ips spp. (Coleoptera: Cuculionidae, Scolytinae),
Acanthocinus aedilis (Col.: Cerambycidae)
Probonomoia opuntiae Wirth
Bonomoia opuntiae Wirth, 2005: 303
Probonomoia opuntiae (Wirth):
Type locality: Sardinia
Distribution: Sardinia, Corsica, Canary Islands (Gomera)
Probonomoia pini (Scheucher)
Bonomoia pini Scheucher, 1957: 366; Mahunka, 1970: 68; Woodring & Moser, 1970: 1256;
Probonomoia pini (Scheucher): Fain & Rack 1987: 71; Khaustov et al., 2018: 62
Type locality: Erlangen (Germany)
Distribution: Croatia, Czech Republic, England, Finland, Germany, Hungary, Japan,
Poland, Russia
Hosts: Hylaster ater, Ips acuminatus, I. cembrae, I. typographus (Coleoptera: Curculionidae,
Scolytinae), Dorcus parallelipipedus (Coleoptera: Lucanidae), Acanthocinus aedilis
(Col.: Cerambycidae), Helops quisquilius (Col.: Tenebrionidae)
Acarorum Catalogus IX 79
Probonomoia sibirica (Wirth)
Bonomoia sibirica Wirth, 2016: 99
Probonomoia sibirica: Khaustov et al., 2018: 64
Type locality: Tyumen Prov., East of Tyumen
Distribution: Russia
Hosts: Dendroctonus micans, Ips typographus (Coleoptera: Curculionidae, Scolytinae)
Probonomoia spinifera (Scheucher)
Bonomoia spinifera Scheucher, 1957: 367
Probonomoia spinifera (Scheucher): Fain & Rack, 1987: 71
Type locality: Erlangen
Distribution: Germany, Slovakia
Genus Prowichmannia Radford
Wichmannia Oudemans, 1929: 450 [praeocc.]
Prowichmannia Radford, 1950: 222
Type species: Histiostoma spiniferum Michael, 1901
Prowichmannia spiniferum (Michael)
Histiostoma spinifera Michael, 1901: 215; Oudemans, 1903:
Anoetus spiniferus Michael: Oudemans, 1902: 46
Wichmannia spiniferum (Michael): Oudemans, 1929:
Wichmannia spiniferus (Michael): Buitendijk, 1945: 361
Wichmannia spinifera (Michael): Scheucher, 1957: 370
Prowichmannia spiniferum (Michael): Radford, 1950: [160]; Beron, 1973: 22
Prowichmannia spinifera (Michael): Mahunka, 1970: 72; Krivolucliy, 1975: 413
Type locality: England
Distribution: Bulgaria, England, Germany, Hungary, Poland, Ukraine
Hosts: Cetonia aurata (Coleoptera: Scarabaeidae); many species of Mammals (Soricomorpha and Rodentia)
Prowichmannia stammeri (Scheucher)(new combination)
Wichmannia stammeri Scheucher, 1957: 374
Prowichmannia stammeri: p.p.
Type locality: Erlangen
Distribution: Germany
Prowichmannia talpae (Scheucher)
Wichmannia talpae Scheucher, 1957: 372
Prowichmannia talpae: Zamec, 2014:
80
Acarorum Catalogus IX
Type locality: England
Distribution: Germany, Slovakia
Host: nest of Talpa europaea (Soricomorpha: Talpidae)
Genus Psyllanoetus Fain et Beaucournu
Psyllanoetus Fain et Beaucournu , 1973: 516
Type species: Psyllanoetus Fain et Beaucournu , 1973
Psyllanoetus afropulicarius Fain et Beaucournu
Psyllanoetus afropulicarius Fain & Beaucournu, 1973: 516
Type locality: Lemera, Kivu
Distribution: DR Congo
Hosts: Listropsylla dolosa (Siphonaptera: Hystrichopsyllidae), ex Praomys jacksoni (Rodentia: Muridae), Stivalius torvus (Siphonaptera), ex Hybomys univittatus (Rodentia),
Dinopsyllus apistus (Siphonaptera), Ctenophthalmus devignati (Siphonaptera), ex H.
univittatus (Rodentia)
Psyllanoetus dinopsyllus Fain et Beaucournu
Psyllanoetus dinopsyllus Fain et Beaucournu, 1973: 517
Type locality: Lemera, Kivu
Distribution: DR Congo
Hosts: Dinopsyllus apistus (Siphonaptera), ex Praomys jacksoni (Rodentia: Muridae),
Ctenophthalmus debrauweri (Siphonaptera), ex Praomys jacksoni (Rodentia), Ct.
calceatus cabirus (Siphonaptera), ex Otomys sp. (Rodentia)
Psyllanoetus melesicola Fain et Beaucournu
Psyllanoetus melesicola Fain & Beaucournu, 1973: 516
Type locality: Les Sieges, Yonne
Distribution: France
Host: Paraceras melis (Siphonaptera: Ceratophyllidae), ex Meles meles (Carnivora: Mustelidae)
Genus Pteranoetus Mahunka
Pteranoetus Mahunka, 1978: 128
Type species: Pteranoetus kaszabi Mahunka, 1978
Pteranoetus incertus Mahunka
Pteranoetus incertus Mahunka, 1978a: 162
Type locality: Puerto Viejo, 10o26’N, 83o56’
Distribution: Costa Rica
Host: Eciton burchelli (Hymenoptera: Formicidae, Dorylinae)
Acarorum Catalogus IX 81
Pteranoetus kaszabi Mahunka
Pteranoetus kaszabi Mahunka, 1978: 128
Type locality: NE Nondugl, 2200–2700 m
Distribution: Papua New Guinea
Pteranoetus latus Mahunka
Pteranoetus latus Mahunka, 1978a: 162
Type locality: Oriente, 00o24S, 76o36’W, Limoncocha
Distribution: Ecuador
Pteranoetus rettenmeyeri Mahunka
Pteranoetus rettenmeyeri Mahunka, 1978a: 166
Type locality: Oriente, 00o24S, 76o36’W, Limoncocha
Distribution: Ecuador
Host: Neivamyrmex gradualis (Hymenoptera: Formicidae, Dorylinae)
Genus Rhaphidothrix Mahunka
Rhaphidothrix Mahunka, 1967: 173
Type species: Rhaphidothrix gozmanyi Mahunka, 1967
Rhaphidothrix gozmanyi Mahunka
Rhaphidothrix gozmanyi Mahunka, 1967a: 173
Type locality: Kindamba, Meya, Louolo river
Distribution: Rep. of the Congo (Brazzaville)
Rhaphidothrix punctolineatus Mahunka
Rhaphidothrix punctolineatus Mahunka, 1973a: 106; 1979: 128
Type locality: Pretsea
Distribution: Ghana
Host: Coleoptera
Rhaphidothrix simplex Mahunka
Rhaphidothrix simplex Mahunka, 1974b: 390
Type locality: Kumasi
Distribution: Ghana
Genus Rhopalanoetus Scheucher
Rhopalanoetus Scheucher, 1957: 362
Type species: Anoetus lanceocrinus Oudemans, 1914
82
Acarorum Catalogus IX
Rhopalanoetus bifurcatus Mahunka
Rhopalanoetus bifurcatus Mahunka, 1961b: 529; 1965: 449; 1969b: 371
Rhopalanoetus furcatus Mahunka: Samšiṅák, 1962
Type locality: Mt Meru, eastern slope (Tanzania)
Distribution: Rep. of the Congo (Brazzaville), Tanzania, DR Congo
Host: elephant excrement; Onitis sphinx (Coleoptera: Scarabaeidae)
Rhopalanoetus chinensis Samšiṅák
Rhopalanoetus chinensis Samšiṅák, 1962b: 188
Type locality: Kanton [Guandong]
Distribution: China
Host: Oniticellus cinctus (Coleoptera: Scarabaeidae)
Rhopalanoetus endroedyyoungai Mahunka
Rhopalanoetus endroedyyoungai Mahunka, 1979: 144
Type locality: Transvaal, Nylsvley Res., 35 km NE from Pretoria
Distribution: South Africa
Rhopalanoetus fimetarius (G. Canestrini et Berlese)
Histiostoma fimetarium Canestrini & Berlese, 1881: 150 (not Histiostoma fimetarius sensu
Hughes & Jackson, 1958)
Rhopalanoetus fimetarius (G. Canestrini et Berlese): Samsinak, 1962: 96; Mahunka, 1970:
66; Bongers et al., 1985: 48
Anoetus lanceocrinus Oudemans, 1914: 72
Histiostoma lanceocrinus: Buitendijk, 1945: 362; Hughes & Jackson, 1958: 82
Rhopalanoetus lanceocrinus: Scheucher, 1957: 362; Mahunka, 1962: 117
Anoetus tienhoveni Oudemans, 1926: 123 – Type loc.: Naardelmeer, Holland
Histiostoma tienhoveni: Buitendijk, 1945: 363
Type locality: Padova (Italy)
Distribution: Holland, Hungary, Italy
Rhopalanoetus gladifer Mahunka
Rhopalanoetus gladifer Mahunka, 1969b: 372
Type locality: Mt Meru, eastern slope
Distribution: Tanzania
Rhopalanoetus heterotrichus Mahunka
Rhopalanoetus heterotrichus Mahunka, 1973: 111
Type locality: Ghana
Distribution: Ghana
Acarorum Catalogus IX 83
Rhopalanoetus hirsutus Mahunka
Rhopalanoetus hirsutus Mahunka, 1973a: 305
Type locality: Nakpanduri
Distribution: Ghana
Rhopalanoetus lanceocrinus (Oudemans)
Anoetus lanceocrinus Oudemans, 1914: 72
Rhopalanoetus lanceocrinus (Oudemans): Scheucher, 1957: 362
Type locality: Holland
Distribution: Germany, Holland
Hosts: Aphodius fimetarius, A. fossor, A. merdarius, Geotrupes stercorarius (Coleoptera:
Scarabaeidae), Sphaeridium sp. (Col.: Hydrophilidae)
Rhopalanoetus robustus Mahunka
Rhopalanoetus robustus Mahunka, 1973a: 112
Type locality: Meru
Distribution: Tanzania
Rhopalanoetus simplex Samšiṅák
Rhopalanoetus simplex Samšiṅák, 1962b: 187
Type locality: Kanton [Guandong]
Distribution: China
Host: Oniticellus cinctus (Coleoptera: Scarabaeidae)
Rhopalanoetus trogicola (Fain, Vangeluwe, Degreef et Wauthy)
Histiostoma trogicola Fain, Vangeluwe, Degreef & Wauthy, 1993: 13
Rhopalanoetus trogicola (Fain, Vangeluwe, Degreef et Wauthy): Mašan, 2000:
Type locality: near Namur (Belgium)
Distribution: Belgium, Slovakia
Hosts: Trox scaber, T. cadaverinus (Coleoptera: Trogidae)
Genus Richardanoetus Samšinák
Richardanoetus Samšinák, 1989: 99
Type species: Richardanoetus demel Samšinák, 1989
Richardanoetus demel Samšiṅák
Richardanoetus demel Samšiṅák, 1989: 99
Type locality: Fuvo
Distribution: Ecuador
Host: Richardia sp. (plant)
84
Acarorum Catalogus IX
Genus Sarraceniopus Fashing et OConnor
Sarraceniopus Fashing & OConnor, 1984: 217
Type species: Zwickia gibsoni Nesbitt, 1954
Sarraceniopus darlingtoniae Fashing et OConnor
Sarraceniopus darlingtoniae Fashing & OConnor, 1984: 219
Type locality: pitchers of Darlingtonia californica, Darlingtonia wayside, Lane County,
Oregon
Distribution: USA (Oregon)
Sarraceniopus gibsoni (Nesbitt)
Zwickia gibsoni Nesbitt, 1954: 454
Anoetus gibsoni (Nesbitt): Atyeo, 1978: 244
Sarraceniopus gibsoni (Nesbitt): Fashing & OConnor, 1984: 219
Type locality: Danford Lake Area, Quebec
Distribution: Canada (N.S., Quebec, Ontario)
Host: the pitchers of Sarracenia purpurea (Sarraceniaceae)
Sarraceniopus hughesi (Hunter et Hunter)
Anoetus hughesi Hunter & Hunter, 1964: 39
Sarraceniopus hughesi (Hunter & Hunter): Fashing & OConnor, 1984: 219; Tagami, 2004:
169
Type locality: Georgia and North Carolina (USA)
Distribution: Japan (imported with plants), USA
Host: Sarracenia plants
Genus Scheucheria Mahunka
Scheucheria Mahunka, 1969a: 184
Type species: Scheucheria mongolica Mahunka, 1969
Scheucheria mongolica Mahunka
Scheucheria mongolica Mahunka, 1969a: 184
Type locality: Mongolia
Distribution: Mongolia
Genus Scutanoetus Mahunka
Scutanoetus Mahunka, 1969b: 367 [placement?]
Type species: Scutanoetus permixtus Mahunka, 1969
Scutanoetus permixtus Mahunka
Scutanoetus permixtus Mahunka, 1969b: 367
Acarorum Catalogus IX 85
Type locality: Mt Meru, eastern slope
Distribution: Tanzania
Genus Sellea Oudemans
Sellea Oudemans, 1929: 449
Type species: Histiostoma pulchrum Kramer, 1886
Sellea pulchrum (Kramer)
Histiostoma pulchrum Kramer, 1886: 259; Li et al., 1992
Sellea pulchrum (Kramer): Oudemans, 1929: 449; Hughes & Jackson, 1958: 175
Type locality: Germany
Distribution: England, Germany, China
Genus Semianoetus Mahunka
Semianoetus Mahunka, 1976b: 313
Type species: Semianoetus mrcziaki Mahunka, 1976
Semianoetus banksi Mahunka
Semianoetus banksi Mahunka, 1979: 341
Type locality: Puntarenas, Monteverde, 1400 m
Distribution: Costa Rica
Semianoetus mrcziaki Mahunka
Semianoetus mrcziaki Mahunka, 1976b: 313
Type locality: Budapest, Hüvõsvõlgy
Distribution: Hungary
Genus Spinanoetus Scheucher
Spinanoetus Scheucher, 1957: 357
Type species: Spinanoetus pelznerae Scheucher, 1957
Spinanoetus heterotrichus Mahunka
Spinanoetus heterotrichus Mahunka, 1962: 432; Kaluz, 1992:
Type locality: Dunazuggebirge, Sikáros
Distribution: Hungary, Slovakia
Spinanoetus pelznerae Scheucher
Spinanoetus pelznerae Scheucher, 1957: 358; Káluz, 1993; Haitlinger, 2004:
Type locality: Erlangen
Distribution: Germany, Hungary, Poland
86
Acarorum Catalogus IX
Hosts: Necrophorus interruptus, Oeceoptoma thoracica, Thanatophilus rugosus, Th. sinuatus (Coleoptera: Silphidae)
Spinanoetus variabilis Mahunka
Spinanoetus variabilis Mahunka, 1973a: 109
Type locality: Nakpanduri
Distribution: Ghana
Spinanoetus weingaertnerae Scheucher
Spinanoetus weingaertnerae Scheucher, 1957: 360; Haitlinger, 2004; Zamec, 2014
Type locality: Erlangen
Distribution: Germany, Hungary, Poland, Slovakia
Hosts: Emus hirtus, Ontolestes sp., Philontus sp. (Coleoptera: Staphylinidae)
Genus Stercoranoetus Mahunka et Mahunka-Papp
Stercoranoetus Mahunka & Mahunka-Papp, 1991: 285
Type species: Stercoranoetus harteni Mahunka et Mahunka-Papp, 1991
Stercoranoetus harteni Mahunka et Mahunka-Papp
Stercoranoetus harteni Mahunka & Mahunka-Papp, 1991: 285
Type locality: Santiago: Santa Cruz
Distribution: Cabo Verde
Stercoranoetus woolleyi (Mahunka et Eraky)
Histiostoma woolleyi Mahunka & Eraky, 1987: 135
Stercoranoetus woolleyi (Mahunka et Eraky): Mahunka & Mahunka- Papp, 1991: 285
Type locality: Hungary
Distribution: Hungary
Genus Synanoetus Mahunka
Synanoetus Mahunka, 1972d: 311
Type species: Synanoetus baloghi Mahunka, 1972
Synanoetus baloghi Mahunka
Synanoetus baloghi Mahunka, 1972d: 311
Type locality: Las Cardas (Prov. Coquimbo)
Distribution: Chile
Genus Syringanoetus Fain
Syringanoetus Fain, 1980: 116
Type species: Syringanoetus vandenberghi Fain, 1980
Acarorum Catalogus IX 87
Syringanoetus faini Mahunka et Eraky
Syringanoetus faini Mahunka & Eraky, 1987: 137
Type locality: Hungary
Distribution: Hungary
Syringanoetus vandenberghi Fain
Syringanoetus vandenberghi Fain, 1980: 116; Mahunka & Eraky 1987: 137
Type locality: Rugege Forest, 2500 m, 75 km from Butare
Distribution: Rwanda
Genus Teinokyra Mahunka
Teinokyra Mahunka, 1973a: 306
Type species: Teinokyra lineata Mahunka, 1973
Teinokyra lineata Mahunka
Teinokyra lineata Mahunka, 1973a: 307
Type locality: Pretsea
Distribution: Ghana
Genus Tensiostoma Wurst et Kovac
Tensiostoma Wurst & Kovac, 2003: 65
Type species: Tensiostoma veliaphilum Wurst et Kovac, 2003
Tensiostoma veliaphilum Wurst et Kovac, 2003
Tensiostoma veliaphilum Wurst & Kovac, 2003: 66
Type locality: Selangor, Darul Ehsan, Ulu Gombak Field Studies Centre, Univ. of Malaya
Distribution: Malaysia (Selangor)
Hosts: Lathriovelia rickmersi, L. capitata (Heteroptera: Veliidae)
Genus Traskorchestianoetus Fain et Colloff
Traskorchestianoetus Fain & Colloff, 1990: 667
Type species: Traskorchestianoetus spiceri Fain et Colloff, 1990
Traskorchestianoetus brevipes Fain et Colloff
Traskorchestianoetus brevipes Fain & Colloff, 1990: 672
Type locality: Dianna Island, Dodger Sound, British Columbia
Distribution: Canada
Host: Traskorchestia traskiana (Crustacea: Talitridae); marine
Traskorchestianoetus spiceri Fain et Colloff
Traskorchestianoetus spiceri Fain & Colloff, 1990: 668
88
Acarorum Catalogus IX
Type locality: Dianna Island, Dodger Sound, British Columbia
Distribution: Canada
Host: Traskorchestia traskiana (Crustacea: Talitridae); marine
Genus Xenanoetus Mahunka
Xenanoetus Mahunka, 1969: 359
Type species: Xenanoetus vestigialis Mahunka, 1969
Xenanoetus afrotropicalis Fain
Xenanoetus afrotropicalis Fain, 1984: 281
Type locality: Kamobuga, Rugege Forest, 2500 m
Distribution: Rwanda
Host: elephant dung
Xenanoetus grandiceps Mahunka
Xenanoetus grandiceps Mahunka, 1973: 61
Type locality: Mongolia
Distribution: Mongolia
Xenanoetus kabrt Samšiṅák
Xenanoetus kabrt Samšiṅák, 1989: 103
Type locality: DR Congo
Distribution: DR Congo
Host: Gymnometopina kibokoensis (Diptera: Sphaeroceridae)
Xenanoetus kofrankova Samšiṅák
Xenanoetus kofrankova Samšiṅák, 1989: 101
Type locality: Washington
Distribution: USA
Host: Coproica acutangula (Diptera: Sphaeroceridae)
Xenanoetus vestigialis Mahunka
Xenanoetus vestigialis Mahunka, 1969a: 359
Type locality: Finland
Distribution: Finland
Host: Sphaeridium sp. (Coleoptera: Hydrophiilidae)
Genus Zwickia Oudemans
Zwickia Oudemans, 1924: 310 [Diagnostic, Nesbitt 1979: 1205]
Type species: Anoetus guentheri Oudemans, 1915
Acarorum Catalogus IX 89
Zwickia colocasiae (Vitzthum)
Anoetus colocasiae Vitzthum, 1931: 118
Zwickia colocasiae (Vitzthum): Hughes & Jackson, 1958: 182
Type locality: Ngebal Lake
Distribution: Indonesia (Java)
Zwickia guentheri (Oudemans)
Anoetus guentheri Oudemans, 1915: 242
Zwickia guentheri (Oudemans): Oudemans, 1924: 310; Vitzthum, 1931: 123; Hughes &
Jackson, 1958: 177
Type locality: Sri Lanka
Distribution: Sri Lanka
Host: in pitchers of Nepenthes distillatoria
Zwickia nepenthesiana Hirst
Zwickia nepenthesiana Hirst, 1928: 19; Vitzthum, 1931: 131
Type locality: Singapore
Distribution: Singapore
Host: from the pitchers of Nepenthes ampullaria
Subfamily Myianoetinae Scheucher
Myianoetini Scheucher, 1957: 337
Myianoetinae: Fain, 1974: 58
Type genus: Myianoetus Oudemans, 1929
Genus Myianoetus Oudemans
Myianoetus Oudemans, 1929: 499
Anoetostoma Womersley, 1941: 485 [Type is oudemansi; syn. by Fain & Domrow 1980: 41]
Type species: Acarus muscarum Linnaeus, 1758
Myianoetus africanus Mahunka
Myianoetus africanus Mahunka, 1979d: 141
Type locality: Kenya
Distribution: Kenya
Myianoetus antipodus Fain et Galloway
Myianoetus antipodus Fain & Galloway, 1993: 106; Clark, 2009: 31; 2010a: 9
Type locality: Motunau Island
Distribution: New Zealand
Host: Eudyptula minor albosignata (Aves: Spheniscidae)
90
Acarorum Catalogus IX
Myianoetus assiuti Eraky, Abdel-Aziz et Abo-Elmaged
Myianoetus assiuti Eraky, Abdel-Aziz & Abo-Elmaged, 2020: 78
Type locality: Assiut, Upper Egypt
Distribution: Egypt
Myianoetus brevis Fain
Myianoetus brevis Fain, 1984: 287
Type locality: Rugege forest, 75 km from Butare, 2300 m
Distribution: Rwanda
Host: elephant dung
Myianoetus clavatus Mahunka
Myianoetus clavatus Mahunka, 1963d: 369; 1970: 51
Type locality: Hortobágy-Máta
Distribution: Hungary
Host: Spermophilus [Citellus] citellus (Rodentia: Sciuridae)
Myianoetus copromyzae Fain, Britt et Molyneux
Myianoetus copromyzae Fain, Britt & Molyneux, 1980: 402
Type locality: Scotland
Distribution: Scotland
Host: Copromyza sp. (Diptera: Sphaeroceridae)
Myianoetus digitiferus (Trägårdh)
Histiostoma digitifera Trägårdh, 1904: 67
Anoetus digitiferus (Trägårdh): Vitzthum, 1925: 155
Myianoetus digitiferus (Trägårdh): Willmann, 1937: 183
Type locality: Greenland
Distribution: Greenland, Germany
Host: Heleomyza [Blepharoptera] serrata (Diptera: Helomyzidae)
Myianoetus dionychus (Oudemans)
Anoetus dionychus Oudemans, 1910: 50
Myianoetus diadematus Willmann, 1937: 182
Myianoetus dionychus (Oudemans): Scheucher, 1957: 339
Type locality: Holland (Maastricht)
Distribution: France, Germany, Holland, Hungary, Romania, Slovakia
Hosts: flies (Diptera: Heleomyzidae); Plecotus auritus (Chiroptera: Vespertilionidae)
Myianoetus domrowi (Fain)
Anoetostoma domrowi Fain, 1968: 246
Acarorum Catalogus IX 91
Myianoetus domrowi (Fain)
Type locality: Butemu Village, Finisterre Range, Madang Central District
Distribution: Papua New Guinea
Myianoetus dycei Fain et Domrow
Myianoetus dycei Fain & Domrow, 1980: 41
Type locality: Queensland, Rockhampton
Distribution: Australia
Host: Culicoides brevitarsis (Diptera: Ceratopogonidae)
Myianoetus filicum (Dujardin)
Hypopus filicum Dujardin, 1841: 261
Myianoetus filicum (Dujardin):
Type locality: France
Distribution: France, Hungary
Myianoetus granatum Eraky
Myianoetus granatum Eraky, 2020: 153
Type locality: Assiut
Distribution: Egypt
Myianoetus heterotrichus Mahunka
Myianoetus heterotrichus Mahunka, 1968: 252
Type locality: Hungary
Distribution: Hungary
Myianoetus kaszabi Mahunka
Myianoetus kaszabi Mahunka, 1967c: 19
Type locality: Chentej aimak, 7 km NE of Somon Mörön
Distribution: Mongolia
Myianoetus lili Eraky
Myianoetus lili Eraky, 1993: 47; Negm & Alatawi, 2011: 97; Abdelgayed et al., 2019: 32
Type locality: Assiut, upper Egypt
Distribution: Egypt, Saudi Arabia
Host: manure
Myianoetus longisetosus Mašan et Krištofik
Myianoetus longisetosus Mašan & Krištofik, 1992: 84
Type locality: Slovakia
Distribution: Slovakia
92
Acarorum Catalogus IX
Host: Hydrotaea sp. (Diptera: Muscidae)
Myianoetus manurei Eraky, Abdel-Aziz et Abo-Elmaged
Myianoetus manurei Eraky, Abdel-Aziz & Abo-Elmaged, 2020: 76
Type locality: Assiut, Upper Egypt
Distribution: Egypt
Myianoetus micromaculatus Fain et Philips
Myianoetus micromaculatus Fain & Philips, 1979: 151
Type locality: Syracuse, N.Y.
Distribution: USA
Host: in the nest of Otus asio (Aves: Strigiformes)
Myianoetus microti Sevastianov et Vysotskaya
Myianoetus microti Sevastianov & Vysotskaya, 1971: 41
Type locality: Transcarpatian
Distribution: Ukraine
Host: in the nest of rodents
Myianoetus mlcoch Samšiṅák
Myianoetus mlcoch Samšiṅák, 1989: 94
Type locality: Pullman, Washington
Distribution: USA
Host: Copromyza (Borborillus) sordidus (Diptera: Sphaeroceridae)
Myianoetus muscarum (Linnaeus)
Acarus muscarum Linnaeus, 1758: 617
Myianoetus muscarum (Linnaeus): Buitendijk, 1945: 361; Scheucher, 1957: 338; Nuorteva,
1971; Sevastianov, 1975: 401; Oconnor et al., 2015: 1; Pimsler et al., 2016: 290
Type locality: Europe
Distribution: Belgium, Germany, England, Holland, Finland, ?France, Italy, Canada, USA
Hosts: Synthesiomyia nudiseta and other flies (Diptera: Muscidae), sometime hosts not
sure (Pimsler et al., 2016).
Myianoetus mystacis Mahunka
Myianoetus mystacis Mahunka, 1972b: 363
Type locality: Vértes-Geb., Fánivölgy
Distribution: Hungary
Host: Fannia sp. (Diptera: Fanniidae)
Myianoetus nerminka (Eraky et Shoker)
Anoetostoma nerminka Eraky & Shoker, 1995: 21
Acarorum Catalogus IX 93
Myianoetus nerminka (Eraky et Shoker):
Type locality: Egypt
Distribution: Egypt
Myianoetus oudemansi (Womersley)
Anoetostoma oudemansi Womersley, 1941: 485; Hughes & Jackson, 1958: 126
Myianoetus oudemansi (Womersley): Fain & Domrow 1980: 41
Type locality: Sidney, NSW
Distribution: Australia
Host: Musca domestica (Diptera: Muscidae)
Myianoetus ovatus Mašan et Kristofik
Myianoetus ovatus Mašan & Kristofik, 1992: 81
Type locality: Slovakia
Distribution: Slovakia
Host: Hydrotaea sp. (Diptera: Muscidae)
Myianoetus paganus Sevastianov
Myianoetus paganus Sevastianov, 1973: 856
Type locality: Tchemerovtsi Village, Hmelnitskaya Obl., Ukraine
Distribution: Ukraine
Host: horse dung
Myianoetus pappi Mahunka
Myianoetus pappi Mahunka, 1972b: 361
Type locality: Aranyosgadány, Kuhfladen
Distribution: Hungary
Host: Azelia parva (Diptera: Muscidae)
Myianoetus parvus Mahunka
Myianoetus parvus Mahunka, 1969a: 360; 1972b: 363
Type locality: Finland
Distribution: Finland, Hungary, Slovakia
Hosts: Ophyra leucostoma (Diptera: Muscidae), Borboropsis fulviceps (Diptera: Heleomyzidae)
Myianoetus pseudomuscarum Sevastianov
Myianoetus pseudomuscarum Sevastianov, 1973: 856
Type locality: Tchemerovtsi Village, Hmelnitskaya Obl., Ukraine
Distribution: Russia (Sakhalin, Tatarstan, Bashkortistan), Ukraine
Host: cow’s dung
94
Acarorum Catalogus IX
Myianoetus reductus Mahunka
Myianoetus reductus Mahunka, 1969d: 368
Type locality: Mt Meru, eastern slope
Distribution: Tanzania
Myianoetus rohaceki Samšiṅák
Myianoetus rohaceki Samšiṅák, 1984: 57; 1989: 91
Type locality: Hrubý Jesenik Mts., Skřitek
Distribution: Czech Rep., Hungary, USA
Hosts: Limosina rufilabris, Coproica vistripennis, Lotophila atra (Diptera: Sphaeroceridae)
Myianoetus rwandanus Fain
Myianoetus rwandanus Fain, 1984: 283
Type locality: Kamobuga, Rugege Forest, 2500 m
Distribution: Rwanda
Host: elephant dung
Myianoetus scheucherae Mahunka
Myianoetus scheucheri Mahunka, 1972: 371 (as nom.nov. for the deutonymphs recorded
as “M. muscarum” by Scheucher, 1957: 338)
Myianoetus scheucherae Mahunka: Klimov & Khaustov, 2018
Type locality: Erlangen
Distribution: Germany
Myianoetus simplex Mahunka
Myianoetus simplex Mahunka, 1972: 363; Bongers, OConnor & Lukoschus, 1985: 19
Type locality: Aranyosgadány (Hungary)
Distribution: Czech Rep., France, Holland, Hungary
Hosts: Coproica lugubris (Diptera: Sphaeroceridae), Hebecnema umbratica (Diptera:
Muscidae), Sepsis thoracica (Diptera: Sepsidae)
Myianoetus simunek Samšiṅák
Myianoetus simunek Samšiṅák, 1989: 91
Type locality: Kiskunsagi N. P. Kunfehérto (lake)
Distribution: Hungary
Host: Coproica lugubris (Diptera: Sphaeroceridae)
Myianoetus sovat Zamec
Myianoetus sovat Zamec, 2014: 916
Type locality: Vlčie Diery Cave
Distribution: Slovakia
Host: phoretic on Gymnomus caesius (Diptera: Heleomyzidae)
Acarorum Catalogus IX 95
Myianoetus szaboi Mahunka
Myianoetus szaboi Mahunka, 1978: 109
Type locality: Kócs (Hortobágy Nat. Park)
Distribution: Hungary
Myianoetus szunyoghyi Mahunka
Myianoetus szunyoghyi Mahunka, 1969d: 369
Type locality: Mt Meru, eastern slope
Distribution: Tanzania
Myianoetus thaleri Mahunka
Myianoetus thaleri Mahunka, 1972: 374
Type locality: Unterengadin, Ramosch, Charbunera
Distribution: Austria
Myianoetus travei Fain
Myianoetus travei Fain, 1976: 309
Type locality: Kerguelen Island
Distribution: Kerguelen Island
Myianoetus tuerkorum Scheucher
Myianoetus tuerkorum Scheucher, 1957: 341; Mahunka, 1970: 48; Fain et al., 1993: 14
Type locality: Erlangen (Germany)
Distribution: Belgium, Germany, Slovakia
Host: bird’s nests
Myianoetus tuomikoskii Mahunka
Myianoetus tuomikoskii Mahunka, 1969: 361
Type locality: Finland
Distribution: Finland, Slovakia, Romania, Mongolia
Hosts: Hydrotaea occulta, H. tuberculata (Diptera: Muscidae)
Myianoetus uncinatus Fain
Myianoetus uncinatus Fain, 1984: 285
Type locality: Kamobuga, Rugege Forest, 2500 m
Distribution: Rwanda
Host: elephant dung
Myianoetus undulatus Hughes et Jackson
Myianoetus undulatus Hughes & Jackson, 1958: 158
Type locality: Chickahominy Swamp, Ellerson, Virginia
Distribution: USA
96
Acarorum Catalogus IX
Myianoetus vesparum Scheucher
Myianoetus vesparum Scheucher, 1957: 343
Type locality: Erlangen
Distribution: Germany
Host: nests of Vespidae (Hymenoptera)
Myianoetus virgatus Scheucher
Myianoetus virgatus Scheucher, 1957: 345; Mahunka, 1970: 49
Type locality: Germany
Distribution: Czech Rep., Germany, Slovakia
Hosts: Diptera (Borboridae, Scatophagidae), Limosina clunipes (Diptera: Sphaeroceridae)
Myianoetus zdena Samšiṅák
Myianoetus zdena Samšiṅák, 1984: 60; 1989: 91
Type locality: Kremnicke pohoria, Mts Tuček, 850 m
Distribution: Austria, Hungary, Slovakia
Hosts: Elachisoma aterrima, Coproica lugubris, C. pallifrons, C. vagans (Diptera: Sphaeroceridae)
Subfam. Scolianoetinae Fain
Scolianoetinae Fain, 1974: 60
Type genus: Scolianoetus Fain, 1974
Genus Scolianoetus Fain
Scolianoetus Fain, 1974: 60
Type species: Scolianoetus scheucherae Fain, 1974
Scolianoetus brasiliensis Fain
Scolianoetus brasiliensis Fain, 1974: 62
Type locality: Novo Friburgo
Distribution: Brazil
Host: Spartocera cinnamomea (Heteroptera: Coreidae)
Scolianoetus faini Mahunka
Scolianoetus faini Mahunka, 1975a: 321
Type locality: Kiunga
Distribution: Papua New Guinea
Scolianoetus javanus Mahunka
Scolianoetus javanus Mahunka, 1976b: 319
Type locality: Java, Pangerango
Acarorum Catalogus IX 97
Distribution: Indonesia (Java)
Host: Pelopides tridens (Lepidoptera: Hesperiidae)
Scolianoetus ramozae Fain et Camerik
Scolianoetus ramozae Fain & Camerik, 1978: 169
Type locality: Rio de Janeiro
Distribution: Brazil
Host: Passalus sp. (Coleoptera: Passalidae)
Scolianoetus scheucherae Fain
Scolianoetus scheucherae Fain, 1974: 62
Type locality: Lisala
Distribution: DR Congo
Host: Campsomeris mansueta (Hymenoptera: Scoliidae)
Subfam. Spinanoetinae Scheucher
Spinanoetini Scheucher, 1957: 357
Spinanoetinae: Fain, 1974: 58
Type genus: Spinanoetus Scheucher, 1957
Genus Spinanoetus Scheucher
Spinanoetus Scheucher, 1957: 357
Type species: Spinanoetus pelznerae Scheucher, 1957
Spinanoetus heterotrichus Mahunka
Spinanoetus heterotrichus Mahunka, 1962: 432; Kaluz, 1992:
Type locality: Dunazuggebirge, Sikáros
Distribution: Hungary, Slovakia
Spinanoetus pelznerae Scheucher
Spinanoetus pelznerae Scheucher, 1957: 358; Káluz, 1993; Haitlinger, 2004:
Type locality: Erlangen (Germany)
Distribution: Germany, Hungary, Poland
Hosts: Nicrophorus interruptus, Oeceoptoma thoracica, Thanatophilus rugosus, Th. sinuatus (Coleoptera: Silphidae)
Spinanoetus variabilis Mahunka
Spinanoetus variabilis Mahunka, 1973a: 109
Type locality: Nakpanduri
Distribution: Ghana
98
Acarorum Catalogus IX
Spinanoetus weingaertnerae Scheucher
Spinanoetus weingaertnerae Scheucher, 1957: 360; Haitlinger, 2004; Zamec, 2014
Type locality: Erlangen (Germany)
Distribution: Germany, Hungary, Poland, Slovakia
Hosts: Emus hirtus, Ontolestes sp., Philontus sp. (Coleoptera: Staphylinidae)
Index of Histiostomatidae
(Names in bold are accepted as valid)
abietis, Histiostoma 39
abietis, Anoetus 39
acanthotarsus, Peripatetes 77
acrocini, Histiostoma 39
adarcuale, Histiostoma 39
adsimile, Histiostoma 40
aelleni, Anoetus 23
aeschlimanni, Hormosianoetus 73
africana, Bonomoia 36
africana, Probonomoia 77
africanus, Anoetoglyphus 21
africanus, Glyphanoetus 18
africanus, Myianoetus 89
afropulicarius, Psyllanoetus 80
afrotropicalis, Xenanoetus 88
akodoni, Anoetus 23
alicola, Anoetus 23
alicola, Hypopus 23
alii, Histiostoma 40
alphaticum, Histiostoma 40
amargynus, Histiostoma 40
Ameranoetus 20
americanum, Histiostoma 40
amieti, Histiostoma 40
Amyzanoetus 20
Ancyranoetus 21
andrei, Histiostoma 40
angolensis, Histiostoma 40
anguillarum, Histiostoma 72
angularis, Anoetus 24
Anoetidae 14
Anoetinae 20
Anoetoglyphus 21
anoetopsis, Anoetus 24
Anoetostoma 89
Anoetus 23
antennatus, Anoetoglyphus 21
antillense, Histiostoma 41
antipodus, Myianoetus 89
Aphodanoetus 35
arcuatus, Histiostoma 41
arcuata, Histiostoma 41
arenapus, Histiostoma 41
asper, Capronomoia 15
asper, Copronomoia 16
assiuti, Myianoetus 90
ateuchi, Anoetoglyphus 21
Auricanoetus 36
australiense, Histiostoma 41
australis, Kanoetus 75
Austranoetus 36
bacilligerum, Histiostoma 41
bakeri, Histiostoma 41
baloghi, Histiostoma 41
baloghi, Synanoetus 86
bananai, Histiostoma 42
banjuwangicus, Anoetus 41
banjuwangicus, Histiostoma 41
banksi, Semianoetus 85
bassoni, Loxanoetus 75
Acarorum Catalogus IX 99
bellulus, Capronomoia 16
bellulus, Copronomoia 16
berghi, Histiostoma 42
berghii, Histiostoma 42
berndhauseri, Histiostoma 42
bernoni, Histiostoma 42
bifastigatus, Anoetoglyphus 21
bifoveolatum, Histiostoma 42
bifurcatus, Rhopalanoetus 82
blomquisti, Histiostoma 42
Bonomoia 36
Bothyanoetus 38
brachypodus, Insulanoetus 74
bradymeri, Anoetus 24
brasiliensis, Scolianoetus 96
brevimanum, Histiostoma 42
brevimana, Lipstorpia 42
brevimanus, Anoetus 43
brevimanus, Histiostoma 42
brevipes, Histiostoma (nomen dubium) 43
brevipes, Traskorchestianoetus 87
brevipodum, Histiostoma 43
brevis, Myianoetus 90
brevisetus, Glyphanoetus 18
bushlandi, Anoetus 24
cacuminatus, Anoetus 24
calicifera, Histiostoma 43
caljubiensis, Anoetus 24
camerunensis, Histiostoma 43
campanula, Anoetus 43
campanula, Histiostoma 43
campanulae, Histiostoma 43
camphori, Histiostoma 43
capillatum, Histiostoma 43
capillata, Histiostoma 43
capillatus, Anoetus 24
Capronomoia 15
caro, Histiostoma 44
castrii, Anoetus 24
cataglyphi, Histiostoma 44
Cavanoetus 72
Cederhjelmia 38
certa, Bonomoia 36
ceylanensis, Anoetus 25
Ceylanoetinae 14
Ceylanoetus 14
chalinolobus, Chiropteranoetus 15
chilensis, Anoetus 25
Chiloanoetus 38
chilopophila, Histiostoma 44
chinensis, Rhopalanoetus
Chiropteranoetus 15
chrysis, Probonomoia 77
cirratus, Anoetus 44
cirratum, Histiostoma 44
cirrosus, Anoetus 25
clavatum, Histiostoma 44
clavatus, Myianoetus 90
clavifer, Histiostoma 44
collis, Anoetus 25
colocasiae, Anoetus 89
colocasia, Zwickia 89
colocasiae, Zwickia 89
columbiana, Probonomoia 78
Comyianoetus 15
conclavicola, Anoetus 44
conclavicola, Histiostoma 44
Conglanoetus 15
congoensis, Anoetus 25
congoensis, Bonomoia 78
congoensis, Probonomoia 78
conjuncta, Anoetus 45
conjuncta, Histiostoma 45
conjunctum, Histiostoma 45
conoidalis, Hexanoetus 39
continuatum, Histiostoma 45
contrasetosus, Kanoetus 75
coprae, Histiostoma 45
copridis, Anoetoglyphus 22
copromyzae, Myianoetus 90
coprophilus, Anoetus 25
coronatum, Histiostoma 45
100
Acarorum Catalogus IX
cossi, Histiostoma 45
crantor, Histiostoma 45
crassipes, Anoetus 45
crassipes, Histiostoma 45
Creutzeria 16
crypturgi, Anoetus 46
crypturgi, Histiostoma 46
cubanus, Kanoetus 75
cubanum, Histiostoma 46
Curculanoetus 17
cyrtandrae, Anoetus 46
cyrtandrae, Histiostoma 46
dactyloscopicum, Histiostoma 46
darlingtoniae, Sarraceniopus 84
darwishii, Histiostoma 46
debilis, Anoetus 25
decemvirgae, Histiostoma 46
decorus, Anoetus 25
demel, Richardanoetus 83
densus, Histiostoma 47
denticulatus, Comyianoetus 15
denticulatus, Conglanoetus 15
diademata, Histiostoma 47
diadematum, Histiostoma 47
diadematus Myianoetus 90
digitifera, Histiostoma 90
digitiferus, Anoetus 90
digitiferus, Myianoetus 90
dinopsyllus, Psylanoetus 80
dionychus, Anoetus 90
dionychus, Myianoetus 90
discrepans, Anoetus 23
disiunctus, Anoetus 47
disiuncta, Histiostoma 47
disiunctum, Histiostoma 47
distentoseta, Histiostoma 47
distentosetum, Histiostoma 47
dolifer, Anoetus 26
domrowi, Anoetostoma 26
domrowi, Anoetus 26
domrowi, Myianoetus 90
dryocoeti, Anoetus 47
dryocoeti, Histiostoma 47
dudichi, Anoetus 26
dugesi, Histiostoma 47
dugesii, Hypopus 47
duplex, Histiostoma 47
dycei, Myianoetus 91
eboris, Glyphanoetus 18
edax, Histiostoma 48
edita, Histiostoma 48
editum, Histiostoma 48
egyptiacum, Histiostoma 48
eickworti, Histiostoma 48
ellipticus, Anoetoglyphus 22
elzingai, Histiostoma 48
endroedii, Anoetus 48
endroedii, Histiostoma 48
endroedyyoungai, Anoetus 26
endroedyyoungai, Rhopalanoetus 82
ensifer, Anoetus 48
ensifer, Histiostoma 48
erugo, Histiostoma 48
excavatus, Ceylanoetus 14
exornatus, Anoetus 26
faini, Scolianoetus 96
faini, Syringanoetus 87
farghali, Histiostoma 49
feroniarum, Anoetus 49
feroniarum, Histiostoma 49
feroniarum, Hypopus 49
fetters, Anoetus 26
Fibulanoetinae 17
Fibulanoetus 17
fibulatus, Anoetus 26
filicum, Hypopus 91
filicum, Myianoetus 91
fimetarium, Histiostoma 82
fimetarius, Rhopalanoetus 82
flaccus, Histiostoma 49
flagellifer, Histiostoma 49
flagellatus, Glyphanoetus 18
Acarorum Catalogus IX 101
flabelliforme, Histiostoma 49
flagellifer, Histiostoma 49
formicarum, Histiostoma 50
formosana, Histiostoma 50
fortis, Histiostoma 50
foveosimile, Histiostoma 50
fragosus, Histiostoma 50
fucatifer, Histiostoma 50
fuliginosi, Histiostoma 50
fulmeki, Glyphanoetus 19
furcata, Histiostoma 50
furcatus, Rhopalanoetus 82
galerus, Histiostoma 51
gambiense, Histiostoma 51
gampsoceri, Histiostoma 51
genavensium, Histiostoma 51
genetica, Histiostoma 74
gervaisi, Histiostoma 51
ghanensis, Anoetus 26
Ghanoetus 18
gibsoni, Anoetus 84
gibsoni, Sarraceniopus 84
gibsoni, Zwickia 84
gladifer, Rhopalanoetus 82
gladiger, Anoetus 51
gladiger, Histiostoma 51
globotuberum, Histiostoma 51
globulus, Anoetoglyphus 22
Glyphanoetinae 14
Glyphanoetus 18
gordius, Anoetus 52
gordius, Histiostoma 52
gorgo, Histiostoma 52
gozmanyi, Rhaphidothrix 81
gracilipes, Histiostoma 52
gracilis, Histiostoma 52
gradatus, Capronomoia 16
gradatus, Copronomoia 16
graeca, Hauserania 38
granatum, Myianoetus 91
grandiceps, Xenanoetus 88
granulatum, Histiostoma 52
grapefruit, Anoetus 27
guanophilum, Histiostoma 52
guentheri, Anoetus 89
guentheri, Zwickia 89
halicticola, Histiostoma 52
halicticolum, Histiostoma 52
halictonida, Anoetus 27
halictonida, Histiostoma 27
halophilus, Amyzanoetus 21
harteni, Stercoranoetus 86
Hauserania 38
hauseri, Anoetus 27
heinemanni, Histiostoma 53
helena, Anoetus 27
heliocopridis, Anoetus 27
helobius, Acarus 27
helobius, Anoetus 27
hematobii, Ameranoetus 20
herbali, Histiostoma 53
heriades, Histiostoma 53
heterotrichus, Myianoetus 91
heterotrichus, Spinanoetus 85
Hexanoetus 39
himalayae, Anoetus 53
himalayae, Histiostoma 53
hirsutus, Rhopalanoetus 83
histeri, Histiostoma 53
Histiostoma 39
Histiostomatidae 14
Histiostomatinae 20
Histiostomatini 20
Histiostomatinae 20
Histiostomatoidea 11
Hormosianoetus 73
horridum, Histiostoma 53
horridus, Anoetus 27
hospitus, Anoetus 28
huettleri, Histiostoma 54
hughesi, Anoetus 84
hughesi, Sarraceniopus 84
102
Acarorum Catalogus IX
humiditatis, Anoetus 54
humiditatis, Histiostoma 54
humilis, Anoetus 28
humprechti, Bonomoia 78
humprechti, Probonomoia 78
hungarica, Histiostoma 54
Hymenanoetus 74
Ichtanoetus 72
imbecillus, Histiostoma 54
immaturus, Anoetoglyphus 22
impressum, Histiostoma 28
impressus, Anoetus 28
incanus, Glyphanoetus 18
incertus, Pteranoetus 80
incurvata, Histiostoma 54
incurvatum, Histiostoma 54
indetonsum, Histiostoma 54
indetonsus, Anoetus 54
indica, Histiostoma 28
indicus, Anoetus 28
inquilinum, Histiostoma 54
insolita, Anoetus 28
insulare, Histiostoma 55
insularis, Anoetus 55
Insulanoetus 74
ionicum, Histiostoma 55
irinum, Histiostoma 55
irinus, Anoetus 55
janosi, Histiostoma 55
javanus, Scolianoetus 96
julorum, Anoetus 52
julorum, Histiostoma 55
julorum, Hypopus 55
junctus, Histiostoma 55
kabrt, Xenanoetus 88
Kanoetus 75
karachiensis, Histiostoma 56
Kaszabanoetinae 14
Kaszabanoetus 75
kaszabi, Anoetus 29
kaszabi, Myianoetus 91
kaszabi, Pteranoetus 81
kerguelenense, Histiostoma 56
kerguelenensis, Austranoetus 36
kofrankova, Xenanoetus 88
komarpictoris, Anoetus 29
labiatus, Fibulanoetus 17
laboratorium, Histiostoma 74
laboratorium, Hormosianoetus 74
laevis, Histiostoma 56
laevis, Zschachia (nomen dubium) 56
lanceocrinus, Anoetus 83
lanceocrinus, Rhopalanoetus 83
langi, Anoetoglyphus 22
latus, Anoetoglyphus 22
latus, Pteranoetus 81
lekiensis, Histiostoma 56
lenae, Loxanoetus 76
leptos, Glyphanoetus 19
levis, Anoetoglyphus 22
lili, Myianoetus 91
lineata, Teinokyra 87
lineatula, Histiostoma 56
lineosculpturatum, Histiostoma 56
Lipstorpia
litorale, Histiostoma 56
litoralis, Anoetus 56
loebli, Histiostoma 57
loksai, Histiostoma 57
longipes, Anoetus 29
longipes, Histiostoma 29
longisetosus, Myianoetus 91
longisetum, Histiostoma 57
longisetus, Anoetus 29
longispina, Glyphanoetus
longitarsus, Fibulanoetus 17
longitarsus, Auricanoetus 36
longitarsus, Loxanoetus (A.) 36
longitubus, Anoetus 29
longus, Glyphanoetus 19
lootsi, Anoetoglyphus 22
lorentzi, Anoetus 57
Acarorum Catalogus IX 103
lorentzi, Histiostoma 57
louisianae, Histiostoma 57
Loxanoetus 75
lunulatus, Anoetus 29
luxus, Histiostoma 57
machadoi, Anoetus 29
machadoi, Porrhanoetus 77
macrosuctus, Anoetus 30
magdolnae, Histiostoma 57
mahunkai, Capronomoia 16
mahunkai, Fibulanoetus 17
mahunkai, Glyphanoetus 19
mahunkai, Histiostoma 57
mahunkai, Hormosianoetus 74
mallotae, Hormosianoetus 74
mallotae, Mauduytia 74
manicae, Histiostoma 58
mannai, Histiostoma 58
mantelli, Histiostoma 58
manurei, Myuanoetus 92
manuri, Histiostoma 58
marani, Histiostoma ovalis 61
margaritata, Histiostoma 58
margaritatum, Histiostoma 58
marianae, Histiostoma 58
maritimus, Anoetus 58
maritimum, Histiostoma 58
marmoreus, Ghanoetus 18
matsakisi, Histiostoma 59
Mauduytia 76
mayumbensis, Chiloanoetus 38
media, Anoetus 59
medium, Histiostoma 59
melesicola, Psyllanoetus 80
melolonthae, Histiostoma 59
meszarosorum, Histiostoma 59
metwallii, Histiostoma 59
micidus, Glyphanoetus 19
micromaculatus, Myianoetus 92
microti, Anoetus 30
microti, Myianoetus 92
minimum, Histiostoma 59
mirabile, Histiostoma 59
mlkoch, Myianoetus 92
mongolica, Scheucheria 84
montanum, Histiostoma 60
moralesi, Histiostoma 60
moshtohorensis, Anoetus 30
mrcziaki, Semianoetus 85
Munduytia 76
murchiei, Histiostoma 60
muscae, Histiostoma 60
muscarum, Acarus 92
muscarum, Myianoetus 92
Myianoetinae 89
Myianoetini 89
Myianoetus 89
myrmicarum, Anoetus 30
myrmicarum, Histiostoma 30
mystacis, Myianoetus 92
nasseri, Histiostoma 60
necopinus, Anoetus 60
necopinus, Histiostoma 60
necrophori, Anoetus
necrophori, Histiostoma
necrophori, Hypopus
neglectus, Anoetus 30
negmi, Histiostoma 60
Nepenthacarus 76
nepenthesiana, Anoetus 89
nepenthesiana, Zwickia 89
nerminka, Anoetostoma 92
nerminka, Myianoetus 92
nichollsi, Histiostoma 62
nigrellii, Histiostoma 73
Nodipalpidae 14
Nodipalpus 23
nodosus, Hymenanoetus 74
nomiensis, Glyphanoetus 19
notabilis, Anoetus 30
novamsterdamense, Histiostoma 61
ocellatum, Histiostoma 73
104
Acarorum Catalogus IX
onioni, Histiostoma 61
onthophagae, Histiostoma 61
opertus, Anoetus 30
opuntiae, Bonomoia 78
opuntiae, Probonomoia 78
orientalis, Histiostoma 61
ornata, Hauserania 39
ornatissimus, Anoetus 31
ornatofoveolatus, Conglanoetus 15
ornatus, Anoetus 31
Otanoetus 76
oudemansi, Anoetostoma 93
oudemansi, Anoetus 61
oudemansi, Histiostoma 61
oudemansi, Myianoetus 93
ovalis, Hypopus 51
ovale, Histiostoma 61
ovalis, Histiostoma 61
ovalis, Uropoda 61
Ovanoetus 77
ovatus, Myianoetus 93
oxfordianum, Histiostoma
paganus, Myianoetus 93
palustre, Histiostoma 62
pango, Capronomoia 16
pango, Copronomoia 16
pannonicum, Histiostoma
pannonicus, Anoetus 31
papillata, Histiostoma 62
papillatum, Histiostoma 62
pappi, Glyphanoetus 19
pappi, Myianoetus 93
parmatus, Anoetus 31
parvus, Myianoetus 93
pauciputeolum, Histiostoma 62
pauliani, Anoetus 31
pectineum, Histiostoma 49
pelznerae, Spinanoetus 85
pereirai, Bothyanoetus 38
Peripatetes 77
permixtus, Scutanoetus 84
perruptum, Histiostoma 62
phyllophorum, Histiostoma 62
phyllophorus, Anoetus 62
phyllophorus, Histiostoma 62
phyllotrichus, Anoetus 19, 52
phyllotrichus, Glyphanoetus 19
phyllotrichus, Hypopus 19
phytophaga, Histiostoma 62
piceae, Anoetus 63
piceae, Histiostoma 63
pickaxei, Histiostoma 63
picturata, Bonomoia 37
pierrestrinatii, Histiostoma (Cavanoetus)
72
piloseta, Histiostoma 63
pini, Anoetus 31
pini, Bonomoia 78
pini, Histiostoma 31
pini, Probonomoia 78
piscium, Histiostoma 73
pocsi, Anoetus 31
polaki, Anoetus 63
polaki, Histiostoma 63
polygonatus, Anoetoglyphus 23
polynesiacus, Anoetus 32
polypori, Anoetus 63
polypori, Histiostoma 63
pontigerum, Histiostoma 63
porosus, Anoetus 32
Porrhanoetus 77
primitiva, Bonomoia 37
privorai, Anoetus 32
probata, Bonomoia 37
Probonomoia 77
processum, Glyphanoetus 20
prodectoris, Anoetus 63
prodectoris, Histiostoma 63
prophthalmi, Anoetus 64
prophthalmi, Histiostoma 64
protuberans, Histiostoma 64
Prowichmannia 79
Acarorum Catalogus IX 105
pseudolongipes, Histiostoma 64
pseudomuscarum, Myianoetus 93
Psyllanoetus 80
Pteranoetus 80
pulchrum, Histiostoma 85
pulchrum, Sellea 85
punctaticeps, Histiostoma 64
punctolineatus, Rhaphidothrix 81
punctulatus, Anoetus 32
puteanus, Comyianoetus 15
puteanus, Conglanoetus 15
pyriforme, Histiostoma 64
quadriuncinata, Cederhjelmia 38
radiferum, Histiostoma 64
rakha, Histiostoma 65
ramozae, Scolianoetus 97
recondita, Bonomoia 37
reductus, Myianoetus 94
reticulatissimum, Histiostoma 65
reticulatus, Anoetus 32
reticulata, Bonomoia 37
reticulofoveolatum, Histiostoma 65
retractus, Anoetus 32
rettenmeyeri, Pteranoetus 81
rettenmeyerorum, Histiostoma 65
Rhaphidothrix 81
Rhopalanoetus 81
rhynchophorus, Curculanoetus 17
Richardanoetus 83
rimosa, Histiostoma 65
rimosum, Histiostoma 65
rohaceki, Myianoetus 94
rizkii, Histiostoma 65
rostrata, Histiostoma 65
rostratum, Histiostoma 65
rostro-serratum, Histiostoma 49
rostroserratum, Histiostoma 49
rostro-serratus, Histiostoma 49
rostro-serratus, Rhizoglyphus 49
rostro-serratus, Tyroglyphus 49
ruehmi, Anoetus 66
ruehmi, Histiostoma 66
rugosa, Bonomoia 37
rugosum, Histiostoma 66
rwandanus, Myianoetus 94
sachsi, Anoetus 66
sachsi, Histiostoma 66
samiensis, Anoetus 32
sammari, Histiostoma
samsinaki, Kanoetus 75
sapromyzarum, Anoetus 66
sapromyzarum, Histiostoma 66
sapromyzarum, Hypopus 66
Sarraceniopus 84
sarrai, Histiostoma 66
schauenbergi, Histiostoma 66
scheucherae, Myianoetus 94
scheucherae, Scolianoetus 97
scheucheri, Myianoetus 94
Scheucheria 84
schoutedeni, Histiostoma 67
scindita, Histiostoma 67
scinditum, Histiostoma 67
Scolianoetinae 96
Scolianoetus 96
Scutanoetus 84
Sellea 85
Semianoetus 85
serrata, Histiostoma 32
serratus, Anoetus 32
setipes, Anoetus 94
setipes, Hypopus 94
setipes, Myianoetus 94
sextoni, Histiostoma 67
seychellensis, Creutzeria 17
shamyi, Anoetus 33
shanensis, Anoetus 33
sharae, Anoetus 33
shiramba, Histiostoma 67
sibirica, Bonomoia 79
sibirica, Probonomoia 79
simile, Histiostoma 67
106
Acarorum Catalogus IX
simplex, Myianoetus 94
simplex, Rhaphidothrix 81
simplex, Rhopalanoetus 83
simunek, Myianoetus 94
sinuosus, Anoetus 33
siphunculinae, Histiostoma 67
solimani, Histiostoma 67
solomoniensis, Anoetus 33
sordida, Anoetus 33
sovat, Myianoetus 94
sphaerocerae, Bonomoia 16
sphaerocerae, Capronomoia 16
spiceri, Traskorchestianoetus 87
Spinanoetinae 97
Spinanoetini 97
Spinanoetus 85
spinifera, Bonomoia 79
spinifera, Probonomoia 79
spinifera, Prowichmannia 79
spinifera, Wichmannia 79
spinifera, Histiostoma 79
spiniferum, Prowichmannia 79
spiniferus, Anoetus 79
spiniferus, Wichmannia 79
spinipes, Anoetus 33
spinitarsus, Anoetus 68
spinitarsus, Histiostoma 67
spinosus, Anoetus 34
stammeri, Anoetus 68
stammeri, Histiostoma 68
stammeri, Wichmannia 79
stammeri, Prowichmannia 79
Stercoranoetus 86
strenzkei, Histiostoma 68
striata, Histiostoma 34
striatosimile, Histiostoma 68
striatus, Anoetus 34
subsimile, Histiostoma 68
sumatrensis, Anoetus 34
sylvestris, Anoetus 68
sylvestre, Histiostoma 68
Synanoetus 86
Syringanoetus 86
szaboi, Myianoetus 95
szelenyii, Anoetus 68
szelenyii, Histiostoma 68
szentivanyi, Histiostoma 69
szunyoghyi, Anoetoglyphus 23
szunyoghyi, Glyphanoetus 20
szunyoghyi, Myianoetus 95
talpae, Wichmannia 79
talpae, Prowichmannia 79
tanypus, Histiostoma 69
tarsalis, Histiostoma (nomen dubium) 69
tauffliebi, Anoetus 34
Teinokyra 87
teinophallus, Aphodanoetus 36
telatum, Histiostoma 69
Tensiostoma 87
tesselata, Histiostoma 69
thaleri, Myianoetus 95
tibetana, Myianoetus (nomen dubium)
tienhoveni, Anoetus 82
tienhoveni, Histiostoma 82
timidum, Histiostoma 69
tinydorsalis, Histiostoma 69
titanolabi, Histiostoma 69
tobaica, Creutzeria (nomen dubium) 17
topali, Anoetus 34
toxici, Anoetus 34
toxopei, Anoetus 70
toxopei, Histiostoma 70
Traskorchestianoetus 87
travei, Myianoetus 95
trichophorum, Histiostoma 70
trichophorus, Anoetus 70
trichophorus, Histiostoma 70
trogicola, Histiostoma 83
trogicola, Rhopalanoetus 83
tropicus, Anoetus 76
tropicus, Munduytia 76
trossis, Capronomoia 16
Acarorum Catalogus IX 107
tuerkorum, Myianoetus 95
tunesiensis, Anoetoglyphus 23
tuomikoskii, Myianoetus 95
turcastanae, Anoetus 34
ulmi, Anoetus 70
ulmi, Histiostoma 70
ulomae, Kaszabanoetus 75
umbrosa, Histiostoma 70
uncinatus, Myianoetus 95
undulatus, Myianoetus 95
unidentatum, Histiostoma 70
urcerosus, Anoetus 35
uvafer, Histiostoma 70
uxoria, Histiostoma 71
valida, Histiostoma 71
validum, Histiostoma 71
vandenberghi, Syringanoetus 87
varia, Anoetus 71
varia, Histiostoma 71
varium, Histiostoma 71
variabilis, Histiostoma 71
variabilis, Spinanoetus 86
variolatus, Anoetus 35
veliaphilum, Tensistoma 87
venustissimus, Anoetus 20
venustissimus, Glyphanoetus 20
verruca, Histiostoma 71
verrucogenicula, Histiostoma 71
vesparum, Myianoetus 96
vestigialis, Xenanoetus 88
vilhenaorum, Anoetus 35
virgatus, Myianoetus 96
virginiana, Anoetus 35
virginiana, Histiostoma 71
virginianum, Histiostoma 71
virkkii, Ancyranoetus 21
vitzthumi, Anoetus 35
vitzthumi, Histiostoma 35
volgini, Histiostoma 71
vulpis, Hypopus 72
vulpis, Histiostoma 72
warreni, Nepenthacarus 76
wastis, Anoetus 35
watanabei, Histiostoma 72
wauense, Histiostoma 72
wautersi, Ovanoetus 77
weingaertnerae, Spinanoetus 86
wetzeli, Otanoetus 77
Wichmannia 79 (praeoccup.)
wooleyi, Histiostoma 86
woolleyi, Stercoranoetus 86
Xenanoetus 88
zaheri, Histiostoma 72
zdena, Myianoetus 96
zoheirii, Bonomoia 37
Zschachia 39
Zwickia 88
108
Acarorum Catalogus IX
Distribution of Histiostomatidae
EUROPE
Austria (Vitzthum, 1926a, 1927; Scheucher, 1957; Mahunka, 1972b; Samšinák, 1958, 1962,
1989) – Anoetoglyphus copridis, Histiostoma feroniarum, H. gladiger, H. gordius, H.
humiditatis, H. julorum, H. vitzthumi, Myianoetus thaleri, M. zdena
Belgium (Cooreman, 1963; Fain et al., 1993) – Anoetus pini, Conglanoetus denticulatus,
C. puteanus, Myianoetus tuercorum, Histiostoma gladiger, H. gordius, H. trogicola
Bulgaria (Samšinák, 1958; Beron, 1973; Takov, Pilarska & Moser, 2009) – Histiostoma
ovale (= gladiger), H. piceae, Prowichmannia spiniferum
Croatia (Pernek et al. 2008; Pernek et al. 2012; Wirth & Pernek. 2012; Wirth et al., 2016)
– Histiostoma piceae, H. ulmi
Czech Rep. (Müller, 1860; Samšinák, 1956, 1958, 1984, 1989) – Capronomoia sphaerocerae, Histiostoma manica, H. ovale, Myianoetus simplex, M. virgatus, Probonomoia
humprechti
Denmark (Jensen, 1895; Haarløv, 1957) – Histiostoma berghii, H. julorum
Greenland (Trägårdh, 1905) – Myianoetus digitiferus
Finland (Mahunka, 1969, 1972b; Nuorteva, 1971; Penttinen et al., 2013) – Histiostoma
sp., Myianoetus muscarum, M. parvus, M. tuomikoskii, Probonomoia pini, Xenanoetus
vestigialis
France (contin.) (Dufour, 1839; Gervais, 1844; Theodorides¸1955; Cooreman, 1954 ;
Samšinák, 1958; Levieux et al., 1989; Fain & Beaucournu, 1973; Mahunka, 1972b,
1977) – Anoetus dolifer, Histiostoma feroniarum, H. gervaisi, H. gladiger, H. piceae,
H. sapromysarum, Myianoetus dionychus [= diadematus], M. simplex, Psyllanoetus
melesicola
Corsica (Samšinák, 1958; Wirth, 2005) - Histiostoma ovale (= gladiger), Probonomoia
opuntiae
Germany (Koch, 1843; Kramer, 1876, 1886; Oudemans, 1902; Willmann, 1937; Scheucher,
1957; Fain & Erteld, 1998; Wirth, 2003) – Anoetus myrmicarum, neglectus, A. pini,
Capronomoia sphaerocerae, Glyphanoetus phyllotrichus, G. venustissimus, Histiostoma crypturgi, H. gladiger, H. gordius, H. halicticolum, H. julorum, H. oudemansi,
H. palustre, H. piceae, H. phyllophorum, H. polypori, H. sapromyzarum, H. stammeri,
H. strenzkei, H. trichophorum, H. ulmi, H. vitzthumi, Myianoetus dionychus [= diadematus], Probonomoia pini, Prowichmannia spiniferum, Rhopalanoetus lanceocrinus,
Sellea pulchrum, Spinanoetus pelznerae, S. weingaertnerae
Great Britain (Michael, 1901; Fain, Britt & Molyneux, 1980; Behura, 1950; Hyatt, 1990)
– Histiostoma polypori, Myianoetus copromyzae, Prowichmannia spiniferum, Selea
pulchrum
Greece (Mahunka, 1972e, 1977) – Anoetus hauseri, A. samiensis, Histiostoma feroniarum,
H. sapromyzarum, H. ionicum, H. matsakisi
Acarorum Catalogus IX 109
Kephallinia (Mahunka, 1971, 1972e) – Anoetus hauseri, A. lunulatus, A. sapromyzarum,
A. samiensis, Hauserania graeca, H. ornata, Histiostoma feroniarum, H. insulare
Zakynthos (Mahunka, 1972e) - Histiostoma feroniarum,
Levkas (Mahunka, 1972e) - Histiostoma sapromyzarum,
Holland (Oudemans, 1914; Bongers, OConnor & Lukoschus, 1985; Fain & Lambrechts,
1987) – Aphodanoetus teinophallus, Histiostoma brevimanum, H. insulare, H. ocellatum, H. phyllophorum, H. polypori, H. sapromyzarum, Myianoetus simplex, Rhopalanoetus lanceocrinus
Hungary (Mahunka, 1961, 1962a, 1962b, 1963d, 1968, 1970, 1972b, 1976, 1977b, 1978;
Mahunka & Eraky, 1987; Samšinák, 1989) – Anoetoglyphus copridis, Anoetus dudichi,
A. myrmicarum, A. pannonicus, A. pini, Capronomoia sphaerocerae, Glyphanoetus
longispina, Histiostoma baloghi, H. dactyloscopicum, H. disiunctum, H. gordius, H.
histeri, H. hungaricum, H. lineatula, H. loksai, H. magdolnae, H. margaritatum, H.
ovalis, H. polypori, H. sapromyzarum, H. serrata, Myianoetus clavatus, M. dionychus
(= diadematus), M. heterotrichus, M. mystacis, M. pappi, M. parvus, M. rohaceki, M.
simplex, M. simunek, M. szaboi, M. tuomikoskii, M. zdena, Peripatetes acanthotarsus,
Probonomoia pini, P. humprechti, Prowichmannia spiniferum, Rhopalanoetus fimetarius, Semianoetus mrcziaki, Spinanoetus heterotrichus, S. pelznerae, S. weingaertnerae,
Stercoranoetus woolleyi
Iceland (Sellnick, 1940) – Anoetus notabilis
Ireland (Curry & Momen, 1988) - Histiostoma feroniarum
Italy (contin.) (Canestrini & Berlese, 1881; Berlese, 1881, 1883, 1918; Oudemans, 1927)
– Anoetoglyphus ateuchi, Glyphanoetus phyllotrichus, G. venustissimus, Histiostoma
feroniarum, H. julorum, H. pyriforme, Rhopalanoetus fimetarius
Sicily (Berlese, 1910; Samšinák, 1962) – Anoetoglyphus ateuchi, Anoetus curtipes, A.
privorai, Histiostoma brevipodum
Sardinia (Wirth, 2005) – Probonomoia opuntiae
Norway (Fain et al., 1993) – Conglanoetus denticulatus
Poland (Chmielewski, 1984; Kiełczewski & Wiśniewski, 1980; Michalski & Ratajczak, 1989;
Haitlinger, 2002, 2004; Witaliński, Rożej-Pabijan & Podkowa, 2014) – Histiostoma
crypturgi, H. ovalis, H. piceae, H. polypori, Spinanoetus pelznerae, S. weingaertnerae,
Probonomoia pini, P. spinifera,
Portugal (cont.) (Vissa et al., 2019) – Histiostoma ovatus
Romania (Cooreman, 1941; Mahunka, 1970, 1977b) – Histiostoma gordius, H. histeri, H.
lineatula, H. margaritatum, H. ovalis, Myianoetus tuomikoskii, Probonomoia humprechti, Xenanoetus vestigialis
Russia (all, continent.) (Vitzthum, 1026; Sevastianov, 1973; Krivoluckiy, 1975; Wirth,
1916; Khaustov, Trach & Bobylev, 2016; Khaustov et al., 2018) – Anoetus exornatus,
Bonomoia recondita, Histiostoma gladiger, H. ovale, H piceae, H. pini, Probonomoia
pini, P. sibirica
Crimea (Krivoluckiy, 1975)
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Sakhalin (Sevastianov, 1973) - Myianoetus pseudomuscarum
Kuril Islands (Klimov, 1998) – Scheucheria sp.
Slovakia (Müller, 1859; Mahunka, 1972b, Samšinák, 1984, 1989; Mangová et al., 2017;
Kaluz, 1992; Mašan & Országhi, 1995; Mašan, 2000; Mašan & Krištofik, 1992; Zamec,
2013, 2014) – Bonomoia recondita, Capronomoia sphaerocerae, Histiostoma crypturgi,
H. berghi, H. crypturi, H. dryocoeti, H. feroniarum, H. maritimus, H. myrmicarum, H.
ovalis, H. pannonicus, H. piceae, H. pini, H. polypori, H. pulchrum, H. sapromyzarum,
Myianoetus diadematus, M. dionychus, M. longisetosus, M. ovatus, M. parvus, M. sovat,
M. tuerkorum, M. tuomikoskii, M. virgatus, M. zdena, Rhopalanoetus lanceocrinus,
Spinanoetus heterotrichus, S. pelznerae, S. weingaertnerae
Slovenia (Moraza, Fernandez & Jurc, 2013) – Histiostoma ovale
Spain (cont.) (Dimitrieff & Fain, 1994; Rebolledo & Arroyo, 1996; Fernandez, Diez &
Moraza, 2013) – Histiostoma feroniarum, Probonomoia chrysis
Canary Islands (Sanchez-Covisa et al., 1999; Wirth, 2005) – Probonomoia opuntiae
(Gomera), Histiostoma feroniarum (Tenerife)
Sweden (Moser et al. 1989; Moser, Perry et al., 1989) – Histiostoma piceae
Switzerland (Mahunka, 1977; Fain, 1980) – Anoetus dolifer, Hormosianoetus aeschlimanni
Turkey (all) (Mahunka, 1976) – Kaszabanoetus ulomae
Ukraine (Sevastianov, 1970, 1971, 1973; Sevastianov & Vysotskaya, 1971) – Anoetus
helobius, A. hospitus, A. opertus, A. parmatus, A. retractus, Bonomoia picturata, B.
probata, Histiostoma indetonsum, H. necopinum, Myianoetus microti, M. paganus,
M. pseudomuscarum, Prowichmannia spinifera
AFRICA
Angola (Mahunka, 1963a, 1963b, 1967) – Anoetus machadoi, A. striatus, A. vilhenaorum, Anoetoglyphus (?) bifastigatus, Glyphanoetus brevisetus, Histiostoma andrei, H.
angolensis, H. arcuatum, H. bacilligerum, H. clavatum, H. impressum, H. minimum,
H. pontigerum, H. puncaticeps, H. simile, H. subsimile, Hymenanoetus nodosus, Porrhanoetus machadoi
Botswana (Camerik, Lukoschus & Kniest, 1980) – Fibulanoetus longitarsus, F. mahunkai
Cabo Verde (Mahunka & Mahunka-Papp, 1991) – Glyphanoetus phyllotrichus, Histiostoma
phyllophorum, Stercoranoetus harteni
Cameroon (Mahunka, 1979b) - Histiostoma amieti, H. camerunensis, H. chilopophila,
H. variabile
Congo Rep. of the (Brazzaville) (Mahunka, 1965, 1967a;) – Anoetoglyphus africanus, A.
(?) immaturus, Anoetus collis, A. congoensis, A. coprophilus, A. impressus, A. longisetus, A. machadoi, A. pauliana, A. punctulatus, A. striatus, A. tauffliebi, Glyphanoetus
africanus, Rhaphidothrix gozmanyi, Rhopalanoetus bifurcatus
Congo DR (Oudemans, 1929; Fain & Elsen, 1972; Fain & Beaucournu, 1973; Fain, 1974;
Samšinák, 1962, 1989) – Anoetus fetters, Chiloanoetus mayumbensis, Curculanoetus
rhynchophorus, Histiostoma conclaviculum, H. schoutedeni, Probonomoia congoen-
Acarorum Catalogus IX 111
sis, Psyllanoetus afropulicarius, Rhopalanoetus bifurcatus, Scolianoetus scheucherae,
Xenanoetus kabrt
Egypt (Yousef & Metwally, 1973; Metwali & Ahmad, 1987; Hassan & Sevastianov, 1991;
Eraky & Shoker, 1993a, 1993b, 1994, 1995; Eraky & Nasser, 1993; Eraky, 1993, 1994a,
1994b, 1997, 1998, 1999a, 1999b, 1999c, 2000, 2003; Abu El-Maged, 1998; AbdelAziz, 1999; Eraky, Abdel-Galil & Bohibah, 2010; Fawzy & Mtewallii, 2007; Eraky,
Abdelgayed, Negm, Helal & Moussa, 2017; Abdelgayed et al., 2019; Eraky, AbdelAziz & Abo-Elmaged, 2020; Desoky et al., 2020) – Anoetus caljubiensis, A. irinus, A.
moshtohorensis, A. sharae, A. wastis, Bonomoia zoherii, Coprobonomoia mahunkai,
C. sphaerocerae, Glyphanoetus mahunkai, G. omari, G. processum, Histiostoma alii,
H. arcuatus, H. bananai, H. camphori, H. cataglyphi, H. darwishii, H. diadematum,
H. egyptiacum, H. essami, H. farghali, H. herbali, H. irinum, H. mannai, H. manuri,
H. nasseri, H. negmi, H. onioni, H. pickaxei, H. rizkii, H. sammari, H. sarrai, H. solimani, H. rizkii, H. tinydorsalis, H. zaheri, Hormosianoetus mahunkai, Myianoetus
granatum, M. lili, M. manuri, M. nerminka
Fiji (Mahunka, 1992) – Histiostoma pseudolongipes
Gambia (Mahunka, 1975, 1979) - Anoetoglyphus szunyoghyi, Ghanoetus marmoreus,
Histiostoma endroedii, H. fucatifer, H. gambiense, H. umbrosa
Ghana (Mahunka, 1973a, 1973d, 1974a, 1975, 1979) – Anoetus heliocopridis, A. sinuosus,
Anoetoglyphus antennatus, A. globulus, A. latus, A. polygonatus, Bonomoia rugosa,
Histiostoma distentosetum, H. editum, H. endroedii, H. furcatum, H. gorgo, H. rugosum,
H. striatosimile, Rhaphidothrix punctolineatus, Rh. simplex, Rhopalanoetus hirsutus,
Spinanoetus variabilis, Teinokyra lineata
Ivory Coast (Fain, 1974) – Anoetus humilis, A. macrosuctus, Histiostoma adsimile, H.
alphaticum, Glyphanoetus eboris
Kenya (Mahunka, 1979) – Histiostoma bernoni, Myianoetus africanus
Madagascar (Fain & Pauly, 2001) – Histiostoma heriades
Morocco (Fain, Athias-Binche & Carton, 1993) – Hormosianoetus laboratorium
Namibia (Fain, 1974) – Loxanoetus bassoni
Rwanda (Fain, 1980, 1984) – Myianoetus brevis, M. rwandanus, M. uncinatus, Syringanoetus vandenberghi, Xenanoetus afrotropicalis,
Seychelles (Nesbitt, 1979) – Creutzeria seychellensis
South Africa (Fain, 1970; Mahunka, 1979; ) – Anoetoglyphus lootsi, Histiostoma volgini,
Loxanoetus bassoni, Rhopalanoetus endroedyyoungai
Tanzania (Oudemans, 1912, 1914; Mahunka, 1961, 1969b, 1974a, 1979) – Anoetus heliocopridis, A. longitubus, A. reticulatus, Glyphanoetus flagellates, G. longus, G. szunyoghyi,
Histiostoma bifoveolatum, H. phyllophorum, H. polaki, H. telatum, H. trichophorum,
Myianoetus reductus, Rhopalanoetus bifurcates, Rh. gladifer
Tunisia (Mahunka, 1981) – Anoetoglyphus tunesiensis
Uganda (Fain, 1974; Fain & Zumpt, 1974) – Auricanoetus longitarsus, Loxanoetus bassoni, Otanoetus wetzeli
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Acarorum Catalogus IX
Indian Ocean Islands (Mahunka, 1978) – Histiostoma schauenbergi
ASIA
Afghanistan (Mahunka, 1974c) – Histiostoma szelenyii
China (Samšinák, 1962b; Li et al., 1992; Wang et al., 2002; Fan, Chen & Wang, 2010)
– Histiostoma feroniarum, H. formosanum, Rhopalanoetus chinensis, Rh. simplex,
Seliea pulchrum
Taiwan (Mahunka, 1976; Phillipsen, 1977; Tseng Y.-H., 1982, 1984) – Histiostoma formosanum, H. gampsoceri, H. humiditatis
India (Vitzthum, 1926; Chinniah & Mohanasundaram, 1995) – Histiostoma coprae, H.
himalayae, H. muscae, H. onthophagae H. siphunculinae
Indonesia (Oudemans, 1903, 1911, 1914; Vitzthum, 1921, 1931, 1932; Mahunka, 1976) –
Histiostoma banjuwangicus, H. campanula, H. cyrtandrae, H. indicus, H. prodectoris,
Munduytia tropicus, Scolianoetus javanus, Zwickia colocasiae
Iran (Lotfollahi, Haddad Irani Nejad, Bagheri, Valizadeh, 2010) – Histiostoma polypori,
H. litorale
Israel and Palestina (Costa, 1967; Mahunka, 1979a) – Histiostoma feroniarum, H. loebli
Japan (Kurosa, 2000, 2002; Takami & Kuwahara, 2005; Tagami, 2016; Kurosa & Tagami,
2006) – Histiostoma globotuberum, H. mahunkai, H. piscium, H. watanabei, Probonomoia pini
Malaysia (Vitzthum, 1921; Hirst, 1928; Kurosa, 1976; Wurst & Kovac, 2003; Azmiera,
Mariana & Heo C.C., 2019) – Anoetus myrmicarum, A. virginiana, Histiostoma sp.,
Tensiostoma veliaphilum
Mongolia (Mahunka, 1967c, 1969c, 1972b, 1973) – Anoetus cacuminatus, A. insulanus, A.
kaszabi, A. microti, A. porosus, Histiostoma insulare, Myianoetus kaszabi, Myianoetus
tuomikoskii, Scheucheria mongolica, Xenanoetus grandiceps
Pakistan (Khan, 1969; Chauhdri, 1986; Ashfaq, 1986, 1988, 2001; Ashfaq, Chauhdri
& Aheer, 1985; Ashfaq & Chauhdri, 1988; Sarwar, Ashfaq & Islam, 1998; Ashfaq,
Sarwar & Akbar, 1999; Ashfaq, Sarwar & Ali, 2000; Ashfaq, Sarwar & Amiad, 2000;
Ashfaq & Sarwar, 2001) – Capronomoia asper, C. bellulus, C. gradates, C. pango, C.
trossis, Glyphanoetus incanus, G. leptos, G. micidus, Histiostoma caro, H. crantor, H.
densus, H. edax, H. erigo, H. fortis, H. fragosus, H. galerus, H. gracilis, H. imbecillum,
H. junctus, H. karachiensis, H. luxus,
Philippines (Vitzthum, 1921) - Histiostoma sapromyzarum
Saudi Arabia (Samšinák, 1979; Negm & Alatawi, 2011) – Capronomoia sphaerocerae,
Histiostoma pickaxei, H. camphori, Myianoetus lili
Singapore (Hirst, 1928) – Zwickia nepenthesiana
Sri Lanka (Oudemans, 1915; Mahunka, 1974d) – Anoetus aelleni, A. angularis, A. ceylanensis, Ceylanoetus excavatus, Zwickia guentheri
Syria () – Histiostoma [Anoetus] sapromysarum, Caprononomoia [Bonomoia] sphaerocerae
Taiwan (Mahunka, 1976b) – Histiostoma gampsoceri
Acarorum Catalogus IX 113
Thailand (Fain & Lambrechts, 1985; Fain, Athias-Binche & Carton, 1993; ) – Histiostoma
piscium, H. unidentatum
Vietnam (Mahunka, 1967d, 1978; Fain & Beaucournu, 1972) – Anoetoglyphus langi,
Anoetus capillatus, A. decorus, A. longipes, A. ornatus, A. pocsi, A. spinosus, Bonomoia
reticulata
NORTH AMERICA
Canada (Woodring & Moser, 1970; Lindquist et al., 1979; Mullen et al., 1989; Fain & Colloff, 1990; Walter & Latonas, 2012; OConnor et al., 2015; Achtymichuk, 2017; Baulieu
et al., 2019) – Anoetus sordida, Bonomoia sp., Histiostoma halictonida, Myianoetus
muscarum, Traskorchestianoetus brevipes, T. spiceri. According to Baulieu et al., 2019,
in Canada are known 21 sp. of Histiostomatidae and the number expected is 80.
Mexico (Woodring & Moser, 1970) – Histiostoma varium
USA (Banks, 1906, 1910, Ewing, 1911; Hughes & Jackson, 1958; Cross, 1968; Woodring,
1971; Moser & Roton, 1971; Woodring & Moser, 1970, 1975; Moser, 1976; Hill &
Deahl, 1978; Fain & Philips, 1979; Mahunka, 1982; Fashing & OConnor, 1984;
Samšinák, 1989; Wirth & Moser, 2010; Uppstrom & Klompen, 2011; Oconnor et al.,
2015; Pimsler et al., 2016; Achtymichuk, 2017) – Ameranoetus hematobii, Anoetus
cossi, A. grapefruit, A. media, A. sordida, Capronomoia sphaerocerae, Glyphanoetus
nomiensis, Histiostoma bakeri, H. blomquisti, H. conjunctum, H. gracilipes, H. brevipes, H. guanophilum, H. halictonida, H. heinemanni, H. inquilinus, H. murchiei, H.
nigrellii, H. pauciputeolum, H. pilosetum, H. phytophaga, H. protuberans, H. sextoni,
H. tarsalis, H. varium, H. virginianum, Myianoetus micromaculatus, M. mlkoch, M.
muscarum, M. rohaceki, Sarraceniopus darlingtoniae, Xenanoetus kofrankova
SOUTH and CENTRAL AMERICA
Argentina (Mahunka, 1963c, 1963d) – Anoetus akodoni, Histiostoma [Anoetus] sapromysarum
Bolivia (Vitzthum, 1921) – Histiostoma sapromyzarum
Brazil (Vitzthum, 1926a; Fain, 1974; Fain & Camerik, 1978; Mahunka, 1982; Souza,
Bernardi & Ferreira, 2018) – Bothyanoetus pereirai, Histiostoma acrocini, H. sapromyzarum, H. tanypus, Scolianoetus brasiliensis, S. ramozae, Spartocera cinammomea
Chile (contin.) (Mahunka, 1963, 1972d) – Anoetus castrii, A. chilensis, A. cirrosus, Synanoetus baloghi
Juan Fernandez Islands (Mahunka, 1982) – Glyphanoetus pappi
Colombia (Fain & Rack, 1987) – Probonomoia columbiana
Costa Rica (Woodring, 1973; Mahunka, 1976, 1978a, 1979) – Histiostoma adarcuale,
H. claviger, H. coronatum, H. duplex, H. eickworthi, H. marianae, H. moralesi, H.
perruptum, H. rostratum, Pteranoetus incertus, Semianoetus banksi
Cuba (Samšinák, 1966, Mahunka, 1978) – Histiostoma cubanum, H. meszarosorum,
Kanoetus cubanus
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Acarorum Catalogus IX
Ecuador (contin.) (Mahunka, 1978; Samšinák, 1989) – Anoetus helena, Conglanoetus
ornatofoveolatus, Histiostoma capillatum, H. continuatum, H. diadematum, H. elzingai, H. rettenmeyerorum, H. rimosa, H. scinditum, Pteranoetus latus, P. rettenmayeri,
Richardanoetus demel
Guadeloupe (Fain, Athias-Binche & Carton, 1993) – Hormosianoetus laboratorium
Guatemala (Woodring & Moser, 1975) – Histiostoma conjunctum
Honduras (Woodring & Moser, 1970, 1975) – Histiostoma conjunctum, H. varium
Panama (Mahunka, 1978a) – Histiostoma incurvatum
Peru (Caceres & Fain, 1975) – Histiostoma ferroniarum
Puerto Rico (Fain & Santiago-Blay, 1993) – Ancyranoetus virkkii
St. Vincent Republic, Grenadines Isl., Bequia Id. (Windward Group)(Fain & Van
Goethem, 1985) – Ovanoetus wautersi
Windward Islands (St. Lucia) (Mahunka, 1981) – Histiostoma antillense, H. mirabile
OCEANIA
Australia (continent) (Womersley, 1941, 1942; Mahunka, 1975; Fain & Domrow, 1980;
Halliday, 1998; Halliday & Collins, 2002; Tagami & Halliday, 2013) – Chiropteranoetus
chalinolobus, Histiostoma australiense, H. feroniarum, H. humiditatis, H. papillata, H.
phyllophorum (= nichollsi Womersley), H. sapromyzarum, H. titanolabi, H. unidentatum, H. varium, Loxanoetus lenae, Myianoetus dycei, M. oudemansi
New Zealand (Fain & Galloway, 1993; Qin & Rohitha, 1996; Clark, 2009, 2010a, 2010b)
– Histiostoma feroniarum, H. mantelli, Myianoetus antipodus
Papua New Guinea (Fain, 1968; Mahunka, 1975, 1976, 1978) – Histiostoma amargynus,
H. foveosimile, H. lineosculpturatum, H. reticulatissimum, H. reticulofoveolatum, H.
szentivanyi, H. wauense, Myianoetus domrowi
French Polynesia (Tahiti) (Mahunka, 1992) – Histiostoma berndhauseri
French Polynesia (Hivaoa) (Vitzthum, 1935) – Histiostoma granulatum
Fiji (Mahunka, 1995,1992) – Anoetus peloponesiacus, Histiostoma genavensium, H. pseudolongipes
Marquesan Islands (Vitzthum, 1935) – Histiostoma granulatum
Salomon Islands (Mahunka, 1972c) – Anoetus bradymeri, A. horridus, A. solomoniensis,
A. toxici
ANTARCTICA
Kerguelen Islands (Fain, 1976) – Amyzanoetus halophilus, Austranoetus kerguelenensis,
Histiostoma kerguelenense, Myianoetus travei
Acarorum Catalogus IX 115
Hosts of Histiostomatidae
Hirudinea
Fam. Aulostomidae
Aulastoma gulo (cocoons) – Histiostoma berghi
Crustacea
Amphipoda
Fam. Talitridae
Traskorchestia traskiana – Traskorchestianoetus brevipes
Isopoda
Fam. Oniscidae
Oniscus asellus – Histiostoma phyllophorum
Oniscus sp. – Histiostoma feroniarum
Decapoda
Fam. Paguridae
Coenobita clypeata – Ovanoetus wautersi
Arachnida
Acari
Fam. Gamasidae
Eugamasus cornutus – Histiostoma polypori
Myriapoda
Diplopoda
Fam. Iulidae
Julus scandinavius - Histiostoma feroniarum, H. litorale, H. sapromyzarum
Chilopoda
Fam. Lithobiidae
Lithobius forficatus – Histiostoma gervaisi
Fam. Scolopendridae
Cormocephalus aurantiipes - Histiostoma feroniarum
Insecta
Orthoptera
Fam. Phalangopsidae
Endecous (Endecous) alejomesai – Histiostoma sp. (cave)
Coleoptera
Fam. Carabidae
Demetrias imperialis – Histiostoma disiunctum, H. serratum
Dichaetodrilus incrassatus - Histiostoma striatum
Fam. Brenthidae
Ectocemus cinnamomeus – Histiostoma prodectoris
Eubactrus semiaeneus – Anoetus polynesiacus
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Acarorum Catalogus IX
Prodector fruhstorferi – Histiostoma prodectoris
Prophthalmus potens – Histiostoma prophthalmi
Schizotrachelus bakeri – Histiostoma sapromyzarum
Fam. Platypodidae
Platypus dispar - Histiostoma crassipes
Fam. Cerambycidae
Acanthocinus aedilis – Probonomoia humprechti, P. pini
Acrocinus longimanus – Histiostoma acrocini
Rhagium sp. – Histiostoma crypturgi
Fam. Chrysomelidae
Galeruca sp. – Histiostoma necopinus
Lema nigripes – Ancyranoetus virkkii
Fam. Coccinellidae
Gen. sp. – Histiostoma lorentzi
Fam. Curculionidae
Curculioninae s.l.
Rhynchophorus phoenicis – Anoetoglyphus bifastigatus, Curculanoetus rhynchophorus, Histiostoma striatum
Scolytinae
Crypturgus pusillus – Histiostoma crypturgi
Dendroctonus brevicomis - Bonomoia certa
D. frontalis - Histiostoma conjunctum
D. micans – Histiostoma vitzthumi
D. pseudotsugae - Bonomoia certa
Dryocoetus autographus – Histiostoma dryocoeti
Dryocoetes villosus – Anoetus pannonicus
Hylaster ater – Probonomoia pini
Ips avulsus - Histiostoma conjunctum
I. cembrae – Histiostoma oudemansi
I. curvidens – Histiostoma abietis
I. grandicollis – Histiostoma varium
I. laricis – Histiostoma gordius
I. sexdentatus – Histiostoma ovalis (= gladiger)
I. typographus – Histiostoma piceae
Mycetophilus piniperda – Probonomoia pini
Polygraphus minor - Histiostoma himalayae
Scolytus ventralis – Bonomoia certa
Fam. Elateridae
Melanotus rufipes – Histiostoma ruehmi, H. sapromyzarum
Fam. Histeridae
Carcinops 14-striata – Histiostoma hungaricum
Carcinops sp. – Anoetus sordida
Acarorum Catalogus IX 117
Hister merdarius – Histiostoma feroniarum
H. stercorarius - Histiostoma feroniarum
H. unicolor - Histiostoma feroniarum
Hololepta africanae - Histiostoma crassipes
Platysoma compressum – Histiostoma histeri, H. lineatula, H. margaritatum,
H. ovale
Fam. Hydrophilidae
Hydrophilus caraboides – Histiostoma maritimus
Sphaeridium sp. - Rhopalanoetus lanceocrinus, Xenanoetus vestigialis
Fam. Lucanidae
Dorcus parallelopipedus – Histiostoma dryocoeti, Probonomoia pini
Fam. Nitidulidae
Brachypeplus sp. – Histiostoma granulatum
Fam. Passalidae
Passalus sp. - Scolianoetus ramozae
Fam. Scarabaeidae
Aphodius fimetarius – Rhopalanoetus lanceocrinus
A. fossor – Rhopalanoetus lanceocrinus
A. frater – Aphodanoetus teinophallus
A. merdarius – Rhopalanoetus lanceocrinus
Bothynus deiphobus - Bothyanoetus pereirai
Bubas bubalus - Histiostoma sapromyzarum
Copris lunaris - Anoetoglyphus copridis
Cetonia aurata – Prowichmannia spiniferum
Geotrupes stercorarius - Rhopalanoetus lanceocrinus
Melolontha hippocastani – Histiostoma melolonthae
Oniticellus cinctus – Rhopalanoetus chinensis, Rh. simplex
Onitis sphinx – Rhopalanoetus bifurcatus
Onthophagus pactolus – Histiostoma onthophagae
Oxythyrea funesta – Histiostoma phyllophorum
Polyphyla fullo - Histiostoma melolonthae
Scarabaeidae - Anoetoglyphus langi
Fam. Staphylinidae
Bledius bicornis – Insulanoetus brachypodus
B. sarmaticus - Insulanoetus brachypodus
Emus hirtus - Spinanoetus weingaertnerae
Lathrobium (Lathrobium) sp. – Histiostoma watanabei
Ontolestes sp. - Spinanoetus weingaertnerae
Philontus sp. - Spinanoetus weingaertnerae
Fam. Silphidae
Nicrophorus interruptus - Spinanoetus pelznerae
Oeceoptoma thoracica - Spinanoetus pelznerae
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Acarorum Catalogus IX
Thanatophilus rugosus - Spinanoetus pelznerae
Th. sinuatus - Spinanoetus pelznerae
Fam. Tenebrionidae
Amarygmus [a “Amargynus”, or “Amargymus”] cf. capillatus - Histiostoma
amargynum, H. szentivanyi
Amarygmus [as “Amargynus”] cf. deformipes - Histiostoma reticulofoveolatum
Androsus greensladei – Anoetus bradymeri
Bradymerus leveri - Anoetus bradymeri
Corticeus coynei – Anoetus sordida
Helops quisquilius - Histiostoma ulmi
Hypophloeus bicolor – Histiostoma ulmi
Leiochrodes suturalis – Anoetus horridus, A. solomoniensis
Platolenes femoratus – Histiostoma wauensis
Platolenes sp. - Histiostoma lineosculpturatum
Prioscelis serrata – Chiloanoetus mayumbensis
Scyophagus piceus - Anoetus horridus
Toxicum quadricorne rufipes – Anoetus toxici
Uloma culinaria – Kaszabanoetus ulomae
Fam. Trogidae
Trox scaber – Rhopalanoetus trogicola
T. cadaverinus - Rhopalanoetus trogicola
Dermaptera
Fam. Labiduridae
Labidura riparia – Anoetus moshtohorensis, A. wastis, Histiostoma camphori
Fam. Anisolabididae
Anisolabella (Gonolabis) marginalis – Histiostoma mahunkai, H. piscium
A. maritima – Histiostoma piscium
Titanolabis colossea – Histiostoma australiense, H. humiditatis, H. titanolabi
Fam. Forficulidae
Forficula auricularia – Histiostoma polypori
Heteroptera
Fam. Coreidae
Spartocera cinammomea - Scolianoetus brasiliensis
Fam. Veliidae
Lathriovelia rickmersi – Tensiostoma veliaphilum
L. capitata - Tensiostoma veliaphilum
Hymenoptera
Fam. Megachilidae
Heriades erythrosoma – Histiostoma heriades
Fam. Apidae
Xenoglossa strenua – Histiostoma inquilinus
Acarorum Catalogus IX 119
Fam. Halictidae
Augochlora spp. – Histiostoma eickworthi
Halictus holtzi – Histiostoma szelenyii
H. sexcinctus – Histiostoma halicticolum
H. rubicundus - Histiostoma halictonida
H. tatrazonianellus – Histiostoma orientalis
Megalopta spp. - Histiostoma halictonida
Nomia melanderi – Glyphanoetus nomiensis
N. nortoni - Glyphanoetus nomiensis
Fam. Formicidae
Aphaenogaster testaceopilosa – Anoetus privorai
Camponotus sp. - Histiostoma myrmicarum
Eciton burchelli – Histiostoma incurvatum, Pteranoetus incertus
E. hamatum – Histiostoma clavifer, H. moralesi, H. rostratum
E. lucanoides – Histiostoma elzingai, H. scinditum
Formica fusca - Histiostoma myrmicarum
F. lemani – Anoetus komarpictoris, A. shamyi
F. rufa – Anoetus exornatus
Gymnometopina kibokoensis – Anoetus caljubiensis
Labidus praedator - Conglanoetus ornatofoveolatus, Histiostoma capillatum,
H. rettenmeyerorum
Labidus sp. – Histiostoma continuatum, H. rimosum
Lasius flavus - Histiostoma myrmicarum
L. niger - Histiostoma myrmicarum
Myrmica laevinodis - Histiostoma myrmicarum
M. ruginodis – Anoetus dolifer, Bonomoia recondita, Histiostoma myrmicarum
M. rubra - Anoetus opertus
M. sabuleti – Anoetus dolifer
Neivamyrmex gibbatus – Histiostoma diadematum
N. gradualis – Pteranoetus rettenmeyeri
Solenopsis invicta – Histiostoma blomquisti
Fam. Scoliidae
Campsomeris mansueta – Scolianoetus scheucherae
Fam. Chrysididae
Chrysis sp. – Probonomoia chrysis
Fam. Vespidae
Vespidae gen. sp. – Histiostoma tanypus
Diptera
Fam. Asilidae
Laphria flava – Probonomoia humprechti
Fam. Ceratopogonidae
Culicoides brevitarsis – Myianoetus dycei
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Fam. Chloropidae
Gampsocera mutata - Histiostoma gampsoceri
Scoliophthalmus sp. – Anoetus domrowi
Siphunculina sp. – Histiostoma siphunculinae
Fam. Drosophilidae
Drosophyla bipectinata – Histiostoma unidentatum
D. willstoni – Hormosianoetus laboratorium
D. subobscura - Hormosianoetus laboratorium
Drosophila sp. - Hormosianoetus aeschlimanni
Fam. Glossinidae
Glossina fuscipes quanzensis –Histiostoma schoutedeni, Probonomoia congoensis
Fam. Muscidae
Azelia parva – Myianoetus pappi
Haematobia irritans – Ameranoetus haematobii
Hebecnema umbratica – Myianoetus simplex
Hydrotaea occulta – Myianoetus tuomikoskii
H. tuberculata – Myianoetus tuomikoskii
Hydrotaea sp. – Myianoetus longisetosus, M. ovatus
Musca domestica – Histiostoma muscae
Ophyra leucostoma – Myianoetus parvus
Synthesiomyia nudiseta – Myianoetus muscarum
Fam. Fanniidae
Fannia sp. – Myianoetus mystacis
Fam. Sepsidae
Sepsis thoracica – Myianoetus simplex
Fam. Heleomyzidae
Amoebaleria cacsia - Myianoetus diadematus
A. spectabilis - Myianoetus diadematus
Borboropsis fulviceps - Myianoetus parvus
Gymnomus caesius - Myianoetus sovat
Helomyza brachypterna - Myianoetus diadematus
H. serrata - Myianoetus diadematus
Fam. Sphaeroceridae
Borborillus sordidus - Myianoetus mlcoch
Coproica acutangula – Xenanoetus kofrankova
C lugubris – Myianoetus simplex, M. simunek, M. zdena
C. pallifrons - Myianoetus zdena
C. vagans - Myianoetus zdena
C. viripennis – Myianoetus rohaceki
Elachisoma aterrima - Myianoetus zdena
Gymnometopina kibokoensis – Xenanoetus kabrt
Acarorum Catalogus IX 121
G. magna - Anoetus fetters, Syringanoetus vandenberghi
Ischiolepta pusilla - Capronomoia sphaerocerae
Limosina clunipes – Myianoetus virgatus
L. rufilabris – Myianoetus rohaceki
L. talparum - Glyphanoetus phyllotrichus
Lotobia pallidiventris - Myianoetus zdena
Lotophila atra - Myianoetus rohaceki
Metaborborus calcaratus – Anoetus fetters
Philocoprella quadrispina - Myianoetus zdena
Physiophora sp. - Capronomoia sphaerocerae
Pilocoprella quadrispina - Myianoetus mlcoch
Richardia sp. – Anoetus helena, Richardanoetus demel
Spelobia clunipes - Myianoetus virgatus
Sphaerocera curvipes - Capronomoia sphaerocerae
S. subsultans - Capronomoia sphaerocerae
Fam. Syrphidae
Mallota bautius – Hormosianoetus mallotae
M. posticata - Hormosianoetus mallotae
Fam. Tabanidae
Hybomitra affinis – Bonomoia sp.
H. epistates - Bonomoia sp.
H. illota - Bonomoia sp.
H. lasiophthalmus - Bonomoia sp.
H. nuda - Bonomoia sp.
Tabanus quinquestriatus - Bonomoia sp.
Lepidoptera
Fam. Cossidae
Cossus cossus – Histiostoma cossi, H. dryocoeti
C. ligniperda - Histiostoma cossi
Fam. Hesperiidae
Pelopides tridens - Scolianoetus javanus
Pisces
Perciformes
Fam. Percichthyidae
Maccullochella peelii peelii – Histiostoma papillatum
Siluriformes
Fam. Pangasiidae
Pangasius sutchi – Histiostoma piscium
Anguilliformes
Fam. Anguillidae
Anguilla anguilla – Histiostoma anguillarum
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Aves
Sphenisciformes
Fam. Spheniscidae
Eudyptula minor albosignata – Histiostoma mantelli, Myianoetus antipodus
Mammalia
Rodentia
Fam. Sciuridae
Spermophilus [Citellus] citellus – Myianoetus clavatus
Fam. Arvicolidae
Myodes [Clethrionomys] glareolus – Histiostoma sapromyzarum
Proboscidea
Fam. Elephantidae
Elephas maximus – Loxanoetus lenae
Loxodonta africana – Loxanoetus bassoni, Otanoetus wetzeli
Artiodactyla
Fam. Bovidae
Syncerus caffer – Auricanoetus longitarsus, Loxanoetus bassoni, Otanoetus
wetzeli
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Zamec R., V.Košel. 2014. A new species of mite (Acari: Histiostomatidae) phoretic on Gymnomus caesius (Diptera: Heleomyzidae) from Vlčie Diery Cave. – Biologia (Bratislava),
69(7): 916–918.
The Mite Family Guanolichidae Fain
Guanolichidae Fain, 1968: 199
Small family of four known species in Africa (Gabon) and America (Suriname, Panama
and Mexico). These completely sclerotized mites live in the bat guano, some were found
on bats (Noctilionidae, Natalidae).
Composition of Guanolichidae
Family Guanolichidae Fain, 1968–3 gen., 4 sp.
Guanolichoides Fain, 1979–1 sp.
Guanolichus Fain, 1968–1 sp.
Neoguanolichus Fain, 1979–2 sp.
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Catalogue of Guanolichidae
Family Guanolichidae Fain
Guanolichidae Fain, 1968: 199
Type genus: Guanolichus Fain, 1968
Genus Guanolichoides Fain
Guanolichoides Fain, 1979: 117
Type species: Guanolichoides lukoschusi Fain, 1979
Guanolichoides lukoschusi Fain
Guanolichoides lukoschusi Fain, 1979: 117
Type locality: Phedra
Distribution: Suriname
Host: Noctilio labialis (Chiroptera: Noctilionidae)
Genus Guanolichus Fain
Guanolichus Fain, 1968: 203
Type species: Guanolichus gabonensis Fain, 1968
Guanolichus gabonensis Fain
Guanolichus gabonensis Fain, 1968: 203
Type locality: bat guano, cave Faucon, Belinga, Gabon
Distribution: Gabon
Substrate: bat guano
Genus Neoguanolichus Fain
Neoguanolichus Fain, 1979: 117 [Redescription, Fashing 1980: 109]
Type species: Neoguanolichus mexicanus Fain, 1979
Neoguanolichus mexicanus Fain
Neoguanolichus mexicanus Fain, 1979: 118
Type locality: Durango
Distribution: Mexico
Substrate: from a bat, Natalus stramineus ventanus (Chiroptera: Natalidae)
Neoguanolichus panamensis Fashing
Neoguanolichus panamensis Fashing, 1980: 109
Type locality: Cocle 1 Cave, 0.8 mile North of Penonomé
Distribution: Panama
Substrate: bat guano
Acarorum Catalogus IX 143
Index of Guanolichidae
(Names in bold are accepted as valid)
gabonensis, Guanolichus 142
Guanolichidae 142
Guanolichus 142
lukoschusi, Guanolichoides 142
mexicanus, Neoguanolichus 142
Neoguanolichus 142
panamensis, Neoguanolichus 142
Distribution of Guanolichidae
AFRICA
Gabon (Fain, 1968) - Guanolichus gabonensis
NORTH AMERICA
Mexico (Fain, 1979) – Neoguanolichus mexicanus
CENTRAL AMERICA
Panama (Fashing, 1980) – Neoguanolichus panamensis
SOUTH AMERICA
Suriname (Fain, 1979) – Guanolichoides lukoschusi
Hosts of Guanolichidae
Chiroptera
Fam. Natalidae
Natalus stramineus ventatus – Neoguanolichus mexicanus
Fam. Noctilionidae
Noctilio labialis – Guanolichoides lukoschusi
Bibliography of Guanolichidae
Fain A. 1968. Deux nouveaux Acariens cavernicoles du Gabon (Sarcoptiformes). – Biologia
Gabonica, 4(2): 195–205.
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Fain A. 1979. Notes sur les Acariens astigmates cavernicoles. Description de deux genres
nouveaux et de quatre espèces nouvelles. - Acarologia (Paris), 20(1): 116–126.
Fashing N.J.1980. Neoguanolichus panamensis, a new species of Guanolichidae from bat caves
in Panama (Acarina: Sarcoptiformes). – Acarologia (Paris), 1979, 21(1): 108–116.
The Mite Superfamily Canestrinioidea
Superfamily of four families (Canestriniidae, Chetochelacaridae, Lophonotacaridae
and Heterocoptidae) of relatively large mites (113 genera, ca. 376 valid species) living as
commensals or parasites on beetles (Coleoptera: Carabidae, Chrysomelidae, Elateridae,
Scarabaeidae, Tenebrionidae, Passalidae and others), mostly under elytrae. One genus
and species of Canestriniidae (Diplopodocoptes transkeiensis) and the mites of the families
Chetochelacaridae and Lophonotacaridae are associated with Myriapods (Diplopoda).
The Mite Family Canestriniidae
Canestriniidae Berlese, 1884
As early as 1878 G. Canestrini described “Dermaleichus” [now Dicanestrinia] cerambycis. The study of the Canestriniidae (as of many other groups of mites) starts with some
papers of Berlese (1881, 1882, 1884, 1910, 1911, 1920), with descriptions of the genera
Canestrinia, Canestriniella, Coleopterophagus, Dicanestrinia and Percanestrinia and of the
family Canestriniidae. Oudemans (1904) described the important genus Photia.
Until the work of Cooreman (1950, 1955, 1958) and Lombardini (1950) have been
described only five more genera (by Trägårdh, 1906, Banks, 1915, Lombardini, 1943, Turk,
1948) and 16 valid species. The publications of Cooreman, and especially the one of 1950,
laid the base of the modern study of Canestriniidae (including nine new genera and several
new species). In the period between 1950 and 1988 several other authors contributed to
the study of these mites (Lavoipierre, Vomero, Fain, Nesbitt, Lindquist, Beron and others).
Should be marked especially the results obtained by K. Samšinák (1963–1971) and Summers and Schuster (1979–1982). In the important paper of Samšinák (1971) have been
described many new genera and new species from Carabidae. In five papers Summers
and Schuster published the subfamily Melisiinae, 11 new genera and many new species.
Until the beginning of the work of R. Haitlinger (1988, but including the paper of Fain,
1989) have been published 34 valid genera and 146 species of the family Canestriniidae.
From the 100 valid genera almost half (42) are monotypic.
In more than 450 papers since 1988 R. Haitlinger contributed most of all to the study
of Canestriniidae. He described 53 of the 100 valid genera and 200 of the 348 valid species
in the family (including the subfamily Vereoxiinae). The papers of Haitlinger deal with
Acarorum Catalogus IX 145
material from more than 50 countries and changed the overall picture of the distribution
and the list of hosts of these mites. Most of the new genera are so far monotypic.
Recently some newer authors joint the research efforts of Haitlinger. Kim et al. (2006)
described the new genus Haitlingeria, Trach, Eidelberg and Khaustov contributed to the
study of Canestriniids of Ukraine, including descriptions of four new genera.
Composition of Canestriniidae
Fam. Canestriniidae - 100 gen., 348 sp.
Subfam. Canestriniinae
Acrotocarus Banks, 1915–3 sp.
Afrocanestrinia Cooreman, 1955–3 sp.
Albinorattia Haitlinger, 1989–1 sp.
Amansiella Khaustov et Eidelberg, 2001–1 sp.
Ambilohylla Haitlinger, 1990–1 sp.
Amboinophela Haitlinger, 1993–1 sp.
Anaspistes Summers et Schuster, 1982–1 sp.
Apalotacarus Summers et Schuster, 1981–17 sp.
Arraphosoma Summers et Schuster, 1981–1 sp.
Arrunsithiana Haitlinger, 1992–10 sp.
Athogavia Haitlinger, 1989–7 sp.
Aurilossongia Haitlinger, 1989–6 sp.
Averithiana Haitlinger, 1999–1 sp.
Barbiangia Haitlinger, 1993–3 sp.
Beeriphotia Haitlinger, 1994–1 sp.
Bibulothiana Haitlinger, 1993–6 sp.
Bircericola Haitlinger, 2000–1 sp.
Boetophela Haitlinger, 1989–10 sp.
Boleohylla Haitlinger, 1991–2 sp.
Camirohylla Haitlinger, 1991–1 sp.
Canestrinia Berlese, 1881–17 sp.
Canestriniella Berlese, 1910–6 sp.
Caraboecius Cooreman, 1950–1 sp.
Cetonicola Cooreman, 1955–5 sp.
Chelinochroa Summers et Schuster, 1982–1 sp.
Cnecoderma Summers et Schuster, 1981–2 sp.
Coleoglyphus Berlese, 1910b - 2 sp.
Coleopterophagus Berlese, 1882–11 sp.
Cooremania Nesbitt, 1976–2 sp.
Danaithiana Haitlinger, 1991b – 4 sp.
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Acarorum Catalogus IX
Dicanestrinia Berlese, 1911–3 sp.
Diplognathophilus Cooreman, 1955–1 sp.
Diplopodocoptes Fain, 1987–1 sp.
Donnelaphontia Lavoipierre, 1958–2 sp.
Dorynothiana Haitlinger, 1999–2 sp.
Ennodithiana Haitlinger, 1993–6 sp.
Eunorithiana Haitlinger, 1999–2 sp.
Gasuthiana Haitlinger, 1989–5 sp.
Gioharattia Haitlinger, 1989–2 sp.
Globosophotia Haitlinger, 1991–1 sp.
Grandiellina Fain, 1989 (= Grandiella Lombardini, 1939 praeocc.) - 21 sp.
Grandiellopsis Fain, 1989–7 sp.
Haitlingeria Kim et al., 2006–1 sp.
Hargeinella Haitlinger, 1989–1 sp.
Hatohylla Haitlinger, 1990–4 sp.
Hyllosihemia Haitlinger, 1994–1 sp.
Hypopteryx Summers et Schuster, 198–1 sp.
Ibizella Haitlinger et Šundić, 1918–3 sp.
Irmongia Haitlinger, 1990b – 1 sp.
Jowitella Haitlinger, 1990b – 1 sp.
Jullongia Haitlinger, 1990b – 1 sp.
Kahoorangia Haitlinger, 1991–1 sp.
Lidiophela Haitlinger, 1990b – 7 sp.
Megacanestrinia Trägårdh, 1906–2 sp.
Subfam. Melisiinae
Contramelisia Samšinák, 1969–1 sp.
Melisia Lombardini, 1944–22 sp.
Mesophotia Samšinak, 1971–1 sp.
Metriothiana Haitlinger, 1999–1 sp.
Mossongia Haitlinger, 1989–2 sp.
Neophotia Samšinák, 1971–3 sp.
Noemiphela Haitlinger, 1992–2 sp.
Oceogavia Haitlinger, 1989–1 sp.
Olafithiana Haitlinger, 1993–5 sp.
Olgattia Haitlinger, 1990–1 sp.
Ovarrothiana Haitlinger, 1993–6 sp.
Paracanestrinia Haitlinger, 2017–1 sp.
Paramansia Cooreman, 1950–3 sp.
Paramelisia Summers et Schuster, 1982–1 sp.
Paraphagella Cooreman, 1955–6 sp.
Paraphotia Khaustov et Eidelberg, 2001–1 sp.
Acarorum Catalogus IX 147
Passalophagus Nesbitt, 1976–1 sp.
Percanestrinia Berlese, 1911–8 sp.
Percanestriniella Khaustov et Eidelberg, 2001–4 sp.
Peruthiana Haitlinger, 2008–1 sp.
Phaleratus Nesbitt, 1976–2 sp.
Phelliculophela Haitlinger, 1993–1 sp.
Photia Oudemans, 1904–18 sp.
Phriknodora Summers et Schuster, 1982–3 sp.
Procericola Cooreman, 1950–2 sp.
Pseudamansia Cooreman, 1950–2 sp.
Pseudocanestrinia Khaustov et Eidelberg, 2001–5 sp.
Pseudoparaphagella Haitlinger, 2010–1 sp.
Rugoniphela Haitlinger, 1992–1 sp.
Sabothiana Haitlinger, 1993–3 sp.
Sajanophela Haitlinger, 1990–1 sp.
Sandrophela Haitlinger, 1990–7 sp.
Saniothiana Haitlinger, 1990–2 sp.
Scarabiphotia Haitlinger, 1994–1 sp.
Shillongia Haitlinger, 1990–1 sp.
Sorbinophela Haitlinger, 1993–1 sp.
Tamarangia Haitlinger, 1991–4 sp.
Tamarophela Haitlinger, 1990–1 sp.
Taseopus Summers et Schuster, 1982–1 sp.
Teklathiana Haitlinger, 1999–3 sp.
Teophyssa Haitlinger, 1990b – 3 sp.
Thopia Summers et Schuster, 1982–6 sp.
Uriophela Haitlinger, 1992–1 sp.
Vandeinella Haitlinger, 1989–2 sp.
Subfam. Vereoxiinae
Vereoxia Haitlinger, 1995–1 sp.
Catalogue of the Family Canestriniidae
Family Canestriniidae Berlese
Canestriniidae Berlese, 1884: 92
Type genus: Canestrinia Berlese, 1881
Subfamily Canestriniinae Berlese
Canestriniidae Berlese, 1884: 92
Type genus: Canestrinia Berlese, 1881
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Genus Acrotocarus Banks
Acrotocarus Banks, 1915: 152
Type species: Acrotocarus mirabilis Banks, 1915
Acrotocarus alutaceus (Turk)
Amansia alutacea Turk, 1948: 88
Acrotocarus alutaceus (Turk): Summers & Schuster 1979b: 480
Type locality: St. Augustine
Distribution: Trinidad
Host: Physonota alutacea (Coleoptera: Chrysomelidae, Cassidinae)
Acrotocarus crataepus (Summers et Schuster)
Acrotocarus crataepus Summers & Schuster, 1979b: 484
Type locality: unknown
Distribution: unknown
Host: Physonota dilata (Coleoptera: Chrysomelidae, Cassidinae)
Acrotocarus mirabilis Banks
Acrotocarus mirabilis Banks, 1915: 152; Summers & Schuster, 1979b: 476
Type locality: Cordoba (Mexico)
Distribution: Costa Rica, Guatemala, Mexico
Hosts: Physsonota alutacea, Physsonota sp. (Coleoptera: Chrysomelidae)
Genus Afrocanestrinia Cooreman
Afrocanestrinia Cooreman, 1955: 25
Type species: Afrocanestrinia straeleni Cooreman, 1955
Afrocanestrinia rufinae Haitlinger
Afrocanestrinia rufinae Haitlinger, 1995a: 85
Type locality: Tanzania
Distribution: Tanzania
Host: Smaragdesthes oertzeni (Coleoptera: Scarabaeidae, Cetoniinae)
Afrocanestrinia samsoni Haitlinger
Afrocanestrinia samsoni Haitliriger, 1995a: 87
Type locality: Senegal
Distribution: Senegal
Host: Smaragdesthes africana (Coleoptera: Scarabaeidae, Cetoniinae)
Afrocanestrinia straeleni Cooreman
Afrocanestrinia straeleni Cooreman, 1955: 25; Haitlinger, 1995: 85
Acarorum Catalogus IX 149
Type locality: May-ya-Moto, Rutshuru
Distribution: D.R. Congo
Hosts: Smaragdesthes africana mutica, Dicronorhina micans (Coleoptera: Scarabaeidae,
Cetoniinae)
Genus Albinorattia Haitlinger
Albinorattia Haitlinger, 1989: 588
Type species: Albinorattia igori Haitlinger, 1989
Albinorattia igori Haitlinger
Albinorattia igori Haitlinger, 1989: 588
Type locality: Batjan
Distribution: Indonesia
Host: Ischipsophia olivacea (Coleoptera: Sarabaeidae, Cetoniinae)
Genus Amansiella Khaustov et Eidelberg
Amansiella Khaustov & Eidelberg, 2001: 41
Type species: Amansiella trilobata Khaustov et Eidelberg, 2001
Amansiella trilobata Khaustov et Eidelberg
Amansiella trilobata Khaustov & Eidelberg, 2001: 41; Trach, 2006: 176; Trach & Khaustov,
2007: 268
Type locality: Crimea, Yalta
Distribution: Crimea, Ukraine
Host: Timarcha tenebricosa (Coleoptera: Chrysomelidae)
Genus Ambilohylla Haitlinger
Ambilohylla Haitlinger, 1990: 316
Type species: Ambilohylla Haitlinger, 1993
Ambilohylla favosa Haitlinger
Ambilohylla favosa Haitlinger, 1990: 316
Type locality: Madagascar
Distribution: Madagascar
Host: Damatria formosus (Coleoptera: Tenebrionidae)
Genus Amboinophela Haitlinger
Amboinophela Haitlinger, 1993: 339
Type species: Amboinophela bernae Haitlinger, 1993
Amboinophela bernae Haitlinger
Amboinophela bernae Haitlinger, 1993b: 340
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Type locality: Moluccas, Amboina Is.
Distribution: Indonesia, New Guinea
Host: Metapodonotus bison, M. cinctus (Coleoptera: Lucanidae)
Genus Anaspistes Summers et Schuster
Anaspistes Summers & Schuster, 1982b: 43
Type species: Anapistes unguiculatus Summers et Schuster, 1982
Anaspistes unguiculatus Summers et Schuster
Anaspistes unguiculatus Summers & Schuster, 1982b: 43
Type locality: Kisarawe
Distribution: Tanzania
Host: Sternotomis bohemanni (Coleoptera: Cerambycidae)
Genus Apalotacarus Summers et Schuster
Apalotacarus Summers & Schuster, 1981a: 28
Type species: Apalotacarus fusulus Summers et Schuster, 1981
Apalotacarus aristatus Summers et Schuster
Apalotacarus aristatus Summers & Schuster, 1981a: 28
Type locality: Mucha
Distribution: Taiwan, Indonesia (Java), India (Assam)
Host: Aceraius grandis hirsutus (Coleoptera: Passalidae)
Apalotacarus cidaris Summers et Schuster
Apalotacarus cidaris Summers & Schuster, 1981a: 25
Type locality: Daru
Distribution: Papua New Guinea
Host: Protomocoelus australis (Coleoptera: Passalidae)
Apalotacarus echinatus Summers et Schuster
Apalotacarus echinatus Summers & Schuster, 1981a: 33
Type locality: 12 mi.W of Munnar, S. India
Distribution: India
Host: Pleurarius brachyphilus (Coleoptera: Passalidae)
Apalotacarus fulgens Summers et Schuster
Apalotacarus fulgens Summers & Schuster, 1981a: 31
Type locality: Batavia [Djakarta], Java
Distribution: Indonesia (Java)
Host: Pelopides tridens (Coleoptera: Passalidae)
Acarorum Catalogus IX 151
Apalotacarus fusticulus Summers et Schuster
Apalotacarus fusticulus Summers & Schuster, 1981a: 31
Type locality: Ulong, NSW
Distribution: Australia (NSW)
Host: Pharochilus nitidulus (Coleoptera: Passalidae)
Apalotacarus fusulus Summers et Schuster
Apalotacarus fusulus Summers & Schuster, 1981a: 20
Type locality: Finschhafen, Papua New Guinea
Distribution: Papua New Guinea, Philippines (Mindoro)
Host: Labienus plox (Coleoptera: Passalidae)
Apalotacarus glaber Summers et Schuster
Apalotacarus glaber Summers & Schuster, 1981a: 25
Type locality: Russel Island
Distribution: Solomon Islands
Host: Protomocoelus triumphator (Coleoptera: Passalidae)
Apalotacarus gracilis Summers et Schuster
Apalotacarus gracilis Summers & Schuster, 1981a: 33
Type locality: 40 mi. SE Walcha, Stop 418, NSW, Australia
Distribution: Australia
Host: Coleoptera (Passalidae)
Apalotacarus luroris Summers et Schuster
Apalotacarus luroris Summers & Schuster, 1981a: 33
Type locality: Ulong, NSW
Distribution: Australia (NSW, Queensland)
Hosts: Mastochilus australasicus, M. quaestionis, Pharochilus dilatatus, Ph. nitidulus (Coleoptera: Passalidae)
Apalotacarus paxillus Summers et Schuster
Apalotacarus paxillus Summers & Schuster, 1981a: 21
Type locality: Wau, Morobe
Distribution: Papua New Guinea
Host: Coleoptera (Passalidae)
Apalotacarus petilus Summers et Schuster
Apalotacarus petilus Summers & Schuster, 1981a: 23
Type locality: Finschhafen
Distribution: Papua New Guinea, incl. Bougainville I.
Host: Protomocoelus australis (Coleoptera: Passalidae)
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Apalotacarus protensus Summers et Schuster
Apalotacarus protensus Summers & Schuster, 1981a: 28
Type locality: Kurseong, Darjeeling
Distribution: India
Host: Macrolinus nicobaricus (Coleoptera: Passalidae)
Apalotacarus rigescens Summers et Schuster
Apalotacarus rigescens Summers & Schuster, 1981a: 26
Type locality: Wau, Morobe
Distribution: Papua New Guinea
Host: Coleoptera (Passalidae)
Apalotacarus sajanae Haitlinger
Apalotacarus sajanae Haitlinger, 1990b: 536
Type locality: Java
Distribution: Indonesia (Java)
Host: Eriocnemis tridens (Coleoptera: Passalidae)
Apalotacarus scaurus Summers et Schuster
Apalotacarus scaurus Summers & Schuster, 1981a: 30
Type locality: Kabasalan, Mindanao
Distribution: Philippines (Mindanao)
Host: Macrolinus weberi (Coleoptera: Passalidae)
Apalotacarus scissus Summers et Schuster
Apalotacarus scissus Summers & Schuster, 1981a: 37
Type locality: Anomali Hills, Cinchona, S. India
Distribution: India
Host: Plurarius brachyphyllus (Coleoptera: Passalidae)
Apalotacarus trullus Summers et Schuster
Apalotacarus trullus Summers & Schuster, 1981a: 23
Type locality: Hassensan
Distribution: Taiwan
Host: Leptaulax bicolor formosanus (Coleoptera: Passalidae)
Genus Arraphosoma Summers et Schuster
Arraphosoma Summers & Schuster, 1981b: 43
Type species: Arraphosoma minax Summers & Schuster, 1981
Arraphosoma minax Summers et Schuster
Arraphosoma minax Summers & Schuster, 1981b: 44
Acarorum Catalogus IX 153
Type locality: New Guinea
Distribution: New Guinea
Host: (Coleoptera: Passalidae)
Arraphosoma paraminax Summers et Schuster
Arraphosoma paraminax Summers & Schuster, 1981b: 44
Type locality: Mollucan Islands
Distribution: Indonesia
Host: Labiens gigas (Coleoptera: Passalidae)
Genus Arrunsithiana Haitlinger
Arrunsithiana Haitlinger, 1991e: 392
Type species: Arrunsithiana artemoni Haitlinger, 1991
Arrunsithiana arcuata (Fain)
Grandiella arcuata Fain, 1987: 109
Grandiellina arcuata (Fain): Fain, 1989: 67
Arrunsithiana arcuata (Fain): Haitlinger, 1991e: 397
Type locality: Bahia, Santarem
Distribution: Brazil
Hosts: Stolas decemguttata, S. esignata (Coleoptera: Chrysomelidae)
Arrunsithiana artemoni Haitlinger
Arrunsithiana artemoni Haitlinger, 1991e: 393
Type locality: Ecuador
Distribution: Ecuador
Host: Stolas latevittata (Coleoptera: Chrysomelidae)
Arrunsithiana battosi Haitlinger
Arrunsithiana battosi Haitlinger, 1991e: 396
Type locality: Bolivia, Coroica
Distribution: Bolivia, Brazil, Colombia, Ecuador, Peru
Hosts: Stolas pleurosticha, S. lenis, S. oblita, S. erichsoni, S. inexculta, S. huanocensis, S.
cruentata, S. scoparia, S. funebris, S. niobe, S. ignita, S. conspersa, S. gibbosa, S. paranensis, S. chalybaea (Coleoptera: Chrysomelidae)
Arrunsithiana erinnae Haitlinger
Arrunsithiana erinnae Haitlinger, 1991e: 394
Type locality: Santa Inez
Distribution: Ecuador
Host: Stolas pectinata (Coleoptera: Chrysomelidae)
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Arrunsithiana esacaudata (Lombardini)
Grandiella esacaudata Lombardini, 1939: 118; 1950: 784; Summers & Schuster, 1979a:
335; Fain, 1987: 109
Grandiellina esacaudata (Lombardini): Fain, 1989: 67
Arrunsithiana esacaudata (Lombardini): Haitlinger, 1991e : 398
Type locality: Brazil
Distribution: Bolivia, Brazil, Costa Rica, French Guyana, Guatemala, Mexico
Hosts: Omoplata marginata (Coleoptera: Cerambycidae), Phaeneum sp., Stolas punicea
(Coleoptera: Chrysomelidae)
Arrunsithiana ilosi Haitlinger
Arrunsithiana ilosi Haitlinger, 1991e: 395
Type locality: Bogota
Distribution: Colombia
Host: Cyrtonota dissecta (Coleoptera: Chrysomelidae)
Arrunsithiana latipes (Fain)
Grandiellina latipes Fain, 1989: 67
Arrunsithiana latipes (Fain): Haitlinger, 1991e: 398
Type locality: South America
Distribution: South America
Host: Stolas inaequalis (Coleoptera: Chrysomelidae)
Arrunsithiana quadrata (Fain)
Grandiella quadrata Fain, 1987: 106
Grandiellina quadrata (Fain): Fain, 1989: 67
Arrunsithiana quadrata (Fain): Haitlinger, 1991e: 397
Type locality: Brazil, Bolivia (Santa Cruz), Colombia (Muzo)
Distribution: Colombia, Bolivia, Brazil
Host: Stolas aenea, S. indigacea, S.conspersa, S. chalybaea, S. funebris, Eugenysa columbiana
(Coleoptera: Chrysomelidae)
Arrunsithiana spicantis (Summers et Schuster)
Grandiella spicantis Summers & Schuster, 1979a: 341; Fain, 1987: 94
Grandiellina spicantis (Summers et Schuster): Fain, 1989: 67
Arrunsithiana spicantis (Summers et Schuster): Haitlinger, 1991e : 397
Type locality: Santa Catarina
Distribution: Brazil
Hosts: Canistra plagosa, Stolas chalybaea, S. conspersa (Coleoptera: Chrysomelidae)
Arrunsithiana squarrosa (Summers et Schuster)
Grandiella squarrosa Summers & Schuster, 1979a: 338; Fain, 1987: 94
Acarorum Catalogus IX 155
Grandiellina squarosa (Summers et Schuster): Fain, 1989: 67
Arrunsithiana squarrosa (Summers et Schuster): Haitlinger, 1991e : 398
Type locality: Chanchamayo Valley
Distribution: Peru, French Guiana
Host: Stolas sp. (Coleoptera: Chrysomelidae)
Genus Athogavia Haitlinger
Athogavia Haitlinger, 1989d: 337
Type species: Athogavia tanzanica Haitlinger, 1989
Athogavia arybbasi Haitlinger
Athogavia arybbasi Haitlinger, 1989d: 592
Type locality: Papua New Guinea
Distribution: Papua New Guinea
Host: Dynastinae (Coleoptera: Scarabaeidae)
Athogavia assamica Haitlinger
Athogavia assamica Haitlinger, 1989d: 348
Type locality: Assam
Distribution: India
Host: Trichogomphus nartolani (Coleoptera: Scarabaeidae, Dynastinae)
Athogavia borneoica Haitlinger
Athogavia borneoica Haitlinger, 1989d: 346
Type locality: Borneo
Distribution: Borneo
Host: Oryctes rhinoceros (Coleoptera: Scarabaeidae, Dynastinae)
Athogavia gamana Haitlinger
Athogavia gamana Haitlinger, 1989d: 342
Type locality: Tanzania
Distribution: Tanzania
Host: Oryctes monoceros (Coleoptera: Scarabaeidae, Dynastinae)
Athogavia nosiana Haitlinger
Athogavia nosiana Haitlinger, 1989: 340
Type locality: Madagascar
Distribution: Madagascar
Host: Oryctes (Coleoptera: Scarabaeidae, Dynastinae)
Athogavia scagana Haitlinger
Athogavia scagana Haitlinger, 1989d: 342
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Acarorum Catalogus IX
Type locality: Borneo
Distribution: Borneo
Host: Oryctes sp. (Coleoptera: Scarabaeidae, Dynastinae)
Athogavia tanzanica Haitlinger
Athogavia tanzanica Haitlinger, 1989d: 338
Type locality: Tanzania
Distribution: Tanzania
Host: Oryctes monoceros (Coleoptera: Scarabeidae, Dynastinae)
Genus Aurilossongia Haitlinger
Aurilossongia Haitlinger, 1989a: 43
Type species: Aurilossongia hoaria Haitlinger, 1989
Aurilossongia bolanica Haitlinger
Aurilossongia bolanica Haitlinger, 1989a: 47
Type locality: Bolan Mts.
Distribution: Papua New Guinea
Host: Xylotrupes asperulus (Coleoptera: Cerambycidae)
Aurilossongia dioni Haitlinger
Aurilossongia dioni Haitlinger, 1989: 593
Type locality: Indonesia
Distribution: Indonesia
Host: Chalcosoma atlas (Coleoptera: Scarabaeidae, Dynastinae)
Aurilossongia fileasi Haitlinger
Aurilossongia fileasi Haitlinger, 1989: 593
Type locality: Papua New Guinea
Distribution: Papua New Guinea
Host: Scarabaeidae, Dynastinae (Coleoptera)
Aurilossongia gaurana Haitlinger
Aurilossongia gaurana Haitlinger, 1989a: 48
Type locality: New Britain, Rabaul
Distribution: Papua New Guinea
Host: ?Xylotrupes sp. (Coleoptera: Cerambycidae, Dynastinae)
Aurilossongia hoaria Haitlinger
Aurilossongia hoaria Haitlinger, 1989a: 44
Type locality: Borneo (Kalimantan)
Acarorum Catalogus IX 157
Distribution: Indonesia (Kalimantan)
Host: Xylotrupes sp. (Coleoptera: Cerambycidae, Dynastinae)
Aurilossongia norae Haitlinger
Aurilossongia norae Haitlinger, 1989: 594
Type locality: Papua New Guinea
Distribution: Papua New Guinea
Host: Dynastes gideon (Coleoptera: Scarabaeidae, Dynastinae)
Genus Averithiana Haitlinger
Averithiana Haitlinger, 1999: 42
Type species: Averithiana anoni Haitlinger, 1999
Averithiana anoni Haitlinger
Averithiana anoni Haitlinger, 1999: 42
Type locality: French Guyana
Distribution: French Guyana
Host: Zatrephina lineata (Coleoptera: Chrysomelidae, Cassidinae)
Genus Barbiangia Haitlinger
Barbiangia Haitlinger, 1993a: 5
Type species: Barbiangia alvari Haitlinger, 1993
Barbiangia alvari Haitlinger
Barbiangia alvari Haitlinger, 1993a: 6
Type locality: Zanzibar
Distribution: Tanzania (Zanzibar)
Host: undetermined Cetoniinae (Coleoptera: Scarabaeidae)
Barbiangia elongata Khaustov
Barbiangia elongata Khaustov, 2006: 80
Type locality: Ethiopia
Distribution: Ethiopia
Host: Diplognatha gagates (Coleoptera: Scarabaeidae, Cetoniinae)
Barbiangia ethiopica (Haitlinger)
Diplognatophilus ethiopicus Haitlinger, 1990: 321
Barbiangia ethiopica (Haitlinger): Haitlinger, 1993a: 7
Type locality: Ethiopia
Distribution: Ethiopia
Host: Scarabaeidae, Cetoniinae (Coleoptera)
158
Acarorum Catalogus IX
Genus Beeriphotia Haitlinger
Beeriphotia Haitlinger, 1994a: 207
Type species: Beeriphotia volkeri Haitlinger, 1994
Beeriphotia volkeri Haitlinger
Beeriphotia volkeri Haitlinger, 1994a: 207
Type locality: Guandong (Kanton)
Distribution: China
Host: Carabus angulicollis antaeus (Coleoptera: Carabidae)
Genus Bibulothiana Haitlinger
Bibulothiana Haitlinger, 1993e: 278
Type species: Grandiella brevispinata Summers et Schuster, 1979
Bibulothiana albertae Haitlinger
Bibulothiana albertae Haitlinger, 1994: 82
Type locality: Turrialba
Distribution: Costa Rica
Host: Chelymorpha comata (Coleoptera: Chrysomelidae, Cassidinae)
Bibulothiana brevispinata (Summers et Schuster)
Grandiella brevispinata Summers & Schuster, 1979: 346
Bibulothiana brevispinata (Summers et Schuster): Haitlinger, 1993e: 278
Type locality: 9 km N .El Cope, Panama
Distribution: Bolivia, Brazil, Colombia, Honduras, Panama
Hosts: Stolas bifenestrata, S. bipustulata, S. coerulescens, S. ephippium, S. lineaticollis, S.
plagiata (Coleoptera: Chrysomelidae, Cassidinae)
Bibulothiana cyriaki Haitlinger
Bibulothiana cyriaki Haitlinger, 1993e: 280
Type locality: Parana, Iguacu
Distribution: Brazil, Mexico
Hosts: Chelymorpha infecta, Ch. nigricollis, Ch. vermiculata (Coleoptera: Chrysomelidae,
Cassidinae)
Bibulothiana everittae Haitlinger
Bibulothiana everittae Haitlinger, 1993e: 281
Type locality: Santa Catarina (Brazil)
Distribution: Brazil, Peru
Hosts: Chelymorpha inflate, Ch. rufoguttata (Coleoptera: Chrysomelidae, Cassidinae)
Acarorum Catalogus IX 159
Bibulothiana oswaldi Haitlinger
Bibulothiana oswaldi Haitlinger, 1993e: 279
Type locality: Bahia
Distribution: Brazil
Host: Stolas plagicollis (Coleoptera: Chrysomelidae, Cassidinae)
Bibulothiana vatia (Summers et Schuster)
Grandiella vatia Summers & Schuster, 1979: 333
Bibulothiana vatia (Summers et Schuster): Haitlinger 1993e: 281
Type locality: Lago, Catemaco, Vera Cruz
Distribution: Mexico
Host: Cyrtonota tristigma (Coleoptera: Chrysomelidae, Cassidinae)
Genus Bircericola Haitlinger
Bircericola Haitlinger, 2000: 69
Type species: Bicercicola bertrami Haitlinger, 2000
Bircericola bertrami Haitlinger
Bircericola bertrami Haitlinger, 2000: 70
Type locality: Bir Bon Ragba, nr. Hammamet
Distribution: Tunisia
Host: undetermined small Tenebrionidae (Coleoptera)
Genus Boetophela Haitlinger
Boetophela Haitlinger, 1989: 589
Type species: Boetophela ginae Haitlinger, 1989
Boetophela cassandrae Haitlinger et Chmielewski
Boetophela cassandrae Haitlinger & Chmielewski, 2004: 196
Type locality: Durban
Distribution: South Africa
Host: undetermined Cetoniinae (Coleoptera: Scarabeidae)
Boetophela erhardi Haitlinger
Boetophela erhardi Haitlinger, 1991d: 304
Type locality: Nyangassy
Distribution: Cameroon
Host: Mecynorrhina [Chelorrhina] savagei (Coleoptera: Scarabeidae, Cetoniinae)
Boetophela ephraimi Haitlinger
Boetophela ephraimi Haitlinger, 1993: 12
Type locality: Ashanti
160
Acarorum Catalogus IX
Distribution: Ghana
Host: undetermined Cetoniinae (Coleoptera: Scarabaeidae)
Boetophela ginae Haitlinger
Boetophela ginae Haitlinger, 1989: 591; 1993: 10
Type locality: Natal, Transvaal [described ex errore from Sumatra, Batocera rubus,
Cerambycidae, corrected by Haitlinger, 1993a]
Distribution: Republic of South Africa
Host: Eudicella smithi (Coleoptera: Scarabaeidae, Cetoniinae)
Boetophela robertsoni (Lavoipierre)
Cetonicola robertsoni Lavoipierre, 1958: 616
Boetophela robertsoni: Haitlinger, 1993: 14
Type locality: Tanzania
Distribution: Tanzania
Host: Dicronorhina oberthuri (Coleoptera: Scarabaeidae, Cetoniinae)
Boetophela silvanae Haitlinger
Boetophela silvanae Haitlinger, 1991d: 303
Type locality: Tanzania
Distribution: Tanzania
Host: Geynodonta laeviplaga (Coleoptera: Scarabaeidae, Cetoniinae)
Boetophela sybillae Haitlinger
Boetophela sybillae Haitlinger, 1991d: 299
Type locality: “surroundings of Nyassa lake”
Distribution: Tanzania
Host: Goliathus alborignathus (Coleoptera: Scarabaeidae, Cetoniinae)
Boetophela symeoni Haitlinger
Boetophela symeoni Haitlinger, 1991d: 300
Type locality: probably Tanzania
Distribution: probably Tanzania
Host: Elasocheilon heterospila (Coleoptera: Scarabeidae, Cetoniinae)
Boetophela werneri Haitlinger
Boetophela werneri Haitlinger, 1993a: 14
Type locality: Bujumbura
Distribution: Burundi
Host: Plaesiorrhina mhondana (Coleoptera: Scarabeidae, Cetoniinae)
Acarorum Catalogus IX 161
Genus Boleohylla Haitlinger
Boleohylla Haitlinger, 1991a: 274
Type species: Boleohylla kiogana Haitlinger, 1991
Boleohylla garrica Haitlinger
Boleohylla garrica Haitlinger, 1991a: 276
Type locality: Ghana
Distribution: Ghana
Host: Metallonotus metallicus (Coleoptera: Tenebrionidae)
Boleohylla kiogana Haitlinger
Boleohylla kiogana Haitlinger, 1991a: 274
Type locality: nr. Lake Kiega
Distribution: Uganda
Host: Psammodes carbonarius (Coleoptera: Tenebrionidae)
Genus Camirohylla Haitlinger
Camirohylla Haitlinger, 1991a: 273
Type species: Camirohylla feziana Haitlinger, 1991
Camirohylla feziana Haitlinger
Camirohylla feziana Haitlinger, 1991a: 274; 2001: 25
Type locality: Morocco
Distribution: Morocco, Spain (Ibiza, Formentera)
Host: Blaps appendiculata (Coleoptera: Tenebrionidae)
Camirohylla sp.
Type locality: Alashtar, Lorestan Province
Distribution: Iran
Host: Dila sp. (Coleoptera: Tenebrionidae)
Genus Canestrinia Berlese
Canestrinia Berlese, 1881: 747
Type species: Canestrinia dorcicola Berlese, 1881
Canestrinia carabicola (Berlese)
Coleopterophagus carabicola Berlese, 1882: 10
Canestrinia (Percanestrinia) sardica Vitzthum, 1924: 41
Percanestrinia carabicola: Samšinak, 1965:
Canestrinia carabicola: Samšinak, 1971: 176
Type locality: Italy
162
Acarorum Catalogus IX
Distribution: Italy (incl. Sardinia and Sicily), Corsica, Tunisia
Host: Carabus (Macrothorax) morbillosus (Coleoptera: Carabidae)
Canestrinia damiani Haitlinger
Canestrinia damiani Haitlinger, 1989: 353
Type locality: Hungary
Distribution: Hungary, Ukraine, Crimea
Host: Pentodon idiota (Coleoptera)
Canestrinia dorcicola Berlese
Canestrinia dorcicola Berlese, 1881: 748; Canestrini, 1897: 910; Canestrini & Kramer, 1899:
22; Leonardi, 1900: 15; Buitendijk, 1945: 364; Turk, 1953: 82; Cooreman, 1954: 86;
Haitlinger, 1988a; 2004: 13; 2012: 186; Khaustov & Edelberg, 2001: 40; Trach, 2006: 176
Type locality: Italy
Distribution: Great Britain, Holland, Italy, Poland, Ukraine, Crimea
Hosts: Oxythyrea funesta (Coleoptera: Scarabaeidae, Cetoniinae), Dorcus parallelopipedus
(Coleoptera: Lucanidae)
Canestrinia hispanica Samšinák
Canestrinia hispanica Samšinák, 1971: 173
Type locality: “Hispania”
Distribution: Morocco, Portugal, Spain
Hosts: Carabus (Rhabdotocarabus) melancholicus, C. (Ctenocarabus) galicianus, C. (Oreocarabus) guadarramus (Coleoptera: Carabidae).
Canestrinia illegalis Samšinák
Canestrinia illegalis Samšinák, 1971: 162
Type locality: Hualien (Taiwan)
Distribution: Taiwan
Hosts: Carabus (Coptolabrus) hankotaizanus, C. (Coptolabrus) longipennis mandarinus
(Coleoptera: Carabidae)
Canestrinia kacperi Haitlinger
Canestrinia kacperi Haitlinger, 1998: 3
Type locality: Guanzhou
Distribution: China
Host: Carabus subformosus (Coleoptera: Carabidae)
Canestrinia kamelskyi Samšinák
Canestrinia kamelskyi Samšinák, 1970: 75
Type locality: Tunisia
Acarorum Catalogus IX 163
Distribution: Tunisia
Host: Blaps sp. (Coleoptera: Tenebrionidae)
Canestrinia macgillavryi Oudemans
Canestrinia macgillavryi Oudemans, 1923: 204; Cooreman, 1954: 86
Type locality: Soember Pakel Station, Dampit, Probolingo
Distribution: Indonesia (Java)Canestrinia manicata Berlese
Canestrinia manicata Berlese, 1910: 269
Type locality: Brazil
Distribution: Brazil
Host: Doryphora tessellata (Coleoptera: Chrysomelidae)
Canestrinia microdisca Berlese
Canestrinia microdisca Berlese, 1910: 269
Type locality: Java
Distribution: Indonesia
Host: Dorcus saiga (Coleoptera: Lucanidae)
Canestrinia nepalensis Berlese
Canestrinia nepalensis Berlese, 1910: 269
Type locality: Java
Distribution: Indonesia
Host: Dorcus nepalensis (Coleoptera: Lucanidae)
Canestrinia paavoi Haitlinger
Canestrinia paavoi Haitlinger, 1994a: 208
Type locality: South China, Tschekiang (= Zhejiangsheng)
Distribution: China
Host: Carabus grandis (Coleoptera: Carabidae)
Canestrinia pekingensis Samšinák
Canestrinia pekingensis Samšinák, 1971: 164
Type locality: Pekin
Distribution: China
Hosts: Carabus (Acathaicus) brandti, C. (Cychrostomus) anchocephalus (Tibet, 3200
m), C. (Eupachys) glyptopterus, C. (Coptolabrus) lafossei saturatus, C. augustus, C.
augustus ignigena, C. pustulifer, C. pustulifer pratti, C. p. querryi, C. p. szechwanensis,
C. ignimetalla angusticollis, C. gemmifer (Coleoptera: Carabidae)
164
Acarorum Catalogus IX
Canestrinia pentodontis Berlese
Canestrinia dorcicola var. pentodontis Berlese, 1897:
Canestrinia pentodontis Berlese: Vitzthum, 1925: 47
Type locality: Italy
Distribution: Italy, Sicily
Host: Pentodon (Coleoptera: Scarabaeidae, Dynastinae)
Canestrinia pictura Samšinák
Canestrinia pictura Samšinák, 1971: 171
Type locality: Mt Kurama near Kyoto
Distribution: Japan
Host: Carabus (Damaster) blaptoides (Coleoptera: Carabidae)
Canestrinia ramoni Haitlinger
Canestrinia ramoni Haitlinger, 1994a: 212
Type locality: South China, Fukien (= Fujiansheng)
Distribution: China
Host: Carabus coelestis giganteus (Coleoptera: Carabidae)
Canestrinia remigans Berlese
Canestrinia remigans Berlese, 1910: 387
Type locality: Africa
Distribution: Africa
Genus Canestriniella Berlese
Canestriniella Berlese, 1910: 270
Type species: Canestriniella amplexans Berlese, 1910
Canestriniella amplexans Berlese
Canestriniella amplexans Berlese, 1910: 270
Type locality: “Batavia” (Jakarta)
Distribution: Indonesia (Java)
Host: Batocera hector (Coleoptera: Cerambycidae)
Canestriniella batocerae (Vishnupriya et Mohanasundaram)
Grandiella batocerae Vishnupriya & Mohanasundaram, 1988: 253
Grandiellina batocerae (Vishnupriya et Mohanasundaram): Fain, 1989: 67
Type locality: Tamil Nadu
Distribution: India
Host: Batocera rufomaculata (Coleoptera: Cerambycidae, Lamiinae)
Acarorum Catalogus IX 165
Canestriniella elae Haitlinger
Canestriniella elae Haitlinger, 1989: 599
Type locality: Sumatra
Distribution: Indonesia
Host: Batocera ferruginea (Coleoptera: Cerambycidae)
Canestriniella phoebinae Haitlinger
Canestriniella phoebinae Haitlinger, 1995a: 89
Type locality: Java
Distribution: Indonesia
Host: Batocera albofasciata octomaculata (Coleoptera: Cerambycidae)
Canestriniella physsana Haitlinger
Canestriniella physsana Haitlinger, 1989: 596
Type locality: Java
Distribution: Indonesia (Java, Sulawesi)
Hosts: Batocera celebiana, B. humilidens (Coleoptera: Cerambycidae)
Canestriniella togata Berlese
Canestriniella togata Berlese, 1910: 270
Type locality: Java
Distribution: Indonesia (Java)
Host: Batocera lunulata (Coleoptera: Cerambycidae)
Genus Caraboecius Cooreman
Caraboecius Cooreman, 1950: 18
Type species: Caraboecius coriacei Cooreman, 1950
Caraboecius coriacei Cooreman
Caraboecius coriacei Cooreman, 1950: 19
Type locality: Tangelanga
Distribution: Belgium
Host: Carabus (Procrustes) coriaceus (Coleoptera: Carabidae)
Genus Cetonicola Cooreman
Cetonicola Cooreman, 1955: 15
Type species: Cetonicola hispidus Cooreman, 1955
Cetonicola hispidus Cooreman
Cetonicola hispidus Cooreman, 1955: 16
Type locality: May-ya-Moto (Riv. Rutshuru)
166
Acarorum Catalogus IX
Distribution: D.R. Congo
Hosts: Dicronorhina micans, Smaragdesthes africana mutica, Eudicella gralli var. meckowi
(Coleoptera: Scarabaeidae, Cetoniinae)
Cetonicola isadorae Haitlinger
Cetonicola isadorae Haitlinger, 1991d: 296
Type locality: Nyangassy
Distribution: Cameroon
Host: Mecynorrhina [Chelorrhina] savagei (Coleoptera: Scarabaeidae, Cetoniinae)
Cetonicola maritae Haitlinger
Cetonicola maritae Haitlinger, 1992d: 297
Type locality: Distr. Kassai
Distribution: Congo-Brazzaville
Host: Stephanorrhina guttata (Coleoptera: Scarabaeidae, Cetoniinae)
Cetonicola orestesi Haitlinger
Cetonicola orestesi Haitlinger, 1992d: 294
Type locality: Liberia
Distribution: Ghana, Liberia
Host: Tmesorrhina iris (Coleoptera: Scarabaeidae, Cetoniinae)
Cetonicola vatus Haitlinger
Cetonicola vatus Haitlinger, 1990 : 323
Type locality: Tanzania
Distribution: Tanzania
Host: Poecilophila hebraea (Coleoptera: Scarabaeidae, Cetoniinae)
Genus Chelinochroa Summers et Schuster
Chelinochroa Summers & Schuster, 1982b: 41
Type species: Chelinochroa dictyophora Summers et Schuster, 1982
Chelinochroa dictyophora Summers et Schuster
Chelinochroa dictyophora Summers & Schuster, 1982b: 41
Type locality: Wajir
Distribution: Kenya
Host: Coleoptera: Cerambycidae
Genus Cnecoderma Summers et Schuster
Cnecoderma Summers & Schuster, 1981b: 39
Type species: Cnecoderma ovalis Summers et Schuster, 1981
Acarorum Catalogus IX 167
Cnecoderma dolichopoda Summers et Schuster
Cnecoderma dolichopoda Summers & Schuster, 1981b: 41
Type locality: Queensland
Distribution: Australia
Host: Passalidae (Coleoptera)
Cnecoderma ovalis Summers et Schuster
Cnecoderma ovalis Summers & Schuster, 1981b: 39
Type locality: Australia
Distribution: Australia
Host: (Coleoptera: Passalidae)
Genus Coleogyphus Berlese
Coleogyphus Berlese, 1910: 268
Type species: Coleogyphus fuscipes Berlese, 1910
Coleoglyphus fuscipes Berlese
Coleoglyphus fuscipes Berlese, 1910: 268
Type locality: Africa
Distribution: Africa
Hosts: Tefflus “megerlei” (Coleoptera: Carabidae). Nymph on Copris hamadryas (Coleoptera: Scarabaeidae)
Coleoglyphus simplex Berlese
Coleoglyphus (?) simplex Berlese, 1910b: 387
Type locality: Ecuador
Distribution: “Equatore” (Ecuador)
Host: Mitrocephala humboldti ()
Genus Coleopterophagus Berlese
Coleopterophagus Berlese, 1882: 9
Type species: Dermoglyphus megninii Berlese, 1881
Coleopterophagus albini Haitlinger
Coleopterophagus albini Haitlinger, 1990d: 328; 2002: 69; 2012: 187; 2015: 239; Trach &
Khaustov, 2011: 223
Type locality: Zaborze n. Polczyn Zdroj (Poland)
Distribution: Austria, Czech Rep., Estonia, Germany, Norway, Poland, Romania, Slovakia,
Ukraine
Hosts: Protaetia (Cetonischema) [Potosia] aeruginosa, P. (Liocola) marmorata, P. metallica
(Coleoptera: Scarabaeidae, Cetoniinae)
168
Acarorum Catalogus IX
Coleopterophagus baali Haitlinger
Coleopterophagus baali Haitlinger, 1990d: 330
Type locality: North Iran, Elburs Mts.
Distribution: Iran
Host: Potosia speciosa (Coleoptera: Scarabaeidae, Cetoniinae)
Coleopterophagus belzebubi Haitlinger
Coleopterophagus belzebubi Haitlinger, 1990d: 332
Type locality: Taihorim
Distribution: Taiwan
Host: Potosia aerata (Coleoptera: Scarabaeidae, Cetoniinae)
Coleopterophagus blattophagus (Banks)
Canestrinia blattophaga Banks, 1914: 61
Coleopterophagus blattophagus (Banks): Vitzthum, 1925: 48
Type locality: Panama
Distribution: Panama
Coleopterophagus dionizyi Haitlinger
Coleopterophagus dionizyi Haitlinger, 1990d: 330
Type locality: Syria
Distribution: Syria
Host: Potosia cuprea (Coleoptera: Scarabaeidae, Cetoniinae)
Coleopterophagus donaldi Haitlinger
Coleopterophagus donaldi Haitlinger, 1990d: 332; Trach & Khaustov, 2011: 223
Type locality: Bolzano (Italy)
Distribution: Italy, Ukraine
Host: Protaetia (Eupotosia) [Potosia] affinis (Coleoptera: Scarabaeidae, Cetoniinae)
Coleopterophagus maroni Haitlinger
Coleopterophagus maroni Haitlinger, 1990d: 336; Trach & Khaustov, 2011: 220
Type locality: Syria
Distribution: Romania, Syria, Ukraine
Hosts: Protaetia (Cetonischema) [Potosia] aeruginosa, P. speciosa jousselini (Coleoptera:
Scarabaeidae, Cetoniinae)
Coleopterophagus megnini (Berlese)
Dermoglyphus megnini Berlese, 1881: 750
Dermoglyphus megninii Berlese: Radford, 1950: 161
Acarorum Catalogus IX 169
Coleopterophagus megninii (Berlese): Berlese, 1882: A.M.S., fasc. I, No 9; Turk, 1953: 82;
Oudemans, 1923 : 205; 1937: 2155; Buitendijk, 1945: 364; Haitlinger, 1988d: 535;
2002: 69; 2012: 186
Coleopterophagus megnini (Berlese): Trach & Khaustov, 2011: 214; Haitlinger, 2012: 186
Coleopterophagus cetoniae Oudemans, 1923; Buitendijk, 1945: 264; Cooreman, 1954: 85
Type locality: Italy
Distribution: Croatia, Czech Rep., Great Britain, Holland, Hungary, Italy, Sicily, Poland,
Serbia, Turkey, Ukraine
Hosts: Cetonia aurata, Protaetia (Potosia) cuprina, Protaetia (Potosia) fieberi, Protaetia
(Potosia) metallica, Protaetia cuprea cuprea, P. cuprea ignicollis, P. cuprea obscura,
Protaetia (Eupotosia) affinis (Coleoptera: Scarabaeidae, Cetoniinae)
Coleopterophagus neglectus Berlese
Coleopterophagus neglectus Berlese, 1920: 147
Type locality: Africa
Distribution: Africa
Host: “Scarabeus” centaurus (Coleoptera: Scarabaeidae)
Coleopterophagus pulcher Cooreman
Coleopterophagus pulcher Cooreman, 1955: 23
Type locality: Rutshuru
Distribution: D.R. Congo
Host: Porphyronota cinnamonea (Coleoptera: Scarabaeidae, Cetoniinae)
Coleopterophagus quadrisetosus Trägårdh
Coleopterophagus quadrisetosus Trägårdh, 1904: 113
Type locality: Gebelein
Distribution: Sudan
Host: Coleoptera (Scarabaeidae, Cetoniinae)
Coleopterophagus rudolfi Haitlinger
Coleopterophagus rudolfi Haitlinger, 1990d: 334
Type locality: China
Distribution: China
Host: Potosia brevitarsis (Coleoptera: Scarabaeidae, Cetoniinae)
Genus Cooremania Nesbitt
Cooremania Nesbitt, 1976: 227
Type species: Cooremania wauensis Nesbitt, 1976
170
Acarorum Catalogus IX
Cooremania kietaensis Haitlinger et Šundić
Cooremania kietaensis Haitlinger & Šundić, 2020: 155
Type locality: Kieta
Distribution: Papua New Guinea
Host: Coleoptera (Lucanidae indet.)
Cooremania wauensis Nesbitt
Cooremania wauensis Nesbitt, 1976: 227
Type locality: Wau, Njau
Distribution: Papua New Guinea
Host: Eurytrachus arfakianus (Coleoptera: Lucanidae)
Genus Danaithiana Haitlinger
Danaithiana Haitlinger, 1991b: 253
Type species: Danaithiana chanchamayoica Haitlinger, 1991
Danaithiana archidonaica Haitlinger
Danaithiana archidonaica Haitlinger, 1991b: 257
Type locality: Archidona (Ecuador)
Distribution: Colombia, Ecuador
Host: Semiotus furcatus (Coleoptera: Elateridae)
Danaithiana ayraica Haitlinger
Danaithiana ayraica Haitlinger, 1991b: 254
Type locality: Laguayra (Ecuador)
Distribution: Colombia, Ecuador
Host: Semiotus virgatus (Coleoptera: Elateridae)
Danaithiana chanchamayoica Haitlinger
Danaithiana chanchamayoica Haitlinger, 1991b: 254
Type locality: Chanchamayo (Peru)
Distribution: Costa Rica, Cuba, Peru
Host: Semiotus imperialis (Coleoptera: Elateridae)
Danaithiana ecuadorica Haitlinger
Danaithiana ecuadorica Haitlinger, 1991b: 257
Type locality: Archidona
Distribution: Ecuador
Host: Semiotus furcatus (Coleoptera: Elateridae)
Acarorum Catalogus IX 171
Genus Dicanestrinia Berlese
Dicanestrinia Berlese, 1911: 184
Type species: Dermaleichus cerambycis G.Canestrini, 1878
Dicanestrinia cerambycis (G. Canestrini)
Dermaleichus cerambycis G. Canestrini, 1878: 69
Canestrinia (Dicanestrinia) cerambicis [sic]: Berlese, 1911: 184
Type locality: Italy
Distribution: Italy
Host: Cerambyx cerdo (Coleoptera: Cerambycidae)
Dicanestrinia huberti Haitlinger
Dicanestrinia huberti Haitlinger, 1994: 200; 2012: 186; Khaustov & Edelberg, 2001: 40
Type locality: Bystre n. Baligród (Bieszcady Mountains – voi. Krosno)(Poland)
Distribution: Byelorussia, Poland
Host: Carabus variolosus (Coleoptera: Carabidae)
Dicanestrinia knobi Samšinák
Dicanestrinia knobi Samšinák, 1971: 177; Haitlinger, 1988a: 535; 2012: 186; Trach, 2006: 175
Type locality: Carpathia, Užok (Ukraine)
Distribution: Czech Rep., Bosnia and Herzegovina, Italy, Poland, Ukraine
Host: Carabus variolosus, C.v. nodulosus (Coleoptera: Carabidae)
Genus Diplognathophilus Cooreman
Diplognathophilus Cooreman, 1955: 14
Type species: Diplognathophilus africanus Cooreman, 1955 (56?)
Diplognathophilus africanus Cooreman
Diplognathophilus africanus Cooreman, 1955: 14
Type locality: Ndeko (Katanda)
Distribution: D.R. Congo
Host: Rhabdotis sobrina (Coleoptera: Scarabaeidae, Cetoniinae)
Genus Diplopodocoptes Fain
Diplopodocoptes Fain, 1987: 167
Type species: Diplopodocoptes transkeiensis Fain, 1987
Diplopodocoptes transkeiensis Fain
Diplopodocoptes transkeiensis Fain, 1987: 168
Type locality: Transkei, District Lusikisiki, Mouth of Riv. Mzimhlava
Distribution: Kenya, South Africa
172
Acarorum Catalogus IX
Hosts: millipede (Diplopoda: Glomeridae); paratypes from Odontopygidae (Diplopoda)
from Kenya
Genus Donnelaphontia Lavoipierre
Donnelaphontia Lavoipierre, 1959: 611
Farahanella Haitlinger, 1990: 314; [Type is suatoha; Syn. Haitlinger 2001: 104]
Type species: Donnelaphontia calostegis Lavoipierre, 1959
Donnelaphontia calostegis Lavoipierre
Donnelaphontia calostegis Lavoipierre, 1959: 611
Type locality: Kumba
Distribution: Cameroon
Host: Calostega crassicornis (Coleoptera: Tenebrionidae)
Donnelaphontia suatoha (Haitlinger)
Farahanella suatoha: Haitlinger, 1990: 315
Donnelaphontia suatoha: Haitlinger, 2001: 104
Type locality: “Zaire”
Distribution: DR Congo
Host: Odontopezus cupreus (Coleoptera: Tenebrionidae)
Genus Dorynothiana Haitlinger
Dorynothiana Haitlinger, 1999: 43
Type species: Dorynothiana jurandi Haitlinger, 1999
Dorynothiana delmari Haitlinger
Dorynothiana delmari Haitlinger, 2008: 66
Type locality: Brazil
Distribution: Brazil
Host: Dorynota sp. (Coleoptera: Chrysomelidae, Cassidinae)
Dorynothiana jurandi Haitlinger
Dorynothiana jurandi Haitlinger, 1999: 44
Type locality: Santa Catarina
Distribution: Brazil
Host: Dorynota pugionata (Coleoptera: Chrysomelidae, Cassidinae)
Genus Ennodithiana Haitlinger
Ennodithiana Haitlinger, 1993d: 66
Type species: Ennodithiana messalinae Haitlinger, 1993
Acarorum Catalogus IX 173
Ennodithiana maniusi Haitlinger
Ennodithiana maniusi Haitlinger, 1993d: 68
Type locality: Brazil, Parana
Distribution: Brazil, Mexico
Hosts: Omaspides basilica, Psalidonota contemta, P. dorsoplagiata (Coleoptera: Chrysomelidae, Cassidinae)
Ennodithiana medoni Haitlinger
Ennodithiana medoni Haitlinger, 1993d: 71
Type locality: Brazil
Distribution: Brazil
Host: Acromis spinifex (Coleoptera: Chrysomelidae, Cassidinae)
Ennodithiana megesi Haitlinger
Ennodithiana megesi Haitlinger, 1993d: 71
Type locality: Goyaz
Distribution: Brazil
Host: Crambelea illuderis (Coleoptera: Chrysomelidae, Cassidinae)
Ennodithiana messalinae Haitlinger
Ennodithiana messalinae Haitlinger, 1993d: 66
Type locality: San Carlos
Distribution: Costa Rica, Cuba
Hosts: Psalidonota dorsopunctata, P. leprosa (Coleoptera: Chrysomelidae, Cassidinae)
Ennodithiana penelopae Haitlinger
Ennodithiana penelopae Haitlinger, 1993: 68
Type locality: Paramaribo (Suriname)
Distribution: Brazil, Suriname
Host: Acromis nebulosa (Coleoptera: Chrysomelidae, Cassidinae)
Ennodithiana platysma (Summers et Schuster)
Grandiella platysma Summers & Schuster, 1979: 335
Ennodithiana platysma (Summers & Schuster): Haitlinger, 1993d: 73
Type locality: Barro Colorado Isl.
Distribution: Panama
Host: Coptocycla sp. (Coleoptera: Chrysomelidae, Cassidinae)
Genus Eunorithiana Haitlinger
Eunorithiana Haitlinger, 1999: 46
Type species: Eunorithiana aidae Haitlinger, 1999
174
Acarorum Catalogus IX
Eunorithiana aidae Haitlinger
Eunorithiana aidae Haitlinger, 1999: 47; 2001: 103
Type locality: Rio Cantino
Distribution: Colombia
Host: Plagiometriona phoebe (Coleoptera: Chrysomelidae, Cassidinae)
Eunorithiana heliodori Haitlinger
Eunorithiana heliodori Haitlinger, 1999: 49
Type locality: Santa Cruz, Santa Catarina
Distribution: Brazil
Host: Plagiometriona ludiera, P. punctipennis (Coleoptera: Chrysomelidae, Cassidinae)
Genus Gasuthiana Haitlinger
Gasuthiana Haitlinger, 1989: 281
Type species: Gasuthiana abapoica Haitlinger, 1989
Gasuthiana abapoica Haitlinger
Gasuthiana abapoica Haitlinger, 1989: 282
Type locality: Brazil
Distribution: Brazil
Host: Alurnus elysianus (Coleoptera: Chrysomelidae, Hispinae)
Gasuthiana abufarica Haitlinger
Gasuthiana abufarica Haitlinger, 1989: 287
Type locality: Bolivia
Distribution: Bolivia
Host: Meristomala nigripes (Coleoptera: Chrysomelidae, Hispinae)
Gasuthiana coarica Haitlinger
Gasuthiana coarica Haitlinger, 1989: 289
Type locality: Brazil
Distribution: Brazil
Host: Mecistomala marginatus (Coleoptera: Chrysomelidae, Hispinae)
Gasuthiana melillae Haitlinger
Gasuthiana melillae Haitlinger, 1995a: 83
Type locality: Brazil
Distribution: Brazil
Host: Tenebrionidae indet. (Coleoptera)
Gasuthiana olareica Haitlinger
Gasuthiana olareica Haitlinger, 1989: 284
Acarorum Catalogus IX 175
Type locality: Colombia
Distribution: Colombia, French Guiana
Hosts: Alurnus bipunctatus, A. boulardi (Coleoptera: Chrysomelidae, Hispinae)
Genus Gioharattia Haitlinger
Gioharattia Haitlinger, 1989a: 57
Type species: Gioharattia maharae Haitlinger, 1989
Gioharattia caroae Haitlinger
Gioharattia caroae Haitlinger, 1989a: 59
Type locality: “South Ceylon”
Distribution: Sri Lanka
Host: Coryphocera sp. (Coleoptera: Scarabaeidae, Cetoniinae)
Gioharattia maharae Haitlinger
Gioharattia maharae Haitlinger, 1989a: 57
Type locality: Borneo
Distribution: Borneo (?Indonesia)
Host: Coglodera diaroi (Coleoptera: Scarabaeidae, Cetoniinae)
Genus Globosophotia Haitlinger
Globosophotia Haitlinger, 1991a: 279
Type species: Globosophotia bettinae Haitlinger, 1991
Globosophotia bettinae Haitlinger
Globosophotia bettinae Haitlinger, 1991a: 279
Type locality: Morocco
Distribution: Morocco
Host: Pseudosteropus globosus (Coleoptera: Carabidae, Pterostichinae)
Genus Grandiella Lombardini
Grandiella Lombardini, 1939: 45 [considered by Fain, 1989 praeocc. from Williams 1928
Hymenoptera, but being nomen nudum, Grandiella Lombardini has been restored
by OConnor, 2004]
Grandiellina Fain, 1989: 67
Type species: Grandiella tetracaudata Lombardini, 1939
Grandiella cooremani Fain
Grandiella cooremani Fain, 1987: 106; OConnor, 2004: 120
Grandiellina cooremani (Fain): Fain, 1989: 67
Type locality: Ecuador
176
Acarorum Catalogus IX
Distribution: Ecuador
Host: Canistra osculatii (Coleoptera: Chrysomelidae)
Grandiella decemcaudata Lombardini
Grandiella decemcaudata Lombardini, 1950: 289; OConnor, 2004: 120
Grandiellina decemcaudata (Lombardini): Fain, 1989: 67
Type locality: Bolivia, Zapirangai
Distribution: Bolivia, Paraguay
Hosts: Echoma [as Omoplata] marginata, Coptocycla contemta (Coleoptera: Chrysomelidae, Cassidinae)
Grandiella ecuadorensis Fain
Grandiella ecuadorensis Fain, 1987: 101; OConnor, 2004: 120
Grandiellina ecuadorensis (Fain): Fain, 1989: 67
Type locality: Ecuador
Distribution: Ecuador
Host: Canistra osculatii (Coleoptera: Chrysomelidae)
Grandiella gambosa Summers et Schuster
Grandiella gambosa Summers & Schuster, 1979: 343; Fain, 1987: 94
Grandiellina gambosa (Summers & Schuster): Fain, 1989: 67
Type locality: El Llano Carti Rd., 10 km N El Llano, Pina, Panama
Distribution: Belize, Colombia, Panama
Hosts: Discomorpha batesi, D. lanuginosa pubescens, D. depilata, Dolichotoma
conspersipennis (Coleoptera: Chrysomelidae)
Grandiella joliveti Fain
Grandiella joliveti Fain, 1987: 97
Grandiellina joliveti (Fain): Fain, 1989: 67
Type locality: Vigosa, Minas Gerais
Distribution: Brazil
Host: Elytrosphaera xanthopyga (Coleoptera: Chrysomelidae)
Grandiella kimseyi Summers et Schuster
Grandiella kimseyi Summers & Schuster, 1979a: 349; Fain, 1987: 94; OConnor, 2004: 120
Grandiellina kimseyi (Summers et Schuster): Fain, 1989: 67
Type locality: El Llano Carti Rd., Pina (Panama)
Distribution: Belize, Colombia, Panama
Hosts: Discomorpha batesi, Dolichotoma conspersipennis (Coleoptera: Chrysomelidae)
Acarorum Catalogus IX 177
Grandiella longini (Haitlinger)
Grandiellina longini Haitlinger, 1991e: 389
Grandiella longini (Haitlinger): OConnor, 2004: 120
Type locality: Bolivia
Distribution: Bolivia
Host: Goniochenia (Baranosa) flavosparsa (Coleoptera: Chrysomelidae, Cassidinae)
Grandiella marinae (Haitlinger)
Grandiellina marinae Haitlinger, 1991e: 390
Grandiella marinae (Haitlinger): OConnor, 2004: 120
Type locality: Chiapas
Distribution: Mexico
Host: Ehoma confluens (Coleoptera: Chrysomelidae)
Grandiella minor Fain
Grandiella minor Fain, 1987: 106; OConnor, 2004: 120
Grandiellina minor (Fain): Fain, 1989: 67
Type locality: Santarem
Distribution: Brazil
Host: Canistra dohrni (Coleoptera: Chrysomelidae)
Grandiella octocaudata Lombardini
Grandiella octocaudata Lombardini, 1950: 288; OConnor, 2004: 120
Grandiellina octocaudata (Lombardini): Fain, 1989: 67
Type locality: Estado de Paraná, Bariqui
Distribution: Brazil
Host: Stolas chalybaea (Coleoptera: Chrysomelidae)
Grandiella pentagona Lombardini
Grandiella pentagona Lombardini, 1950: 286; OConnor, 2004: 120
Grandiellina pentagona (Lombardini): Fain, 1989: 67
Type locality: Distrito Federal, Cerro de Naiguatá
Distribution: Venezuela
Host: Coleoptera: Chrysomelidae, Cassidinae
Grandiella rogeri (Haitlinger)
Grandiellina rogeri Haitlinger, 1991e: 388
Grandiella rogeri (Haitlinger): OConnor, 2004: 120
Type locality: Chanchamayo
Distribution: Peru
Host: Goniochenia (Baranosa) heroldi (Coleoptera: Chrysomelidae, Cassidinae)
178
Acarorum Catalogus IX
Grandiella rosascostai (Lombardini)
Coleoglyphus rosascostai Lombardini, 1950: 279
Hatohylla rosascostai (Lombardini): Haitlinger 1991: 311
Grandiella rosascostai (Lombardini): OConnor, 2004: 120
Type locality: Distrito Federal Cerro del Naiguatá
Distribution: Venezuela
Host: Nyctobates sp. (Coleoptera: Tenebrionidae)
Grandiella rugosita Summers et Schuster
Grandiella rugosita Summers & Schuster, 1979: 343; OConnor, 2004: 120
Type locality: Panama
Distribution: Panama
Host: Coleoptera
Grandiellina sicula Summers et Schuster
Grandiella sicula Summers & Schuster, 1979a: 333; OConnor, 2004: 120
Grandiellina sicula (Summers & Schuster): Fain, 1989: 67; Haitlinger, 1991e: 392
Type locality: Depto. Del Cesar, 0.3 km N Castilla, Inez Ranch
Distribution: Colombia
Hosts: Coptocycla nigropunctata, C. nr. wagneri (Coleoptera: Chrysomelidae, Cassidinae)
Grandiella soniae (Haitlinger)
Grandiellina soniae Haitlinger, 1991e: 391
Grandiella soniae (Haitlinger): OConnor, 2004: 120
Type locality: Rio Negro
Distribution: Colombia
Hosts: Cyrtonota punicea, C. textilis, Stolas coalita, S. coerulescens, S. erichsoni, S. placida
(Coleoptera: Chrysomelidae)
Grandiella tacjannae (Haitlinger)
Grandiellina tacjannae Haitlinger, 1991e: 383
Grandiella tacjannae (Haitlinger): OConnor, 2004: 120
Type locality: Brazil, Candido de Abreu
Distribution: Argentina, Brazil, Paraguay, Peru
Hosts: Anacassis fuscata, Botanochara angulata, B. bonariensis, B. convexiuscula, B. duodecimverrucata, B. impressa, B. macularia, B. nervosa, B. regina, B. ruforeticulata, B.
sanguinea, B. sedecimpustulata, B. tesselata, B. texta (Coleoptera: Chrysomelidae,
Cassidinae)
Grandiella tertia Lombardini
Grandiella tertia Lombardini, 1942: 46; OConnor, 2004: 120
Acarorum Catalogus IX 179
Grandiellina tertia (Lombardini): Fain, 1989: 67
Type locality:Brazil
Distribution: Brazil
Host: Cassids (Coleoptera: Chrysomelidae)
Grandiella tetracaudata Lombardini
Grandiella tetracaudata Lombardini, 1939: 45; 1950: 283; Summers & Schuster, 1979a:
330; Fain, 1987: 94; OConnor, 2004: 120
Grandiellina tetracaudata (Lombardini): Fain, 1989: 67; Haitlinger, 1991e : 383
Type locality: Brazil
Distribution: Brazil, Colombia, Peru
Hosts: [Botanochara sedecimpustulata, B. subnervosa], Coptocycla adamantina, Cyrtonota
cyanea, C. thalassina, C. tigrina, Eumolpus fulgidus, Stolas chalybaea, S. subrugosa
(Coleoptera: Chrysomelidae). According to Haitlinger (1992e), the mites recorded
on genus Botanochara belong actually to Grandiellina tacjannae.
Grandiella tetracaudata omoplata Fain
Grandiella tetracaudata omoplata Fain, 1987 : 94; OConnor, 2004: 120
Grandiellina tetracaudata omoplata (Fain): Fain, 1989: 67
Type locality: Cayenne, St. Jean de Maroni, Gourdonville
Distribution: French Guyana
Host: Eichoma clypeata (= Omoplata marginata) (Coleoptera: Chrysomelidae, Cassidinae)
Genus Grandiellopsis Fain
Grandiellopsis Fain, 1989: 68
Type species: Grandiellopsis cyclosoma Fain, 1989
Grandiellopsis alani Haitlinger
Grandiellopsis alani Haitlinger, 1992: 52
Type locality: Archidona
Distribution: Ecuador
Host: Canistra osculatii (Coleoptera: Chrysomelidae)
Grandiellopsis ariani Haitlinger
Grandiellopsis ariani Haitlinger, 1992: 53
Type locality: Campirias
Distribution: Brazil
Host: Canistra carbonaria (Coleoptera: Chrysomelidae)
Grandiellopsis canistra Fain
Grandiellopsis canistra Fain, 1989: 70
180
Acarorum Catalogus IX
Type locality: Ecuador
Distribution: Ecuador
Host: Canistra osculatii (Coleoptera: Chrysomelidae)
Grandiellopsis cyclosoma Fain
Grandiellopsis cyclosoma Fain, 1989: 68
Type locality: Bas Maroni
Distribution: French Guyana
Host: Cyclosoma palliata (Coleoptera: Chrysomelidae)
Grandiellopsis odiosi Haitlinger
Grandiellopsis odiosi Haitlinger, 1992: 56
Type locality: Brazil
Distribution: Brazil
Host: Polychalca (P.) punctatissima (Coleoptera: Chrysomelidae)
Grandiellopsis orsenae Haitlinger
Grandiellopsis orsenae Haitlinger, 1992: 58
Type locality: Puerto Inca
Distribution: Peru
Host: Cyclosoma mirabilis (Coleoptera: Chrysomelidae)
Grandiellopsis violae Haitlinger
Grandiellopsis violae Haitlinger, 1992: 55
Type locality: Bolivia
Distribution: Bolivia
Host: Canistra cruentata (Coleoptera: Chrysomelidae)
Genus Haitlingeria Wontae Kim et al.
Haitlingeria Kim et al., 2006: 13
Type species: Haitlingeria longilobata Kim et al., 2006
Haitlingeria longilobata Kim et al.
Haitlingeria longilobata Kim et al., 2006: 13
Type locality: South Korea, near the Bung-Dong Reservoir Seonbukcong Yuseongku
Daejeon
Distribution: Korea
Host: Serrognathus platymelus (Coleoptera: Lucanidae)
Genus Hargeinella Haitlinger
Hargeinella Haitlinger, 1989a: 53
Acarorum Catalogus IX 181
Type species: Hargeinella poppana Haitlinger, 1989
Hargeinella poppana Haitlinger
Hargeinella poppana Haitlinger, 1989a: 53
Type locality: Sumatra
Distribution: Indonesia (Sumatra)
Host: Trichogomphus milon (Coleoptera: Scarabaeidae, Dynastinae)
Genus Hatohylla Haitlinger
Hatohylla Haitlinger, 1990a: 307 1991 ?
Type species: Hatohylla morrata Haitlinger, 1990
Hatohylla bahiana Haitlinger
Hatohylla bahiana Haitlinger, 1990a: 308
Type locality: Santa Catarina
Distribution: Brazil
Host: Nyctobates maximus (Coleoptera: Tenebrionidae)
Hatohylla bibiannae Haitlinger
Hatohylla bibiannae Haitlinger, 1990a: 311
Type locality: Santa Catarina
Distribution: Brazil
Host: undeternined Tenebrionidae (Coleoptera)
Hatohylla morrata Haitlinger
Hatohylla morrata Haitlinger, 1990a: 308
Type locality: Amazonas
Distribution: Brazil
Host: Nyctobates gigas (Coleoptera: Tenebrionidae)
Genus Hyllosihemia Haitlinger
Hyllosihemia Haitlinger, 1994: 93
Type species: Hyllosihemia belerofoni Haitlinger, 1994
Hyllosihemia belerofoni Haitlinger
Hyllosihemia belerofoni Haitlinger, 1994: 93
Type locality: Sumatra
Distribution: Indonesia (Sumatra)
Host: Coleoptera: Passalidae
Genus Hypopteryx Summers et Schuster
Hypopteryx Summers & Schuster, 1981b: 41
182
Acarorum Catalogus IX
Type species: Hypopteryx collaris Summers et Schuster, 1981
Hypopteryx collaris Summers et Schuster
Hypopteryx collaris Summers & Schuster, 1981b: 42
Type locality: Australia
Distribution: Australia
Host: Coleoptera
Genus Ibizella Haitlinger et Šundić
Ibizella Haitlinger & Šundić, 1916: 75
Type species: Ibizella balearica Haitlinger & Šundić, 1916
Ibizella asilahica Haitlinger et Šundić
Ibizella asilahica Haitlinger et Šundić, 1918: 413
Type locality: Asilah (31 km south of Tangier)
Distribution: Morocco
Host: Coleoptera: Tenebrionidae indet.
Ibizella balearica Haitlinger et Šundić
Ibizella balearica Haitlinger & Šundić, 1916: 75
Type locality: Ibiza
Distribution: Spain (Ibiza)
Host: (Coleoptera, Tenebrionidae: Blaps sp.)
Ibizella samsinaki (Beron)
Canestrinia samsinaki Beron, 1975: 83; Haitlinger, 1993c: 87; 2001a: 26; Haitlinger &
Šundić, 1916: 75
Ibizella samsinaki (Beron): Haitlinger & Šundić, 1916: 75
Type locality: Bulgaria, Sozopol
Distribution: Bulgaria, Cyprus, Spain (Ibiza)
Host: Blaps sp., Gnaptor sp. (Coleoptera: Tenebrionidae)
Genus Irmongia Haitlinger
Irmongia Haitlinger, 1990b: 547
Type species: Irmongia helgae Haitlinger, 1990
Irmongia helgae Haitlinger
Irmongia helgae Haitlinger, 1990b: 547
Type locality: Tanzania
Distribution: Tanzania
Host: undetermined Passalidae (Coleoptera)
Acarorum Catalogus IX 183
Genus Jowitella Haitlinger
Jowitella Haitlinger, 1990b: 547
Type species: Jowitella barbarae Haitlinger, 1990
Jowitella barbarae Haitling
Jowitella barbarae Haitlinger, 1990b: 547
Type locality: Santa Catarina
Distribution: Brazil
Host: undetermined Passalidae (Coleoptera)
Genus Jullongia Haitlinger
Jullongia Haitlinger, 1990b: 538
Type species: Jullongia izae Haitlinger, 1990
Jullongia izae Haitlinger
Jullongia izae Haitlinger, 1990b: 539
Type locality: Stephansort
Distribution: Papua New Guinea
Host: undetermined Passalidae (Coleoptera)
Genus Kahoorangia Haitlinger
Kahoorangia Haitlinger, 1992: 288
Type species: Kahoorangia almae Haitlinger, 1992
Kahoorangia almae Haitlinger
Kahoorangia almae Haitlinger, 1991d: 289
Type locality: ? South Africa
Distribution: ? South Africa
Host: Pachnoda carmelita (Coleoptera: Scarabaeidae, Cetoniinae)
Genus Lidiophela Haitlinger
Lidiophela Haitlinger, 1990b: 540
Type species: Lidiophela pelagiae Haitlinger, 1990
Lidiophela aruica Haitlinger
Lidiophela aruica Haitlinger, 1990b: 542
Type locality: Aru Is.
Distribution: Indonesia (Aru Is.)
Host: Labienus ptox (Coleoptera: Passalidae)
Lidiophela bognae Haitlinger
Lidiophela bognae Haitlinger, 1990b: 543
184
Acarorum Catalogus IX
Type locality: Moluccas, Batjan Is.
Distribution: Indonesia (Moluccas, Batjan Is.)
Host: undetermined Passalidae (Coleoptera)
Lidiophela celinae Haitlinger
Lidiophela celinae Haitlinger, 1990b: 541
Type locality: Moluccas, Ambon Is.
Distribution: lndonesia, Moluccas (Ambon Is.)
Host: undetermined Passalidae (Coleoptera)
Lidiophela igae Haitlinger
Lidiophela igae Haitlinger, 1990b: 542
Type locality: Moluccas, Batjan Is.
Distribution: Indonesia (Batjan Is.)
Host: Labienus gigas (Coleoptera: Passalidae)
Lidiophela pauliana Haitlinger
Lidiophela pauliana Haitlinger, 1991a: 276
Type locality: Australia
Distribution: Australia
Host: Amarygmus striatus (Coleoptera: Tenebrionidae)
Lidiophela pecki (Nesbitt)
Coleopterophagus pecki Nesbitt, 1976: 220
Lidiophela pecki: Haitlinger, 1990b: 540
Type locality: Baiyer River Sanctuary, Mt.Hagen, Papua New Guinea
Distribution: Papua New Guinea, Solomon Islands (Gizo Is.),
Hosts: Labienus sp., Obiensus sp., Protomocoelus australis (Coleoptera: Passalidae)
Lidiophela pelagiae Haitlinger
Lidiophela pelagiae Haitlinger, 1990b: 540
Type locality: Moluccas, Batjan Is.
Distribution: Indonesia, Moluccas (Batjan Is.)
Host: Labienus gigas (Coleoptera: Passalidae)
Genus Megacanestrinia Trägårdh
Megacanestrinia Trägårdh, 1906: 877
Type species: Megacanestrinia mucronata Trägårdh, 1906
Megacanestrinia beloniana Haitlinger
Megacanestrinia beloniana Haitlinger, 1990e: 312
Type locality: Tanzania
Acarorum Catalogus IX 185
Distribution: Tanzania
Host: Tefflus purpuripennis (Coleoptera: Carabidae)
Megacanestrinia mucronata Trägårdh
Megacanestrinia mucronata Trägårdh, 1906: 877
Type locality: Umfolozi, Zululand
Distribution: South Africa
Host: «Auf einen grossen Carabide« (Coleoptera: Carabidae)
Subfam. Melisiinae Summers et Schuster
Melisiinae Summers & Schuster, 1982a: 23
Type genus : Melisia Lombardini, 1944
Genus Melisia Lombardini
Melisia Lombardini, 1944: 2
Type species: Melisia melisii Lombardini, 1944
Melisia anomala Samšinák
Melisia anomala Samšinák, 1963: 257
Type locality: Mts Singalang
Distribution: Indonesia (Sumatra)
Hosts: Epiphanus indicus, E. stoliczkae (Coleoptera: Passalidae)
Melisia ariadnae Haitlinger
Melisia ariadnae Haitlinger, 1989: 529
Type locality: Soekaranda (Sumatra); Borneo
Distribution: lndonesia (Sumatra, Borneo)
Host: Aceraius laevicollis (Passalidae, Coleoptera)
Melisia baloghi Samšinák
Melisia baloghi Samšinák, 1969: 407
Type locality: Cuc-phuong Bong
Distribution: Vietnam
Host: Coleoptera indet.
Melisia gabrielae Haitlinger
Melisia gabrielae Haitlinger, 1990: 531
Type locality: Soekaranda, Sumatra
Distribution: Indonesia (Sumatra)
Host: Aceraius laevicollis (Coleoptera: Passalidae)
186
Acarorum Catalogus IX
Melisia helenae Haitlinger
Melisia helenae Haitlinger, 1990b: 533
Type locality: North Vietnam
Distribution: Vietnam
Host: undetermined Passalidae (Coleoptera)
Melisia hieronimae Haitlinger
Melisia hieronimae Haitlinger, 1990b: 535
Type locality: holotype from North Vietnam; paratypes from Thailand and Sumatra
Distribution: Indonesia (Sumatra), Thailand, Vietnam
Host: undetermined Passalidae (Coleoptera)
Melisia ingeborgae Haitlinger
Melisia ingeborgae Haitlinger, 1990b: 535
Type locality: Soekaranda, Sumatra
Distribution: Indonesia (Sumatra)
Host: undetermined Passalidae (Coleoptera)
Melisia irenae Haitlinger
Melisia irenae Haitlinger, 1990b: 531
Type locality: Sinabong, Sumatra
Distribution: Indonesia (Sumatra)
Host: Aceraius grandis (Coleoptera: Passalidae)
Melisia jadwigae Haitlinger
Melisia jadwigae Haitlinger, 1990b: 532
Type locality: North Vietnam
Distribution: Vietnam
Host: undetermined Passalidae (Coleoptera)
Melisia klapaleki Samšinák
Melisia klapaleki Samšinák, 1963: 260
Type locality: India mer.: Coimbatore
Distribution: India
Host: Epiphenus stoliczkae (Coleoptera: Passalidae)
Melisia leptinotarsa Summers et Schuster
Melisia leptinotarsa Summers & Schuster, 1982a: 25
Type locality: Paluma, Stop 388, Queensland
Distribution: Australia
Host: Coleoptera Passalidae (gen. sp. Indet.)
Acarorum Catalogus IX 187
Melisia lombardinii Samšinák
Melisia lombardinii Samšinák, 1963: 255
Type locality: Mts Singalang
Distribution: Indonesia (Sumatra)
Host: Pelopides mniszecki (Coleoptera: Passalidae)
Melisia mahunkai Samšinák
Melisia mahunkai Samšinák, 1969: 409
Type locality: Cuc-phuong, Bong
Distribution: Vietnam
Host: Coleoptera indet.
Melisia mariettae Haitlinger
Melisia mariettae Haitlinger, 1990b: 528
Type locality: Sinabong, Sumatra
Distribution: Indonesia (Sumatra)
Host: Aceraius grandis (Coleoptera: Passalidae)
Melisia melaniae Haitlinger
Melisia melaniae Haitlinger, 1990b: 532
Type locality: Sumatra
Distribution: Indonesia (Sumatra)
Host: Aceraius mayeri (Coleoptera: Passalidae)
Melisia melisii Lombardini
Melisia melisii Lombardini, 1944: 2; Samšinák, 1963: 253
Type locality: America
Distribution: America
Host: Passalus sp. (Coleoptera: Passalidae)
Melisia occidii Samšinák
Melisia occidii Samšinák, 1969: 410; Haitlinger, 1990b: 532
Type locality: Cuc-phuong, Bong
Distribution: Vietnam
Host: Coleoptera indet.
Melisia rutae Haitlinger
Melisia rutae Haitlinger, 1990b: 533
Type locality: North Vietnam
Distribution: Vietnam
Host: undetermined Passalidae (Coleoptera)
188
Acarorum Catalogus IX
Melisia salomeae Haitlinger
Melisia salomeae Haitlinger, 1990b: 529
Type locality: Sinabong, Sumatra
Distribution: Indonesia (Sumatra)
Host: Aceraius grandis (Coleoptera: Passalidae)
Melisia samsinaki Vomero
Melisia samsinaki Vomero, 1972: 95
Type locality: Kon Techore
Distribution: Cambodja
Host: Aceraius helferi (Coleoptera: Passalidae)
Melisia sentosa Summers et Schuster
Melisia sentosa Summers & Schuster, 1982a: 27
Type locality: Ta Rau, Hainan
Distribution: China (Hainan Isl.)
Host: Passalidae gen. sp. indet. (Coleoptera)
Melisia superba Summers et Schuster
Melisia superba Summers & Schuster, 1982a: 27
Type locality: Musha, Taiwan
Distribution: Taiwan
Host: Aceraius grandis hirsutus (Coleoptera: Passalidae)
Genus Mesophotia Samšinák
Mesophotia Samšinák, 1971: 209
Type species: Mesophotia penicillata Samšinák, 1971
Mesophotia penicillata penicillata Samšinák
Mesophotia penicillata penicillata Samšinák, 1971: 209
Type locality: Sevilla
Distribution: Spain
Host: Carabus (Rhabdotocarabus) melancholicus, C.m. costatus (Coleoptera: Carabidae)
Mesophotia penicillata similis Samšinák
Mesophotia penicillata similis Samšinák, 1971: 214
Type locality: Andalusia
Distribution: Spain
Host: Carabus (Rhabdotocarabus) melancholicus costatus (Coleoptera: Carabidae)
Acarorum Catalogus IX 189
Mesophotia penicillata tertia Samšinák
Mesophotia penicillata tertia Samšinák, 1971: 214
Type locality: Sevilla
Distribution: Morocco, Spain
Host: Carabus (Rhabdotocarabus) melancholicus (Coleoptera: Carabidae)
Genus Metriothiana Haitlinger
Metriothiana Haitlinger, 1999: 40
Type species: Metriothiana bolebori Haitlinger, 1999
Metriothiana bolebori Haitlinger
Metriothiana bolebori Haitlinger, 1999: 40
Type locality: Tegusigalpa
Distribution: Honduras
Host: Metrionella bilimeki (Coleoptera: Chrysomelidae, Cassidinae)
Genus Mossongia Haitlinger
Mossongia Haitlinger, 1989a: 49
Type species: Mossongia ssavina Haitlinger, 1989
Mossongia bissaina Haitlinger
Mossongia bissaina Haitlinger, 1989a: 52
Type locality: Bougainville, Kieta
Distribution: Papua New Guinea
Host: Scapanus solomonensis (Coleoptera: Scarabaeidae, Dynastinae)
Mossongia ssavina Haitlinger
Mossongia ssavina Haitlinger, 1989a: 50
Type locality: New Guinea
Distribution: New Guinea
Host: Scapanus australis (Coleoptera: Scarabaeidae, Dynastinae)
Genus Neophotia Samšinák
Neophotia Samšinák, 1971: 201
Type species: Neophotia drvotocka Samsinák, 1971
Neophotia drvotocka Samšinák
Neophotia drvotocka Samšinák, 1971: 202
Type locality: San Ildefonso
Distribution: Portugal, Spain
Hosts: Carabus (Iniopachus) pyrenaeus, C. (Oreocarabus) amplipennis, C. (Oreocarabus)
guadarramus (Coleoptera: Carabidae)
190
Acarorum Catalogus IX
Neophotia jureceki Samšinák
Neophotia jureceki Samšinák, 1971: 204
Type locality: Pyrenees, Pic du Midi
Distribution: France
Host: Carabus (Iniopachus) pyrenaeus (Coleoptera: Carabidae)
Neophotia latissima Samšinák
Neophotia latissima Samšinák, 1971: 206
Type locality: Guadarrama, Cercedilla
Distribution: Spain
Host: Carabus (Mesocarabus) lusitanicus (Coleoptera: Carabidae)
Genus Noemiphela Haitlinger
Noemiphela Haitlinger, 1992b: 536
Type species: Noemiphela izabelae Haitlinger, 1992
Noemiphela barryi Haitlinger
Noemiphela barryi Haitlinger, 1992b: 538
Type locality: Sumatra
Distribution: Indonesia
Host: Eurytrachelus purpurescens (Coleoptera: Lucanidae)
Noemiphela izabelae Haitlinger
Noemiphela izabelae Haitlinger, 1992b: 537
Type locality: Celebes (Sulawesi)
Distribution: Indonesia
Host: Cladognathus giraffa (Coleoptera: Lucanidae)
Genus Oceogavia Haitlinger
Oceogavia Haitlinger, 1989: 586
Type species: Oceogavia ubaldi Haitlinger, 1989
Oceogavia ubaldi Haitlinger
Oceogavia ubaldi Haitlinger, 1989: 586
Type locality: unknown
Distribution: unknown
Host: Papuana woodlarkiana (Coleoptera: Scarabaeidae, Dynastinae)
Genus Olafithiana Haitlinger
Olafithiana Haitlinger, 1993e: 274
Type species: Olafithiana adelinae Haitlinger, 1993
Acarorum Catalogus IX 191
Olafithiana adelinae Haitlinger
Olafithiana adelinae Haitlinger, 1993e: 274
Type locality: Parana
Distribution: Brazil
Host: [Stolas] Anacassis cribrum (Coleoptera: Chrysomelidae, Cassidinae)
Olafithiana foliacea (Fain)
Grandiella foliacea Fain, 1987: 101
Grandiellina foliacea (Fain): Fain, 1989: 67
Olafithiana foliacea (Fain): Haitlinger 1993e: 277
Type locality: Brazil, Nova Teutonia
Distribution: Argentina, Brazil, Panama
Hosts: Stolas areolata, S. chalybea, S. conspersa, S. complexa, S. festiva, S. impluviata, S.
lacunose, S. lenis, S. nudicollis, S. oblita, S. paranensis, S. subreticulata (Coleoptera:
Chrysomelidae, Cassidinae)
Olafithiana iwettae Haitlinger
Olafithiana iwettae Haitlinger, 1994b: 83
Type locality: Costa Rica
Distribution: Costa Rica, Venezuela
Host: Charidotella (= Metriona) testudinacea (Coleoptera: Chrysomelidae, Cassidinae)
Olafithiana joliveti (Fain)
Grandiella joliveti Fain, 1987: 97
Olafithiana joliveti (Fain): Haitlinger 1993e: 278
Type locality: Viçosa, Minas Gerais
Distribution: Brazil
Host: Elytrosphaera xanthopyga (Coleoptera: Chrysomelidae)
Olafithiana multifida (Fain)
Grandiella multifida Fain, 1987: 101
Olafithiana multifida (Fain): Haitlinger, 1993e: 276
Type locality: Brazil
Distribution: Argentina, Bolivia, Brazil, Ecuador
Hosts: Stolas antiqua, S. aurosetosa, S. cruentata (Coleoptera: Chrysomelidae, Cassidinae)
Genus Olgattia Haitlinger
Olgattia Haitlinger, 1990b: 538
Type species: Olgattia useguaica Haitlinger, 1990
192
Acarorum Catalogus IX
Olgattia useguaica Haitlinger
Olgattia useguaica Haitlinger, 1990b: 538
Type locality: Usegua
Distribution: Tanzania
Host: Eumelasomus sansibaricus (Coleoptera: Passalidae)
Genus Ovarrothiana Haitlinger
Ovarrothiana Haitlinger, 1993d: 60
Type species: Ovarrothiana nikodemi Haitlinger, 1993
Ovarrothiana echinoderma (Summers et Schuster)
Grandiella echinoderma Summers & Schuster, 1979: 346; Fain, 1987: 94
Grandiellina echinoderma (Summers & Schuster): Fain, 1989: 67
Ovarrothiana echinoderma (Summers & Schuster): Haitlinger, 1993d: 65
Type locality: Cerro Compana, Panama
Distribution: Colombia, Panama
Host: Eugenysa colombiana (Coleoptera: Chrysomelidae, Cassidinae)
Ovarrothiana erazmi Haitlinger
Ovarrothiana erazmi Haitlinger, 1993d: 61
Type locality: Cayenne
Distribution: French Guyana
Host: Eugenysa venosa (Coleoptera: Chrysomelidae, Cassidinae)
Ovarrothiana leopoldi Haitlinger
Ovarrothiana leopoldi Haitlinger, 1993d: 65
Type locality: Yungaz de La Paz
Distribution: Bolivia
Host: Miocalaspis gentilis (Coleoptera: Chrysomelidae, Cassidinae)
Ovarrothiana ludomiri Haitlinger
Ovarrothiana ludomiri Haitlinger, 1993d: 64
Type locality: Weise
Distribution: Bolivia
Host: Cyclosoma tristis (Coleoptera: Chrysomelidae, Cassidinae)
Ovarrothiana nikodemi Haitlinger
Ovarrothiana nikodemi Haitlinger, 1993d: 60
Type locality: Peru
Distribution: Brazil, Peru
Host: Eugenysa divalis (Coleoptera: Chrysomelidae, Cassidinae)
Acarorum Catalogus IX 193
Ovarrothiana ozannae Haitlinger
Ovarrothiana ozannae Haitlinger, 1993d: 63
Type locality: Peru, Chanchamayo
Distribution: Brazil, Ecuador, Peru
Hosts: Agenyssa caedemadens, A. crassicornis, Eugenysa andicola (Coleoptera: Chrysomelidae, Cassidinae)
Genus Paracanestrinia Haitlinger
Paracanestrinia Haitlinger, 2017: 658
Type species: Paracanestrinia denmarkica Haitlinger, 2017
Paracanestrinia denmarkica Haitlinger
Paracanestrinia denmarkica Haitlinger, 2017: 659
Type locality: 11 km north of Hvidbjerg
Distribution: Denmark
Host: Carabus sp. (Coleoptera: Carabidae)
Genus Paramansia Cooreman
Paramansia Cooreman, 1950: 13
Type species: Paramansia menthastri Cooreman, 1950
Paramansia bicornis Khaustov et Eidelberg
Paramansia bicornis Khaustov & Eidelberg, 2001: 43; Trach & Khaustov, 2007: 268
Type locality: Crimea, Yalta
Distribution: Crimea, Ukraine
Hosts: Chrysolina menthastri, Ch. polita, Ch. fastuosa (Coleoptera: Chrysomelidae)
Paramansia giardi (Trouessart)
Canestrinia giardi Trouessart, in Berlese, 1887:
Paramansia giardi (Trouessart): Cooreman, 1950: 18
Type locality: ? Brazil
Distribution: ? Brazil
Host: ? Megistomela reticulata (Coleoptera: Chrysomelidae)
Paramansia menthastri Cooreman
Paramansia menthastri Cooreman, 1950: 14
Type locality: Bray-sous-Fay nr. Richelieu, Indre et Loire, France
Distribution: France
Host: Chrysolina menthastri (Coleoptera: Chrysomelidae)
Genus Paramelisia Summers et Schuster
Paramelisia Summers & Schuster, 1982a: 29
194
Acarorum Catalogus IX
Type species: Paramelisia lacuna Summers & Schuster, 1982
Paramelisia lacuna Summers et Schuster
Paramelisia lacuna Summers & Schuster, 1982a: 29
Type locality: Anomal Hills, Cinchona, S. India
Distribution: India
Host: Pleurarius brachyphyllus (Coleoptera: Passalidae)
Genus Paraphagella Cooreman
Paraphagella Cooreman, 1955: 28
Type species: Paraphagella princeps Cooreman, 1955
Paraphagella eudicellae Cooreman
Paraphagella eudicellae Cooreman, 1955: 33
Type locality: Kibga, Sud Visoke, 2400 m; Rutshuru
Distribution: D.R. Congo
Hosts: Eudicella smithi var. bertherandi, E. gralli var. meckowi (Coleoptera: Chrysomelidae)
Paraphagella indirae Haitlinger
Paraphagella indirae Haitlinger, 1991d: 291
Type locality: Nyangassy
Distribution: Cameroon
Host: Mecynorrhina [Chelarrhina – recte Chelorrhina] savagei (Coleoptera)
Paraphagella ingridae Haitlinger
Paraphagella ingridae Haitlinger, 1991d: 290
Type locality: Liberia
Distribution: Liberia
Host: Tmesorrhina iris (Coleoptera: Scarabaeidae, Cetoniinae)
Paraphagella minor Cooreman
Paraphagella minor Cooreman, 1955: 36
Type locality: Kibga, Sud Visoke, 2400 m
Distribution: D.R. Congo
Host: Eudicella smithi var. bertherandi (Coleoptera: Chrysomelidae)
Paraphagella odae Haitlinger
Paraphagella odae Haitlinger, 1991d: 292
Type locality: Distr. Kassai
Distribution: Congo (Brazzaville)
Host: Stephanorrhina guttata (Coleoptera: Scarabaeidae, Cetoniinae)
Acarorum Catalogus IX 195
Paraphagella princeps Cooreman
Paraphagella princeps Cooreman, 1955: 29
Type locality: May-ya-Moto, riv. Rutshuru
Distribution: D.R. Congo
Hosts: Dicronorhina micans, Smaragdesthes africana mutica (Coleoptera: Chrysomelidae)
Genus Paraphotia Khaustov et Eidelberg
Paraphotia Khaustov & Eidelberg, 2001: 37
Type species: Paraphotia calosomae Khaustov et Eidelberg, 2001
Paraphotia calosomae Khaustov et Eidelberg
Paraphotia calosomae Khaustov & Eidelberg, 2001: 39; Trach, 2006: 175
Type locality: Crimea, Opuk Mount.
Distribution: Crimea
Host: Calosoma auropunctatum (Coleoptera: Carabidae)
Genus Passalophagus Nesbitt
Passalophagus Nesbitt, 1976: 230
Type species: Passalophagus georgei Nesbitt, 1976
Passalophagus georgei Nesbitt
Passalophagus georgei Nesbitt, 1976: 231
Type locality: Baiyer River Bird of Paradise Sanctuary, Mt. Hagen Area
Distribution: Papua New Guinea
Host: Labienus sp. (Coleoptera: Passalidae)
Genus Percanestrinia Berlese
Canestrinia (Percanestrinia) Berlese, 1911: 184
Percanestrinia Berlese: Samšinák, 1965 (syn. with Canestrinia); Haitlinger, 1992a: 535
(restored as genus)
Type species: Alloptes blaptis Canestrini et Berlese, 1880
Percanestrinia blaptis (Canestrini et Berlese)
Alloptes blaptis Canestrini & Berlese, 1880: 148
Percanestrinia blaptis: Berlese, 1911: 184; Turk, 1953: 82; Rack, 1968: 265; Haitlinger,
1992a: 541; 2012: 187; Trach, 2006: 176
Canestrinia blaptis: Berlese, 1884: A.M.S., fasc. XXIX, No1; Haitlinger, 1992a: 541
Coleopterophagus blaptis (Canestrini et Berlese): Oudemans, 1923: 187
Percanestrinia (Percanestrinia) blaptis: Cooreman, 1954: 85
Canestrinia blaptis rackae Samšinák, 1970: 72 (syn. by Haitlinger, 1992a)
Type locality: ? „Italia septentrionalis“
196
Acarorum Catalogus IX
Distribution: Bulgaria, Georgia, Germany, Great Britain, Hungary, Italy (incl. Sardinia),
Poland, Ukraine
Hosts: Blaps gibbosa, B. lethifera, B. occulta, B. punctata (Coleoptera: Tenebrionidae),
Sphodrus leucophthalmus (Coleoptera: Carabidae)
Percanestrinia egypti El Bishlawi et Allam
Percanestrinia egypti El Bishlawi & Allam, 2003: 267
Type locality: Egypt
Distribution: Egypt
Percanestrinia izabelae Haitlinger
Percanestrinia izabelae Haitlinger, 1992a: 536
Type locality: Cyprus
Distribution: Cyprus
Host: Blaps gibbosa (Coleoptera: Tenebrionidae)
Percanestrinia maroccana Cooreman
Percanestrinia maroccana Cooreman, 1953: 6; Haitlinger, 1992a: 540
Type locality: Morocco
Distribution: Corsica (France), Italy, Morocco
Hosts: Blaps mucronata, B. pinguis, B. producta (Coleoptera: Tenebrionidae)
Percanestrinia norodomi Haitlinger
Percanestrinia norodomi Haitlinger, 1992a: 538
Type locality: Tashkent
Distribution: Uzbekistan
Host: Blaps hejconota cortosa (Coleoptera: Tenebrionidae)
Percanestrinia sellnicki Samšinák
Percanestrinia sellnicki Samšinák, 1965: 145; Beron, 1971: 205; Haitlinger, 1988d, 2012:
188; Canestrinia sellnicki: Samsinak, 1971: 155; Haitlinger, 1989; Trach, 2006: 176
Type locality: “Hungaria (Com. Bihar)” (now in Romania)
Distribution: Czech Rep., Hungary, former Jugoslavia, Moldova, Poland, Romania,
Slovakia, Ukraine
Hosts: Carabus (Procrustes) coriaceus rugifer, C. c. coriaceus, C. coriaceus pseudorugifer,
C. coriaceus banaticus, C. coriaceus subrugosus, C. coriaceus gressorius, C. coriaceus
imminutus, C. violaceus (Coleoptera: Carabidae)
Percanestrinia viviannae Haitlinger
Percanestrinia viviannae Haitlinger, 1992a: 540
Type locality: China
Acarorum Catalogus IX 197
Distribution: China
Host: Tenebrionidae (Coleoptera)
Genus Percanestriniella Khaustov et Eidelberg
Percanestriniella Khaustov & Eidelberg, 2001: 34
Type species: Percanestriniella caucasica Khaustov et Eidelberg, 2001
Percanestriniella caucasica Khaustov et Eidelberg
Percanestriniella caucasica Khaustov & Eidelberg, 2001: 34
Type locality: Caucasus, Mzymsha river bank near Kardyvach
Distribution: Russia (Caucasus)
Host: Carabus sp. (Coleoptera: Carabidae)
Percanestriniella levis (Samšinák)
Canestrinia levis Samšinák, 1971: 166
Percanestriniella levis: Khaustov & Eidelberg, 2001: 36
Type locality: ‘Turkestan’(Sussamyr – GB, Ketmen-Tjube), Buchara
Distribution: Kirghizstan, Uzbekistan
Hosts: Carabus (Pantophyrtus) turcomannorum, C.(P.) tanypedilus, C. (Cratocephalus)
balassogloi, C. (C.) cicatricosus, C. (Ulocarabus) sturowskyi, C. (Cratocechenus) akinini
(Coleoptera: Carabidae)
Percanestriniella vlachiana (Samšinák)
Canestrinia vlachiana Samšinák, 1971: 169
Percanestriniella vlachiana Khaustov & Eidelberg, 2001: 34
Type locality: Jalloori, Kulu (Kashmir)
Distribution: India (Kashmir)
Hosts: Carabus (Imaibius) boysi, C. (I.) stoliczkanus (Coleoptera: Carabidae)
Genus Peruthiana Haitlinger
Peruthiana Haitlinger, 2008: 69
Type species: Peruthiana briannae Haitlinger, 2008
Peruthiana briannae Haitlinger
Peruthiana briannae Haitlinger, 2008: 69
Type locality: Peru
Distribution: Peru
Host: undetermined Hispinae (Coleoptera: Chrysomelidae)
Genus Phaleratus Nesbitt
Phaleratus Nesbitt, 1976: 233
Type species: Phaleratus fentoni Nesbitt, 1976
198
Acarorum Catalogus IX
Phaleratus alatus Summers et Schuster
Phaleratus alatus Summers & Schuster, 1982a: 29
Type locality: Anomali Hills, Madras
Distribution: India
Host: Pleurarius brachyphyllus (Coleoptera: Passalidae)
Phaleratus fentoni Nesbitt
Phaleratus fentoni Nesbitt, 1976: 233
Type locality: Baiyer River Bird of Paradise Sanctuary, Mt. Hagen Area
Distribution: Papua New Guinea
Host: Coleoptera
Genus Phelliculophela Haitlinger
Phelliculophela Haitlinger, 1993a: 9
Type species: Phelliculophela roaldi Haitlinger, 1993
Phelliculophela roaldi Haitlinger
Phelliculophela roaldi Haitlinger, 1993a: 9
Type locality: Cameroon
Distribution: Cameroon
Host: Goliathus goliathus (Coleoptera : Scarabaeidae, Cetoniinae)
Genus Photia Oudemans
Photia Oudemans, 1904: 192
Type species: Canestrinia procustidis Berlese, 1881
Eucanestrinia Berlese, 1911: 184
Type species: “Canestrinia procrusti”
Photia adolfinae Haitlinger
Photia adolfinae Haitlinger, 1988 (sub “polymorpha”; 1994a: 202)
Photia adolphinae Haitlinger: Khaustov & Edelberg, 2001: 30; Trach, 2006: 175
Type locality: Narewka, voi. Bialystok, and many other localities in Poland
Distribution: Byelorussia, Poland, Crimea
Hosts: Carabus (C.) glabratus, C. gyllenhali (Coleoptera: Carabidae)
Photia bardoica Haitlinger
Photia bardoica Haitlinger, 1988: 321; 1995b: 261; 2012: 189
Type locality: Poland
Distribution: Austria, Poland
Host: Carabus nemoralis (Coleoptera: Carabidae)
Acarorum Catalogus IX 199
Photia bilkorum Samšinák
Photia bilkorum Samšinák, 1971: 199; Haitlinger, 2012: 190
Type locality: Abhasia
Distribution: Abhasia, Russia
Hosts: Carabus (Tribax) protensis, C. (T.) komarovi, C. (T.) shamyli, C. (T.) plasoni
(Coleoptera: Carabidae)
Photia chrysocarabi Cooreman
Photia chrysocarabi Cooreman, 1950: 34; Samšinák, 1971: 193; Haitlinger, 1988d; 1995b:
262; 2012: 188; Trach, 2006: 176
Type locality: Arlon (Belgium)
Distribution: Austria, Belgium, Czech Rep., England, France, Germany, Poland, Slovakia,
Ukraine
Hosts: Carabus auratus, C. irregularis, C. nemoralis, C. scheidleri, Chrysocarabus auronitens
(Coleoptera: Carabidae)
Photia graeca Cooreman
Photia graeca Cooreman, 1958: 1; Samšinák, 1965: 143; 1971: 189
Type locality: Philothée nr. Athens
Distribution: Belgium, England, France, Greece (incl. Rhodos), Holland, Turkey
Hosts: Carabus c. coriaceus, C. coriaceus foudrasi, C. c. kindermanni, C. c. impudicus, C.
violaceus purpurascens (Coleoptera: Carabidae)
Photia hejniana Samšinák
Photia hejniana Samšinák, 1971: 193; Haitlinger, 1988d; 1995b: 261; 2012: 189; Trach, 2006: 176
Type locality: Sobotka
Distribution: Austria, Croatia, Czech Rep., Moldova, Poland, Slovakia, Slovenia, Ukraine
Hosts: Carabus alpestris hoppei, C. coriaceus, C. croaticus, C. violaceus, C. v. exasperates,
C. v. fioris, C. v. obliquus, C. v. savinicus, C. v. styriensis (Coleoptera: Carabidae)
Photia hermengildae Haitlinger
Photia hermengildae Haitlinger, 1988: 319; 2012: 189
Type locality: Huta Szklana, Gory Swietokrzyskie, Voivodeship Kielce
Distribution: Poland, Ukraine
Host: Carabus linnaei (Coleoptera: Carabidae)
Photia howdeni Nesbitt
Photia howdeni Nesbitt, 1976: 223
Type locality: Baiyer River Bird of Paradise Sanctuary, Mt. Hagen Area
Distribution: Papua New Guinea
Host: Coleoptera
200
Acarorum Catalogus IX
Photia javensis (Oudemans)
Canestrinia javensis Oudemans, 1923b: 203
Photia javensis (Oudemans)
Type locality: Java, Soember Pakel, Probolingo
Distribution: Indonesia (Java)
Host: Coleoptera
Photia lopatini Khaustov et Eidelberg
Photia lopatini Khaustov & Eidelberg, 2001: 26
Type locality: Sakhalin Is., Krasnoselskaya river bank
Distribution: Russia (Sakhalin)
Host: Carabus (Acoptolabrus) lopatini (Coleoptera: Carabidae)
Photia lusitanica Samšinak
Photia lusitanica Samšinak, 1971: 189
Type locality: Sierra de Queija, Prov. Orense, Spain
Distribution: Portugal, Spain
Hosts: Carabus (Oreocarabus) amplipennis, Carabus (Mesocarabus) lusitanicus breuningi,
C. (M.) lusitanicus helluo, C.(M.) lusitanicus castilianus, C. (M.) lusitanicus logronicus,
C. (M.) lusitanicus schaumi (Coleoptera: Carabidae)
Photia melchiori Haitlinger
Photia melchiori Haitlinger, 1998: 2; 2012: 191
Type locality: China
Distribution: China
Host: Carabus sp. (Coleoptera: Carabidae)
Photia pacifica Khaustov et Eidelberg
Photia pacifica Khaustov & Eidelberg, 2001: 28
Type locality: Kuril Is. (Kunashir Is., Golovnin volcano)
Distribution: Russia (Kuril Is.)
Host: Carabus (Megodontus) kolbei (Coleoptera: Carabidae)
Photia polymorpha Samšinák
Photia polymorpha Samšinak, 1971: 184; Khaustov & Eidelberg, 2001: 31; Trach, 2006:
175; Haitlinger, 2012: 191
Type locality: Ulbinskoe, Siberia
Distribution: Japan, Korea, Russia (Siberia), Ukraine, Crimea
Host: holotype from Carabus (Megodontus) imperialis (Coleoptera: Carabidae). Reported
also from other species of the genus Carabus, but, according to Khaustov & Eidelberg
(2001), the mites collected from them may represent other species.
Acarorum Catalogus IX 201
Photia procera (Berlese)
Canestrinia (Eucanestrinia) procera Berlese, 1911: 184
Photia procera procera (Berlese): Samšinák, 1964: 35
Photia procera (Berlese): Cooreman, 1950: 26; Beron, 1971: 205; Haitlinger, 2012: 192
Type locality: “Carniolia” (Slovenia)
Distribution: Bulgaria, Slovenia (“Carniolia”), Italy
Hosts: Carabus (Procerus) caucasicus, C. (P.) gigas, C. (P.) scabrosus, Carabus (Procrustes)
coriaceus (Coleoptera: Carabidae)
Photia procera adanensis Samšinák
Photia procera adanensis Samšinák, 1964: 36
Type locality: Adana
Distribution: Turkey
Host: Carabus (Procerus) scabrosus transversalis (Coleoptera: Carabidae)
Photia procera marani Samšinák
Photia procera mařani Samšinák, 1964: 37
Type locality: Lebanon
Distribution: Lebanon
Host: Carabus (Procerus) syriacus (Coleoptera: Carabidae)
Photia procera turcica Samšinák
Photia procera turcica Samšinák, 1964: 35
Type locality: Turkey
Distribution: Turkey
Host: Carabus (Procerus) scabrosus caucasicus (Coleoptera: Carabidae)
Photia procustidis (Berlese)
Canestrinia procustidis Berlese, 1881: 749
Canestrinia procrusti Berlese, 1892; Canestrini, 1897
Photia procrusti (Berlese): Oudemans, 1904: 192
Photia procustidis (Berlese): Oudemans, 1904; 1923b: 203; Samšinák, 1965: 140; 1971;
Haitlinger, 2012: 190
Eucanestrinia procrusti Berlese, 1911: 184
Eucanestrinia procrustidis Vitzthum, 1925: 48
Type locality: Italy
Distribution: Albania, Cyprus, Greece (incl. Crete, Rhodes), Israel, Italy, Lebanon, Slovakia, Turkey, former Yugoslavia
Hosts: Carabus anatolicus, C. coriaceus, C. impressus (Coleoptera: Carabidae)
202
Acarorum Catalogus IX
Photia saetolata (Cooreman)
Percanestrinia saetolata Cooreman, 1950: 42; 1954: 85
Photia saetolata (Cooreman): Samšinák, 1965: 145; 1971: 197
Type locality: Belgium
Distribution: Belgium, Czech Rep., France, Germany, Great Britain, Holland
Hosts: Carabus (Procrustes) c. coriaceus, C. violaceus purpurascens, C.v. exasperaus, C.v.
bicinctus, C.v. fulgens (Coleoptera: Carabidae)
Photia sibirica Khaustov et Eidelberg
Photia sibirica Khaustov & Eidelberg, 2001: 23
Type locality: Novosibirskaya Obl., Togoochinskiy Reg., Kotorovo Village
Distribution: Russia
Host: Carabus (Megodontus) schoenherri (Coleoptera: Carabidae)
Genus Phriknodora Summers et Schuster
Phriknodora Summers & Schuster, 1982b: 43
Type species: Phriknodora pustulosa Summers & Schuster, 1982
Phriknodora avae Haitlinger
Phriknodora avae Haitlinger, 1994b: 81
Type locality: Costa Rica
Distribution: Costa Rica
Host: Stolas lebesi (Coleoptera: Chrysomelidae)
Phriknodora pustulosa Summers et Schuster
Phriknodora pustulosa Summers & Schuster, 1982b: 43
Type locality: Oaxaca
Distribution: Mexico, Panama
Hosts: Brazilianus mexicanus, B. batus (Coleopera: Cerambycidae)
Phriknodora scabra Summers et Schuster
Phriknodora scabra Summers & Schuster, 1982b: 46
Type locality: South Africa
Distribution: South Africa
Host: Trachyderes succinctus (Coleoptera: Cerambycidae)
Genus Procericola Cooreman
Procericola Cooreman, 1950: 44
Type species: Procericola ichthyoides Cooreman, 1950
Procericola bourgognei (Oudemans)
Photia bourgognei Oudemans, 1923b: 203
Acarorum Catalogus IX 203
Caraboecius coriacei Cooreman, 1950: 19
Caraboecius bourgognei Cooreman, 1954: 87
Procericola bourgognei (Oudemans): Samšinák, 1965: 138; 1971; Beron, 1971: 205; Lundquist, 1985: 381; Khaustov & Eidelberg, 2001: 39; Trach, 2006: 175
Type locality: France (Bourgogne)
Distribution: Albania, Belgium, Bulgaria, Caucasus, former Jugoslavia, France, Greece,
Hungary, Italy, Moldova, Romania, Russia, Slovakia, Sweden, Turkey, Ukraine
Hosts: Carabus coriaceus, Carabus (Procrustes) ch. chevrolati, C. (P.) clypeatus clypeatus,
C. (Megodontus) exaratus (Coleoptera: Carabidae)
Procericola ichthyoides Cooreman
Procericola ichthyoides Cooreman, 1950: 44
Procericola ichthyoides ichthyoides Cooreman: Samšinák, 1964: 38
Type locality: “Carniolia, Yougoslavie et Tolmezzo, Torrente Vinadia, Carnia”
Distribution: Italy, Slovenia
Host: Carabus (Procerus) gigas (Coleoptera: Carabidae)
Procericola ichthyoides scabricola Samšinák
Procericola ichthyoides scabricola Samšinák, 1964: 38
Type locality: Caucasus, Mekish Mt.
Distribution: Bulgaria, Russia, Turkey
Hosts: Carabus (Procerus) scabrosus transversalis (Coleoptera: Carabidae)
Procericola ichthyoides transversicola Samšinák
Procericola ichthyoides transversicola Samšinák, 1964: 40
Type locality: “Cilicien”
Distribution: Turkey
Hosts: Carabus (Procerus) scabrosus transversalis (Coleoptera: Carabidae)
Genus Pseudamansia Cooreman
Pseudamansia Cooreman, 1950: 5
Type species: Dermaleichus chrysomelinus Koch, 1841
Pseudamansia chrysomelinus (C.L. Koch)
Dermaleichus chrysomelinus C.L. Koch, 1841: 4
Amansia chrysomelinus: Oudemans, 1923:
Pseudamansia chrysomelinus (C.L. Koch): Cooreman, 1950: 5; Jolivet, 1952, 1954; Theodorides, 1955; Santiago-Blay & Fain, 1994: 412
Pseudamansia chrysomelina (C.L. Koch): Bernini,
Type locality: various places in France (Neotype)
204
Acarorum Catalogus IX
Distribution: Belgium, France, Algeria, Morocco, Mayorca Is., Italy
Hosts: Timarcha atlantica, T. balearica, T. goettingensis, T. interstitialis, T. nicaeensis, T.
pauperata, T. punctella, T. rugosa, T. tenebricosa (Coleoptera: Chrysomelidae)
Pseudamansia decorata Trach et Khaustov
Pseudamansia decorata Trach & Khaustov, 2007: 269
Type locality: Odessa and Vinnica regions
Distribution: Ukraine
Host: Timarcha goettingensis (Coleoptera: Chrysomelidae)
Genus Pseudocanestrinia Khaustov et Eidelberg
Pseudocanestrinia Khaustov & Eidelberg, 2001: 31
Type species: Pseudocanestrinia carabi Khaustov et Eidelberg, 2001
Pseudocanestrinia berndi (Haitlinger)
Canestrinia berndi Haitlinger, 1995: 260
Pseudocanestrinia berndi (Haitlinger): Khaustov & Eidelberg, 2001: 34
Type locality: Jablanica (Herzegovina)
Distribution: Bosna and Herzegovina, Crimea
Hosts: Carabus latenulatus plassensis, C. gyllenhali (Coleoptera: Carabidae)
Pseudocanestrinia carabi Khaustov et Eidelberg
Pseudocanestrinia carabi Khaustov & Eidelberg, 2001: 33; Trach, 2006: 175
Type locality: Crimean mountains ‘Ay-Petri’
Distribution: Crimea
Host: Carabus (C.) bessarabicus (Coleoptera: Carabidae)
Pseudocanestrinia mahunkai (Samšinák)
Canestrinia mahunkai Samšinák, 1971: 160; Haitlinger, 1988
Pseudocanestrinia mahunkai (Samšinák): Khaustov & Eidelberg, 2001: 32; Haitlinger,
2012: 186
Type locality: Balatonaracs (Hungary)
Distribution: Hungary, Poland
Host: Carabus coriaceus (Coleoptera: Carabidae)
Pseudocanestrinia occidentalis (Samšinák)
Canestrinia occidentalis Samšinák, 1971: 155
Pseudocanestrinia occidentalis (Samšinák): Khaustov & Eidelberg, 2001: 32
Type locality: La Lourde
Distribution: France
Host: Carabus (Chrysocarabus) auronitens (Coleoptera: Carabidae)
Acarorum Catalogus IX 205
Pseudocanestrinia orientalis (Samšinák)
Canestrinia orientalis Samšinák, 1971: 157
Pseudocanestrinia orientalis (Samšinák): Khaustov & Eidelberg, 2001: 32
Type locality: “Carniolia”
Distribution: Slovenia
Hosts: Carabus (Megodontus) caelatus, C. (M.) croaticus (Coleoptera: Carabidae)
Genus Pseudoparaphagella Haitlinger
Pseudoparaphagella Haitlinger, 2010: 58
Type species: Pseudoparaphagella gerardi Haitlinger, 2010
Pseudoparaphagella gerardi Haitlinger
Pseudoparaphagella gerardi Haitlinger, 2010: 58
Type locality: Guinea
Distribution: Guinea
Host: Caelorrhina thoreyi (Coleoptera: Scarabaeidae, Cetoniinae)
Genus Rugoniphela Haitlinger
Rugoniphela Haitlinger, 1992b: 533
Type species: Rugoniphela marloni Haitlinger, 1992
Rugoniphela marloni Haitlinger
Rugoniphela marloni Haitlinger, 1992b: 534
Type locality: holotype from Borneo, paratypes from Java and Sumatra
Distribution: Indonesia (Borneo, Java, Sumatra)
Host: Eurytrachelus bucephalus (Coleoptera: Lucanidae)
Genus Sabothiana Haitlinger
Sabothiana Haitlinger, 1993e: 269
Type species: Sabothiana riccardoi Haitlinger, 1993
Sabothiana boi Haitlinger
Sabothiana boi Haitlinger, 1993e: 272
Type locality: Bahia
Distribution: Brazil
Host: Charidotis sp.(Coleoptera: Chrysomelidae, Cassidinae)
Sabothiana reidari Haitlinger
Sabothiana reidari Haitlinger, 1993e: 273
Type locality: Bocaina
Distribution: Brazil
Host: Charidotella kesseli (Coleoptera: Chrysomelidae, Cassidinae)
206
Acarorum Catalogus IX
Sabothiana riccardoi Haitlinger
Sabothiana riccardoi Haitlinger, 1993e: 271
Type locality: Brazil
Distribution: Brazil
Host: Charidotis furunculus (Coleoptera: Chrysomelidae, Cassidinae)
Genus Sajanophela Haitlinger
Sajanophela Haitlinger, 1990b: 545
Type species: Sajanophela alfredae Haitlinger, 1990
Sajanophela alfredae Haitlinger
Sajanophela alfredae Haitlinger, 1990b: 545
Type locality: Papua New Guinea
Distribution: Papua New Guinea
Host: Protomocelus australis (Coleoptera: Passalidae)
Genus Sandrophela Haitlinger
Sandrophela Haitlinger, 1990: 315
Type species: Sandrophela kokodaica Haitlinger, 1990
Sandrophela amarae Haitlinger
Sandrophela amarae Haitlinger, 1994a: 197
Type locality: East Java, Sumatra
Distribution: Indonesia (Java, Sumatra)
Host: Eurytrachelus eurycephalus, E. gypeatus (Coleoptera: Lucanidae)
Sandrophela dittae Haitlinger
Sandrophela dittae Haitlinger, 1992: 282
Type locality: Borneo
Distribution: Indonesia
Host: Colydium sp. (Coleoptera: Colydiidae)
Sandrophela hugoni Haitlinger
Sandrophela hugoni Haitlinger, 1994a: 194
Type locality: Sumatra
Distribution: Indonesia (Sumatra)
Host: Odontolabis wollastoni (Coleoptera: Lucanidae)
Sandrophela kokodaica Haitlinger
Sandrophela kokodaica Haitlinger, 1990e: 316
Type locality: Kokoda
Acarorum Catalogus IX 207
Distribution: Papua New Guinea
Host: Passandra trigemina (Coleoptera: Passandridae)
Sandrophela rodioni Haitlinger
Sandrophela rodioni Haitlinger, 1993b: 342
Type locality: North Vietnam, Monte Mouson
Distribution: Vietnam
Host: Eurytrachelus platymelus (Coleoptera: Lucanidae)
Sandrophela sophiae Haitlinger
Sandrophela sophiae Haitlinger, 1994a: 196
Type locality: India
Distribution: India
Host: Hemisodorcus chevrilati (Coleoptera: Lucanidae)
Sandrophela spectanda (Berlese)
Canestrinia spectanda Berlese, 1910: 269
Sandrophela spectanda (Berlese): Haitlinger, 1994a: 199
Type locality: Java
Distribution: Indonesia (Java, Sumatra)
Host: Dorcus bucephalus (Coleoptera: Lucanidae)
Genus Saniothiana Haitlinger
Saniothiana Haitlinger, 1990e: 317
Type species: Saniothiana pycnosa Haitlinger, 1990
Saniothiana barumbaica Haitlinger
Saniothiana barumbaica Haitlinger, 1990e: 320
Type locality: Barumba
Distribution: Cameroon
Host: Odontopezus cupreus lucens (Coleoptera: Tenebrionidae)
Saniothiana pycnosa Haitlinger
Saniothiana pycnosa Haitlinger, 1990e: 318
Type locality: Africa
Distribution: Africa
Host: Pycnocerus sp. (Coleoptera: Tenebrionidae)
Genus Scarabiphotia Haitlinger
Scarabiphotia Haitlinger, 1994a: 205
Type species: Scarabiphotia vignoni Haitlinger, 1994
208
Acarorum Catalogus IX
Scarabiphotia vignoni Haitlinger
Scarabiphotia vignoni Haitlinger, 1994a: 205
Type locality: unknown
Distribution: unknown
Host: Scarabaeus variolosus (Coleoptera: Scarabaeidae)
Genus Shillongia Haitlinger
Shillongia Haitlinger, 1990b: 546
Type species: Shillongia marzenae Haitlinger, 1990
Shillongia marzenae Haitlinger
Shillongia marzenae Haitlinger, 1990b: 546
Type locality: Assam, Shillong
Distribution: India (Assam)
Host: Leptaulus indicus (Coleoptera: Passalidae)
Genus Sorbinophela Haitlinger
Sorbinophela Haitlinger, 1993a: 7
Type species: Sorbinophela sandyi Haitlinger, 1993
Sorbinophela sandyi Haitlinger
Sorbinophela sandyi Haitlinger, 1993a: 7
Type locality: nr. Lake Ngami
Distribution: Botswana
Host: Poecilophila maculatissima (Coleoptera : Scarabaeidae, Cetoniinae)
Genus Tamarangia Haitlinger
Tamarangia Haitlinger, 1991d: 284
Type species: Tamarangia nimfae Haitlinger, 1992
Tamarangia ethiopica Khaustov
Tamarangia ethiopica Khaustov, 2006: 79
Type locality: Ethiopia
Distribution: Ethiopia
Host: Pachnoda marginata (Coleoptera: Scarabaeidae, Cetoniinae)
Tamarangia fabiolae Haitlinger
Tamarangia fabiolae Haitlinger, 1991d: 286
Type locality: Freetown
Distribution: Sierra Leone
Host: Pachnoda marginata (Coleoptera: Scarabaeidae, Cetoniinae)
Acarorum Catalogus IX 209
Tamarangia flawiani Haitlinger
Tamarangia flawiani Haitlinger, 1991d: 287
Type locality: Freetown
Distribution: Sierra Leone
Host: Pachnoda marginata (Coleoptera: Scarabaeidae, Cetoniinae)
Tamarangia nimfae Haitlinger
Tamarangia nimfae Haitlinger, 1991d: 284; Haitlinger & Chmielewski, 2004: 199
Type locality: Ethiopia
Distribution: Ethiopia, South Africa
Host: Pachnoda thoracica (Coleoptera: Scarabaeidae, Cetoniinae)
Genus Tamarophela Haitlinger
Tamarophela Haitlinger, 1990b: 537
Type species: Tamarophela batjanica Haitlinger, 1990
Tamarophela batjanica Haitlinger
Tamarophela batjanica Haitlinger, 1990b: 537
Type locality: Moluccas, Batjan Is.
Distribution: Indonesia (Moluccas, Batjan Is.)
Host: undetermined Passalidae (Coleoptera)
Genus Taseopus Summers et Schuster
Taseopus Summers & Schuster, 1982a: 27
Type species: Taseopus platythrix Summers et Schuster, 1982
Taseopus platythrix Summers et Schuster
Taseopus platythrix Summers & Schuster, 1982a: 27
Type locality: Queensland
Distribution: Australia
Host: Passalidae gen. sp. (Coleoptera)
Genus Teklathiana Haitlinger
Teklathiana Haitlinger, 1999: 36
Type species: Teklathiana onnae Haitlinger, 1999
Teklathiana conula (Summers et Schuster)
Grandiella conula Summers & Schuster, 1979: 341
Grandiellina conula: Santiago-Blay & Fain, 1994:
210
Acarorum Catalogus IX
Ennodithiana conula (Summers & Schuster): Haitlinger 1993: 72
Teklathiana conula (Summers et Schuster): Haitlinger, 1999: 40
Type locality: Panama (El Cope)
Distribution: Panama
Host: Omaspides bistriata (Coleoptera: Chrysomelidae, Cassidinae)
Teklathiana onnae Haitlinger
Teklathiana onnae Haitlinger, 1999: 37; 2001: 103
Type locality: holotype from Colombia, paratype from Bolivia
Distribution: Bolivia, Colombia
Host: Acromis sparsa (Coleoptera: Chrysomelidae, Cassidinae)
Teklathiana rugosita (Summers et Schuster)
Grandiella rugosita Summers & Schuster, 1979: 343
Teklathiana rugosita (Summers et Schuster): Haitlinger, 1999: 40
Type locality: Panama (Canal Zone and Panama City)
Distribution: Panama
Host: Acromis sparsa (Coleoptera: Chrysomelidae, Cassidinae)
Genus Teophyssa Haitlinger
Teophyssa Haitlinger, 1990b: 544
Type species: Teophyssa honoratae Haitlinger, 1990
Teophyssa beatae Haitlinger
Teophyssa beatae Haitlinger, 1990b: 544
Type locality: Sumatra, Bekantiang
Distribution: Indonesia (Sumatra)
Host: undetermined Passalidae (Coleoptera)
Teophyssa honoratae Haitlinger
Teophyssa honoratae Haitlinger, 1990b: 544
Type locality: North Sumatra
Distribution: Indonesia (Sumatra)
Host: Gonatus noviculator (Coleoptera: Passalidae)
Teophyssa teresae Haitlinger
Teophyssa teresae Haitlinger, 1990b: 545
Type locality: Sumatra, Soekaranda
Distribution: Indonesia (Sumatra)
Host: undetermined Passalidae (Coleoptera)
Acarorum Catalogus IX 211
Genus Thopia Summers et Schuster
Thopia Summers & Schuster, 1982b: 35
Type species: Thopia tenax Summers et Schuster, 1982
Thopia amedae Haitlinger
Thopia amedae Haitlinger, 1993e: 268
Type locality: Cayenne
Distribution: French Guiana
Host: Eugenysa venosa (Coleoptera: Chrysomelidae, Cassidinae)
Thopia boguwolae Haitlinger
Thopia boguwolae Haitlinger, 1993e: 266
Type locality: Brazil
Distribution: Brazil
Host: Polychalca punctatissima (Coleoptera: Chrysomelidae, Cassidinae)
Thopia bulbocaudata Summers et Schuster
Thopia bulbocaudata Summers & Schuster, 1982b: 37
Type locality: Panama
Distribution: Panama
Host: Neoptychodes trilineatus (Coleoptera: Cerambycidae, Lamiinae)
Thopia corinnae Haitlinger
Thopia corinnae Haitlinger, 1993e: 268
Type locality: Bahia
Distribution: Brazil
Host: Polychalca dentipennis (Coleoptera: Chrysomelidae, Cassidinae)
Thopia pannucea Summers et Schuster
Thopia pannucea Summers & Schuster, 1982b: 41
Type locality: Panama
Distribution: Panama
Host: Lagocheirus foveolatus (Coleoptera: Cerambycidae, Lamiinae)
Thopia tenax Summers et Schuster
Thopia tenax Summers & Schuster, 1982b: 37
Type locality: Canal Zone
Distribution: Panama
Host: Taeniodes scalaris (Coleoptera: Cerambycidae)
Genus Uriophela Haitlinger
Uriophela Haitlinger, 1992b: 539
212
Acarorum Catalogus IX
Type species: Uriophela arieli Haitlinger, 1992
Uriophela arieli Haitlinger
Uriophela arieli Haitlinger, 1992b: 540
Type locality: Amur District (Russia)
Distribution: Russia (Far East)
Host: Hemisodorcus [Macrodora] rubrofemoratus (Coleoptera: Lucanidae)
Genus Vandeinella Haitlinger
Vandeinella Haitlinger, 1989a: 54
Type species: Vandeinella adriasae Haitlinger, 1989
Vandeinella adriasae Haitlinger
Vandeinella adriasae Haitlinger, 1989a: 54
Type locality: Sumatra
Distribution: Indonesia (Sumatra)
Host: Trichogomphus milon (Coleoptera: Scarabaeidae, Dynastinae)
Vandeinella opressa Haitlinger
Vandeinella opressa Haitlinger, 1989a: 55
Type locality: Borneo
Distribution: ? Indonesia (Borneo)
Host: Trichogomphus lunicollis (Coleoptera: Scarabaeidae, Dynastinae)
Subfam. Vereoxiinae Haitlinger
Vereoxiinae Haitlinger, 1995a: 80
Type genus: Vereoxia Haitlinger, 1995
Genus Vereoxia Haitlinger
Vereoxia Haitlinger, 1995a: 80
Type species: Vereoxia bogeri Haitlinger, 1995
Vereoxia bogeri Haitlinger
Vereoxia bogeri Haitlinger, 1995a: 80
Type locality: unknown
Distribution: unknown
Host: Lucanidae gen. sp. (Coleoptera)
Acarorum Catalogus IX 213
Index of Canestriniidae
(Names in bold are accepted as valid)
abapoica, Gasuthiana 174
abufarica, Gasuthiana 174
Acrotocarus 148
adanensis, Photia procera 201
adelinae, Olafithiana 191
adolfinae, Photia 198
adolphinae, Photia 198
adriasae, Vandeleinella 212
africanus, Diplognathophilus 171
Afrocanestrinia 148
aidae, Eunorithiana 174
alani, Grandiellopsis 179
alatus, Phaleratus 198
albertae, Bibulothiana 158
albini, Coleoglyphus 167
Albinorattia 149
tacea, Amansia
almae, Kahoorangia 183
alutacea, Amansia 148
alutacea, Grandiella 148
alutaceus, Acrotocarus 148
alvari, Barbiangia 157
Amansia
Amansiella 149
amarae, Sandrophela 206
Ambilohylla 149
Amboinophela 149
amedae, Thopia 211
amplexans, Canestriniella 164
Anapistes 150
anomala, Melisia 185
anoni, Averithiana 157
Apalotacarus 150
archidonaica, Danaithiana 170
arcuata, Arrunsithiana 153
arcuata, Grandiella 153
arcuata, Grandiellina 153
ariadnae, Melisia 185
ariani, Grandiellopsis 179
arieli, Uriophela 212
aristatus, Apalotacarus 150
Arraphosoma 152
Arrunsithiana 153
artemoni, Arrunsithiana 153
aruica, Lidiophela 183
arybbasi, Athogavia 155
asahinai, Ibizella
asilahica, Ibizella 182
assamica, Athogavia 155
Athogavia 155
Aurilossongia 156
Averithiana 157
avae, Phiknodorae 202
ayraica, Danaithiana 170
baali, Coleopterophagus 168
bahiana, Hatohylla 181
balearica, Ibizella 182
baloghi, Melisia 185
barbarae, Jovitella 183
barryi, Noemiphela 190
barumbaica, Saniothiana 207
batjanica, Tamarophela 209
batocerae, Canestriniella 164
batocerae, Grandiella 164
batocerae, Grandiellina 164
battosi, Arrunsithiana 153
beatae, Teophyssa 210
Beeriphotia 158
belerofoni, Hyllosihemia 181
beloniana, Megacanestrinia 184
belzebubi, Coleopterophagus 168
berlesei, Coleopterophagus
214
Acarorum Catalogus IX
bernae, Amboinophela 149
berndi, Canestrinia 204
berndi, Pseudocanestrinia 204
bertrami, Bircericola 159
bettinae, Globosophotia 175
bibiannae, Hatohylla 181
Bibulothiana 158
bicornis, Paramansia 193
bilkorum, Photia 199
Bircericola 159
bissaina, Mossongia 189
blaptis, Alloptes 195
blaptis, Canestrinia 195
blaptis, Coleopterophagus 195
blaptis, Percanestrinia 195
blattophaga, Canestrinia 168
blattophagus, Coleopterophagus 168
Boetophela 159
bogeri, Vereoxia 212
bognae, Lidiophela 183
boguwolae, Thopia 211
boi, Sabothiana 211
bolanica, Aurilossongia 156
bolebori, Metriothiana 189
Boleohylla 161
borneoica, Athogavia 155
bourgognei, Caraboecius 203
bourgognei, Photia 202
bourgognei, Procericola 202
brevispinata, Bibulothiana 158
brevispinata, Grandiella 158
briannae, Peruthiana 197
bulbocaudata, Thopia 211
calosomae, Paraphotia 195
calostegis, Donnelafontia 172
Camirohylla 161
Canestrinia 161
Canestrinia (Dicanestrinia) 171
Canestrinia (Percanestrinia) 161
Canestriniella 164
Canestriniidae 147
Canestriniinae 147
canistra, Grandiellopsis 179
carabi, Pseudocanestrinia 204
carabicola, Canestrinia 161
carabicola, Coleopterophagus 161
carabicola, Percanestrinia 161
Caraboecius 165
caroae, Gioharattia 175
cassandrae, Boetophela 159
caucasica, Percanestriniella 197
celinae, Lidiophela 184
cerambicis, Canestrinia (Dicanestrinia)
171
cerambycis, Dermaleichus 171
cerambycis, Dicanestrinia 171
cetoniae, Coleopterophagus 169
Cetonicola 165
chanchamayoica, Danaithiana 170
Chelynochroa 166
chrysocarabi, Photia 199
chrysomelinus, Amansia 203
chrysomelinus, Dermaleichus 203
chrysomelinus, Pseudomansia 203
cidaris, Apalotacarus 150
Cnecoderma 166
coarica, Gasuthiana 174
Coleoglyphus 167
Coleopterophagus 167
collaris, Hypopteryx 181
conula, Ennodithiana 210
conula, Grandiella 209
conula, Grandiellina 209
conula, Teklathiana 209
cooremani, Grandiella 175
cooremani, Grandiellina 175
Cooremania 169
coriacei, Caraboecius 165
corinnae, Thopia 211
crataepus, Acrotocarus 148
cyclosoma, Grandiellopsis 180
cyriaki, Bibulothiana 158
damiani, Canestrinia 162
Danaithiana 170
Acarorum Catalogus IX 215
decemcaudata, Grandiella 176
decemcaudata, Grandiellina 176
decorata, Pseudamansia 204
delmari, Dorynothiana 172
denmarkica, Paracanestrinia 193
Dicanestrinia 171
dolichopoda, Cnecoderma 167
dictyophora, Chelinochroa 166
dioni, Aurilossongia 156
dionizyi, Coleopterophagus 168
Diplognathophilus 171
Diplopodocoptes 171
dittae, Sandrophela 206
dolichopoda, Cnecoderma 167
donaldi, Coleopterophagus 168
Donnelafontia 172
dorcicola, Canestrinia 162
Dorynothiana 172
drvotocka, Neophotia 189
echinatus, Apalotacarus 150
echinoderma, Grandiella 192
echinoderma, Grandiellina 192
echinoderma, Ovarrothiana 192
ecuadorica, Danaithiana 170
ecuadorensis, Grandiella 176
ecuadorensis, Grandiellina 176
egypti, Percanestrinia 156
elae, Canestriniella 165
elongata, Barbiangia 157
Ennodithiana 172
Ennorithiana 173
ephraimi, Boetophela 159
erazmi, Ovarrothiana 192
erhardi, Boetophela 159
erinnae, Arrunsithiana 154
escaudata, Arrunsithiana 154
esacaudata, Grandiella 154
esacaudata, Grandiellina 154
ethiopica, Barbiangia 157
ethiopicus, Diplopodophilus 157
ethiopica, Tamarangia 208
Eucanestrinia 198
eudicellae, Paraphagella 194
Eunorithiana 173
everittae, Bibulothiana 158
fabiolae, Tamarangia 208
favosa, Ambilohylla 149
fentoni, Phaleratus 198
feziana, Camirohylla 161
fileasi, Aurilossongia 156
flawiani, Tamarangia 209
foliacea, Grandiella 191
foliacea, Grandiellina 191
foliacea, Olafithiana 191
fuscipes, Coleoglyphus 167
fulgens, Apalotacarus 150
fusticulus, Apalotacarus 151
fusulus, Apalotacarus 151
gabrielae, Melisia 185
gamana, Athogavia 155
gambosa, Grandiella 176
gambosa, Grandiellina 176
garrica, Boleohylla 161
Gasuthiana 174
guarana, Aurilossongia 156
gentilis, Percanestrinia (Lombardiniella)
georgei, Passalophagus 195
gerardi, Pseudoparaphagella 205
giardi, Canestrinia 193
giardi, Paramansia 193
ginae, Boetophela 160
Gioharattia 175
glaber, Apalotacarus 151
Globosophotia 175
gracilis, Apalotacarus 151
graeca, Photia 199
Grandiella 175
Grandiellina 175
Grandiellopsis 179
Haitlingeria 180
Hargeinella 180
Hatohylla 181
hejniana, Photia 199
helenae, Melisia 186
216
Acarorum Catalogus IX
helgae, Irmongia 182
heliodori, Eunorithiana 173
hermengildae, Photia 199
hieronimae, Melisia 186
hispanica, Canestrinia 162
hispanica, Melisia
hispidus, Cetonicola 165
hoaria, Aurilossongia 156
honoratae, Teophyssa 210
howdeni, Photia 199
huberti, Dicanestrinia 171
hugoni, Sandrophela 206
Hyllosihemia 181
Hypopteryx 181
Ibizella 182
ichtyoides, Procericola 203
igae, Lidiophela 184
igori, Albinorattia 149
illegalis, Canestrinia 162
ilosi, Arrunsithiana 154
indirae, Paraphagella 194
ingeborgae, Melisia 186
ingridae, Paraphagella 194
irenae, Melisia 186
Irmongia 182
isadorae, Cetonicola 166
iwettae, Olafithiana 191
izabelae, Noemiphela 190
izabelae, Percanestrinia 196
izae, Jullongia 183
jadwigae, Melisia 186
javensis, Canestrinia 200
javensis, Photia 200
joliveti, Grandiella 191
joliveti, Grandiellina 191
joliveti, Olafithiana 191
Jovitella 183
Jullongia 183
jurandi, Dorynothiana 172
jureceki, Neophotia 190
kacperi, Canestrinia 162
Kahoorangia 183
kamelskyi, Canestrinia 162
kietaensis, Cooremania 170
kimseyi, Grandiella 176
kimseyi, Grandiellina 176
kiogana, Boleohylla 161
klapaleki, Melisia 186
knobi, Dicanestrinia 171
kokodaica, Sandrophela 206
lacuna, Paramelisia 194
latipes, Arrunsithiana 154
latipes, Grandiellina 154
latissima, Neophotia 190
leopoldi, Ovarrothiana 192
leptinotarsa, Melisia 186
levis, Canestrinia 197
levis, Percanestriniella 197
Lidiophela 183
lombardinii, Melisia 187
longilobata, Haitlingeria 180
longini, Grandiella 177
longini, Grandiellina 177
lopatini, Photia 200
ludomiri, Ovarrothiana 192
luroris, Apalotacarus 151
lusitanica, Photia 200
macgillavryi, Canestrinia 163
maharae, Gioharattia 175
mahunkai, Canestrinia 204
mahunkai, Melisia 187
mahunkai, Pseudocanestrinia 204
manicata, Canestrinia 163
maniusi, Ennodithiana 173
marani, Photia procera 201
mariettae, Melisia 187
marinae, Grandiella 177
marinae, Grandiellina 177
maritae, Cetonicola 166
marloni, Rugoniphela 205
maroccana, Percanestrinia 196
maroni, Coleopterophagus 168
marzenae, Shillongia 208
medoni, Ennodithiana 173
Acarorum Catalogus IX 217
Megacanestrinia 184
megesi, Emmodithiana 173
megninii, Coleopterophagus 169
megnini, Coleopterophagus 169
megninii, Dermogliphus 168
megnini, Dermoglyphus 168
melaniae, Melisia 187
melchiori, Photia 200
melillae, Gasuthiana 174
Melisia 185
melisii, Melisia 187
Melisiinae 185
menthastri, Paramansia 193
Mesophotia 188
messalinae, Ennodithiana 173
Metriothiana 188
microdisca, Canestrinia 163
minax, Araphosoma 152
minor, Grandiella 177
minor, Grandiellina 177
minor, Paraphagella 194
mirabilis, Acrotocarus 148
morrata, Hatohylla 181
Mossongia 189
mucronata, Megacanestrinia 185
multifida, Grandiella 191
multifida, Olafithiana 191
neglectus, Coleopterophagus 169
Neophotia 189
nepalensis, Canestrinia 163
nikodemi, Ovarrothiana 192
nimfae, Tamarangia 209
Noemiphela 190
norae, Aurilossongia 157
norodomi, Percanestrinia 196
nosiana, Athogavia 155
occidentalis, Canestrinia 204
occidentalis, Pseudocanestrinia 204
occidii, Melisia 187
Oceogavia 190
octocaudata, Grandiella 177
octocaudata, Grandiellina 177
odae, Paraphagella 194
odiosi, Grandiellopsis 180
Olafithiana 190
olareica, Gasuthiana 174
Olgattia 191
omoplata, Grandiella tetracaudata 179
omoplata, Grandiellina tetracaudata 179
onnae, Teklathiana 210
opressa, Vandeleinella 212
orestesi, Cetonicola 166
orientalis, Canestrinia 205
orientalis, Pseudocanestrinia 205
orsenae, Grandiellopsis 180
oswaldi, Bubulothiana 159
ovalis, Cnecoderma 167
Ovarrothiana 192
ozannae, Ovarrothiana 193
paavoi, Canestrinia 163
pacifica, Photia 200
pannucea, Thopia 211
Paracanestrinia 193
Paramansia 193
Paramelisia 193
paraminax, Araphosoma 153
Paraphagella 194
Paraphotia 195
Passalophagus 195
pauliana, Lidiophela 184
paxillus, Apalotacarus 151
pecki, Coleopterophagus 184
pecki, Lidiophela 184
pekingensis, Canestrinia 163
pelagiae, Lidiophela 184
penelopae, Ennodithiana 173
penicilata p., Mesophotia 187
penicillata similis, Mesophotia 187
pentagona, Grandiella 177
pentagona, Grandiellina 177
pentodontis, Canestrinia 164
Percanestrinia 195
Percanestriniella 197
Peruthiana 197
218
Acarorum Catalogus IX
petilus, Apalotacarus 151
Phaleratus 197
Phelliculophela 198
phoebinae, Canestriniella 165
Photia 198
Phriknodora 202
physsana, Canestriniella 165
pictura, Canestrinia 164
platysma, Ennodithiana 173
platysma, Grandiella 173
platythrix, Taseopus 209
polymorpha, Photia 200
poppana, Hargeinella 181
princeps, Paraphagella 195
procera, Canestrinia (Eucanestrinia) 201
procera, Photia 201
Procericola 202
procrusti, Eucanestrinia 201
procrusti, Photia 201
procrustidis, Eucanestrinia 201
procustidis, Canestrinia 201
procustidis, Photia 201
protensus, Apalotacarus 152
Pseudamansia 203
Pseudocanestrinia 204
Pseudoparaphagella 205
pulcher, Coleopterophagus 169
pustulosa, Phriknodora 202
pycnosa, Saniothiana 207
quadrata, Arrunsithiana 154
quadrata, Grandiella 154
quadrata, Grandiellina 154
quadrisetosus, Coleopterophagus 169
rackae, Canestrinia blaptis 195
ramoni, Canestrinia 164
reidari, Sabothiana 205
remigans, Canestrinia 164
riccardoi, Sabothiana 206
rigescens, Apalotacarus 151
roaldi, Pheliculophela 198
robertsoni, Cetonicola 160
robertsoni, Boetophela 160
rodioni, Sandrophela 207
rogeri, Grandiella 177
rogeri, Grandiellina 177
rosascostai, Coleoglyphus 178
rosascostai, Grandiella 178
rosascostai, Hatohylla 178
rosascostai, Donnelafontia 178
rudolfi, Coleopterophagus 169
rufinae, Afrocanestrinia 148
Rugoniphela 305
rugosita, Grandiella 210
rugosita, Teklathiana 210
rutae, Melisia 187
Sabothiana 205
saetolata, Percanestrinia 202
saetolata, Photia 202
sajanae, Apalotacarus 152
Sajanophela 206
salomeae, Melisia 188
samsinaki, Canestrinia 182
samsinaki, Ibizella 182
samsinaki, Melisia 188
samsoni, Afrocanestrinia 148
Sandrophela 206
sandyi, Sorbinophela 208
Saniothiana 206
sardica, Canestrinia (Percanestrinia) 161
scabra, Phriknodora 202
scabricola, Procericola ichthioides 203
scagana, Athogavia 155
Scarabiphotia 208
scaurus, Apalotacarus 152
scissus, Apalotacarus 152
sellnicki, Canestrinia 196
sellnicki, Percanestrinia 196
sentosa, Melisia 188
Shilongia 208
sibirica, Photia 202
sicula, Grandiella 178
sicula, Grandiellina 178
silvanae, Boetophela 160
simplex, Coleoglyphus 167
Acarorum Catalogus IX 219
soniae, Grandiella 178
soniae, Grandiellina 178
sophiae, Sandrophela 207
Sorbinophela 208
spectanda, Canestrinia 207
spectanda, Sandrophela 207
spicantis, Arrunsithiana 154
spicantis, Grandiella 154
spicantis, Grandiellina 154
squarrosa, Arrunsithiana 154
squarrosa, Grandiella 154
ssavina, Mossongia 189
straeleni, Afrocanestrinia 148
suatoha, Donelaphontia 172
suatoha, Farahanella 172
superba, Melisia 187
sybiellae, Boetophela 160
symeoni, Boetophela 160
tacjannae, Grandiella 178
tacjannae, Grandiellina 178
Tamarangia 208
Tamarophela 209
tanzanica, Athogavia 156
Taseopus 209
Teklathiana 209
tenax, Thopia 211
Teophyssa 210
teresae, Teophyssa 210
tertia, Grandiella 178
tertia, Grandiellina 179
tertia, Mesophotia penicillata 189
tetracaudata, Grandiella 179
tetracaudata, Grandiellina 179
Thopia 211
togata, Canestriniella 165
transkeiensis, Diplopodocoptes 171
transversicola, Procericola ichtihyoides 203
trilobata, Amansiella 149
trullus, Apalotacarus 152
turcica, Photia procera 201
ubaldi, Oceogavia 190
unguiculata, Anaspistes 150
Uriophela 211
useguaica, Olgattia 192
Vandeinella 212
vatia, Bibulothiana 159
vatia, Grandiella 159
vatus, Cetonicola 166
Vereoxia 212
Vereoxiinae 212
vietnamensis, Contramelisia
vignoni, Scarabiphotia 208
violae, Grandiellopsis 180
viviannae, Percanestrinia 196
vlachiana, Canestrinia 197
vlachiana, Percanestriniella 197
volkeri, Beeriphotia 158
wauensis, Cooremania 170
werneri, Boetophela 160
Distribution of Canestriniidae
ЕUROPE
Albania (Samšinák, 1965) – Photia procustidis, Procericola bourgognei
Austria (Samšinák, 1971; Haitlinger, 1990d, 1995b) - Coleopterophagus albini, Photia
bardoica, Ph. chrysocarabi, Ph. hejniana
Belgium (Cooreman, 1950; Fain, Noti & Dufrêne, 1995) - Photia chrysocarabi, Ph. saetolata, Procericola bourgognei, Pseudamansia chrysomelinus
Bosna and Herzegovina (Samšinák, 1971; Haitlinger, 1995b) – Dicanestrinia knobi,
Pseudocanestrinia berndi, P. carabi
220
Acarorum Catalogus IX
Bulgaria (Samšinák, 1970; Beron, 1971, 1975) – Ibizella [Canestrinia] samsinaki, Percanestrinia blaptis, Procericola bourgognei, P. ichthyoides, Photia procera
Byelorussia (Khaustov & Eidelberg, 2001) – Dicanestrinia huberti, Photia adolphinae
Crimea (Khaustov & Eidelberg, 2001) – Amansiella trilobata, Canestrinia damiani, C.
dorcicola, Paramansia bicornis, Paraphotia calosomae, Photia polymorpha, Pseudocanestrinia berndti, P. carabi
Croatia (Samšinák, 1971; Haitlinger, 1990d ) - Coleopterophagus megnini, Photia hejniana
Czech Rep. (Samšinák, 1965, 1971; Haitlinger, 1990d) - Coleopterophagus albini, C. megnini, Dicanestrinia knobi, Percanestrinia sellnicki, Photia chrysocarabi, Ph. hejniana,
Ph. saetolata
Denmark (Haitlinger, 2017) - Paracanestrinia denmarkica
Estonia (Haitlinger, 2015) – Coleopterophagus albini
France (contin.) (Oudemans, 1923; Cooreman, 1950; Jolivet, 1950, 1954; Samšinák, 1970, 1971)
– N. jureceki, Paramansia menthastri, Percanestrinia blaptis, Photia chrysocarabi, Ph. saetolata, Procericola bourgognei, Pseudocanestrinia occidentalis, Pseudamansia chrysomelinus
Corsica (Haitlinger, 1992a) – Canestrinia carabicola, Percanestrinia maroccana
Georgia (Samšinák, 1971; Haitlinger, 1992a) – Percanestrinia blaptis, Photia bilkorum
Germany (Poppe, 1907; Rack, 1968; Samšinák, 1965, 1970; Haitlinger, 1992a) – Coleopterophagus albini, Percanestrinia blaptis, Procericola bourgognei, Photia chrysocarabi,
Ph. saetolata
Great Britain (Turk, 1953; Evans, Sheals & Macfarlane, 1961; Samšinák, 1965, 1971)
– Canestrinia dorcicola, Coleopterophagus megnini, Percanestrinia blaptis, Photia
chrysocarabi, Ph. saetolata
Greece (Cooreman, 1958; Samšinák, 1965) - Photia graeca, Ph. procustidis,, Procericola
bourgognei
Crete (Samšinák, 1965) – Photia procustidis
Rhodos (Samšinák, 1965) - Photia procustidis
Holland (Buitendijk, 1945; Cooreman, 1954; Samšinák, 1965; Siepel, Cremers & Vierbergen, 2016) – Canestrinia dorcicola, Coleopterophagus megnini, Photia saetolata
Hungary (Samšinák, 1965; Haitlinger, 1989, 1990d; 1992a) – Canestrinia damiani, Coleopterophagus megnini, Percanestrinia blaptis, P. sellnicki, Procericola bourgognei,
Italy (contin.) (Berlese, 1881; Lombardini, 1943; Cooreman, 1950; Haitlinger, 1990d,
1992a, 2004) – Canestrinia carabicola, C. dorcicola, Coleopterophagus donaldi, C.
megninii, Dicanestrinia knobi, Percanestrinia blaptis, P. maroccana, Photia procera,
Ph. procustidis, Procericola bourgognei, P. ichthyoides, Pseudamansia chrysomelinus
Sicily (Haitlinger, 1990d) - Canestrinia carabicola, Coleopterophagus megnini,
Sardinia (Canestrini & Berlese, 1880; Vitzthum,1924) – Canestrinia carabicola, Percanestrinia blaptis, P. sardica
Moldova (Trach, 2006) – Percanestrinia sellnicki, Photia hejniana, Procericola bourgognei
Norway (Haitlinger, 2015) - Coleopterophagus albini
Poland (Haitlinger, 1988a, 1988b, 1988c, 1988d, 1989, 1990c, 1990d, 1991c, 1992a, 1994a,
2002, 2004, 2012) - Canestrinia dorcicola, Coleopterophagus albini, C. megnini, Di-
Acarorum Catalogus IX 221
canestrinia huberti, D. knobi, Percanestrinia blaptis, P. sellnicki, Photia adolfinae (as
“polymorpha”), P. hejniana, P. chrysocarabi, P. hermengildae, P. bardoica, Pseudocanestrinia mahunkai, Procericola bourgognei
Portugal (cont.) (Samšinák, 1971) – Canestrinia hispanica, Neophotia drvotocka
Romania (Samšinák, 1965; Haitlinger, 1990d) – Coleopterophagus albini, C. maroni,
Percanestrinia sellnicki, Procericola bourgognei
Russia (all, continent.)(Samšinák, 1971; Haitlinger, 1992b; Khaustov & Eidelberg, 2001) –
Percanestriniella caucasica, Photia bilkorum, Ph. polymorpha, Ph. sibirica, Procericola
bourgognei, Uriophela arieli
Kunashir I. (Khaustov & Eidelberg, 2001) – Photia pacifica
Sakhalin I. (Khaustov & Eidelberg, 2001) – Photia lopatini
Serbia (Haitlinger, 19 ) – Coleopterophagus megnini
Slovakia (Haitlinger, 1990 ; Samšinák, 1965, 1971; Zamec, 2014) – Coleopterophagus
albini, Percanestrinia selnicki, Photia chrysocarabi, Ph. hejniana, Ph. procustidis,
Procericola bourgognei
Slovenia (Cooreman, 1950; Samšinák, 1971) – Photia hejniana, Ph. procera, Procericola
ichthyoides, Pseudocanestrinia orientalis
Spain (cont.) (Jolivet, 1954; Samšinák, 1971) - Canestrinia hispanica, Mesophotia p.
penicillata, M. p. similis, Neophotia drvotocka, N. latissima, Photia lusitanica, Pseudamansia chrysomelinus
Balearic Is. (Haitlinger, 1991a, 2001; Haitlinger & Šundić, 1918) - Camirohylla feziana
[Ibiza, Formentera], Ibizella samsinaki [Ibiza], I. balearica [Ibiza], Pseudamansia
chrysomelinus [Mayorca]
Sweden (Lundquist, 1985) – Procericola bourgognei
Turkey (all) (Samšinák, 1964, 1965, ; Haitlinger, 1990d) - Coleopterophagus megnini,
Photia graeca, Ph. procera adanensis, Ph. procera turcica, Ph. procustidis, Procericola
bourgognei, P. ichthyoides scabricola, P. i. transversicola
Ukraine (Haitlinger, 1990, 2002; Khaustov & Eidelberg, 2001; Samšinák, 1971; Trach,
2006; Trach & Khaustov, 2007, 2011) – Amansiella trilobata, Canestrinia dorcicola,
Coleopterophagus albini, C. donaldi, C. megnini, C. maroni, Dicanestrinia knobi, Paramansia bicornis, Paraphotia calosomae, Percanestrinia blaptis, P. sellnicki, Photia
adolfinae, Ph. hejniana, Ph. hermengildae, Ph. polymorpha, Procericola bourgognei,
Pseudamansia decorata, Pseudocanestrinia carabi, P. berndi
AFRICA
Algeria (Jolivet, 1954) - Pseudamansia chrysomelinus
Botswana (Haitlinger, 1993) - Sorbinophela sandyi
Burundi (Haitlinger, 1993) - Boetophila werneri
Cameroon (Haitlinger, 1990, 1991, 1991d, 1993, 2011) - Boetophela erhardi, Cetonicola
isadorae, Paraphagella grimaldi, P. indirae, P. odae, Pheliculophela roaldi, Saniothiana
barumbaica
Congo-Brazaville (Haitlinger, 1991d) – Cetonicola maritae, Paraphagella odae
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Acarorum Catalogus IX
Congo DR (Cooreman, 1953, 1955; Haitlinger, 1995) - Afrocanestrinia straeleni, Cetonicola hispidus, Coleopterophagus pulcher, Diplognatophilus africanus, Donnelaphontia
suatoha
Egypt (El-Bishlawy & Allam, 2003) – Percanestrinia egypti
Ethiopia (Haitlinger, 1990, 1991d, 1993; Khaustov, 2006) – Barbiangia elongata, B. ethiopica, Diplognatophilus ethiopicus, Tamarangia ethiopica, T. nimfae
Ghana (Haitlinger, 1991a, 1991d, 1993) - Boetophela ephraimi, Boleohylla garrica, Cetonicola orestesi
Guinea (Haitlinger, 2010) - Pseudoparaphagella gerardi
Kenya (Fain, 1987b) – Diplopodocoptes transkeiensis
Liberia (Haitlinger, 1991d) - Cetonicola orestesi
Madagascar (Haitlinger, 1989a, 1990) – Ambilohylla favosa, Athogavia nosiana
Morocco (Cooreman, 1953; Samšinák, 1971; Haitlinger, 1991; Haitlinger & Šundić, 1918)
– Canestrinia hispanica,Camirohylla feziana, Globosophotia bettinae, Ibizella asilahica
Mesophotia penicillata tertia, Percanestrinia maroccana, Pseudamansia chrysomelinus
Senegal (Haitlinger, 1995a) - Afrocanestrinia samsoni
Sierra Leone (Haitlinger, 1992) - Tamarangia fabiolae
South Africa (Trägårdh, 1906; Fain, 1987; Haitlinger, 1989, Haitlinger & Chmielewski,
2004) – Boetophela cassandrae, B. ginae, Diplopodocoptes transkeyensis, Megacanestrinia mucronata, Tamarangia nimfae
Sudan (Trägårdh, 1904) – Coleopterophagus quadrisetosus
Tanzania (cont.)(Lavoipierre, 1958; Haitlinger, 1989, 1990b, 1991, 1995a) - Afrocanestrinia
rufinae, Athogavia gamana, A. tanzanica, Barbiangia alvari [Zanzibar], Megacanestrinia beloniana, Boetophela robertsoni, B. silvanae, B. sybillae, B. symeoni, Cetonicola
vatus, Irmongia helgae, Olgattia useguaica
Zanzibar (Haitlinger, 1993) - Barbiangia alvari
Tunisia (Samšinák, 1970; Haitlinger, 2000) - Bircericola bertrami, Percanestrinia kamelskyi
Uganda (Haitlinger, 1991) - Boleohylla kiogana
“Africa” (Berlese, 1910, 1920; Haitlinger, 1990) – Canestrinia remigans, Coleopterophagus
neglectus, Saniothiana pycnosa
ASIA
China (Samšinák, 1971; Haitlinger, 1990d, 1992a, 1994a, 1998; Fan, Chen & Wang, 2010)
- Beeriphotia volkeri, Canestrinia kacperi, C. paavoi, C. pekingensis, C. ramoni, Coleopterophagus belzebubi, C. rudolfi, Gioharattia maharae, Percanestrinia melchiori, P.
viviannae, Photia melchiori
Hainan (Summers & Schuster, 1982a;) – Melisia sentosa
Taiwan (Samšinák, 1971; Summers & Schuster, 1981a; Haitlinger, 1990d; Fan, Chen &
Wang, 2010) – Apalotacarus aristatus, C. cristatus, A. trullus, Canestrinia illegalis,
Coleopterophagus belzebubi, Melisia superba
Cyprus (Samšinák, 1965; Haitlinger, 1992a) – Ibizella [Canestrinia] samsinaki, Percanestrinia blaptis, Photia procustidis
Acarorum Catalogus IX 223
India (Samšinák, 19 1971; Summers & Schuster, 1981a; Haitlinger, 1990, 1994a; Vishnupriya & Mohanasundaram, 1988) - Apalotacarus aristatus, A. echinatus, A. protensus,
A. scissus, “Coleopterophagus procerus”, Canestriniella batocerae (Tamil Nadu), Melisia
anomala, M. klapaleki, Percanestriniella vlachiana (Kashmir), Shillongia marzenae
(Assam), Sandrophela sophiae
Indonesia (Berlese, 1910; Samšinák, 1963, 1944; Summers & Schuster, 1981a; Haitlinger,
1989a,1990b, 1991, 1992b, 1993b, 1994a, 1995) – Albinorattia igori [Batjan], Amboinophela bernae [Amboina, New Guinea], Apalotacarus aristatus [Java], A. fulgens
[Java], A. sajanae [Java], Arraphosoma paraminax [Mollucas], Athogavia borneoica
[Borneo, presumably Kalimantan], A. scagana [Borneo, presumably Kalimantan],
Aurilossongia dioni, A. hoaria [Kalimantan], Canestriniella amplexans [Java], C.
physsana [Java], C. elae, C. phoebinae [Java], C. togata phoebinae [Java], C. phoebinae
[Java], Gioharattia maharae [Borneo, presumably Kalimantan], Hargeinella poppana
[Sumatra], Hyllosihemia belerofoni [Sumatra], Lidiophela bognae, L. celinae, L. pelagiae, Melisia jadwigae [Moluccas], Melisia ariadnae, M. gabrielae, M. hieronimae, M.
ingeborgae, M. irenae, M. lombardinii, M. mariettae, M. melaniae, M. salomeae, M.
sumatrae, Noemiphela barryi [Sumatra], N. izabelae [Sulawesi], Rugoniphela marloni
[Borneo, Java, Sumatra], Sandrophela amarae [Java, Sumatra], S. hugoni [Sumatra], S.
spectanda [Java, Sumatra], Tamarophela batianica [Moluccas], Vandeinella adriasae
[Sumatra], V. opressa [Borneo]
Iran (Haitlinger, 1990c; Ardeshir et al., 2016) – Camirohylla sp., Coleopterophagus baali
Israel and Palestina (Samšinák, 1965) - Dicanestrinia knobi, Photia procustidis
Japan (Kishida, 1924, Samšinák, 1971; Haitlinger, 1990d) - Canestrinia pictura, Coleopterophagus belzebubi, Photia polymorpha
Kambodia (Vomero, 1972) – Melisia samsinaki
Kirghizstan (Khaustov & Eidelberg, 2001) - Percanestriniella levis
Korea (Samšinák, 1971; Wontae Kim et al., 2006) - Haitlingeria longilobata, Photia polymorpha
Lebanon (Samšinák, 1964, 1965) - Photia procera marani, Ph. procustidis
Malaysia (cont.)
Sarawak and Sabah (Haitlinger, 1991) - Rugoniphela marioni [Borneo]
Philippines (Summers & Schuster, 1981a) - Apalotacarus fusulus [Mindoro], A. scaurus
[Mindanao]
Sri Lanka (Haitlinger, 1989a) – Gioharattia caroae
Syria (Haitlinger, 1990d) – Coleopterophagus dionizyi, C. maroni,
Taiwan (Summers & Schuster, 1981a) – Apalotacarus aristatus, A. trullus
Thailand (Haitlinger, 1990) - Melisia hieronimae
Uzbekistan (Samšinák, 1971) – Percanestrinia levis
Vietnam (Samšinák, 1969; Haitlinger, 1990b, 1993b) – Melisia baloghi, M. helenae, M.
jadwigae, M.hieronimae, M. mahunkai, M.occidii, M. rufae, Sandrophela rodioni
Asia (Haitlinger, 1989)
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Acarorum Catalogus IX
NORTH AMERICA
Mexico (Banks, 1915; Summers & Schuster, 1979a, 1982b; Haitlinger, 1992e, 1993d,
1993e) – Acrotacarus mirabilis, Arrunsithiana escaudata, Bibulothiana cyriaki, B.
vatia,Grandiellina marinae, G. soniae, Ennodithiana maniusi, Phriknodora pustulosa
SOUTH and CENTRAL AMERICA
Argentina (Haitlinger, 1992e, 1993e) - Grandiella tacjannae, Olafithiana foliacea, O.
multifida
Belize (Fain, 1987a) - Grandiella gambosa, G. kimsleyi
Bolivia (Lombardini, 1950; Fain, 1987a; Haitlinger, 1989, 1992e, 1993d, 1993e, 1999) –
Arrunsithiana escaudata, Bibulothiana brevispinata, Grandiella decemcaudata, G.
longini, G. quadrata, G. rosascostai, Olafithiana multifida, Ovarrothiana leopoldi, O.
ludomiri, Teklathiana onnae
Brazil (Lombardini, 1939, 1950; Summers & Schuster, 1979a; Fain, 1987a; Haitlinger,
1989, 1990a, 1991e, 1992e, 1993d, 1993e, 1995a, 2008) – Arrunsithiana arcuata, A.
battosi, A. escaudata, A. quadrata, A. spicantis, Bibulothiana brevispinata, B. cyriaki,
B. everittae, B. oswaldi, Dorynothiana delmari, D. jurandi, Ennodithiana maniusi, E.
medoni, E. megesi, Ennorithiana heliodori, Gasuthiana abapoica, G. coarica, G. melillae, Grandiella minor, G. tacjannae, G. tetracaudata, Hatohylla bahiana, H. bibiannae,
H. morrata, Olafithiana adelinae, O. multifida, O. foliacea, O. joliveti, Ovarrothiana
erazmi, O. ozannae, Sabothiana boi, S. reidari, S. riccardoi, Thopia boguwolae, Th.
corinnae
Colombia (Summers & Schuster, 1979a; Fain, 1987; Haitlinger, 1997, 1989, 1991b, 1991e,
1993d, 1993e, 2001) - Arrunsithiana ilosi, A. battosi, A. quadrata, Bibulothiana brevispinata, Danaithiana ayraica, Ennorithiana aidae, Gasuthiana olareica, Grandiella
gambosa, G. kimseyi, G. sicula, G. soniae, G. tetracaudata, Ovarrothiana echinoderma,
Teklathiana onnae,
Costa Rica (Haitlinger, 1991b, 1994b; Summers & Schuster, 1979b) – Acrotocarus mirabilis, Arrunsithiana escaudata, Bibulothiana albertae, Danaithiana chanchamayoica,
Olafithiana iwettae, Phriknodora avae
Cuba (Haitlinger, 1991b, 1993d) - Danaithiana chanchamayoica, Ennodithiana messalinae
Ecuador (contin.) (Fain, 1987a; Haitlinger, 1991b, 1991e, 1993d, 1993e,) – Arrunsithiana
artemoni, A. battoni, A. erinnae, Bibulothiana brevispinata, Danaithiana archidonaica;
D. ayraica, D. ecuadorica, Grandiella cooremani, G. ecuadorensis, G. soniae, Olafithiana
multifida, Ovarrothiana ozannae,
French Guyana (Lombardini, 1950; Fain, 1987a; Haitlinger, 1993e, 1993d, 1999) – Averithiana anoni, Arrunsithiana esacaudata, A. quadrata, A. squarrosa, Gasuthiana
olareica, Grandiella tetracaudata, G. t. omoplata, Ovarrothiana erazmi, Thopia amedae
Guatemala (Summers & Schuster, 1979a, 1979b) - Acrotocarus mirabilis, Arrunsithiana
esacaudata
Honduras (Haitlinger, 1993e, 1999) - Bibulothiana brevispinata, Metriothiana bolebori
Acarorum Catalogus IX 225
Panama (Banks, 1915; Summers & Schuster, 1979b, 1982b; Haitlinger, 1993d, 1993e, ) –
Bibulothiana brevispinata, Coleopterophagus blattophagus, Ennodithiana platysma,
Grandiella gambosa,G. kimseyi, Ovarrothiana echinoderma, Phriknodora pustulosa,
Teklathiana conula, T. rugosita, Thopia tenax
Paraguay (Lombardini, 1950; Haitlinger, 1992e) – Grandiella decemcaudata, G. tacjannae
Peru (Summers & Schuster, 1979; Fain, 1987a; Haitlinger, 1991b, 1992e, 1993d, 1993e,
2008) – Bibulothiana everittae, Danaithiana chanchamayoica, Grandiella rogeri, G.
soniae, G. tetracaudata, Ovarrothiana ozannae, Peruthiana briannae
Suriname (Haitlinger, 1993) – Ennodithiana penelopae
Trinidad and Tobago (Turk, 1948) – Acrotacarus alutaceus,
Venezuela (Lombardini, 1950; Haitlinger, 1994b) - Olafithiana iwettae, Grandiella pentagona
South America (Fain, 1989) – Arrunsithiana latipes
OCEANIA
Australia (continent) (Summers & Schuster, 1981a, 1981b, 1982a; Haitlinger, 1991) —
Apalotacarus fusticulus, A. gracilis, A. luroris, Cnecoderma dolichopoda, C. ovalis,
Hypopteryx collaris, Lidiophela pauliana, Melisia leptinotarsa, Taseopus platythrix
Papua New Guinea (Nesbitt, 1976; Summers & Schuster, 1981a, b; Haitlinger, 1989a,
1989b, 1990a, 1990b, 1993b, Haitlinger & Šundić, 2020) – Apalotacarus cidaris, A.
fusulus, A. paxillus, A. petilus, A. rigescens, Aurilossongia bolanica, A. fileasi, A. gaurana
[New Britain], A. norae, Lidiophela [Coleopterophagus] pecki, Cooremania kietaensis
[Bougainville], C. wauensis, Jullongia izae, Passalophagus georgei, Phaleratus fentoni,
Photia howdeni, Sajanophela alfredae
New Guinea (Summers & Schuster, 1981b) – Arraphosoma minax
Salomon Islands (Nesbitt, 1976; Summers & Schuster, 1981a) – Apalotacarus
glaber [Russel I.], Lidiophela [Coleopterophagus] pecki
Hosts of Canestriniidae
Myriapoda
Diplopoda
Glomerida
Fam. Glomeridae – Diplopodocoptes transkeiensis
Spirostrepsida
Fam. Odontopygidae - Diplopodocoptes transkeiensis
Insecta
Coleoptera
Fam. Carabidae
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Calosoma (Campalita) auropunctatum – Paraphotia calosomae
Carabus (Oreocarabus) amplipennis - Photia lusitanica
C. (Cychrostomus) anchocephalus – Canestrinia pekingensis
C. (Eupachys) glyptopterus - Canestrinia pekingensis
C. (Coptolabrus) lafossei saturatus - Canestrinia pekingensis
C. augustus - Canestrinia pekingensis
C. augustus ignigena - Canestrinia pekingensis
C. pustulifer - Canestrinia pekingensis
C. pustulifer pratti - Canestrinia pekingensis
C. p. querryi - Canestrinia pekingensis
C. p. szechwanensis - Canestrinia pekingensis
C. ignimetalla angusticollis - Canestrinia pekingensis
C. gemmifer - Canestrinia pekingensis
C. (Cratocephenus) akinini – Percanestriniella levis
C. angulicollis antaeus – Beeriphotia volkeri
C. (Tachypus) auratus – Photia chrysocarabi
C. (Chrysocarabus) auronitens - Photia chrysocarabi, Pseudocanestrinia
occidentalis
C. (Cratocephalus) balassogloi – Percanestriniella levis
C. (C.) bessarabicus – Pseudocanestrinia carabi
C. (Damaster) blaptoides – Canestrinia pictura
C. (Imaibius) boysi – Percanestriniella vlachiana
C. (Acanthaicus) brandti – Canestrinia pekingensis
C. (Megodontus) caelatus – Pseudocanestrinia orientalis
C. (C.) cicatricosus - Percanestriniella levis
C. (Procrustes) coriaceus - Photia procera, Percanestrinia saetolata, Procericola
bourgognei
C. (Megodontus) croaticus – Pseudocanestrinia orientalis
C. coelestes giganteus – Canestrinia ramoni
C. (Procrustes) a. anatolicus – Photia procustidis
C. (Procrustes) auticollis – Procericola bourgognei
C. (Procrustes) b. banoni – Photia procustidis
C. (Procrustes) ch. chevrolati – Procericola bourgognei
C. (Procrustes) clypeatus clypeatus - Procericola bourgognei
C. (Procrustes) coriaceus banaticus - Percanestrinia sellnicki
C. (Procrustes) c. coriaceus - Percanestrinia sellnicki, Photia saetolata
C. (Procrustes) coriaceus foudrasi – Photia graeca
C. (Procrustes) coriaceus gressorius - Percanestrinia sellnicki
C. (Procrustes) coriaceus imminutus - Percanestrinia sellnicki
C. (Procrustes) coriaceus impudicus - Photia graeca
C. (Procrustes) coriaceus kindermanni - Photia graeca
Acarorum Catalogus IX 227
C. (Procrustes) coriaceus pseudorugifer - Percanestrinia sellnicki
C. (Procrustes) coriaceus rugifer - Percanestrinia sellnicki
C. (Procrustes) coriaceus subrugosus - Percanestrinia sellnicki
C. (Procrustes) coriaceus – Pseudocanestrinia mahunkai
C. (Ctenocarabus) galicianus – Canestrinia hispanica
C. (Procerus) gigas – Photia procera, Procericola ichthyoides
C. (Procerus) scabrosus transversalis - Photia procera adanensis
C. (Procerus) scabrosus caucasicus - Photia procera
C. (Procerus) syriacus - Photia procera marani
C. (C.) glabratus – Photia adolphinae
C. grandis – Canestrinia paavoi
C. (Oreocarabus) guadarramus - Canestrinia hispanica, Neophotia drvotocka
C. (Megodontus) gyllenhali – Photia adolfinae, Pseudocanestrinia berndi
C. (Coptolabrus) hankotaizanus – Canestrinia illegalis
C. (Procrustes) impressus carmelita – Photia procustidis
C. (Megodontus) kolbei – Photia pacifica
C. (Tribax) komarovi – Photia bilkorum
C. (Oreocarabus) linnaei – Photia hermengildae
C. latenulatus plassensis - Pseudocanestrinia berndi
C. (Coptolabrus) longipennis mandarinus - Canestrinia illegalis
C. (Acoptolabrus) lopatini – Photia lopatini
C. (Mesocarabus) lusitanicus – Neophotia latissima
C. (M.) lusitanicus breuningi – Photia lusitanica
C. (M.) lusitanicus castilianus – Photia lusitanica
C. (M.) lusitanicus logronicus - Photia lusitanica
C. (M.) lusitanicus schaumi - Photia lusitanica
C. (Rhabdotocarabus) melancholicus – Canestrinia hispanica, Mesophotia p.
penicillata
C. (Macrothorax) morbillosus – Canestrinia carabicola
C. (Procrustes) m. mulsatianus – Photia procustidis
C. (Archicarabus) nemoralis – Photia bardoica, Ph. chrysocarabi
C. (Procrustes) payafa - Procericola bourgognei
C. (Tribax) plasoni – Photia bilkorum
C. (Tribax) protensis – Photia bilkorum
C. (Iniopachus) pyrenaeus – Neophotia drvotocka, N. jureceki
C. (Procerus) scabrusus - Photia procera
C. (Megodontus) schoenherri – Photia sibirica
C. (Tribax) shamyli – Photia bilkorum
C. (I.) stoliczkanus - Percanestriniella vlachiana
C. (Ulocarabus) sturowskyi - Percanestriniella levis
C. (P.) tanypedilus - Percanestriniella levis
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C. (Pantophyrtus) turcomannorum - Percanestriniella levis
C. (Hygrocarabus) variolosus – Dicanestrinia huberti, D. knobi
C. (Megodontus) violaceus – Photia hejniana, Ph. saetolata
C. v. bicinctus – Photia saetolata
C. v. exasperatus – Photia hejniana, Ph. saetolata
C. v. fioris – Photia hejniana
C. v. fulgens – Photia saetolata
C. v. obliquus - Photia hejniana
C. v. purpurascens – Photia graeca, Ph. saetolata
C. v. savinicus - Photia hejniana
C. v. styriensis - Photia hejniana
Pseudosteropus globosus – Globosophotia bettinae
Sphodrus leucophthalmus - Percanestrinia blaptis
Tefflus purpuripennis – Megacanestrinia beloniana
Carabus sp. – Percanestriniella caucasica, Paracanestrinia denmarkica
Carabidae gen. sp. – Megacanestrinia mucronata
Fam. Cerambycidae
Batocera albofasciata octomaculata – Canestriniella phoebinae
B. celebiana - Canestriniella physsana
B. ferruginea – Canestriniella elae
B. hector – Canestriniella amplexans
B. humilidens – Canestriniella physsana
B. lunulata – Canestriniella togata
B. procera - Canestriniella togata
Brazilianus batus – Phriknodora pustulosa
B. brazilianus - Phriknodora pustulosa
Cerambyx cerdo – Dicanestrinia cerambycis
Lagocheirus foveolatus – Thopia pannucea
Neoptychodes trilineatus – Thopia bulbocaudata
Sternotomis bohemanni – Anaspistes unguiculatus
Taeniotes scalatus – Thopia tenax
Trachyderes succinctus – Phriknodora scabra
Xylotrupes asperulus – Aurilossongia bolanica
Xylotrupes sp. – Aurilossongia gaurana, A. hoarica
Cerambycidae gen. sp. – Chelinophora dictyophora
Fam. Chrysomelidae
Acromis nebulosa – Ennodithiana penelopae
A. sparsa - Teklathiana onnae, T. rugosita
A. spinifex – Ennodithiana medoni
Agenyssa caedemadens – Ovarrothiana ozannae
A. crassicornis - Ovarrothiana ozannae
Acarorum Catalogus IX 229
Alurnus bipunctatus - Gasuthiana olareica
A. boulsardi – Gasuthiana olareica
A. clysianus – Gassuthiana abapoica
Anacassis cribrum – Olafithiana adelinae
A. fuscata - Grandiella tacjannae
Botanochara angulata – Grandiella tacjannae
B. bonariensis - Grandiella tacjannae
B. convexiuscula - Grandiella tacjannae
B. duodecimverrucata - Grandiella tacjannae
B. impressa - Grandiella tacjannae
B. macularia - Grandiella tacjannae
B. nervosa - Grandiella tacjannae
B. regina - Grandiella tacjannae
B. ruforeticulata - Grandiella tacjannae
B. sanguinea - Grandiella tacjannae
B. sedecimpustulata - Grandiella tacjannae
B. tesselata - Grandiella tacjannae
B. texta - Grandiella tacjannae
Canistra carbonaria – Grandiellopsis arriani
C. cruentata – Grandiellopsis violae
C. osculatii – Grandiellopsis alani, G. canistra
C. plagosa – Arrunsithiana spicantis
Charidotella kesseli – Sabothiana reidari
Ch. (= Metriona) testudinacea – Olafithiana iwettae
Charidotis furunculus - Sabothiana riccardoi
Charidotis sp. – Sabothiana boi
Chelymorpha comata – Bibulothiana albertae
Ch. Infecta - Bibulothiana cyriaki
Ch. inflata – Bibulothiana everittae
Ch. nigricollis – Bibulothiana cyriaki
Ch. rufoguttata - Bibulothiana everittae
Ch. vermiculata - Bibulothiana cyriaki
Chrysolina fastuosa – Paramansia bicornis
Ch. mentastri – Paramansia bicornis, P. mentastri
Ch. polita - Paramansia bicornis
Coptocycla adamantina – Grandiella tetracaudata
C. contenta - Grandiella decemcaudata
C. nigropunctata - Grandiella sicula
C. wagneri - Grandiella sicula
Coptocycla sp. – Ennodithiana platysma
Crambelea illuderis – Ennodithiana megesi
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Acarorum Catalogus IX
Cyclosoma mirabilis – Grandiellopsis orsenae
C. paliata – Grandiellopsis cyclosoma
C. tristis – Ovarothiana ludomiri
Cyrtonota cyanea - Grandiella tetracaudata
C. dissecta – Arrunsithiana ilosi
C. latevittata – Arrunsithiana artemoni
C. punicea – Grandiella soniae
C. textilis - Grandiella soniae
C. thalassina - Grandiella tetracaudata
C. tigrina - Grandiella tetracaudata
C. tristigma – Bibulothiana vatia
Discomorpha batesi - Grandiella gambosa
D. lanuginosa pubescens - Grandiella gambosa
D. depilata - Grandiella gambosa
Dolichotoma conspersipennis - Grandiella gambosa
Dorynota pugionata – Dorynothiana jurandi
Dorynota sp. – Dorynothiana delmari
Echoma [as Omoplata] marginata – Arrunsithiana escaudata, Grandiellina
decemcaudata
E. clypeata – Grandiella tetracaudata
E. confluens – Grandiella marinae
Elytrosphaera xanthopyga – Olafithiana joliveti
Eudicella gralli var. meckowi – Paraphagella eudicellae
E. smithi var. bertherandi – Paraphagella eudicellae, P. minor
Eugenysa andicola – Ovarrothiana ozannae
E. bacchus - Ovarrothiana erazmi
E. columbiana – Ovarrothiana echinoderma, Arrunsithiana quadrata
E. delicata - Ovarrothiana nikodemi
E. divalis – Ovarrothiana nikodemi
E. grossa - Ovarrothiana erazmi, Arrunsithiana squarrosa
E. venosa – Ovarrothiana erazmi, Thopia amedae
Eumolpus fulgidus - Grandiella tetracaudata
Goniochenia (Baranosa) decolor – Grandiella rogeri
G. (B.) heroldi - Grandiella rogeri
G. (B.) flavosparsa - Grandiella longini
Mecistomala marginatus – Gassuthiana coarica
M. nigripes – Gassuthiana abufarica
Metriona testudinacea – Olafithiana iwettae
Metrionella bilimeki - Metriothiana bolebori
Myocalaspis gentilis – Ovarrothiana leopoldi
Omaspides basilica - Ennodithiana messalinae
O. bistriata – Teklathiana conula
Acarorum Catalogus IX 231
Physonota alutacea – Acrotacarus alutaceus, A. mirabilis
Ph. diluta – Acrotacarus crataepus
Plagiometriona ludiera – Ennorithiana heliodori
P. punctipennis - Ennorithiana heliodori
Polychalca dentipennis – Thopia corinnae
P. punctatissima – Grandiellopsis odiosi, Thopia boguwolae
Psalidonota contemta – Ennodithiana maniusi
P. dorsoplagiata - Ennodithiana messalinae
P. dorsopunctata – Ennodithiana messalinae
P. leprosa - Ennodithiana messalinae
Pycnocerus sp. – Saniothiana pycnosa
Stolas aenea - Arrunsithiana quadrata
S. antiqua – Olafithiana multifida
S. areolata - Olafithiana foliacea
S. aurosetosa - Olafithiana multifida
S. bifenestrata - Bibulothiana brevispinata
S. bipunctata - Arrunsithiana quadrata
S. bipustulata - Bibulothiana brevispinata
S. chalybaea – Arrunsithiana spicantis, A. battosi, Grandiella tetracaudata,
Olafithiana foliacea
S. coalita - Grandiella soniae
S. coerulescens – Grandiella soniae, Bibulothiana brevispinata
S. conspersa - Olafithiana foliacea, Arrunsithiana battosi, A. spicantis
S. cruentata - Olafithiana multifida, Arrunsithiana battosi
S. decemguttata - Arrunsithiana arcuata
S. demissa - Arrunsithiana quadrata
S. ephippium - Bibulothiana brevispinata
S. erichsoni - Grandiella soniae, Arrunsithiana battosi
S. esignata - Arrunsithiana arcuata
S. funebris - Arrunsithiana battosi, A. spicantis
S. gibbosa - Arrunsithiana battosi
S. huanocensis - Arrunsithiana battosi
S. ignita - Arrunsithiana battosi
S. imitans - Arrunsithiana quadrata
S. implexa - Olafithiana foliacea
S. impluviata - Olafithiana foliacea
S. inaequalis – Arrunsithiana latipes, A. quadrata
S. indigacea - Arrunsithiana quadrata
S. inexculta - Arrunsithiana battosi
S. lacunosa - Olafithiana foliacea
S. latevittata – Arrunsithiana artemoni
S. lebesi – Arrunsithiana escaudata, Phriknodora avae
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S. lenis - Olafithiana foliacea, Arrunsithiana battosi
S. lineaticollis - Bibulothiana brevispinata
S. niobe - Arrunsithiana battosi
S. nudicollis - Olafithiana foliacea
S. oblita - Olafithiana foliacea, Arrunsithiana battosi
S. paranensis - Olafithiana foliacea, Arrunsithiana battosi
S. pectinata – Arrunsithiana erinnae
S. placida - Grandiella soniae
S. plagiata - Bibulothiana brevispinata
S. plagicollis – Bibulothiana oswaldi
S. pleurosticha - Arrunsithiana battosi
S. punicea – Arrunsithiana escaudata, Grandiellina soniae
S. redtenbacheri - Arrunsithiana quadrata
S. scoparia - Arrunsithiana battosi
S. subreticulata - Olafithiana foliacea
S. subrugosa - Grandiella tetracaudata
S. thoreyi - Arrunsithiana quadrata
Stolas sp. – Arrunsithiana squarrosa
Timarcha atlantica - Pseudamansia chrysomelinus
T. balearica - Pseudamansia chrysomelinus
T. goettingensis - Pseudamansia chrysomelinus, P. decorata
T. interstitialis - Pseudamansia chrysomelinus
T. nicaensis - Pseudamansia chrysomelinus
T. pauperata - Pseudamansia chrysomelinus
T. punctella - Pseudamansia chrysomelinus
T. rugosa - Pseudamansia chrysomelinus
T. tenebricosa – Amansiella trilobata, Pseudamansia chrysomelinus
Zatrephina lineata – Averithiana anoni
Hispinae gen. sp. – Peruthiana briannae
Chrysomelidae gen. sp. – Acrotacarus mirabilis
Fam. Colydiidae
Colydium sp. – Sandrophela dittae
Fam. Elateridae
Semiotus furcatus - Danaithiana ecuadorica, D. archidonaica
S. imperialis - Danaithiana chanchamayoica
S. virgatus - Danaithiana ayraica
Fam. Lucanidae
Cladognathus giraffa - Noemiphela izabelae
Dorcus bucephalus – Sandrophela spectanda
D. paralellipipedus – Canestrinia dorcicola
Eumelasomus sansibaricus -Olgattia useguaica
Acarorum Catalogus IX 233
Eurytrachelus anthaeus – Rugoniphela marloni
Eurytrachelus bucephalus – Rugoniphela marloni
Eurytrachelus eurycephalus - Rugoniphela marloni, Sandrophela amarae
Eurytrachelus gypeatus - Sandrophela amarae
Eurytrachelus platymelus - Sandrophela rodioni
Eurytrachelus purpurescens - Noemiphela barryi
Hemisodorcus chevrilati – Sandrophela sophiae
Hemisodorcus [Macrodora] rubrofemoratus - Uriophela arieli
Metapodonotus bison - Amboinophela bernae
Metapodonotus cinctus - Amboinophela bernae
Odontolabis wollastoni – Sandrophela hugoni
Serrognathus platymelus – Haitlingeria longilobata
Telodorcus saiga – Canestrinia microdisca
Lucanidae indet. - Vereoxia bogeri, Cooremania kietaensis
Fam. Passalidae
Aceraius grandis – Apalotacarus aristatus, Melisia mariettae, M. salomeae
A. grandis hirsutus – Apalotacarus cristatus, Melisia superba
A. helferi – Melisia samsinaki
A. laevicollis – Melisia gabrielae
A. mayeri – Melisia melaniae
Epiphanus indicus – Melisia anomala
E. stoliczkae - Melisia anomala, M. klapaleki
Eriocnemis tridens – Apalotacarus sajanae
Eumelasomus sansibaricus – Olgattia useguaica
Gonatus noviculator - Teophyssa honoratae
Labienus gigax – Lidiophela igae, L. pelagiae
L. ptox – Apalotacarus fusulus, Lidiophela arnica
Labienus sp. - Passalophagus georgei, Lidiophela [Coleopterophagus] pecki
Leptaulax bicolor formosanus - Apalotacarus trullus
L. indicus – Shillongia marzenae
Mastochilus australasiens – Apalotacarus luroris
M. quaestionis - Apalotacarus luroris
Macrolinus nicobaricus – Apalotacarus protensus
M. weberi – Apalotacarus scaurus
Obiensus sp. – Lidiophela [Coleopterophagus] pecki
Pelopides mniszecki – Melisia lombardinii
P. tridens – Apalotacarus fulgens
Pharochilus dilatus – Apalotacarus luroris
Ph. nitidulus – Apalotacarus fusticulus, A. luroris
Pleurarius brachyphyllus – Apalotacarus echinatus, A. scissus, Phaleratus
alatus
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Protomocoelus australis – Apalotacarus cidaris, A. petilus, Sajanophela alfredae,
Lidiophela [Coleopterophagus] pecki
P. triumphator - Apalotacarus glaber
Passalidae gen. sp. – Apalotacarus gracilis, A. paxillus, A. rigescens, Arraphosoma minax, A. paraminax, Cnecoderma dolichopoda, C. ovalis, Hyllosihemia belerofoni, Jowitella barbarae, Jullongia izae, Lidiophela bognae,
L. celinae, Melisia helenae, M. ingeborgae, M. irenae, M. jadwigae, M.
leptinotarsa, M. rutae, M. sentosa, Tamarophela batjanica
Fam. Passandridae
Passandra trigemina – Sandrophela kokodaica
Fam. Scarabaeidae
Caelorrhina thoreyi – Pseudoparaphagella gerardi
Cetonia aurata – Coleopterophagus megnini
Coglodera diaroi – Gioharattia caroae
Coryphocera sp. – Gioharattia caroae
Dicronorhina micans – Afrocanestrinia straeleni, Cetonicola hispidus, Paraphagella princeps
D. oberthuri – Boetophela robertsoni
Diplognatha gagates – Barbiangia elongata
Elasocheilon haterospila – Boetophela symeoni
Eudicella smithi – Boetophela ginae
E. gralli var. meckowi – Canestrinia hispidus
Geynodonta laeviplaga – Boetophela silvanae
Goliathus alborignathus – Boetophela sybillae
Goliathus goliathus - Phelliculophela roaldi
Ischipsophia olivacea – Albinorattia igori
Mecynorrhina [Chelarrhina – recte Chelorrhina] savagei – Boetophela erhardi,
Cetonicola isadorae, Paraphagella indirae
Meristomala marginatus – Gasuthiana coarica
M. nigriceps – Gasuthiana abufarica
Oryctes monoceros – Athogavia gamana, A. tanzanica
O. pyrrhus - Athogavia nosiana
Oryctes sp. – Athogavia scagana
Oxythirea funesta – Canestrinia dorcicola
Pachnoda abyssinica – Tamarangia ethiopica
P. carmelita – Kahoorangia almae
P. marginata – Tamarangia ethiopica, T. fabiolae, T. flawiani
P. thoracica – Tamarangia nimphae
Papuana woodlarkiana – Oceogavia ubaldi
Plaesiorrhina mhondana – Boetophela werneri
Acarorum Catalogus IX 235
Poecilophila hebraea – Cetonicola vatus
P. maculatissima – Sorbinophela sandyi
Porphyroneta cinnamonea – Coleopterophagus pulcher
Protaetia (Potosia) aerata - Coleopterophagus belzebubi
Protaetia (Potosia) affinis – Coleopterophagus donaldi, C. megnini
Protaetia (Potosia) brevitarsis - Coleopterophagus rudolfi
Protaetia (Potosia) cuprea – Coleopterophagus dionizyi
Protaetia (Potosia) cuprea ignicollis - Coleopterophagus megnini
Protaetia (Potosia) cuprea cuprea - Coleopterophagus megnini
Protaetia (Potosia) cuprea obscura - Coleopterophagus megnini
Protaetia (Potosia) cuprina - Coleopterophagus megnini
Protaetia (Potosia) fieberi - Coleopterophagus megnini
Protaetia (Potosia) metallica - Coleopterophagus megnini, C. albini
Protaetia (Potosia) speciosa – Coleopterophagus baali
Protaetia (Potosia) speciosa jousselini – Coleopterophagus maroni
Protaetia (Cetonischema) aeruginosa – Coleopterophagus albini, C. maroni
Protaetia (Eupotosia) affinis – Coleopterophagus donaldi, C. megnini
Protaetia (Liocola) marmorata - Coleopterophagus albini
Rhabdotis sobrina – Diplognathophilus africanus
Scarabaeus variolosus – Scarabiphotia vignoni
Smaragdesthes africana mutica – Paraphagella princeps
S. oertzeni – Afrocanestrinia rufinae
Stephanorrhina guttata – Cetonicola maritae, Paraphagella odae
Tmesorrhina iris – Cetonicola orestesi, Paraphagella ingridae
Trichogomphus lunicollis – Vandeinella opressa
T. milon – Hargeinella poppana, Vandeinella adriassae
T. nartolani – Athogavia assamica
“Scarabeus” centaurus – Coleopterophagus neglectus
Scarabaeidae (Cetoniinae) – Babiangia alvari, B. ethiopica, Boetophela
cassandrae, B. ephraimi, Coleopterophagus quadrisetosus
Scarabaeidae (Dynastinae) – Athogavia arybbasi, Aurilossongia fileasi
Scarabaeidae gen. sp. – Anaspistes unguiculatus
Fam. Tenebrionidae
Amarygmus striatus – Lidiophela pauliana
Blaps appendiculata – Camirohylla feziana
B. gibbosa – Percanestrinia blaptis, P. izabelae
B. hejconota cortosa – Percanestrinia norodomi
B. lethifera - Percanestrinia blaptis
B. mucronata – Percanestrinia maroccana
B. occulta - Percanestrinia blaptis
B. pinguis - Percanestrinia maroccana
B. polychresta – Percanestrinia egypti
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B. producta - Percanestrinia maroccana
B. punctata - Percanestrinia blaptis
Blaps sp. – Ibizella [Canestrinia] samsinaki, Ibizella balearica
Damatria formosus – Ambylohylla favosa
Dila sp. - Camirohylla sp.
Gnaptor sp. – Ibizella [Canestrinia] samsinaki
Metallonotus metallicus – Boleohylla garrica
Nyctobates gigas - Hatohylla morrata
N. maximus – Hatohylla bahiana
Nyctobates sp. – Grandiella [Hatohylla] rosascostai
Odontopezus cupreus – Donnelaphontia cuatoha
Odontopezus cupreus lucens - Saniothiana barumbaica
Psammodes carbonarius – Boleohylla kiogana
Tenebrionidae gen. sp. – Bicercicola bertrami, Gasuthiana melillae, Hatohylla
bibiannae
Coleoptera indet. – Ennodithiana platysma, Grandiella rugosita, Melisia baloghi, M.
helenae, Phaleratus fentoni
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Santiago-Blay J.A., A. Fain. 1994. Phoretic and ectoparasitic mites (Acari) of the Chrysomelidae. In: Jolivet P.H., Cox M.L., Petitpierre E. (Eds). Novel aspects of the biology of
Chrysomelidae, Series Entomologica book series (SENT, volume 50): 407–417.
Siepel H., H. Cremers, B. Vierbergen. 2016. Provisional checklist of the astigmatic mites of the
Netherlands (Acari: Oribatida: Astigmatina). – Nederlandse Faunistische Mededelingen,
47: 49–88.
Summers F.M., R.O. Schuster. 1979a. Eleven new species of Grandiella Lombardini (Acari,
Canestriniidae). – Intern. J. of Acarology, 5(4): 325–350.
Summers F.M., R.O. Schuster. 1979b. Revision of Acrotacarus Banks (Acari: Canestriniidae).
- Proc. Entom. Soc. Washington, 81: 475–485.
Summers F.M., R.O. Schuster. 1981a. Apalotacarus, a new genus of canestriniid mites from
passalid beetles. – Internat. J. of Acarology, 7(1–4): 17–38.
Summers F.M., R.O. Schuster. 1981b. Three new genera of canestriniid mites associated with
passalid beetles. - Internat. J. of Acarology, 7(1–4): 39–46.
Summers F.M., R.O. Schuster. 1982a. The Melisiinae, new subfamily of mites in the Canestriniidae associated with passalid beetles. - Internat. J. of Acarology, 8(1): 23–31.
Summers F.M., R.O. Schuster. 1982b. New canestriniid mites from beetles of the family Cerambycidae. – Int. J. Acarol., 8(1): 33–46.
Théodoridès J. 1955. Contribution à l’étude des parasites et phorétiques de coléoptères terrestres. – Actualités scientifiques et industrielles 1217, Paris, Hermann & Cie, Eds. - Vie
et Milieu, Supplements 4: 1–310.
Acarorum Catalogus IX 243
Trach V.A. 2006. The host-parasite associations of the mites family Canestriniidae (Acari:
Astigmata) in Ukraine and Moldova. – Vestnik odesskogo nat. universiteta. Biologiya,
11(9): 174–180 (in Ukrainian, sum. Russ., Engl.).
Trach V.A., A.A. Khaustov. 2007. A review of mites of the family Canestriniidae (Acari, Astigmata) associated with the leaf beetles (Coleoptera, Chrysomelidae) in Ukraine with description of a new species of the genus Pseudamansia. – Vestnik Zoologii, 41(3): 267–272.
Trach V.A., A.A. Khaustov. 2011. A review of the genus Coleopterophagus Berlese, 1882 (Acari:
Astigmata: Canestriniidae) of Ukraine. – Acarina, 19(2): 213–230.
Trägårdh I. 1906. Neue Acariden aus Natal and Zululand. – Zool. Anzeiger, 30: 870–887.
Turk F. A. 1948. Insecticolous Acari from Trinidad, B. W. I. – Proc. Zool. Soc. London, 118,
art 1: 82–125.
Turk F.A. 1953. A synonymic catalogue of British Acari: Part II. – Annals and Magazine of
Nat. History, 12 ser., 62, 6: 81–99.
Vishnupriya R., M. Mohanasundaram. 1988. Mites associated with insects in Tamil Nadu,
India. – Entomon, 13(3–4): 247–257.
Vitzthum H.G. 1925. Acarologische Beobachtungen. 8. Reihe. – Arch. Naturgesch., 90 (1924):
A(10): 1–86.
Vomero V. 1972. Una nuova specie del genere Melisia Lombardini, 1944, associata a coleotteri
passalidi (Acarina – Sarcoptiformes). – Acarologia, 14(1): 95–108.
The Mite Family Chetochelacaridae Fain
Chetochelacaridae Fain, 1987: 161
One genus and species of African mites associated with Diplopoda.
Fam. Chetochelacaridae Fain
Chetochelacaridae Fain, 1987: 161
Type genus: Chetochelacarus Fain, 1987
Genus Chetochelacarus Fain
Chetochelacarus Fain, 1987: 162
Type species: Chetochelacarus mamillatus Fain, 1987
Chetochelacarus mamillatus Fain
Chetochelacarus Fain, 1987: 162
Type locality: Kivu, Djuma Valley
Distribution: D.R. Congo (former Zaire)
Host: millipede (Diplopoda: Julidae)
244
Acarorum Catalogus IX
Index of Chetochelacaridae
(Names in bold are accepted as valid)
Chetochelacaridae 243
Chetochelacarus 243
mamillatus, Chetochelacarus 243
Distribution of Chetochelacaridae
AFRICA
D.R. Congo (Fain, 1987) – Chetochelacarus mamillatus
Hosts of Chetochelacaridae
Myriapoda
Diplopoda
Julida
Fam. Julidae
Gen. sp. indet. - Chetochelacarus mamillatus
Bibliography of Chetochelacaridae
Fain A. 1987. Notes on mites associated with Myriapoda 1. Three new astigmatic mites from
Afrotropical Myriapoda (Acari, Astigmata). – Bull. Inst. R. Sci. Nat. de Belgique, Entomologie, 57: 161–172.
The Mite Family Lophonotacaridae Fain
Lophonotacaridae Fain, 1987: 163
One genus and species of African mites associated with Diplopoda.
Family Lophonotacaridae Fain
Lophonotacaridae Fain, 1987: 163
Type genus: Lophonotacarus Fain, 1987
Acarorum Catalogus IX 245
Genus Lophonotacarus Fain
Lophonotacarus Fain, 1987: 164
Type species: Lophonotacarus minutus Fain, 1987
Lophonotacarus minutus Fain
Lophonotacarus minutus Fain, 1987: 164
Type locality: Transkei, District Lusikisiki, Mouth of Riv. Mzimhlava
Distribution: South Africa
Host: millipede (Diplopoda: Glomeridae)
Index of Lophonotacaridae
(Names in bold are accepted as valid)
Lophonotacaridae 244
Lophonotacarus 245
minutus, Lophonotacarus 245
Distribution of Lophonotacaridae
AFRICA
South Africa (Fain, 1987) – Lophonotacarus minutus
Hosts of Lophonotacaridae
Myriapoda
Diplopoda
Glomerida
Fam. Glomeridae
Gen. sp. indet. - Chetochelacarus mamillatus
Bibliography of Lophonotacaridae
Fain A. 1987. Notes on mites associated with Myriapoda 1. Three new astigmatic mites from
Afrotropical Myriapoda (Acari, Astigmata). – Bull. Inst. R. Sci. Nat. de Belgique, Entomologie, 57: 161–172.
246
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The Mite Family Heterocoptidae Fain
Heterocoptidae Fain, 1961: 91
Mites associated with chrysomelid beetles (Cassidinae and Hispinae), other Coleoptera
(Lycidae, Erotylidae, Passandridae) and Hymenoptera (Apidae, Sphecidae), mostly in the
Afritropical and Indomalayan Regions. The first genus and species (Heterocoptes tarsii
Fain, 1967) has been found on Tarsius (Mammalia: Primates)(chance host ?). Haitlinger
described since 1990 10 of the 12 genera, known so far, and most of the species from
Africa, Madagascar, South Asia, Peru, Australia, New Guinea and other tropical areas.
Composition of Heterocoptidae
Fam. Heterocoptidae Fain, 1967–13 gen., 39 sp.
Abboticoptes Haitlinger, 1996–1 sp.
Amarucoptes Haitlinger, 2001–1 sp.
Bahrucoptes Haitlinger, 1999 – 1 sp.
Cassiocoptes Haitlinger, 1996–1 sp.
Conchycoptes Haitlinger, 2002–1 sp.
Erotylocoptes Fain, 1987–7 sp.
Gascaricoptes Haitlinger, 2002–1 sp.
Heterocoptes Fain, 1967–15 sp.
Honiarrea Haitlinger, 1990–1 sp.
Kamerucoptes Haitlinger, 2002–5 sp.
Nolaecoptes Haitlinger, 1996–2 sp.
Tannuicoptes Haitlinger, 1999–2 sp.
Uvallicoptes Haitlinger, 1999–1 sp.
Catalogue of Heterocoptidae
Family Heterocoptidae Fain
Heterocoptidae Fain, 1967: 91
Type genus: Heterocoptes Fain, 1967
Genus Abboticoptes Haitlinger
Abboticoptes Haitlinger, 1996: 988
Type species: Abboticoptes eddae Haitlinger, 1996
Acarorum Catalogus IX 247
Abboticoptes eddae Haitlinger
Abboticoptes eddae Haitlinger, 1996: 988
Type locality: Philippines
Distribution: Philippines
Host: Aspidomorpha quadrilobata (Coleoptera: Chrysomelidae, Cassidinae)
Genus Amarucoptes Haitlinger
Amarucoptes Haitlinger, 2001: 112
Type species: Amarucoptes eddae Haitlinger, 2001
Amarucoptes busannae Haitlinger
Amarucoptes busannae Haitlinger, 2001: 113
Type locality: 30 km east from Puerto Maldonado, at river Madre de Dios
Distribution: Peru
Host: Coleoptera: Passalidae indet.
Genus Bahrucoptes Haitlinger
Bahrucoptes Haitlinger, 1999: 60
Type species: Bahrucoptes amaliae Haitlinger, 1999
Bahrucoptes amaliae Haitlinger
Bahrucoptes amaliae Haitlinger, 1999: 60
Type locality: Kota Bahru
Distribution: Malaysia
Host: Aspidomorpha elevata (Coleoptera: Chrysomelidae, Cassidinae)
Genus Cassiocoptes Haitlinger
Cassiocoptes Haitlinger, 1996: 986
Type species: Cassiocoptes mikki Haitlinger, 1996
Cassiocoptes mikki Haitlinger
Cassiocoptes mikki Haitlinger, 1996: 986
Type locality: Soekaranda (Sumatra)
Distribution: Indonesia (Sumatra)
Host: Lasiochila fallax (Coleoptera: Chrysomelidae, Hispinae)
Genus Conchycoptes Haitlinger
Conchycoptes Haitlinger, 2002: 25
Type species: Conchycoptes tangaicus Haitlinger, 2002
Conchycoptes tangaicus Haitlinger
Conchycoptes tangaicus Haitlinger, 2002: 25
248
Acarorum Catalogus IX
Type locality: Tanga
Distribution: Tanzania
Host: Conchyloctenia hybrida (Coleoptera: Chrysomelidae, Hispinae)
Genus Erotylocoptes Fain
Erotylocoptes Fain, 1987: 244
Type species: Erotylocoptes linodesmus Fain, 1987
Erotylocoptes anthophora Fain
Erotylocoptes anthophora Fain, 1987: 251
Type locality: Mobwasa
Distribution: DR Congo [Zaire]
Host: Anthophora acraensis (Hymenoptera, Apidae)
Erotylocoptes helenae Haitlinger
Erotylocoptes helenae Haitlinger, 1996: 980
Type locality: ?Radburg (RSA)
Distribution: Mozambique, South Africa
Hosts: Laccoptera rugosicollis (Coleoptera: Chrysomelidae, Cassidinae)
Erotylocoptes linodesmus Fain
Erotylocoptes linodesmus Fain, 1987: 244
Type locality: Mediakoko
Distribution: DR Congo [Zaire]
Host: Linodesmus coccus apicalis (Coleoptera, Erotylidae)
Erotylocoptes luluensis Fain
Erotylocoptes luluensis Fain, 1987: 252
Type locality: Lulua-Kapanga
Distribution: DR Congo [Zaire]
Host: Chlorion (Isodontia) simoni (Hymenoptera, Sphecidae)
Erotylocoptes taorettae Haitlinger
Erotylocoptes taorettae Haitlinger, 1996: 981
Type locality: ?Pugu
Distribution: Tanzania
Host: Laccoptera aurosa (Coleoptera: Chrysomelidae, Cassidinae)
Erotylocoptes verenae Haitlinger
Erotylocoptes verenae Haitlinger, 1996: 982
Type locality: Congo (Brazaville)
Distribution: Congo (Brazaville)
Acarorum Catalogus IX 249
Host: Laccoptera caduca (Coleoptera: Chrysomelidae, Cassidinae)
Genus Gascaricoptes Haitlinger
Gascaricoptes Haitlinger, 2002: 22
Type species: Gascaricoptes analevelonaicus Haitlinger, 2002
Gascaricoptes analevelonaicus Haitlinger
Gascaricoptes analevelonaicus Haitlinger, 2002: 22
Type locality: Analavelona
Distribution: Madagascar
Host: Cassida fuscomacula (Coleoptera: Chrysomelidae, Cassidinae)
Genus Heterocoptes Fain
Heterocoptes Fain, 1967: 94
Type species: Heterocoptes tarsii Fain, 1967
Heterocoptes barbarae Haitlinger
Heterocoptes barbarae Haitlinger, 2009: 334
Type locality: New Britain, Gazelle Peninsula
Distribution: Papua New Guinea (New Britain)
Host: Aspidomorpha aurata (Coleoptera: Chrysomelidae, Cassidinae)
Heterocoptes floresiensis Haitlinger
Heterocoptes floresiensis Haitlinger, 2009: 62
Type locality: Flores
Distribution: Indonesia (Flores)
Host: Aspidimorpha indica (Coleoptera: Chrysomelidae, Cassidinae)
Heterocoptes hannae Haitlinger
Heterocoptes hannae Haitlinger, 1998: 5
Type locality: Shanghai
Distribution: China
Host: Basipronota bisignata (Coleoptera: Chrysomelidae, Cassidinae)
Heterocoptes ingeri Haitlinger
Heterocoptes ingeri Haitlinger, 1999: 67
Type locality: South Luzon
Distribution: Philippines
Host: Laccoptera tredecimguttata (Coleoptera: Chrysomelidae, Cassidinae)
Heterocoptes kamillae Haitlinger
Heterocoptes kamillae Haitlinger, 1998: 8
250
Acarorum Catalogus IX
Type locality: ? Solim-Ho, Yunnan
Distribution: China (Yunnan)
Host: Aspidimorpha miliaris (Coleoptera: Chrysomelidae, Cassidinae)
Heterocoptes lombokensis Haitlinger
Heterocoptes lombokensis Haitlinger, 2009: 326
Type locality: Lombok, Senggigi
Distribution: Indonesia (Lombok, Bali, Java)
Hosts: Aspidimorpha sanctaecrucis, A. miliaris (Coleoptera: Chrysomelidae, Cassidinae)
Heterocoptes lottae Haitlinger
Heterocoptes lottae Haitlinger, 1996: 985; 1999: 72
Type locality: North Vietnam
Distribution: Malaysia, Vietnam
Host: Laccoptera hospita, Aspidomorpha elevata (Coleoptera: Chrysomelidae, Cassidinae)
Heterocoptes marcellae Haitlinger
Heterocoptes marcellae Haitlinger, 2009: 336
Type locality: South Sulawesi
Distribution: Indonesia (Sulawesi)
Host: Aspidimorpha amabilis (Coleoptera: Chrysomelidae, Cassidinae)
Heterocoptes nolae Haitlinger
Heterocoptes nolae Haitlinger, 1996: 983
Type locality: Andaman Is.
Distribution: India (Andaman Is.)
Host: Aspidomorpha inquinata (Coleoptera: Chrysomelidae, Cassidinae)
Heterocoptes rukaensis Haitlinger
Heterocoptes rukaensis Haitlinger, 1999: 70
Type locality: Ruka
Distribution: Papua New Guinea
Host: Aspidimorpha punctum (Coleoptera: Chrysomelidae, Cassidinae)
Heterocoptes ruperti Haitlinger
Heterocoptes ruperti Haitlinger, 2009: 64
Type locality: ? Liangages, Sumatra
Distribution: Indonesia (Sumatra)
Host: Platypria echinogale (Coleoptera: Chrysomelidae, Hispinae)
Heterocoptes samueli Haitlinger
Heterocoptes samueli Haitlinger, 2009: 332
Acarorum Catalogus IX 251
Type locality: “New Territory”, Darwin
Distribution: Australia
Host: Aspidimorpha quadriradiata (Coleoptera: Chrysomelidae, Cassidinae)
Heterocoptes simonettae Haitlinger
Heterocoptes simonettae Haitlinger, 1999: 70
Type locality: Papua New Guinea
Host: Aspidimorpha punctata (Coleoptera: Chrysomelidae, Cassidinae)
Heterocoptes sumatrensis Haitlinger
Heterocoptes sumatrensis Haitlinger, 2009: 329
Type locality: Sumatra, Soekaranda
Distribution: Indonesia (Sumatra)
Host: Basipronota decemmaculata (Coleoptera: Chrysomelidae, Cassidinae)
Heterocoptes tarsii Fain
Heterocoptes tarsii Fain, 1967: 94; Haitlinger, 1996: 985; 2009: 69
Type locality: Sarawak (Malaysia)
Distribution: India (Assam), Indonesia (Bali, Sanur, Lombok, Java, Sumatra, Ceram),
Malaysia (Sarawak, Perak), Philippines
Hosts: Aspidomorpha elevata, A. sanctaecrucis (Coleoptera: Chrysomelidae, Cassidinae);
described from Tarsius sp. (Primates: Tarsiidae)
Genus Honiarrea Haitlinger
Honiarrea Haitlinger, 1990: 343
Type species: Honiarrea lagossa Haitlinger, 1990
Honiarrea lagossa Haitlinger
Honiarrea lagossa Haitlinger, 1990: 343
Type locality: Hainan Is.
Distribution: China (Hainan Is.)
Host: Passandra heros (Coleoptera: Passandridae)
Genus Kamerucoptes Haitlinger
Kamerucoptes Haitlinger, 2002: 8
Type species: Kamerucoptes doubeicus Haitlinger, 2002
Kamerucoptes doubeicus Haitlinger
Kamerucoptes doubeicus Haitlinger, 2002: 8
Type locality: Doube
Distribution: Cameroon
252
Acarorum Catalogus IX
Host: Aspidimorpha obovata (Coleoptera: Chrysomelidae, Cassidinae)
Kamerucoptes lycius Haitlinger
Kamerucoptes lycius Haitlinger, 2002: 13
Type locality: D.R. Congo
Distribution: D.R. Congo
Host: Lycidae gen. sp. (Coleoptera)
Kamerucoptes ruvenzoricus Haitlinger
Kamerucoptes ruvenzoricus Haitlinger, 2002: 13
Type locality: Ruvenzori, Mutwanga
Distribution: Uganda
Host: Aspidimorpha dissentonea (Coleoptera: Chrysomelidae, Cassidinae)
Kamerucoptes sibuticus Haitlinger
Kamerucoptes sibuticus Haitlinger, 2002: 16
Type locality: N’Chari, Fort Sibut
Distribution: Tshad
Host: Laccoptera (Patrisma) murrayi (Coleoptera: Chrysomelidae, Cassidinae)
Kamerucoptes tsaratananaicus Haitlinger
Kamerucoptes tsaratananaicus Haitlinger, 2002: 19
Type locality: Mts Tsaratanana, 2000 m
Distribution: Madagascar
Host: Cassida pretiosa (Coleoptera: Chrysomelidae, Cassidinae)
Genus Nolaecoptes Haitlinger
Nolaecoptes Haitlinger, 1996: 987
Type species: Nolaecoptes vonettae Haitlinger, 1996
Nolaecoptes tanahlotions Haitlinger
Nolaecoptes tanahlotions Haitlinger, 2009: 324
Type locality: Bali, Tanahlot
Distribution: Indonesia (Bali)
Nolaecoptes vonettae Haitlinger
Nolaecoptes vonettae Haitlinger, 1996: 987
Type locality: Sikkim
Distribution: India (Sikkim)
Host: Aspidomorpha sanctaecrucis (Coleoptera: Chrysomelidae, Cassidinae)
Acarorum Catalogus IX 253
Genus Tannuicoptes Haitlinger
Tannuicoptes Haitlinger, 1999: 62
Type species: Tannuicoptes ulfi Haitlinger, 1999
Tannuicoptes jeroni Haitlinger
Tannuicoptes jeroni Haitlinger, 1999: 65
Type locality: Indonesia
Distribution: Indonesia
Host: Corynodes compressicornis (Coleoptera: Chrysomelidae, Eumolpinae)
Tannuicoptes ulfi Haitlinger
Tannuicoptes ulfi Haitlinger, 1999: 62
Type locality: Sikkim
Distribution: India (Sikkim)
Host: Aspidomorpha dorsata (Coleoptera: Chrysomelidae, Cassidinae)
Genus Uvallicoptes Haitlinger
Uvallicoptes Haitlinger, 1999: 62
Type species: Uvallicoptes peeteri Haitlinger, 1999
Uvallicoptes peeteri Haitlinger
Uvallicoptes peeteri Haitlinger, 1999: 62
Type locality: ? Small Gela, Florida Is., Solomon Islands
Distribution: Solomon Islands, Indonesia [Sumatra]
Host: Aspidomorpha adhaerens (Coleoptera: Chrysomelidae, Cassidinae)
Index of Heterocoptidae
(Names in bold are accepted as valid)
Abboticoptes 246
amaliae, Bahrucoptes 247
Amarucoptes 247
analevelonaicus, Gascaricoptes 249
anthophora, Erotylocoptes 248
Bahrucoptes 247
barbarae, Heterocoptes 249
busannae, Amarucoptes 247
Cassiocoptes 247
Conchycoptes 247
doubeicus, Kamerucoptes 251
eddae, Abboticoptes 247
Erotylocoptes 248
floresiensis, Heterocoptes 249
Gascaricoptes 249
hannae, Heterocoptes 249
helenae, Erotylocoptes 248
Heterocoptes 249
Heterocoptidae 246
Honiarrea 251
254
Acarorum Catalogus IX
ingeri, Heterocoptes 249
jeroni, Tannuicoptes 253
Kamerucoptes 251
kamillae, Heterocoptes 249
lagossa, Honiarrea 251
linodesmus, Erotylocoptes 248
lombokensis, Heterocoptes 250
lottae, Heterocoptes 250
luluensis, Erotylocoptes 248
lycius, Kamerucoptes 252
marcellae, Heterocoptes 250
mikki, Cassiocoptes 247
nolae, Heterocoptes 250
Nolaecoptes 252
peeteri, Uvallicoptes 252
rukaensis, Heterocoptes 250
ruperti, Heterocoptes 250
ruvenzoricus, Kamerucoptes 252
samueli, Heterocoptes 250
sibuticus, Kamerucoptes 252
simonettae, Heterocoptes 251
sumatrensis, Heterocoptes 251
tanahlotions, Nolaecoptes 252
tangaicus, Conchycoptes 247
Tannuicoptes 253
taorettae, Erotylocoptes 248
tarsii, Heterocoptes 251
tsaratananaicus, Kamerucoptes 252
ulfi, Tannuicoptes 253
Uvallicoptes 253
verenae, Erotylocoptes 248
vonettae, Nolaecoptes 252
Distribution of Heterocoptidae
AFRICA
Cameroon (Haitlinger, 2002) – Kamerucoptes doubeicus
Congo (Brazzaville) (Haitlinger, 1996) - Erotylocoptes verenae
Congo DR (Fain, 1987; Haitlinger, 2002) – Erotylocoptes anthophora, E. linodesmus, E.
luluensis, Kamerucoptes lycius
Madagascar (Haitlinger, 2002) - Gascaricoptes analevelonaicus, Kamerucoptes tsaratananaicus
Mozambique (Haitlinger, 1996) - Erotylocoptes helenae
South Africa (Haitlinger, 1996) – Erotylocoptes helenae
Tanzania (Haitlinger, 1996, 2002) – Conchycoptes tangaicus, Erotylocoptes taorettae
Tshad (Haitlinger, 2002) - Kamerucoptes sibuticus
Uganda (Haitlinger, 2002) - Kamerucoptes ruvenzoricus
ASIA
China (Haitlinger, 1998; Fan, Chen & Wang, 2010) – Heterocoptes hannae, H. kamillae
Hainan (Haitlinger, 1990) - Honiarrea lagossa
India (Fain, 1967; Haitlinger, 1996 , 1999) – Heterocoptes tarsii, Nolaecoptes vonettae,
Tannuicoptes ulfi
Andaman Is. (Haitlinger, 1996) – Heterocoptes nolae
Indonesia (Haitlinger, 1996, 1999; 2009a) – Cassiocoptes mikki [Sumatra], Heterocoptes
floresiensis [Flores], H. lombokensis [Lombok, Bali, Java], H. marcellae [Sulawesi],
Acarorum Catalogus IX 255
H. ruperti [Sumatra], H. sumatrensis [Sumatra], H. tarsii (Bali, Sanur, Lombok, Sumatra, Java, Ceram), Nolaecoptes tanahloticus [Bali], Uvallicoptes peeteri [Sumatra],
Tannuicoptes jeroni
Malaysia (Fain, 1967; Haitlinger, 1999) – Bahrucoptes amaliae, Heterocoptes tarsii
Sarawak and Sabah (Fain, 1967) – Heterocoptes tarsii
Philippines (Haitlinger, 1996, 1999) - Abboticoptes eddae, Heterocoptes ingeri, H. tarsi,
Vietnam (Haitlinger, 1996) – Heterocoptes lottae
NORTH AMERICA
Canada (Beaulieu et al., 2019) – one unidentified sp.
SOUTH and CENTRAL AMERICA
Peru (Haitlinger, 2001) – Amarucoptes busannae
OCEANIA
Australia (Haitlinger, 2009a) – Heterocoptes samueli
Papua New Guinea (Haitlinger, 1999, 2009) - Heterocoptes simonettae, H. rukaensis
New Britain (Haitlinger, 2009a) – Heterocoptes barbarae
New Ireland - Uvallicoptes peeteri
Solomon Islands (Haitlinger, 1999) - Uvallicoptes peeteri
Hosts of Heterocoptidae
Insecta
Coleoptera
Fam. Chrysomelidae
Subfam. Hispinae
Conchyloctenia hybrida - Conchycoptes tangaicus
Lasiochila sp. – Cassiocoptes mikki
Platypria echinogale - Heterocoptes ruperti
Subfam. Cassidinae
Aspidimorpha adhaerens – Uvallicoptes peeteri
Aspidimorpha amabilis – Heterocoptes marcellae
Aspidimorpha dissentonea - Kamerucoptes ruvenzoricus
Aspidimorpha dorsata – Tannuicoptes ulfi
Aspidimorpha elevata – Bahrucoptes amaliae, Heterocoptes tarsii
Aspidimorpha fuscopunctata - Uvallicoptes peeteri
Aspidimorpha inquinata – Heterocoptes nolae
Aspidimorpha miliaris – Heterocoptes kamillae, H. lombokensis
Aspidimorpha obovata - Kamerucoptes doubeicus
Aspidimorpha punctum – Heterocoptes simonettae
256
Acarorum Catalogus IX
Aspidimorpha quadrilobata – Abboticoptes edda
Aspidimorpha quadriradiata – Heterocoptes samueli
Aspidimorpha sanctaecrucis – Heterocoptes lombokensis, H. tarsii, Nolaecoptes tanahloticus, N. vonettae
Aspidimorpha socia – Heterocoptes barbarae
Basipronota bisignata - Heterocoptes hannae
Basipronota decemmaculata - Heterocoptes sumatrensis
Cassida fuscomacula - Gascaricoptes analevelonaicus
Cassida pretiosa - Kamerucoptes tsaratananaicus
Laccoptera aurosa - Erotylocoptes taorettae
Laccoptera caduca – Erotylocoptes verenae
Laccoptera hospita – Heterocoptes lottae
Laccoptera (Patrisma) murrayi - Kamerucoptes sibuticus
Laccoptera rugosicollis - Erotylocoptes helenae
Laccoptera tredecimguttata – Heterocoptes ingeri
Subfam. Eumolpinae
Corynodes compressicornis - Tannuicoptes jeroni
Fam. Erotylidae
Linodesmus coccus apicalis – Erotylocoptes linodesmus
Fam. Lycidae gen. sp. - Kamerucoptes lycius
Fam. Passandridae
Passandra heros – Honiarrea lagossa
Fam. Passalidae gen.sp. - Amarucoptes busannae
Hymenoptera
Fam. Apidae
Anthophora acraensis - Erotylocoptes anthophora
Fam. Sphecidae
Chlorion (Isodontia) simoni – Erotylocoptes luluensis
Mammalia
Primates
Fam. Tarsiidae
Tarsius sp. - Heterocoptes tarsii
Bibliography of Heterocoptidae
Beaulieu et al. 2019. Acari of Canada. – ZooKeys, 819: 77–168.
Fain A. 1967. Un Acarien rémarquable recolté sur un Tarsier (Heterocoptidae f. n.: Sarcoptiformes). – Zool. Anzeiger, 178 (1/2): 90–94.
Fain A. 1987. Nouveau taxa dans la famille Heterocoptidae Fain, 1967 (Acari, Astigmata). –
Bull. Annls Soc. r. belge Ent., 123: 243–253.
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Fan Qing-Hai, Yan Chen, Zi-Qing Wang. 2010. Acaridia (Acari: Astigmatina) of China: A
review of research progress. – p.225–259. In: Zhang Z.-Q., Hong X.-Y. & Fan Q.-H. (eds)
Xin Jie-Lin Centenary: Progress in Chinese Acarology. Zoosystema 4, 1–345.
Haitlinger R. 1990. Honiarrea gen. nov., a new genus of the family Heterocoptidae (Acari,
Astigmata). – Annales Zoologici, Warszawa, 43(16): 343–345.
Haitlinger R. 1996. New heterocoptid mites (Acari, Astigmata, Heterocoptidae) associated with
Cassidinae and Hispinae (Coleoptera, Chrysomelidae) from Africa and Asia. – Linzer
Biol. Beitr., 28: 979–998.
Haitlinger R. 1998. Four new species of canestriniid and heterocoptid mites from China (Acari,
Astigmata, Canestriniidae, Heterocoptidae). – Spixiana, 21: 1–10.
Haitlinger R. 1999. Six new heterocoptid mites (Acari, Astigmata, Heterocoptidae) associated
with Cassidinae and Eumolpinae (Coleoptera, Chrysomelidae) from Indo-Australian
Region. – Misc. Zool., 22: 59–73.
Haitlinger R. 2001. Amarucoptes, a new genus of the family Heterocoptidae (Acari, Astigmata).
– Syst. Appl. Acarol., 6: 111–114.
Haitlinger R. 2002. Seven new Heterocoptid mites (Acari: Astigmata: Heterocoptidae) from
Africa. – Zeszyty naukowe Akademii Rol. we Wroclawiu, Zootechnika XLIX, 447: 7–29.
Haitlinger R. 2009a. New species of mites (Acari: Astigmata: Heterocoptidae) from IndoAustralian region. – Zesz. Nauk. UP Wroc., Biol. Hod. Zwierz., 58(572): 61–72.
Haitlinger R. 2009b. Nine new species and new records of mites (Acari: Astigmata: Heterocoptidae) from tropical Africa. – Revista Ibérica de Aracnología, 17: 97–115.
Haitlinger R. 2009c. New species of mites (Acari: Astigmata: Heterocoptidae) from the
Australian and Oriental regions, with a description of the female of Uvalicoptes peeteri
Haitlinger, 1999. – Biologia, 64(2): 324–339.
OConnor B.M. 1979. A review of the family Heterocoptidae. – Proc. V Int. Congr. Acar.
(August 6–12, 1978, East Lansing). Recent Advances of Acarology II: 429–433.
The Mite Superfamily Hemisarcoptoidea
According to O’Connor (2009), the superfamily contains the families Chaetodactylidae,
Hyadesiidae, Carpoglyphidae, Algophagidae, Hemisarcoptidae and Winterschmidtiidae, or
in total 51 genera and ca. 345 species. Some of them (Chaetodactylidae) live on Hymenoptera, some (Hyadesiidae, Algophagidae) prefer the sea shore and the saline habitats. Hemisarcoptidae are found to live as parasites on Insects (Coccodea, Coleoptera – Chrysomelidae, Coccinellidae ) and some of them are used for the pest control. Carpoglyphidae and
Winterschmidtiidae are inhabitants of rotten materials, sometimes found also on insects.
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The Mite Family Chaetodactylidae
Glyciphagidae (Chaetodactylinae) Zachvatkin, 1941: 347
Chaetodactylidae Zachvatkin: Baker, 1962: 229 (part.); Klimov & OConnor, 2008: 98
Obligate associates of long-tongued bees (OConnor & Klimov, 2009). From the five known
genera in the family the first (Chaetodactylus Rondani) has been described as early as in
1866, but remained the only one until Oudemans (1905) described (based on Pediculus
cerambycinus Scopoli, 1763) the large genus Sennertia, revised much later by Fain (from
1966 to 2001). The thorough study of the family starts with the results of B. OConnor
and P. Klimov, who published generic revision of the group (OConnor, 1993) and very
detailed monograph (Klimov & OConnor, 2008), containing the history of the research
on Chaetodactylidae until this time. As early as in 1903 Tschitscherin separated these
mites in a group Chaetodactylini. Zachvatkin (1941) considered Chaetodactylinae as
subfamily of Glycyphagidae and Baker (1962) raised them in a family rank. Meanwhile
some other authors contributed to the taxonomy and/or the biology of Chaetodactylidae
(Womersley, 1941, Vitzthum, 1941, Elbadry, 1971, Delfinado & Baker, 1976, Sherbef &
El-Duweini, 1980, Baker & Delfinado-Baker, 1983, Haitlinger, 1999, 2000, Kurosa, 1987,
2003, Ramaraju & Mohanasundaram, 2001, and others).
Composition of Chaetodactylidae
Fam. Chaetodactylidae Zachvatkin, 1941–5 genera, 109 sp.
Achaetodactylus Fain, 1981–3 sp.
Centriacarus Klimov et OConnor, 2007–2 sp.
Chaetodactylus Rondani, 1866–26 sp.
Roubikia OConnor, 1993–4 sp.
Sennertia Oudemans, 1905–72 sp.
Catalogue of Chaetodactylidae
Fam. Chaetodactylidae Zachvatkin
Glycyphagidae (Chaetodactylinae) Zachvatkin, 1941: 347
Chaetodactylidae Zachvatkin; Baker, 1962: 229 (par.); Klimov & OConnor, 2008 : 98
Type genus: Chaetodactylus Rondani, 1866
Genus Achaetodactylus Fain
Achaetodactylus Fain, 1981a: 2 (as subgenus)
Acarorum Catalogus IX 259
Achaetodactylus Fain: Klimov et al., 2007a: 1370 (as genus)
Chaetodactylus (Ochaetodactylus) Fain, 1981b: 2. Type species Chaetodactylus
decellei Fain, 1974
Type species: Chaetodactylus leleupi Fain, 1974
Achaetodactylus ceratinae (Fain)
Chaetodactylus ceratinae Fain, 1974a: 214
Chaetodactylus (Achaetodactylus) ceratinae Fain, 1981a: 4
Achaetodactylus ceratinae (Fain): Klimov et al., 2007a: 1371; Klimov & OConnor, 2008: 215
Type locality: Nairobi
Distribution: Kenya, South Africa, Tanzania
Hosts: Ceratina apaca, Ceratina (Hirashima) lativentris, C. (H.) nigriceps (Hymenoptera:
Apidae, Xylocopinae)
Achaetodactylus decellei (Fain)
Chaetodactylus decellei Fain, 1974: 215
Chaetodactylus (Ochaetodactylus) decellei: Fain, 1981a: 8
Ochaetodactylus decellei: Klimov et al., 2007a: 1371
Achaetodactylus decellei (Fain): Klimov & OConnor, 2008: 216
Type locality: Ngurdoto Nat. Park, Tanzania
Distribution: Cameroon, DR Congo, Tanzania
Hosts: Ceratina aereola, C. excavata, C. spilota (Hymenoptera: Apidae, Xylocopinae)
Achaetodactylus leleupi (Fain)
Chaetodactylus leleupi Fain, 1974: 214
Chaetodactylus (Achaetodactylus) leleupi: Fain, 1981a: 4; Fain & Pauly, 2001: 131
Achaetodactylus leleupi (Fain): Klimov et al., 2007a: 1371; Klimov & OConnor, 2008: 215
Type locality: Karen, Nairobi (Kenya)
Distribution: Cameroon, DR Congo, Kenya, Tanzania
Hosts: Ceratina (Hirashima) lativentris, Ceratina diloloensis, C. ruwenzorica, C. spilota
(Hymenoptera: Apidae, Xylocopinae)
Genus Centriacarus Klimov et OConnor
Centriacarus Klimov & OConnor, 2007: 816
Type species: Centriacarus turbator Klimov et OConnor, 2007
Centriacarus guahibo Klimov et OConnor
Centriacarus guahibo Klimov & OConnor, 2007: 817; Klimov et al., 2007a: 1371; 2008: 216
Type locality: Territorio Amazonas
Distribution: Venezuela
Host: Centris sp. (Hymenoptera: Apidae, Apinae)
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Centriacarus turbator Klimov et OConnor
Centriacarus turbator Klimov & OConnor, 2007: 816; Klimov et al., 2007a: 1371; Klimov
& OConnor, 2008: 216
Type locality: Mato Grosso de Sul, Aquidauana (Brazil)
Distribution: Brazil, Colombia, Mexico, Panama
Host: Centris (Heterocentris) vittata (Hymenoptera: Apidae, Apinae)
Genus Chaetodactylus Rondani
Chaetodactylus Rondani, 1866: 19
Type species: Trichodactylus osmiae Dujardin, 1839
Subgenus Chaetodactylus Rondani
Chaetodactylus Rondani, 1866: 19
Type species: Trichodactylus osmiae Dujardin, 1839
Chaetodactylus abditus Klimov et OConnor
Chaetodactylus abditus Klimov & OConnor, 2004: 175; 2008: 124
Type locality: Arizona, Pima Co., Continental
Distribution: Mexico, USA (Arizona)
Hosts: Lithurgus (Lithurgopsis) echinocacti, L. (Lithurgopsis) planifrons (Hymenoptera:
Megachilidae)
Chaetodactylus anthidii (Oudemans)
Trichotarsus anthidii Oudemans, 1911a: 165; Vitzthum, 1919: 31
Chaetodactylus anthidii: Oudemans, 1924: 328; Zachvatkin, 1941: 398; Klimov & OConnor, 2008: 110
Chaetodactylus (Chaetodactylus) anthidii: Fain, 1981a: 2; OConnor, 1993a: 62
Type locality: Tunisia
Distribution: Tunisia
Host: Rhodanthidium sticticum (as Anthidium sticticum)(Hymenoptera: Megachilidae)
Chaetodactylus antillarum Klimov et OConnor
Chaetodactylus antillarum Klimov & OConnor, 2008: 113
Type locality: Jamaica, Hellshire Hills
Distribution: Dominican Republic, Jamaika
Host: Lithurgus antilleorum antilleorum (Hymenoptera: Megachilidae)
Chaetodactylus azteca Klimov et OConnor
Chaetodactylus azteca Klimov & OConnor, 2008: 127
Type locality: Hidalgo: Tepeapulco
Distribution: Mexico
Acarorum Catalogus IX 261
Host: Osmia (Diceratosmia) azteca (Hymenoptera: Megachilidae)
Chaetodactylus birulai Zachvatkin
Chaetodactylus birulai Zachvatkin, 1941: 396; Samšinák, 1973: 404; Fain, 1974a: 213;
Klimov & OConnor, 2008: 216
Chaetodactylus (Chaetodactylus) birulai Zachvatkin: Fain, 1981a: 3; Kurosa, 1987: 373
Chaetodactylus poetae Samšinák, 1973: 401; Fain, 1981a: 2. Type: Czech Republic: Sobotka;
syn. by Klimov & OConnor, 2008: 109
Chaetodactylus (Chaetodactylus) poetae: Kurosa, 1987: 373
Type locality: Kirovskaya Oblast’ (Russia)
Distribution: Czech Rep., Holland, Russia
Host: Chelostoma florisomne (Hymenoptera: Megachilidae), also found phoretic on cleptoparasites: Sapyga quinquepunctata, Sapyga clavicornis (Hymenoptera: Sapygidae)
Chaetodactylus chrysidis Fain et Baugnée
Chaetodactylus chrysidis Fain & Baugnée, 1996: 23; Van Asselt, 2000: 225; Klimov &
OConnor, 2008: 110
Chaetodactylus chrysidis aurulenticola Fain & Baugnée,1996: 28. Type: Belgium, TreignesRivelottes, ex Osmia aurulenta; syn. by Klimov & OConnor, 2008: 110; Van Asselt,
2000: 225
Type locality: Treignes-Saumières
Distribution: Belgium
Host: Chrysura trimaculata (Hymenoptera: Chrysididae), Osmia aurulenta and O. bicolor
(Hymenoptera: Megachilidae)
Chaetodactylus claudus Klimov et OConnor
Chaetodactylus claudus Klimov & OConnor, 2008: 133
Type locality: Utah, Cache Co., Green Canyon
Distribution: USA (Utah, Texas)
Host: Osmia (O.) rubiflorus (Hymenoptera: Megachilidae)
Chaetodactylus claviger Oudemans
Chaetodactylus claviger Oudemans, 1924: 328; Zachvatkin, 1941: 396; Klimov & OConnor, 2008: 111
Chaetodactylus (Spinodactylus) claviger: Fain, 1981a: 2 (part., excluding Chaetodactylus
krombeini)
Chaetodactylus osmiae: Abou Senna, 1997: 667 (misidentification, Egypt, from Apis
melifera)
Type locality: Promontorio del Gargano (“Monte Gargano, Apulië”), Italy
Distribution: Egypt, France, Italy
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Hosts: Osmia tricornis (Hymenoptera: Megachilidae), Apis melifera (Hymenoptera:
Apidae)
Chaetodactylus dalyi Fain
Chaetodactylus dalyi Fain, 1974: 213; Klimov & OConnor, 2007: 826; 2008: 213
Chaetodactylus (Chaetodactylus) dalyi: Fain, 1981a: 6
Type locality: Natal
Distribution: Mozambique, South Africa, Zimbabwe
Host: Ceratina (Pithitis) turneri (Hymenoptera: Apidae, Xylocopinae)
Chaetodactylus dementjevi Zachvatkin
Chaetodactylus dementjevi Zachvatkin, 1941: 399; Klimov & OConnor, 2008: 103
Chaetodactylus (Achaetodactylus) dementjevi: Fain, 1981a: 2
Type locality: Uzbekistan: Toshkent (Gora Aktash)
Distribution: Kazakhstan, Russia, Uzbekistan
Hosts: Megachile bombycina, M. ligniseca (Hymenoptera: Megachilidae), also on Vespula
germanica (Vespidae)
Chaetodactylus furunculus Klimov et OConnor
Chaetodactylus furunculus Klimov & OConnor, 2008: 116
Type locality: California, San Bernardino Co., Cedar Canyon
Distribution: USA (California)
Host: Lithurgus listrotus (Hymenoptera: Megachilidae)
Chaetodactylus gibbosi Klimov et OConnor
Chaetodactylus gibbosi Klimov & OConnor, 2004: 178; 2008: 124
Type locality: Florida, Liberty Co.
Distribution: USA (Florida)
Host: Lithurgus gibbosus (Hymenoptera: Megachilidae)
Chaetodactylus hirashimai Kurosa
Chaetodactylus (Chaetodactylus) hirashimai Kurosa, 1987: 374
Saproglyphus sp.: Hirashima, 1957: 200 (Host: Osmia excavata)
Chaetodactylus sp.: Krombein, 1962: 239; Host: Osmia excavata, Japan
Chaetodactylus hirashimai: Qu et al., 2002: 121; Klimov & OConnor, 2008: 110
Type locality: Japan: Honshu, Nara Pref., Nara-shi, Furuichi-chô
Distribution: Japan (Honshu, Kyushu)
Hosts: Osmia (Osmia) excavata, O. (O.) cornifrons, O. (O.) pedicornis, Osmia (Helicosmia)
imaii
(Hymenoptera: Megachilidae)
Acarorum Catalogus IX 263
Chaetodactylus hopliti Klimov et OConnor
Chaetodactylus hopliti Klimov & OConnor, 2008: 137
Type locality: New York, Onondaga Co., Syracuse
Distribution: Canada, USA
Hosts: Hoplitis (Alcidamea) producta (type host); Hoplitis (Alcidamea) pilosifrons; Hoplitis
(Monumetha) spoliata; Megachile (Megachile) relativa; Osmia (Melanosmia) brevis;
Osmia (Centrosmia) bucephala; Osmia (Chenosmia) inermis (Megachilidae); Isodontia
mexicana (Hymenoptera: Sphecidae)
Chaetodactylus kouboy Klimov et OConnor
Chaetodactylus kouboy Klimov & OConnor, 2008: 121
Type locality: New Mexico, Colfax Co., Cimarron Canyon
Distribution: USA (New Mexico)
Host: Lithurgus (Lithurgopsis) apicalis (Hymenoptera: Megachilidae)
Chaetodactylus krombeini Baker
Chaetodactylus krombeini Baker, 1962: 229; Klimov & OConnor, 2008: 141; McKinney
& Park, 2012:
Chaetodactylus (Spinodactylus) krombeini Baker: Fain, 1981 : 2; OConnor, 1993a: 362
Type locality: Maryland
Distribution: USA
Hosts: Osmia cornifrons, O. rubifloris, O. r. biedermanni, O. bucephala, O. chlybaea, O. lignaria propinqua, O. montana, O. californica, O. grindeliae (Hymenoptera: Magachilidae)
Chaetodactylus lassulus Klimov et OConnor
Chaetodactylus lassulus Klimov & OConnor, 2007: 824; 2008: 111
Type locality: Coquimbo
Distribution: Chile
Hosts: Trichothurgus dubius, T. herbsti (Hymenoptera: Magachilidae)
Chaetodactylus lithurgi Klimov et OConnor
Chaetodactylus lithurgi Klimov & OConnor, 2004: 173; 2008: 121
Type locality: New Mexico, Colfax Co., Cimarron Canyon
Distribution: USA (New Mexico, Idaho, Texas, Arizona)
Hosts: Lithurgus apicalis, L. gibbosus, L. littoralis (Hymenoptera: Megachilidae)
Chaetodactylus ludwigi (Trouessart)
Trichotarsus Ludwigi Trouessart, 1904a: 234; 1904b: 365; Ludwig, 1904: 216; Vitzthum,
1912d: 292; 1919; Knülle, 1959: 385
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Chaetodactylus ludwigi (Trouessart): Zachvatkin, 1941: 42; Baker, 1962a: 229; 1987: 65;
OConnor, 1993a: 353; Pauly & Munzinger, 2003: 160; Klimov & OConnor, 2007: 826;
Klimov et al., 2007a: 1371; Klimov & OConnor, 2008: 111
Chaetodactylus (Chaetodactylus) ludwigi: Fain & Pauly, 2001: 128
Type locality: Carolines Islands: Ponapé
Distribution: Federated States of Micronesia: Pohnpei Is. (“Carolines Islands: Ponapé”);
New Caledonia; French Polynesia: Moorea Is. Nr. Taiti; India, Indonesia (Java),
Madagascar
Hosts: Lithurgus (Lithurgus) atratus (as Megachile lonalap), nest; Lithurgus scabrosus
(Hymenoptera: Megachilidae)
Chaetodactylus melitomae Klimov et OConnor
Chaetodactylus melitomae Klimov & OConnor, 2007: 821; 2008: 113
Type locality: Mexico: Tabasco, Cardenas
Distribution: Honduras, Mexico
Host: Melitoma sp., Melitoma segmentaria (Hymenoptera: Apidae, Apinae)
Chaetodactylus micheneri Klimov et OConnor
Chaetodactylus micheneri Klimov & OConnor, 2008: 127
Type locality: California, Fresno Co., Mt. Crocker, 11800 ft.
Distribution: USA (California)
Host: Osmia subaustralis (Hymenoptera: Megachilidae)
Chaetodactylus nipponicus Kurosa
Chaetodactylus sp.: Yamada et al., 1971: 32
Chaetodactylus (Chaetodactylus) nipponicus Kurosa, 1987: 377
Chaetodactylus nipponicus: Van Asselt, 2000: 221; Bosh & Kemp, 2001: 62; Qu et al., 2002:
121; Qu et al., 2003: 55; Klimov & OConnor, 2008: 110
Type locality: Tsugaru-gun (Tsushima Is.)
Distribution: Japan (Tsushima Is., Honshu)
Hosts: Osmia (Osmia) excavata, O. (O.) cornifrons, O. (O.) taurus, O. (O.) pedicornis
(Hymenoptera: Megachilidae)
Chaetodactylus osmiae (Dufour)
Trichodactylus osmiae Dufour, 1839: 276; Gervais, 1844: 266 (part.); Murray, 1877: 252
(part.); Mégnin, 1880: 147; Canestrini, 1888a: 395; 1888b: 23
Chaetodactylus osmiae: Rondani, 1866: 183; Zachvatkin, 1941: 394 (part.); Turk, 1953:
82; Türk & Türk, 1957: 207; Lith, 1957: 197; Baker, 1962a: 229; 1987: 65; Krombein,
1962: 238; Fain, 1966: 249; Elbadry, 1971: 88; De Jong et al., 1982: 245 (Apis); Bosch,
1992: 77; Chmielewski, 1993: 133; Fain & Baugnée, 1996: 23; Van Asselt, 2000: 221;
Acarorum Catalogus IX 265
Fain & Pauly, 2001: 127; Qu et al., 2003: 59; Krunic´ et al., 2005: 143; Klimov et al.,
2007a: 1371; Klimov & OConnor, 2008: 110
Trichotarsus osmiae: Berlese, 1897: 105; Canestrini & Kramer, 1899: 149; Giard, 1900: 377;
Oudemans, 1900: 117; 1901: 81; 1903a: 147; 1905a: 22; Oudemans, 1905b: 80; 1911a: 165;
Michael, 1903: 17; Ludwig, 1904: 216; Trouessart, 1904a: 235; 1904b: 365; Vitzthum,
1912d: 291; 1919: 38; Popovici-Baznosanu, 1913: 32; Knülle, 1959: 385
Chaetodactylus mahunkai Samsinák, 1973: 404. Type: Hungary; synonymized by Fain,
1981b.
Chaetodactylus (Chaetodactylus) osmiae: Fain, 1981b: 4; Kurosa, 1987: 373; OConnor,
1993a:353
Misidentifications (acc. to Klimov & OConnor, 2008): Donnadieu, 1868 (Chaetodactylus
claviger), Banks, 1902: 176 (Chaetodactylus krombeini or Ch. rozeni), Abou Senna,
1997 (Chaetodactylus claviger), Fain et al., 1992 (Chaetodactylus chrysidis aurulenticola)
Type locality: France, Département des Landes (Gervais)
Distribution: Belgium, Croatia, England, France, Georgia, Germany, Holland, Hungary,
Italy, Luxemburg, Poland, Romania, Russia
Hosts: Osmia bicornis, O. cornuta, O. fronticornis, O. kohlii, O. niveata, O. rufa, O. rufiventris, O. tricornis (Hymenoptera: Megachilidae); Andrena spp., Megachile sp. (Berlese);
Anthophora sp.
Chaetodactylus reaumuri (Oudemans)
Trichotarsus reaumuri Oudemans, 1905b: 81; 1905a: 22; 1911a: 165; Vitzthum, 1919: 31
Chaetodactylus reaumuri (Oudemans): Oudemans, 1924: 328; Vitzthum, 1929: 77;
Zachvatkin, 1941: 396; Türk & Türk, 1957: 210; Fain & Baugnée, 1996: 28; Klimov
& OConnor, 2008: 110
Chaetodactylus (Chaetodactylus) reaumuri: Fain, 1981b: 2; Kurosa, 1987: 373 (reamuri);
OConnor, 1993a: 362 (reaumerii)
Type locality: Kolin (Czech Rep.)
Distribution: Czech Rep., Georgia, Germany, Greece (Corfu), ? Poland (“Odran, Oostenrijksch Silezië”); Crimea.
Hosts: Osmia rufiventris, O. brevicornis, O. leucogastra (as Osmia panzeri)(Hymenoptera:
Megachilidae)
Chaetodactylus rozeni Klimov et OConnor
Chaetodactylus rozeni Klimov & OConnor, 2008: 132
Type locality: North Carolina, Swain Co., Bryson City
Distribution: USA (North Carolina, Texas, Tennessee)
Hosts: Osmia georgica (Hymenoptera: Megachilidae); Chrysis coerulans (Hymenoptera:
Chrysididae)
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Chaetodactylus zachvatkini Klimov et OConnor
Chaetodactylus zachvatkini Klimov & OConnor, 2008: 110 (nom. nov. pro Chaetodactylus
osmiae sensu Zachvatkin, 1941(part., non Dufour, 1839)
Type locality: Italy
Distribution: Italy, Spain, Tunisia
Hosts: Osmia cornuta, O. latreillei, O. niveata, O. tricornis (Hymenoptera: Megachilidae)
Genus Roubikia OConnor
Roubikia OConnor, 1993: 347
Type species: Chaetodactylus panamensis Baker, Roubik et Delfinado-Baker, 1987
Roubikia imberba Klimov et OConnor
Roubikia imberba Klimov & OConnor, 2007: 819; 2008: 221
Type locality: Tucumán, Salta
Distribution: Argentina
Host: Tetrapedia sp. (Hymenoptera: Apidae, Apinae) and its cleptoparasites Coelioxoides
waltheriae and C. exulans
Roubikia latebrosa Klimov et OConnor
Roubikia latebrosa Klimov & OConnor in Klimov et al., 2007b: 118; Klimov et al., 2007a:
1371; Klimov & OConnor, 2008: 103
Type locality: Loreto
Distribution: Peru
Host:Tetrapedia sp. (Hymenoptera: Apidae, Apinae)
Roubikia officiosa Klimov et OConnor
Roubikia officiosa Klimov & OConnor, 2007: 821; 2008: 107
Type locality: Jalisco
Distribution: Mexico
Host: Tetrapedia maura (Hymenoptera: Apidae, Apinae)
Roubikia panamensis (Baker, Roubik et Delfinado-Baker)
Chaetodactylus panamensis Baker, Roubik & Delfinado-Baker, 1987: 67; Roubik, 1987: 75
Roubikia panamensis (Baker, Roubik et Delfinado-Baker): OConnor, 1993 : 345; Van
Asselt, 2000: 225; Klimov & OConnor, 2007: 819; Klimov et al., 2007a: 1371; 2007b:
116; Klimov & OConnor, 2008: 103
Type locality: Panama, Curundú
Distribution: Bolivia, Brazil, French Guiana, Mexico, Panama
Hosts: Tetrapedia diversipes, T. peckoltii (Hymenoptera: Apidae, Apinae), also phoretic on
Coelioxoides waltheriae (cleptoparasite of Tetrapedia diversipes)
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Genus Sennertia Oudemans
Sennertia Oudemans, 1905: 21
Type species: Pediculus cerambycinus Scopoli, 1763
Subgenus Afrosennertia Fain
Afrosennertia Fain, 1981b: 147
Asiosennertia Fain, 1981b: 147, type species: Sennertia (Afrosennertia) delfinadoae Fain,
1981,synonymized by Klimov & OConnor, 2008: 147
Type species: Sennertia monicae Fain, 1971
Sennertia (Afrosennertia) basilewskyi Fain
Sennertia basilewskyi Fain, 1974: 215; Klimov & OConnor, 2008: 223
Sennertia (Afrosennertia) basilewskyi: Fain, 1981b: 180; Fain, 1982: 67
Type locality: Bambesa, Haut-Congo Prov.
Distribution: DR Congo
Hosts: Hymenoptera (Megachilidae); Chedrion nigrihirtum, Mesotrichia africana
Sennertia (Afrosennertia) delfinadoae Fain
Sennertia (Asiosennertia) delfinadoae Fain, 1981b: 172
Sennertia delfinadoae Fain: Klimov & OConnor, 2008: 227
Sennertia bakeri Ramaraju & Mohanasundaram, 2001: 109 (Type: Tamil Nadu Coimbatore;
Klimov & OConnor, 2008: 227)
Type locality: India
Distribution: India
Host: Xylocopa (Nodula) amethystina (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Afrosennertia) jeanalexi Fain
Sennertia jeanalexi Fain, 1971: 269; Fain, 1974a: 216; Klimov & OConnor, 2008: 230
Sennertia (Afrosennertia) jeanalexi: Fain, 1981b: 178; Fain, 1982: 67; OConnor, 1993a: 362
Type locality: Bubulu river, near Mvungu, Mayumbe
Distribution: DR Congo
Hosts: Xylocopa (Koptortosoma) nigrita, X. (K.) lepeletieri (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Afrosennertia) monicae Fain
Sennertia monicae Fain, 1971: 269; 1974a: 215; Klimov & OConnor, 2008: 233
Sennertia (Afrosennertia) monicae: Fain, 1981b: 178; 1982: 67
Type locality: Kivu, Uvira
Distribution: DR Congo
Host: Xylocopa (Xenoxylocopa) inconstans (as Mesotrichia inconstans) (Hymenoptera:
Apidae, Xylocopinae)
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Sennertia (Afrosennertia) oudemansi Zachvatkin
Sennertia oudemansi Zachvatkin, 1941: 390; Turk, 1948: 84; Klimov & OConnor, 2008: 233
Sennertia (Asiosennertia) oudemansi: Fain, 1981b: 176
Type locality: Japan
Distribution: Japan
Host: Xylocopa (Biluna) nasalis (as Xylocopa dissimilis) (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Afrosennertia) queenslandica Womersley
Sennertia queenslandica Womersley, 1941: 479; Klimov & OConnor, 2008: 234
Sennertia (Asiosennertia) queenslandica: Fain, 1981b: 176
Sennertia (Afrosennertia) queenslandica: Fain, 1982: 68
Type locality: Queensland, «Moa Is., Torres Straits”
Distribution: Australia
Host: Xylocopa (Koptortosoma) bryorum (as Mesotrichia bryorum) (Hymenoptera: Apidae,
Xylocopinae)
Sennertia (Afrosennertia) vitzthumi Fain
Sennertia (Asiosennertia) vitzthumi Fain, 1981b: 172
Sennertia vitzthumi Fain: Klimov & OConnor, 2008: 237
Type locality: Unknown
Distribution: Unknown (probably Oriental)
Host: Unknown
Subgenus Amsennertia Fain
Amsennertia Fain, 1981a: 147
Type species: Sennertia frontalis Vitzthum, 1941
Sennertia (Amsennertia) americana Delfinado et Baker
Sennertia americana Delfinado & Baker, 1976: 84; OConnor, 1988: 341; 1993b: 164; Ramaraju & Mohanasundaram, 2001: 107; Klimov & OConnor, 2008: 173
Sennertia (Amsennertia) americana (Delfinado et Baker): Fain, 1981b: 147; Alzuet &
Abrahamovich, 1987: 350; Lombert et al., 1987: 113; OConnor, 1993a: 362
Type locality: New York (Albany Co., Albany)
Distribution: USA
Host: Xylocopa (Xylocopoides) virginica, X. (Xylocopoides) virginica texana (Hymenoptera:
Apidae, Xylocopinae)
Sennertia (Amsennertia) faini Baker et Delfinado-Baker
Sennertia faini Baker & Delfinado-Baker, 1983: 117; Alzuet & Abrahamovich, 1987: 350;
Ramaraju & Mohanasundaram, 2001: 107 (Senertia); Klimov & OConnor, 2008: 170
Acarorum Catalogus IX 269
Type locality: Sololá, San Lucas Tolimán
Distribution: Guatemala
Host: Apis mellifera (Hymenoptera: Apidae, Apinae)
Sennertia (Amsennertia) frontalis Vitzthum
Sennertia frontalis Vitzthum, 1941: 308; Baker & Delfinado-Baker, 1983: 119; Ramaraju
& Mohanasundaram, 2001: 109; Klimov & OConnor, 2008: 179
Sennertia (Amsennertia) frontalis Vitzthum : Fain, 1981b: 176; Alzuet & Abrahamovich,
1987: 346; 1990: 630; OConnor, 1993a: 362
Type locality: Argentina, Entre Rios
Distribution: Argentina, Brazil, Honduras, Mexico, Panama, Peru, Suriname, Venezuela
Hosts: Xylocopa (Neoxylocopa) frontalis, X. (N.) nautlana (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Amsennertia) ignota Delfinado et Baker
Sennertia ignota Delfinado & Baker, 1976: 85; Baker & Delfinado-Baker, 1983: 119; Klimov
& OConnor, 2008: 211
Senertia ignota [sic !]: Ramaraju & Mohanasundaram, 2001: 107
Sennertia (Amsennertia) ignota: Fain, 1981a: 178; Alzuet & Abrahamovich, 1987: 346
Type locality: Peru: Piura, Talara
Distribution: Peru
Host: Xylocopa sp. (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Amsennertia) longipilis Alzuet et Abrahamovich
Sennertia (Amsennertia) longipilis Alzuet & Abrahamovich, 1987: 346
Sennertia longipilis Alzuet & Abrahamovich, 1989: 115; Klimov & OConnor, 2008: 232;
Montalva & Allendes, 2010: 55
Type locality: Misiones Prov. (Argentina)
Distribution: Argentina, Chile
Host: Xylocopa (Schonnherria) splendidula splendidula (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Amsennertia) splendidulae Alzuet et Abrahamovich
Sennertia (Amsennertia) brevipilis Alzuet & Abrahamovich, 1987: 345 (praeoccup. from
Sennertia vanderhammeni brevipilis Fain,1974)
Sennertia splendidulae Alzuet & Abrahamovich, 1989: 236; 1990: 630
Sennertia (Amsennertia) splendidulae Alzuet & Abrahamovich, 1991: 1; OConnor, 1993a.
362
Type locality: La Plata, Buenos Aires Prov.
Distribution: Argentina
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Acarorum Catalogus IX
Host: Xylocopa (Schonnherria) splendidula splendidula (Hymenoptera: Apidae, Xylocopinae)
Subgenus Sennertia Oudemans
Sennertia Oudemans, 1905: 21 (Fain, 1981b: 146)
Eosennertia Kurosa, 2003: 25. Type species: Sennertia (Eosennertia) bifida Kurosa, 2003,
synonymized by Klimov & OConnor, 2008
Type species: Pediculus cerambycinus Scopoli, 1763
Sennertia (Sennertia) aldeodadi Haitlinger
Sennertia (Sennertia) aldeodadi Haitlinger, 2000: 18
Sennertia aldeodadi : Klimov & OConnor, 2008: 222
Type locality: Mohebourg
Distribution: Mauricius
Host: undetermined Anthophoridae (Hymenoptera)
Sennertia (Sennertia) alfkeni (Oudemans)
Trichotarsus alfkeni Oudemans, 1900: 115; 1901: 82; 1903a: 147; Vitzthum, 1912c: 233
~part., only specimens from Xylocopa circumvolans – see Klimov & OConnor (2008).
Sennertia alfkeni Oudemans, 1905a: 22; 1924: 329; Vitzthum, 1914: 323; 1919: 31; Vitzthum, 1941: 308; Zachvatkin, 1941: 389; Fain, 1974b: 229; Delfinado & Baker, 1976: 85;
Okabe & Makino, 2002: 73; Klimov & OConnor, 2008: 222; Kontschán et al., 2016: 3
Trichotarsus japonicus Oudemans, 1900: 117, synonymized by Zachvatkin, 1941; Oudemans, 1901: 83; 1903a: 147; Trägårdh, 1904: 156
Sennertia japonica (Oudemans): Oudemans, 1905a: 22; Vitzthum, 1914: 323; 1919: 43;
1941: 308; Okabe & Makino, 2002: 73
Sennertia japonicus: Fain, 1974b: 224
Sennertia (Sennertia) japonicus: Fain, 1981b: 163
Sennertia (Sennertia) alfkeni: Fain, 1981b: 163; Fain, 1982: 70; OConnor, 1993a: 362
Sennertia ?bifilis: Womersley, 1941: 480 (after Fain, 1982); from Xylocopa (Koptortosoma)
bryorum (as “Mesotricha bryorum“), Australia: Queensland
Type locality: Japan
Distribution: Australia, China, Japan, Korea
Host: Xylocopa (Alloxylocopa) circumvolans [in Zachvatkin, 1941 as “X. kalinovskii]
(Hymenoptera: Apidae, Xylocopinae)
Misidentifications (after Klimov & OConnor, 2008): Trichotarsus Alfkeni: Vitzthum,
1912c: 233, from Xylocopa (Koptortosoma) aestuans (East Sumatra), Xylocopa (Koptortosoma) caerulea (Java)
Sennertia (Sennertia) antarctica (Trägårdh)
Trichotarsus antarcticus Trägårdh, 1907: 12
Acarorum Catalogus IX 271
Chaetodactylus antarcticus: Pugh, 1993: 373
Sennertia antarctica (Trägårdh): Klimov & OConnor, 2008: 222
Type locality: Booth Is. (as Wandel Is.)
Distribution: Antarctica (The geographic locality is, most likely, erroneous because
carpenter bees, the only hosts of Sennertia, do not occur in Antarctica - Klimov &
Oconnor, 2008)
Host: marin algae
Sennertia (Sennertia) augustii Alzuet et Abrahamovich
Sennertia augustii Alzuet & Abrahamovich, 1990: 628; Klimov & OConnor, 2008: 192
Type locality: Buenos Aires, Berazategui
Distribution: Argentina, Mexico, Uruguay
Hosts: Xylocopa (Neoxylocopa) augusti, X. fimbriata (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) benoiti Fain
Sennertia benoiti Fain, 1974: 218; 1980: 988; Klimov & OConnor, 2008: 223
Sennertia (Sennertia) benoiti Fain: Fain, 1981a: 157; Haitlinger, 2000: 17
Type locality: Bafing, 7 km ESE Touba (Ivory Coast)
Distribution: DR Congo, Ivory Coast
Host: Ceratina sp. (type host), Ceratina atopura (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) bifida Kurosa
Sennertia (Eosennertia) bifida Kurosa, 2003: 26
Sennertia (Sennertia) bifida: Klimov & OConnor, 2008: 224
Type locality: Hitsujigaoka, Sapporo, Hokaido
Distribution: Japan (Hokkaido, Honshu and Ryukyus)
Hosts: Ceratina (Ceratinidia) japonica, C. flavipes, C. okinawana (Hymenoptera: Apidae,
Xylocopinae)
Sennertia (Sennertia) bifilis (Canestrini)
Trichotarsus bifilis Canestrini, 1897: 474; Canestrini & Kramer, 1899: 149; Giard, 1900:
377; Oudemans, 1901: 83; 1903: 147; 1905a: 22
Sennertia bifilis (Canestrini): Vitzthum, 1919: 61; Zachvatkin, 1941: 42; Fain, 1981b: 180;
1982: 70; Lombert et al., 1987: 113; Klimov & OConnor, 2008: 224
Type locality: Madang: Astrolabe Bay (Erima)
Distribution: Papua New Guinea
Host: Xylocopa (Koptortosoma) combinata (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) caffra Vitzthum
Sennertia caffra Vitzthum, 1919: 53; 1941: 310; Zachvatkin, 1941: 42; Klimov & OConnor,
2008: 224
Sennertia (Sennertia) caffra: Fain, 1981b: 159; Lombert et al., 1987: 113; Haitlinger, 2000: 17
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Acarorum Catalogus IX
Type locality: Eastern Cape (Willowmore)
Distribution: South Africa
Host: Xylocopa (Koptortosoma) caffra (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) cantabrica Zachvatkin
Sennertia cantabrica Zachvatkin, 1941: 385; Lombert et al., 1987: 113; Klimov & OConnor,
2008: 224
Sennertia (Sennertia) cantabrica: Fain, 1981b: 163; Haitlinger, 2000: 17
Type locality: Algeria
Distribution: Algeria
Host: Xylocopa (X.) cantabrita (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Senneria) capensis Fain
Sennertia capensis Fain, 1971: 266; Klimov & OConnor, 2008: 225
Sennertia (Sennertia) capensis: Fain, 1981b: 157; Haitlinger, 2000: 17
Type locality: Eastern Cape (Willowmore)
Distribution: South Africa
Host: Xylocopa (Gnathoxylocopa) sicheli (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) cerambycina (Scopoli)
Pediculus cerambycinus Scopoli, 1763: 386
Trichodactylus osmiae: Gervais, 1844: 266 (part.)
Trichodactyle Xylocopae Donnadieu, 1868: 84
Trichodactylus xylocopae: Murray, 1877: 252 (part.); Canestrini & Fanzago, 1878: 205;
Mégnin, 1880: 147; Berlese, 1885: XVIII, No 1; Canestrini & Berlese, 1885: 206
Trichotarsus xylocopae: Dugès,1834; Canestrini, 1888a: 394; 1888b: 23; 1897: 474; Berlese,
1892: fasc. LXV, n 1; 1897: 105; 1903a: 322; Canestrini & Kramer, 1899: 149; Giard,
1900: 377; Oudemans, 1900: 116; 1901: 81; 1902: 44; 1903a: 322; Trägårdh, 1907: 12;
Popovici-Baznosanu, 1913: 32; Vitzthum, 1912c: 232; 1912d: 292
Trichotarsus cerambycinus : Oudemans, 1905b: 80
Sennertia cerambycina (Scopoli): Oudemans, 1905: 21; Vitzthum, 1919: 31; 1929: 77; 1933:
152; Vitzthum, 1941: 307; Zachvatkin, 1941: 382; Eyndhoven, 1941: 325; 1952: 34;
Skaife, 1952: 76; Fain, 1971: 265; Elbadry, 1971: 87; Vicidomini & Meloni, 1999: 138;
Klimov & OConnor, 2008: 226
Sennertia xylocopae: Türk & Türk, 1957: 210
Sennertia (Sennertia) cerambycina (Scopoli): Fain, 1981b: 152; Lombert et al., 1987:
113; OConnor, 1993a: 362; Vicidomini, 1996: 71
Type locality: “Carniola” (now Slovenia)
Acarorum Catalogus IX 273
Distribution: France, Germany, Greece, Holland, Italy, Slovenia, Spain, China; “All southern, central, and eastern Europe, up to 55, 8 N; West and Central Asia up to Tibet
and Gashun Gobi Desert China, on the east and up to northern Iran to the south”.
Hosts: Xylocopa valga, X. varentzowi, X. violacea, Xylocopa (Proxylocopa) nitidiventris,
Xylocopa (P.) przewalskyi, Xylocopa (Copoxyla) turanica (Hymenoptera: Apidae,
Xylocopinae). Also phoretic on Polochrum repandum (Hymenoptera, Sapygidae,
cleptoparasite of Xylocopa, occasionally found on Hoplitis princeps (as Osmia princeps), Osmia (Monosmia) apicata (as Osmia macroglossa), Megachile (Eutricharaea)
leachella (as Megachile argentata), and Anthophora (Paramegilla) balassogloi(as
Anthidium christofi)
Sennertia (Sennertia) ceratinarum Fain
Sennertia ceratinarum Fain, 1974 : 216; 1980: 983; Klimov & OConnor, 2008: 226
Sennertia (Sennertia) ceratinarum Fain: Fain, 1981b: 156; Haitlinger, 2000: 17
Type locality: Salt Rock, Natal
Distribution: Cameroon, D.R. Congo, South Africa
Hosts: Ceratina atopura, C. acutipyga, C. congoensis, C. nilotica, C. turneri, Pithitis viridis
(Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) congoicola Fain
Sennertia congoicola Fain, 1971: 265; Klimov & OConnor, 2008: 227
Sennertia (Sennertia) congoicola: Fain, 1981b: 161; Haitlinger, 1999: 59; 2000: 17
Type locality: Moanda
Distribution: DR Congo
Hosts: “Mesotrichia striata”, Xylocopa (Mesotrichia) torrida, Xylocopa (Koptortosoma)
imitator (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) dalyi Fain
Sennertia dalyi Fain, 1980: 990; Klimov & OConnor, 2008: 227
Sennertia (Sennertia) dalyi: Fain, 1981b: 163; Haitlinger, 2000: 17
Type locality: 3.5 km S Tabarka
Distribution: Tunisia
Hosts: Ceratina (Euceratina) dallatorreana, C.(E.) chalybea, C. (E.) mocsaryi, C. (E.) callosa
(Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) dalyi nilotica Fain
Sennertia dalyi nilotica Fain, 1980: 990; Klimov & OConnor, 2008: 227
Sennertia (Sennertia) dalyi nilotica: Fain, 1981b: 163
Type locality: Al QalyuS bıyah, Banha (“Benha, Nile Delta”)
Distribution: Egypt
Host: Ceratina (Pithitis) tarsata (Hymenoptera: Apidae, Xylocopinae)
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Acarorum Catalogus IX
Sennertia (Sennertia) devincta Klimov et OConnor
Sennertia devincta Klimov & OConnor, in Klimov et al., 2007: 121; Klimov & OConnor,
2008: 148
Type locality: Huanuco
Distribution: Peru
Host: Ceratina sp. (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) dissimilis Zachvatkin
Sennertia dissimilis Zachvatkin, 1941: 385; Fain, 1981b: 182; Klimov & OConnor, 2008: 227
Type locality: Japan
Distribution: Japan
Host: Xylocopa (Biluna) nasalis (as Xylocopa dissimilis) (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) duweinii Sherbef et Duweini
Sennertia duweinii Sherbef & Duweini, 1980: 245; Klimov & OConnor, 2008: 228
Type locality: not indicated
Distribution: ? Egypt
Host: Xylocopa (Koptortosoma) aestuans (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) egyptiaca Elbadry
Sennertia egyptiaca Elbadry, 1971: 87; Sherbef & Duweini, 1980: 246; Fain, 1981b: 182;
Klimov & OConnor, 2008: 228
Type locality: Shoubra Elkheima, Cairo
Distribution: Egypt, Sudan
Host: Xylocopa (Koptortosoma) aestuans (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) elseni Fain
Sennertia elseni Fain, 1971: 266; Klimov & OConnor, 2008: 228
Sennertia (Sennertia) elseni: Fain, 1981b: 161; OConnor, 1993a: 362; Haitlinger, 1999: 59;
2000: 17; Fain & Pauly, 2001: 132
Type locality: Bas-Congo (Muanda)
Distribution: DR Congo, Madagascar
Hosts: Xylocopa (Koptortosoma) caffra (as Mesotrichia olivacea) (Hymenoptera: Apidae,
Xylocopinae); on Madagascar: except Ranomafana: Melanempis sp.; Ranomafana:
cleptoparasitic bee of the family Nomadinae
Acarorum Catalogus IX 275
Sennertia (Sennertia) flabellifera Oudemans
Sennertia flabellifera Oudemans, 1924: 331; Lombert et al., 1987: 113; Klimov & OConnor, 2008: 228
Type locality: Bogor (Java)
Distribution: Indonesia (Java)
Host: Xylocopa (Koptortosoma) sp., nest (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) frontalis Vitzthum
Sennertia frontalis Vitzthum, 1941: 308; Klimov & OConnor, 2008: 228
Type locality: Entre Rios, Argentina
Distribution: Argentina, Brazil, Mexico, Peru, Panama, Suriname, Venezuela
Hosts: Xylocopa frontalis, X. nautlana (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) gargantua Zachvatkin
Sennertia gargantua Zachvatkin, 1941: 385; Klimov & OConnor, 2008: 228
Sennertia (Sennertia) gargantua: Fain, 1981b: 154
Type locality: Turkey
Distribution: Tajikistan, Turkey, Uzbekistan
Hosts: Xylocopa (Nodula) punctilabris, X. (X.) valga (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) greeni (Oudemans)
Hericia greeni Oudemans, 1917: 345
Sennertia greeni (Oudemans): Oudemans, 1924: 330; Zachvatkin, 1941: 42; Lombert et
al., 1987: 113; Klimov & OConnor, 2008: 228
Type locality: Eton Estate, Punduloga
Distribution: Sri Lanka
Host: Xylocopa (Mesotrichia) tenuiscapa (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) haustrifera Klimov et OConnor
Sennertia haustrifera Klimov & OConnor, 2008: 151
Type locality: Jalisco, Estacion de Biologia Chamela
Distribution: Mexico (Jalisco, Chiapas)
Host: Ceratina captiosa (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) herminae Haitlinger
Sennertia (Sennertia) herminae: Haitlinger, 1999: 57; 2000: 17
Sennertia herminae: Klimov & OConnor, 2008: 229
Type locality: Antananarivo
Distribution: Madagascar
Host: Xylocopa sp. (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) hipposideros (Oudemans)
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Acarorum Catalogus IX
Trichotarsus hipposideros Oudemans, 1902: 44; 1903: 145
Trichotarsus hipposiderus Oudemans, 1903a: 145 (unjust. emend.); Trägårdh, 1904: 156;
1907: 12; Vitzthum, 1912d: 290
Sennertia hipposiderus (Oudemans): Oudemans, 1905a: 22; Vitzthum, 1941: 310 (hipposideruns); Fain, 1974b: 226; OConnor, 1993b: 161; Krantz, 1998: 298
Sennertia hipposidera: Vitzthum, 1919: 43 (unjust. emend.)
Sennertia hipposideros: Zachvatkin, 1941: 42; Fain, 1974 : 226; Klimov & OConnor, 2008:
229
Sennertia (Sennertia) hipposiderus (Oudemans): Fain, 1981b: 156
Type locality: India
Distribution: India, Malaysia
Hosts: Xylocopa (Platynopoda) latipes, X. (Mesotrichia) tenuiscapa (Hymenoptera: Apidae,
Xylocopinae)
Sennertia (Sennertia) horrida (Vitzthum)
Trichotarsus horridus Vitzthum, 1912: 290
Sennertia horrida (Vitzthum): Vitzthum, 1919: 57; 1941: 308; Zachvatkin, 1941: 387;
Klimov et al., 2007b: 124; Klimov & OConnor, 2008: 229
Sennertia (Sennertia) horrida: Fain, 1981b: 170
Sennertia sumatrensis Oudemans, 1924: 329; Fain, 1974b: 228 (synonymized by Fain,
1981b)
Sennertia leucothorae Ramaraju & Mohanasundaram, 2001: 107 (Type: Tamil Nadu,
Coimbatore; synonymized by Klimov et al., 2007b)
Type locality: Jakarta (Java)
Distribution: India, Indonesia (Java, Sumatra), Japan, Taiwan
Hosts: Xylocopa (Biluna) nasalis (as Xylocopa (Koptorthosoma?) dissimilis), Xylocopa (Koptortosoma) aestuans (as Xylocopa leucothorae), X. auripennis, X. fallax, X. iridipennis,
X. mcgregori, X. mimetica (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) hurdi Klimov et OConnor
Sennertia hurdi Klimov & OConnor, 2008: 164
Type locality: Oaxaca, Presa Benito Juárez, nr. Jalapa de Marquez
Distribution: Mexico
Hosts: Xylocopa t. tabaniformis, X. tabaniformis azteca (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) indica Delfinado et Baker
Sonnertia [sic !] indica Delfinado & Baker, 1976: 87
Seneria [sic !] indica: Ramaraju & Mohanasundaram, 2001: 107
Sennertia indica: Fain, 1981b: 182 (inadequately described); Klimov & OConnor, 2008: 230
Type locality: Hissar
Acarorum Catalogus IX 277
Distribution: India
Host: Ceratina binghami (as Pithitis binghami) (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) koptorthosomae (Oudemans)
Trichotarsus koptorthosomae Oudemans, 1901: 82; 1903a: 47; Vitzthum, 1912d: 290 (part.)
Trichotarsus coptorthosomae Trägårdh, 1904: 156 (unjust. emend.)
Sennertia coptorthosomae: Oudemans, 1905a: 22 (unjust. emend.)
Sennertia koptorthosomae (Oudemans): Oudemans, 1924: 329; Zachvatkin, 1941: 42; Fain,
1974: 219; OConnor 1993b: 161; Krantz, 1998: 298; Klimov & OConnor, 2008: 192
Sennertia (Sennertia) koptorthosomae (Oudemans): Fain, 1981b: 156
Type locality: Java (Bogor)
Distribution: India, Indonesia (Java, Sumatra), Malaysia, Sri Lanka
Hosts: nest of Xylocopa (Mesotrichia) latipes, X. (M.) tenuiscapa [Koptorthosoma] (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) latipilis Fain
Sennertia latipilis Fain, 1974: 217; 1980: 986; Klimov & OConnor, 2008: 231
Sennertia (Sennertia) latipilis: Fain, 1981a: 156; Haitlinger, 2000: 17
Type locality: Malindi
Distribution: Kenya
Host: Ceratina (Ctenoceratina) penicilligera (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) lauta Klimov and OConnor
Sennertia lauta Klimov & OConnor, in Klimov et al., 2007b: 125; Klimov & OConnor,
2008: 231
Type locality: Davao (Philippines)
Distribution: Malaysia (Sarawak), Philippines (Mindanao, Palawan)
Hosts: Xylocopa (Zonohirsuta) fuliginata, X. (Z.) dejeanii (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) leclercqi Fain
Sennertia leclercqi Fain, 1971: 268; Klimov & OConnor, 2008: 232
Sennertia (Sennertia) leclercqi: Fain, 1981b: 157; Haitlinger, 2000: 17
Type locality: Sud-Kivu, Uvira
Distribution: DR Congo
Host: Xylocopa (Xenoxylocopa) inconstans (as Mesotrichia inconstans) (Hymenoptera:
Apidae, Xylocopinae)
Sennertia (Sennertia) leei Fain
Sennertia (Sennertia) leei Fain, 1982: 67; OConnor, 1993a: 362
Sennertia leei Fain: Klimov & OConnor, 2008: 150, 192
278
Acarorum Catalogus IX
Type locality: Ku-rin-gai, New South Wales
Distribution: Australia (N.S.W., Queensland, SA)
Hosts: Xylocopa (Lestis) bombylans, X. aerata (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) loricata Klimov et OConnor
Sennertia loricata Klimov & OConnor, 2008: 181
Type locality: Florida, Alachua Co., Gainesville
Distribution: Mexico (Chiapas, Jalisco), USA (Florida, Louisiana, Puerto Rico, Texas,
Virginia)
Hosts: Xylocopa b. barbatella, X. micans, X. mordax, X. muscaria (Hymenoptera: Apidae,
Xylocopinae)
Sennertia (Sennertia) lucrosa Klimov and OConnor
Sennertia lucrosa Klimov & OConnor, 2008: 164
Type locality: California, Los Angeles Co., Los Angeles, Crenshaw Distr.
Distribution: Mexico, USA
Hosts: Xylocopa californica, X. clarionensis, X. guatemalensis, X. mexicanorum, X. varipuncta (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) madagascarensis Fain
Sennertia madagascarensis Fain, 1971: 270; Fain & Pauly, 2001: 125; Klimov et al., 2007b;
Klimov & OConnor, 2008: 232
Sennertia (Sennertia) madagascarensis: Fain, 1981b: 170; OConnor, 1993a: 362
Type locality: Antananarivo (“Tananarive”)
Distribution: Madagascar
Hosts: Xylocopa (Prosopoxylocopa) mirabilis, Xylocopa (Koptortosoma) calens (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) mesotrichia Fain
Sennertia mesotrichia Fain, 1971: 265; Klimov & OConnor, 2008: 232
Sennertia (Sennertia) mesotrichia: Fain, 1981b: 154; Haitlinger, 2000: 17
Type locality: Bambesa, Uele
Distribution: D.R. Congo
Hosts: Xylocopa (Koptortosoma) africana (as Mesotrichia africana), X. (K.) lepeletieri,
Xylocopa (Xylomelissa) tarsata (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) micheli Fain
Sennertia micheli Fain, 1971: 268; Klimov & OConnor, 2008: 232
Sennertia (Sennertia) micheli: Fain, 1981b: 164
Type locality: Java
Distribution: Indonesia (Java)
Host: Bembix borrei (Hymenoptera: Sphecidae)
Acarorum Catalogus IX 279
Sennertia (Sennertia) morstatti Vitzthum
Sennertia morstatti Vitzthum, 1914: 323; 1919: 42; Zachvatkin, 1941: 42; Klimov &
OConnor, 2008: 233
Sennertia (Sennertia) morstatti: Fain, 1981b: 164; Lombert et al., 1987: 113; Sherbef &
Duweini, 1980: 245
Sennertia moandensis Fain, 1971: 267 (synonymized by Fain, 1981b). Type: Moanda, DR
Congo
Sennertia tanythrix Fain, 1971: 267 (part., specimens from X. nigrita)
Type locality: Tanzania, Amani
Distribution: DR Congo, Sudan, Tanzania
Hosts: Xylocopa (Koptortosoma) nigrita (Hymenoptera: Apidae, Xylocopinae), Megachilidae
Sennertia (Sennertia) perturbans Vitzthum
Sennertia perturbans Vitzthum, 1919: 48; Zachvatkin, 1941: 42; Fain, 1981b: 145; Lombert
et al., 1987: 113; Klimov & OConnor, 2008: 233
Type locality: Amani
Distribution: Tanzania
Host: Xylocopa (Koptortosoma) nigrita (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) pirata Klimov et OConnor
Sennertia pirata Klimov & OConnor, 2008: 144
Type locality: Bahamas, Bimini, South Bimini Island
Distribution: Bahamas, British Virgin Islands (Guana Is.), Cuba, Virgin Islands of the
United States, Puerto Rico
Hosts: Xylocopa cubaecola, X. mordax (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) potanini Zachvatkin
Sennertia potanini Zachvatkin, 1941: 390; Klimov & OConnor, 2008: 233
Sennertia (Sennertia) potanini: Fain, 1981b: 170; Klimov et al., 2007b: 125
Type locality: Sichuan, Yazhou
Distribution: China
Host: Xylocopa (Biluna) tranquebarorum or Xylocopa (Bomboixylocopa)
rufipes (as Xylocopa pictifrons) (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) ratiocinator Klimov et OConnor
Sennertia ratiocinator Klimov et OConnor, in Klimov et al., 2007b: 128; Klimov &
OConnor, 2008: 234
Type locality: NW Sri Lanka
Distribution: Sri Lanka
280
Acarorum Catalogus IX
Hosts: Xylocopa (Zonohirsuta) bhowara, X. (Z.) dejeanii, Xylocopa (Nodula) amethystina
(Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) recondita Klimov et OConnor
Sennertia recondita Klimov & OConnor, 2008: 151
Type locality: Quintana Roo, Vallarta, 17 km W Pto. Morelos
Distribution: Mexico
Host: Ceratina (Calloceratina) exemia (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) robusta Delfinado et Baker
Sennertia robusta Delfinado et Baker, 1976: 87; Klimov & OConnor, 2008: 234
Sennertia (Sennertia) robusta: Fain, 1981b: 157; Haitlinger, 2000: 18
Sennertia carpenteri Ramaraju & Mohanasundaram, 2001: 109 – Type: Tamil Nadu,
Coimbatore
Type locality: Hissar
Distribution: India
Hosts: «Megachilid bee«, Xylocopa (Nodula) amethystina (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) roepkei Oudemans
Sennertia roepkei Oudemans, 1924: 330; Lombert et al., 1987: 113; Klimov & OConnor,
2008: 234
Type locality: Java (Bogor)
Distribution: Indonesia (Java)
Host: On eggs of Horia (as “Cissites)”(Coleoptera: Meloidae) in nest of Xylocopa sp.
(Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) sayutara Klimov et OConnor
Sennertia sayutara Klimov & OConnor, in Klimov et al., 2007: 122; 2008: 161
Type locality: San José
Distribution: Costa Rica
Host: Ceratina laticeps (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) scutata Fain
Sennertia scutata Fain, 1974: 217; 1980: 986; Klimov & OConnor, 2008: 234
Sennertia (Sennertia) scutata: Fain, 1981b: 159; Haitlinger, 2000: 17
Type locality: Cape Town
Distribution: South Africa, El Salvador, Guatemala
Hosts: Ceratina truncata, Ceratina (Pithitis) nasalis (as Ceratina viridior) (Hymenoptera:
Apidae, Xylocopinae)
Acarorum Catalogus IX 281
Sennertia (Sennertia) segnis Klimov et OConnor
Sennertia segnis Klimov & OConnor, 2008: 170
Type locality: California, Tulare Co., 2 mi N Hartland
Distribution: Mexico, USA (California, Arizona)
Hosts: Xylocopa c. californica, X. californica arizonensis, X. cyanea, X. tabaniformis orpifex
(Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) shimanukii Baker et Delfinado-Baker
Sennertia shimanukii Baker & Delfinado-Baker, 1983: 117; Klimov & OConnor, 2008: 176
Type locality: Sololá, San Lucas Tolimán (Guatemala)
Distribution: Guatemala, Mexico, USA
Hosts: Xylocopa (Neoxylocopa) fimbriata, X. (N.) mexicanorum, X. (N.) sonorina, X. (N.)
varipuncta, Xylocopa (Xylocopoides) californica arizonensis (Hymenoptera: Apidae,
Xylocopinae); Apis mellifera (Hymenoptera: Apidae, Apinae)(type host, accidental)
Sennertia (Sennertia) simplex (Trägårdh)
Trichotarsus simplex Trägårdh, 1904: 156
Sennertia simplex (Trägårdh): Vitzthum, 1919: 32; 1941: 308; Zachvatkin, 1941: 42; Klimov
& OConnor, 2008: 235
Sennertia (Sennertia) simplex: Fain, 1981b: 159; Haitlinger, 2000: 17
Type locality: Cameroon
Distribution: Cameroon
Host: Xylocopa (Koptortosoma) nigrita (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) sodalis Klimov et OConnor
Sennertia sodalis Klimov & OConnor, 2008: 156
Type locality: San Juan de la Punta (Veracruz)
Distribution: Costa Rica, Mexico, Panama
Host: Ceratina sp., Ceratina (Calloceratina) eximia (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) spinifera Fain
Sennertia ceratinarum spinifera Fain, 1974: 217
Sennertia (Sennertia) spinifera Fain, 1980: 984; 1981b: 156; Haitlinger, 2000: 17
Sennertia spinifera Fain: Klimov & OConnor, 2008: 235
Type locality: N. E. Lusambo
Distribution: D.R. Congo
Host: Ceratina (Propithitis) aereola (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) surinamensis Fain et Lukoschus
Sennertia surinamensis Fain & Lukoschus, 1971: 270; Klimov et al., 2007a: 1371; Klimov
& OConnor, 2008: 237
282
Acarorum Catalogus IX
Sennertia (Sennertia) surinamensis (Fain et Lukoschus): Fain, 1981b: 170; Alzuet & Abrahamovich, 1987: 348
Type locality: Paramaribo
Distribution: French Guiana, Panama, Suriname
Host: Ceratina (Calloceratina) chloris (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) tanythrix Fain
Sennertia tanythrix Fain, 1971: 267; Klimov & OConnor, 2008: 236
Sennertia (Sennertia) tanythrix: Fain, 1981b: 167; Haitlinger, 2000: 17
Type locality: Uele, Haut-Congo Prov.
Distribution: DR Congo
Hosts: Xylocopa (Mesotrichia) torrida, Xylocopa (Koptortosoma) imitator, X. (K.) nigrita
(Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) tunisiana Fain
Sennertia tunisiana Fain, 1980: 991; Klimov & OConnor, 2008: 236
Sennertia (Sennertia) tunisiana: Fain, 1981b: 163; Haitlinger, 2000: 17
Type locality: Silya nah Maktar
Distribution: Tunisia
Hosts: Ceratina (Ceratina) cucurbitina, Ceratina (Euceratina) albosticta, Ceratina (Euceratina) mocsaryi, Ceratina dallatorreana, C. chalybea (Hymenoptera: Apidae,
Xylocopinae)
Sennertia (Sennertia) vaga Klimov et OConnor
Sennertia vaga Klimov & OConnor, 2008: 150
Type locality: California, Los Angeles Co., Rustic Canyon
Distribution: Mexico, USA
Hosts: Xylocopa (Notoxylocopa) tabaniformis orpifex, X. (N.) tabaniformis androleuca, X.
(Xylocopoides) californica (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) vanderhammeni Fain
Trichotarsus koptorthosomae Oudemans, 1901: 82 (part.)
Sennertia vanderhammeni Fain, 1974: 222; Klimov & OConnor, 2008: 236
Sennertia (Sennertia) vanderhammeni: Fain, 1981b: 156
Type locality: Peradeniya
Distribution: India, Indonesia (Java), Sri Lanka
Host: Xylocopa (Koptorthosoma) tenuiscapa (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) vanderhammeni brevipilis Fain
Trichotarsus koptorthosomae Oudemans, 1901: 82 (part.)
Acarorum Catalogus IX 283
Sennertia vanderhammeni brevipilis Fain, 1974: 224; 1981b: 156; Klimov & OConnor,
2008: 236
Sennertia (Sennertia) vanderhammeni brevipilis: Fain, 1981b: 156; Alzuet
& Abrahamovich, 1989: 236; 1990: 319
Type locality: Java
Distribution: Indonesia (Java)
Host: Xylocopa (Koptorthosoma) tenuiscapa (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) varicosa Fain .
Sennertia varicosa Fain, 1971: 268; Klimov & OConnor, 2008: 237
Sennertia (Sennertia) varicosa: Fain, 1981b: 167; Haitlinger, 2000: 17
Type locality: Katanga – Monts Kundelungu (DR Congo)
Distribution: Angola, DR Congo
Hosts: Xylocopa (Mesotrichia) sp., X. (Xenoxylocopa) inconstans (as Mesotrichia
flavescens inconstans) (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Sennertia) zhelochovtsevi Zachvatkin
Sennertia zhelochovtsevi Zachvatkin, 1941: 388; Turk, 1948: 84; Klimov et al., 2007a: 1371;
Klimov & OConnor, 2008: 237; Nazan et al., 2019
Sennertia (Sennertia) zhelochovtsevi Zachvatkin: Fain, 1981b: 148
Type locality: Greece
Distribution: Armenia, Greece, Iran, Turkey, Uzbekistan
Hosts: Xylocopa olivieri, X. rufa (Hymenoptera: Apidae, Xylocopinae)
Subgenus Spinosennertia Fain
Spinosennertia Fain, 1981b: 147; Klimov & OConnor, 2008: 147
Type species: Sennertia argentina Vitzthum, 1941
Sennertia (Spinosennertia) argentina Vitzthum
Sennertia argentina Vitzthum, 1941: 309; Klimov & OConnor, 2008: 156
Sennertia (Spinosennertia) argentina Vitzthum: Fain, 1981b: 147
Type locality: Argentina
Distribution: Argentina, Brazil, Costa Rica, Guatemala, Mexico, Nicaragua, Panama,
Peru, Suriname
Hosts: Xylocopa (Neoxylocopa) fimbriata, X. (N.) frontalis, X. (N.) mexicanorum, X. (N.)
nasica, X. (N.) nautlana (Hymenoptera: Apidae, Xylocopinae)
Sennertia (Spinosennertia) donaldi Turk
Sennertia donaldi Turk, 1948: 84; Klimov & OConnor, 2008: 148
Sennertia (Spinosennertia) argentina: Fain, 1981b: 176 (part.); Alzuet & Abrahamovich,
1987: 350 (part.); Haitlinger, 1999: 59 (part.)
284
Acarorum Catalogus IX
Sennertia (Spinosennertia) donaldi: Klimov et al., 2007b: 130
Type locality: Piarco
Distribution: Brazil, Trinidad and Tobago, Venezuela
Hosts: Xylocopa (Neoxylocopa) fimbriata, X. (N.) frontalis (Hymenoptera: Apidae, Xylocopinae)
Speciae inquirendae and Taxa Incorrectly attributed to Chaetodactylidae – see
in Klimov & OConnor, 2008
Index of Chaetodactylidae
(Names in bold are accepted as valid)
abditus, Chaetodactylus 258
Achaetodactylus 257
Afrosennertia 265
aldeodadi, Sennertia 268
alfkeni, Trichotarsus 268
alfkeni, Sennertia 268
americana, Sennertia 267
Amsennertia 267
antarctica, Sennertia 269
antarcticus, Chaetodactylus 269
antarcticus, Trichotarsus 269
anthidii, Chaetodactylus 269
anthidii, Trichotarsus 269
antillarum, Chaetodactylus 269
argentina, Sennertia 282
Asiosennertia 265
augustii, Sennertia 269
aurulenticola, Chaetodactylus chrysidis
259
azteca, Chaetodactylus 259
bakeri, Sennertia 265
basilewskyi, Sennertia 265
basilewskyi, Sennertia (Afrosennertia) 283
benoiti, Sennertia 269
bifida, Sennertia 269
bifida, Sennertia (Eosennertia) 269
bifilis, Sennertia 270
bifilis, Trichotarsus 270
birulai, Chaetodactylus 259
brevipilis, Sennertia vanderhammeni 281
caffra, Sennertia 270
cantabrica, Sennertia 270
capensis, Sennertia 270
carpenteri, Sennertia 277
Centriacarus 258
cerambycina, Sennertia 270
cerambycinus, Pediculus 270
cerambycinus, Trichotarsus 271
ceratinae, Achaetodactylus 257
ceratinae, Chaetodactylus 257
ceratinarum, Sennertia 271
Chaetodactylidae 257
Chaetodactylinae 257
Chaetodactylini 256
Chaetodactylus 257
chrysidis, Chaetodactylus 259
claviger, Chaetodactylus 260
claviger, Chaetodactylus (Spinodactylus) 260
claudus, Chaetodactylus 260
congoicola, Sennertia 271
coptorthosomae, Sennertia 275
dalyi, Chaetodactylus 260
dalyi, Sennertia 272
decellei, Achaetodactylus 257
Acarorum Catalogus IX 285
decellei, Chaetodactylus (Ochaetodactylus) 257
decellei, Ochaetodactylus 257
delfinadoae, Sennertia 265
dementjevi, Chaetodactylus 260
devincta, Sennertia 272
dissimilis, Sennertia 272
donaldi, Sennertia 282
duweinii, Sennertia 272
egyptiaca, Sennertia 272
elseni, Sennertia 273
Eosennertia 268
faini, Sennertia 267
flabellifera, Sennertia 273
frontalis, Sennertia 267
furunculus, Chaetodactylus 260
gargantua, Sennertia 273
gibbosi, Chaetodactylus 261
greeni, Sennertia 273
greeni, Hericia 273
guahibo, Centriacarus 258
haustrifera, Sennertia 274
herminae, Sennertia 274
hipposidera, Sennertia 274
hipposideros, Sennertia 274
hipposiderus, Sennertia 274
hipposideros, Trichotarsus 274
hipposiderus, Trichotarsus 274
hirashimai, Chaetodactylus 261
hopliti, Chaetodactylus 261
horrida, Sennertia 274
horridus, Trichotarsus 274
hurdi, Sennertia 275
ignota, Sennertia 267
imberba, Roubikia 264
indica, Sennertia 275
japonica,Sennertia 268
japonicus, Sennertia 269
japonicus, Trichotarsus 269
jeanalexi, Sennertia 266
koptorthosomae, Sennertia 275
koptorthosomae, Trichotarsus 275
kouboy, Chaetodactylus 261
krombeini, Chaetodactylus 261
krombeini, Chaetodactylus (Spinodactylus) 261
lassulus, Chaetodactylus 262
latebrosa, Roubikia 264
latipilis, Sennertia 275
lauta, Sennertia 276
leclercqi, Sennertia 276
leei, Senertia 276
leleupi, Achaetodactylus 258
leleupi, Chaetodactylus 258
leucothorae, Sennertia 274
lithurgi, Chaetodactylus 262
longipilis, Sennertia 268
loricata, Sennertia 276
lucrosa, Sennertia 276
ludwigi, Chaetodactylus 262
ludwigi, Trichotarsus 262
madagascarensis, Sennertia 277
mahunkai, Chaetodactylus 261
melitomae, Chaetodactylus 262
mesotrichia, Sennertia 277
micheli, Sennertia 277
micheneri, Chaetodactylus 262
moandensis, Sennertia 277
monicae, Sennertia 266
morstatti, Sennertia 277
nilotica, Sennertia dalyi 272
nipponicus, Chaetodactylus 263
Ochaetodactylus 257
officiosa, Roubikia 265
osmiae, Chaetodactylus 263
osmiae, Trichodactylus 263
osmiae, Trichotarsus 263
oudemansi, Sennertia 266
panamensis, Chaetodactylus 265
panamensis, Roubikia 265
perturbans, Sennertia 277
pirata, Sennertia 278
poetae, Chaetodactylus 259
potanini, Sennertia 278
286
Acarorum Catalogus IX
queenslandica, Sennertia 266
ratiocinator, Sennertia 278
reaumuri, Trichotarsus 263
reaumuri, Chaetodactylus 263
recondita, Sennertia 277
robusta, Sennertia 277
roepkei, Sennertia 279
Roubikia 264
rozeni, Chaetodactylus 264
sayutara, Sennertia 279
scutata, Sennertia 279
segnis, Sennertia 279
Sennertia 265
shimanukii, Sennertia 279
simplex, Sennertia 279
simplex, Trichotarsus 279
sodalis, Sennertia 280
spinifera, Sennertia 280
Spinodactylus
Spinosennertia 282
splendidulae, Sennertia 268
sumatrensis, Sennertia 274
surinamensis, Sennertia 280
tanythrix, Sennertia 280
Trichotarsus
tunisiana, Sennertia 280
turbator, Centriacarus 258
vaga, Sennertia 281
vanderhammeni, Sennertia 281
varicosa, Sennertia 281
vitzthumi, Sennertia 266
xylocopae, Sennertia 271
xylocopae, Trichodactylus 270
xylocopae, Trichotarsus 271
zachvatkini, Chaetodactylus 264
zhelochovtsevi, Sennertia 281
Distribution of Chaetodactylidae
EUROPE
Armenia (Zachvatkin, 1941) – Sennertia zhelochovtsevi
Belgium (Fain & Baugnée, 1996 ; Van Asselt, 2000; Klimov & OConnor, 2008) – Chaetodactylus chrysidis, Ch. osmiae
Bulgaria (Beron, 2010) – Sennertia sp.
Croatia (Klimov & OConnor, 2008) – Chaetodactylus osmiae
Czech Rep. (Oudemans, 1905) – Chaetodactylus birulai (= poetae), Ch. reaumuri
France (contin.) (Dufour, 1839;) – Chaetodactylus anthidii, Ch. claviger, Ch. osmiae,
Sennertia cerambycina
Georgia (Klimov & OConnor, 2008) - Chaetodactylus osmiae, Ch. reaumuri,
Germany (Oudemans, 1905) – Chaetodactylus chrysidis, Ch. osmiae, Ch. reaumuri, Sennertia cerambycina
Great Britain (Michael, 1901; Turk, 1953) – Chaetodactylus osmiae
Greece (Zachvatkin, 1941) – Ch. reaumuri, Sennertia cerambycina, S. zhelochovtsevi
Holland (Eyndhoven, 1952; Siepel, Cremers & Vierbergen, 2016) – Chaetodactylus birulai,
Ch. osmiae, Sennertia cerambycina
Hungary (Samšinak, 1973; Klimov & OConnor, 2008) – Chaetodactylus osmiae (=
mahunkai)
Acarorum Catalogus IX 287
Italy (contin.) (Oudemans, 1924, 1928; Fain, 1981b; Klimov & OConnor, 2008) – Chaetodactylus claviger, Ch. osmiae, Ch. reaumuri, Ch. zachvatkini, Sennertia cerambycina
Luxemburg - Chaetodactylus osmiae
Poland (Chmielewski, 1993) – Chaetodactylus osmiae
Romania (Popovici-Baznosanu, 1913; Klimov & OConnor, 2008) – Chaetodactylus osmiae
Russia (all, continent.) (Zachvatkin, 1941) – Chaetodactylus birulai, Ch. dementjevi,
Ch. osmiae
Crimea (Zachvatkin, 1941) - Chaetodactylus reaumuri
Serbia (Krunić et al., 2005) – Chaetodactylus osmiae
Slovenia (Scopoly, 1763) – Sennertia cerambycina
Spain (cont.) (Fain, 1981b; Bosch, 1992) – Chaetodactylus osmiae, Ch. reaumuri, Ch.
zachvatkini, Sennertia cerambycina
Turkey (all) (Zachvatkin, 1941; Fain, 1981b) – Ch. reaumuri, Sennertia gargantua, S.
zhelochovtsevi
Western Palaearctic (Zachvatkin, 1941) – Chaetodactylus dementjevi
AFRICA
Algeria (Zachvatkin, 1941) – Sennertia cantabrica
Angola Sennertia varicosa
Cameroon (Trägårdh, 1904; Klimov & OConnor, 2008) – Achaetodactylus decellei, A.
leleupi, Sennertia af. basilewskyi, S. simplex
Congo DR (Fain, 1981b) – Achaetodactylus decellei, A. leleupi, Sennertia basilewskyi, S.
benoiti, S. ceratinarum, S. elseni, S. jeanalexi, S. leclercki, S. mesotrichia, S. monicae,
S. morstatti, S. spinifera, S. tanythrix, S.varicosa
Egypt (Elbadry, 1971; Sherbef & El-Duweini, 1980; Faini, 1981a) – Chaetodactylus anthidii,
Ch. claviger, Sennertia dalyi nilotica, S. duweinii, S. egyptiaca
Ivory Coast (Fain, 1974) – Sennertia benoiti
Kenya (Fain, 1974) – Achaetodactylus ceratinae, A. leleupi, Sennertia latipilis
Madagascar (Fain, 1971; Haitlinger, 1999; Fain & Pauly, 2001) – Chaetodactylus dalyi,
Ch. ludwigi, Sennertia elseni, S. herminae, S. madagascarensis
Mauricius (Haitlinger, 2000) - Sennertia aldeodadi
Mozambique (Fain, 1974) – Chaetodactylus dalyi
South Africa (Fain, 1974; Klimov & OConnor, 2008) – Achaetodactylus ceratinae, Chaetodactylus dalyi, Senertia caffra, S. capensis, S. ceratinarum, S. scutata
Sudan – Sennertia egyptiaca, S. morstatti
Tanzania (Vitzthum, 1914, 1919) – Achaetodactylus ceratinae, A. decellei, A. leleupi,
Sennertia morstatti, S. perturbans
Tunisia (Oudemans, 1911; Fain, 1980; ) – Chaetodactylus anthidii, Ch. reaumuri, Ch.
zachvatkini, Sennertia dalyi, S. tunisiana
Zimbabwe (Fain, 1974) – Chaetodactylus dalyi
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ASIA
China (Zachvatkin, 1941) – Sennertia alfkeni, S. cerambycinus, S. potanini
Taiwan (Kawazoe et al., 2010) – Sennertia horrida
India (Oudemans, 1902 ; Ramaraju & Mohanasundaram, 2001; Fain & Pauly, 2001) –
Chaetodactylus ludwigi, Sennertia delfinadoae, S. hipposideros, S. horrida, S. indica,
S. koptorthosomae, S. robusta (= carpenteri), S. vanderhammeni
Indonesia (Vitzthum, 1912; Oudemans, 1924; Fain, 1971, 1981b; ) - Chaetodactylus ludwigi, Sennertia flabellifera, S. horrida (= sumatrensis), S. koptorthosomae, S. micheli,
S. roepkei, S. vanderhammeni, S. vanderhammeni brevipilis
Japan (Oudemans, 1901 ; Hirashima, 1957; Krombein, 1962; Fain, 1974 ; Kurosa, 1987,
2003; Qu et al., 2002; Okabe & Makino, 2002, 2005; Okabe, Makino & Endo, 2008;
Kawazoe et al., 2010) – Chaetodactylus hirashimai, Ch. nipponicus (Tsushima Is.,
Honshu), Sennertia alfkeni, S. horrida, S. japonicus, S. oudemansi
Kazakhstan (Zachvatkin, 1941) – Chaetodactylus dementjevi
Korea (Kontschán et al., 2016) – Sennertia alfkeni
Malaysia (Fain, 1981b; Klimov & OConnor, 2007) – Sennertia hipposideros, S. kopthortosomae, S. lauta
Philippines (Klimov & OConnor, 2007) - Sennertia lauta
Sri Lanka (Oudemans, 1917; Klimov et OConnor, 2007) – Sennertia greeni, S. kopthortosomae, S. ratiocinator, S. vanderhammeni
Tajikistan (Zachvatkin, 1941) - Sennertia gargantua
Uzbekistan (Zachvatkin, 1941) – Chaetodactylus dementjevi, Sennertia gargantua, S.
zhelochovtsevi
NORTH AMERICA
Canada (Klimov & OConnor, 2008) – Chaetodactylus hopliti, Ch. krombeini, Ch. micheneri
Mexico (Klimov & OConnor, 2007, 2008) – Centriacarus turbator, Chaetodactylus abditus,
Ch. melitomae, Roubikia officiosa, R. panamensis, Sennertia argentina, S. augustii,
S. frontalis, S. haustrifera, S. hurdi, S. loricata, S. lucrosa, S. recondita, S. segnis, S.
shimanukii, S. sodalis, S. vaga
USA (Osborn, 1893; Baker, 1962; Klimov & OConnor, 2008; McKinney & Park, 2012)
– Chaetodactylus abditus, Ch. azteca, Ch. claudus, Ch. furunculus, Ch. gibbosi, Ch.
hopliti, Ch. kouboy, Ch. krombeini, Ch. lithurgi, Ch. micheneri, Ch. rozeni, Sennertia
americana, S. loricata, S. lucrosa, S. segnis, S. shimanukii, S. vaga
Puerto Rico (Klimov & OConnor, 2008) - Sennertia loricata, S. pirata
Virgin Islands of the USA (Klimov & OConnor, 2008) - Sennertia pirata
SOUTH and CENTRAL AMERICA
Argentina (Vitzthum, 1941; Alzuet & Abrahamovich, 1989) – Roubikia imberba, Sennertia
argentina, S. augustii, S. frontalis, S. longipilis, S. splendidulae
Bahamas (Klimov & OConnor, 2008) - Sennertia pirata
Acarorum Catalogus IX 289
Belize (Klimov & OConnor, 2008) – Sennertia recondita
Bolivia (Klimov & OConnor, 2008) – Roubikia panamensis
Brazil (Klimov & OConnor, 2008) – Centriacarus turbator, Roubikia panamensis, Sennertia
argentina, S. donaldi, S. frontalis
British Virgin Islands (Klimov & OConnor, 2008) - Sennertia pirata
Chile (contin.) (Klimov & OConnor, 2007; Montalva & Allendes, 2010) – Chaetodactylus
lassulus, Sennertia longipilis
Colombia (Klimov & OConnor, 2008) – Centriacarus turbator
Costa Rica (Klimov & OConnor, 2007, 2008) – Sennertia argentina, S. sayutara, S. sodalis
Cuba (Klimov & OConnor, 2008) – Sennertia pirata
Dominican Republic (Klimov & OConnor, 2008) – Chaetodactylus antillarum
El Salvador (Klimov & OConnor, 2008) – Sennertia scutata
French Guiana (Klimov & OConnor, 2008) – Roubikia panamensis, Sennertia surinamensis
Guatemala (Baker & Delfinado-Baker, 1983; ) – Sennertia faini, S. scutata S. shimanukii
Honduras (Klimov & OConnor, 2007) – Chaetodactylus melitomae, Sennertia frontalis
Jamaica (Klimov & OConnor, 2008) – Chaetodactylus antillarum
Nicaragua (Klimov & OConnor, 2008) – Sennertia argentina
Panama (Baker, Roubik & Delfinado-Baker, 1987; Klimov & OConnor, 2008) – Centriacarus turbator, Roubikia panamensis, Sennertia argentyina, S. frontalis, S. sodalis,
S. surinamensis
Peru (Delfinado & Baker, 1976; Klimov & OConnor, 2007, 2008) – Roubikia latebrosa,
Sennertia argentina, S. devincta, S. frontalis, S. ignota
Suriname (Fain & Lukoschus, 1971) - Sennertia argentina, S. frontalis, S. surinamensis
Trinidad (Turk, 1948) – Sennertia donaldi
Uruguay (Klimov & OConnor, 2008) – Sennertia augustii
Venezuela (Klimov & OConnor, 2007) – Centriacarus guahibo, Sennertia donaldi, S. frontalis
Virgin Islands of the USA (Klimov & OConnor, 2008) - Sennertia pirata
OCEANIA
Australia (continent) (Womersley, 1941; Fain, 1982; Houston, 1992) – Sennertia queenslandica, S. alfkeni, S. leei
Papua New Guinea (Canestrini, 1897) – Sennertia bifilis
New Caledonia (Klimov & OConnor, 2008; Fain & Pauly, 2001) – Chaetodactylus ludwigi
Micronesia (Klimov & OConnor, 2008) - Chaetodactylus ludwigi
French Polynesia (Klimov & OConnor, 2008; Fain & Pauly, 2001) - Chaetodactylus ludwigi
? ANTARCTICA
(Trägårdh, 1907) - Sennertia antarctica
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Hosts of Chaetodactylidae
Insecta
Hymenoptera
Fam. Apidae
Subfam. Xylocopinae
Ceratina aereola - Achaetodactylus decellei
Ceratina apaca – Achaetodactylus ceratinae
Ceratina cucurbitina – Sennertia tunisiana
Ceratina diloloensis – Achaetodactylus leleupi
Ceratina excavata – Achaetodactylus decellei
Ceratina exemia – Sennertia recondita
Ceratina lativentris – Achaetodactylus ceratinae
Ceratina nigriceps - Achaetodactylus ceratinae
Ceratina penicilligera – Sennertia latipilis
Ceratina ruwenzorica - Achaetodactylus leleupi
Ceratina spilota - Achaetodactylus leleupi, A. decellei
Ceratina turneri – Chaetodactylus dalyi, Sennertia ceratinarum
Ceratina sp. – Sennertia benoiti
Xylocopa aerata - Sennertia leei
Xylocopa aestuans – Sennertia duweinii, S. egyptiaca, S. horrida
Xylocopa amethystina – Sennertia delfinadoae (= bakeri), S. ratiocinator,
S. robusta
Xylocopa auripennis - Sennertia horrida
Xylocopa b. barbatella – Sennertia loricata
Xylocopa bhowara - Sennertia ratiocinator
Xylocopa bombylans – Sennertia leei
Xylocopa calens – Sennertia elseni
Xylocopa californica - Sennertia lucrosa
Xylocopa californica arizonensis - Sennertia segnis
Xylocopa clarionensis - Sennertia lucrosa
Xylocopa cubaecola – Sennertia pirata
Xylocopa dejeanii - Sennertia ratiocinator
Xylocopa fallax - Sennertia horrida
Xylocopa fimbriata - Sennertia argentina, S. donaldi
Xylocopa frontalis – Sennertia argentina, S. donaldi
Xylocopa guatemalensis - Sennertia lucrosa
Xylocopa imitator - Sennertia tanythrix
Xylocopa inconstans – Sennertia leclercqi
Xylocopa iridipennis - Sennertia horrida
Acarorum Catalogus IX 291
Xylocopa latipes - Sennertia hipposideros
Xylocopa lepeletieri - Sennertia jeanalexi
Xylocopa mcgregori - Sennertia horrida
Xylocopa mexicanorum – Sennertia argentina, S. lucrosa
Xylocopa micans – Sennertia loricata
Xylocopa mimetica - Sennertia horrida
Xylocopa mirabilis - Sennertia madagascarensis
Xylocopa mordax – Sennertia loricata, S. pirata
Xylocopa muscaria – Sennertia loricata
Xylocopa nasalis – Sennertia dissimilis, S. horrida, S. oudemansi
Xylocopa nautlana – Sennertia argentina
Xylocopa nigrita - Sennertia jeanalexi, S. perturbans, S. simplex, S. tanythrix
Xylocopa nitidiventris - Sennertia cerambycina
Xylocopa olivieri - Sennertia zhelochovtsevi
Xylocopa przewalskyi - Sennertia cerambycina
Xylocopa punctilabris - Sennertia gargantua
Xylocopa rufa - Sennertia zhelochovtsevi
Xylocopa scioensis (= Mesotrichia bryorum) – Sennertia queenslandica
Xylocopa sonorina - Sennertia shimanukii
Xylocopa splendidula splendidula Xylocopa tenuiscapa - Sennertia greeni, S. hipposideros, S. koptorthosomae, S. vanderhammeni
Xylocopa torrida - Sennertia tanythrix
Xylocopa tranquebarorum – Sennertia potanini
Xylocopa turanica - Sennertia cerambycina
Xylocopa valga – Sennertia cerambycina, S. gargantua
Xylocopa varentzowi – Sennertia cerambycina
Xylocopa varipuncta - Sennertia lucrosa
Xylocopa violacea – Sennertia cerambycina
Subfam. Apinae
Apis mellifera – Chaetodactylus claviger, Sennertia faini
Melitoma segmentaria – Chaetodactylus melitomae
Tetrapedia diversipes – Roubikia panamensis
T. maura - Roubikia officiosa
T. peckoltii - Roubikia panamensis
Tetrapedia sp. – Roubikia imberba, R. latebrosa
Fam. Megachilidae
Chelostoma florisomne – Chaetodactylus birulai
Hoplitis pilosifrons - Chaetodactylus hopliti
H. producta - Chaetodactylus hopliti
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Acarorum Catalogus IX
Lithurgus antilleorum – Chaetodactylus antillarum
L. apicalis – Chaetodactylus kouboy, Ch. lithurgi
L. atratus – Chaetodactylus ludwigi
L. echinocacti – Chaetodactylus abditus
L. gibbosus – Chaetodactylus gibbosi, Ch. lithurgi
L. littoralis - Chaetodactylus lithurgi
L. planifrons – Chaetodactylus abditus
L. scabrosus - Chaetodactylus ludwigi
Megachile bombycina – Chaetodactylus dementjevi
M. liguseca - Chaetodactylus dementjevi
Megachile sp. – Chaetodactylus osmiae
Osmia aurulenta – Chaetodactylus chrysidis
O. azteca – Chaetodactylus azteca
O. bicolor – Chaetodactylus chrysidis
O. bicornis – Chaetodactylus osmiae
O. brevicornis – Chaetodactylus reaumuri
O. brevis - Chaetodactylus hopliti
O. bucephala - Chaetodactylus hopliti
O. cornifrons – Chaetodactylus hirashimai, Ch. krombeini, Ch. nipponicus
O. cornuta – Chaetodactylus osmiae, Ch. zachvatkini
O. excavata - Chaetodactylus hirashimai, Ch. nipponicus
O. fronticornis – Chaetodactylus osmiae
O. georgica – Chaetodactylus rozeni
O. imaii - Chaetodactylus hirashimai
O. inermis - Chaetodactylus hopliti
O. kohlii – Chaetodactylus osmiae
O. latreillei – Chaetodactylus zachvatkini
O. leucogastra – Chaetodactylus reaumuri
O. niveata - Chaetodactylus osmiae, Ch. zachvatkini
O. pedicornis - Chaetodactylus hirashimai, Ch. nipponicus
O. rubriflorus – Chaetodactylus claudus
O. rufa – Chaetodactylus osmiae
O. rufiventris – Chaetodactylus osmiae, Ch. reaumuri
O. subaustralis – Chaetodactylus micheneri
O. taurus - Chaetodactylus nipponicus
O. tricornis - Chaetodactylus claviger, Ch. osmiae, Ch. zachvatkini
Rhodanthidium sticticum – Chaetodactylus anthidii
Trichorhurgus dubius - Chaetodactylus lassulus
Fam. Chrysididae
Chrysis coerulans – Chaetodactylus rozeni
Chrysis trimaculata – Chaetodactylus chrysidis
Fam. Vespidae
Acarorum Catalogus IX 293
Vespula germanica – Chaetodactylus dementjevi
Fam. Sphecidae
Isodontia mexicana – Chaetodactylus hopliti
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The Mite Family Hyadesiidae
Hyadesiidae Halbert, 1915: 53
The mites of the genera Hyadesia and Amhyadesia (total of 47 species, see Proches and
Marshall, 2001), inhabit marine intertidal to littoral habitats, occurring on all coastlines and often associated with barnacles, algal mats. The Hyadesiids feed primarily on
marine algae, but other organic debris may also be consumed. (Pugh & King, 1985a).
Deutonymphs are never formed in the family, and dispersal is accomplished by feeding
forms. The start of the studies of Hyadesiidae belongs to Mégnin (1891), who described
the genus Hyadesia, to Michael (1893) and to Halbert (1915) – the author of the family.
Major taxonomic contributions are due also to Fain, Fain & Schuster, Fain & Ganning,
Marshall & Nunkumar, Luxton, Lohmann, Marshall & Ugrasen, Viets, Womersley and
others. Important biological data are due to Schuster, Pugh, Matshall and other researchers of coastal habitats.
Composition of Hyadesiidae
Family Hyadesiidae Halbert, 1915–2 gen., 47 sp.
Amhyadesia Fain et Schuster, 1984–15 sp.
Hyadesia Mégnin, 1891–32 sp.
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Catalogue of Hyadesiidae
Fam. Hyadesiidae Halbert
Hyadesidae Halbert, 1915: 53
Hyadesiidae: Fain, 1981: 47
Lentungulidae Berlese, 1897
Type genus: Hyadesia Mégnin, 1889
Genus Amhyadesia Fain et Schuster
Amhyadesia Fain & Schuster, 1984: 15
Type species: Amhyadesia californica Fain et Schuster, 1984
Amhyadesia atlantica Fain et Schuster
Amhyadesia atlantica Fain & Schuster, 1983: 420
Type locality: Penhurst Park, Main Island
Distribution: Bermudas
Amhyadesia austrafricana Marshall et Nunkumar
Amhyadesia austrafricana Marshall & Nunkumar, 1999: 173
Type locality: Sardinia Bay near Port Elizabeth
Distribution: South Africa
Amhyadesia bartschae Luxton
Amhyadesia bartschae Luxton, 1992: 229
Type locality: Hong Kong
Distribution: China
Amhyadesia bermudana Fain et Schuster
Amhyadesia bermudana Fain & Schuster, 1983: 418
Type locality: Crawl Hill, Main Island
Distribution: Bermudas
Amhyadesia brasiliensis Fain et Schuster
Amhyadesia brasiliensis Fain & Schuster, 1984b: 274
Type locality: south of São Sebastião (São Paulo State)
Distribution: Brazil
Amhyadesia bursaria Fain et Schuster
Amhyadesia bursaria Fain & Schuster, 1984a: 106
Type locality: Noumea, between Baie des Citrons and Anse Vata
Acarorum Catalogus IX 305
Distribution: New Caledonia
Amhyadesia californica Fain et Ganning
Amhyadesia californica Fain & Ganning, 1979: 139
Type locality: California
Distribution: USA
Amhyadesia costaricensis Fain et Schuster
Amhyadesia costaricensis Fain & Schuster, 1984: 271
Type locality: Pacific coast, northern part of the Nat. park “Manuel Antonio”, Quepos
Distribution: Costa Rica
Amhyadesia elizabethensis Marshall et Nunkumar
Amhyadesia elizabethensis Marshall & Nunkumar, 1999: 176
Type locality: Sardinia Bay near Port Elizabeth
Distribution: South Africa
Amhyadesia heteromorpha Luxton
Amhyadesia heteromorpha Luxton, 1992a: 232
Type locality: Hong Kong
Distribution: China
Amhyadesia heterophallus Fain et Schuster
Amhyadesia heterophallus Fain & Schuster, 1984a: 103; Marshall & Nunkumar, 1999: 177
Type locality: Villingili Island, Male Atoll
Distribution: Maledives, Philippines, Mozambique, South Africa
Amhyadesia longipillis Fain et Schuster
Amhyadesia longipillis Fain & Schuster, 1984a: 109
Type locality: Pang-an Island
Distribution: Philippines
Amhyadesia madeirensis Fain et Schuster
Amhyadesia madeirensis Fain & Schuster, 1986: 4
Type locality: East of Caniçal
Distribution: Madeira (incl. Porto Santo)
Amhyadesia pacifica Fain et Schuster
Amhyadesia pacifica Fain & Schuster, 1984a: 109
Type locality: Coconut Island, Camotes Sea
Distribution: Philippines
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Amhyadesia punctulata Luxton
Amhyadesia punctulata Luxton, 1989a: 73; 1990: 374
Type locality: Auckland
Distribution: New Zealand
Genus Hyadesia Mégnin
Hyadesia Mégnin, 1891: 53
Lentungula Michael, 1893: 262
Type species: Hyadesia uncinifer Mégnin, 1891
Subgenus Hyadesia Mégnin
Hyadesia Mégnin, 1891: 53; made subgenus by Fain, 1981
Parahyadesia (unavailable name? No type species fixation)
Type species: Hyadesia uncinifer Mégnin, 1891
Hyadesia (H.) agulhensis Marshall et Ugrasen
Hyadesia (Parahyadesia) agulhensis Marshall & Ugrasen, 2000: 88
Type locality: South Africa
Distribution: South Africa
Hyadesia (H.) alexi Marshall et Chown
Hyadesia alexi Marshall & Chown, 2003: 120
Type locality: Gough Island
Distribution: Gough Island (nr. Tristan da Cunha)
Hyadesia (H.) algivorans (Michael)
Lentungula algivorans Michael, 1893: 262
Hyadesia algivorans (Michael): Turk, 1953: 82; Fain, 1981: 55
Type locality: Cornwall, Land’s End
Distribution: Great Britain,
Hyadesia (H.) arabica Fain et Schuster
Hyadesia (H.) arabica Fain & Schuster, 1985: 68
Type locality: Red Sea at Al Ghardaqa
Distribution: Egypt
Hyadesia (H.) australiana Fain et Synnot
Hyadesia australiana Fain & Synnot, 1981: 195
Type locality: Gunnamatta Beach, Mornington Peninsula, nr. Melbourne, Victoria
Distribution: Australia (Victoria)
Acarorum Catalogus IX 307
Hyadesia (H.) benguelensis Marshall et Ugrasen
Hyadesia (Parahyadesia) benguelensis Marshall & Ugrasen, 2000
Type locality: Angola
Distribution: Angola, South Africa
Hyadesia (H.) curassaviensis Viets
Hyadesia curassaviensis Viets, 1936: 426; Fain, 1981: 53
Type locality: Curaçao, Riftwal nr. Willemstad
Distribution: Curaçao
Hyadesia (H.) fusca (Lohmann)
Lentungula fusca Lohmann, 1894: 85; Canestrini & Kramer, 1899: 136; Halbert, 1915: 108
Hyadesia fusca (Lohmann): Halbert, 1920: 137; Willmann, 1937: 167; Sellnick, 1949: 127;
Turk, 1953: 82
Hyadesia (H.) fusca (Lohmann): Fain & Ganning, 1989: 163
Hyadesia furcillipes Benard, 1961: 81. Type: France (syn. by Fain, 1981)
Type locality: Helgoland Island (Germany)
Distribution: Belgium, France, Germany, Great Britain, Finland, Holland, Ireland, Norway, Sweden, USA
Hyadesia (H.) glynni Manson
Hyadesia glynni Manson, 1963: 163
Type locality: Hopkins Marine Station, Pacific Grove, California
Distribution: USA (California)
Hyadesia (H.) halophila Fain
Hyadesia (Hyadesia) halophila Fain, 1974: 699
Type locality: Kerguelen Island
Distribution: Kerguelen Island
Hyadesia (H.) heteromorpha Marshall et Ugrasen
Hyadesia (Parahyadesia) heteromorpha Marshall & Ugrasen, 2000: 87
Type locality: South Africa
Distribution: South Africa
Hyadesia (H.) maxima Fain, Sømme et Block
Hyadesia maxima Fain, Sømme & Block, 1983: 171
Type locality: South Georgia, Maiviken and Sooty Bluff
Distribution: South Georgia
308
Acarorum Catalogus IX
Hyadesia (H.) microseta Luxton
Hyadesia (H.) microseta Luxton, 1989a: 81; 1990: 376
Type locality: Auckland
Distribution: New Zealand
Hyadesia (H.) mollis Luxton
Hyadesia (Parahyadesia) mollis Luxton, 1989a: 81; 1990: 378
Type locality: Auckland
Distribution: New Zealand (Auckland)
Hyadesia (H.) nearctica Fain et Ganning
Hyadesia (H.) nearctica Fain & Ganning, 1978: 215
Type locality: from Saint John, New Brunswick, Canada throughout the coast of Main,
New Hampshire to Cape Ann, Massachusetts, USA
Distribution: Canada, USA
Hyadesia (H.) pakistanensis Fain et Schuster
Hyadesia pakistanensis Fain & Schuster, 1985: 73
Type locality: Paradise Point near Karachi
Distribution: Pakistan
Hyadesia (H.) paulensis Fain
Hyadesia (H.) paulensis Fain, 1975b: 157
Type locality: St. Paul Is.
Distribution: St. Paul Is.
Hyadesia (H.) plicata Luxton
Hyadesia (Parahyadesia) plicata Luxton, 1989a: 84; 1990: 378
Type locality: Chatham Islands
Distribution: New Zealand (Chatham Islands)
Hyadesia (H.) reticulata Luxton
Hyadesia (Parahyadesia) reticulata Luxton, 1989a: 88; 1990: 378
Type locality: Kaikoura
Distribution: New Zealand
Hyadesia (H.) sanjuanensis Fain et Ganning
Hyadesia (H.) sanjuanensis Fain & Ganning, 1989: 162
Type locality: San Juan Archip., Friday Harbor Marine Lab.
Acarorum Catalogus IX 309
Distribution: USA (State of Washington)
Hyadesia (H.) sellai Viets
Hyadesia sellai Viets, 1937: 3; Fain, 1981: 54
Type locality: Rovinj, Split
Distribution: Croatia
Hyadesia (H.) subantarctica Fain
Hyadesia (Hyadesia) subantarctica Fain, 1974: 702
Type locality: Kerguelen Island
Distribution: Kerguelen Island
Hyadesia (H.) tenuipilis Fain
Hyadesia fusca tenuipilis Fain, 1979: 131
Type locality: Abo
Distribution: Finland
Hyadesia (H.) tesselata Luxton
Hyadesia (Parahyadesia) tesselata Luxton, 1989a: 91; 1990: 378
Type locality: Dunedin
Distribution: New Zealand
Hyadesia (H.) travei Fain
Hyadesia (H.) travei Fain, 1975b: 153
Type locality: Saint-Paul Isl.
Distribution: Saint-Paul and New Amsterdam Islands
Hyadesia (H.) tumida Benard
Hyadesia tumida Benard, 1961: 71
Type locality: Roscoff, Bloscon (Brittany), Cotentin (Normandy)
Distribution: France
Hyadesia (H.) uncinifer Mégnin
Hyadesia uncinifer Mégnin, 1889: 51
Type locality: Tierra del Fuego
Distribution: Tierra del Fuego
Hyadesia (H.) verrucosa Fain et Schuster
Hyadesia verrucosa Fain & Schuster, 1985: 70
Type locality: Rovinj
Distribution: Croatia
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Acarorum Catalogus IX
Hyadesia (H.) vietsi Womersley
Hyadesia vietsi Womersley, 1961: 204; Fain, 1979: 127
Type locality: Biak Is.
Distribution: Indonesia (Papua)
Hyadesia (H.) womersleyi Luxton
Hyadesia womersleyi Luxton, 1989b: 407
Type locality: Macquarie Island
Distribution: Macquarie Island
Hyadesia (H.) zelandica Luxton
Hyadesia zelandica Luxton, 1984: 293; 1990: 376
Type locality: Auckland
Distribution: New Zealand, Auckland
Subgenus Hyadesiella Fain
Hyadesiella Fain, 1974: 693
Type species: Hyadesia kerguelensis Lohmann, 1907
Hyadesia (Hyadesiella) kerguelensis Lohmann
Hyadesia kerguelensis Lohmann, 1907: 369
Hyadesia (Hyadesiella) kerguelensis Lohmann: Fain, 1974: 693
Type locality: Kerguelen Island
Distribution: Kerguelen Island
Index of Hyadesiidae
(Names in bold are accepted as valid)
agulhensis, Hyadesia 306
alexi, Hyadesia 306
algivorans, Hyadesia 306
algivorans, Lentungula 306
Amhyadesia 304
arabica, Hyadesia 306
atlantica, Amhyadesia 304
austrafricana, Amhyadesia 304
australiana, Hyadesia 306
bartschae, Amhyadesia 304
benguelensis, Hyadesia 307
bermudana, Amhyadesia 304
brasiliensis, Amhyadesia 304
bursaria, Amhyadesiа 304
californica, Amhyadesia 305
costaricensis, Amhyadesia 305
curassаviensis, Hyadesia 307
elizabethensis, Amhyadesia 305
furcillipes, Hyadesia 307
fusca, Hyadesia 307
fusca, Lentungula 307
glynni, Hyadesia 307
halophila, Hyadesia 307
heteromorpha, Amhyadesia 305
Acarorum Catalogus IX 311
heteromorpha, Hyadesia 307
heterophallus, Amhyadesia 305
Hyadesia 306
Hyadesiella 310
Hyadesiidae 304
Hyadesidae 304
kerguelensis, Hyadesia 310
Lentungula 306
Lentungulidae 304
longipilis, Amhyadesia 305
madeirensis, Amhyadesia 305
maxima, Hyadesia 307
microseta, Hyadesia 308
mollis, Hyadesia 308
nearctica, Hyadesia 308
pacifica, Amhyadesia 305
pakistanensis, Hyadesia 308
Parahyadesia 306
paulensis, Hyadesia 308
plicata, Hyadesia 308
punctulata, Amhyadesia 306
reticulata, Hyadesia 308
sanjuanensis, Hyadesia 308
sellai, Hyadesia 309
subantarctica, Hyadesia 309
tenuipilis, Hyadesia 309
tenuipilis, Hyadesia fusca 309
tesselata, Hyadesia 309
travei, Hyadesia 309
tumida, Hyadesia 309
uncinifer, Hyadesia 309
verrucosa, Hyadesia 309
vietsi, Hyadesia 309
womersleyi, Hyadesia 310
zelandica, Hyadesia 310
Distribution of Hyadesiidae
EUROPE
Belgium (Fain, 1979) – Hyadesia fusca
Croatia (Viets, 1937; Fain & Schuster, 1985) – Hyadesia (H.) sellai, H. (H.) verrucosa
Finland (Fain, 1979) – Hyadesia fusca, H. tenuipilis
France (contin.) (Benard, 1961) – Hyadesia fusca (= furcillipes), H. tumida
Germany (Lohmann, 1894; Willmann, 1937) – Hyadesia fusca
Great Britain (Michael, 1893; Turk, 1953) – Hyadesia algivorans, H. fusca
Holland (Oudemans, 1927; Pugh & King, 1985) – Hyadesia fusca
Ireland (Halbert, 1920) – Hyadesia fusca
Madeira (Fain & Schuster, 1986) – Amhyadesia madeirensis, Hyadesia (H.) verrucosa
Sweden (Sellnick, 1949; Ganning, 1970) – Hyadesia fusca
AFRICA
Angola (Marshall & Ugrasen, 2000) - Hyadesia benguelensis
Egypt (Fain & Schuster, 1985) – Hyadesia (H.) arabica
South Africa (Marshall & Nunkumar, 1999; Marshall & Ugrasen, 2000) – Amhyadesia
austrafricana, A. elizabethensis, Hyadesia agulhensis, H. benguelensis
ASIA
China (Luxton, 1992a) – Amhyadesia bartschae, A. heteromorpha [Hong Kong]
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Indonesia (Womersley, 1961) – Hyadesia vietsi [Papua, Biak Isl.]
Maledive Islands (Fain & Schuster, 1984a) - Amhyadesia heterophallus
Pakistan (Fain & Schuster, 1985) – Hyadesia (H.) pakistanensis
Philippines (Talker, Alberti & Sotto, 1981; Fain & Schuster, 1984a) - Amhyadesia heterophallus, A. longipilis, A. pacifica
NORTH AMERICA
Bermuda Islands (Fain & Ganning, 1989; Fain & Schuster, 1983) – Amhyadesia atlantica,
A. bermudana
Canada (Fain & Ganning, 1978, 1979, 1989) – Hyadesia nearctica
USA (Manson, 1963; Fain & Ganning, 1979, 1989) – Amhyadesia californica, A. glynni,
Hyadesia fusca, H. sanjuanensis, H. nearctica
SOUTH and CENTRAL AMERICA
Argentina (Megnin, 1891) – Hyadesia uncinifer (Tierra del Fuego)
Brazil (Fain & Schuster, 1984b) – Amhyadesia brasiliensis
Costa Rica (Fain & Schuster, 1984b) – Amhyadesia costaricensis
Curaçao (Viets, 1936) – Hyadesia curassaviensis
ATLANTIC OCEAN
Gough Island (Marshall & Chown, 2003) – Hyadesia alexi
Saint Paul and New Amsterdam Islands (Fain, 1975a, b) – Hyadesia paulensis, H. travei
OCEANIA
Australia (continent) (Fain & Synnot, 1981) – Hyadesia australiana
New Zealand (Luxton, 1984, 1989, 1990) – Amhyadesia punctulata, Hyadesia microseta,
H. mollis, H. reticulata, H. tesselata, H. zelandica
New Caledonia (Fain & Schuster, 1984a) - Amhyadesia bursaria
ANTARCTICA
South Georgia (Fain, Sømme & Block, 1983) – Hyadesia maxima
Kerguelen Island (Lohmann, 1907; Fain, 1974) – Hyadesia halophila, H. subantarctica,
H. (Hyadesialla) kerguelensis
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Acarorum Catalogus IX 313
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The Mite Family Algophagidae Fain
Algophaginae Fain, 1974: 703
Algophagidae Fain, 1981: 49
The small family Algophagidae (eight genera, 17 species) was proposed by A. Fain, first
as subfamily (1974), than as family (1981). The first genera (Hericia G. Canestrini, 1888,
Fusohericia Vitzthum, 1931, Algophagus Hughes, 1955) and species have been described
much earlier, but only with the intensifying of the Antarctic and subantarctic research
many new taxa (mainly of Algophaginae) were discovered by Fain & Johnston (1975),
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Hughes & Goodman (1969), Cruz & Socarras (1992 - Cuba), Fashing, OConnor & Kitching (2000), Marshall, OConnor & Pugh (2001, 2003), and others.
These peculiar mites are aquatic, with various species inhabiting sap flows on trees
(Hericia, Fusohericia), water filled tree holes {Lamingtonacarus), rivers and lakes (Algophagopsis), or other aquatic or subaquatic habitats (O’Connor, 2009) up to 100 m (Anderson
& Krantz, 1993). Species of Algophagus and Neohyadesia are generally restricted to subantarctic islands. They range from coastal brackish pools to inland tussock grass soils,
where they feed primarily on algae (Goddard 1982; Marshall, OConnor & Pugh, 2003,
generalized by O’Connor, 2009). Genital papillae are strongly reduced in Algophagidae,
but the uniquely developed axillary organs between legs I—II characterize the family.
The sap inhabiting taxa of the algophagid subfamily Hericiinae form deutonymphs
for dispersal on sap frequenting Coleoptera (Nitidulidae, Nosodendridae) and Lepidoptera (O’Connor, 2009). The algophagine genus Lamingtonacarus oreillyorum is known to
form deutonymphs, but these have not been collected from any insect host. OConnor
and Moser (1985) proposed generic phylogenetic analysis.
Composition of Algophagidae
Fam. Algophagidae Fain, 1981–9 gen., 24 sp.
Subfam. Algophaginae Fain, 1974–14 sp.
Algophagopsis Fain et Johnston, 1975–1 sp.
Algophagus Hughes, 1955–6 sp.
Lamingtonacarus Fashing, OConnor et Kitching, 2000–2 sp.
Neohyadesia Hughes et Goodman, 1969–3 sp.
Terraphagus Clark et Andrews, 2012–1 sp.
Thalassophagacarus J. de la Cruz et Socarras, 1992–1 sp.
Subfam. Hericiinae OConnor et Moser, 1985–10 sp.
Fusohericia Vitzthum, 1931–3 sp.
Hericia G. Canestrini, 1888–6 sp.
Prohericia OConnor et Moser, 1985–1 sp.
Catalogue of Algophagidae
Fam. Algophagidae Fain
Algophaginae Fain, 1974: 703
Algophagidae Fain: Fain, 1981: 49
Type genus: Algophagus Hughes, 1955
Subfam. Algophaginae Fain
Algophaginae Fain, 1974: 703
Acarorum Catalogus IX 317
Type genus: Algophagus Hughes, 1955
Genus Algophagopsis Fain et Johnston
Algophagopsis Fain & Johnston, 1975: 67
Type species: Algophagopsis pneumatica Fain et Johnston, 1975
Algophagopsis pneumatica Fain et Johnston
Algophagopsis pneumatica Fain & Johnston, 1975: 67
Type locality: Kings River, California
Distribution: USA (California)
Genus Algophagus Hughes
Algophagus Hughes, 1955: 1
Type species: Algophagus antarcticus Hughes, 1955
Algophagus antarcticus Hughes
Algophagus antarcticus Hughes, 1955: 1; OConnor & Pugh, 2003: 39
Type locality: Heard Island
Distribution: Heard Island, Kerguelen Island
Algophagus brachytarsus OConnor et Pugh
Algophagus brachytarsus OConnor & Pugh, 2003: 41
Type locality: Prince Edward Islands
Distribution: Prince Edward Islands
Algophagus laticollaris Fain
Algophagus antarcticus laticollaris Fain, 1974: 704
Algophagus laticollaris Fain: OConnor & Pugh, 2003: 39
Type locality: Kerguelen I.
Distribution: Kerguelen I.
Algophagus macrolithus OConnor et Pugh
Algophagus macrolithus OConnor & Pugh, 2003: 43
Type locality: Prince Edward Islands
Distribution: Prince Edward Islands
Algophagus pennsylvanicus Fashing et Wiseman
Algophagus pennsylvanicus Fashing & Wiseman, 1980: 79
Type locality: Cook Forest State Park, Forest County, Pennsylvania
Distribution: USA
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Acarorum Catalogus IX
Algophagus semicollaris Fain
Algophagus semicollaris Fain, 1974: 704; Fedorova, Fedorov, Stavsky, Matyukhina & Voronin 1980: 82
Type locality: Kerguelen Island
Distribution: Kerguelen Island
Genus Lamingtonacarus Fashing, OConnor et Kitching
Lamingtonacarus Fashing, OConnor & Kitching, 2000: 592
Type species: Lamingtonacarus oreillyorum Fashing, OConnor et Kitching, 2000
Lamingtonacarus oreillyorum Fashing, OConnor et Kitching
Lamingtonacarus oreillyorum Fashing, OConnor & Kitching, 2000: 592
Type locality: Lamington National Park, Box Forest; Queensland
Distribution: Australia (Queensland)
Lamingtonacarus posidonis Fashing, OConnor et Kitching
Lamingtonacarus posidonis Fashing, OConnor & Kitching, 2000: 599
Type locality: Lamington National Park, Box Forest; Queensland
Distribution: Australia (Queensland)
Genus Neohyadesia Hughes et Goodman
Neohyadesia Hughes & Goodman, 1969: 39
Type species: Neohyadesia signyi Hughes et Goodman, 1969
Neohyadesia microtricha Marshall, OConnor et Pugh
Neohyadesia microtricha Marshall, OConnor & Pugh, 2001: 101
Type locality: Marion Island (Prince Edward Islands, South Georgia, Trypot beach)
Distribution: Indian Ocean islands
Neohyadesia minor Clark et Andrews
Neohyadesia minor Clark & Andrews, 2012: 1478
Type locality: Lake Forsyth/Wairewa, Canterbury
Distribution: New Zealand
Neohyadesia signyi Hughes et Goodman
Neohyadesia signyi Hughes & Goodman, 1969: 39
Type locality: Signy Island
Distribution: South Orkney Islands
Neohyadesia signyi punctulata Fain
Neohyadesia signyi punctulata Fain, 1974: 706
Acarorum Catalogus IX 319
Type locality: Kerguelen Island
Distribution: Kerguelen Island
Genus Terraphagus Clark et Andrews
Terraphagus Clark & Andrews, 2012: 1476
Type species: Terraphagus antipodus Clark et Andrews, 2012
Terraphagus antipodus Clark et Andrews
Terraphagus antipodus Clark & Andrews, 2012: 1476
Type locality: Antipodes Islands
Distribution: Antipodes Islands
Host: from grey petrel Procellaria cinerea burrow soil
Genus Thalassophagacarus J. de la Cruz et Socarras
Thalassophagacarus J. de la Cruz & Socarras, 1992: 1
Type species: Thalassophagacarus faime J. de la Cruz et Socarras, 1992
Thalassophagacarus faime J. de la Cruz et Socarras
Thalassophagacarus faime J. de la Cruz & Socarras, 1992: 2
Type locality: “playa cercana al Faro Roncali, Cabo de San Antonio, Pinar del Rio”
Distribution: Cuba
Subfam. Hericiinae OConnor et Moser
Hericiinae OConnor & Moser, 1985: 786
Type genus: Hericia G. Canestrini, 1888
Genus Fusohericia Vitzthum
Fusohericia Vitzthum, 1931: 98
Type species: Fusohericia incredibilis Vitzthum, 1931
Fusohericia incredibilis Vitzthum
Fusohericia incredibilis Vitzthum, 1931: 98
Type locality: South Sumatra
Distribution: Indonesia (Sumatra)
Fusohericia heliconiae Fashing et Glist
Fusohericia heliconiae Fashing & Glist, 2012: 591
Type locality: La Selva Biological Station and Reserve, Heredia Province
Distribution: Costa Rica
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Fusohericia lawrencei Baker et Crossley
Fusohericia lawrencei Baker & Crossley, 1964: 4
Type locality: treehole, trunk of yellow poplar, Liriodendron tulipijera, Roane County,
Tennessee
Distribution: USA (Tennessee)
Genus Hericia G.Canestrini
Hericia G. Canestrini, 1888: 6
Type species: Glyciphagus hericius Robin, 1868
Hericia fermentationis Vitzthum
Hericia fermentationis Vitzthum, 1931: 85
Type locality: Sumatra
Distribution: Indonesia (Sumatra)
Hericia georgei Michael
Hericia georgei Michael, 1903: 38; Samšinák, 1972: 65; Haitlinger, 2002: 68; Fashing, 2008: 5
Type locality: Great Britain
Distribution: Great Britain, Finland, France, Poland, Russia
Hosts: Cetonia aurata, Potosia cuprea metallica (Coleoptera: Scarabaeidae), Catocala
sponsa, C. nupta, C. adultera, C. elocata, C. neonympha, C. promissa, C. pacta, C.
fraxini (Lepidoptera: Noctuidae), Cossus cossus (Lep.: Cossidae),
Nymphalis antiopa, N. atalanta, Polygonia c-album, Acronicta [Apatele] psi, Europis occulta
(Nymphalidae), Scoliopteryx libatrix (Erebidae)
Hericia hericia (Robin)
Glyciphagus hericius Robin, 1868: 607
Hericia robini Michael, 1903 (synonymy 1868–1903)
Hericia hericia (Robin): Zachvatkin, 1941
Type locality: France
Distribution: England, France, Germany, Italy
Hericia janehenleyi Fashing
Hericia janehenleyi Fashing, 2008: 7
Type species: Newport News, Eastern Virginia
Distribution: USA
Host: Glischrochilus obtusus (Coleoptera, Nitidulidae).
Hericia paradoxa Türk et Türk
Hericia (?) paradoxa Türk & Türk, 1957: 180
Type species: Erlangen
Distribution: Germany
Acarorum Catalogus IX 321
Hericia sanukiensis Fashing
Hericia sanukiensis Fashing & Okabe, 2006: 3
Type locality: Takamatsu City, Kagawa Prefecture
Distribution: Japan
Host: Librodor japonicus (Coleoptera: Nitidulidae)
Genus Prohericia OConnor et Moser
Prohericia OConnor & Moser, 1985: 783
Type species: Prohericia longipes OConnor et Moser, 1985
Prohericia longipes OConnor et Moser
Prohericia longipes OConnor & Moser, 1985: 783
Type locality: Germany
Distribution: Germany
Index of Algophagidae
(Names in bold are accepted as valid)
Algophagidae 316
Algophaginae 316
Algophagopsis 317
Algophagus 317
antarcticus, Algophagus 317
antipodus, Terraphagus 319
brachytarsus, Algophagus 317
faime, Thalassophagacarus 319
fermentationis, Hericia 320
Fusohericia 319
georgei, Hericia 320
heliconiae, Fusohericia 319
Hericia 320
hericia, Hericia 320
Hericiinae 319
hericius, Glyciphagus 320
incredibilis, Fusohericia 319
janehenleyi, Hericia 320
Lamingtonacarus 318
laticollaris, Algophagus 317
laticollaris, Algophagus antarcticus 317
lawrencei, Fusohericia 320
longipes, Prohericia 321
macrolithus, Algophagus 317
microtricha, Neohyadesia 318
minor, Neohyadesia 318
Neohyadesia 318
oreillyorum, Lamingtonacarus 318
paradoxa, Hericia 320
pennsylvanicus, Algophagus 317
pneumatica, Algophagopsis 317
posidonis, Lamingtonacarus 318
Prohericia 321
punctulata, Neohyadesia signyi 318
robini, Hericia 320
sanukiensis, Hericia 321
semicollaris, Algophagus 318
signyi, Neohyadesia 318
Terraphagus 319
Thalassophagacarus 319
322
Acarorum Catalogus IX
Distribution of Algophagidae
EUROPE
Finland (Samšinák, 1972) – Hericia georgei
France (contin.) (Türk & Türk, 1957) – Hericia georgei, H. hericia
Germany (Türk & Türk, 1957; OConnor & Moser, 1985) – Hericia hericia, H. paradoxa,
Prohericia longipes
Great Britain (Michael, 1903) – Hericia georgei, H. hericia
Italy (Bernini et al., 1995) – Hericia hericia
Poland (Haitlinger, 2002, 2004) – Hericia georgei
Russia (all, continent.) (Samšinák, 1972) – Hericia georgei
ASIA
Indonesia (Vitzthum, 1931) - Hericia fermentationis (Sumatra), Fusohericia incredibilis
(Sumatra)
Japan (Fashing & Okabe, 2006; Hayashi, Ichikawa & Yasui, 2010) – Hericia sanukiensis
NORTH AMERICA
USA (Baker & Crossley, 1964; Fain & Johnston, 1975; Fashing, 2008) – Algophagopsis
pneumatica, Hericia janehenleyi, Fusohericia lawrencei
SOUTH and CENTRAL AMERICA
Costa Rica (Fashing, 2012) – Fusohericia heliconiae
Cuba (Cruz & Socarras, 1992) – Thalassophagacarus faime
OCEANIA
Australia (continent) (Fashing, OConnor & Kitching, 2000) – Lamingtonacarus oreillyorum, L. posidonis
New Zealand (Clark & Andrews, 2012) - Neohyadesia minor
Antipodes Islands, New Zealand (Clark & Andrews, 2012) – Terraphagus antipodus
ANTARCTICA
Heard Island (Hughes, 1955) - Algophagus antarcticus
Kerguelen Island (Fain, 1974) - Algophagus antarcticus, A. laticollaris, A. semicollaris,
Neohyadesia signyi punctulata
South Orkney Islands (Hughes & Goodman, 1969) – Neohyadesia signyi
Marion island (Marshall, OConnor & Pugh, 2001) - Neohyadesia microtricha
South Georgia (Marshall, OConnor & Pugh, 2001) - Neohyadesia microtricha
Prince Edward Islands (Marshall et al., 1999; Marshall, OConnor & Pugh, 2003) - Algophagus brachytarsus, A. macrolithus
Acarorum Catalogus IX 323
Hosts of Algophagidae
Insecta
Coleoptera
Fam. Scarabaeidae
Cetonia aurata - Hericia georgei
Potosia cuprea metallica - Hericia georgei
Fam. Nitidulidae
Glischrochilus obtusus – Hericia janehenleyi
Librodor japonicus – Hericia sanukiensis
Lepidoptera
Fam. Noctuidae
Catocala sponsa – Hericia georgei
C. nupta - Hericia georgei
C. adultera - Hericia georgei
C. elocata - Hericia georgei
C. neonympha - Hericia georgei
C. promissa - Hericia georgei
C. pacta - Hericia georgei
C. fraxini - Hericia georgei
Nymphalis antiopa - Hericia georgei
N. atalanta - Hericia georgei
Polygonia c-album - Hericia georgei
Acronicta [Apatele] psi - Hericia georgei
Fam. Nymphalidae
Europis occulta - Hericia georgei
Fam. Erebidae
Scoliopteryx libatrix - Hericia georgei
Bibliography of Algophagidae
Baker E.W., D.A. Crossley. 1964. A new species of mite, Fusohericia lawrencei, from an artificial
tree-hole (Acarina: Glycyphagidae). – Annals of the Natal Museum, 16: 4.
Bernini F., M. Castagnoli, R. Nannelli. 1995. Checklist delle specie fauna d’Italia. Fascicolo
24 Arachnida: Acari. Ed. Calderini.
Canestrini G. 1888. Prospetto dell’Acarofauna Italiana. Famiglie: Tarsonemini e Tiroglofini e
Appendice ogli Analgesini. Vol.3: 309–425.
Clark J.M.; N. Andrews. 2012: New mites Terraphagus antipodus gen. n., sp. n. and Neohyadesia
minor sp. n. (Acarina: Astigmata: Algophagidae) from islands of the Southern Ocean. Polar biology, 35(10): 1473–1485
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Cruz J. de la, A.A. Socarras. 1992. Nuevos genero y especie de acaro litoral (Acarina: Algophagidae) de Cuba. – Poeyana, 412: 1–4. Fain A. 1974. Acariens récoltés par le Dr. J.
Travé aux îles subantarctiques I. Familles Saproglyphidae et Hyadesidae (Astigmates).
– Acarologia, 16(4): 684–708.
Fain A. 1981. Notes on the Hyadesiidae Halbert, 1915 and Algophagidae Fain, 1974, nov. tax.
(Acari, Astigmata), with a redescription of Hyadesia curassaviensis Viets, 1936 and H.
sellai Viets, 1937. - Acarologia, 22(1): 47–61.
Fain A., D. Johnston. 1975. A new algophagid mite Algophagopsis pneumatica n.g., n.sp. living
in a river (Astigmata: Hyadesidae). – Bull. Ann. Soc. r. belge Ent., 111: 67–70.
Fashing N. J. 1991. Deutonymphal dimorphism in the genus Hericia (Astigmata: Algophagidae). In: Dusbabek, F. & Bukva, V. (eds.) Modern Acarology Vol. 2. Prague, Academia
and The Hague, SPB Academic Publishing bv. pp. 287–291.
Fashing N. J. 1992. Observations on the feeding biology of Algophagus pennsylvanicus (Astigmata: Algophagidae), a mite restricted to water-filled treeholes. - International Journal
of Acarology, 18(2): 77–81.
Fashing N.J. 2001. Morphological adaptations associated with mating behaviour in the genus
Hericia (Algophagidae: Astigmata). In R. B. Halliday, D. E. Walter, H. C. Proctor, R. A.
Norton and M. J. Colloff (eds.) Proceedings of the 10th International Congress of Acarology. CSIRO Publishing, Collingwood, Victoria, Australia. pp. 176–179.
Fashing N. J. 2008. Biology of the genus Hericia (Algophagidae: Astigmata), with the description of a new species from the eastern United States. - Systematic and Applied Acarology,
13(1): 3–25.
Fashing N. J. 2016. Fine structure of the axillary organs of Hericia janehenleyi Fashing (Algophagidae: Astigmatina : Sarcoptiformes) with a discussion of a potential correlation
between axillary organ size and habitat. - Journal of the Acarological Society of Japan,
25(S1): 3–17
Fashing N. J., A. Glist. 2012. A new Fusohericia (Acari: Astigmata: Algophagidae) from Costa
Rica. - Systematic and Applied Acarology, 17(3): 591–606.
Fashing N. J., K. Okabe. 2006. Hericia sanukiensis, a new species of Algophagidae (Astigmata)
inhabiting sap flux in Japan. - Systematic and Applied Acarology Special Publications,
22(1): 1–14.
Fashing N. J., L.L. Wiseman. 1980. Algophagus pennsylvanicus a new species of Hyadesiidae
from water-filled treeholes. – Internat. J. Acarology, 6(1): 79–84.
Hayashi K., T. Ichikawa, Y. Yasui. 2010. Life history of the newly discovered Japanese tree
sap mite, Hericia sanukiensis (Acari, Astigmata, Algophagidae). – Exp. Appl. Acarol.,
50(1): 35–49.
Haitlinger R. 2002. Mites (Acarina) associated with Cetoniinae and Trichiinae (Insecta:
Coleoptera: Scarabaeidae) in Poland. – Postępy polskiej acarologii: 63–73.
Haitlinger R. 2004. Mites (Acari) occuring on some Coleoptera (Insecta) in Poland. – Polish
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Hughes A.M. 1955. A new genus and species of hyadesid mites – Algophagus antarcticus from
Heard Island. – A.N.A.R.E. Rep., B1 (Zool.): 1–19.
Hughes A.M., B.J.A. Goodman. 1969. Neohyadesia signyi (Hyadesidae: Acarina): a new genus and
species from Signy Island, South Orkney Islands. - British Antarctic Survey Bulletin, 22: 39–48.
Marshall D. J., N.G.M. Gremmen, L. Coetzee, B.M. OConnor, P.J.A. Pugh, P.D. Theron, E.A.
Ueckermann. 1999. New records of Acari from the sub-antarctic Prince Edward Islands.
– Polar Biology, 21: 84–89.
Marshall D.J., B.M. OConnor, Ph.J.A. Pugh. 2006. Neohyadesia microtricha (Acari: Astigmata:
Algophagidae): A new species from the sub-Antarctic. - Polar Biology, 24(2): 101–104.
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taxonomic descriptions. - Journal of Zoology, 259(01): 31–47
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1903–2: 1–183.
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Anthophoridae: Xylocopinae) with description of a new type of acarinarium. – International Journal of Acarology, 19(2): 159–166.
OConnor M., J.C. Moser. 1985. Phylogenetic relationships of the Algophagidae (Acari:
Astigmata), with descriptions of a new subfamily, genus, and species. - Annals of the
Entomological Society of America, 78(6): 783–789.
Robin M.C. 1868. Sur une espèce nouvelle de Sarcoptides du genre Glyciphage. Journal of
Anatomy and Physiology (Paris), 5: 603–625.
Samšinák K. 1972. Redescription of Hericia georgei Michael, 1903 (Acarina, Tyroglyphydae)
phoretic on Lepidoptera. – Ann. Zool. Fenn., 9: 65–68.
Schenker R. 1986. Biological and ecological notes on Neohyadesia signyi (Acari: Astigmata).
- British Antarctic Survey Bulletin, 70: 85–91.
Türk E., F. Türk. 1957. Systematik und Ökologie der Tyroglyphiden Mitteleuropas in H. J.
Stammer: Beiträge zur Systematik und Ökologie Mitteleuropäischer Acarina. – Akad.
Verlagsges., Leipzig, 1, Teil 1, Abh. 1: 1–231.
Vazquez I., M.G. Lopez-Campos, M. G. 1996. Prostigmatid mites (Prostigmata) from a littoral
zone in Veracruz, Mexico. - International congress of acarology, 575–578.
Vitzthum H. 1931. Terrestrische Acarinen (unter Ausschluße der Oribatiden und Ixodiden)
der Deutschen Limnologischen Sunda-Expedition. - Archiv für Hydrobiologie (Supplement), 9(1): 59–134.
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The Mite Family Hemisarcoptidae
Hemisarcoptidae Oudemans, 1908: 60
Hemisarcoptoid mites are common inhabitants of various arboreal or aquatic habitats,
with many species specialized as inquilines in nests of Hymenoptera.
Linobia coccinellae is ectoparasite (hematophagous) of Chrysomela populi and Ch.
saliceti (Coleoptera: Chrysomelidae). Other Linobia species have been described on the
plant Eichhornia crassipes in Egypt (Soliman, Hussein & Ramadan, 1991). According to
OConnor (1982), “Deutonymphs of Hemisarcoptes are known to be phoretic only on coccinellid beetles of the genus Chilocorus, which feeds exclusively on scale insects ”. Hemisarcoptes species are used for control of scale insects (Coccodea), as they are predaceous
on the eggs and adults of these insects (Gerson, 1967). The peculiar Espeletiacarus andinus
Fain, 1987 is known from the high Andes in Colombia on the flowers of Espeletia incana.
Composition of Hemisarcoptidae
Fam. Hemisarcoptidae - 8 gen., 24 sp.
Divilia Sevastjanov, 1969–3 sp.
Espeletiacarus Fain, 1987–1 sp.
Hemisarcoptes Lignières, 1893–5 sp.
Huronopus OConnor et Houck, 1989–1 sp.
Linobia Berlese, 1884–4 sp.
Nanacarus Oudemans, 1902–8 sp.
Pasohopus OConnor et Houck, 1989–1 sp.
Superioropus OConnor et Houck, 1989–1 sp.
Catalogue of Hemisarcoptidae
Family Hemisarcoptidae Oudemans
Hemisarcoptidae Oudemans, 1908: 60
Nanacaridae Oudemans, 1923: 206
Linobiidae Oudemans, 1908
Type genus: Hemisarcoptes Lignières, 1893
Genus Divilia Sevastjanov
Divilia Sevastjanov, 1969: 447 [Diagnostic, Fain, Hurst, Tweddle, Lachlan, Majerus &
Britt 1995: 103]
Acarorum Catalogus IX 327
Type species: Divilia oculata Sevastjanov, 1969
Divilia exigua Fain, Hurst, Fassotte, Webberley, Sloggett et Majerus
Divilia exigua Fain, Hurst, Fassotte, Webberley, Sloggett & Majerus, 1997: 91
Type locality: Torun
Distribution: Poland
Host: Vibidia duodecimguttata (Coleoptera: Coccinellidae)
Divilia occidentalis Fain, Hurst, Tweddle, Lachlan, Majerus et Britt
Divilia occidentalis Fain, Hurst, Tweddle, Lachlan, Majerus & Britt, 1995: 103
Type locality: England
Distribution: England
Host: Chilocorus renipustulatus (Coleoptera: Coccinellidae, Chilocorinae)
Divilia oculata Sevastjanov
Divilia oculata Sevastjanov, 1969: 447
Type locality: Ukraine
Distribution: Russia, Ukraine
Hosts: Lasius fuliginorus, Formica polyctena, F. rufa (Hymenoptera: Formicidae)
Divillia sp.
Hosts: Chrysops univittatus, Hybomitra epistates, H. frontalis, H. lasiophthalma, H. nuda,
Tabanus quinquevittatus (Canada)
Genus Espeletiacarus Fain
Espeletiacarus Fain, 1987: 37
Type species: Espeletiacarus andinus Fain, 1987
Espeletiacarus andinus Fain
Espeletiacarus andinus Fain, 1987: 38
Type locality: Páramo La Rusia, ca. 200 km NNW of Bogotá, 10 km N of Duitame, alt.
3550 m
Distribution: Colombia
Host: from a flower of Espeletia incana
[Genus Giardius Perraud, 1896; placement? Nanacaridae]
[Giardius vitis Perraud, 1896]
Genus Hemisarcoptes Lignières
Hemisarcoptes Lignières, 1893: 16; [Diagnostic: Fain, 1988: 127]
Subgenus Coleovidia H. A. Thomas
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Vidia (Coleovidia) H. A. Thomas, 1961: 461 [syn. of Hemisarcoptes by Gerson 1967: 637].
Type species: Vidia cooremani H. A. Thomas, 1961
Type species: Acarus malus Shimer, 1868
Hemisarcoptes budensis Fain et Ripka
Hemisarcoptes budensis Fain & Ripka, 1998: 35
Type locality: Budapest
Distribution: Hungary
Hosts: Chilocorus renipustulatus (Coleoptera: Coccinellidae) & Coccoidea (Hemiptera)
Hemisarcoptes coccophagus Meyer
Hemisarcoptes coccophagus Meyer, 1962: 414; Charles, Hill & Allan, 1995: 297; Houk,
1999: 100
Type locality: Letaba, Transvaal, South Africa
Distribution: New Zealand, South Africa
Hemisarcoptes cooremani (Thomas)
Vidia (Coleovidia) cooremani Thomas, 1961: 461
Hemisarcoptes cooremani (Thomas): syn. by Gerson, 1967: 637; Houck & OConnor, 1990: 870
Type locality: Weslaco, Texas
Distribution: USA
Host: Chilocorus cacti (Coleoptera: Coccinellidae)
Hemisarcoptes dzhashii K. Dzhibladze
Hemisarcoptes dzhashii K. Dzhibladze, 1969: 435; Rekk, 1976: 40
Type locality: Zelenyy Cape
Distribution: Georgia
Hemisarcoptes malus (Shimer)
Acarus malus Shimer, 1868: 368
Hemisarcoptes malus (Shimer): André, 1942: 173; Cooreman, 1951: 30; Redescription:
Vacante 1985: 750; Abdelgayed et al., 2019: 32
Hemisarcoptes coccisugus Lignières, 1893: 75
Type locality: Mt. Carroll, Illinois
Distribution: Algeria, Canada, Egypt, Italy, Morocco, USA
Hosts: Chrysomphalus dictyospermi, Lepidosaphes beckii, L. gloveri, Parlatoria blanchardi,
P. zizyphi (Coccodea: Diaspididae)
Genus Huronopus OConnor et Houck
Huronopus OConnor & Houck, 1989b: 6
Type species: Huronopus michiganensis OConnor et Houck, 1989
Acarorum Catalogus IX 329
Huronopus michiganensis OConnor et Houck
Huronopus michiganensis OConnor & Houck, 1989b: 6
Type locality: Michigan
Distribution: USA
Genus Linobia Berlese
Linobia Berlese, 1884: 29
Type species: Acarus coccinellae Scopoli, 1763
Linobia coccinellae (Scopoli)
Acarus coccinellae Scopoli, 1763: 389
Berlese, 1884: 29; Théodoridès, 1955: 71; Haitlinger, 1999: 181; Bernini et al., 1995; Tarasi,
Saboori & Sadeghi, 2001; Trach, 2016; Pohlink, Georgi & Müller, 2020: 723
Type locality: Europe
Distribution: Czech Rep., France, Germany, Great Britain, Iran, Italy, Poland, Ukraine
Hosts: Chrysomela populi, C. saliceti, Leptinotarsa decemlineata (Coleoptera: Chrysomelidae)
Linobia eichhornia Soliman, Hussein et Ramadan
Linobia eichhornia Soliman, Hussein & Ramadan, 1991: 22
Type locality: Egypt
Distribution: Egypt
Host: Eichhornia crassipes (plant)
Linobia nilotica Soliman, Hussein et Ramadan
Linobia nilotica Soliman, Hussein & Ramadan, 1991: 17
Type locality: Egypt
Distribution: Egypt
Host: Eichhornia crassipes (plant)
Linobia replacementica Soliman, Hussein et Ramadan
Linobia replacementica Soliman, Hussein & Ramadan, 1991: 19
Type locality: Egypt
Distribution: Egypt
Host: Eichhornia crassipes (plant)
Genus Nanacarus Oudemans
Nanacarus Oudemans, 1902: 7, 45 [Diagnostic, Fain 1988: 127; Redescription, Fain,
1985: 280]
Froweinia Oudemans, 1923: 183
Congovidiella Fain, 1971: 282] [type is hieroglypha; syn. Fain 1988: 126]
Nanacaroides Volgin & Mironov, 1979: 93 [type is elongatus; syn. Purvis & Evans 1982: 820]
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Acarorum Catalogus IX
Type species: Hypopus minutus Oudemans, 1901
Nanacarus collembolicola (Fain et Johnston)
Congovidiella collembolicola Fain & Johnston, 1974: 418
Nanacarus collembolicola (Fain et Johnston)
Type locality: England, Domleo, Spinney, Sutton Bonington
Distribution: Great Britain
Host: Sminthurus fuscus (Collembola, Symphypleona: Sminthuridae)
Nanacarus elongatus (Volgin et Mironov)
Nanacaroides elongatus Volgin & Mironov, 1979: 93
Nanacarus elongatus (Volgin et Mironov): Purvis & Evans, 1982: 820
Type locality: Habarovskiy Kray, Bolshe-Hehcirskiy Zapovednik (the Far East of Russia)
Distribution: Russia
Nanacarus hieroglypha (Fain et Elsen)
Congovidiella hieroglypha Fain et Elsen, in Fain, 1971: 282
Nanacarus hieroglypha (Fain et Elsen):
Type locality: Haut Lopori
Distribution: D.R. Congo
Host: Glossina fuscipes fuscipes (Diptera: Glossinidae)
Nanacarus hungaricus Halmai et Mahunka
Nanacarus hungaricus Halmai & Mahunka, 1980: 266 [mammalian host]
Type locality: Hungary
Distribution: Hungary
Nanacarus longisetosus Volgin et Mironov
Nanacarus longisetosus Volgin & Mironov, 1979: 91
Type locality: Buryatian ASSR, Gremyachenskiy Rayon, Istok
Distribution: Russia
Nanacarus manus Purvis et Evans
Nanacarus manus Purvis & Evans, 1982: 815
Type locality: S.E. Ireland
Distribution: Ireland
Nanacarus minutus (Oudemans)
Hypopus minutus Oudemans, 1901: 85
Nanacarus minutus (Oudemans): Krivoluckiy, 1975: 474; Fain 1985: 280; Fain et al., 1997:
94; Li et al., 1992: 23
Acarorum Catalogus IX 331
Type locality: Indonesia (Java)
Distribution: China, Germany, Indonesia (Java, Buru), Poland, Russia
Hosts: Adalia sp., Calvia duodecimguttata, Exochomus quadripustulus (Coleoptera: Coccinellidae); Xylocopa (Koptorthosoma) tenuiscapa (Hymenoptera: Apidae, Xylocopinae),
Eptesicus serotinus, Pipistrellus pipistrellus, Sorex araneus (Mammalia), Megapodius
reinwardt buruensis (Aves: Megapodiidae)
Nanacarus nominis Woodring
Nanacarus nominis Woodring, 1966: 82
Type locality: Holden, Utah
Distribution: USA
Host: «from the cells of the alkali bee Nominia melanderi«(Hymenoptera: Halictidae,
Nomininae)
Nanacarus pragensis (Samšinák et Vobrazkova)
Congovidiella pragensis Samšinák & Vobrazkova, 1983: 119
Nanacarus pragensis (Samšinák & Vobrazkova): Khaustov et al., 2018: 48
Type locality: Prague
Distribution: Czech Rep.
Genus Pasohopus OConnor et Houck
Pasohopus OConnor & Houck, 1989a: 17
Type species: Pasohopus labrynthus OConnor et Houck, 1989
Pasohopus labrynthus OConnor et Houck
Pasohopus labrynthus OConnor & Houck, 1989a: 17
Type locality: Pasoh Forest Reserve
Distribution: Malaysia
Genus Superioropus OConnor et Houck
Superioropus OConnor & Houck, 1989b: 2
Type species: Superioropus huronmontanus OConnor & Houck, 1989
Superioropus huronmontanus OConnor et Houck
Superioropus huronmontanus OConnor & Houck, 1989b: 2; OConnor, 1991; Khaustov
et al., 2018: 49
Type locality: the Huron Mountains of northern Michigan
Distribution: Russia, USA
Hosts: Exeristes comstockii (Hymenoptera: Ichneumonidae), galleries of Ips typographus
(Coleoptera: Curculionidae, Scolytinae)
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Index of Hemisarcoptidae
(Names in bold are accepted as valid)
andinus, Espeletiacarus 327
budensis, Hemisarcoptes 328
coccinellae, Acarus 329
coccinellae, Linobia 329
coccisugus, Hemisarcoptes 328
coccophagus, Hemisarcoptes 328
Coleovidia 327
collembolicola, Congovidiella 330
collembolicola, Nanacarus 330
Congovidiella 329
cooremani, Hemisarcoptes 328
cooremani, Vidia (Coleovidia) 328
Divilia 326
dzhashii, Hemisarcoptes 327
eichhornia, Linobia 327
elongatus, Nanacarus 330
elongatus, Nanacaroides 328
Espeletiacarus 326
exigua, Divilia 325
Froweinia 329
Hemisarcoptes 327
Hemisarcoptidae 326
hieroglypha, Congovidiella 330
hieroglypha, Nanacarus 330
hungaricus, Nanacarus 330
huronmontanus, Superioropus 331
Huronopus 328
labrynthus, Pasohopus 331
Linobia 329
Linobiidae 326
longisetosus, Nanacarus 330
malus, Acarus 328
malus, Hemisarcoptes 328
manus, Nanacarus 330
michiganensis, Huronopus 329
minutus, Hypopus 330
minutus, Nanacarus 330
Nanacaridae 326
Nanacaroides 329
Nanacarus 329
nilotica, Linobia 329
nominis, Nanacarus 331
occidentalis, Divilia 327
oculata, Divilia 327
Pasohopus 331
pragensis, Congovidiella 331
pragensis, Nanacarus 331
replacementica, Linobia 329
Superioropus 331
Distribution of Hemisarcoptidae
EUROPE
Belgium (Fain, Hurst et al. 1997) – Nanacarus minutus
Czech Rep. (Samšinak & Vobrazkova, 1983) – Nanacarus pragensis
France (contin.)(Théodoridès, 1955) – Linobia coccinellae
Georgia (Dzhibladze, 1969) – Hemisarcoptes dzhashii
Germany (Pohlink, Georgi & Müller, 2020) – Linobia coccinellae, Nanacarus minutus
Acarorum Catalogus IX 333
Great Britain (Türk & Türk, 1952; Fain & Johnston, 1974; Fain, Hurst, Tweddle et al., 1995;
Fain, Hurst, Fassotte et al. 1997) – Congovidia coccinellidarum, Linobia coccinellae,
Nanacarus (= Congovidiella) collembolicola
Hungary (Halmai & Mahunka, 1980; Fain & Ripka, 1998) – Hemisarcoptes budensis,
Nanacarus hungaricus
Ireland (Purvis & Evans, 1982) – Nanacarus manus
Italy (contin.) (Bernini et al., 1995) – Hemisarcoptes malus, Linobia coccinellae
Sicily (Bernini et al., 1995) – Hemisarcoptes malus
Poland (Fain et al., 1997) – Divilia exigua, Nanacarus minutus
Russia (all, continent.) (Sevastjanov, 1969; Krivoluckiy, 1975: 474; Volgin & Mironov,
1979) – Divilia oculata, Divilia aff. occidentalis, Hemisarcoptes sp., Nanacarus longisetosus, N. elongatus, N. minutus, Superioropus huronmontanus
Ukraine (Sevastjanov, 1969; Trach, 2007) - Divilia oculata, Linobia coccinellae
AFRICA
Congo DR (Fain, 1971) – Nanacarus hieroglypha
Egypt (Soliman, Hussein & Ramadan, 1991; Abdelgayed et al., 2019) – Hemisarcoptes
malus, Linobia eichhornia, L. nilotica, L. replacementica Morocco (Cooreman, 1951)
– Hemisarcoptes malus
South Africa (Meyer, 1962) – Hemisarcoptes coccophagus
ASIA
China (Fan, Chen & Wang, 2010; Ji, L.-Z., Gerson & Izraylevich, 1994; Li, L.-S., Xuan,
J.-Y. & Fan, Q.-H., 1992) – Hemisarcoptes sp., Nanacarus minutus
Indonesia (Oudemans, 1901, 1928) – Nanacarus minutus
Iran (Tarasi, Saboori & Sadeghi, 2001) – Linobia coccinellae
Israel and Palestina (Gerson, 1964, 1967; Gerson & Schneider, 1981, 1982; Izraylevich
& Gerson, 1993) – Hemisarcoptes coccophagus
Malaysia (OConnor & Houck, 1989a) – Pasohopus labrynthus
NORTH AMERICA
Canada (Mullen et al., 1989; Tothill, 1918; Baulieu et al., 2019) – Hemisarcoptes malus
and two other sp., Divilia sp.
USA (Shimer, 1868; Woodring, 1966; OConnor, 1991, OConnor & Houck, 1989b; Luck,
Jiang & Houck, 1999) – Huronopus michiganensis, Nanacarus nominis, Superioropus
huronmontanus
SOUTH and CENTRAL AMERICA
Colombia (Fain, 1987) – Espeletiacarus andinus
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Acarorum Catalogus IX
OCEANIA
Australia (continent) (Gerson, 1994) – Hemisarcoptes 2 spp.
New Zealand (Charles, Hill & Allan, 1995a, 1995b; Hill et al., 1993) – Hemisarcoptes
coccophagus
Hosts of Hemisarcoptidae
Insecta s.l.
Collembola
Fam. Sminthuridae
Sminthurus fuscus – Nanacarus collembolicola
Coleoptera
Fam. Chrysomelidae
Chrysomela populi – Linobia coccinellae
C. saliceti - Linobia coccinellae
Fam. Coccinellidae
Adalia sp. - Nanacarus minutus
Chilocorus cacti – Hemisarcoptes cooremani
Ch. renipustulatus - Hemisarcoptes budensis
Exochomus quadripustulus - Nanacarus minutus
Vibidia duodecimguttata - Nanacarus minutus
Homoptera
Coccodea
Fam. Diaspididae
Chrysomphalus dictyospermi – Hemisarcoptes malus
Hemiberlesia lataniae – Hemisarcoptes coccophagus
Lepidosaphes beckii – Hemisarcoptes malus
L. gloveri - Hemisarcoptes malus
Parlatoria blanchardi – Hemisarcoptes malus
P. zizyphi - Hemisarcoptes malus
Hymenoptera
Fam. Apidae
Xylocopa (Koptorthosoma) tenuiscapa – Nanacarus minutus
Fam. Halictidae
Nominia melanderi – Nanacarus nominis
Fam. Formicidae
Lasius fuliginorus – Divilia oculata
Formica polyctena - Divilia oculata
F. rufa - Divilia oculata
Fam. Ichneumonidae
Exeristes comstockii - Superioropus huronmontanus
Acarorum Catalogus IX 335
Diptera
Fam. Glossinidae
Glossina fuscipes fuscipes – Nanacarus hieroglypha
Aves
Fam. Megapodiidae
Megapodius reinwardt buruensis – Nanacarus minutus
Mammalia
Soricomorpha
Fam. Soricidae
Sorex araneus – Nanacarus minutus
Chiroptera
Fam. Vespertilionidae
Eptesicus serotinus – Nanacarus minutus
Pipistrellus pipistrellus - Nanacarus minutus
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Trach V.A. 2007. On the record of Linobia coccinellae (Astigmata, Hemisarcoptidae) in Ukraine.
-Vestnik Zoologii, 41: 510.
Türk P., S. Türk. 1952. LV.—Studies of Acari.—7th series: “Records and descriptions of mites
new to the British fauna, together with short notes on the biology of sundry species”. Annals and Magazine of Natural History, 5: 475– 506.
Türk P., S. Türk. 1957. Systematik und Ökologie der Tyroglyphiden Mitteleuropas in H.- J.
Stammer: Beiträge zur Systematik und Ökologie Mitteleuropäischer Acarina. – Akad.
Verlagsges., Leipzig, 1, Teil 1, Abh. 1: 1–231.
Volgin V. I., S.V. Mironov. 1979. New species and new genus of mites of the family Saproglyphidae (Acarina, Acaroidea). – Trudy Zool. Inst. Akad. Nauk U.S.S.R., 85: 91–98 (in Russian).
The Mite Family Carpoglyphidae
Carpoglyphidae Oudemans, 1923: 206
The best known representative of the Carpoglyphidae is the dried fruit mite, Carpoglyphus
lactis (L.). This species is a cosmopolitan pest of stored materials with a high sugar content,
such as dried fruit, powdered milk, and honey, as well as of other stored food substrates
(Chmielewski 1971; Hughes 1976). Deutonymphs have been collected from Lepidoptera
and Coleoptera (Türk & Türk, 1957).
Composition of Carpoglyphidae
Fam. Carpoglyphidae Oudemans, 1923–4 gen., 12 sp.
Carpoglyphus Robin, 1869–9 sp.
Coproglyphus Türk et Türk, 1957–1 sp.
Dichotomiopus Fain et Camerik, 1978–1 sp.
Pullea Canestrini, 1884–1 sp.
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Catalogue of Carpoglyphidae
Family Carpoglyphidae Oudemans
Carpoglyphidae Oudemans, 1923: 206
Type genus: Carpoglyphus Robin, 1869
Genus Carpoglyphus Robin
Carpoglyphus Robin, 1869: 197
Type species: Acarus lactis Linnaeus, 1758
Carpoglyphus afzali Khan
Carpoglyphus afzali Khan, 1967: 398
Type locality: Pakistan
Distribution: Pakistan
Carpoglyphus alienus Banks
Carpoglyphus alienus Banks, 1905: 42
Type locality: “Batavia” (Java)
Distribution: Indonesia (Java)
Carpoglyphus ganzhouensis Jiang
Carpoglyphus ganzhouensis Jiang, 1991: 82
Type locality: Jiangxi
Distribution: China
Carpoglyphus lactis (Linnaeus, 1758) [ 1767?]
Acarus lactis Linnaeus, 1767: 1024
Carpoglyphus lactis (Linnaeus): Jeleva & Angelkova, 1964:
Carpoglyphus passularum Robin, 1869
Glycyphagus anonymus Haller, 1882
Type locality: Europe
Distribution: Argentina, Australia, Bulgaria, China, England, Germany, Italy
Carpoglyphus munroi Hughes
Carpoglyphus munroi Hughes, 1962: 168
Type locality: Imperial College, Field Station, Silwood Park, Ascot, Surrey
Distribution: England
Carpoglyphus nidicolous Hubard et Fashing
Carpoglyphus nidicolous Hubard & Fashing, 1996: 215
Acarorum Catalogus IX 341
Type locality: Oregon
Distribution: USA
Host: Petrochelidon pyrrhonota (Aves) nests
Carpoglyphus sturmi Fain et Rack
Carpoglyphus sturmi Fain & Rack, 1987: 12
Type locality: Páramo de Montserrate, 3250 m
Distribution: Colombia
Host: Espeletiopsis corymbosa (plant)
Carpoglyphus vagabundus Berlese
Carpoglyphus vagabundus Berlese, 1910: 267
Type locality: Rosignano (Pisa)
Distribution: Italy
Host: nest of Polystes gallicus (Hymenoptera: Vespidae)
Carpoglyphus wardleorum Clark
Carpoglyphus wardleorum Clark, 2010: 454
Type locality: Ashley Gorge, Canterbury
Distribution: New Zealand
Host: from sooty mould fungi on the bark of black beech, Nothofagus solandri.
Genus Coproglyphus Türk et Türk
Coproglyphus Türk & Türk, 1957: 181
Type species: Coproglyphus stammeri Türk et Türk, 1957
Coproglyphus stammeri Türk et Türk
Coproglyphus stammeri Türk & Türk, 1957: 181; Li, 1999: 72
Type locality: Erlangen (Germany)
Distribution: China, Germany
Genus Dichotomiopus Fain et Camerik
Dichotomiopus Fain & Camerik, 1978: 172
Type species: Dichotomiopus biaggioi Fain et Camerik, 1978
Dichotomiopus biaggioi Fain et Camerik
Dichotomiopus biaggioi Fain & Camerik, 1978: 173
Type locality: Cotia, Sao Paulo
Distribution: Brazil
Host: Dichotomius anaglypticus (Coleoptera: Scarabaeidae)
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Genus Pullea Canestrini
Pullea Canestrini, 1884: 720
Type species: Pullea discoidalis Canestrini, 1884
Pullea discoidalis Canestrini
Pullea discoidalis Canestrini, 1884: 720
Type locality: Australia
Distribution: Australia
Index of Carpoglyphidae
(Names in bold are accepted as valid)
afzali, Carpoglyphus 340
alienus, Carpoglyphus 340
anonymus, Glyciphagus 340
biaggioi, Dichotomiopus 341
Carpoglyphidae 340
Carpoglyphus 340
Coproglyphus 341
Dichotomiopus 341
discoidalis, Pullea 342
ganzhouensis, Carpoglyphus 340
lactis, Acarus 340
lactis, Carpoglyphus 340
munroi, Carpoglyphus 340
nidicolous, Carpoglyphus 340
passularum, Carpoglyphus 340
Pullea 342
stammeri, Coproglyphus 341
sturmi, Carpoglyphus 341
vagabundus, Carpoglyphus 341
wardleorum, Carpoglyphus 341
Distribution of Carpoglyphidae
EUROPE
Bulgaria (Jeleva & Angelkova, 1964) – Carpoglyphus lactis
Germany (Türk & Türk, 1957) – Carpoglyphus lactis, Coproglyphus stammeri
Great Britain (Hughes, 1962) – Carpoglyphus lactis, C. munroi
Holland (Siepel, Cremers & Vierbergen, 2016) - Carpoglyphus lactis
Italy (contin.) (Berlese, 1910) – Carpoglyphus lactis, C. vagabundus
Turkey (Turgu & Kumral, 2019) - Carpoglyphus lactis
ASIA
China (Jiang, 1991; Li, 1999; Fan, Chen & Wang, 2010) – Carpoglyphus ganzhouensis, C.
lactis, Coproglyphus stammeri
Indonesia (Banks, 1905) – Carpoglyphus alienus
Pakistan (Khan, 1967) – Carpoglyphus afzali
Acarorum Catalogus IX 343
NORTH AMERICA
USA (Hubard & Fashing, 1996) – Carpoglyphus nidicolous
SOUTH and CENTRAL AMERICA
Argentina (Hughes, 1978) – Carpoglyphus lactis
Brazil (Fain & Camerik, 1978) – Dichotomiopus biaggioi
Colombia (Fain & Rack, 1987) – Carpoglyphus sturmi
OCEANIA
Australia (continent) (Womersley, 1941) – Carpoglyphus lactis, Pullea discoidalis
New Zealand (Clark, 2010) – Carpoglyphus wardleorum
Hosts of Carpoglyphidae
Hymenoptera
Fam. Vespidae
Polystes gallicus - Carpoglyphus vagabundus
Bibliography of Carpoglyphidae
Baker E.W., M.D. Delfinado. 1978. Notes on dried fruit mite Carpoglyphus lactis (Acarina:
Carpoglyphidae) infesting honeybee combs. - Jour. Agr. Res., 17: 52–54.
Banks N. 1905. An alleged parasitic Tyroglyphid. - Proceedings of the Entomological Society
of Washington, 7(1): 40–42.
Berlese A. 1910. Lista di nuove specie e nuovi generi di Acari. – Redia, 6(2): 242–271.
Chmielewski W. 1971. Morfologia, biologia i ekologia Carpoglyphus lactis (L., 1758) (Glycyphagidae, Acarina). - Prace Nauk. Inst. Ochr. Rostlin, 13: 63–166.
Chmielewski W. 1999. Bionomics of Carpoglyphus lactis (Acari: Carpoglyphidae) on honey. - Ecology and Evolution of the Acari, Series Entomologica book series (SENT, volume 55), 423–424
Clark J.M. 2010. A new species of Carpoglyphus (Astigmatina: Carpoglyphidae) from the
bark of black beech (Nothofagus) honeydew in New Zealand. – Int. J. of Acarology, 36(5):
Fain A., A.M. Camerik. 1978. New hypopi phoretic on Brazilian Coleoptera and Hymenoptera
(Acari: Astigmata). – Intl. J. Acar., 4(3): 169–178.
Fain A., G. Rack. 1987. Notes on the mites living in the flowers of Espeletia spp. (Asteraceae)
in Colombia. 1. Carpoglyphus sturmi sp. n. (Acari, Carpoglyphidae). - Entomologische
Mitteilungen aus dem Zoologischen Staatsinstitut und Zoologischen Museum Hamburg,
9(130): 9–19.
Fan Qing-Hai, Yan Chen, Zi-Qing Wang. 2010. Acaridia (Acari: Astigmatina) of China: A
review of research progress. – p.225–259. In: Zhang Z.-Q., Hong X.-Y. & Fan Q.-H. (eds)
Xin Jie-Lin Centenary: Progress in Chinese Acarology. Zoosystema 4, 1–345.
344
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Hubard C.K., N.J. Fashing. 1996. Carpoglyphus nidicolus - a new species of Carpoglyphidae
(Acarina: Astigmata) inhabiting the nests of swallows. -International Journal of Acarology, 22(3): 215–220.
Hughes A.M. 1952. A new glycyphagid mite - Carpoglyphus munroi. - Journal of the Linnean
Society of London, Zoology, 42(284): 168–175.
Hughes A.M. 1976. The mites of stored food and houses (No. 2nd edition). Her Majesty’s
Stationery Office, London, 400 pp.
Jeleva M., E.B. Angelkova. 1964. Contribution to the study of the Tyroglyphid fauna in
Bulgaria. – Ann. Univ. Sofia, 56, Fac. Biol., Géol., Géogr., Livre 1 – Biologie (Zoologie):
141–146 (in Bulg., summ. Russ., Engl.).
Jiang Z.-T. 1991. A new species of Carpoglyphidae from China (Acarina: Acaroidea). - Journal
of Nanchang University (Natural Science), 15(1): 82–86.
Li C.P. 1999. Preliminary study of the acaridid mites breeding in stored Chinese traditional
medical materials. – Chinese Journal of Parasitic Diseases Control, 12(1): 72–73.
ODonnovan W.J. 1920. Carpoglyphus Passularum causing Dermatitis. - Proc R Soc Med. 1920;
13 (Dermatol Sect): 150–151.
Oudemans A.C. 1923. Studie over de sedert 1877 ontworpen systemen der Acari; nieuwe
classificatie; phylogenetische beschouwingen. - Tijdschrift voor entomologie, 66: 49—85.
Robin C. 1869. Note sur un nouveau genre d‘acariens. - Journal de l‘Anatomie et de la
Physiologie Normales et Pathologiques de l‘Homme et des Animaux, 6: 197–204, plates
7–8.
Siepel H., H. Cremers, B. Vierbergen. 2016. Provisional checklist of the astigmatic mites of the
Netherlands (Acari: Oribatida: Astigmatina). – Nederlandse Faunistische Mededelingen,
47: 49–88.
Tomaševič B. 1956. Carpoglyphus lactis L. na suvim sljivama. - Zastita Bilja, 33: 57–65.
Turgu V., N.A. Kumral. 2019. Alternative control agents of the dried fruit mite, Carpoglyphus
lactis (L.) (Acari: Carpoglyphidae) on dried apricots. - Acarological Studies, 1(2): 59–64.
Türk E., F. Türk. 1957. Systematik und Ökologie der Tyroglyphiden Mitteleuropas in H. J.
Stammer: Beiträge zur Systematik und Ökologie Mitteleuropäischer Acarina. – Akad.
Verlagsges., Leipzig, 1, Teil 1, Abh. 1: 1–231.
Womersley H. 1941. Studies in Australian Acarina. (2) Tyroglyphidae (s.l.). – Records of the
South Australian Museum, 6(4): 481–487.
The Mite Family Winterschmidtiidae
(Saproglyphidae)
Winterschmidtiidae Oudemans, 1923
Saproglyphinae Oudemans, 1924
Winterschmidtiidae (= Saproglyphidae) is the largest family in the Hemisarcoptoidea,
with 28 genera and 139 known species with a wide range of ecological associations.
Acarorum Catalogus IX 345
Species of Saproglyphus are fungivores in decaying materials, bark crevices, and woody
fungi, and deutonymphs are dispersed by a wide variety of insecthosts. Other species are
common fungivores on leaf surfaces. In temperate regions, the tritonymph is a resistant
overwintering stage. Species of Winterschmidtia are also predators of the eggs of bark
beetles (Scolytinae). Some taxa are known from deutonymphs associated with various
families of subcortical Coleoptera. Other winterschmidtiid genera are found in various
types of decaying material such as dung, carrion and in vertebrate nests. Procalvolia zacheri
(Oudemans) and Acalvolia squamata (Oudemans) occasionally infest stored food products,
P. zacheri was associated with severe dermatitis in a horse. The subfamily Ensliniellinae
includes obligate associates of Hymenoptera. Some taxa are associated with sphecid wasps
(Crabrovidia and others), most species are associates of Vespidae. Other genera are associated with solitary wasps of the subfamilies Masarinae (pollen wasps) and Eumeninae. A
partial list of associations and a phylogeny for the subfamily was provided by OConnor
and Klompen (1999). Some species such as Monobiacarus quadridens Baker and Kurosaia
jiju OConnor et Okabe feed on hemolymph of paralyzed arthropod prey provisioned by
the female wasp. Other species, in the genera Ensliniella, Kennethiella, and Vespacarus
feed directly on hemolymph of the wasp host larva or pupa. Many female (or rarely male)
wasp hosts have more or less elaborated acarinaria in different locations on the body that
harbor dispersing deutonymphs. Based on their phylogeny, OConnor and Klompen (1999)
concluded that acarinaria have evolved independently in many host groups.
For many years was used the family name Saproglyphidae Oudemans, 1924, but
OConnor (1984) insisted to use the antedating name Winterschmidtiidae Oudemans,
1923 and was followed by other researchers. Now both names are used on subfamily level.
Composition of Winterschmidtiidae
Fam. Winterschmidtiidae (Saproglyphidae) - 28 gen., 139 sp.
Subfam. Ensliniellinae Vitzthum, 1924–71 sp.
Crabrovidia Zakhvatkin, 1941–20 sp.
Ensliniella Vitzthum, 1925–7 sp.
Kennethiella Cooreman, 1954–1 sp.
Kurosaia Okabe et OConnor, 2002–1 sp.
Macroharpa Mostafa, 1970–4 sp.
Monobiacarus Baker et Cunliffe, 1960–3 sp.
Sphexicozela Mahunka, 1970–1 sp.
Vespacarus Baker, 1960–10 sp.
Vidia Oudemans, 1905–2 sp.
Zethacarus Mostafa, 1971–6 sp.
Zethovidia Mostafa, 1970–16 sp.
Subfam. Saproglyphinae Oudemans, 1924–18 sp.
Acalvolia Fain, 1971–2 sp.
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Congovidia Fain et Elsen, in Fain, 1971–6 sp.
Gambacarus Mahunka, 1975–1 sp.
Neocalvolia Hughes, 1970–3 sp.
Neosuidasia Ranganath et Channabasa,1983–1 sp.
Neottiglyphus Volgin, 1974–1 sp.
Saproglyphus Berlese, 1890–4 sp.
Subfam. Oulenziinae Oudemans, 1928– 7 sp.
Oulenzia Radford, 1950–1 sp.
Oulenziella Fan et al., 2015–2 sp.
Procalvolia Fain, 1971–3 sp.
Trypetacarus Fain, 1971–1 sp.
Subfam. Winterschmidtiinae Oudemans, 1923–50 sp.
Allocalvolia Fain et Rack, 1987–2 sp.
Calvolia Oudemans, 1911–34 sp.
Czenspinskia Oudemans, 1927–2 sp.
Parawinterschmidtia Khaustov, 2000–1 sp.
Riemia Oudemans, 1925–1 sp.
Winterschmidtia Oudemans, 1923 (= Afrocalvolia Fain et Elsen, 1971) – 10 sp.
Catalogue of Winterschmidtiidae
Fam. Winterschmidtiidae Oudemans
Winterschmidtiidae Oudemans, 1923: 206
Saproglyphidae Oudemans, 1924
Czenspinskiidae Oudemans, 1927
Oulenziidae Oudemans, 1928
Calvoliinae Turk & Turk, 1957: 155
Type genus: Winterschmidtia Oudemans, 1923
Subfam. Ensliniellinae Vitzthum
Genus Crabrovidia Zakhvatkin
Vidia (Crabrovidia) Zakhvatkin, 1941: 371
Crabrovidia Zachvatkin: Fain, 1971: 284
Euvidia Delfinado & Baker, 1976: 81] [Type is utahensis; syn. OConnor & Eickwort 1988:
172]
Type species: Vidia (Crabrovidia) gussakovskii Zachvatkin, 1941
Crabrovidia africana Fain
Crabrovidia africana Fain, 1973: 183
Acarorum Catalogus IX 347
Type locality: Kibombo
Distribution: DR Congo
Host: Sphex hemorrhoidalis (Hymenoptera: Sphecidae)
Crabrovidia brasiliensis Fain
Crabrovidia brasiliensis Fain, 1973: 169
Type locality: Merretes, Alto de Sierra, Sta Catarina, Nova Teutonia
Distribution: Brazil
Host: Ectemnius cubiceps (Hymenoptera: Crabronidae)
Crabrovidia concellaria (Cooreman)
Vidia concellaria Cooreman, 1948a: 1
Crabrovidia concellaria Fain, 1973: 179
Type locality: Uccle (Bruxelles)
Distribution: Belgium, France
Host: Cerceris arenaria (Hymenoptera: Crabronidae); ? accidental host: Hedichrum nobile
(Chrysididae)
Crabrovidia corsicana Fain
Crabrovidia corsicana Fain, 1973: 183
Type locality: Bonifacio
Distribution: Corsica
Host: Crossocercus elongatus (Hymenoptera: Crabronidae)
Crabrovidia crossocerus Fain
Crabrovidia crossocerus Fain, 1973: 167
Type locality: Sta Catarina, Nova Teutonia
Distribution: Brazil
Host: Crossocerus porexus (Hymenoptera: Crabronidae)
Crabrovidia cubensis Fain
Crabrovidia cubensis Fain, 1973: 170
Type locality: Guantanamo
Distribution: Cuba
Host: Lestica cubensis (Hymenoptera: Crabronidae)
Crabrovidia dasyproctus Fain
Crabrovidia dasyproctus Fain, 1973: 177
Type locality: Taiherin (Taiwan)
Distribution: India, Philippines, South Africa (Natal), Taiwan
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Hosts: Dasyproctus agilis orientalis, D. bipunctatus lugubris, D. buddha, D. cevirus, D.
westermanni (Hymenoptera: Crabronidae)
Crabrovidia ectemnii Fain
Vidia gussakovskii: Cooreman, 1948: 8 (nec Zachvatkin, 1941)
Crabrovidia ectemnii Fain, 1973: 163
Type locality: La Panne (Belgium)
Distribution: Argentina, Austria, Belgium, Bolivia, Brazil, Colombia, Costa Rica, Croatia, Cyprus, France, Greece (Crete), Guiana, Honduras, Jamaica, Japan [Okinawa],
Mexico, Morocco, Puerto Rico, Spain, Switzerland
Hosts: Ectemnius carinatus, E. cavifrons, E. centralis, E. continuus, E. craesus, E. guttatus,
E. hypsae, E. impressus, E. laevigatus, E. lapidaries, E. schettereri, E. semipunctatus, E.
singularis, E. sonorensis, E. sexcinctus (Hymenoptera: Crabronidae)
Crabrovidia gussakovskii (Zachvatkin)
Vidia (Crabrovidia) gussakovskii Zachvatkin, 1941: 270
Crabrovidia gussakovskii (Zachvatkin): Fain, 1973: 160; Klimov, 1998: 25
Type locality: Crimea (Karadag)
Distribution: Russia, Crimea, Kuril Isl. (Iturup)
Hosts: Crabro solenius, Ectemnius (Metacrabro) fossorius, E. (Hypocrabro) impressus, E.
(Hypocrabro) continuus (= vagus auct.)(Hymenoptera: Crabronidae)
Crabrovidia hirsuta (OConnor et Eickwort)
Vidia hirsuta OConnor & Eickwort, 1988: 161
Crabrovidia hirsuta (OConnor et Eickwort):
Type locality: New Mexico, Sandoval Co., Jemez Springs
Distribution: USA
Host: Megachile (Sayapis) fidelis (Hymenoptera: Megachilidae)
Crabrovidia julianii (Sevastianov)
Vidia julianii Sevastianov, in Kadzhaya & Sevastianov, 1967: 199
Crabrovidia julianii (Sevastianov): OConnor & Eickwort, 1988: 172
Type locality: Odessa
Distribution: Ukraine
Crabrovidia latimanus (OConnor et Eickwort)
Vidia latimanus OConnor & Eickwort, 1988: 160
Crabrovidia latimanus (OConnor et Eickwort):
Type locality: Nova Scotia, Cape Breton Highlands National Park, Ingonish Beach
Distribution: Canada
Acarorum Catalogus IX 349
Host: Megachile (Xanthosaurus) latimanus (Hymenoptera: Megachilidae)
Crabrovidia leclercqi Fain
Crabrovidia leclercqi Fain, 1973: 179
Type locality: Yermasoyia
Distribution: Cyprus
Host: Cerceris rubida (Hymenoptera: Crabronidae)
Crabrovidia ninae (Sevastianov)
Vidia ninae Sevastianov, in Kadzhaya & Sevastianov 1967: 198
Crabrovidia ninae (Sevastianov): OConnor & Eickwort 1988: 172
Type locality: Kherson Prov.
Distribution: Ukraine
Host: Pseudovespa sp. (Hymenoptera: Vespidae)
Crabrovidia oudemansi Fain
Crabrovidia oudemansi Fain, 1971: 284; 1972: 244; 1973: 175; Nazan et al., 2019
Type locality: Corfu (Greece)
Distribution: Greece (Corfu), Iran
Host: Cerceris sp. (Hymenoptera: Crabronidae)
Crabrovidia popovi (Zachvatkin)
Vidia (Crabrovidia) popovi Zachvatkin, 1941: 375
Crabrovidia popovi (Zachvatkin): Fain, 1973: 161
Type locality: Krasnograd (Ukraine)
Distribution: Abhasia, Croatia, Italy, Spain, Turkey, Ukraine, Uzbekistan
Hosts: Lestica (Clypeocrabro) clypeata, L. (Ceratocolus) alata, L. (Clypeocrabro) wollmanni,
Ectemnius impressus (Hymenoptera: Crabronidae)
Crabrovidia rubi (OConnor et Eickwort)
Vidia rubi OConnor & Eickwort, 1988: 148
Crabrovidia rubi (OConnor et Eickwort): Fain, 1971: 284
Type locality: Georgia, Liberty Co., St. Catherine‘s Island
Distribution: USA (Georgia)
Host: Megachile (Xeromegachile) rubi (Hymenoptera: Megachilidae)
Crabrovidia texana (OConnor et Eickwort)
Vidia texana OConnor & Eickwort, 1988: 162
Crabrovidia texana (OConnor et Eickwort):
Type locality: New York, Albany Co., Rensselaerville, Edmund Niles Huyck Preserve
Distribution: USA
Host: Megachile (Litomegachile) texana (Hymenoptera: Megachilidae)
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Crabrovidia thomasi (Delfinado et Baker)
Vidia thomasi Delfinado & Baker, 1976: 81 (nom. nom. for V. cooremani Baker)
Crabrovidia thomasi (Delfinado et Baker): OConnor & Eickwort 1988: 172 (nom. nov.
for Vidia cooremani Baker, 1964 nec V. cooremani Thomas, 1961)
Type locality: Plummers Island, Maryland
Distribution: USA (Maryland)
Crabrovidia utahensis (Delfinado et Baker)
Vidia (Euvidia) utahensis Delfinado & Baker, 1976: 81
Crabrovidia utahensis (Delfinado et Baker): OConnor & Eickwort 1988: 172
Type locality: Beaver, Utah
Distribution: USA
Host: Hymenoptera (Sphecidae)
Genus Ensliniella Vitzthum
Ensliniella Vitzthum, 1925 [1924?]: 289
Type species: Ensliniella aegyptiana Baker et Cunliffe, 1960
Ensliniella aegyptiana Baker et Cunliffe
Ensliniella aegyptiana Baker & Cunliffe, 1960: 214; Klompen & OConnor 1995: 126
Type locality: Abu Suttan
Distribution: Egypt
Hosts: Allodynerus koenigi, A. vinciquerrae (Hymenoptera: Vespidae, Eumeninae)
Ensliniella asiatica Abe et Okabe
Ensliniella asiatica Abe & Okabe, 2006: 929
Type locality: Sakurazawa, Yorii town, Osato District, Saitama Prefecture, central Honshu,
36°07’49”N, 139°12’28”E; 102 m
Distribution: Japan
Host: Allodynerus mandschuricus (Hymenoptera: Vespidae, Eumeninae)
Ensliniella dignotus Klompen et OConnor
Ensliniella dignotus Klompen & OConnor, 1995: 126
Type locality: Turkey
Distribution: Turkey
Host: Allodynerus dignotus (Hymenoptera: Vespidae, Eumeninae)
Ensliniella floricola Klompen et OConnor
Ensliniella floricola Klompen & OConnor, 1995: 127
Type locality: Pula (Croatia)
Distribution: Croatia, Austria, Bulgaria, France, Greece, Holland, Turkey
Hosts: Allodynerus floricola, A. rossii (Hymenoptera: Vespidae, Eumeninae)
Acarorum Catalogus IX 351
Ensliniella koenigi Baker et Cunliffe
Ensliniella koenigi Baker & Cunliffe, 1960: 216; Klompen & OConnor 1995: 127
Type locality: Ijonkak
Distribution: Morocco
Host: Allodynerus koenigi (Hymenoptera: Vespidae, Eumeninae)
Ensliniella kostylevi Zakhvatkin
Ensliniella kostylevi Zakhvatkin, 1941: 267; Klompen & OConnor, 1995
Type locality: Byalovezh (Belarus)
Distribution: Belarus, Crimea, Holland, Algeria, Austria, Belgium, Bulgaria, France,
Greece, Turkey
Host: Allodynerus rossii (Hymenoptera: Vespidae, Eumeninae)
Ensliniella parasitica Vitzthum
Ensliniella parasitica Vitzthum, 1925: 289; Cooreman, 1942; Baker & Cunliffe, 1960: 214;
Krivoluckiy, 1975: 476; Klompen & OConnor 1995: 128
Type locality: Germany (Bayern, Nürnberg)
Distribution: Armenia, Italy, Germany, Holland, Slovakia, Uzbekistan, Belgium, Czech
Rep., Spain, Gibraltar, Tadjikistan, Kyrghizstan, Kazakhstan
Hosts: Allodynerus delphinalis, A. floricola (Hymenoptera: Vespidae, Eumeninae; Melittobia acasta (Eulophidae)
Genus Kennethiella Cooreman
Kennethiella Cooreman, 1954: 2
Type species: Ensliniella trisetosa Cooreman, 1942
Kennethiella trisetosa (Cooreman)
Ensliniella trisetosa Cooreman, 1942: 1; Cooper, 1955
Kennethiella trisetosa (Cooreman): Cooreman, 1954; Baker & Cunliffe, 1960: 218
Type locality: USA
Distribution: Finland, USA
Host: Ancistrocerus antilope (Hymenoptera: Vespidae, Eumeninae)
Genus Kurosaia Okabe et OConnor
Kurosaia Okabe & OConnor, 2002: 69
Type species: Kurosaia jiju Okabe et OConnor, 2002
Kurosaia jiju Okabe et OConnor
Kurosaia jiju Okabe & OConnor, 2002: 69
Distribution: Japan
Host: Anterhynchium flavomarginatum (Hymenoptera, Vespidae)
352
Acarorum Catalogus IX
Genus Macroharpa Mostafa
Macroharpa Mostafa, 1970b: 395
Type species: Macroharpa amazonica Mostafa, 1970
Macroharpa amazonica Mostafa
Macroharpa amazonica Mostafa, 1970b: 397
Type locality: Faro, Estado de Para
Distribution: Brazil
Host: Zethus biglumis ferrugineus (Hymenoptera: Vespidae, Zethinae)
Macroharpa arota Mostafa
Macroharpa arota Mostafa, 1970b: 399
Type locality: Amazonas, Manaus
Distribution: Brazil
Host: Zethus arotus (Hymenoptera: Vespidae, Zethinae)
Macroharpa pachymapoda Mostafa
Macroharpa pachymapoda Mostafa, 1970b: 398
Type locality: Obidos, Estado de Para
Distribution: Brazil
Host: Zethus lobulatus (Hymenoptera: Vespidae, Zethinae)
Macroharpa palpognatha Mostafa
Macroharpa palpognatha Mostafa, 1970b: 400
Type locality: Tefe
Distribution: Brazil
Host: Zethus roridus (Hymenoptera: Vespidae, Zethinae)
Genus Monobiacarus Baker et Cunliffe
Monobiacarus Baker & Cunliffe, 1960: 210
Type species: Monobiacarus quadridens Baker et Cunliffe, 1960
Monobiacarus funebris Baker et Cunliffe
Monobiacarus funebris Baker & Cunliffe, 1960: 211
Type locality: Santa Rosa, Veracruz
Distribution: Mexico
Host: Monobia apicalipennis (Hymenoptera: Vespidae)
Monobiacarus insularis Baker et Cunliffe
Monobiacarus insularis Baker & Cunliffe, 1960: 212
Type locality: San Rafael, Jicoltepee
Distribution: Mexico
Acarorum Catalogus IX 353
Host: Monobia angulosa (Hymenoptera: Vespidae)
Monobiacarus quadridens Baker et Cunliffe
Monobiacarus quadridens Baker & Cunliffe, 1960: 211
Type locality: Kill Devis Hills, North Carolina
Distribution: USA
Host: Monobia quadridens (Hymenoptera: Vespidae)
Genus Sphexicozela Mahunka
Sphexicozela Mahunka, 1970: 77
Type species: Sphexicozela connivens Mahunka, 1970
Sphexicozela connivens Mahunka
Sphexicozela connivens Mahunka, 1970: 77; Rusina &· Orlova, 2011: 685
Type locality: Budapest
Distribution: Hungary, Ukraine, Crimea
Hosts: Polistes dominula, P. gallica, P. nimpha (Hymenoptera: Vespidae, Polistinae)
Genus Vespacarus Baker et Cunliffe
Vespacarus Baker & Cunliffe, 1960: 220
Type species: Vespacarus rufovestis Baker et Cunliffe, 1960
Vespacarus anacardivorus Baker et Cunliffe
Vespacarus anacardivorus Baker & Cunliffe, 1960: 224
Type locality: Paradise Key, Everglades National Park, Florida
Distribution: USA (Florida)
Host: Stenodynerus (Parancystrocerus) perennis anacardovora (Hymenoptera: Vespidae,
Eumeninae)
Vespacarus fulvipes Baker et Cunliffe
Vespacarus fulvipes Baker & Cunliffe, 1960: 222
Type locality: Kill Devil Hills, North Carolina
Distribution: USA
Host: Stenodynerus (Parancystrocerus) f. fulvipes (Hymenoptera: Vespidae, Eumeninae)
Vespacarus histrio Baker et Cunliffe
Vespacarus histrio Baker & Cunliffe, 1960: 222
Type locality: Kill Devil Hills, North Carolina
Distribution: USA
Hosts: Stenodynerus (Parancystrocerus) histrio, S. (P.) f. fulvipes (Hymenoptera: Vespidae,
Eumeninae)
354
Acarorum Catalogus IX
Vespacarus minimoferus Mostafa
Vespacarus minimoferus Mostafa, 1967: 618
Type locality: California
Distribution: USA
Host: Parancystrocercus minimoferus (Hymenoptera: Eumenidae)
Vespacarus pedestris Baker et Cunliffe
Vespacarus pedestris Baker & Cunliffe, 1960: 226
Type locality: Derby, New York
Distribution: USA
Host: Stenodynerus (Parancystrocerus) p. pedestris (Hymenoptera: Vespidae, Eumeninae)
Vespacarus rufovestis Baker et & Cunliffe
Vespacarus rufovestis Baker & Cunliffe, 1960: 220
Type locality: Lake Placid, Florida
Distribution: USA
Host: Stenodynerus (Parancystrocerus) fulvipes rufovestis (Hymenoptera: Vespidae, Eumeninae)
Vespacarus saecularis Baker et Cunliffe
Vespacarus saecularis Baker & Cunliffe, 1960: 224
Type locality: Lake Placid, Florida
Distribution: USA
Host: Stenodynerus (Parancystrocerus) saecularis (Hymenoptera: Vespidae, Eumeninae)
Vespacarus tigris Baker et Cunliffe
Vespacarus tigris Baker & Cunliffe, 1960: 228
Type locality: Arlington, Virginia
Distribution: USA
Host: Ancistrocerus t. tigris (Hymenoptera: Vespidae, Eumeninae)
Vespacarus toltecus Baker et Cunliffe
Vespacarus toltecus Baker & Cunliffe, 1960: 226
Type locality: Chisos Mts., Brewster Co., Texas
Distribution: USA (Texas)
Host: Stenodynerus (Parancystrocerus) toltecus (Hymenoptera: Vespidae, Eumeninae)
Vespacarus vagus Baker et Cunliffe
Vespacarus vagus Baker & Cunliffe, 1960: 228
Type locality: Toronto
Distribution: Canada
Host: Stenodynerus v. vagus (Hymenoptera: Vespidae, Eumeninae)
Acarorum Catalogus IX 355
Genus Vidia Oudemans
Vidia Oudemans, 1905: 22 [OConnor & Eickwort 1988]
Type species: Vidia undulata Oudemans, 1905
Vidia lineata Oudemans
Vidia lineata Oudemans, 1917: 391; Zachvatkin, 1941: 373; Buitendijk, 1945: 356; Fain,
1972: 240; Nazan et al., 2019
? Vidia stoeckherti Türk & Türk, 1957: 152 – Type: Erlangen, Germany.
Type locality: Bozen
Distribution: Belgium, Croatia, Germany, Greece (Corfu), Holland, Iran, Italy, Russia
Host: Megachile pilicrus (Hymenoptera: Megachilidae)
Vidia undulata Oudemans
Vidia undulata Oudemans, 1905: 22; Zachvatkin, 1941: 374; Fain, 1972: 239
Type locality: Maggianico
Distribution: Italy
Host: Hylaeus (Prosopis) conformis (Hymenoptera: Colletidae)
Genus Zethacarus Mostafa
Zethacarus Mostafa, 1971: 566
Type species: Zethacarus boharti Mostafa, 1971
Zethacarus andinus Mostafa
Zethacarus andinus Mostafa, 1971: 572
Type locality: R.A. Cordoba, Totoral
Distribution: Argentina
Host: Zethus andinus (Hymenoptera: Vespidae, Zethinae)
Zethacarus argentinae Mostafa
Zethacarus argentinae Mostafa, 1971: 571
Type locality: Chaco de Santiago del Estro Bords du Rio Salado env. d’Icano
Distribution: Argentina
Host: Zethus haywardi (Hymenoptera: Vespidae, Zethinae)
Zethacarus boharti Mostafa
Zethacarus boharti Mostafa, 1971: 567
Type locality: Tamaulipas
Distribution: Mexico
Host: Zethus gurreroi mayorum (Hymenoptera: Vespidae, Zethinae)
Zethacarus ellipsus Mostafa
Zethacarus ellipsus Mostafa, 1971: 575
356
Acarorum Catalogus IX
Type locality: Orizaba, Ver.
Distribution: Mexico
Host: Zethus orizabae (Hymenoptera: Vespidae, Zethinae)
Zethacarus miscogaster Mostafa
Zethacarus misco gaster Mostafa, 1971: 569
Type locality: 38 mi. W Jiquilpan Mich.
Distribution: Mexico
Host: Zethus miscogaster (Hymenoptera: Vespidae, Zethinae)
Zethacarus paranicus Mostafa
Zethacarus paranicus Mostafa, 1971: 574
Type locality: Parana
Distribution: Brazil
Host: Zethus smithii (Hymenoptera: Vespidae, Zethinae)
Genus Zethovidia Mostafa, 1970
Zethovidia Mostafa, 1970a: 178 (subgenus of Vidia)
Zethovidia Mostafa: Fain, 1972: 245 (genus)
Type species: Vidia (Zethovidia) diminuta Mostafa, 1970
Zethovidia acaris (Mostafa)
Vidia (Zethovidia) acaris Mostafa, 1970b: 385
Zethovidia acaris (Mostafa)
Type locality: Maroni
Distribution: French Guiana
Host: Zethus carbonarius (Hymenoptera: Vespidae, Zethinae)
Zethovidia arizonensis (Mostafa)
Vidia (Zethovidia) arizonensis Mostafa, 1970a: 187
Zethovidia arizonensis (Mostafa)
Type locality: San Simeon, Cochise Co., Arizona
Distribution: USA
Host: Zethus guerreroi arizonensis (Hymenoptera: Vespidae, Zethinae)
Zethovidia brasilensis (Mostafa)
Vidia (Zethovidia) brasilensis Mostafa, 1970a: 389
Zethovidia brasilensis (Mostafa)
Type locality: Chapa
Distribution: Brazil
Host: Zethus coeruleopennis (Hymenoptera: Vespidae, Zethinae)
Acarorum Catalogus IX 357
Zethovidia brevapodemis (Mostafa)
Vidia (Zethovidia) brevapodemis Mostafa, 1970b: 387
Zethovidia brevapodemis (Mostafa)
Type locality: Suriname
Distribution: Suriname
Host: Zethus magnus (Hymenoptera: Vespidae, Zethinae)
Zethovidia compactilis (Mostafa)
Vidia (Zethovidia) compactilis Mostafa, 1970a: 188
Zethovidia compactilis (Mostafa)
Type locality: 3 mi. N.W. Petlalcingo, Puebla
Distribution: Mexico
Host: Zethus guerreroi guerreroi (Hymenoptera: Vespidae, Zethinae)
Zethovidia diminuta (Mostafa)
Vidia (Zethovidia) diminuta Mostafa, 1970a: 181
Zethovidia diminuta (Mostafa)
Type loc/ality: San Bernardino
Distribution: Brazil, Paraguay
Host: Zethus diminutus diminutus, Z. diminutus santaremae (Hymenoptera: Vespidae,
Zethinae)
Zethovidia floridae (Mostafa)
Vidia (Zethovidia) floridae Mostafa, 1970b: 392
Zethovidia floridae (Mostafa)
Type locality: Royal Palm Park, Florida
Distribution: USA
Host: Zethus slossonae (Hymenoptera: Vespidae, Zethinae)
Zethovidia fusca (Mostafa)
Vidia (Zethovidia) fusca Mostafa, 1970b: 394
Zethovidia fusca (Mostafa)
Type locality: Barro Colorado, Canal Zone
Distribution: Panama
Host: Zethus fuscus (Hymenoptera: Vespidae, Zethinae)
Zethovidia laevapodemis (Mostafa)
Vidia (Zethovidia) laevapodemis Mostafa, 1970a: 183
Zethovidia laevapodemis (Mostafa)
358
Acarorum Catalogus IX
Type locality: 23 mi. S. Matias Romero, Oaxaca
Distribution: Mexico
Host: Zethus menkei (Hymenoptera: Vespidae, Zethinae)
Zethovidia mexicana (Mostafa)
Vidia (Zethovidia) mexicana Mostafa, 1970a: 184
Zethovidia mexicana (Mostafa)
Type locality: 30 mi. W. Acayucan, Veracruz
Distribution: Mexico
Host: Zethus parkeri (Hymenoptera: Vespidae, Zethinae)
Zethovidia oculabsensa (Mostafa)
Vidia (Zethovidia) oculabsensa Mostafa, 1970a: 191
Zethovidia oculabsensa (Mostafa)
Type locality: Chapada de Aunr Mato Grosso
Distribution: Brazil
Host: Zethus inconstans (Hymenoptera: Vespidae, Zethinae)
Zethovidia paraguayensis (Mostafa)
Vidia (Zethovidia) paraguayensis Mostafa, 1970a: 182
Zethovidia paraguayensis (Mostafa)
Type locality: San Bernardino
Distribution: Paraguay
Host: Zethus diminutus diminutus (Hymenoptera: Vespidae, Zethinae)
Zethovidia parvula (Mostafa)
Vidia (Zethovidia) parvula Mostafa, 1970b: 392
Zethovidia parvula (Mostafa)
Type locality: Corrientes
Distribution: Argentina
Host: Zethus parvulus (Hymenoptera: Vespidae, Zethinae)
Zethovidia promina (Mostafa)
Vidia (Zethovidia) promina Mostafa, 1970a: 388
Zethovidia promina (Mostafa)
Type locality: Chapa
Distribution: Brazil
Host: Zethus prominens (Hymenoptera: Vespidae, Zethinae)
Zethovidia reticulata (Mostafa)
Vidia (Zethovidia) reticulata Mostafa, 1970b: 384
Acarorum Catalogus IX 359
Zethovidia reticulata (Mostafa)
Type locality: 42 km SE Maturin Monagas
Distribution: Venezuela
Host: Zethus romandinus (Hymenoptera: Vespidae, Zethinae )
Zethovidia rugocoxa (Mostafa)
Vidia (Zethovidia) rugocoxa Mostafa, 1970a: 189
Zethovidia rugocoxa (Mostafa)
Type locality: 3 mi. N.W. Petlalcingo, Puebla
Distribution: Mexico
Host: Zethus guerreroi guerreroi (Hymenoptera: Vespidae, Zethinae )
Zethovidia salvadorensis (Mostafa)
Vidia (Zethovidia) salvadorensis Mostafa, 1970a: 186
Zethovidia salvadorensis (Mostafa)
Type locality: 4 mi. N. Quezaltepeque
Distribution: El Salvador
Host: Zethus irwini (Hymenoptera: Vespidae, Zethinae )
Subfamily Saproglyphinae Oudemans
Saproglyphinae Oudemans, 1924
Type genus: Saproglyphus Berlese, 1890
Genus Acalvolia Fain
Acalvolia Fain, 1971: 283
Type species: Vidia squamata Oudemans, 1909
Acalvolia americana Fan Qing-Hai, George & Kumarasinghe
Acalvolia americana Fan Qing-Hai, George & Kumarasinghe, 2010: 42
Type locality: intercepted in New Zealand from oranges from the USA
Distribution: USA
Acalvolia squamata (Oudemans)
Vidia squamata Oudemans, 1909: 317; Zachvatkin, 1941: 371
Acalvolia squamata (Oudemans): Fain, 1971: 283; 1972: 234; Fain & Knülle, 1981: 140;
Barbosa & de Moraes, 2021
Type locality: Semarang (Java)
Distribution: Germany, Holland, (?) Indonesia (probably an error), Brazil
Host: Mus jerdoni (Rodentia: Muridae); house dust
Genus Congovidia Fain et Elsen, in Fain, 1971
360
Acarorum Catalogus IX
Congovidia Fain & Elsen, in Fain 1971: 282; Fain, Hurst, Tweddle, Lachlan, Majerus &
Britt 1995: 100; see also Fain & Camerik, 1977 description of adult]
Type species: Congovidia glossinae Fain, 1971
Congovidia benelux Fain, Hurst, Fassotte, Webberley, Sloggett et Majerus
Congovidia benelux Fain, Hurst, Fassotte, Webberley, Sloggett & Majerus, 1997: 89
Type locality: Wageningen
Distribution: Holland
Host: Adalia bipunctata (Coleoptera: Coccinellidae)
Congovidia brasiliensis Fain et Camerik
Congovidia brasiliensis Fain & Camerik, 1977: 46; Fain, Hurst, Tweddle, Lachlan, Majerus
& Britt 1995: 103
Type locality: Itatiaia (State of Rio de Janeiro)
Distribution: Brazil
Host: in the nest of Trypoxylon (Trypargilum) aestivale (Hymenoptera: Sphegidae)
Congovidia coccinellidarum Fain, Hurst, Tweddle, Lachlan, Majerus et Britt
Congovidia coccinellidarum Fain, Hurst, Tweddle, Lachlan, Majerus & Britt, 1995: 100
Type locality: England
Distribution: England, Belgium
Host: Halyzia sedecimguttata (Coleoptera: Coccinellidae)
Congovidia glossinae Fain
Congovidia glossinae Fain & Elsen , 1971: 282; 1972: 77; Fain, Hurst, Tweddle, Lachlan,
Majerus & Britt, 1995: 103
Type locality: Luputa, Mabenga, Benzali
Distribution: DR Congo
Host: Glossina fuscipes quanzensis (Diptera: Glossinidae)
Congovidia lomamiensis Fain et Elsen
Congovidia lomamiensis Fain & Elsen, 1972: 81; Fain, Hurst, Tweddle, Lachlan, Majerus
& Britt, 1995: 103
Type locality: Luputa
Distribution: DR Congo
Host: Glossina fuscipes quanzensis (Diptera: Glossinidae)
Congovidia servalina Fain et Elsen
Congovidia servalina Fain & Elsen, 1972: 83; Fain, Hurst, Tweddle, Lachlan, Majerus &
Britt, 1995: 103
Type locality: Lundu, Mayumbe
Distribution: DR Congo
Acarorum Catalogus IX 361
Host: Glossina palpalis palpalis (Diptera: Glossinidae)
Congovidia sp.
Hybomitra epistates H. lasiophthalma, H. microcephala, H. nuda, H. sodalist, Tabanus
lineata, T. reinwardii (Canada)
Genus Gambacarus Mahunka
Gambacarus Mahunka, 1975: 43
Type species: Gambacarus huettleri Mahunka, 1975
Gambacarus huettleri Mahunka
Gambacarus huettleri Mahunka, 1975: 43
Type locality: Gambia
Distribution: Gambia
Genus Neocalvolia Hughes
Neocalvolia Hughes, 1970: 153
Type species: Neocalvolia claggi Hughes, 1970
Neocalvolia claggi Hughes
Neocalvolia claggi Hughes, 1970: 153
Type locality: grass, Bird I., South Georgia
Distribution: South Georgia
Neocalvolia coperminensis Rosas et Casanueva
Neocalvolia coperminensis Rosas & Casanueva, 1993: 66
Type locality: South Shetlands
Distribution: South Shetlands
Neocalvolia inermis Fain et Schuster
Neocalvolia inermis Fain & Schuster, 1983: 421
Type locality: Harrington Sound, Main Island
Distribution: Bermudas
Genus Neosuidasia Ranganath et Channabasa, in Ranganath, Channabasavanna
& Krishna-Rao
Neosuidasia Ranganath & Channabasa, in Ranganath, Channabasavanna & Krishna-Rao,
1983: 63
Type species: Neosuidasia faini Ranganath et Channabasa, in Ranganath, Channabasavanna & Krishna-Rao, 1983
362
Acarorum Catalogus IX
Neosuidasia faini Ranganath et Channabasa, in Ranganath, Channabasavanna
& Krishna-Rao
Neosuidasia faini Ranganath & Channabasa, in Ranganath, Channabasavanna & KrishnaRao, 1983: 64
Type locality: unknown (India?)
Distribution: on imported animal feed by Vegetal Quarantine of Cuba
Genus Neottiglyphus Volgin
Neottiglyphus Volgin, 1974: 223
Type species Neottiglyphus alatavicus Volgin, 1974
Neottiglyphus alatavicus Volgin
Neottiglyphus alatavicus Volgin, 1974: 224
Type locality: Kazakhstan
Distribution: Kazakhstan
Genus Saproglyphus Berlese
Saproglyphus Berlese, 1890: 6
Type species: Saproglyphus neglectus Berlese, 1890 (Diagnostic, Volgin 1980: 161)
Saproglyphus cocciphagus Womersley
Saproglyphus cocciphagus Womersley, 1941: 471
Type locality: Australia
Distribution: Australia
Saproglyphus neglectus Berlese
Saproglyphus neglectus Berlese, 1890: 6 [Diagnostic, Volgin 1980: 161]
Type locality: Sahalin
Distribution: Russia (Sahalin), Italy
Saproglyphus setosus Volgin
Saproglyphus setosus Volgin, 1980: 159
Type locality: Yuzhno-Sakhalinsk (Sakhalin Island)
Distribution: Russia (Far East)
Host: on birch bark
Saproglyphus waldorfae (Fain et Johnston)
Calvolia waldorfae Fain & Johnston, 1974: 416
Saproglyphus waldorfae (Fain et Johnston): Khaustov et al., 2018: 53
Type locality: Domleo, Spinney, Sutton Bonington
Acarorum Catalogus IX 363
Distribution: England, Russia (Tyumen Region)
Host: Sminthurus fuscus (Collembola: Sminthuridae), galleries of Ips typographus (Coleoptera: Curculionidae, Scolytinae)
Subfam. Oulenziinae Oudemans
Oulenziidae: Oudemans, 1928
Type genus: Oulenzia Radford, 1950
Genus Oulenzia Radford
Lenzia Oudemans, 1928: 327 [praeocc.!]
Oulenzia Radford, 1950: 152; [syn. by Hughes 1962: 60; Type is arboricola; nomen nova
for Lenzia?; restaured by Fan et al., 2012]
Type species: Lenzia arboricola Oudemans, 1928
Oulenzia arboricola (Oudemans)
Lenzia arboricola Oudemans, 1928: 327
Calvolia arboricola (Oudemans): Hughes, 1962: 61
Oulenzia arboricola (Oudemans): Radford, 1950: 222; Fan, George & Kumarasinghe,
2012: 334; Barbosa & de Moraes, 2021
Type locality: Medan, Deli, Sumatra
Distribution: Indonesia (Sumatra), Brazil
Host: Hevea leaves
Genus Oulenziella Fan, George, Li et Zhang
Oulenziella Fan, George, Li & Zhang, 2015: 191
Type species: Calvolia bakeri Hughes, 1962
Oulenziella bakeri (Hughes)
Calvolia bakeri Hughes, 1962: 59 (as nom. nov.); Woodring, 1966: 80
Oulenzia bakeri (Hughes): Radford, 1950
Oulenziella bakeri (Hughes): Fan et al., 2015: 191; Barbosa & de Moraes, 2021
Oulenzia gossypii Meyer & Rodrigues, 1965: (Mozambique, Kaidundjua, on Gossypium
sp., syn. by Fan et al., 2015)
Type locality: India
Distribution: India, Mozambique, Brazil, Indonesia (Sumatra)
Oulenziella longiseta Barbosa et de Moraes
Oulenziella longiseta Barbosa & de Moraes, 2021:
Type locality: Brazil
Distribution: Brazil
364
Acarorum Catalogus IX
Genus Procalvolia Fain
Procalvolia Fain, 1971: 283
Type species: Calvolia zacheri Oudemans, 1929
Procalvolia aviophilus Sevastianov et Hassan
Procalvolia aviophilus Sevastianov & Hassan, 1989: 141
Type locality: Ukraine
Distribution: Ukraine
Host: Pica pica (Aves: Corvidae)
Procalvolia coccinella Sevastianov et Hassan
Procalvolia coccinella Sevastianov & Hassan, 1989: 141
Type locality: Ukraine
Distribution: Ukraine
Host: Coccinella (Coleoptera: Cocinellidae)
Procalvolia hamadi Sevastianov et Hassan
Procalvolia hamadi Sevastianov & Hassan, 1989: 141
Type locality: Egypt
Distribution: Egypt
Host: Labidura riparia (Dermaptera: Labiduridae)
Procalvolia zacheri (Oudemans)
Calvolia zacheri Oudemans, 1929: 36; Zachvatkin, 1941: 361
Calvolia romanovae Zachvatkin, 1941: 345; Hughes, 1962: 54 (syn. by Fain, 1972)(type
from Moskow); Angelkova, 1988:
Calvolia tarsoinfracta Türk & Türk, 1957: 167 (syn. with Calvolia romanovae by Hughes,
1962)
Procalvolia zacheri (Oudemans): Fain, 1971: 283; 1972: 235; Rack, 1980: 304
Type locality: Berlin
Distribution: Bulgaria, Germany, New Zealand (cheese), Russia
Genus Psylloglyphus Fain
Psylloglyphus Fain, 1966: 159
Type species: Psylloglyphus u. uilenbergi Fain, 1966
Subgenus Psylloglyphus (s. str.) Fain
Psylloglyphus Fain, 1966: 159
Type species: Psylloglyphus u. uilenbergi Fain, 1966
Acarorum Catalogus IX 365
Psylloglyphus (Psylloglyphus) australiensis Fain, Bartholomaeus, Cooke et
Beaucournu
Psylloglyphus (Psylloglyphus) australiensis Fain, Bartholomaeus, Cooke & Beaucournu,
1990: 97
Type locality: Forrest (Victoria)
Distribution: Australia
Hosts: Pygiopsylla hoplia, Bibikovana rainbowi, Acanthopsylla r. rothschildi, Macropsylla
hercules (Siphonaptera: Pygiopsyllidae)
Psylloglyphus (Psylloglyphus) crenulatus Fain
Psylloglyphus (Psylloglyphus) crenulatus Fain & Beaucournu, 1984: 100
Type locality: Ile de la Possession, Pointe Lieutard, alt. 50 m, Arch. Crozet
Distribution: Arch. Crozet
Psylloglyphus (Psylloglyphus) parapsyllus Fain et Galloway
Psylloglyphus (Psylloglyphus) parapsyllus Fain & Galloway, 1993a: 160
Type locality: Motunau Island, Plateau Region
Distribution: New Zealand
Host: fleas in nests of Eudyptula minor and Pachyptila turtur
Psylloglyphus (Psylloglyphus) maculatus Fain et Beaucournu
Psylloglyphus (Psylloglyphus) maculatus Fain & Beaucournu, 1976: 184
Type locality: Ulugombak, Selangor, Malaysia
Distribution: Malaysia
Host: Medwayella robinsoni (Siphonaptera: Pygiopsyllidae)
Psylloglyphus (Psylloglyphus) reticulatus Fain et Beaucournu
Psylloglyphus reticulatus Fain & Beaucournu, 1976: 182
Type locality: Buhengere, Kivu, DR Congo
Distribution: DR Congo
Host: Ctenophthamus calceatus cabirus (Siphonaptera)
Psylloglyphus (Psylloglyphus) u. uilenbergi Fain
Psylloglyphus (Psylloglyphus) uilenbergi Fain, 1966: 162; Fain & Beaucournu, 1976: 181
Type locality: Madagascar, Iwato
Distribution: Madagascar
Hosts: Paractenophyllus petiti, Synopsyllus fonquerniei (Siphonaptera from tenrecs - Microgale and Setifer setosus)
Psylloglyphus (Psylloglyphus) uilenbergi kivuensis Fain et Beaucournu
Psylloglyphus (Psylloglyphus) uilenbergi kivuensis Fain & Beaucournu, 1976: 182; Fain &
Schwan, 1976: 634
366
Acarorum Catalogus IX
Type locality: Kivu (DR Congo)
Distribution: DR Congo, Gabon, Kenya
Hosts: Ctenophthalmus cabirus, Dinopsyllus apistus, D. echinus, D. lypusus, Listropsylla
dolosa, Stivalius sitianus, Xiphiopsylla levis (Siphonaptera)
Psylloglyphus (Psylloglyphus) vietnamensis Fain et Beaucournu
Psylloglyphus (Psylloglyphus) vietnamensis Fain & Beaucournu, 1972: 529
Type locality: Dalat, South Vietnam
Distribution: Vietnam
Host: Stivalius aporus rectodigitus (Siphonaptera) from Rattus rattus (Rodentia)
Subgenus Psylloglyphus (Hemimeropus) Fain et Beaucournu
Psylloglyphus (Hemimeropus) Fain & Beaucournu, 1976: 184
Type species: Psylloglyphus (Hemimeropus) hemimerus Fain et Beaucournu, 1976
Psylloglyphus (Hemimeropus) hemimerus Fain et Beaucournu
Psylloglyphus (Hemimerus) hemimerus Fain & Beaucournu, 1976: 184
Type locality: Butare
Distribution: Rwanda
Host: on Hemimerus bouvieri (Dermaptera: Hemimeridae), parasite on Cricetomys gambianus (Rodentia: Nesomyidae)
Subgenus Psyllobia Fain, Bartholomaeus, Cooke et Beaucournu
Psyllobia Fain, Bartholomaeus, Cooke et Beaucournu, 1990:
Type species: Psylloglyphus (Psylloglyphus) foveolatus Fain et Mason, 1989
Psylloglyphus (Psyllobia) foveolatus Fain et Mason
Psylloglyphus (Psylloglyphus) foveolatus Fain & Mason, 1989: 43; Fain et al., 1990:
Type locality: 12 km S of Launceston, Tasmania
Distribution: Australia (Tasmania, Victoria)
Hosts: phoretic on Pygiopsylla hoplia from Rattus rattus (Muridae), Bibikovana rainbowi,
Acanthopsylla r. rothschildi (Siphonaptera: Pygiopsyllidae)
Subgenus Tetrapsyllopus Fain et Beaucournu
Psylloglyphus (Tetrapsyllopus): Fain & Beaucournu, 1986: 257
Type species: Psylloglyphus (Tetrapsyllopus) micronychus Fain et Beaucournu, 1986
Psylloglyphus (Tetrapsyllopus) chiliensis Fain et Beaucournu
Psylloglyphus (Tetrapsyllopus) chiliensis Fain & Beaucournu, 1989: 139
Type locality: Isla Quila (Chiloë)
Distribution: Chile
Acarorum Catalogus IX 367
Hosts: Sphinctopsylla ares (Stephanocircidae), Ctenoparia inopinata (Hystrichopsyllidae)
(Siphonaptera)
Psylloglyphus (Tetrapsyllopus) micronychus Fain et Beaucournu
Psylloglyphus (Tetrapsyllopus) micronychus Fain & Beaucournu, 1986: 258
Type locality: Galletué (Prov. Malleco)
Distribution: Chile
Genus Trypetacarus Fain
Trypetacarus Fain, 1971: 283
Type species: Calvolia trypetidarum Cooreman, 1942
Trypetacarus trypetidarum (Cooreman)
Calvolia trypetidarum Cooreman, 1942: 1
Trypetacarus trypetidarum (Cooreman): Fain, 1971: 283
Type locality: Kutu. District des Bangala
Distribution: DR Congo
Host: Celidodacus fenestratus (Diptera: Trypetidae)
Subfamily Winterschmidtiinae Oudemans
Winterschmidtiidae Oudemans, 1923: 206
Calvoliinae Türk & Türk, 1957: 155 – Type: Calvolia Oudemans, 1911
Type genus: Winterschmidtia Oudemans, 1923
Genus Allocalvolia Fain et Rack
Allocalvolia Fain & Rack, 1987: 310
Type species: Allocalvolia habrocytus Fain et Rack, 1987
Allocalvolia bisculpturata (Mahunka)
Calvolia bisculpturata Mahunka, 1974a: 137; Eraky 1998: 248
Allocalvolia bisculpturatus (Mahunka): Fain & Rack 1987: 311
Type locality: Amedzofe
Distribution: Ghana
Allocalvolia habrocytus Fain et Rack
Allocalvolia habrocytus Fain & Rack, 1987: 311
Type locality: Eastern Austria
Distribution: Austria
Host: Habrocytus elevatus (Hymenoptera: Pteromalidae)
368
Acarorum Catalogus IX
Genus Calvolia Oudemans
Calvolia Oudemans, 1911: 187
Type species: Calvolia hagensis Oudemans, 1911
Calvolia antarctica Hughes
Calvolia antarctica Hughes, 1966: 45
Type locality: South Sandwich Islands
Distribution: South Sandwich Islands
Calvolia astoma Türk et Türk
Calvolia astoma Türk & Türk, 1957: 160; Hughes, 1962: 61
Type locality: Erlangen (Germany)
Distribution: Germany, Slovakia
Hosts: Formica rufa, Lasius fuliginosus (Hymenoptera: Formicidae)
Calvolia bulgarica Štorkan
Calvolia bulgarica Štorkan, 1935: 22; Zachvatkin, 1941: 257; Hughes, 1962: 61
Type locality: Bulgaria, Botev Summit (= Yumrukchal)
Distribution: Bulgaria, Russia
Host: Aphthona euphorbiae (Coleoptera: Chrysomelidae, Alticinae)
Calvolia calliphorae Vitzthum
Calvolia calliphorae Vitzthum, 1922: 539; Zachvatkin, 1941: 256; Hughes, 1962: 61
Type locality: Weimar
Distribution: Germany
Host: Calliphora vomitoria (Diptera: Calliphoridae)
Calvolia domicola Oshima
Calvolia domicola Oshima, 1979: 240
Type locality: Minami-ku, Yokohama
Distribution: Japan
Host: house dust
Calvolia elisabethae Zachvatkin
Calvolia elisabethae Zachvatkin, 1941: 254
Type locality: Staraya Bukhara (Uzbekistan)
Distribution: Georgia, Uzbekistan
Hosts: Ellobius talpinus, Prometheomys shaposhnikovi (Mammalia: Rodentia)
Calvolia elongata Volgin
Calvolia elongata Volgin, 1980: 167
Acarorum Catalogus IX 369
Type locality: Alma-Atinskaya Obl., Kaskelenskiy Rayon
Distribution: Kazakhstan
Host: manure
Calvolia fraxini Türk et Türk
Calvolia fraxini Türk & Türk, 1957:
Type locality: Munich
Distribution: Germany
Host: Leperisinus orni (Coleoptera: Curculionidae)
Calvolia furnissi Woodring
Calvolia furnissi Woodring, 1966: 76
Type locality: Moskow, Idaho
Distribution USA
Host: from the galleries and frass: “from the galleries and frass therein of Dendroctonus
pseudotsugae in Douglas fir”
Calvolia fusiformis Zachvatkin
Calvolia fusiformis Zachvatkin, 1941: 259; Hughes, 1962: 61
Type locality: Klinskiy Rayon, Moskovskaya Obl.
Distribution: Russia
Host: Blastophagus minor (Coleoptera: Curculionidae, Scolytinae)
Calvolia glabra Womersley
Calvolia glabra Womersley, 1941: 474
Type locality: Adelaide Zoo
Distribution: Australia
Host: “from a branchium of a Boa”
Calvolia goetzae Türk et Türk
Calvolia goetzae Türk & Türk, 1957: 163; Hughes, 1962: 61
Type locality: Moskow
Distribution: Russia, Slovakia
Hosts: Aphodius minor, A. prodromus (Coleoptera: Scarabaeidae)
Calvolia hagensis Oudemans
Calvolia hagensis Oudemans, 1911: 187; Vitzthum, 1920: 58; Zachvatkin, 1941: 359;
Hughes, 1962: 61; Fain, 1972: 229
Type locality: Hague
Distribution: Holland
370
Acarorum Catalogus IX
Calvolia hebeclinii (Sicher)
Glycyphagus hebeclinii Sicher, in Canestrini, 1899: 971
Calvolia hebeclinii (Sicher): Zachvatkin, 1941: 253; Hughes, 1962: 61
Type locality: Sicily
Distribution: Italy (Sicily)
Host: on plants
Calvolia heterocoma (Michael)
Tyroglyphus heterocomus Michael, 1903: 106
Czenspinskia heterocoma: Oudemans, 1927:
Calvolia heterocoma (Michael): Zachvatkin, 1941: 248; Hughes, 1962: 61
Type locality: New Forest, Hampshire
Distribution: England, Bulgaria, Slovakia
Calvolia kamenskii Zachvatkin
Calvolia kamenskii Zachvatkin, 1941: 254; Hughes, 1962: 61
Type locality: Bashkortistan
Distribution: Russia
Calvolia longipes Türk et Türk
Calvolia longipes Türk & Türk, 1957: 170
Type locality: Erlangen (Germany)
Distribution: Germany, Slovakia
Host: Chelidura acanthopygia (Dermaptera: Forficulidae)
Calvolia longireticulatae Ramaraju et Mohanasundaram
Calvolia longireticulatae Ramaraju & Mohanasundaram, 1999: 47
Type locality: Chidambaram, Tamil Nadu
Distribution: India
Host: Coleoptera, Meloidae gen. sp.
Calvolia marie Samšinák
Calvolia marie Samšinák, 1989: 105
Distribution: DR Congo
Host: Gymnometopina magna (Diptera: Sphaeroceridae)
Calvolia minima Türk et Türk
Calvolia minima Türk & Türk, 1957: 170; Hughes, 1962: 61
Type locality: Erlangen
Distribution: Germany
Acarorum Catalogus IX 371
Host: Scolytus intricatus (Coleoptera: Curculionidae, Scolytinae)
Calvolia miniopteri Dusbábek
Calvolia miniopteri Dusbábek, 1964: 228
Type locality: Ciertova diera
Distribution: Slovakia
Host: Miniopterus schreibersi (Chiroptera: Miniopteridae)
Calvolia priapus Volgin
Calvolia priapus Volgin, 1980: 161; Faradonbeh et al., 2018: 12
Type locality: Semirabad (Azerbaidzhan)
Distribution: Azerbaijan, Iran, Ukraine (Odessa)
Host: bird’s nests
Calvolia reticulata Zachvatkin
Calvolia reticulata Zachvatkin, 1941: 254; Hughes, 1962: 61
Type locality: Sussillon (Switzerland)
Distribution: Slovakia, Switzerland
Host: Osmia (Anthocopa) dalmatina (Hymenoptera:
Calvolia romanovae Zachvatkin
Calvolia romanovae Zachvatkin, 1941: 252
Calvolia tarsoinfracta Türk & Türk, 1957: 167 (syn. by Hughes, 1962)
Type locality: Moskow (Russia)
Distribution: Russia, Germany, New Zealand
Calvolia ruehmi Türk et Türk
Calvolia ruehmi Türk & Türk, 1957: 162; Hughes, 1962: 61; Woodring, 1966: 80
Type locality: Erlangen
Distribution: Germany
Host: Dryocoetes villosus (Coleoptera: Curculionidae, Scolytinae)
Calvolia solimani Eraky
Calvolia solimani Eraky, 1999: 215
Type locality: Aswan, Botanical Island
Distribution: Egypt
Host: Phoretic on Amitermes desertorum (Insecta)
Calvolia striata Vitzthum
Calvolia striata Vitzthum, 1914: 324; Zachvatkin, 1941: 260; Hughes, 1962: 61
Type locality: Ivory Coast
372
Acarorum Catalogus IX
Distribution: Italy (?), Ivory Coast
Hosts: Taphrorhynchus sp. (?), Pityogenes lepidus (Coleoptera: Curculionidae, Scolytinae)
Calvolia summersi Mostafa
Calvolia summersi Mostafa, 1971: 577
Type locality: Amazonas, Manaus
Distribution: Brazil
Host: Zethus binodis (Hymenoptera: Vespidae, Eumeninae)
Calvolia thraca Vitzthum
Calvolia thraca Vitzthum, 1922: 542; Zachvatkin, 1941: 255; Hughes, 1962: 61
Type locality: Bulgaria, Sofia
Distribution: Bulgaria, Russia
Host: Diptera
Calvolia tuberculata Zachvatkin
Calvolia tuberculata Zachvatkin, 1941: 248; Hughes, 1962: 61; Woodring, 1966: 80
Type locality: Georgia
Distribution: Georgia, Slovakia
Calvolia wichmanni Türk et Türk
Calvolia wichmanni Türk & Türk, 1957: 164
Type locality: Erlangen
Distribution: Germany
Host: Pyrrhidium sanguineum (Coleoptera: Cerambycidae)
Calvolia zachvatkini Volgin
Calvolia zachvatkini Volgin, 1980: 165
Type locality: Russia (Orichevsky Rayon, Kirovskaya Obl.)
Distribution: Kazakhstan, Russia
Host: hay and rodent’s nests
Calvolia zacheriana nom. nov. Eraky
Calvolia zaheri Eraky, 1998: 246; 1999: 217
Type locality: 47 km North of El-Kharga city, New Valley
Distribution: Egypt
Host: Psammotermes hypostoma (Isoptera: Rhinotermitidae)
Calvolia sp.
Hosts: Hybomitra affinis, H. epitates, H. lasiophthalma, H. nuda, Tabanus atratus (Canada)
Acarorum Catalogus IX 373
Genus Czenspinskia Oudemans
Genus Czenspinskia Oudemans, 1923 [syn. with Calvolia by Hughes 1962: 60; separate
by OConnor]
Donndorffia Oudemans, 1931: 203; [syn. by Hughes 1962: 60; Type is transversostriata]
Type species: Tyroglyphus heterocomus Michael, 1903
Czenspinskia transversostriata (Oudemans)
Donndorffia transversostriata Oudemans, 1931: 203
Czenspinskia transversostriata (Oudemans): Ripka et al., 2002: 278; Barbosa & de Moraes,
2021
Czenspinskia lordi Nesbitt, 1946: syn. by Hughes, 1962
Calvolia transversostriata (Oudemans): Hughes 1962: 48
Type locality: “Mares niet gevonen”
Distribution: Croatia, Holland, France, Hungary, Italy, Switzerland, Spain, Canada, USA,
Brazil, Australia
Genus Parawinterschmidtia Khaustov
Parawinterschmidtia Khaustov, 2000: 57
Type species: Calvolia kneissli Krausse, 1919
Parawinterschmidtia kneissli (Krausse)
Calvolia kneissli Krausse, 1919: 123; Zachvatkin, 1941: 255; Türk & Türk, 1957: 157;
Samšinák, 1957; Hughes, 1962: 61; Cooreman, 1963: 24; Woodring, 1966: 80
Parawinterschmidtia kneissli (Krausse): Khaustov, 2000: 57; Khaustov et al., 2018: 49
Type locality: Germany
Distribution: Belgium, Germany; “Czechoslovakia”, Poland, Crimea, Russia, Slovakia
Hosts: Lithobius forficatus (Chilopoda, Lithobiomorpha: Lithobiidae); Raphidia sp. (larva)
(Raphidioptera: Raphidiidae); Ips [Orthomicus] laricis, Brachyderes incanus Hylobius
abietis, Pissodes pini, P. piniphilus (Coleoptera: Curculionidae, Scolytinae), Thanasimus
formicarius (Coleoptera: Cleridae); Mesosa nebulosi, Rhagium sycophanta, R. inquisitor, Plagionotus detritus, Monochamus galloprovincialis, Spondylis buprestoides (Col.:
Cerambycidae), Crypticus quisquilius (Col.: Tenebrionidae), Myzia oblongoguttata
(Col.: Coccinellidae), Acanthocinus aedilis, Spondylis buprestoides, Sericus brunneus
(Elateridae)
Tomicus piniperda, Hylastes attenuates, Crossocerus vagabundus, C. leucostoma, Ectemnius cavifrons, E. cephalotes (Crabronidae), Sceliphron deforme (Sphecidae), Astata
kashmirensis (Astatinae)
Parawinterschmidtia sp. – Finland (Penttinen, Viiri & Moser, 2013)
374
Acarorum Catalogus IX
Genus Riemia Oudemans
Riemia Oudemans, 1925: 90
Type species: Riemia hesperidum Oudemans, 1925
Riemia hesperidum Oudemans
Riemia hesperidum Oudemans, 1925: 90
Type locality: Arnhem
Distribution: Holland
Genus Winterschmidtia Oudemans
Winterschmidtia Oudemans, 1923: 207
Afrocalvolia Fain & Elsen, 1971: 281 [Type is glossinarum; Syn. Khaustov 2000: 50]
Type species of Winterschmidtia Oudemans: Suidasia hamadryas Vitzthum, 1923
Winterschmidtia athiasae (Fain)
Afrocalvolia athiasae Fain, 1974: 70
Type locality: La Savane de Lamto
Distribution: Ivory Coast
Winterschmidtia brenyi (Cooreman)
Calvolia brenyi Cooreman, 1963: 32
Winterschmidtia brenyi (Cooreman): Khaustov 2000: 53
Type locality: Gembloux, Prov. de Namur)
Distribution: Belgium, Crimea
Host: Pteleobius vittatus (Coleoptera: Curculionidae, Scolytinae)
Winterschmidtia chaetoptelii Khaustov
Winterschmidtia chaetoptelii Khaustov, 2000: 55
Type locality: Crimea, Alushta
Distribution: Crimea
Host: Phoretic on Chaetoptelius vestitus (Coleoptera: Curculionidae)
Winterschmidtia circumspectans (Vitzthum)
Calvolia circumspectans Vitzthum, 1920: 56; Zachvatkin, 1941: 258; Hughes, 1962: 61
Afrocalvolia circumspectans (Vitzthum): Fain, 1974: 70
Type locality: Himalaya, Nilam-Dshong, NW Abhang .des Gaurisankar
Distribution: Tibet
Host: Ips stebbingi (Coleoptera: Curculionidae, Scolytinae)
Winterschmidtia crassisetosa Willmann
Winterschmidtia crassisetosa Willmann, 1941: 65
Acarorum Catalogus IX 375
Type locality: Italy
Distribution: Italy
Host: Phloeotribus scarabaeoides (Coleoptera: Curculionidae, Scolytinae)
Winterschmidtia elongata Türk et Türk
Winterschmidtia elongata Türk & Türk, 1957: 154
Type locality: Erlangen
Distribution: Germany
Winterschmidtia glossinarum (Fain et Elsen)
Afrocalvolia glossinarum Fain & Elsen, in Fain 1971: 281; Fain & Elsen, 1972: 74
Winterschmidtia glossinarum (Fain et Elsen): [Khaustov, 2000]
Type locality: Benzali nr. Kinshasa
Distribution: DR Congo
Hosts: Glossina fuscipes quanzensis, G. palpalis palpalis (Diptera: Glossinidae)
Winterschmidtia hamadryas (Vitzthum)
Suidasia hamadryas Vitzthum, 1923: 156
Winterschmidtia hamadryas (Vitzthum): Oudemans, 1923: 206; 1927: 246; Zachvatkin,
1941: 240; Khaustov, 2000: 51; Ahadyiat & Ostovan, 2005
Type locality: Germany
Distribution: Germany, Crimea, Iran
Host: burrows of Scolytus rugulosus; on Scolytus amyggdali (Coleoptera: Curculionidae,
Scolytinae)
Winterschmidtia lamtoensis (Fain)
Afrocalvolia lamtoensis Fain, 1974: 72
Type locality: La Savane de Lamto
Distribution: Ivory Coast
Winterschmidtia nataliae (Zachvatkin)
Calvolia nataliae Zachvatkin, 1941: 262; Cooreman, 1963: 28
Calvolia elliptica Zachvatkin, 1941: 262; Hughes, 1962: 61 – Type: Sviblovo near Moskow,
Russia, on Ips acuminatus (Coleoptera: Curculionidae, Scolytinae) Calvolia fraxini
Turk & Turk, 1957: 1957: 166; Hughes, 1962: 61 (syn. by Cooreman, 1963)
Winterschmidtia nataliae (Zachvatkin): Khaustov, 2000: 52
Type locality: Alekseevskoe Village near Moskow
Distribution: Belgium, Germany, Poland, Russia, Crimea
Hosts: Leperesinus [Hylesinus] fraxini, L. omi (Coleoptera: Curculionidae, Scolytinae)
Winterschmidtia tsetse (Fain et Elsen)
376
Acarorum Catalogus IX
Afrocalvolia tsetse Fain & Elsen, 1972: 75
Type locality: Kaniama, Lomami
Distribution: DR Congo
Hosts: Glossina fuscipes quanzensis (Diptera: Glossinidae)
Winterschmidtia villifronsi Khaustov
Winterschmidtia villifronsi Khaustov, 2000: 56
Type locality: Crimea, Yaltinskiy Gornolesnoy reserve
Distribution: Crimea
Host: Phoretic on Taphrorychus villifrons (Coleoptera)
Winterschmidtia zachvatkini Khaustov
Winterschmidtia zachvatkini Khaustov, 2000: 57
Type locality: Yalta
Distribution: Crimea
Host: Phoretic on Pityogenes calcaratus (Coleoptera: Curculionidae, Scolytinae)
Acarorum Catalogus IX 377
Index of Winterschmidtiidae
(Names in bold are accepted as valid)
Acalvolia 359
acaris, Vidia (Zethovidia) 356
acaris, Zethovidia 356
aegyptiana, Ensliniella 350
africana, Crabrovidia 346
Afrocalvolia 374
alatavicus, Neottiglyphus 362
Allocalvolia 367
amazonica, Macroharpa 352
americana, Acalvolia 359
anacardivorus, Vespacarus 353
andinus, Zethacarus 355
antarctica, Calvolia 368
arboricola, Calvolia 363
arboricola, Lenzia 363
arboricola, Oulenzia 363
argentinae, Zethacarus 355
arizonensis, Vidia (Zethovidia) 256
arizonensis, Zethovidia 356
arota, Macroharpa 352
asiatica, Ensliniella 350
astoma, Calvolia 368
athiasae, Afrocalvolia 374
athiasae, Winterschmidtia 374
australiensis, Psylloglyphus 365
aviophilus, Procalvolia 364
bakeri, Calvolia 363
bakeri, Oulenzia 363
bakeri, Oulenziella 363
benelux, Congovidia 360
bisculpturata, Calvolia 367
bisculpturata, Allocalvolia 367
bisculpturatus, Allocalvolia 367
boharti, Zethacarus 355
brasiliensis, Congovidia 360
brasiliensis, Crabrovidia 347
brasilensis, Vidia (Zethovidia) 356
brasilensis, Zethovidia 356
brenyi, Calvolia 375
brenyi, Winterschmidtia 375
brevapodemis, Vidia (Zethovidia) 357
brevapodemis, Zethovidia 357
bulgarica, Calvolia 368
calliphorae, Calvolia 368
Calvolia 368
Calvoliinae 346
chaetoptelii, Winterschmidtia 375
chiliensis, Psylloglyphopus 366
circumspectans, Afrocalvolia 375
circumspectans, Calvolia 375
circumspectans, Winterschmidtia 375
claggi, Neocalvolia 361
coccinella, Procalvolia 364
coccinellidarum, Congovidia 360
cocciphagus, Saproglyphus 362
compactilis, Vidia (Zethovidia) 357
compactilis, Zethovidia 357
concellaria, Crabrovidia 347
concellaria, Vidia 347
Congovidia 360
connivens, Sphexicozela 353
cooremani, Vidia 350
coperminensis, Neocalvolia 361
corsicana, Crabrovidia 347
Crabrovidia 346
crassisetosa, Winterschmidtia 375
crenulatus, Psylloglyphus 365
crossocerus, Crabrovidia 347
cubensis, Crabrovidia 347
Czenspinskia 373
Czenspinskiidae 346
dasyproctus, Crabrovidia 347
378
Acarorum Catalogus IX
dignotus, Ensliniella 350
diminuta, Vidia (Zethovidia) 257
diminuta, Zethovidia 357
domicola, Calvolia 367
Donndorffia 373
ectemnii, Crabrovidia 348
elisabethae, Calvolia 368
ellipsus, Zethacarus 355
elliptica, Calvolia 375
elongata, Calvolia 368
elongata, Winterschmidtia 375
Ensliniella 350
Ensliniellinae 346
Euvidia 346
faini, Neosuidasia 359
floricola, Ensliniella 350
floridae, Vidia (Zethovidia) 357
floridae, Zethovidia 357
foveolatus, Psylloglyphus 366
fraxini, Calvolia 369
fulvipes, Vespacarus 353
funebris, Monobiacarus 352
furnissi, Calvolia 369
fusca, Vidia (Zethovidia) 357
fusca, Zethovidia 357
fusiformis, Calvolia 369
Gambacarus 361
glabra, Calvolia 369
glossinae, Congovidia 360
glossinarum, Afrocalvolia 375
glossinarum, Winterschmidtia 375
goetzae, Calvolia 369
gossypii, Oulenzia 363
gussakovskii, Crabrovidia 348
gussakovskii, Vidia (Crabrovidia) 348
habrocytus, Allocalvolia 367
hagensis, Calvolia 369
hamadi, Procalvolia 364
hamadryas, Suidasia 375
hamadryas, Winterschmidtia 375
hebeklinii, Calvolia 370
hebeklinii, Glycyphagus 370
Hemimeropus 366
hemimerus Psylloglyphus 366
hesperidum, Riemia 374
heterocoma, Calvolia 370
heterocoma, Czenspinskia 370
heterocomus, Tyroglyphus 370
hirsuta, Crabrovidia 348
hirsuta, Vidia 348
histrio, Vespacarus 353
huettleri, Gambacarus 361
inermis, Neocalvolia 361
insularis, Monobiacarus 352
jiju, Kurosaia 351
juliani, Crabrovidia 348
juliani, Vidia 348
kamenskii, Calvolia 367
Kennethiella 351
kivuensis, Psylloglyphus uilenbergi 362
kneissli, Calvolia 373
kneissli, Parawinterschmidtia 373
koenigi, Ensliniella 351
kostylevi, Ensliniella 351
Kurosaia 351
laevapodemis, Vidia (Zethovidia) 357
laevapodemis, Zethovidia 357
lamtoensis, Afrocalvolia 375
lamtoensis, Winterschmidtia 375
latimanus, Crabrovidia 348
latimanus, Vidia 348
leclercqi, Crabrovidia 349
Lenzia (praeoccup.) 363
lineata, Vidia 355
lomamiensis, Congovidia 360
longipes, Calvolia 370
longireticulatae, Calvolia 370
longiseta, Oulenziella 363
lordi, Czenspinskia 373
Macroharpa 352
maculatus, Psylloglyphus 365
marie, Calvolia 370
mexicana, Vidia (Zethovidia) 358
mexicana, Zethovidia 358
Acarorum Catalogus IX 379
micronychus, Psylloglyphus (Tetrapsyllopus) 367
minimoferus, Vespacarus 354
miniopteri, Calvolia 371
miscogaster, Zethacarus 355
Monobiacarus 352
nataliae, Calvolia 375
nataliae, Winterschmidtia 375
neglectus, Saproglyphus 362
Neocalvolia 361
Neosuidasia 361
Neottiglyphus 362
ninae, Crabrovidia 349
ninae, Vidia 349
oculabsensa, Vidia (Zethovidia) 358
oculabsensa, Zethovidia 358
oudemansi, Crabrovidia 349
Oulenzia 363
Oulenziella 363
Oulenziidae 346
Oulenziinae 363
pachymapoda, Macroharpa 352
palpognatha, Macroharpa 352
paraguayensis, Vidia (Zethovidia) 358
paraguayensis, Zethovidia 358
paranicus, Zethacarus 356
parapsyllus, Psylloglyphus 365
parasitica, Ensliniella 351
Parawinterschmidtia 373
parvula, Vidia (Zethovidia) 358
parvula, Zethovidia 358
pedestris, Vespacarus 351
popovi, Vidia (Crabrovidia) 349
popovi, Crabrovidia 349
priapus, Calvolia 371
Procalvolia 364
promina, Vidia (Zethovidia) 358
promina, Zethovidia 358
Psyllobia 366
Psylloglyphus 364
quadridens, Monobiacarus 353
reticulata, Calvolia 371
reticulata, Vidia (Zethovidia) 358
reticulata, Zethovidia 358
reticulatus, Psylloglyphus 365
Riemia 374
romanovae, Calvolia 364, 371
rubi, Crabrovidia 349
rubi, Vidia 349
ruehmi, Calvolia 371
rufovestis, Vespacarus 354
rugocoxa, Vidia (Zethovidia) 359
rugocoxa, Zethovidia 359
saecularis, Vespacarus 354
salvadorensis, Vidia (Zethovidia) 359
salvadorensis, Zethovidia 359
Saproglyphidae 346
Saproglyphinae 359
Saproglyphus 362
servalina, Congovidia 360
setosus, Saproglyphus 362
solimani, Calvolia 371
Sphexicozela 353
squamata, Acalvolia 359
squamata, Vidia 359
stoeckherti, Vidia 355
striata, Calvolia 371
summersi, Calvolia 372
tarsoinfracta, Calvolia 364
Tetrapsyllopus 366
texana, Crabrovidia 349
texana, Vidia 349
thomasi, Crabrovidia 350
thomasi, Vidia 350
thraca, Calvolia 372
tigris, Vespacarus 354
toltecus, Vespacarus 354
transversostriata, Calvolia 373
transversostriata, Czenspinskia 373
transversostriata, Donndorffia 370
trisetosa, Ensliniella 351
trisetosa, Kennethiella 351
Trypetacarus 367
trypetidarum, Calvolia 367
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Acarorum Catalogus IX
trypetidarum, Trypetacarus 367
tsetse, Afrocalvolia 376
tsetse, Winterschmidtia 376
tuberculata, Calvolia 372
uilenbergi, Psylloglyphus 365
undulata, Vidia 355
utahensis, Crabrovidia 350
utahensis, Vidia (Euvidia) 350
vagus, Vespacarus 354
Vidia 355
vietnamensis, Psylloglyphus 366
villifronsi, Winterschmidtia 376
waldorphae, Calvolia 362
waldorphae, Saproglyphus 362
wichmanni, Calvolia 372
Winterschmidtia 374
Winterschmidtiidae 346
Winterschmidtiinae 367
zachvatkini, Calvolia 372
zachvatkini, Winterschmidtia 376
zaheri, Calvolia Oud. 364
zaheri, Calvolia Eraky (praeoccup.) 372
zaheri, Procalvolia 364
zaheriana, Calvolia 372 (nom. nov.)
Zethacarus 352
Zethovidia 353
Distribution of Winterschmidtiidae
EUROPE
Armenia (Klompen & OConnor, 1995) - Ensliniella parasitica
Austria (Fain, 1973; Fain & Rack, 1987; Klompen & OConnor, 1995; Moser et al., 2005)
- Allocalvolia habrocytus, Crabrovidia ectemnii, Ensliniella kostylevi
Azerbaidjan (Volgin, 1980) - Calvolia priapus
Belarus (Vitzthum, 1925) – Ensliniella kostylevi
Belgium (Cooreman, 1948, 1963; Fain, 1972, 1973; Fain, Hurst et al. 1997; Klompen &
OConnor, 1995) – Congovidia coccinellidarum, Crabrovidia concellaria, C. ectemnii,
Ensliniella floricola, E. kostylevi Winterschmidtia brenyi, W. nataliae
Bulgaria (Vitzthum, 1922; Štorkan, 1935; Angelkova, 1988; Klompen & OConnor, 1995)
– Calvolia bulgarica, C. heterocoma, C. thraca, C. tuberculata, Ensliniella kostylevi,
Procalvolia zacheri (= Calvolia romanovae)
Croatia (Fain, 1972, 1973; Klompen & OConnor, 1995; Ripka et al., 2002; ) – Crabrovidia
ectemnii, C. popovi, Czenspinskia transversostriata, Ensliniella floricola
Finland (Penttinen, Viiri & Moser, 2013; Linnakoski et al., 2016) – Probonomoia pini
France (Fain, 1972, 1973; Klompen & OConnor, 1995) – Crabrovidia concellaria, C.
ectemnii, Ensliniella floricola, Е. kostylevi
Corsica (Fain, 1973) – Crabrovidia corsicana
Georgia (Zachvatkin, 1941) – Calvolia elisabethae, C. tuberculata
Germany (Vitzthum, 1922, 1923, 1925; Türk & Türk, 1957) – Calvolia calliforae,C. fraxini,
C. minima, C. romanovae (= tarsoinfracta), Ensliniella parasitica, Procalvolia zacheri,
Winterschmidtia hamadryas, W. nataliae
Great Britain (Fain & Johnston, 1974; Fain, Hurst et al., 1997; Fain & Greenwood, 1991)
– Calvolia waldorfae, Congovidia coccinellidarum, Probonomoia pini
Greece (Klompen & OConnor, 1995) - Ensliniella floricola, Е. kostylevi
Acarorum Catalogus IX 381
Crete (Fain, 1973) - Crabrovidia ectemnii
Corfu (Fain, 1971) – Crabrovidia oudemansi, Vidia lineata
Holland (Oudemans, 1909, 1911, 1917, 1925, 1931; Klompen & OConnor, 1995) – Acalvolia squamata, Calvolia hagensis, Czenspinskia transversostriata, Ensliniella kostylevi,
E. parasitica, Myianoetus dionychus, M. simplex, Riemia hesperidum, Vidia lineata
Hungary (Mahunka, 1970; Ripka et al, 2002) - Sphexicozela connivens, Czenspinskia
transversostriata
Italy (Oudemans, 1905; Fain, 1973; Baker & Cunliffe, 1960) – Calvolia hebeklinii, C. striata,
Crabrovidia popovi, Czenspinskia transvertostriata, Ensliniella parasitica, Saproglyphus
neglectus, Vidia lineata, V. undulata
Sicily (Sicher, in Canestrini, 1899) – Calvolia hebeclinii
Sardinia - Calvolia hebeclinii
Poland (Fain, Hurst et al., 1997; Kiełczewski, Moser & Wiśniewski, 1983; Kiełczewski
& Wiśniewski, 1971, 1980; Haitlinger, 1904; Michalski & Ratajczak, 1989) – Divilia
exigua, Probonomoia pini
Russia (Zachvatkin, 1941; Türk & Türk, 1957; Krivoluckiy, 1975; Bugrov, 1997; Klimov,
1998; Ermilov & Mokrousov, 2008, 2010; Khaustov et al., 2018) – Calvolia elisabethae,
C. goetzae, C. kamenskii, C. romanovae, C. thraca, Crabrovidia gussakovskii, Parawinterschidtia kneissli, Probonomoia pini, P. sibirica, Winterschmidtia nataliae
Crimea (Zachvatkin, 1941; Khaustov, 2000) - Crabrovidia gussakovskii, Parawinterschidtia
kneissli, Winterschmidtia brenyi, W. chaetoptelii, W. villifronsi, W. zachvatkini
Kuril Islands (Klimov, 1998) - Crabrovidia gussakovskii (Iturup)
Slovakia (Dusbabek, 1964; Kaluz & Žuffa, 1986; Sidor, 1986; Zamec, 2013, 2014) – Calvolia
astoma, C. fraxini, C. goetzae, C. heterocoma, C. kneissli, C. longipes, C. miniopteri,
C. reticulata, C. tarsoinfracta, Ensliniella parasitica
Spain (Fain, 1973, Klompen & OConnor, 1995) - Crabrovidia ectemnii, C. popovi, Ensliniella parasitica
Switzerland (Fain, 1973) - Crabrovidia ectemnii
Turkey (Fain, 1973; Klompen & OConnor, 1995) - Crabrovidia popovi, Ensliniella dignotus,
E. floricola, Е. kostylevi
Ukraine (Zachvatkin, 1941; Khaustov, 2000; Rusina & Orlova, 2011) - Crabrovidia popovi,
Ensliniella kostylevi, Sphexicozela connivens, Winterschmidtia hamadryas, W. natalyae,
AFRICA
Algeria (Klompen & OConnor, 1995) – Ensliniella kostylevi
DR Congo (Cooreman, 1942; Fain, 1971; Fain & Elsen, 1972, 1973; Fain & Beaucournu,
1976; Samšinák, 1989) – Calvolia marie, Congovidia glossinae, Congovidiella hieroglypha, Psylloglyphus uilenbergi kivuensis, Winterschmidtia (= Afrocalvolia) glossinarum, Trypetacarus trypetidarum
Egypt (Baker & Cunliffe, 1960; Sevastianov & Gad-Hamada-Hassa, 1989; Hassan & Sevastianov, 1989; Eraky, 1998; El-Kawas & Negm, 2018) – Calvolia solimani, C. zaheri,
Ensliniella aegyptiaca, Procalvolia hamadi
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Acarorum Catalogus IX
Gambia (Mahunka, 1975) - Gambacarus huettleri
Ghana (Mahunka, 1974) – Allocalvolia bisculpturata
Ivory Coast (Fain, 1974) – Winterschmidtia (= Afrocalvolia) athiasae, W. lamtoensis
Madagascar (Fain, 1966; Fain & Beaucournu, 1976) - Psylloglyphus uilenbergi
Kenya (Fain & Schwan, 1976) – Psylloglyphus uilenbergi kivuensis
Morocco (Fain, 1972, 1973; Baker & Cunliffe, 1960) – Crabrovidia ectemnii, Ensliniella
koenigi
Mozambique (Meyer & Rodrigues, 1965) – Oulenziella bakeri (syn. Oulenzia gossypii)
Rwanda (Fain & Beaucournu, 1976) - Psylloglyphus (Hemimerus) hemimerus
South Africa (Fain, 1973) - Crabrovidia dasyproctus
ASIA
China (Chang et al., 2017, 2020) - Winterschmidtiidae
Taiwan (Fain, 1973) – Crabrovidia dasyproctus
Cyprus (Fain, 1973) - Crabrovidia ectemnii, C. leclercqi
India (Hughes, 1962; Fain, 1973) - Crabrovidia dasyproctus, Calvolia bakeri
Indonesia (Oudemans, 1928; Hughes, 1962) – Oulenzia arboricola, Calvolia bakeri
Iran (Ahadiyat & Ostovan, 2006; Faradonbeh et al., 2018; Nazari, Monfared, Nemati
& Azhari, 2019) – Calvolia priapus, Crabrovidia oudemansi, Vidia lineata, Winterschmidtia hamadryas
Japan (Fain, 1973; Okabe & OConnor, 2002; Abe & Okabe, 2006) – Crabrovidia ectemnii
[Okinawa], Ensliniella asiatica, Kurosaia jiju
Kazakhstan (Volgin, 1974) - Neottiglyphus alatavicus
Malaysia (Fan et al., 2012) – Oulenzia sp.
Philippines (Fain, 1973) - Crabrovidia dasyproctus
Singapour (Fain, 1973) – Crabrovidia dasyproctus
Tadjikistan (Klompen & OConnor, 1995) - Ensliniella parasitica
Uzbekistan (Zachvatkin, 1941) - Crabrovidia popovi, Ensliniella parasitica
Vietnam (Fain & Beaucournu, 1972) – Psylloglyphus vietnamensis
NORTH AMERICA
Bermudas (Fain & Schuster, 1983) – Neocalvolia inermis
Canada (Nesbitt, 1946; Baker & Cunliffe, 1960; OConnor & Eickwort, 1988; Mullen,
Leprince & OCoonor, 1989) – Crabrovidia latimanus, Vespacarus vagus, Vidia sp. A
and B, Calvolia sp.
Mexico (Baker & Cunliffe, 1960; Mostafa, 1970b, 1971; Fain, 1972, 1973) – Crabrovidia
ectemnii, Monobiacarus funebris, M. insularis, Zethacarus boharti, Z. ellipsus, Z.
miscogaster, Zethovidia floridae
USA (Baker & Cunliffe, 1960; Baker, 1964; Woodring, 1966; Mostafa, 1967; Delfinado
& Baker, 1976; Fain & Philips, 1978; OConnor & Eickwort, 1988; Fan, George &
Kumarasinghe, 2010) – Acalvolia americana, Calvolia furnissi, Crabrovidia rubi, C.
Acarorum Catalogus IX 383
texana, C. thomasi, C. utahensis, Monobiacarus quadridens, Sapracarus tuberculatus,
Vespacarus anacardivorus, V. histrio, V. minimoferus, V. pedestris, V. tigris, V. toltecus
SOUTH AND CENTRAL AMERICA
Argentina (Mostafa, 1971; Fain, 1972, 1973) - Crabrovidia ectemnii, Zethacarus andinus,
Z. argentinae, Z. haywardi, Zethovidia parvula
Bolivia (Fain, 1972, 1973) - Crabrovidia ectemnii
Brazil (Mostafa, 1970b; 1971; Fain, 1972, 1973; Fain & Camerik, 1978; Barbosa & de
Moraes, 2021) – Acalvolia squamata, Calvolia summersi, Crabrovidia brasiliensis,
C. crossocerus, C. ectemnii, Czenspinskia transversostriata, Dichotomiopus biiaggioi,
Macroharpa amazonica, M. arota, M. pachymapoda, M. palpognatha, Oulenzia arboricola, Oulenziella bakeri, O. longiseta, Zethacarus paranicus, Z. smithii, Zethovidia
oculabsena, Z. brasilensis, Z. diminuta, Z. promina
Chile (Fain & Beaucournu, 1986, 1989) – Psylloglyphus (Tetrapsyllopus) chiliensis, P. (T.)
micronychus
Colombia (Fain, 1972, 1973) - Crabrovidia ectemnii
Costa Rica (Fain, 1972, 1973) - Crabrovidia ectemnii
Cuba (Fain, 1973) - Crabrovidia cubensis
El Salvador (Mostafa, 1970a) - Zethovidia salvadorensis
French Guiana (Mostafa, 1970b) - Zethovidia acaris
Guiana (Fain, 1972, 1973) - Crabrovidia ectemnii
Honduras (Fain, 1972, 1973) - Crabrovidia ectemnii
Jamaica (Fain, 1973) - Crabrovidia ectemnii
Panama (Mostafa, 1970b) – Zethovidia fusca
Paraguay (Mostafa, 1970a) - Zethovidia diminuta, Z. paraguayensis
Puerto Rico (Fain, 1973) - Crabrovidia ectemnii
Suriname (Mostafa, 1970b) - Zethovidia brevapodemis
Venezuela (Mostafa, 1970b) - Zethovidia reticulata
OCEANIA
Australia (Womersley, 1941; Fain, 1989; Fain, Bartholomaeus, Cooke & Beaucornu,
1990) – Calvolia glabra, Czenspinskia transversostriata, Psylloglyphus australiensis,
P. foveolatus [Tasmania, Victoria], Saproglyphus cocciphagus
New Zealand (Fain & Galloway, 1973a, 1973b) - Psylloglyphus (Psylloglyphus) parapsyllus
ANTARCTICA
South Georgia (Hughes, 1970) – Neocalvolia claggi
South Shetlands (Rosas & Casanueva, 1993) - Neocalvolia coperminensis
South Sandwich Islands (Hughes, 1966) - Calvolia antarctica
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Acarorum Catalogus IX
Hosts of Winterschmidtiidae
Insecta
Coleoptera
Fam. Scarabaeidae
Aphodius minor – Calvolia goetzae
prodromus - Calvolia goetzae
Cetonia aurata – Vidia sp.
Fam. Curculionidae
Subfam. Scolytinae
Hylesinus fraxini – Winterschmidtia nataliae
Ips stebbingi – Winterschmidtia circumspectans
Leperisinus omi – Winterschmidtia nataliae, Calvolia fraxini
Orthomicus laricis – Parawinterschmidtia kneissli
Pityogenes calcaratus - Winterschmidtia zachvatkini
Pteleobius vittatus – Winterschmidtia brenyi
Scolytus amygdali – Winterschmidtia hamadryas
S. intricatus – Calvolia minima
Fam. Coccinellidae
Adalia bipunctata - Congovidia benelux
Calvia duodecimguttata – Divilia exigua
Chilocorus renipustulatus – Divilia occidentalis
Halyzia sedecimguttata - Congovidia coccinellidarum
Fam. Cerambycidae
Mesosa nebulosa - Parawinterschmidtia kneissli
Dermaptera
Fam. Labiduridae
Chelidura acanthopygia – Calvolia longipes
Labidura riparia - Procalvolia hamadi
Hymenoptera
Fam. Megachilidae
Megachile pilicrus - Vidia lineata
Megachile (Litomegachile) texana – Crabrovidia texana
Megachile (Sayapis) fidelis - Crabrovidia hirsuta
Megachile (Xeromegachile) rubi – Crabrovidia rubi
Fam. Crabronidae
Crabro solenius - Crabrovidia gussakovskii
Cerceris arenaria – Crabrovidia concellaria
C. rubida – Crabrovidia leclercqi
Cerceris sp. - Crabrovidia oudemansi
Acarorum Catalogus IX 385
Crossocerus elongatus – Crabrovidia corsicana
C. porexus - Crabrovidia crossocerus
Dasyproctus agilis orientalis – Crabrovidia dasyproctus
D. bipunctatus lugubris - Crabrovidia dasyproctus
D. buddha - Crabrovidia dasyproctus
D. cevirus - Crabrovidia dasyproctus
D. westermanni - Crabrovidia dasyproctus
Ectemnius carinatus - Crabrovidia ectemnii
E. cavifrons - Crabrovidia ectemnii
E. centralis - Crabrovidia ectemnii
E. continuus [= Crabro vagus auct.] - Crabrovidia ectemnii, C. gussakovskii
E. craesus - Crabrovidia ectemnii
E. cubiceps – Crabrovidia brasiliensis
E. [Crabro] fossorius - Crabrovidia gussakovskii
E. guttatus - Crabrovidia ectemnii
E. hypsae - Crabrovidia ectemnii
E. impressus - Crabrovidia ectemnii, C. gussakovskii, C. popovi
E. laevigatus - Crabrovidia ectemnii
E. lapidaries - Crabrovidia ectemnii
E. schettereri - Crabrovidia ectemnii
E. semipunctatus - Crabrovidia ectemnii
E. singularis - Crabrovidia ectemnii
E. sonorensis - Crabrovidia ectemnii
E. sexcinctus - Crabrovidia ectemnii
Lestica alata - Crabrovidia popovi
L. clypeata – Crabrovidia popovi
L. cubensis – Crabrovidia cubensis
L. wollmanni - Crabrovidia popovi
Sphex hemorrhoidalis – Crabrovidia africana
Fam. Pteromalidae
Habrocytus elevatus – Allocalvolia habrocytus
Fam. Colletidae
Hylaea (Prosopis) conformis – Vidia undulata
Fam. Vespidae
Subfam. Eumeninae
Allodynerus floricola - Ensliniella floricola
A. delphinalis – Ensliniella parasitica
A. mandschuricus – Ensliniella asiatica
Subfam. Zethinae
Zethus andinus – Zethacarus andinus
Z. carbonarius – Zethovidia acaris
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Acarorum Catalogus IX
Z. coeruleopennis – Z. brazilensis
Z. d. diminutus – Z. diminuta
Z. dimunutus santaremae – Z. diminuta
Z. haywardi – Zethacarus argentinae
Z. guerreroi arizonensis – Zethovidia arizonensis
Z. guerreroi mayorum – Zethacarus boharti
Z. orizabae – Z. ellipticus
Z. miscogaster – Z. miscogaster
Z. smithii – Z. paranicus
Z. arotus – Macroharpa arota
Z. biglumis ferrugineus – Macroharpa amazonica
Z. lobulatus – Macroharpa pachymapoda
Z. orizabae – Zethacarus ellipsus
Z. roridus – Macroharpa palpognatha
Subfam. Polystinae
Polystes dominula - Sphexicozela connivens
P. gallica - Sphexicozela connivens
P. nimpha - Sphexicozela connivens
Fam. Eulophidae
Melittobia acasta – Ensliniella parasitica
Diptera
Fam. Trypetidae
Celidodacus fenestratus - Trypetacarus trypetidarum
Fam. Glossinidae
Glossina fuscipes quanzensis - Winterschmidtia glossinarum, W. tsetse, Congovidia glossinae
Glossina palpalis palpalis - Winterschmidtia glossinarum
Fam. Sphaeroceridae
Gymnometopina magna - Calvolia marie
Fam. Tabanidae
Hybomitra epistates – Vidia sp. A, Calvolia sp.
H. affinis – Calvolia sp.
H. lasiophthalmus - Calvolia sp.
H. microcephala - Vidia sp. A
H. nuda - Calvolia sp.
Tabanus atratus - Calvolia sp.
T. quinquevittatus - Vidia sp. B
Siphonaptera (phoretic acari on parasites)
Notiopsylla kerguelenensis – Notiopsyllopus segermanae, Psylloglyphus crenulatus
Synopsyllus fonquerniei – Psylloglyphus u. uilenbergi
Acarorum Catalogus IX 387
Paractenopsyllus petiti - Psylloglyphus u. uilenbergi
Medwayella robinsoni – Psylloglyphus maculatus
Stivalius aporus rectodigitus – Psylloglyphus vietnamensis
Pygiopsylla hoplia – Psylloglyphus foveolatus
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Acknowledgements
This volume required the support of many distinguished specialists, to mention only a few:
A.Fain, J.Cooreman, R.Haitlinger, S.Mahunka, N.J.Fashing, K.Samšinák, A.A.Khaustov,
U. Gerson, B.M.OConnor, P.B.Klimov, J.de la Cruz, and others. To all of them the author
is immensely grateful.
ACARORUM
CATALOGUS
IX
The ninth volume of this series "Acarorum Catalogus" is
devoted to the Superfamilia Shizoglyphoidea
(Schizoglyphidae). Histiostomatoidea (Histiostomatidae,
Guanolichidae). Canestrinioidea (Canestriniidae,
Chetochelacaridae, Lophonotacaridae, Heterocoptidae).
Hemisarcoptoidea (Chaetodactylidae, Hyadesiidae,
Algophagidae, Hemisarcoptidae, Carpoglyphidae,
Winterschmidtiidae). About 1268 species of 225 genera are
listed therein, with synonymies and citations in various
papers. The book also contains a list of by-country
distributions, type localities, hosts of the parasitic species
and lists of authors who published on any known species of
these families. The last Xth volume of the same series is
under way.