Zhang Li: Hong Kong Bryophytes 1
AN UPDATED AND ANNOTATED INVENTORY OF HONG KONG BRYOPHYTES
ZHANG Li
Department of Ecology & Biodiversity, The University of Hong Kong, Pokfulam Road, Hong Kong
S.A.R., China, and South China Institute of Botany, The Chinese Academy of Sciences, Guangzhou
510650, Guangdong Province, China
ABSTRACT
This updated and annotated inventory of Hong Kong bryophytes is based on available specimens and
related literature. All taxa were critically considered before acceptance. A total of 360 species and 12
infraspecific taxa in 70 families and 159 genera of bryophytes are accepted to occur in Hong Kong. In
addition, nineteen previously reported species are excluded from the local flora, and 20 species are
treated as doubtful records. Four families, 18 genera and 46 taxa are reported as new additions to Hong
Kong, of which Barbella convolvens is new to China, and four species, Acroporium secundum,
Leucophanes glaucum, Mitthyridium fasciculatum, and Syrrhopodon spiculosus, are new to mainland
China (excluding Hainan, Taiwan and South China Sea Islands). Ectropothecium nervosum is reduced
to a synonym of E. zollingeri. For each accepted species, information on examined specimens or
relevant references or both, distribution ranges within China and worldwide are supplied.
INTRODUCTION
Hong Kong is a Special Administrative Region of China lying on the south China coast, 22°9′ 22°37′N, 113°52′ - 114°30′ S, with a total land area of about 1,100 km2. It consists of four
geographical units: a section of the mainland (New Territories and Kowloon), Hong Kong Island,
Lantau Island, and some 230 other small islands (Outlying Islands). The landscape of Hong Kong is
generally rugged and hills occupy about 75% of the total land area. Tai Mo Shan, in the middle of the
New Territories, is the highest mountain with the summit reaching 957 m a.s.l.. Hong Kong has a
subtropical climate with distinct humid and dry seasons. The mean annual rainfall ranges from 1300
mm at Waglan Island in the extreme southeast of Hong Kong waters to more than 3000 mm near the
summit of Tai Mo Shan, with about 80% falling between May and September. Mean annual temperate
ranges from 22.8°C in lowland Kowloon to 18.3°C near the summit of Tai Mo Shan. Subzero
temperatures and frost occur annually at the highest altitudes and less frequently down to around 400 450 m on Tai Mo Shan, and down to sea-level in the northern New Territories. The primary forests
which used to cover Hong Kong were cleared centuries ago and the vegetation today consists of
secondary forests (10%), shrubland (37%), grassland (16%) and plantations (5%) (Ashworth et
al.,1993; Corlett, 1992, 2000; Dudgeon and Corlett, 1994; Hong Kong Observatory, 2000; Zhang et
al.,1989).
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2 Memoirs of the Hong Kong Natural History Society 26, 2003
The earliest bryological study in Hong Kong can be dated back to 1848 (Wilson, 1848). The history of
Hong Kong bryology has been well reviewed recently by Yip (1996) and Zhang and Lin (1997). Zhang
and Lin (1997) also published the first detailed checklist of Hong Kong bryophytes, which was totally
based on publication records up to early 1996 (abbreviated hereafter as the 1997 checklist). Before that,
there existed three incomplete lists compiled by So and Yip (1995b), So and Zhu (1996b) and Zhu and
So (1996). The 1997 checklist recognized 198 species, 3 infraspecific taxa and 3 doubtful species of
mosses in 94 genera and 36 families, and 129 species of liverworts and hornworts in 53 genera and 29
families.
Since the middle of the 1990s, there has been a striking increase in the bryological research in Hong
Kong and a number of papers have been published (cf. inventory text). The present project was started
in November 1997. Since then, extensive and systematic field surveys in most parts of Hong Kong
have been carried out and a lot of new records have been found. At the same time, some previous
records have been confirmed or excluded by checking the historical specimens. These studies in the
past three years have made it worthwhile to make an updated and detailed inventory of the local
bryoflora, although there are still some previous records waiting for verification, and some new
collections waiting for determination. Therefore the present inventory is still in progress and is not yet
finalized.
Although the present inventory is not a checklist, the third of the principles for compiling checklists
suggested by Koponen et al. (1998) was followed. This states: “Popularizing books and plant
sociological papers lacking documentation should be omitted or used very critically as a basis of
checklists …”. Following this principle, I treat species based on such sources as So (1995a) and Zhu
and So (1996) as doubtful records, unless I have found evidence to confirm or reject them during my
study, with the exception of species of liverworts and hornworts in Zhu and So (1996), which cited
voutcher specimens. Therefore, some species accepted in the 1997 checklist are now treated as
doubtful species, for example, Taxiphyllum inundatum.
In summary, two categories of species were included in the present inventory. The first category is
those species based on specimens which I have studied, and the second is species which were based on
publication records with accompanying specimen citations.
The purposes of this inventory are: (1) to present an overall summary of the species diversity of Hong
Kong bryophytes, (2) to provide updated information on their localities, habitats, substrates, and
distributions, i.e. the basic information required for conservation management.
MATERIALS & METHODS
Field surveys
From November 1997 to April 2000, extensive and systematic field surveys were carried out in various
parts of Hong Kong. Two different survey methods, general survey and sample survey, were employed.
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Zhang Li: Hong Kong Bryophytes 3
General surveys were carried out in as many localities and environments as possible (including 61
small outlying islands), whereas sample surveys were carried out in selected localities and
environments, including secondary forests, feng shui woods, shrublands, grasslands, agricultural
habitats, and urban habitats, following standard methods (Zhang, 2001) for the ecology of bryophyte
communities. In total over 3,300 bags of specimens were collected, and they will be deposited in the
Herbarium of The University of Hong Kong (HKU), with duplicates in herbaria of Kadoorie Farm &
Botanic Garden (KFBG), and South China Institute of Botany, The Chinese Academy of Sciences,
Guangzhou (IBSC).
Examination of specimens
Most collections made in field surveys were examined microscopically by myself. Those identified
include most mosses and a small part of the liverwort specimens collected during general surveys, and
almost all collections made in sample plots. In addition, some historical specimens deposited in the
herbaria of Harvard University (FH), British Museum (BM), IBSC, Hong Kong Baptist University
(HKBU), and The Chinese University of Hong Kong (CUHK) have been consulted. In total about
2,370 bags of specimens have been identified.
Literature review
References considered in this present inventory only include those publications which appeared after
1997, and those with specimen citations which were overlooked by Zhang and Lin (1997).
FEATURES OF THE PRESENT INVENTORY
The 1997 checklist was the basis of the present inventory. The present one (especially mosses) is based
mainly on either my own collections and specimens deposited in the herbaria of BM, CUHK, FH,
HKBU, IBSC and KFBG, or pertinent references with documentation of Hong Kong specimens.
Names lacking specimen documentation were excluded from the present inventory and are treated as
doubtful records. I have paid less attention to the liverworts because the Hepatica Flora of Hong Kong
was dealt with by colleagues in the Chinese University of Hong Kong and in press (P. P.-H. But, 2000,
pers. comm.). Therefore I primarily focused on mosses. Only a small part of the liverwort information
used in the present inventory comes from specimen examination, and most comes from publication
records.
Much information has been given in the 1997 checklist, and thus is not repeated here. Therefore the
present inventory should be consulted in conjunction with the 1997 checklist.
The Latin binomials and authorities generally were checked by consulting Wijk, Margadant and
Florschuetz (1959-1969), Crosby, Magill and Bauer (1992), Crosby and Magill (1994, 1997) and
Redfearn et al. (1996) for mosses, and Piippo (1990) for liverworts (including hornworts), in addition
to the new publications.
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Localities in Hong Kong for each species are arranged as follows: New Territories, Kowloon, Hong
Kong Island, Lantau Island, and Outlying Islands. Within each locality, sublocalities are ordered
alphabetically. I have tried to get the locality and habitat information for specimens which were not
collected by myself by checking the pertinent literature and labels on the specimen bags.
The distribution ranges for species within China largely follow Redfearn et al. (1996) for mosses and
Piippo (1990) for liverworts, in addition to consulting some updated references available. They are
ordered alphabetically. For convenience, I treat Hong Kong and Guangdong as different geographical
regions, and do not see Hong Kong as a part of Guangdong. Therefore, the presence of a taxon in Hong
Kong does not mean it occurs in Guangdong. The global distribution of each species mostly comes
from floras and checklists of countries and regions as well as monographs.
There are 360 species and 12 infraspecific taxa in 70 families and 159 genera of bryophytes accepted to
occur in Hong Kong presently. In addition, nineteen previously reported species are excluded from the
local flora, and 20 species are treated as doubtful records. Four families, 18 genera and 46 taxa are
reported as new additions to Hong Kong, of which Barbella convolvens is new to China, and four
species, Acroporium secundum, Leucophanes glaucum, Mitthyridium fasciculatum, and Syrrhopodon
spiculosus, are new to mainland China (excluding Hainan, Taiwan and South China Sea Islands).
Ectropothecium nervosum was reduced to a synonym of E. zollingeri.
The inventory consists of the following parts: (1) List of accepted mosses, (2) Excluded species of
mosses, (3) Doubtful species of mosses, (4) List of accepted liverworts and hornworts, (5) Excluded
species of liverworts, and (6) Doubtful species of liverworts.
Genera and taxa are alphabetically arranged. The information on each species in the inventory is given
in a standard format as follows, except for excluded and doubtful species.
Scientific name (synonyms which relate to Hong Kong species) - references citing.
Note: where needed.
Localities: NEW TERRITORIES. TAI MO SHAN. Tai Shing Stream. 780 m (SE), 2180. [The block
letters in parenthesis are the direction in which the locality faces. This followed by my collection
number if the collector is omitted]. HONG KONG IS. VICTORIA PEAK, 350 m (Zhang et al., 1998b)
[This locality was based on the report of Zhang et al. (1998b)]. LANTAU IS. LANTAU PEAK. 900 m,
Herklots 123 (BM) [The block letters in parenthesis are the acronym of the herbarium in which the
specimen was deposited].
Habitat and Substrate: All specimens were collected in forest. 1 on rocks, 1 on soil, and 1 unknown.
Distribution: China (Anhui, Guangdong, Hainan, Hong Kong, Shandong, Taiwan, Yunnan). India,
Malaysia, Papua New Guinea, Australia, Hawaii, Africa, South America (Gao and Crosby, 1999; Tan
and Jia, 1997).
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Zhang Li: Hong Kong Bryophytes 5
AN UPDATED AND ANNOTATED INVENTORY OF HONG KONG BRYOPHYTES
List of accepted mosses
AMBLYSTEGIACEAE
Platyhypnidium riparioides (Hedw.) Dix. (Rhynchostegium riparioides (Hedw.) Card.)
Locality: NEW TERRITORIES. TAI MO SHAN. Ng Tung Chai Falls. 190 m (N), 1769.
Habitat and Substrate: On rocks in forest.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hebei, Hong Kong, Hubei, Hunan,
Jilin, Liaoning, Shaanxi, Shanxi, Shandong, Shanghai, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang).
Japan, Nepal, Bhutan, Central Asia, Philippines, Africa, North America (Anderson, Crum and Buck,
1990; Gangulee, 1969-1980; Jia and Wang, 1993; Koponen et al., 2000; Noguchi, 1991; O’Shea, 1999;
Redfearn et al., 1996; Tan and Iwatsuki, 1991; Zhao and Cao, 1998).
ARCHIDIACEAE
Archidium ohioense Schimp. ex C. Muell. - So and Zhu (1996c)
Localities: NEW TERRITORIES. TAI LAM. Kiu Keng Shan. 260 m (NW), 345. Lin Fa Shan. 105 m
(SW), 651. - SAI KUNG. Nam Shek Tsim. 440 m (W), 24(1). - TAI MO SHAN. 950 m, So 95124
(HKBU, IBSC). - PAK TAI TO YAN. 480 m (S), 46(1). - TSING SHAN. 470 m (SW), 9(1). HONG
KONG IS. D’AGUILAR PEAK. 10 m (W), 1256. - VIOLET HILL. 225 m, 1366. LANTAU IS. KAU
NGA LING (Western). 420 m (S), 44(1). - NEI LAK SHAN. 160 m (NE), 2069. - SUNSET PEAK.
Wong Lung Hang (East Lung Stream). 185 m (N), 1263. OUTLYING ISLANDS. TSING YI. 275 m,
981.
Habitat and Substrate: There were 12 specimens studied, of which 4 were found in forest, 3 in
grassland, 2 in shrubland, 1 by the seashore, 1 by a waterfall, and 1 unknown. 8 were found on soil, 3
on rocks, 1 unknown.
Distribution: China (Henan, Hong Kong, Shandong). Japan, India, Sri Lanka, Thailand, Australia, New
Caledonia, Africa, North America, West Indies (Anderson, Crum and Buck, 1990; Gao and Crosby,
1999; He, 2000; Noguchi, 1987b; Streimann and Curnow, 1989).
BARTRAMIACEAE
Philonotis hastata (Duby) Wijk. & Marg.
Localities: NEW TERRITORIES. GRASS HILL. Fo Tan Stream. 270 m (SE), 222. - KAI KUNG
LENG. Ha Che Stream. 195 m (S), 1865. - KOWLOON PEAK. Ho Chung Stream. 130 m (E), 1517. TAI MO SHAN. Kwun Yam Shan. 95 m (N), 931. - TAI TO YAN. Lam Tsuen Stream. 100 m (E),
701; 105 m (E), 702.
Habitat and Substrate: All 6 specimens were collected in forest. 3 were found on rocks, and 3 on soil.
Distribution: China (Fujian, Hainan, Hong Kong, Jiangsu, Shanghai, Taiwan, Xizang, Zhejiang). Japan,
India, Myanmar, Vietnam, Thailand, Cambodia, Malaysia, Philippines, Borneo, Indonesia, Australia,
Society Islands, Vanuatu, Africa, Mexico, Brazil (Eddy, 1996; Gangulee, 1969-1980; He, 2000;
Higuchi, 1996; Koponen, 1998; Mohamed and Tan, 1988; Noguchi, 1989; O’Shea, 1999; Redfearn et
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al., 1996; Scott, Stone and Rosser, 1976; Sharp, Crum and Eckel, 1994; Streimann and Curnow, 1989;
Tan and Iwatsuki, 1991, 1993; Touw, 1978; Whittier, 1976; Yano, 1995).
Philonotis thwaitesii Mitt. (Philonotis appressifolia Dix., fide Koponen (1998)).
Localities: NEW TERRITORIES. TAI LAM. Kat Hing Bridge to Tsing Fai Tong. 130 m (NW),
1059. - TAI MO SHAN. Chuen Lung Stream. 725 m (SW), 2435. Ng Tung Chai Falls. 245 m (NW),
1400; 500 m (NW), 3(20). Shek Kong Stream. 320 m (NW), 1973. Tai Shing Stream. 860 m (NE),
1139. HONG KONG IS. ZOOLOGICAL AND BOTANICAL GARDENS. 170 m (N), 1397.
LANTAU IS. SUNSET PEAK. San Shek Wan Stream. 370 m (S), 2042. OUTLYING ISLANDS.
SIU AP CHAU. 5 m, 2484.
Habitat and Substrate: Eight of 9 specimens were collected in forest, and 1 by a stream. 4 were found
on soil, 4 on rocks, and 1 on a concrete wall.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hong Kong, Hunan, Jiangsu,
Jiangxi, Jilin, Shaanxi, Shandong, Shanghai, Sichuan, Taiwan, Yunnan, Zhejiang). Korea, Japan, India,
Nepal, Sri Lanka, Thailand, Philippines, Borneo, Indonesia, Papua New Guinea, Tonga (Fang et al.,
1998; Gangulee, 1969-1980; He, 2000; Koponen et al., 2000; Liu et al., 1999; Redfearn et al., 1996;
Tan and Iwatsuki, 1991; Touw, 1978; Zhao and Cao, 1998).
Philonotis turneriana (Schwaegr.) Mitt.
Locality: NEW TERRITORIES. FAN LING. 105 m, 42.
Habitat and Substrate: On a brick wall by a road.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hong Kong, Jiangsu, Jiangxi,
Shandong, Sichuan, Taiwan, Xizang, Yunnan). Japan, Northern India, Nepal, Bhutan, Sri Lanka,
Myanmar, Thailand, Malaysia, Philippines, Malesia, Vanuatu, Hawaii (Bartram, 1933; Eddy, 1996;
Gangulee, 1969-1980; He, 2000; Higuchi, 1996; Long, 1994; Mohamed and Tan, 1988; Redfearn et
al., 1996; Tan and Iwatsuki, 1991).
BRACHYTHECIACEAE
Brachythecium garovaglioides C. Muell. (B. wichurae (Broth.) Par., fide Ignatov and Koponen
(1996))
Locality: NEW TERRITORIES. TAI MO SHAN. Tai Shing Stream. 915 m (NE), 1112 (Confirmed
by M. Ignatov, HKU, IBSC, KFBG, ).
Habitat and Substrate: On rocks in forest.
Distribution: China (Anhui, Fujian, Guangdong, Guizhou, Hong Kong, Hunan, Inner Mongolia,
Jiangsu, Shandong, Shanghai, Sichuan, Taiwan, Xinjiang, Yunnan, Zhejiang). Japan, Korea, Russian
Far East, Pakistan, India, Nepal, Bhutan, Myanmar, Indonesia (Ignatov and Koponen, 1996; Koponen
et al., 2000; Noguchi, 1991; Redfearn et al., 1996).
Brachythecium moriense Besch.
Locality: NEW TERRITORIES. TAI MO SHAN. Tai Shing Stream. 915 m (NE), 1114.
Habitat and Substrate: On rocks in grassland.
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Zhang Li: Hong Kong Bryophytes 7
Distribution: China (Anhui, Hong Kong, Shaanxi, Zhejiang). Japan (Noguchi, 1991; Redfearn et al.,
1996).
Brachythecium plumosum (Hedw.) B.S.G. - So and Zhu (1996c)
Localities: NEW TERRITORIES. TAI MO SHAN. 950 m (So and Zhu, 1996c). Tai Shing Stream.
885 m (NE), 1353 (Confirmed by M. Ignatov, HKU, IBSC, KFBG); 915 m (NE), 1111 (Confirmed by
M. Ignatov, HKU, IBSC, KFBG).
Habitat and Substrate: On trunks and on soil in forest.
Distribution: China (Anhui, Guangxi, Guizhou, Heilongjiang, Henan, Hong Kong, Hubei, Hunan, Inner
Mongolia, Jiangsu, Jiangxi, Jilin, Liaoning, Qinghai, Shaanxi, Shanghai, Shanxi, Sichuan, Taiwan,
Xinjiang, Xizang, Yunnan, Zhejiang). Japan, India, Nepal, Bhutan, Philippines, Borneo, Australia,
New Zealand, Europe, Africa, North America, Mexico, Central and South America (Anderson, Crum
and Buck, 1990; Buck, 1998; Fang et al., 1998; Fife, 1995; Gangulee, 1969-1980; Hedenas, 1996;
Noguchi, 1991; O’Shea, 1999; Redfearn et al., 1996; Tan and Iwatsuki, 1991; Tan and Jia, 1997;
Touw, 1978).
Eurhynchium asperisetum (C. Muell.) Bartr.
Locality: HONG KONG (Unknown) (So, 1995b).
Habitat and Substrate: Along ditches in vegetable fields (So, 1995b).
Distribution: China (Anhui, Guizhou, Hong Kong, Hubei, Shaanxi, Shandong, Taiwan, Xizang,
Yunnan, Zhejiang). Japan, Thailand, Philippines, Indonesia (He, 2000; Li, Zeng and Su, 1995;
Redfearn et al., 1996; Tan and Iwatsuki, 1991; Zhao and Cao, 1998).
Eurhynchium hians (Hedw.) Sande Lac.
Localities: NEW TERRITORIES. TAI MO SHAN. Ng Tung Chai Falls. 300 m (N), 5(8). LANTAU
IS. SUNSET PEAK. San Shek Wan Stream. 480 m (S), 2048.
Habitat and Substrate: On rocks in forest.
Distribution: China (Fujian, Guizhou, Heilongjiang, Henan, Hong Kong, Hunan, Inner Mongolia, Jilin,
Shaanxi, Taiwan, Zhejiang). Japan, Nepal, Bhutan, Russian Siberia, Europe, North America
(Anderson, Crum and Buck, 1990; Bai, 1996; Gangulee, 1969-1980; Koponen et al., 2000; Noguchi,
1991; Redfearn et al., 1996).
Eurhynchium savatieri Schimp. ex Besch.
Species new to Hong Kong.
Localities: NEW TERRITORIES. HUNG FA LENG. Muk Min Tau Stream. 230 m (E), 499. - TAI
MO SHAN. Ng Tung Chai Falls. 395 m (N), 4(17).
Habitat and Substrate: Both specimens were collected in forest. One was found on rocks, and the other
on soil.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Henan, Hong Kong,
Hubei, Hunan, Inner Mongolia, Jiangsu, Jiangxi, Shaanxi, Shandong, Shanghai, Shanxi, Sichuan,
Xinjiang, Xizang, Yunnan, Zhejiang). Korea, Japan, Vanuatu (Bai, 1996; Fang et al., 1998; Higuchi,
1996; Noguchi, 1991; Rao et al., 1997; Redfearn et al., 1996; Zhao and Cao, 1998; Zhang, 1996).
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Rhynchostegium pallidifolium (Mitt.) Jaeg.
Localities: NEW TERRITORIES. MA ON SHAN. Ma On Shan Tsuen Stream. 615 m (SW), 2173. SAI KUNG. Nam She Tsim. 225 m (N), 1953.
Habitat and Substrate: On rocks and on rotten wood in forest.
Distribution: China (Anhui, Guangdong, Guizhou, Heilongjiang, Henan, Hong Kong, Hunan, Inner
Mongolia, Jilin, Shaanxi, Shandong, Shanxi, Shanghai, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang).
Japan (Koponen et al., 2000; Redfearn et al., 1996; Zhao and Cao, 1998).
BRYACEAE
Brachymenium exile (Dozy & Molk.) Bosch & Sande Lac.
Genus and species new to Hong Kong.
Specimens examined: HONG KONG IS. D’AGUILAR PEAK. 10 m (W), 1255. OUTLYING
ISLANDS. CHEUNG MUK TAU. 35 m, 884. - FAT TONG CHAU. 80 m, 919. - KAI CHAU. 5 m,
439. - PING CHAU. 55 m, 2280. - TAI A CHAU. 10 m, 849; 20 m, 850. - TAI TAU CHAU. 5 m, 436.
- TANG LUNG CHAU. 25 m, 837. – TAU LAU CHAU. 10 m, 841. - TIT CHAN CHAU. 20 m, 924.
Habitat and Substrate: Most studied specimens were found on the seashore. 8 were collected on soil, 2
on rocks, and 1 on concrete steps.
Distribution: China (Hainan, Hong Kong, Hubei, Hunan, Shandong, Sichuan, Taiwan, Xizang,
Yunnan). Korea, Japan, India, Myanmar, Vietnam, Thailand, Malaysia, Philippines, Australia, Hawaii,
Africa, Mexico, Central and South America. (Gangulee, 1969-1980; He, 2000; Koponen et al., 2000;
Mohamed and Tan, 1988; Noguchi, 1988; O’Shea, 1999; Sharp, Crum and Eckel, 1994; Stoneburner et
al., 1993; Tan and Iwatsuki, 1991, 1993).
Bryum argenteum Hedw.
Locality: NEW TERRITORIES. KOWLOON PEAK. Ho Chung Catchwater. 85 m, 1514.
Habitat and Substrate: On a concrete wall.
Distribution: Cosmopolitan.
Bryum atrovirens Brid. (B. erythrocarpum Schwaegr.)
Locality: NEW TERRITORIES. TAI PO (Dixon, 1933).
Habitat and Substrate: On rocks in nullah (Dixon, 1933).
Distribution: China (Hong Kong, Jiangsu, Shandong, Taiwan, Xinjiang, Xizang, Zhejiang). Japan,
Europe, Africa, North America (Noguchi, 1988; Redfearn et al., 1996; Zhao and Cao, 1998).
Bryum billardieri Schwaegr.
Localities: NEW TERRITORIES. MA ON SHAN. Ma On Shan Tsuen Stream. 450 m (SW), 2165.
Ngong Ping. 345 m, 126. - PAT SIN LENG. Hsien Ku Fung. 420 m (NE), 1024. - SAI KUNG. Shek
Uk Shan. 460 m (NW), 312. - TAI LO SHAN. Kwun Yam Shan. 305 m (N), 959; 430 m (N), 972. TAI MO SHAN. Ng Tung Chai Falls. 270 m (NW), 1404; 500 m (N), 3(3); 600 m (N), 2(6); 630 m
(N), 1189; 685 m (N), 1241. Shek Kong Stream. 600 m (NW), 1986. Tai Shing Stream. 840 m (NE),
1422; 865 m (NE), 12. Yin Ngam Stream. 305 m (NE), 1883. LANTAU IS. LANTAU PEAK. 665 m
(S), 1692; 725 m (N), 2460. - SUNSET PEAK. San Shek Wan Stream. 405 m (S), 2046; 485 m (S),
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Zhang Li: Hong Kong Bryophytes 9
2056; 600 m (S), 26(2); 700 m (SE), 27(18). Wong Lung Hang (East Lung Stream). 140 m (N), 1262.
Wong Lung Hang (Main Streams). 260 m (N), 2089; 590 m (N), 2119; 730 m (N), Ng 34.
Habitat and Substrate: Twenty-four of 25 specimens were collected in forest, and 1 in grassland. 15
were found on rocks, 9 on soil, and 1 on humus.
Distribution: China (Anhui, Fujian, Guangdong, Guizhou, Hong Kong, Hubei, Hunan, Shaanxi,
Sichuan, Taiwan, Xinjiang, Xizang, Yunnan, Zhejiang). Japan, India, Nepal, Myanmar, Vietnam,
Thailand, Philippines, Australia, New Zealand, Pacific Islands, Africa, North, Central and South
America (Anderson, Crum and Buck, 1990; Eddy, 1996; Gangulee, 1969-1980; He, 2000; Koponen et
al., 2000; Noguchi, 1988; Redfearn et al., 1996; Sharp, Crum and Eckel,1994; Tan and Iwatsuki, 1991,
1993).
Bryum caespiticium Hedw.
Locality: HONG KONG (Locality unknown) (So, 1995b).
Habitat and Substrate: On moist rock surfaces, soil banks near streams, and urban areas (So, 1995b).
Distribution: Cosmopolitan.
Bryum capillare Hedw.
Localities: NEW TERRITORIES. TAI MO SHAN. Yin Ngam Stream. 200 m (NE), 1878.
OUTLYING ISLANDS. SIU AP CHAU. 5 m, 2489.
Habitat and Substrate: One specimen was growing on soil by a path, and the other on rocks in forest.
Distribution: Cosmopolitan.
Bryum cellulare Hook. in Schwaegr.
Localities: NEW TERRITORIES. KAM SHAN. 110 m, 754. - SHA TAU KOK. Tsiu Hang. 15 m,
2539. - TAI LAM. Lin Fa Shan. 40 m (SW), 645. - TAI PO. Herklots 260a (BM). HONG KONG IS.
HAPPY VALLEY. Ah Nin H.14 (BM). - MOUNT CAMERON. 125 m (S), 1715. - WONG NAI
CHONG. Ah Nin 308 (BM). LANTAU IS. SZE PAK WAN. 5 m, 2129. OUTLYING ISLANDS.
LAMMA IS. Yung Shue Ha. 10 m, Lin et al. 2009 (IBSC, KFBG). - PING CHAU. 15 m, 2269.
Habitat and Substrate: There were 10 specimens studied, of which 4 were collected in forest, 1 in
grassland, and 5 unknown. 5 were found on rocks, 4 on soil, and 1 unknown.
Distribution: China (Anhui, Fujian, Guangdong, Guizhou, Hong Kong, Shaanxi, Shanghai, Sichuan,
Taiwan, Xinjiang, Xizang, Yunnan, Zhejiang). Japan, Thailand, Vietnam, Australia, Southern Africa,
Reunion, Mexico, Central and Northern South America (Eddy, 1996; He, 2000; Liu et al., 1999;
Noguchi, 1988; Redfearn et al., 1996; Sharp, Crum and Eckel, 1994; Tan and Iwatsuki, 1993).
Bryum coronatum Schwaegr.
Localities: NEW TERRITORIES. TAI MO SHAN. 880 m, So 95404 (HKBU, IBSC). HONG
KONG IS. MOUNT CAMERON. 325 m (S), 1722. OUTLYING ISLANDS. SIU MO TO. 15 m,
823. - WAGLAN IS. 70 m, 897. HONG KONG (Locality unknown). Wright s.n. (FH).
Habitat and Substrate: Two of 5 specimens were collected by the seashore, and 3 unknown. 3 were
found on soil, 1 on concrete steps, and 1 unknown.
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Distribution: China (Guangdong, Hong Kong, Shaanxi, Shandong, Sichuan, Taiwan, Xizang, Yunnan).
Japan, Bhutan, Myanmar, Vietnam, Thailand, Cambodia, Malaysia, Singapore, Borneo, Australia,
Africa, Brazil, Colombia, Ecuador (Churchill, 1998, Eddy, 1996; He, 2000; Long, 1994; Mohamed and
Tan, 1988; Noguchi, 1988; Redfearn et al., 1996; Sharp, Crum and Eckel, 1994; Streimann and
Curnow, 1989; Tan and Iwatsuki, 1993; Touw, 1978; Yano, 1995).
Pohlia camptotrachela (Rens. & Card.) Broth. - So and Zhu (1996c).
Locality: NEW TERRITORIES. TAI MO SHAN. 950 m (So and Zhu, 1996c).
Habitat and Substrate: On moist rock (So and Zhu, 1996c).
Distribution: China (Hong Kong, Shaanxi, Xizang). Korea, Japan, Nepal, Russia, Europe, North
Africa, North America, Brazil (Anderson, Crum and Buck, 1990; Gangulee, 1969-1980; Ignatov and
Afonina, 1992; Noguchi, 1988; Redfearn et al., 1996; Yano, 1995).
Pohlia proligera (Kindb.) Broth. - So and Zhu (1996c).
Locality: HONG KONG (Locality unknown). Above 400 m (So and Zhu, 1996c).
Habitat and Substrate: On dripping rock in deep shade (So and Zhu, 1996c).
Distribution: China (Guizhou, Heilongjiang, Hong Kong, Hunan, Inner Mongolia, Jilin, Liaoning,
Shandong, Zhejiang). Russia, North America (Anderson, Crum and Buck, 1990; Koponen et al., 2000;
Ignatov and Afonina, 1992).
Rhodobryum giganteum (Schwaegr.) Par.
Genus and species new to Hong Kong.
Localities: LANTAU IS. SUNSET PEAK. Wong Lung Hang (East Lung Stream). 730 m (N), 1293;
760 m (N), 2310.
Habitat and Substrate: On humus in forest.
Distribution: China (Anhui, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hong Kong, Hubei,
Hunan, Jiangxi, Shaanxi, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Korea, Japan, India, Nepal,
Bhutan, Sri Lanka, Myanmar, Laos, Vietnam, Thailand, Malaysia, Philippines, Borneo, Hawaii,
Madagascar (Eddy, 1996; Fang et al., 1998; Gangulee, 1969-1980; He, 2000; Long, 1994; Mohamed
and Tan, 1988; Redfearn et al., 1996; Tan and Iwatsuki, 1991, 1993; Touw, 1978).
CALYMPERACEAE
Calymperes afzelii Schwaegr. - Li and Crosby (2001)
Localities: NEW TERRITORIES. PAT SIN LENG. Hok Tau (Gao and But, 1992). - TAI MO SHAN
(Li and Crosby (2001) as Da-Mo Mt.). - TAI PO (Gao and But, 1992). - TAI PO KAU (Li and Crosby
(2001) as Da-Po-Kau). KOWLOON (Gao, 1996). HONG KONG IS. TAI TAM RESERVOIR (Gao
and But, 1992).
Habitat and Substrate: On rocks and trunks (Gao and But, 1992).
Distribution: China (Guangdong, Hainan, Hong Kong, Taiwan, Yunnan). Laos, Vietnam, Thailand,
Cambodia, Malaysia, Singapore, Philippines, Borneo, Indonesia, Papua New Guinea, Australia, Africa,
Mexico, West Indies, Northern South America (Gao, 1996; Mohamed and Tan, 1988; Redfearn et al.,
10
Zhang Li: Hong Kong Bryophytes 11
1996; Reese and Mohamed, 1985; Reese and Stone, 1995; Sharp, Crum and Eckel, 1994; Tan and
Iwatsuki, 1993; Yano, 1995).
Calymperes erosum C. Muell. - Li and Crosby (2001)
Localities: NEW TERRITORIES. GRASS HILL. Fo Tan Stream. 430 m (SE), 209. - HUNG FA
LENG. Mu Min Tau Stream. 50 m (E), 486; 55 m (E), 487. - KAU LUNG HANG SHAN. 95 m (NW),
1545. - KAI KUNG LENG. Ha Che Stream. 40 m (S), 1858; 50 m (S), 1860; 160 m (S), 1864. - MA
ON SHAN. Tai Tung Wo Liu Stream. 70 m (NE), 1562. Tiu Shau Ngam. 185 m (NW), 732. PLOVER COVE RESERVOIR. Ngor Kai Teng. 210 m, 132, 133. - SAI KUNG. Pat Tam Chung. 50
m, 828. Kap Man Hang Pool. 25 m (E), 1212. Yung Shue O to Pat Tam Chung. 125 m, 1382. SHATIN. Campus of the Chinese University (Gao and But, 1992). - TAI LAM. Lin Fa Shan. 40 m
(SW), 648. Kat Hing Bridge to Tsing Fai Tong Stream. 90 m (NW), 1044. - TAI LO SHAN. Kwun
Yam Shan. 100 m (N), 937; 370 m (N), 968. - TAI MO SHAN. Chuen Lung Stream. 355 m (SW),
2418. Ng Tung Chai Falls. 200 m (N), 16(13); 500 m (N), 3(12). Tai Shing Stream. 200 m (SE), 17(7);
270 m (SE), 1872; 300 m (SE), 18(7). Tai Yuen Stream. 215 m (S), 1836; 220 m (S), 1837; 230 m (S),
1838. Water-pumping House Stream. 540 m (E), 1969. Yin Ngam Stream. 190 m (NE), 1877. - TAI
PO KAU (Li and Crosby (2001) as Da-Po-Kau). - TAI TO YAN. Lam Tsuen Stream. 75 m (E), 688;
100 m (E), 694, 697; 110 m (E), 704. San Lung Wai Stream. 265 m (W), 766. - WU KAU TANG.
Mirror Pool Feng Shui Wood. 60 m, 1749. So Lo Pun Back Stream. 50 m (NE), 1615; 65 m (NE),
1621. HONG KONG IS. ABERDEEN RESERVOIR (Gao and But, 1992). - CHATHAM PATH. 290
m (N), 1744. - D’AGUILAR PEAK. 85 m (W), 1247, 1248; 110 m (W), 142; 230 m (W), 162. HATTON RD. 185 m, 1487. - MOUNT CAMERON. 125 m (S), 1714. - MOUNT NICHOLSON. 260
m (N), 2393, 2394. - TAI TAM RESERVOIR. 120 m, 1378. - VICTORIA PEAK (Li and Crosby,
2001). LANTAU IS. LANTAU PEAK. 110 m (S), 1656; 125 m (S), 2211, 2212; 250 m (S), 2213; 330
m (SW), 2256. - NEI LAK SHAN. 25 m (NE), 2066. - SUNSET PEAK. San Shek Wan Stream. 90 m
(S), 2014; 100 m (S), 30(3); 120 m (S), 2016; 145 m (S), 2023; 155 m (S), 2024; 200 m (S), 31(8); 300
m (S), 32(5); 400 m (S), 33(1), 33(15). Wong Lung Hang (East Lung Stream). 100 m (N), 39(2). Wong
Lung Hang (Main Streams). 260 m (N), 2090; 615 m (N), 92. OUTLYING ISLANDS. FU WONG
CHAU. 10 m, 2520, 2521. - KAI CHAU. 10 m, 446, 447. - LAMMA IS. So Kwu Wan. 50 m, Lin et al.
1980 (IBSC, KFBG); 70 m, Lin et al. 2000 (IBSC, KFBG). - MA SHI CHAU. 10 m, 384. - SAM
MUN TSAI CHAU (not found on the map). 10 m, 450. - SUNG KONG. 45 m, 910. - WU CHAU. 5 m,
372. - YEUNG CHAU (near Kat O). 20 m, 2528. HONG KONG (Locality unknown). Bowring s.n.
(BM).
Habitat and Substrate: Seventy-two of 77 specimens were collected in forest, 1 by a waterfall, 1 by a
stream, and 3 unknown. 49 were found on rocks, 19 on trunks, 2 on soil, 2 on tree base, 1 on a rotten
trunk, 1 on bark, 1 on a root, 1 on a liana, and 1 unknown.
Distribution: China (Guangdong, Guangxi, Hainan, Hong Kong, Yunnan). India, Sri Lanka, Myanmar,
Thailand, Malaysia, Singapore, Philippines, Borneo, Indonesia, Australia, Africa, Mexico, West Indies,
Northern South America (Gao, 1996; Mohamed and Tan, 1988; Reese and Mohamed, 1985; Reese and
Stone, 1995; Tan and Iwatsuki, 1993; Sharp, Crum and Eckel, 1994; Yano, 1995).
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12 Memoirs of the Hong Kong Natural History Society 26, 2003
Calymperes fasciculatum Dozy & Molk. – Li and Crosby (2001)
Localities: NEW TERRITORIES. MA ON SHAN. Tiu Shau Ngam. 150 m (NW), 253. - PAT SIN
LENG. Wang Shan Keuk Sheung Tsuen. 315 m (NW), 1164. - TAI MO SHAN. Tai Shing Stream. 855
m (NE), 1184. HONG KONG IS. TAI TAM RESERVOIR. Iwatsuki 45 (IBSC, NICH). LANTAU IS.
LANTAU PEAK. 300 m (N), 1303. - SUNSET PEAK. Wong Lung Hang (East Lung Stream). 500 m
(N), 36(16); 505 m (N), 1277; 600 m (N), 37(3).
Habitat and Substrate: Seven of 8 specimens were collected in forest, and 1 unknown. 5 were found on
rocks, 2 on trunks, and 1 unknown.
Distribution: China (Guangdong, Hainan, Hong Kong, Taiwan, Yunnan). Japan, Sri Lanka, Myanmar,
Vietnam, Thailand, Cambodia, Malaysia, Philippines, Borneo, Indonesia, New Caledonia, Fiji,
Solomon Islands, Hawaii (Gao, 1996; Matsui, 1995; Menzel and Schultze-Motel, 1990; Reese and
Mohamed, 1985; Tan and Iwatsuki, 1993).
Calymperes graeffeanum C. Muell.
Species new to Hong Kong.
Localities: OUTLYING ISLANDS. KAU LO CHAU. 5 m, 844.
Habitat and Substrate: On rocks by seashore.
Distribution: China (Hainan, Hong Kong, Taiwan, Yunnan). Thailand, Philippines, Australia, Africa
(Gao, 1996; O’Shea, 1999; Reese and Stone, 1995; Tan and Iwatsuki, 1991, 1993).
Calymperes lonchophyllum Schwaegr.
Locality: HONG KONG IS. TAI TAM RESERVOIR. Iwatsuki 39 (IBSC, NICH).
Habitat and Substrate: Unknown.
Distribution: China (Hainan, Hong Kong, Taiwan). Japan, Myanmar, Thailand, Malaysia, Philippines,
Australia, Africa, Mexico, West Indies, Brazil (Gao, 1996; He, 2000; Matsui, 1995; Menzel and
Schultze-Motel, 1990; Reese and Stone, 1995; Sharp, Crum and Eckel, 1994; Tan and Iwatsuki, 1993;
Yano, 1995).
Calymperes palisotii Schwaegr. ssp. molluccense (Schwaegr.) Menzel & Schultze-Motel (C.
molluccense Schwaegr.)
Localities: HONG KONG IS. TAI TAM RESERVOIR. Iwatsuki 33 (IBSC, NICH). HONG KONG
(Localities unknown). Bowring s.n. (IBSC); Wright s.n. (FH).
Habitat and Substrate: One of 3 specimens were collected on rocks, and 2 unknown.
Distribution: China (Hainan, Hong Kong, Yunnan). Sri Lanka, Myanmar, Thailand, Philippines,
Borneo, Anadamans, Malesia, Papua New Guinea, Australia, Africa (Gao, 1996; He, 2000; Li and
Crosby, 2001; O’Shea, 1999; Reese and Stone, 1995; Tan and Iwatsuki, 1993; Touw, 1978).
Calymperes tenerum C. Muell.
Localities: NEW TERRITORIES. FAN LING (Gao and But, 1992). - SAI KUNG. Kap Man Hang
Pool. 25 m (E), 1215. Sheung Yiu. 15 m, 1413. HONG KONG IS. ABERDEEN RESERVOIR (Gao
and But, 1992). LANTAU IS. LIN FA SHAN. 500 m (NE), 681. OUTLYING ISLANDS. CHA
KWO CHAU. 20 m, 886. - KWUN CHAM CHAU (Not found in the map). 5 m, 472. - LAMMA IS.
12
Zhang Li: Hong Kong Bryophytes 13
Shek Pai Wan (Gao and But, 1992). - NGO CHAU TSAI (Not found in the map). 40 m, 925. - PING
CHAU. 5 m, 2264. - SHEK CHAU. 10 m, 873. - TAI SHUI HANG IS. (Not found in the map). 5 m,
453.
Habitat and Substrate: Most specimens were collected by the seashore. 6 were found on rocks, 1 on
leaves, 1 on a rotten twig, and 1 on a trunk.
Distribution: China (Guangdong, Hainan, Hong Kong, Taiwan, Yunnan). India, Laos, Vietnam,
Thailand, Cambodia, Malaysia, Singapore, Philippines, Borneo, Indonesia, Australia, New Caledonia,
Society Islands, Hawaii, Africa, Mexico, Southern Brazil, Haiti, Puerto Rico (Gao, 1996; He, 2000;
Mohamed and Tan, 1988; O’Shea, 1999; Reese and Stone, 1995; Sharp, Crum and Eckel, 1994; Tan
and Iwatsuki, 1993; Touw, 1978; Whittier, 1976).
Mitthyridium fasciculatum (Hook. & Grev.) Robins.
Species new to mainland China, and genus and species new to Hong Kong.
Locality: NEW TERRITORIES. MA ON SHAN. Ma On Shan Tsuen Stream. 605 m (SW), 2170.
Habitat and Substrate: On trunks in forest.
Distribution: China (Hainan, Hong Kong). India, Nepal, Sri Lanka, Myanmar, Vietnam, Thailand,
Cambodia, Philippines, Malaysia, Singapore, Borneo, Indonesia, Australia, Samoa, Vanuatu, Africa,
Mauritius, Seychelles, South America (Gao, 1996; He, 2000; Higuchi, 1996; Mohamed and Tan, 1988;
Reese and Stone, 1995; Tan and Iwatsuki, 1993; Touw, 1978).
Mitthyridium flavum (C. Muell.) Robins.
Species new to Hong Kong.
Locality: NEW TERRITORIES. WU KAU TANG. Wu Kau Tang Feng Shui Wood. 95 m, 1755.
Habitat and Substrate: On trunks in forest.
Distribution: China (Hainan, Hong Kong, ?Taiwan (Lai and Yang-Wang (1976) and Kuo and Chiang
(1987) listed it, but no further information to confirm its occurrence), Yunnan). Vietnam, Thailand,
Cambodia, Philippines, Malaysia, Singapore, Borneo, Papua New Guinea, Australia, Africa (He, 2000;
Mohamed and Tan, 1988; O’Shea, 1999; Reese and Stone, 1995; Tan and Iwatsuki, 1993; Touw,
1978).
Syrrhopodon armatus Mitt. - Li and Crosby (2001)
Localities: NEW TERRITORIES. KAI KUNG LENG. Ha Che. 35 m, 1856. Ksiu Keng Pang Uk
Stream. 365 m (NE), 1871; 385 m (NE), 1870. - LAM TSUEN. Tai Om Feng Shui Wood. 110 m,
1902, 1903, 1904. - TAI LAM. Kat Hing Bridge to Tsing Fai Tong. 210 m (NW), 1069. - TAI MO
SHAN. Chuen Lung Stream. 400 m (SW), 2422. Ng Tung Chai Tsuen. 110 m, 1781. Shing Mun
Reservoir. 300 m, 1481. Tai Shing Stream. 300 m (SE), 18(10); 345 m (SE), 1855; 385 m (SE), 1854.
Tai Yuen Stream. 215 m (S), 1835; 235 m (S), 1839. - TAI PO KAU. 100 m, 2080. HONG KONG IS.
HATTON RD. 190 m, 1486. - VICTORIA PEAK (Li and Crosby, 2001). LANTAU IS. NGONG
PING. 430 m, 2216.
Habitat and Substrate: Sixteen of 18 specimens were collected in forest, and 2 by roads. 16 were found
on trunks, and 2 on bark.
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14 Memoirs of the Hong Kong Natural History Society 26, 2003
Distribution: China (Guangdong, Guangxi, Hainan, Hong Kong, Sichuan, Taiwan, Yunnan). Japan,
India, Vietnam, Thailand, Malaysia, Singapore, Borneo, Philippines, Australia, New Zealand, New
Caledonia, Hawaii, Africa (Frahm et al., 1990; Gangulee, 1969-1980; Gao, 1996; He, 2000; Mohamed
and Tan, 1988; Reese and Stone, 1995; Tan and Iwatsuki, 1991).
Syrrhopodon chenii Reese & P.-J. Lin
Species new to Hong Kong.
Localities: NEW TERRITORIES. MA ON SHAN. Ma On Shan Tsuen Stream. 380 m (SW), 2159. SAI KUNG. Nam She Tsim. 350 m (N), 1951. - SHEUNG WO HANG FENG SHUI WOOD. 150 m
(NW), 1757, 1758; 180 m (NW), 1764. - WU KAU TANG. Fan Kei Tok Feng Shui Wood. 280 m (N),
1612; 290 m (N), 1609. Fan Kei Tok Summit. 350 m, 1598. HONG KONG IS. D’AGUILAR PEAK.
215 m (E), 527.
Habitat and Substrate: Eight of 9 specimens were collected in forest, and 1 in grassland. 5 were found
on trunks, 2 on a tree base, 1 on rotten wood, and 1 on humus.
Distribution: Endemic to South China (Guangdong, Guangxi, Hong Kong) (Gao, 1996).
Syrrhopodon flammeonnervis C. Muell.
Species new to Hong Kong.
Localities: LANTAU IS. LANTAU PEAK. 575 m (N), 2448; 600 m (N), 1322.
Habitat and Substrate: On trunks and on rocks in forest.
Distribution: China (Hainan, Hong Kong, Guangxi). Vietnam, Thailand, Cambodia, Philippines,
Indonesia, Vanuatu (Gao, 1996; Higuchi, 1996; Tan and Iwatsuki, 1991, 1993).
Syrrhopodon gardneri (Hook.) Schwaegr. - Li and Crosby (2001)
Gao and But (1992) excluded this species from Hong Kong, but Gao (1996) still included Hong Kong
in the locality list. Zhang and Lin (1997) included it in the 1997 checklist too. The specimen on which
this record was based (Iwatsuki 39, IBSC, NICH) was listed twice as authentic material under both S.
gardneri and Calymperes lonchophyllum. I have checked the voucher specimen and confirmed that it is
C. lonchophyllum, therefore the occurrence of S. gardneri in Hong Kong which was reported
previously should be rejected. Recently, Li and Crosby (2001) listed a new specimen for Hong Kong
(Lin et al. 104, IBSC, not seen).
Locality: NEW TERRITORIES: TAI MO SHAN. Kadoorie Farm (Li and Crosby (2001) as Mt.
Kadoorie).
Habitat and Substrate: Unknown.
Distribution: China (Guangdong, Hainan, Hong Kong, Jiangxi, Taiwan, Yunnan). Pantropical.
Syrrhopodon hongkongensis L. Zhang - Li and Crosby (2001); Zhang, Corlett and Chau (1999).
Localities: NEW TERRITORIES. SHEUNG WO HANG FENG SHUI WOOD. 130 m (NW),
1074;145 m (NW), 1078; 150 m (NW), 1759; 160 m (NW), 1079 (Holotype IBSC, Isotypes HKU,
KFBG, LAF); 170 m (NW), 1761.
Habitat and Substrate: All 5 specimens were collected in forest. 3 were found on trunks, 1 on a trunk
and a tree base, and 1 on soil.
14
Zhang Li: Hong Kong Bryophytes 15
Distribution: Endemic to Hong Kong.
Syrrhopodon japonicus (Besch.) Broth. - Li and Crosby (2001)
Localities: NEW TERRITORIES. MA ON SHAN. Ma On Shan Tsuen Stream. 355 m (SW), 2154;
360 m (SW), 2156; 545 m (SW), 2163. Tai Tung Wo Liu Stream. 170 m (NE), 1573. Tiu Shau Ngam.
310 m (NW), 273; 365 m (NW), 114. - PAT SIN LENG. Hsien Ku Fung. 335 m (E), 1006; 370 m
(NE), 1013; 375 m (NE), 1014. Pin Fung Shan. 475 m (SW), 397. Sze Tau Leng. 240 m (NW), 991. SAI KUNG. Nam Shek Tsim. 445 m (N), 1936. - TAI LAM. Lin Fa Shan. 230 m (SW), 564. - TAI
MO SHAN. Kadoorie Farm (Li and Crosby (2001) as Mt. Kadoori). Shek Kong Stream. 755 m (NW),
2295. Tai Shing Stream. 600 m (SE), 21(14); 785 m (NE), 1434. - TAI TO YAN. Summit. 470 m,
2540. LANTAU IS. LANTAU PEAK. 535 m (N), 2449; 645 m (N), 1328; 695 m (S), 1698; 720 m
(N), 2456; 730 m (S), 1702; 880 m, 1344; 915 m, 2542. - LIN FA SHAN. 645 m (E), 1457. - SUNSET
PEAK. San Shek Wan Stream. 675 m (S), 2058. Wong Lung Hang (East Lung Stream). 700 m (N),
38(5); 760 m (N), 2308; 770 m (N), 2248; 775 m (N), 2251.
Habitat and Substrate: Twenty-seven of 30 specimens were collected in forest, and 3 on a summit. 24
were collected on rocks, 4 on trunks, and 2 on rotten twigs.
Distribution: China (Fujian, Guangdong, Guangxi, Hainan, Hong Kong, Hunan, Jiangxi, Sichuan,
Taiwan, Yunnan, Zhejiang). Japan, Vietnam, Thailand, Malaysia, Philippines, Borneo, Indonesia,
Papua New Guinea, New Caledonia, Fiji, Vanuatu (Gao, 1996; Higuchi, 1996; Tan and Iwatsuki, 1991,
1993; Touw, 1978).
Syrrhopodon prolifer Schwaegr. var. tosaensis (Card.) Orban & Reese - Li and Crosby (2001)
Localities: NEW TERRITORIES. MA ON SHAN. Ma On Shan Tsuen Stream. 450 m (SW), 2166.
Tai Tung Wo Liu Stream. 170 m (NE), 1572, 1578. Tiu Shau Ngam. 580 m (NW), 1589. - PAT SIN
LENG. Pin Fung Shan. 480 m (SW), 393. Sze Tau Leng. 330 m (NW), 996, 998. Wang Shan Keuk
Sheung Tsuen. 310 m (NW), 1163; 445 m (NW), 1173. - SAI KUNG. Nam She Tsim. 380 m (N),
1955; 445 m (N), 1935. - TAI MO SHAN. Shek Kong Stream. 775 m (NW), 2007. Tai Shing Stream.
820 m (NE), 19; 835 m (NE), 1359; 855 m (NE), 1185; 860 m (NE), 1138, 1355. Water-pumping
House Stream. 700 m (E), S29. Yin Ngam Stream. 480 m (NE), 1896; 580 m (NE), 1901. - TAI TO
YAN. San Lung Wai Stream. 460 m (W), 755. - WU KAU TANG. Fan Kei Tok Feng Shui Wood. 320
m (N), 1608; 330 (N), 1606. Fan Kei Tok Summit. 345 m, 1597, 1599; 360 m, 1592. Wu Kau Tang
Feng Shui Wood. 85 m, 1753; 105 m, 1756. LANTAU IS. LANTAU PEAK. 360 m (N), 1308, 1309;
560 m (SW), 2259; 610 m (N), 1323; 730 m (S), 1705. - LIN FA SHAN. 585 m (NE), 639; 610 m
(NE), 640. - NGONG PING (Gao and But, 1992). - SHUI LO CHO. 210 m (N), 565; 250 m (N), 570,
572, 577a. - SUNSET PEAK. Wong Lung Hang (East Lung Stream). 585 m (N), 1280; 600 m (N),
37(19); 680 m (N), 1286; 700 m (N), 38(3); 800 m (E), 42(24). Wong Lung Hang (Main Streams). 175
m (N), 2082; 310 m (N), 2102; 560 m (N), 66.
Habitat and Substrate: Forty-six of 47 specimens were collected in forest, and 1 in shrubland. 23 were
found on rocks, 14 on trunks, 3 on tree bases, 2 on rotten wood, 2 on soil, 2 on roots, and 1 on a liana.
Distribution: China (Guangdong, Guangxi, Hainan, Hong Kong). Japan, Vietnam, Thailand (Gao,
1996; He, 2000; Orban and Reese, 1990; Tan and Iwatsuki, 1993).
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16 Memoirs of the Hong Kong Natural History Society 26, 2003
Syrrhopodon spiculosus Hook. & Grev.
Species new to Hong Kong and mainland China.
Localities: OUTLYING ISLANDS. SIU NIM CHAU. 10 m, 2507, 2509. - TSING CHAU. 15 m,
2502.
Habitat and Substrate: On rocks by the seashore.
Distribution: China (Hong Kong, Hainan). Vietnam, Thailand, Cambodia, Philippines, Malaysia,
Singapore, Borneo, Papua New Guinea, Australia, Oceania, Africa (Ellis and Tan, 1999; Gao, 1996;
He, 2000; Mohamed and Tan, 1988; Reese and Stone, 1995; Tan and Iwatsuki, 1993; Touw, 1978).
Syrrhopodon trachyphyllus Mont. - Li and Crosby (2001)
Localities: NEW TERRITORIES. MA ON SHAN. Tiu Shau Ngam. 200 m (NW), 262. - SHEUNG
WO HANG FENG SHUI WOOD. 190 m (NW), 1765. - TAI MO SHAN. Kadoorie Farm (Li and
Crosby (2001) as Mt. Kadoorie). – TAI PO KAU (Li and Crosby (2001) as Da Po Kau).
Habitat and Substrate: On rocks in forest.
Distribution: China (Guangdong, Hainan, Hong Kong, Taiwan). Japan, Sri Lanka, Vietnam, Thailand,
Cambodia, Malaysia, Singapore, Philippines, Borneo, Indonesia, Australia, New Caledonia (Gao,
1996; Mohamed and Tan, 1988; Tan and Iwatsuki, 1993; Touw, 1978).
CRYPHAEACEAE
Pilotrichopsis dentata (Mont.) Besch.
Localities: NEW TERRITORIES. TAI MO SHAN. Chuen Lung Stream. 730 m (SW), 2439.
LANTAU IS. LANTAU PEAK. 565 m (S), 1678. – SUNSET PEAK. Wong Lung Hang (East Lung
Stream). 600 m (N), 1283. Wong Lung Hang (Main Streams). 605 m (N), 89.
Habitat and Substrate: All specimens were collected in forest. 3 were found on rocks, and 1 on a trunk.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hong Kong, Hubei, Hunan,
Jiangxi, Taiwan, Zhejiang). Japan, Vietnam, Philippines (Fang et al., 1998; Li, Liao and Huang, 1998;
Noguchi, 1989; Rao et al., 1997; Redfearn et al., 1996; Tan and Iwatsuki, 1991, 1993).
DICRANACEAE
Campylopus atrovirens De Not.
Localities: NEW TERRITORIES. MA ON SHAN. Tai Tung Wo Liu Stream. 155 m (NE), 1566.
HONG KONG IS. D’AGUILAR PEAK. 305 m (E), 174. LANTAU IS. LIN FA SHAN. 595 m (E),
1449. – SHEUNG TUNG AU. 650 m, 47. – SUNSET PEAK. Wong Lung Hang (Main Streams). 290
m (N), 2097.
Habitat and Substrate: All specimens were collected in forest. 4 were found on rocks, and 1 on a trunk.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hebei, Hong Kong, Hunan,
Jiangsu, Jiangxi, Liaoning, Shaanxi, Xizang, Yunnan, Zhejiang). Japan, Nepal, Europe, North America
(Gao and Crosby, 1999; Redfearn et al., 1996).
Campylopus ericoides (Griff.) Jaeg.
Locality: LANTAU IS. SUNSET PEAK. Wong Lung Hang (East Lung Stream). 720 m (N), 1292.
Habitat and Substrate: On rocks in forest.
16
Zhang Li: Hong Kong Bryophytes 17
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hubei,
Jiangxi, Sichuan, Yunnan). India, Sri Lanka, Nepal, Bhutan, Myanmar, Laos, Vietnam, Thailand,
Cambodia, Malaysia, Philippines, Indonesia (Gangulee, 1969-1980; Gao and Crosby, 1999; Long,
1994; Mohamed and Tan, 1988; Redfearn et al., 1996; Tan and Iwatsuki, 1993).
Campylopus japonicus Broth.
Localities: NEW TERRITORIES. GRASS HILL. Fo Tao Stream. 580 m (SE), 191. – HUNG FA
LENG. Summit. 445 m, 1144. – MA ON SHAN. Ngau Ngak Shan Summit. 680 m (S), 13(5). Tiu Shau
Ngam. 195 m (NW), 736; 205 m (NW), 737. – TAI MO SHAN. Tai Shing Stream. 700 m (SE), 6(16).
HONG KONG IS. HIGH WEST. 430 m (N), 1727; 490 m (N), 1728. – VICTORIA PEAK. 540 m
(E), 15(1). LANTAU IS. LANTAU PEAK. 800 m (N), 2470; 915 m, 2483. – LIN FA SHAN. 705 m
(E), 10(1). – SUNSET PEAK. Wong Lung Hang (East Lung Stream). 245 m (N), 1265.
Habitat and Substrate: There were 13 specimens examined, of which 8 were collected in forest, 4 in
grassland, and 1 unknown. 7 specimens were found on rocks, 4 on soil, 1 on a twig, and 1 on a trunk.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hebei, Hong Kong,
Hubei, Jiangxi, Liaoning, Shaanxi, Sichuan, Taiwan, Yunnan, Zhejiang). Japan, Vietnam, Thailand,
Australia, Society Islands, Mexico, Western North America (Frahm, 1993; Gao and Crosby, 1999; He,
2000; Jia and Wang, 1993; Redfearn et al., 1996).
Campylopus laxitextus Sande Lac.
Localities: NEW TERRITORIES. MA ON SHAN. Ngau Ngak Shan Summit. 680 m (S), 13(4). –
PAT SIN LENG. Hsien Ku Fung. 420 m (NE), 1027. – TAI MO SHAN. Ng Tung Chai Falls. 500 m
(N), 3(16). Summit. 925 m (E), 11(1). Tai Shing Stream. 300 m (SE), 18(16). – PAK TAI TO YAN.
480 m (S), 46(2). LANTAU IS. SUNSET PEAK. San She Wan Stream. 200 m (S), 31(9); 400 m (S),
33(5); 700 m (SE), 27(7). Wong Lung Hang (East Lung Stream). 200 m (N), 40(4); 320 m (N), 41(5).
HONG KONG (Locality unknown). So 726A (IBSC, HKBU).
Habitat and Substrate: Eight of 12 specimens were collected in forest, 2 in grassland, 1 in shrubland,
and 1 unknown. 6 were found on rocks, 5 on soil, and 1 on bark.
Distribution: China (Hainan, Hong Kong, Jiangxi, Taiwan, Yunnan). Malaysia, Philippines, Indonesia,
Papua New Guinea, Australia, Pacific Islands (Gao and Crosby, 1999; Mohamed and Tan, 1988;
Redfearn et al., 1996).
Campylopus serratus Sande Lac. (C. scabridorsus Dix.) – Gao and Crosby (1999) as C. comosus
(Schwaegr.) Bosch & Sande Lac.
Locality: HONG KONG IS. REPULSE BAY. 600 ft. (Dixon, 1933).
Habitat and Substrate: On rock (Dixon, 1933).
Distribution: China (Hong Kong). Vietnam, Thailand, Cambodia, Malaysia, Singapore, Borneo (Frahm
and Mohamed, 1987; Mohamed and Tan, 1988; Tan and Iwatsuki, 1993).
Campylopus taiwanensis Sak. – Gao and Crosby (1999).
Localities: NEW TERRITORIES. MA ON SHAN. Ngau Ngak Shan Summit. 665 m, 1586. – TAI
TO YAN. Summit. 495 m, 31. – TAI MO SHAN. Shek Kong Stream. 915 m (NW), 1740. Summit.
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18 Memoirs of the Hong Kong Natural History Society 26, 2003
920 m, 1174. Tai Shing Stream. 600 m (SE), 21(8); 900 m (NE), 1124; 920 m (NE), 1143. HONG
KONG IS. VICTORIA PEAK, 530 m, 1740. LANTAU IS. LIN FA SHAN. 695 m (NE), 644. – NEI
LAK SHAN. 160 m (NE), 2070; 170 m (NE), 2071. – SHEUNG TUNG AU. 765 m, 45. – SUNSET
PEAK. San Shek Wan Stream. 120 m (S), 2019; 600 m (S), 26(1); 630 m (S), 2057; 700 m (SE),
27(21). OUTLYING ISLANDS. SAM PUI CHAU. 5 m, 362. – TSING YI. 300 m, 976.
Habitat and Substrate: There were 18 specimens studied, of which 11 were found in forest, 5 in
grassland, 1 in shrubland, and 1 by a road. 11 were found on rocks, 3 on soil, 2 on trunks, 1 on a
concrete wall, and 1 on rotten wood.
Distribution: Endemic to China (Anhui, Guangxi, Hong Kong, Hunan, Shandong, Taiwan, Zhejiang)
(Gao and Crosby, 1999).
Campylopus umbellatus (Arn.) Par.
Localities: NEW TERRITORIES. TAI MO SHAN. Shek Kong Stream. 910 m (NW), 1469. Tai
Shing Stream. 400 m (SE), 19(16); 790 m (NE), 21; 900 m (NE), 1119. Tai Yuen Stream. 365 m (S),
1849. – HO PUI RESERVOIR. 220 m, 810. LANTAU IS. SUNSET PEAK. San She Wan Stream. 400
m (S), 33(11). Wong Lung Hang (Main Streams). 615 m (N), 91.
Habitat and Substrate: Six of 8 specimens were collected in forest, 1 in grassland, and 1 unknown. 7
specimens were found on rocks, and 1 on a twig.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hubei,
Hunan, Jiangxi, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Korea, Japan, Bhutan, Sri Lanka,
Myanmar, Vietnam, Thailand, Cambodia, Malaysia, Philippines, Borneo, Indonesia, Australia, Society
Islands, Vanuatu, Hawaii (Bartram, 1933; Gao and Crosby, 1999; He, 2000; Higuchi, 1996; Long,
1994; Mohamed and Tan, 1988; Tan and Iwatsuki, 1991, 1993; Touw, 1978; Whittier, 1976).
Dicranella coarctata (C. Muell.) Bosch. & Sande Lac. Var. coarctata (D. coarctata var. obscura (Sull.
& Lesq.) Iwats.) – Gao and Crosby (1999)
Localities: NEW TERRITORIES. HO CHUNG. 20 m, 2084. – MA ON SHAN. Tiu Shau Ngam. 360
m (NW), 266. – PAT SIN LENG. Shek Au Shan. 210 m, 1553. – SAI KUNG. Nam She Tsim. 410 m
(N), 1956. Shek Uk Shan Summit. 480 m, 304. – SHATIN PASS. Herklots 573 (BM). – TAI MO
SHAN. Shing Mun Reservoir. 415 m, 1480; 450 m, 1479. – TUEN MUN. Tai Shui Hang. 80 m (NW),
552. – WU KAU TANG. Half way from Wu Kau Tang to Fan Kei Tok. 215m (SW), 1590. HONG
KONG IS. BACK OF UNIVERSITY. Herklots 269, 274, 280 (BM). – HIGH WEST. 385 m (N),
1738; 430 m (N), 1726; 540 m (E), 15(2). – MOUNT PARKER. Youngsaye 300A. – VICTORIA
PEAK. 430 m, 1508. LANTAU IS. LANTAU PEAK. 715 m (N), 1333. – NEI LAK SHAN. 335 m
(NE), 2077. OUTLYING ISLANDS. KAU SAI CHAU. Tai Leng Stream. 30 m (NW), 294. – PO TAI
IS. 30 m, Lin et al. 1922 (IBSC, KFBG); 130 m, Lin et al. 1913 (IBSC, KFBG). – SIU MO TO. 5 m,
831; 10 m, 829. – WONG YI CHAU. So 941013 (HKBU, IBSC). HONG KONG (Locality unknown).
Clements 4231; Wright 55, 56, 57, 59, 80, s.n., s.n., s.n. (FH). Herklots H.61. (BM).
Habitat and Substrate: There were 36 specimens studied, of which 5 were found in forest, 5 in
grassland, 3 by a road, 1 in shrubland, 1 by a pool, 1 by a stream, 1 by a waterfall, and 19 unknown. 25
specimens were found on soil, 9 on rocks, and 4 unknown.
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Zhang Li: Hong Kong Bryophytes 19
Distribution: China (Fujian, Guangdong, Guizhou, Hainan, Hong Kong, Hubei, Hunan, Jiangxi,
Shanghai, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Japan, Sri Lanka, Myanmar, Vietnam,
Thailand, Malaysia, Philippines, Indonesia, Australia (Gao and Crosby, 1999; He, 2000; Mohamed and
Tan, 1988; Redfearn et al., 1996; Tan and Iwatsuki, 1993).
Dicranella coarctata (C. Muell.) Bosch. & Sande Lac. Var. torrentium Card. – Gao and Crosby
(1999).
Locality: NEW TERRITORIES. TAI MO SHAN (Gao and Crosby, 1999).
Habitat and Substrate: Unknown (Gao and Crosby, 1999).
Distribution: Endemic to South China (Hong Kong, Taiwan) (Gao and Crosby, 1999).
Dicranella varia (Hedw.) Schimp.
Localities: NEW TERRITORIES. HO CHUNG. 20 m, 1511, 1512, 1513, 1784, 1785. – TAI PO
KAU. 300 m, So 95125A (HKBU, IBSC). OUTLYING ISLANDS. MA WAN. 10 m, 839.
Habitat and Substrate: Five of 7 specimens were collected by road, 1 in forest, and 1 unknown. 6 were
found on soil, and 1 unknown.
Distribution: China (Anhui, Guangdong, Guangxi, Guizhou, Hong Kong, Jiangxi, Liaoning, Shandong,
Shanghai, Sichuan, Xizang, Yunnan, Zhejiang). Japan, Russia, Europe, Africa, North America, Cuba,
Haiti (Anderson, Crum and Buck, 1990; Gao and Crosby, 1999; Ignatov and Afonina, 1992; Noguchi,
1987b; Redfearn et al., 1996; Sharp, Crum and Eckel, 1994).
Dicranodontium asperulum (Mitt.) Broth.
Genus and species new to Hong Kong.
Localities: NEW TERRITORIES. PAT SIN LENG. Wang Shan Keuk Sheung Tsuen. 420 m (NW),
1167, 1168.
Habitat and Substrate: On rocks by waterfalls.
Distribution: China (Anhui, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hunan, Jiangsu,
Sichuan, Taiwan, Xizang, Yunnan). Japan, Russia, Nepal, Thailand, Indonesia, Europe, North America
(Frahm, 1997; Gao and Crosby, 1999; He, 2000; Ignatov and Afonina, 1992; Li, Liao and Huang,
1998; Redfearn et al.,1996).
Dicranodontium didictyon (Mitt.) Jaeg.
Species new to Hong Kong.
Localities: NEW TERRITORIES. MA ON SHAN. Tiu Shau Ngam. 415 m (NW), 115. – TAI MO
SHAN. Tai Shing Stream. 855 m (NE), 1414. LANTAU IS. LANTAU PEAK. 715 m (N), 1336. –
SUNSET PEAK. Wong Lung Hang (East Lung Stream). 800 m (N), 42(23).
Habitat and Substrate: Three of 4 specimens were collected in forest, and 1 by a waterfall. 3 were
found on rocks, and 1 on a trunk.
Distribution: China (Guangxi, Guizhou, Hainan, Hong Kong, Sichuan, Xizang, Yunnan). Japan, India,
Vietnam, Myanmar, Thailand (Frahm, 1997; Gao and Crosby, 1999; He, 2000; Tan and Iwatsuki,
1993).
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20 Memoirs of the Hong Kong Natural History Society 26, 2003
Dicranoloma assimile (Hamp.) Par. (Dicranum sericifolium Dix., fide Gao 1994) – Gao and Crosby
(1999)
Localities: LANTAU IS. LANTAU PEAK. 550 m (SW), 2261; 720 m (N), 2458; SUNSET PEAK.
Wong Lung Hang (East Lung Stream). 775 m (N), 2252; 2 000 – 3 000 ft., Herklots 349, 358 (BM).
Habitat and Substrate: There were 5 specimens examined, of which 3 were collected in forest, and 2
unknown. 3 specimens were found on rocks, and 2 unknown.
Distribution: China (Fujian, Guangxi, Guizhou, Hainan, Hong Kong, Taiwan, Zhejiang). Vietnam,
Thailand, Malaysia, Philippines, Borneo, Indonesia, Papua New Guinea, Solomon Islands (Gao and
Crosby, 1999; He, 2000; Redfearn et al., 1996; Tan and Iwatsuki, 1993; Touw, 1978).
Dicranoloma cylindrothecium (Mitt.) Sak. (D. fragiliforme (Card.) Broth.)
Locality: LANTAU IS. LANTAU PEAK. 3 000 ft. (Dixon, 1933).
Habitat and Substrate: On igneous rock (Dixon, 1933).
Distribution: China (Anhui, Fujian, Guangxi, Guizhou, Hong Kong, Taiwan, Zhejiang). Korea, Japan,
Russian Far East (Gao and Crosby, 1999; Redfearn et al., 1996).
Holomitrium densifolium (Wils.) Wijk. & Marg.
Localities: NEW TERRITORIES. TAI MO SHAN. Chuen Lung Stream. 595 m (SW), 2427. Shek
Kong Stream. 910 m (NW), 1470. Tai Shing Stream. 900 m (NE), 1126; 835 m (NE), 1425. LANTAU
IS. LANTAU PEAK. 860 m (N), 2476. – SUNSET PEAK. Wong Lung Hang (East Lung Stream). 775
m (N), 2250. Wong Lung Hang (Main Streams). 580 m (N), 2117.
Habitat and Substrate: Six of 7 specimens were collected in forest, and 1 unknown. 5 were on rocks, 1
on a trunk, and 1 on a twig.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hong Kong, Sichuan, Taiwan,
Yunnan, Zhejiang). Japan, India, Bhutan, Sri Lanka, Myanmar, Laos, Vietnam, Thailand, Philippines
(Gao and Crosby, 1999; Long, 1994; Redfearn et al., 1996; Tan and Iwatsuki, 1993).
Holomitrium vaginatum (Hook.) Brid.
Species new to Hong Kong. I accept the species concept of Eddy (1988)’s and Whittier (1976)’s and
treat H. cylindraceum (P. Beauv.) Wijk and Marg. And H. vaginatum as distinct species. Gao (1994)
supplied two illustrations for H. cylindraceum (Pl. 99: 1-7, Pl. 102: 1-8), and clearly they are different
species. Pl. 99 is H. vaginatum (under the name of H. cylindraceum), and the latter probably is H.
cylindraceum or H. densifolium. On the other side, Gao and Crosby (1999) only kept Pl. 102 (as Pl. 49)
for H. cylindraceum and gave no explanation.
Localities: NEW TERRITORIES. PAT SIN LENG. Pin Fung Shan. 480 m (SW), 396. – TAI TO
YAN. Summit. 475 m, 28. HONG KONG (Locality unknown). Bowring s.n. (BM).
Habitat and Substrate: There were 3 specimens examined, of which 1 was collected in forest, 1 on the
summit, and 1 unknown. 2 were found on rocks, and 1 unknown.
Distribution: China (Guizhou, Hong Kong, Yunnan). Philippines, Borneo, Indonesia, Society Islands,
Polynesia, Africa, Madagascar, Reunion, Mauritius (Eddy, 1988; Redfearn et al., 1996; Touw, 1978;
Whittier, 1976).
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Zhang Li: Hong Kong Bryophytes 21
Leucoloma molle (C. Muell.) Mitt.
Localities: NEW TERRITORIES. GRASS HILL. Fo Tan Stream. 405 (SE), 212. – KOWLOON
PEAK. Pak Fa Lam. 310 m (E), 1529. – PAT SIN LENG. Wang Chung Stream. 300 m (E), 4. – SAI
KUNG. Nam Shek Tsim. 375 m (N), 1950. Shek Uk Shan. 320 m (NW), 321. – TAI LAM. Hung Shui
Shan. 415 m (N), 417. Lin Fa Shan. 235 m (SW), 664; 240 m (SW), 666. – TAI LO SHAN. Kwun
Yam Shan. 250 m (N), 958. – TAI MO SHAN. Ng Tung Chai Falls. 395 m (N), 4(1a); 600 m (N),
2(27); 700 m (N), 1(7). Tai Shing Stream. 840 m (NE), 1417. Water-pumping House Stream. 700 m
(E), S15, S16. – WU KAU TANG. Fan Kei Tok Feng Shui Wood. 350 m (N), 1604. LANTAU IS.
LANTAU PEAK. 315 m (S), 1663; 325 m (N), 1306; 645 m (S), 1690. – LIN FA SHAN. 260 m (NE),
606; 490 m (NE), 630; 570 m (E), 1451. – SUNSET PEAK. Wong Lung Hang (East Lung Stream).
400 m (N), 1273. Wong Lung Hang (Main Streams). 275 m (N), 2094; 640 m (N), 48.
Habitat and Substrate: All 25 specimens were collected in forest. 22 on rocks, 2 on trunks, and 1 on a
tree base.
Distribution: China (Guangdong, Guangxi, Hainan, Hong Kong, Taiwan). Japan, Vietnam, Thailand,
Cambodia, Malaysia, Philippines, Borneo, Indonesia, Society Islands, Vanuatu, Hawaii (Bartram,
1933; Gao and Crosby, 1999; He, 2000; Higuchi, 1996; Tan and Iwatsuki, 1991, 1993; Whittier, 1976).
Leucoloma walkeri Broth.
Species new to mainland China and Hong Kong.
Localities: NEW TERRITORIES. SAI KUNG. Shek Uk Shan. 335 m (NW), 318. – TAI MO SHAN.
Tai Shing Stream. 300 m (SE), 18(14); 600 m (SE), 21(19). Yin Ngam Stream. 270 m (NE), 1881; 285
m (NE), 1882. HONG KONG IS. D’AGUILAR PEAK. 160 m (W), 154. LANTAU IS. LANTAU
PEAK. 120 m (S), 2210; 200 m (S), 1659; 270 m (S), 1661; 375 m (S), 1664; 575 m (S), 1679. –
SUNSET PEAK. San She Wan Stream. 300 m (S), 32(1); 500 m (S), 34(18); 600 m (S), 26(14). Wong
Lung Hang (Main Streams). 275 m (N), 2092, 2093.
Habitat and Substrate: All specimens were collected in forest. 13 were found on rocks, and 3 on trunks.
Distribution: China (Hainan, Hong Kong). India, Myanmar, Thailand, Malaysia, Philippines (Allen and
Pursell, 1984; Mohamed and Tan 1988).
Microdus brasiliensis (Duby) Ther.
Locality: NEW TERRITORIES. TAI MO SHAN. Ng Tung Chai Falls. 400 m (N), So 95621 (HKBU,
IBSC).
Habitat and Substrate: Unknown.
Distribution: China (Hainan, Henan, Hong Kong, Shandong, Shanghai, Sichuan, Xizang, Yunnan).
India, Sri Lanka, Myanmar, Vietnam, Thailand, Cambodia, Malaysia, Philippines, Borneo, Indonesia,
South America (Gangulee, 1969-1980; Gao and Crosby, 1999; He, 2000; Mohamed and Tan, 1988;
Redfearn et al., 1996; Tan and Iwatsuki; 1991; Touw, 1978; Zhao and Cao, 1998).
Trematodon longicollis Michx.
Localities: NEW TERRITORIES. HO CHUNG. 20 m, 1783. – LUNG KWU TAN POWER
STATION. 80 m, Xing 17. – TAI PO KAU. 120 m, Choi 33. LANTAU IS. NEI LAK SHAN Summit.
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22 Memoirs of the Hong Kong Natural History Society 26, 2003
550 m, 2079. OUTLYING ISLANDS. CHEUNG SOK. 10 m, 836. – TAI MO TO. 5 m, 833, 834. –
SIU MO TO. 5 m, 830; 15 m, 832.
Habitat and Substrate: Five of 9 specimens were collected in grassland, 3 in forest, and 1 by a road. 8
were found on soil, and 1 on a used battery wall.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Henan, Hong Kong,
Hubei, Hunan, Jiangsu, Jiangxi, Liaoning, Shandong, Shanghai, Sichuan, Taiwan, Xizang, Yunnan,
Zhejiang). Japan, India, Sri Lanka, Myanmar, Thailand, Cambodia, Malaysia, Philippines, Papua New
Guinea, Fiji, Society Islands, Australia, Hawaii, Europe, Africa, North, Central and South America
(Anderson, Crum and Buck, 1990; Eddy, 1988; Gangulee, 1969-1980; Gao and Crosby, 1999; He,
2000; Mohamed and Tan, 1988; O’Shea, 1999; Redfearn et al., 1996; Sharp, Crum and Eckel, 1994;
Streimann and Curnow, 1989; Tan and Iwatsuki, 1991, 1993; Whittier, 1976).
DIPHYSCIACEAE
Diphyscium involutum Mitt.
Species new to Hong Kong.
NEW TERRITORIES. CASTLE PEAK. 405 m (NE), 1393. – KAI KUNG LENG. Kiu Keng Pang
Uk Stream. 425 m (NE), 1868. – KOWLOON PEAK. Ho Chung Stream. 380 m, 1528. – MA ON
SHAN. Tiu Shau Ngam. 250 m (NW), 739; 340 m (NW), 98, 104; 390 m (NW), 272. – PAT SIN
LENG. Hsien Ku Fung. 385 m (NE), 1016. Pin Fung Shan. 325 m (SW), 408. Wang Shan Keuk
Sheung Tsuen. 400 m (NW), 1165. Nam Chung Stream. 315 m (NE), 999. – TAI LAM. Hung Shui
Shan. 385 m (N), 423. Lin Fa Shan. 325 m (SW), 673; 380 m (SW), 677. – TAI MO SHAN. Ng Tung
Chai Falls. 500 m (N), 3(15). Shek Kong Stream. 485 m (NW), 1979. Tai Shing Stream. 300 m (SE),
18(5); 855 m (NE), 1186. Water-pumping House Stream. 700 m (E), S36. – TUEN MUN. Tai Shui
Hang. 65 m (NW), 541. – WU KAU TANG. Wu Kau Tang Feng Shui Wood. 85 m, 1754. HONG
KONG IS. D’AGUILAR PEAK. 160 m (W), 155, 158. – VIOLET HILL. 275 m, 1369. LANTAU IS.
LANTAU PEAK. 485 m (N), 1316; 720 m (S), 1700. – LIN FA SHAN. 700 m (E), 1445; 460 m (NE),
629. – SUNSET PEAK. 500 m, Ip, Gao and Woo 623 (CUHK). San She Wan stream. 700 m (SE),
27(12). Wong Lung Hang (East Lung Stream). 335 m (N), 2243 (HKU, IBSC, KFBG, MO, det. By Z.
Magombo); 600 m (N), 1281; 770 m (N), 1300. Wong Lung Hang (Main Streams). 195 m (N), 2086.
Habitat and Substrate: Thirty-three of 34 specimens were collected in forest, and 1 unknown. 33 were
found on rocks, and 1 on soil.
Distribution: China (Guizhou, Hainan, Hong Kong, Taiwan, Yunnan). Japan, India, Sri Lanka,
Thailand, Cambodia, Malaysia, Philippines, Borneo (Gangulee, 1969-1980; He, 2000; Mohamed and
Tan, 1988; Noguchi, 1987b; Redfearn et al., 1996; Tan and Iwatsuki, 1991, 1993; Touw, 1978).
DITRICHACEAE
Ditrichum pallidum (Hedw.) Hampe
So and Yip (1995a) superfluously reported this species new to Hong Kong. Previously, several authors
had reported this species under different names from Hong Kong (cf. Zhang and Lin, 1997).
Locality: HONG KONG (Locality unknown).
Habitat and Substrate: On soil (So and Yip, 1995a).
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Zhang Li: Hong Kong Bryophytes 23
Distribution: China (Anhui, Fujian, Guangdong, Guizhou, Henan, Hong Kong, Hubei, Hunan, Inner
Mongolia, Jiangsu, Jiangxi, Shandong, Xizang, Yunnan, Zhejiang). Japan. Russia, Thailand, Europe,
Central Africa, North America (Anderson, Crum and Buck, 1990; Bai, 1996; Gao and Crosby, 1999;
Ignatov and Afonina, 1992; Tan and Iwatsuki, 1993).
Eccremidium brisbanicum (Broth.) Stone & Scott – Zhang, Corlett and Chau (1998)
This genus in China is only recorded in Hong Kong.
Locality: NEW TERRITORIES. SHEUNG SHUI. Kwu Tong Reservoir. 20 m, 974 (with
Micromitrium tenerum).
Habitat and Substrate: On mud of a dried out reservoir bank
Distribution: China (Hong Kong). Japan, Vietnam, Australia (Zhang, Corlett and Chau, 1998).
Garckea flexuosa (Griff.) Marg. & Nork.
Locality: NEW TERRITORIES. HO CHUNG. 20 m, 1782.
Habitat and Substrate: On soil by a road.
Distribution: China (Guangdong, Hainan, Hong Kong, Taiwan, Xizang, Yunnan, Zhejiang). Japan,
India, Nepal, Bhutan, Sri Lanka, Myanmar, Vietnam, Thailand, Cambodia, Malaysia, Singapore,
Philippines, Indonesia, Papua New Guinea, Australia, Madagascar, Central and South America
(Gangulee, 1969-1980; He, 2000; Mohamed and Tan, 1988; Noguchi, 1987b; Redfearn et al., 1996;
Tan and Iwatsuki, 1991, 1993).
ENTODONTACEAE
Entodon cladorrizans (Hedw.) C. Muell.
Species new to Hong Kong.
Locality: NEW TERRITORIES. TAI MO SHAN. Tai Shing Stream. 915 m (NE), 1116 (Det. By Y.F. Wang, HKU, IBSC, KFBG, HSNU).
Habitat and Substrate: On soil in bamboo forest.
Distribution: China (Anhui, Guizhou, Heilongjiang, Henan, Hong Kong, Inner Mongolia, Jiangsu,
Jiangxi, Jilin, Liaoning, Shandong, Shaanxi, Shanxi, Sichuan, Xizang, Yunnan, Zhejiang). North and
East Asia, Russia, Europe, North America (Anderson, Crum and Buck, 1990; Bai, 1996; Fang et al.,
1998; Hu, 1983; Ignatov and Afonina, 1992; Redfearn et al., 1996; Zhao and Cao, 1998).
Entodon concinnus (De Not.) Par.
Species new to Hong Kong.
Locality: NEW TERRITORIES. TAI MO SHAN. Tai Shing Stream. 525 m (SE), 1917 (Det. By Y.F. Wang, HKU, IBSC, KFBG, HSNU).
Habitat and Substrate: On rocks in forest.
Distribution: China (Anhui, Fujian, Gansu, Guizhou, Hebei, Heilongjiang, Hong Kong, Hubei, Inner
Mongolia, Jilin, Liaoning, Shaanxi, Shanxi, Shandong, Sichuan, Xizang, Yunnan). Korea, Japan,
Russia, Europe, Africa, North America (Hu, 1983; Ignatov and Afonina, 1992; O’Shea, 1999; Redfearn
et al., 1996; Zhao and Cao, 1998).
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24 Memoirs of the Hong Kong Natural History Society 26, 2003
Entodon macropodus (Hedw.) C. Muell.
Localities: NEW TERRITORIES. GRASS HILL. Fo Tan Stream. 355 m (SE), 216. – KOWLOON
PEAK. Ho Chung Stream. 195 m (E), 1520. – SAI KUNG. Hoi Ha. 5 m, 1042. – TAI LO SHAN.
Kwun Yam Shan. 100 m (N), 936. – TAI MO SHAN. Ng Tung Chai Falls. 185 m (NW), 1771. Shek
Kong Stream. 755 m (NW), 2293 (Confirmed by Y.-F. Wang, HKU, IBSC, KFBG, HSNU). Tai Yuen
Stream. 325 (S), 1844. – TAI PO KAU. 330 m (NE), 2340.
Habitat and Substrate: All 8 specimens were collected in forest. 7 were found on rocks, and 1 on rotten
wood.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Heilongjiang, Hong Kong,
Hubei, Hunan, Inner Mongolia, Jilin, Jiangsu, Jiangxi, Shaanxi, Shanxi, Shandong, Sichuan, Taiwan,
Yunnan, Zhejiang). Japan, Nepal, Myanmar, Laos, Vietnam, Thailand, Africa, North, Central and
South America (Anderson, Crum and Buck, 1990; Bai, 1996; Buck, 1998; Fang et al., 1998; He, 2000;
Hu, 1983; O’Shea, 1999; Rao et al.,1997; Redfearn et al., 1996; Tan and Iwatsuki, 1993; Zhao and
Cao, 1998; Zhang, 1996).
Entodon obtusatus Broth.
Localities: NEW TERRITORIES. TAI MO SHAN. Kadoorie Farm. 600 m, So 941122 (IBSC,
HKBU). LANTAU IS. SUNSET PEAK. San She Wan Stream. 300 m (S), 32(2).
Habitat and Substrate: One specimen was collected on rocks in forest, the other unknown.
Distribution: China (Fujian, Guangdong, Guizhou, Henan, Hong Kong, Hubei, Hunan, Jilin, Liaoning,
Shaanxi, Shandong, Shanxi, Sichuan, Taiwan, Xinjiang, Yunnan, Zhejiang). Japan, India (Hu, 1983;
Rao et al., 1997; Redfearn et al., 1996; Zhao and Cao, 1998).
Entodon viridulus Card.
Localities: NEW TERRITORIES. TAI MO SHAN. 880 m, So 95413 (HKBU, IBSC). Ng Tung Chai
Falls. 420 m (NW), 1410 (Confirmed by Y.-F. Wang, HKU, IBSC, KFBG, HSNU).. LANTAU IS.
SUNSET PEAK. San Shek Wan Stream. 220 m (S), 2028.
Habitat and Substrate: Two of 3 were found on rocks in forest, the other unknown.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Heilongjiang, Hong Kong, Jiangsu, Jiangxi,
Liaoning, Shandong, Shanghai, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Korea, Japan (Fang et
al., 1998; Hu, 1983; Redfearn et al., 1996; Zhao and Cao, 1998).
EPHEMERACEAE
Micromitrium tenerum (B. S. G.) Crosby
Genus and species as well as the family Ephemeraceae new to Hong Kong. Li (2000) accepted that
only one genus Ephemerum of Ephemeraceae occurred in China. Previously, Gangulee (1969-1980)
and Redfearn et al. (1996) listed Xizang (Tibet) as one of its localities of Micromitrium tenerum.
Localities: NEW TERRITORIES. SHEUNG SHUI. Kwu Tung Reservoir. 20 m, 822 (with
Eccremidium brisbanicum), 975 (with E. brisbanicum).
Habitat and Substrate: On soft mud of a dried-out reservoir.
Distribution: China (Hong Kong, Xizang). Korea, Japan, India, New Zealand, Europe, North, Central
and South America, West Indies (Bryan, 1999; Redfearn et al., 1996).
24
Zhang Li: Hong Kong Bryophytes 25
FABRONIACEAE
Schwetschkeopsis fabronia (Schwaegr.) Broth.
Family Fabroniaceae and genus and species new to Hong Kong.
Locality: LANTAU IS. LANTAU PEAK. 595 m (S), 1687.
Habitat and Substrate: On trunks in forest.
Distribution: China (Anhui, Guangdong, Guangxi, Guizhou, Heilongjiang, Henan, Hong Kong,
Jiangxi, Jilin, Liaoling, Shaanxi, Shandong, Shanghai, Shanxi, Taiwan, Yunnan, Zhejiang). Korea,
Japan, Nepal, Russian Far East, Eastern North America, Cuba (Anderson, Crum and Buck, 1990; Buck,
1998; Ignatov and Afonina, 1992; Li, Liao and Huang, 1998; Noguchi, 1991; Redfearn et al., 1996).
FISSIDENTACEAE
Fissidens bryoides Hedw. Var. schmidii (C. Muell.) Chopra & Kumar – Zhang et al. (1998b)
Localities: NEW TERRITORIES. TAI MO SHAN. Kadoorie Farm. 395 m (NW), Lin et al. 248
(IBSC, KFBG); 420 m (NW), Lin et al. 1197 (IBSC, KFBG).
Habitat and Substrate: On rocks in forest.
Distribution: China (Guizhou, Heilongjiang, Hong Kong, Taiwan, Xizang, Yunnan). Japan, India,
Pakistan, Sri Lanka, Philippines, Indonesia, Papua New Guinea, Fiji (Gao, 1996; Iwatsuki and Suzuki,
1996; Iwatsuki, Suzuki and Li, 1999; Redfearn et al., 1996; Tan and Iwatsuki, 1991).
Fissidens ceylonensis Dozy & Molk. – Zhang et al. (1998b)
Localities: NEW TERRITORIES. MA ON SHAN. Siu Lek Yuen (Zhang et al., 1998). – PAT SIN
LENG. Tsat Muk Kiu (Zhang et al., 1998). – PLOVER COVE RESERVOIR. Shek Nga Tal. 175 m,
137. – TAI LAM. Kat Hing Bridge to Tsing Fai Tong. 95 m (NW), 1046; 130 m (NW), 1058. Lin Fa
Shan. 45 m (SW), 649. – TAI MO SHAN. Ng Tung Chai Falls. 300 m (N), 5(5). Shing Mun (Zhang et
al., 1998). Tai Shing Stream. 800 m (NE), 7(17) (with Thuidium pristocalyx). – TAI PO KAU (Zhang
et al., 1998). – TUEN MUN. Tai Shui Hang. 170 m (NW), 548. KOWLOON. TSIM SHA TSUI. 5 m
(W), 25(1). HONG KONG IS. CHUNG HOM KOK (Zhang et al., 1998). – D’AGUILAR PEAK. 85
m (W), 1253. – VICTORIA PEAK (Zhang et al., 1998). LANTAU IS. LIN FA SHAN. 65 m (NE),
594. – SHAM WAT (Zhang et al., 1998). – SUNSET PEAK. San She Wan Stream. 100 m (S), 30(2);
200 m (S), 31(4). – TAI O TO SHAM WAT (Zhang et al., 1998). – TAI O (Zhang et al., 1998).
OUTLYING ISLANDS. A CHAU. 5 m, 359. – CHEUNG MUK KAU. 30 m, 880, 881, 882. – FAT
TONG CHAU. 70 m, 918; 80 m, 917. – KAI CHAU. 5 m, 438, 441. – KIU TSUI CHAU. 5 m, 475. –
KWUN CHAM WAN. 5 m, 469. – LAMMA IS. 50 m, Lin et al. 1988 (IBSC, KFBG); 70 m, Lin et al.
1967 (IBSC, KFBG). – MA CHAU. 10 m, 858. – PO TAI IS. 10 m, Lin et al. 1924 (IBSC, KFBG). –
SAM PUI CHAU. 5 m, 363, 369. – TAI SHUI HANG IS (not found in the map). 5 m, 454. – TAP
MUN CHAU. 20 m, Lin et al. 1842 (IBSC, KFBG). – TIT LO CHAU (not found in the map). 20 m,
921. – TSING CHAU. 10 m, 2500. – YEUNG CHAU (near Kat O). 10 m, 2527. – YEUNG CHAU
(near Ma Shi Chau). 5 m, 377.
Habitat and Substrate: Eighteen of 34 specimens were collected in forest, 4 in shrubland, 3 in
grassland, 1 by the seashore, 1 in an urban area, and 7 unknown. 22 were found on rocks, 7 on soil, 1
on a concrete wall, and 4 unknown.
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26 Memoirs of the Hong Kong Natural History Society 26, 2003
Distribution: China (Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Taiwan, Yunnan). India,
Nepal, Sri Lanka, Laos, Vietnam, Thailand, Cambodia, Malaysia, Singapore, Philippines, Borneo,
Indonesia, Australia, New Zealand, Fiji, Africa (Gao, 1996; Iwatsuki and Suzuki, 1996; O’Shea, 1999;
Redfearn et al., 1996; Stoneburner et al., 1993; Tan and Iwatsuki, 1991, 1993; Touw, 1978).
Fissidens crenulatus Mitt. – Zhang et al. (1998b)
Localities: NEW TERRITORIES. HUNG FA LENG. Ma Tseuk Leng Stream. 10 m (S), 2138. San
Kwai Tin Stream. 395 m (N), 1160. – LAU FAU SHAN. 20 m, Lin et al. 1625 (IBSC, KFBG).
LANTAU IS. SUNSET PEAK. San She Wan Stream. 100 m (S), 30(6). OUTLYING ISLANDS.
CHEUNG MUK KAU. 30 m, 883. – LAMMA IS. 50 m, Lin et al. 1979 (IBSC, KFBG). – NG FAN
CHAU. 70 m, 913.
Habitat and Substrate: There were 7 specimens examined, of which 4 were collected in forest, 2 on
grassland, and 1 unknown. 5 were found on soil, and 2 on rocks.
Distribution: China (Guangdong, Hainan, Hong Kong, Taiwan, Yunnan). India, Nepal, Sri Lanka,
Myanmar, Vietnam, Thailand, Malaysia, Borneo, Papua New Guinea (Eddy, 1988; Gao, 1996; He,
2000; Mohamed and Tan, 1988; Redfearn et al., 1996, Tan and Iwatsuki, 1993, Touw, 1978).
Fissidens crispulus Brid. Var. cripulus (F. zippelianus Dozy & Molk.) – Li and Crosby (2001); Zhang
et al. (1998b)
Localities: NEW TERRITORIES. CASTLE PEAK TEMPLE (Zhang et al., 1998). – FAN LING. Kei
Lak Tsai. 205 m, 37. – GRASS HILL. Fo Tan Stream. 330 m (SE), 217; 420 m (SE), 210; 500 m (SE),
197. – HO PUI RESERVOIR. 185 m, 802, 803. – HUNG FA LENG. Muk Min Tau Stream. 20 m (E),
476. San Kwai Tin Stream. 340 m (N), 1155. – LAM TSUEN. Tai Om Feng Shui Wood. 110 m, 1908.
– MA ON SHAN. Tiu Shau Ngam. 60 m (NW), 243. – SAI KUNG. Kap Man Hang. 60 m (NE), 1223.
Sai Wan. 5 m, 1206. Shek Uk Shan. 270 m (NW), 327. Tai Long. 65 m (S), 1385. Tam Chai Shan. 180
m (E), 1041. – SHA TAU KOK. Muk Min Tau Feng Shui Wood. 10 m (S), 29(1). – SHATIN. Man Fo
Chi (Zhang et al. 1998). – TAI LO SHAN. Kwun Yam Shan. 120 m (N), 940. – TAI LAM. Kat Hing
Bridge to Tsing Fai Tong. 155 m (NW), 1066. Kiu Keng Shan. 160 m (NW), 334; 185 m (NW), 337;
210 m (NW), 340. – TAI MO SHAN. Kadoorie Farm (Zhang et al. 1998). Ng Tung Chai Falls. 200 m
(N), 16(19); 300 m (N), 5(3) (with Pseudoleskeopsis zippelii); 395 m (N), 4(10), 4(16); 500 m (N),
3(5); 600 m (N), 2(16), 2(18), 2(19), 2(21), 2(24); 630 m (N), 1190; 700 m (N), 1(12), 1(22). Tai Shing
Stream. 200 m (SE), 17(15); 700 m (SE), 6(20); 900 m (NE), 8(4). Shing Mun. Wun Yiu. Yin Ngam
(Zhang et al. 1998). – TAI PO. Sha Lo Tung (Zhang et al. 1998). – TAI PO KAU (Zhang et al. 1998).
– TAI TO YAN. Lam Tsuen Stream. 245 m (E), 1204. – TUEN MUN. Tai Shui Hang. 90 m (NW),
551; 165 m (NW), 546. WU KAU TANG. Bride’s Pool (Zhang et al. 1998). KOWLOON. TSIM SHA
TSUI. 5 m (W), 25(2). HONG KONG IS. CHUNG HOM KOK (Zhang et al. 1998). – D’AGUILAR
PEAK. 110 m (W), 143; 115 m (W), 146; 135 m (W), 150; 140 m (W), 152. – HONG KONG PARK. –
MOUNT PARKER. – VICINITY OF TAI TAM RESERVOIR. –VICTORIA PEAK (Zhang et al.
1998). LANTAU IS. LIN FA SHAN. 170 m (NE), 597. – LANTAU PEAK. Ngong Ping to Lantau
Peak. Ngong Ping to Tei Tong Tsai (Zhang et al. 1998). – SUNSET PEAK. San She Wan Stream. 200
m (S), 31(5); 300 m (S), 32(7); 400 m (S), 33(3); 600 m (S), 26(5); 700 m (SE), 27(13). Wong Lung
Hang (East Lung Stream). 500 m (N), 36(3); 600 m (N), 37(13). – SZE PAK WAN. 10 m, 2128.
26
Zhang Li: Hong Kong Bryophytes 27
OUTLYING ISLANDS. A CHAU. 5 m, 356, 357; 10 m, 360. – CHING CHAU. 5 m, 462. – KAI
CHAU. 5 m, 442. – LAMMA IS. 30 m, Lin et al. 2006 (IBSC, KFBG); 50 m, Lin et al. 1965, Lin et al.
1991, Lin et al. 1998, Lin et al. 1999 (IBSC, KFBG); 150 m, Lin et al. 1995 (IBSC, KFBG). – NG
FAN CHAU. 80 m, 916. – PAK KWO CHAU. 90 m, 890; 95 m, 892. – PO TAI IS. 180 m, Lin et al.
1916 (IBSC, KFBG); 200 m, Lin et al. 1920 (IBSC, KFBG). – PING CHAU. 5 m, 2271. – SAM MUN
TSAI CHAU (not found in the map). 5 m, 452. – SIU A CHAU. 5 m, 878. – SIU AP CHAU. 5 m,
2488; 10 m, 2491. – SUNG KONG. 55 m, 907, 908. – TAI A CHAU. 5 m, 860; 10 m, 854, 856, 857,
863, 867. – TAP MUN CHAU. 20 m, Lin et al. 1840 (IBSC, KFBG); 50 m, Lin et al. 1832, Lin et al.
1844, Lin et al. 1846, Lin et al. 1848 (IBSC, KFBG). – TAU LAU CHAU. 15 m, 842. – TSING
CHAU. 5 m, 2503. – WONG YI CHAU TSAI. 5 m, 459.
Habitat and Substrate: There were 91 specimens examined, of which 64 were collected in forest, 3 by
the seashore, 2 in grassland, 1 by a path, 1 in an urban area, and 20 unknown. 57 were found on rocks,
19 on soil, 1 on a tree base, 1 on a concrete wall, and 13 unknown.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hubei,
Hunan, Shandong, Sichuan, Taiwan, Yunnan, Zhejiang). Myanmar, Vietnam, Thailand, Cambodia,
Malaysia, Singapore, Philippines, Borneo, Australia, Fiji, Vanuatu, Africa (Gao, 1996; He, 2000;
Higuchi, 1996; Iwatsuki and Suzuki, 1996; Mohamed and Tan, 1988; Pursell, Bruggeman-Nannenga
and Iwatsuki, 1992; Rao et al., 1997; Redfearn et al., 1996; Streimann and Curnow, 1989; Tan and
Iwatsuki, 1991, 1993; Touw, 1978; Zhao and Cao, 1998).
Fissidens crispulus Brid. Var. robinsonii (Broth.) Z. Iwats. & Z.-H. Li
Species new to Hong Kong.
Specimen examined: OUTLYING ISLANDS. CHEUNG SOK. 10 m, 835.
Habitat and Substrate: On soil in forest.
Distribution: China (Fujian, Hainan, Hong Kong, Yunan). India, Thailand, Malaysia, Philippines, Fiji,
Vanuatu (Gao, 1996; He, 2000; Higuchi, 1996; Iwatsuki and Suzuki, 1996; Li and Crosby, 2001;
Mohamed and Tan, 1988).
Fissidens curvatus Hornsch. (F. strictulus C. Muell.) – Zhang et al. (1998b)
Locality: NEW TERRITORIES. SHATIN. Campus of the Chinese University of Hong Kong. So
94803 (HKBU, IBSC).
Habitat and Substrate: On rocks.
Distribution: China (Hong Kong, Shaanxi, Shanghai, Taiwan, Xizang, Yunnan). Japan, India,
Philippines, Australia, New Zealand, New Caledonia, Africa, Mexico, Chile, Argentina, Brazil (Gao,
1996; Iwatsuki, Suzuki and Li, 1999; O’Shea, 1999; Redfearn et al., 1996; Sharp, Crum and Eckel,
1994; Tan and Iwatsuki, 1991).
Fissidens dubius P. Beauv. (F. cristatus Wils. Ex Mitt.) – Zhang et al. (1998b)
Localities: NEW TERRITORIES. GRASS HILL. Fo Tan Stream. 415 m (SE), 211. – KOWLOON
PEAK. Pak Fa Lam. 225 m (E), 1530. – LUK KENG (Zhang et al. 1998). – PAT SIN LENG. Wang
Chung Stream. 380 m (E), 7, 8. – SAI KUNG. Shek Uk Shan. 335 m (NW), 320; 345 m (NW), 316. –
TAI LO SHAN. Kwun Yam Shan. 310 m (N), 961. – TAI MO SHAN. Ng Tung Chai Falls. 200 m (N),
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28 Memoirs of the Hong Kong Natural History Society 26, 2003
16(22); 505 m (N), 1347. 600 m (N), 2(9). Tai Shing Stream. 500 m (SE), 20(9). Water-pumping
House Vicinity. 710 m (E), 727. Yin Ngam (Zhang et al. 1998). LANTAU IS. LIN FA SHAN, 235 m
(NE), 604; 375 m (NE), 619. – SUNSET PEAK. Wong Lung Hang (Mean Streams), 250 m (N), 2087;
405 m (NW), 2404.
Habitat and Substrate: All 16 specimens were collected in forest. 15 were found on rocks, and 1 on
soil.
Distribution: China (Anhui, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Heilongjiang, Henan,
Hong Kong, Hubei, Hunan, Inner Mongolia, Jiangsu, Jiangxi, Jilin, Liaoning, Shaanxi, Shandong,
Shanghai, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Korea, Japan, Russia, India, Nepal, Sri Lanka,
Philippines, Indonesia, Papua New Guinea, New Zealand, Europe, Africa, North America (Anderson,
Crum and Buck, 1990; Bai, 1996; Fang et al., 1998; Fife, 1995; Gao, 1996; Ignatov and Afonina, 1992;
Iwatsuki, Suzuki and Li, 1999; Rao et al., 1997; Redfearn et al., 1996; Tan and Iwatsuki, 1991).
Fissidens flaccidus Mitt. (F. maceratus Mitt., F. splachnobryoides Broth. In Schum. & Lauterb.) – Li
and Crosby (2001); Zhang et al. (1998b) as F. maceratus
Locality: HONG KONG (Locality unknown). 200 m, So 93803 (HKBU, IBSC).
Habitat and Substrate: On damp soil.
Distribution: China (Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Taiwan, Yunnan) Japan,
India, Nepal, Sri Lanka, Myanmar, Vietnam, Malaysia, Philippines, Borneo, Indonesia, Papua New
Guinea, Australia, Caroline Islands, Fiji, Vanuatu, Africa, Americas (Gao, 1996; Higuchi, 1996;
Iwatsuki and Suzuki, 1996; Li and Crosby, 2001; Mohamed and Tan, 1988; Redfearn et al., 1996;
Stoneburner et al., 1993; Tan and Iwatsuki, 1991, 1993; Touw, 1978).
Fissidens gardneri Mitt. (F. brevinervis Broth.) – Zhang et al. (1998b)
Locality: NEW TERRITORIES. TAI MO SHAN. Shing Mun. 350 m, Lin et al. 1228 (IBSC, KFBG).
Habitat and Substrate: On rocks in forest.
Distribution: China (Fujian, Guangdong, Guangxi, Hainan, Hong Kong, Shandong, Sichuan, Taiwan,
Yunnan). Japan, India, Laos, Myanmar, Thailand, Africa, Mexico, Central and South America (He,
2000; Noguchi, 1987b; O’Shea, 1999; Pursell, Bruggeman-Nannenga and Iwatsuki, 1993; Redfearn et
al., 1996; Sharp, Crum and Eckel, 1994; Tan and Iwatsuki, 1993; Zhao and Cao, 1998).
Fissidens geminiflorus Dozy & Molk. – Zhang et al. (1998b)
Localities: NEW TERRITORIES. HO PUI RESERVOIR. 210 m, 807. – TAI LAM. Lin Fa Shan. 205
m (SW), 658. Kiu Keng Shan. 185 m (NW), 336. – TAI MO SHAN. Kadoorie Farm (Zhang et al.,
1998). Ng Tung Chai Falls. 395 m (N), 4(12). Kap Lung Forest Trail (Zhang et al., 1998). – TAI TO
YAN. Lam Tsuen Stream. 100 m (E), 698. – WU KAU TANG. Bride’s Pool (Zhang et al., 1998).
HONG KONG IS. VICTORIA PEAK. 305 m (SW), 1497. LANTAU IS. SHUI LO CHO. 250 m (N),
574, 575.
Habitat and Substrate: Seven of 8 specimens were collected in forest, and 1 unknown. All were found
on rocks.
Distribution: China (Anhui, Fujian, Guangdong, Guizhou, Hainan, Hong Kong, Jiangsu, Jiangxi,
Shandong, Sichuan, Taiwan, Yunnan, Zhejiang). Japan, Vietnam, Thailand, Malaysia, Philippines,
28
Zhang Li: Hong Kong Bryophytes 29
Borneo, Indonesia, Fiji (Gao, 1996; He, 2000; Iwatsuki and Suzuki,1996; Iwatsuki, Suzuki and Li,
1999; Mohamed and Tan, 1988; Redfearn et al., 1996; Tan and Iwatsuki, 1991, 1993; Touw, 1978;
Zhao and Cao, 1998).
Fissidens geppii Fleisch. – Zhang et al. (1998b)
Locality: NEW TERRITORIES. TAI MO SHAN. Kadoorie Farm. 470 m (NW), Lin et al. 186
(IBSC, KFBG); 490 m (NW), Lin et al. 226 (IBSC, KFBG).
Habitat and Substrate: On rocks in forest.
Distribution: China (Guangxi, Guizhou, Hong Kong, Taiwan, Yunnan). Korea, Japan, Eastern India,
Borneo, Indonesia (Gao, 1996; Redfearn et al., 1996; Touw, 1978).
Fissidens guangdongensis Iwats. & Li – Zhang et al. (1998b)
Locality: LANTAU IS. TAI O TO SHAM WAT. 50 m, Lin et al. 861 (IBSC, KFBG).
Habitat and Substrate: On soil in forest.
Distribution: China (Guangdong, Hainan, Hong Kong, Taiwan, Zhejiang). Japan, Malaysia, Singapore,
Philippines, Fiji (Gao, 1996; Iwatsuki and Suzuki, 1996; Redfearn et al., 1996; Tan, Lubos and
Schwarz, 2000).
Fissidens gymnogynus Besch. – Zhang et al. (1998b)
Localities: NEW TERRITORIES. TAI MO SHAN. Yin Ngam (Zhang et al., 1998). LANTAU IS.
LIN FA SHAN. 335 m (NE), 613. – SUNSET PEAK. Wong Lung Hang (Main Streams). 140 m (N),
1261; 250 m (N), 2088.
Habitat and Substrate: All 3 specimens were collected in forest. 2 were found on rocks, and 1 on soil.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hubei,
Hunan, Jilin, Shaanxi, Shandong, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Korea, Japan,
Thailand, Philippines (He, 2000; Rao et al., 1997; Redfearn et al., 1996; Tan and Iwatsuki, 1991).
Fissidens hollianus Dozy & Molk. – Zhang et al. (1998b)
Localities: NEW TERRITORIES. HO PUI RESERVOIR. 285 m, 814. – MA ON SHAN. Tiu Shau
Ngam. 215 m (NW), 264. – SHA TAU KOK. Muk Min Tau Feng Shui Wood. 10 m (S), 29(4). – TAI
MO SHAN. Kadoorie Farm (Zhang et al. 1998). – TAI TO YAN. Lam Tsuen Stream. 170 m (E), 711;
220 m (E), 717. – PAT SIN LENG. Sze Tau Leng. 115 m (NW), 988. – TAI LAM. Hung Shui Shan.
395 m (N), 421. LANTAU IS. SUNSET PEAK. Wong Lung Hang (East Lung Stream). 100 m (N),
39(11). OUTLYING ISLANDS. YEUNG CHAU (near Ma Shi Chau). 15 m, 378.
Habitat and Substrate: All 9 specimens were collected in forest. 6 were found on rocks, 2 on tree bases,
and 1 on rotten wood.
Distribution: China (Guangdong, Guangxi, Hainan, Hong Kong, Taiwan, Yunnan). Japan, Myanmar,
Vietnam, Thailand, Cambodia, Malaysia, Singapore, Philippines, Borneo, Indonesia, Papua New
Guinea, Fiji, Vanuatu (Gao, 1996; Higuchi, 1996; Iwatsuki and Suzuki, 1996; Mohamed and Tan,
1988; Redfearn et al., 1996; Tan and Iwatsuki, 1991; Touw, 1978).
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30 Memoirs of the Hong Kong Natural History Society 26, 2003
Fissidens cf. Incognitus Gangulee
Species new to Hong Kong.
Locality: NEW TERRITORIES. SHEUNG WO HANG FENG SHUI WOOD. 125 m (NW), 1072.
Habitat and Substrate: On rocks in forest.
Distribution: China (Hainan, Hong Kong, Shandong). Eastern India (Gao, 1996; Zhao and Cao, 1998).
Fissidens javanicus Dozy & Molk. – Zhang et al. (1998b)
Localities: NEW TERRITORIES. HUNG FA LENG. Muk Min Tau Stream. 220 m (E), 497. – KAU
LUNG HANG SHAN. 160 m (NW), 1548. – MA ON SHAN. Tiu Shau Ngam. 65 m (NW), 244; 150
m (NW), 254; 360 m (NW), 111, 112. Ma On Shan Tsuen Stream. 360 m (SW), 2158. – SAI KUNG.
Shek Uk Shan. 275 m (NW), 325. – TAI MO SHAN. Chuen Long Stream. 610 m (SW), 2429.
Kadoorie Farm (Zhang et al., 1998). Ng Tung Chai Falls. 700 m (N), 1(28). Shek Kong Stream. 640 m
(NW), 1992. Tai Shing Stream. 200 m (SE), 17(22). Yin Ngam (Zhang et al., 1998). Shing Mun
Reservoir. 200 m, 745. – TAI PO KAU (Zhang et al., 1998). – TAI TO YAN. Lam Tsuen Stream. 265
m (E), 722. – WU KAU TANG. Fan Kei Tok Feng Shui Wood. 200 m (N), 1602. HONG KONG IS.
MOUNT PARKER. 165 m (NW), 2381. LANTAU IS. NEI LAK SHAN. 30 m (NE), 2064.
Habitat and Substrate: All specimens were collected in forest. 14 were found on rocks, and 3 on soil.
Distribution: China (Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Taiwan, Xizang, Yunnan).
Japan, India, Nepal, Sri Lanka, Myanmar, Vietnam, Thailand, Malaysia, Singapore, Philippines,
Borneo, Indonesia, Papua New Guinea (Gao, 1996; Iwatsuki, Suzuki and Li, 1999; Jia and Wang,
1993; Redfearn et al., 1996; Tan and Iwatsuki, 1991; Touw, 1978).
Fissidens jungermannioides Griff.
Species new to Hong Kong, and this is the second report of this species occurred from mainland China.
Localities: NEW TERRITORIES. MA ON SHAN. Tai Tung Wo Liu Stream. 405 m (NE), 1580. Tiu
Shau Ngam. 340 m (NW), 101, 102 ( with Diphyscium involutum). – SAI KUNG. Shek Uk Shan. 260
m (NW), 328. LANTAU IS. LANTAU PEAK. 715 m (N), 1334. – SHUI LO CHO. 210 m (N), 562.
Habitat and Substrate: Five of 6 specimens were collected in forest, and 1 by a waterfall. All were
found on rocks.
Distribution: China (Guangdong, Hong Kong, Taiwan). Eastern India ( Gao, 1996; Yang and Lin,
1992).
Fissidens kinabaluensis Iwats. – Zhang et al. (1998b)
Localities: NEW TERRITORIES. LAM TSUEN. She Shan Tsuen Feng Shui Wood. 70 m, 1364. –
MA ON SHAN. Tiu Shau Ngam. 290 m (NW), 263. – PAT SIN LENG. Hsien Ku Fung. 420 m (NE),
1026. Ping Fung Shan. 340 m (SW), 405. – TAI MO SHAN. Kadoorie Farm (Zhang et al., 1998). –
TAI TO YAN. Lam Tsuen Stream. 425 m (E), 724. San Lung Wai Stream. 315 m (W), 762; 395 m
(W), 756. – WU KAU TANG. Fan Kei Tok Feng Shui Wood. 280 m (N), 1611.
Habitat and Substrate: All 8 specimens were collected in forest. 5 were found on soil, and 3 on rocks.
Distribution: China (Guangdong, Hong Kong, Taiwan, Yunnan). Thailand, Malaysia, Borneo (Gao,
1996; He, 2000; Iwatsuki, Suzuki and Li, 1999; Redfearn et al., 1996; Touw, 1978).
30
Zhang Li: Hong Kong Bryophytes 31
Fissidens linearis Brid. Var. obscurirete (Broth. & Par.) Stone (F. obscurirete Broth. & Par.) – Zhang
et al. (1998b)
Localities: NEW TERRITORIES. TAI MO SHAN. Kadoorie Farm. 350 m (NW), Lin et al. 661
(IBSC, KFBG); 580 m (NW), Lin et al. 1035 (IBSC, KFBG). Shing Mun. Lead Mine Pass. 350 m, Lin
et al. 1239 (IBSC, KFBG). – TAI PO KAU. 300m, Lin et al. 1876 (IBSC, KFBG). LANTAU IS.
LANTAU PEAK. Ngong Ping to Tei Tong Tsai. 500 m (N), Lin et al. 635, 640 (IBSC, KFBG). –
SUNSET PEAK. San She Wan Stream. 500 m (S), 34(14). OUTLYING ISLANDS. PO TAI IS. 200
m, Lin et al. 1919 (IBSC, KFBG).
Habitat and Substrate: Six of 8 specimens were collected in forest, 1 in grassland, and 1 in
Rhododendron shrubs. 6 were found on soil, 1 on rocks, and 1 on a trunk.
Distribution: China (Fujian, Guangdong, Guizhou, Hainan, Hong Kong, Liaoning, Shandong,
Shanghai, Taiwan, Yunnan). Korea, Japan, Philippines, Australia, New Caledonia, Fiji, Vanuatu (Gao,
1996; Higuchi, 1996; Iwatsuki and Suzuki, 1996; Li and Crosby, 2001; Redfearn et al., 1996; Tan and
Iwatsuki, 1991; Zhao and Cao, 1998).
Fissidens minutus Thwait. & Mitt. In Mitt. – Zhang et al. (1998b)
Localities: NEW TERRITORIES. HO CHUNG (Zhang et al. 1998). – HUNG FA LENG. Muk Min
Tau Stream. 170 m (E), 496; 220 m (E), 498. – LAM TSUEN. She Shan Tsuen Feng Shui Wood. 70 m,
1363. – MA ON SHAN. Tiu Shau Ngam. 130 m (NW), 728; 355 m (NW), 107. – PAT SIN LENG.
Vicinity of Ngau Au Stream. 140 m (SE), 1029. – SAI KUNG. Kap Man Hang. 70 m (NE), 1225. –
SHA TAU KOK. Muk Min Tau Feng Shui Wood. 10 m (S), 29(8). – TAI LAM. Hung Shui Shan. 320
m (N), 428. Kat Hing Bridge to Tsing Fai Tong. 120 m (NW), 1054; 135 m (NW), 1062; 140 m (NW),
1063. Kiu Keng Shan. 145 m (NW), 331. Lin Fa Shan. 350 m (SW), 675. – TAI MO SHAN. Kadoorie
Farm. Kap Lung Forest Trail (Zhang et al. 1998). Tai Shing Stream. 400 m (SE), 19(14); 600 m (SE),
21(22b) (with F. oblongifolius). Water-pumping House Stream. 700 m (E), S30. – TAI PO KAU
(Zhang et al. 1998). HONG KONG IS. D’AGUILAR PEAK. 160 m (W), 156, 157. LANTAU IS.
SUNSET PEAK. Wong Lung Hang (East Lung Stream). 800 m (N), 42(21).
Habitat and Substrate: All 19 specimens were collected in forest. 14 were found on rocks, 3 on soil,
and 2 on tree bases.
Distribution: China (Fujian, Guangdong, Hong Kong, Taiwan). Eastern Asia, Malesia, Fiji, Africa,
Mexico, Caribbean, North, Central and South America (Iwatsuki and Suzuki, 1996; Pursell,
Bruggeman-Nannenga and Iwatsuki, 1993; Redfearn et al., 1996; Sharp, Crum and Eckel, 1994).
Fissidens nobilis Griff. – Li and Crosby (2001); Zhang et al. (1998b)
Localities: NEW TERRITORIES. HUNG FA LENG. Muk Min Tau Stream. 35 m (E), 481; 85 m (E),
494. San Kwai Tin Stream. 100 m (N), 2145. – KOWLOON PEAK. Ho Chung Stream. 300 m, 1525. –
MA ON SHAN. Tai Tung Wo Liu Stream. 160 m (NE), 1570. – SAI KUNG. Tam Chai Shan. 55 m
(E), 1038. – TAI LAM. Hung Shui Shan. 355 m (N), 426. – TAI MO SHAN. Kadoorie Farm. Kap
Lung Forest Trail (Zhang et al. 1998). Ng Tung Chai Falls. 420 m (NW), 1409. Shek Kong Stream.
395 m (NW), 1974. Shing Mun (Zhang et al. 1998). Tai Shing Stream. 200 m (SE), 17(11). – TAI PO
KAU (Zhang et al. 1998). – TAI TO YAN. Lam Tsuen Stream. 180 m (E), 714; 390 m (E), 1202. –
WU KAU TANG. Bride’s Pool. Ha Miu Tin. Wu Kau Tang to Bride’s Pool (Zhang et al. 1998).
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32 Memoirs of the Hong Kong Natural History Society 26, 2003
HONG KONG IS. CHATHAM PATH. 245 m (N), 1745. – VICINITY OF TAI TAM RESERVOIR
(Zhang et al. 1998). LANTAU IS. LIN FA SHAN. 375 m (NE), 621. – NEI LAK SHAN. 30 m (NE),
2065. – LANTAU PEAK. Ngong Ping to Tei Tong Tsai (Zhang et al. 1998). Wong Lung Hang (East
Lung Stream). 400 m (N), 1271, 35(16); 500 m (N), 36(5). Wong Lung Hang (Main Streams). 260 m
(NW), 2399; 630 m (N), 49.
Habitat and Substrate: All 20 specimens studied were collected in forest. 16 were found on rocks, and 4
on soil.
Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Hainan, Heilongjiang, Hong Kong, Hubei,
Hunan, Jiangsu, Jiangxi, Shandong, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Korea, Japan,
Russian Far East, India, Nepal, Sri Lanka, Myanmar, Vietnam, Thailand, Cambodia, Malaysia,
Singapore, Philippines, Borneo, Indonesia, Fiji (Fang et al.,1998; Gao, 1996; Ignatov and Afonina,
1992; Mohamed and Tan, 1988; Rao et al., 1997; Redfearn et al., 1996; Tan and Iwatsuki, 1991, 1993;
Touw, 1978).
Fissidens oblongifolius Hook. F. & Wils. – Zhang et al. (1998b)
Localities: NEW TERRITORIES. MA ON SHAN. Tiu Shau Ngam. 145 m (NW), 731. – SAI
KUNG. Shek Uk Shan. 410 m (NW), 314. – TAI LAM. Hung Shui Shan. 385 m (N), 422. Kiu Keng
Shan. 185 m (NW), 333; 280 m (NW), 347; 410 m (NW), 352. – TAI LO SHAN. Kwun Yam Shan.
340 m (N), 965. – TAI MO SHAN. Kadoorie Farm (Zhang et al. 1998). Tai Shing Stream. 600 m (SE),
21(22a) (with F. minutus). Water-pumping House Stream. 700 m (E), S54. Yin Ngam (Zhang et al.
1998). – TAI PO KAU (Zhang et al. 1998). HONG KONG IS. D’AGUILAR PEAK. 135 m (W), 151.
– MOUNT CAMERON. 175 m (S), 1718. – MOUNT PARKER (Zhang et al. 1998). – NAM FUNG
ROAD FENG SHUI WOOD. 140 m (S), 14(3). – TAI TAM RESERVOIR (Zhang et al. 1998). –
VIOLET HILL. 270 m, 1368. LANTAU IS. SUNSET PEAK. Wong Lung Hang (East Lung Stream).
100 m (N), 39(9); 200 m (N), 40(14); 600 m (N), 37(21). Wong Lung Hang (Main Streams). 530 m
(N), 71; 570 m (N), 85. – YI TUNG SHAN (Zhang et al. 1998). OUTLYING ISLANDS. KAU SAI
CHAU. Tai Leng Stream. 130 m (NW), 276, 290; 135 m (NW), 283; 150 m (NW), 284; 155 m (NW),
286. – WONG YI CHAU. 25 m, 455. A small Island situated on NE of Yim Tin Tsai without name on
the map. 5 m, 465.
Habitat and Substrate: Twenty-three of 25 specimens were collected in forest, and 2 unknown. 24 were
found on rocks, and 1 on leaves.
Distribution: China (Fujian, Guangdong, Guizhou, Hainan, Hong Kong, Hunan, Sichuan, Taiwan,
Xizang). Japan, Thailand, Malaysia, Philippines, Australia, New Zealand, Fiji, Vanuatu, Society
Islands, Africa, South and Central America, Mexico (Fife, 1995; Gao, 1996; He, 2000; Higuchi, 1996;
Iwatsuki and Suzuki, 1996; Li and Crosby, 2001; Mohamed and Tan, 1988; O’Shea, 1999; Rao et al.,
1997; Redfearn et al., 1996; Scott, Stone and Rosser, 1976; Tan and Iwatsuki, 1991; Whittier, 1976).
Fissidens pellucidus Hornsch. (F. laxus Sull. & Lesq.) – Zhang et al. (1998b)
Localities: NEW TERRITORIES. MA ON SHAN. Ngau Ngak Shan Summit. 680 m (S), 13(2). –
PAK NAI (Zhang et al., 1998). – PAT SIN LENG. Pin Fung Shan. 330 m (SW), 406. – TAI MO
SHAN. Kadoorie Farm (Zhang et al., 1998). Ng Tung Chai Falls. 500 m (N), 3(17). Shing Mun Tai
Wai Feng Shui Wood. 225 m (W), 12(4) (with Calypogeia arguta). Summit. 925 m (E), 11(2); 945 m,
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Zhang Li: Hong Kong Bryophytes 33
1439. – VICINITY OF HO PUI RESERVOIR. 260 m (NW), 43(2). – WU KAU TANG. Mirror Pool
(Zhang et al., 1998). HONG KONG IS. MOUNT CAMERON. 265 m (S), 1721. – TAI TAM
RESERVOIR. 120 m, 1377. LANTAU IS. LANTAU PEAK. 50 m (SW), 2254; 135 m (SW), 2255. –
SUNSET PEAK. San She Wan Stream. 400 m (S), 33(8). Wong Lung Hang (East Lung Stream). 320
m (N), 41(17). – TAI O TO SHAM WAT (Zhang et al., 1998). OUTLYING ISLANDS. KAT O. 10
m, 2533.
Habitat and Substrate: There were 14 specimens studied, of which 9 were collected in forest, 4 in
grassland, and 1 in shrubland. 10 were found on soil, and 4 on rocks.
Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Shandong, Sichuan,
Taiwan, Yunnan, Zhejiang). Japan, India, Nepal, Sri Lanka, Myanmar, Vietnam, Thailand, Cambodia,
Malaysia, Singapore, Philippines, Borneo, Indonesia, Australia, Fiji, Vanuatu, Africa, North, Central
and South America, West Indies (Gao, 1996; Higuchi, 1996; Iwatsuki and Suzuki, 1996; Iwatsuki,
Suzuki and Li, 1999; Jia and Wang, 1993; O’Shea, 1999; Pursell, Bruggeman-Nannenga and Iwatsuki,
1992; Redfearn et al., 1996; Sharp, Crum and Eckel, 1994; Tan and Iwatsuki, 1991; Touw, 1978; Zhao
and Cao, 1998).
Fissidens polypodioides Hedw. (F. areolatus Griff.) – Zhang et al. (1998b)
Localities: NEW TERRITORIES. GRASS HILL. Fo Tan Stream. 435 m (SE), 208. – TAI MO
SHAN. Yin Ngam (Zhang et al., 1998). LANTAU IS. LIN FA SHAN. 350 m (NE), 614; 380 m (NE),
624. OUTLYING ISLANDS. KAU SAI CHAU. Tai Leng Stream. 135 m (NW), 282.
Habitat and Substrate: All specimens were collected in forest. 2 were found on rock, and 2 on soil
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Heilongjiang, Hong Kong,
Hunan, Jiangxi, Sichuan, Taiwan, Xizang, Yunnan). Japan, India, Nepal, Myanmar, Vietnam,
Thailand, Malaysia, Singapore, Philippines, Borneo, Indonesia, Papua New Guinea, North, Central and
South America, West Indies (Gao, 1996; Iwatsuki, Suzuki and Li, 1999; Pursell, Bruggeman-Nannenga
and Iwatsuki, 1992; Redfearn et al., 1996; Rao et al., 1997; Tan and Iwatsuki, 1991, 1993; Touw,
1978).
Fissidens serratus C. Muell. (F. papillous Sande Lac.) – Zhang et al. (1998b)
Locality: NEW TERRITORIES. TAI MO SHAN. Kadoorie Farm. 420 m (NW), Lin et al. 1196
(IBSC, KFBG).
Habitat and Substrate: On rocks in forest.
Distribution: China (Guizhou, Hainan, Hong Kong, Shandong, Taiwan, Zhejiang). Japan, Malaysia,
Singapore, Philippines, Borneo, Indonesia, Fiji, Africa, North, Central and South America, West Indies
(Gao, 1996; Iwatsuki and Suzuki, 1996; Mohamed and Tan, 1988; O’Shea, 1999; Redfearn et al.,
1996; Sharp, Crum and Eckel, 1994; Touw, 1978; Zhao and Cao, 1998).
Fissidens teysmannianus Dozy & Molk. (F. adelphinus Besch.) – Li and Crosby (2001); Zhang et al.
(1998b) as F. adelphinus
Localities: NEW TERRITORIES. TAI MO SHAN. Water-pumping House Stream. 700 m (E), S22,
S26, S48.
Habitat and Substrate: On rocks in forest.
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34 Memoirs of the Hong Kong Natural History Society 26, 2003
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hunan,
Jiangsu, Jiangxi, Shandong, Sichuan, Taiwan, Yunnan, Zhejiang). Korea, Japan, Russian Far East,
Malaysia, Indonesia (Fang et al., 1998; Gao, 1996; Ignatov and Afonina, 1992; Li and Crosby, 2001;
Rao et al., 1997; Redfearn et al., 1996).
Fissidens tosanensis Broth. – Zhang et al. (1998b)
Localities: NEW TERRITORIES. TAI MO SHAN. Kadoorie Farm. 550 m (NW), Lin et al. 92
(IBSC, KFBG). Summit, 900 m, Lin et al. 1335, Lin et al. 1339 (IBSC, KFBG).
Habitat and Substrate: There were 3 specimens studied, of which 1 was collected in forest, and 2
unknown. 1 was found on rocks, 1 on soil, and 1 on rusted ironware.
Distribution: China (Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hunan.
Jiangsu, Shaanxi, Shanghai, Sichuan, Taiwan, Yunnan, Zhejiang). Japan (Gao, 1996; Iwatsuki, Suzuki
and Li, 1999; Rao et al., 1997; Redfearn et al., 1996).
Fissidens zollingeri Mont. – Zhang et al. (1998b)
Locality: HONG KONG IS. HONG KONG PARK. 40 m, Lin et al. 779 (IBSC, KFBG).
Habitat and Substrate: On rocks.
Distribution: China (Guangdong, Guizhou, Hainan, Hong Kong, Jiangsu, ?Sichuan, Taiwan, Yunnan,
Zhejiang). Japan, Myanmar, Vietnam, Thailand, Cambodia, Malaysia, Singapore, Philippines, Borneo,
Australia, Fiji, Society Islands, Vanuatu, Africa, North, Central and South America, West Indies (Gao,
1996; He, 2000; Higuchi, 1996; Iwatsuki and Suzuki, 1996; Mohamed and Tan, 1988; Noguchi, 1987b;
O’Shea, 1999; Redfearn et al., 1996; Sharp, Crum and Eckel, 1994; Streimann and Curnow, 1989; Tan
and Iwatsuki, 1991; Touw, 1978; Whittier, 1976).
FUNARIACEAE
Funaria hygrometrica Mitt.
Localities: NEW TERRITORIES. TUEN MUN. Lung Kwu Tan Power Station. 80 m, Xing 18.
OUTLYING ISLANDS. TSING YI. 145 m, 985.
Habitat and Substrate: One on soil in grassland, and the other on a concrete wall by a stream.
Distribution: Cosmopolitan.
Physcomitrium eurystomum Sendtn.
Localities: NEW TERRITORIES. SHA TAU KOK. Tsiu Hang. 10 m, 45(3). TAI PO. Wun Yiu. 50
m, Lin et al. 1959 (IBSC, KFBG).
Habitat and Substrate: On soil in agricultural land and a flower pot.
Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Heilongjiang, Henan, Hong Kong, Inner
Mongolia, Jiangsu, Jilin, Liaoning, Shandong, Shanghai, Sichuan, Taiwan, Xinjiang, Xizang, Yunnan,
Zhejiang). Japan, Russian Far East, Central Asia, Europe, Africa (Bai, 1996; Li, 2000; Li, Zeng and Su,
1995; Redfearn et al., 1996; Zhao and Cao, 1998).
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Zhang Li: Hong Kong Bryophytes 35
Physcomitrium japonicum (Hedw.) Mitt.
Localities: NEW TERRITORIES. LAM TSUEN. San Uk Pai Tsuen. 50 m, 1910, 1911. – TAI LUNG
FARM. 20 m, Eddie M3, 3 (HKU, IBSC). HONG KONG (Locality unknown). Ford 150, Wright 19
(FH).
Habitat and Substrate: There were 6 specimens examined, of which 2 were in agricultural land, 1 in
tree seedling plastic tubes, 1 in forest, and 2 unknown. 4 were found on soil, and 2 unknown.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Heilongjiang, Hong Kong, Hubei,
Hunan, Jiangsu, Shanghai, Sichuan, Taiwan, Yunnan). Korea, Japan, India, Bhutan, Myanmar (Li,
2000; Long, 1994; Noguchi 1988; Rao et al., 1997; Redfearn et al., 1996; Tan and Iwatsuki, 1993).
Physcomitrium sphaericum (Ludw.) Fuernr. In Hampe – So and Zhu (1996c)
Locality: NEW TERRITORIES. TAI MO SHAN. Kadoorie Farm. 165 m (N), 47(1).
Habitat and Substrate: On soil in agricultural land.
Distribution: Widespread in North Hemisphere (Eddy 1996).
GRIMMIACEAE
Racomitrium fasciculare (Hedw.) Brid.
Family Grimmiaceae and genus and species new to Hong Kong.
Localities: LANTAU IS. LANTAU PEAK. 660 m (N), 1329; 815 m (N), 2473.
Habitat and Substrate: On rocks in forest.
Distribution: China (Anhui, Fujian, Hong Kong, Jiangxi, Jilin, Liaoling, Shaanxi, Shandong, Sichuan,
Taiwan, Xizang, Yunnan, Zhejiang). Japan, New Zealand, Hawaii, Russia, Europe, North America,
Southern South America (Anderson, Crum and Buck, 1990; Redfearn et al., 1996; Sharp, Crum and
Eckel, 1994).
HOOKERIACEAE
Callicostella papillata (Mont.) Mitt.
Localities: NEW TERRITORIES. GRASS HILL. Fo Tan Stream. 160 m (SE), 228. – HUNG FA
LENG. Ma Tseuk Leng Stream. 270 m (S), 2140. – PAT SIN LENG. Wang Chung Stream. 400 m (E),
5. – SAI KUNG. Shek Uk Shan. 385 m (NW), 315. – TAI LAM. Hung Shui Shan. 305 m (N), 432. Kiu
Keng Shan. 395 m (NW), 351; 435 m (NW), 354. – TAI MO SHAN. Tai Shing Stream. 390 m (SE),
1875; 400 m (SE), 19(3), 19(9); 500 m (SE), 20(16); 520 m (SE), 1915; 700 m (SE), 6(21). – TAI PO
KAU. 155 m (NE), 2327; 225 m (NE), 2336, 2337. HONG KONG IS. VICTORIA PEAK. 280 m
(SW), 1495. LANTAU IS. LIN FA SHAN. 165 m (E), 1443; 315 m (NE), 610; 410 m (E), 1447. –
NEI LAK SHAN. 255 m (NE), 2075. – SHUI LO CHO. 210 m (N), 563. – SUNSET PEAK. San Shek
Wan Stream. 350 m (S), 2038. Wong Lung Hang (Main Streams), 210 m (NW), 2396; 530 m (N), 74.
Habitat and Substrate: All 25 specimens were collected in forest. 24 were found on rocks, and 1 on a
tree root.
Distribution: China (Guangdong, Hainan, Hong Kong, Taiwan, Yunnan, Xizang). India, Sri Lanka,
Myanmar, Vietnam, Thailand, Cambodia, Malaysia, Singapore, Philippines, Borneo, Society Islands,
Vanuatu, Africa (Gangulee, 1969-1980; He, 2000; Higuchi, 1996; Lin and Tan, 1995; Mohamed and
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36 Memoirs of the Hong Kong Natural History Society 26, 2003
Tan, 1988; O’Shea, 1999; Redfearn et al., 1996; Tan and Iwatsuki, 1991, 1993; Touw, 1978; Whittier,
1976).
Distichophyllum collenchymatosum Card.
Localities: NEW TERRITORIES. TAI LAM. Hung Shui Shan. 375 m (N), 424. Kiu Keng Shan. 240
m (NW), 344; 305 m (NW), 349; 385 m (NW), 350. Lin Fa Shan. 150 m (SW), 657; 235 m (SW), 660.
– TAI MO SHAN. Chuen Lung Stream. 580 m (SW), 2426. Tai Shing Stream. 355 m (SE), 1874; 400
m (SE), 19(12); 500 m (SE), 20(14); 520 m (SE), 1916; 640 m (SE), 1920. LANTAU IS. SUNSET
PEAK. Wong Lung Hang (Main Streams). 550 m (N), 81; 595 m (N), 58, 59.
Habitat and Substrate: All specimens were collected in forest. 13 were found on rocks, 1 on soil, and 1
on humus.
Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hunan, Taiwan,
Yunnan, Zhejiang). Korea, Japan, Philippines (Koponen et al., 2000; Lin and Tan, 1995).
Distichophyllum maibarae Besch.
Localities: NEW TERRITORIES. HO PUI RESERVOIR. 235 m, 811. – HUNG FA LENG. San
Kwai Tin Stream. 200 m (N), 2144. – KAU LUNG HANG SHAN. 210 m (NW), 1550. – SHEUNG
WO HANG FENG SHUI WOOD. 135 m (NW), 1075, 1076; 150 m (NW), 1760. – TAI LAM. Hung
Shui Shan. 315 m (N), 430. Lin Fa Shan. 325 m (SW), 672. – TAI MO SHAN. Tai Shing Stream. 670
m (SE), 1921; 785 m (NE), 1358. LANTAU IS. LIN FA SHAN. 185 m (E), 1444. – POR KAI SHAN.
330 m (NE), 2410. – SUNSET PEAK. Wong Lung Hang (Main Streams). 530 m (N), 77; 540 m (N),
78.
Habitat and Substrate: All 14 specimens were collected in forest. 13 were found on rocks, and 1 on
soil.
Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hunan, Jiangxi,
Sichuan, Taiwan, Yunnan, Zhejiang). Japan, Northern India, Indochina, Philippines, Malesia (Koponen
et al., 2000; Lin and Tan, 1995).
Hookeria acutifolia Hook. & Grev.
Localities: NEW TERRITORIES. TAI MO SHAN. Chuen Lung Stream. 610 m (SW), 2428. Tai
Shing Stream. 800 m (NE), 7(7). HONG KONG IS. VICTORIA PEAK. 305 m (SW), 1494.
LANTAU IS. SUNSET PEAK. Wong Lung Hang (East Lung Stream). 500 m (N), 36(2); 550 m (N),
2244; 760 m (N), 2309.
Habitat and Substrate: All specimens were collected in forest. 3 were found on rocks, 2 on humus, and
1 on soil.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hunan,
Jiangsu, Jiangxi, Sichuan, Xizang, Yunnan, Taiwan, Zhejiang). Japan, Russian Caucasus, India, Nepal,
Bhutan, Vietnam, Philippines, Borneo, ?Australia, Hawaii, Africa, North, Central and South America
(Anderson, Crum and Buck, 1990; Bartram, 1933; Buck, 1998; Gangulee, 1969-1980; Ignatov and
Afonina, 1992; Lin and Tan, 1995; Long, 1994; Noguchi, 1991; Streimann and Curnow, 1989; Tan and
Iwatsuki, 1991, 1993; Touw, 1978; Yano, 1995).
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Zhang Li: Hong Kong Bryophytes 37
Hookeriopsis utacamundiana (Mont.) Broth.
Localities: NEW TERRITORIES. MA ON SHAN. Tiu Shau Ngam. 360 m (NW), 108. – TAI MO
SHAN. Shek Kong Stream. 750 m (NW), 2001; 800 m (NW), 2010. Tai Shing Stream. 800 m (NE),
7(16); 900 m (NE), 8(24) (with Heteroscyphus argutus). Water-pumping House Stream. 700 m (E), S2,
S19. LANTAU IS. SUNSET PEAK. Wong Lung Hang (East Lung Stream). 500 m (N), 36(11); 600 m
(N), 37(6), 37(11); 800 m (N), 42(16).
Habitat and Substrate: All specimens were collected in forest. 8 were found on rocks, 2 on submerged
rocks, and 1 on humus.
Distribution: China (Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Taiwan, Yunnan). Japan, Sri
Lanka, Vietnam, Thailand, Philippines, Borneo, the Pacific Islands (Gangulee, 1969-1980; He, 2000;
Lin and Tan, 1995; Tan and Iwatsuki, 1991, 1993; Touw, 1978).
HYLOCOMIACEAE
Macrothamnium macrocarpum (Reinw. & Hornsch.) Fleisch.
Genus and species as well as the family Hylocomiaceae new to Hong Kong.
Localities: NEW TERRITORIES. TAI MO SHAN. Tai Shing Stream. 700 m (SE), 6(7); 770 m (NE),
1356.
Habitat and Substrate: On rocks in forest.
Distribution: China (Fujian, Guangxi, Guizhou, Hong Kong, Taiwan, Yunnan, Xizang, Zhejiang).
Japan, India, Nepal, Bhutan, Sri Lanka, Myanmar, Vietnam, Thailand, Malaysia, Singapore,
Philippines, Borneo, Indonesia, Hawaii (Gangulee, 1969-1980; Mohamed and Tan, 1988; Noguchi,
1994; Redfearn et al., 1996; Tan and Iwatsuki, 1991, 1993).
HYPNACEAE
Ctenidium andoi Nishinuma – So and Zhu (1996c).
Locality: NEW TERRITORIES. TAI MO SHAN. Tai Shing Stream. 800 m (NE), 7(6).
Habitat and Substrate: On rocks in forest.
Distribution: China (Anhui, Hong Kong, Taiwan, Zhejiang). Japan, Philippines, Indonesia, Papua New
Guinea (Nishimura, 1985; Redfearn et al., 1996).
Ctenidium capillifolium (Mitt.) Broth. – So and Zhu (1996c).
Locality: NEW TERRITORIES. TAI MO SHAN. Tai Shing Stream. 600 m (SE), 21(17); 900 m
(NE), 8(16).
Habitat and Substrate: On rocks in forest.
Distribution: China (Anhui, Fujian, Guangxi, Guizhou, Hong Kong, Jiangsu, Liaoning, Sichuan,
Taiwan, Zhejiang). Korea, Japan (Nishimura, 1985; Redfearn et al., 1996).
Ctenidium serratifolium (Card.) Broth. – So and Zhu (1996c).
Locality: NEW TERRITORIES. TAI MO SHAN. Tai Shing Stream. 700 m (SE), 6(15).
Habitat and Substrate: On rocks in forest.
Distribution: China (Guangxi, Hainan, Hong Kong, Taiwan, Zhejiang). Japan, Vietnam, Thailand (He,
2000; Nishimura, 1985; Redfearn et al., 1996; Tan and Iwatsuki, 1993).
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38 Memoirs of the Hong Kong Natural History Society 26, 2003
Ectropothecium dealbatum (Hornsch. & Reinw.) Jaeg.
Localities: HONG KONG IS. WEST POINT. Wong Nin 296G, 299Aa (BM). HONG KONG
(Locality unknown). Herklots H59 (BM).
Habitat and Substrate: Unknown.
Distribution: China (Guangdong, Hainan, Hong Kong, Taiwan). India, Myanmar, Thailand, Malaysia.
Philippines, Borneo, Indonesia (Bartram, 1939; Gangulee, 1969-1980; He, 2000; Mohamed and Tan,
1988; Tan and Iwatsuki, 1991, 1993; Touw, 1978).
Ectropothecium monumentorum (Duby) Jaeg.
Localities: KOWLOON. Ah Nin H39A (BM), Herklots H68 (BM). HONG KONG (Locality
unknown). Ah Nin H.39B, H39C (BM).
Habitat and Substrate: One specimen on a tree near sea level, the other unknown.
Distribution: China (Guangdong, Hong Kong, Taiwan). India, Myanmar, Vietnam, Thailand,
Singapore, Philippines, Borneo, Indonesia, Caroline Islands (Bartram, 1939; Gangulee, 1969-1980; He,
2000; Mohamed and Tan, 1988; Redfearn et al., 1996; Tan and Iwatsuki, 1991, 1993; Touw, 1978).
Ectropothecium obtusulum (Card.) Iwats.
Localities: NEW TERRITORIES. KOWLOON PEAK. Ho Chung Stream. 225m (E), 1522. – PAT
SIN LENG. Nam Chung Stream. 200 m (NE), 1005. – TAI MO SHAN. Chuen Lung Stream. 350 m
(SW), 2419. – TAI TO YAN. Lam Tsuen Stream. 200 m (E), 789.
Habitat and Substrate: All specimens were collected in forest. 3 were found on rocks, and 1 on leaves.
Distribution: China (Guangxi, Hong Kong, Taiwan). Japan (Noguchi, 1994; Redfearn et al., 1996).
Ectropothecium zollingeri (C. Muell.) Jaeg. (Ectropothecium nervosum Dix., syn. Nov.) – Nishimura
(1985), Reimers (1931) as E. planulum Card.
I have studied the type specimen of Ectropothecium nervosum Dix. (Bowring s.n., BM) and authentic
materials (Herklots 254b, 257a, 257b, BM) cited by Dixon (1933), and I could not find any reliable
characteristics to separate it from E. zollingeri.
Localities: NEW TERRITORIES. GRASS HILL. Fo Tan Stream. 160 m (SE), 227. – HUNG FA
LENG. Ma Tseuk Leng Stream. 15 m (S), 2135; 285 m (S), 2142. Mu Min Tau Stream. 30 m (E), 480;
85 m (E), 493. San Kwai Tin Stream. 395 m (N), 1158. – KAU LUNG HANG SHAN. 30 m (NW),
1534; 105 m (NW), 1547. – PAT SIN LENG. Ping Shan Tsai. 130 m, 1556. – SAI KUNG. Shek Uk
Shan. 290 m (NW), 322. Yung Shue O to Pat Tam Chung. 165 m, 1383. – TAI LAM. Kiu Keng Shan.
215 m (NW), 342. Lin Fa Shan. 40 m (SW), 655. – TAI LO SHAN. Kwun Yam Shan. 90 m (N), 930;
140 m (N), 942. – TAI MO SHAN. Ng Tung Chai Falls. 300 m (N), 5(14). Tai Shing Stream. 200 m
(SE), 17(2); 400 m (SE), 19(1); 500 m (SE), 20(13); 700 m (SE), 6(19); 900 m (NE), 8(9). Waterpumping House Stream. 700 m (E), S23, S37. – TAI PO. Herklots 254b, 257a, 257b (BM). Sha Lo
Tung. 70 m, 1559. – WU KAU TANG. Mirror Pool Feng Shui Wood. 60 m, 1750. HONG KONG IS.
MOUNT PARKER. 170 m (NW), 2382. – STANLEY. Herklots 298 (BM). – WEST POINT. Wong
Nin 299Ba (BM). LANTAU IS. HANG PUI CATCHWATER. 110 m, 554. – LIN FA SHAN. 410 m
(E), 1446. – SUNSET PEAK. Wong Lung Hang (East Lung Stream). 500 m (N), 36(4). – TUNG
38
Zhang Li: Hong Kong Bryophytes 39
CHUNG. Mok Ka. 20 m, Ng 20. OUTLYING ISLANDS. LAMMA IS. So Kwu Wan. 30 m, Lin et al.
1982 (IBSC, KFBG). HONG KONG (Locality unknown). Bowring s.n. (Type of E. nervosum, BM).
Habitat and Substrate: 25 of 37 specimens were collected in forest, 4 by stream, 1 by a waterfall, and 7
unknown. 28 were found on rocks, 4 on soil, 1 on a twig, 1 on a tree base, and 3 unknown.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hong Kong, Hunan, Jiangsu,
Jiangxi, Sichuan, Taiwan, Yunnan, Zhejiang). Japan, Laos, Vietnam, Thailand, Malaysia, Singapore,
Philippines, Borneo, Indonesia, New Caledonia, Hawaii, Vanuatu (Fang et al., 1998; He, 2000;
Higuchi, 1996; Higuchi and Iwatsuki, 1994; Iwatsuki, 1967; Mohamed and Tan, 1988; Rao et al.,
1997; Redfearn et al., 1996; Tan and Iwatsuki, 1991, 1993; Touw, 1978).
Hypnum plumaeformae Wils. Var. plumaeformae
Localities: NEW TERRITORIES. HUNG FA LENG. Muk Min Tau Stream. 50 m (E), 489. –
KOWLOON PEAK. Pak Fa Lam. 160 m (E), 1532. – PAT SIN LENG. Pin Shan Tsai. 135 m (N),
1558. – PLOVER COVE RESERVOIR. Shek Nga Tal. 210 m, 138. – TAI MO SHAN. Summit. 930
m, Siu 6. Tai Shing Stream. 800 m (NE), 7(14); 900 m (NE), 9. LANTAU IS. LANTAU PEAK. 915
m, 2481. SHUI LO CHO. 254 m (N), 569. – SUNSET PEAK. Wong Lung Hang (East Lung Stream).
200 m (N), 40(2). Wong Lung Hang (Main Streams). 700 m (N), 2126. OUTLYING ISLANDS. AP
CHAU. 5 m, 2494; Xing 1. – FU WONG CHAU. 10 m, 2511. – KAU SAI CHAU. Tai Leng Stream. 3
m (NW), 295. – PAK SHA CHAU. 10 m, 2524. – SAM PUI CHAU. 5 m, 361, 367. – SIU AP CHAU.
5 m, 2490. – TSING CHAU. 5 m, 2504. – YEUNG CHAU (near Kat O). 15 m, 2530.
Habitat and Substrate: Nine of 21 specimens were collected in forest, 4 in grassland, 3 in shrubland, 1
by the seashore, and 4 unknown. 15 were found on rocks, 3 on soil, 2 on trunks, and 1 on a tree root.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hebei, Heilongjiang,
Henan, Hong Kong, Hubei, Hunan, Inner Mongolia. Jiangsu, Jiangxi, Jilin, Liaoning, Shaanxi,
Shandong, Shanghai, Sichuan, Xizang, Yunnan, Zhejiang). Korea, Japan, Russian Far East, Nepal,
Myanmar, Vietnam, Hawaii (Fang et al., 1998; Gangulee, 1969-1980; Noguchi, 1994; Redfearn et al.,
1996; Rao et al., 1997; Tan and Iwatsuki, 1993; Zhao and Cao, 1998).
Hypnum plumaeformae Wils. Var. minus Broth. Ex Ando
Localities: NEW TERRITORIES. FAN LING. Kei Lak Tsai. 205 m, 39. – PAT SIN LENG. Pin Fung
Shan. 370 m (SW), 404; 460 m (SW), 399. – PLOVER COVE RESERVOIR. Ngor Kai Teng. 210 m,
129. – TAI LAM. Kat Hing Bridge to Tsing Fai Tong. 115 m (NW), 1053. – TAI MO SHAN. Ng Tung
Chai Falls. 200 m (N), 16(5); 700 m (N), 1(24); 750 m (N), 1242. HONG KONG IS. D’AGUILAR
PEAK. 200 m (E), 189; 205 m (E), 537; 230 (E), 517. LANTAU IS. SUNSET PEAK. San Shek Wan
Stream. 200 m (S), 31(2), 31(6); 250 m (S), 2021; 400 m (S), 33(19); 500 m (S), 34(16); 600 m (S),
26(7). OUTLYING ISLANDS. A CHAU. 5 m, 358. – FU WONG CHAU. 10 m, 2514. – LAMMA IS.
So Kwu Wan, 150 m, Lin et al. 1970 (IBSC, KFBG). – SAM PUI CHAU. 5 m, 365, 366. – SIU NIM
CHAU. 10 m, 2506, 2510. – TSING CHAU. 10 m, 2499. – TSING YI. 295 m, 983.
Habitat and Substrate: There were 26 specimens studied, of which 23 were collected in forest, 2 in
shrubland, and 1 unknown. 19 were found on rocks, 4 on trunks, 2 on soil, and 1 on a twig.
Distribution: China (Anhui, Hong Kong, Taiwan). Japan (Noguchi, 1994; Redfearn et al., 1996).
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40 Memoirs of the Hong Kong Natural History Society 26, 2003
Isopterygium albescens (Hook.) Jaeg.
Localities: NEW TERRITORIES. MA ON SHAN. Ma On Shan Tsuen Stream. 580 m (SW), 2168.
Tiu Shau Ngam. 130 m (NW), 729. – PAT SIN LENG. Ping Shan Tsai. 135 m, 1557. – PLOVER
COVE RESERVOIR. Ngor Kai Teng. 210 m, 135. – TAI LAM. Hung Shui Shan. 455 m (N), 416. Lin
Fa Shan. 125 m (SW), 654. – TAI MO SHAN. Chuen Lung Stream. 350 m (SW), 2417. Shek Kong
Stream. 765 m (NW), 2292. Tai Shing Stream. 100 m (SE), 30(7); 600 m (SE), 21(6); 700 m (SE),
6(10), 6(22); 780 m (NE), 1430; 875 m (NE), 1132; 890 m (NE), 1141; 900 m (NE), 8(13), 8(20); 925
m (NE), 1106. Water-pumping House Stream. 700 m (E), S18, S33. – TAI TO YAN. San Lung Wai
Stream. 315 m (W), 763. – WU KAU TANG. Fan Kei Tok Feng Shui Wood. 280 m (N), 1613. Lai Chi
Wo. 15 m, 1614. HONG KONG IS. HATTON RD. 190 m, 1485. – MOUNT CAMERON. 160 m (S),
1717. LANTAU IS. LANTAU PEAK. 540 m (S), 1676. – SUNSET PEAK. San She Wan Stream. 500
m (S), 34(4); 600 m (S), 26(8); 700 m (SE), 27(10), 27(15). Wong Lung Hang (East Lung Stream). 320
m (N), 41(9), 600 m (N), 37(16). OUTLYING ISLANDS. AP CHAU. 10 m, 2495. – KAI CHAU. 20
m, 444. – MA WAN. 10 m, 840. – TSING CHAU. 10 m, 2501. – YEUNG CHAU (near Ma Shi Chau).
15 m, 377.
Habitat and Substrate: Thirty-four of 37 specimens were collected in forest, 1 in bamboo forest, and 2
by the seashore. 12 were found on rocks, 5 on rotten wood, 8 on trunks, 3 on tree bases, 2 on bark, 2 on
lianas, 2 on rotten twigs, 2 on soil, and 1 on a twig.
Distribution: China (Anhui, Guangxi, Hainan, Hong Kong, Taiwan, Xizang, Yunnan). Japan,
Myanmar, Laos, Vietnam, Thailand, Cambodia, Malaysia, Singapore, Philippines, Borneo, Australia,
New Zealand, Vanuatu, Society Islands, Hawaii (Bartram, 1933; Fife, 1995; He, 2000; Higuchi, 1996;
Jia and Wang, 1993; Mohamed and Tan, 1988; Noguchi, 1994; Redfearn et al., 1996; Scott, Stone and
Rosser, 1976; Tan and Iwatsuki, 1991, 993; Touw, 1978; Whittier, 1976).
Isopterygium minutirameum (C. Muell.) Jaeg. (Taxiphyllum minutirameum (C. Muell.) Miller &
Smith)
Localities: NEW TERRITORIES. SHA TAU KOK. Muk Min Tau Feng Shui Wood. 10 m (S), 29(2).
HONG KONG IS. D’AGUILAR PEAK. 220 m (E), 533.
Habitat and Substrate: Both were collected in forest. One on a tree base, and the other on a fern root.
Distribution: China (Anhui, Guangxi, Guizhou, Hong Kong, Jiangxi, Shaanxi, Taiwan, Xizang,
Yunnan). Japan, India, Sri Lanka, Myanmar, Laos, Vietnam, Thailand, Cambodia, Malaysia,
Singapore, Philippines, Borneo, Australia, New Zealand, Vanuatu (Fang et al., 1998; Fife, 1995;
Gangulee, 1969-1980; He, 2000; Higuchi, 1996; Jia and Wang, 1993; Mohamed and Tan, 1988;
Noguchi, 1994; Redfearn et al., 1996; Tan and Iwatsuki, 1991, 1993; Touw, 1978).
Phyllodon lingulatus (Card.) Buck (Glossadelphus lingulatus (Card.) Fleisch.)
Locality: NEW TERRITORIES. TAI TO YAN. Lam Tsuen Stream. 175 m (E), 712.
Habitat and Substrate: On rocks in forest.
Distribution: China (Hong Kong, Taiwan, Xizang, Yunnan). Japan, Vietnam, Philippines (Noguchi,
1994; Redfearn et al., 1996; Tan and Iwatsuki, 1991).
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Zhang Li: Hong Kong Bryophytes 41
Pseudotaxiphyllum pohliaecarpum (Sull. & Lesq.) Iwats.
Localities: NEW TERRITORIES. LAM TSUEN. She Shan Tsuen Feng Shui Wood. 70 m, 1361. –
MA ON SHAN. Tiu Shau Ngam. 135 m (NW), 249; 180 m (NW), 258; 215 m (NW), 265; 330 m
(NW), 97. – PAT SIN LENG. Sze Tau Leng. 245 m (NW), 992. – TAI LAM. Hung Shui Shan. 346 m
(N), 427. Kat Hing Bridge to Tsing Fai Tong. 95 m (NW), 1045. Kiu Keng Shan. 210 m (NW), 341. –
TAI LO SHAN. Kwun Yam Shan. 150 m (N), 948. – TAI MO SHAN. Midway to Lead Mine Pass.
915 m, 1472. Ng Tung Chai Falls. 500 m (N), 3(2); 700 m (N), 1(2), 1(17), 1(30). Shing Mun Tai Wai
Feng Shui Wood. 225 m (W), 12(2), 12(3). Summit. 925 m (E), 11(3). Tai Shing Stream. 300 m (SE),
18(12); 795 m (NE), 20; 800 m (NE), 7(9); 900 m (NE), 8(23). Water-pumping House Stream. 700 m
(E), S38, S51. – TAI TO YAN. San Lung Wai Stream. 335 m (W), 758. HONG KONG IS.
CENTRAL. 45 m (NE), 23(2). – HIGH WEST. 470 m (N), 1735. – VICTORIA PEAK. 460 m, 1739;
540 m, 1742. LANTAU IS. LANTAU PEAK. 430 (N), 1312. – SUNSET PEAK. San She Wan
Stream. 400 m (S), 33(7); 500 m (S), 34(9); 600 m (S), 26(3); 700 m (SE), 27(3), 27(8). Wong Lung
Hang (East Lung Stream). 320 m (N), 41(2), 41(3); 600 m (N), 37(4), 37(22); 700 m (N), 38(17); 800
m (N), 42(25). OUTLYING ISLANDS. FU WONG CHAU. 15 m, 2517. – KAT O. 20 m, 2537. –
TSING YI. 270 m, 982; 300 m, 979. – WONG YI CHAU. 25 m, 457.
Habitat and Substrate: Forty-two of 46 specimens were collected in forest, 1 by a road, 1 in grassland,
1 in an urban area, and 1 unknown. 30 were found on rocks, 10 on soil, 3 on tree bases, 1 on rocks and
soil,1 on a concrete wall, and 1 on humus.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hubei,
Hunan, Jiangsu, Jiangxi, Shandong, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Japan, India, Sri
Lanka, Myanmar, Laos, Vietnam, Thailand, Cambodia, Malaysia, Philippines, Borneo, Vanuatu (Fang
et al., 1998; Gangulee, 1969-1980; He, 2000; Higuchi, 1996; Mohamed and Tan, 1988; Rao et al.,
1997; Redfearn et al., 1996; Tan and Iwatsuki, 1991, 1993; Touw, 1978; Zhao and Cao, 1998).
Stereodontopsis pseudorevoluta (Reim.) Ando – Ando (1963)
Localities: LANTAU IS. LANTAU PEAK. 915 m, Herklots H27B, H35B (BM). HONG KONG
(Locality unknown). Herklots H29C (BM).
Habitat and Substrate: One on igneous rocks, and 2 unknown.
Distribution: China (Anhui, Guangxi, Hong Kong). Japan (Noguchi, 1994).
Taxiphyllum arcuatum (Bosch. & Sande Lac.) He (T. subarcuatum (Broth.) Iwats.) – So and Zhu
(1996c) as T. subarcuatum
Localities: NEW TERRITORIES. SHEUNG WO HANG FENG SHUI WOOD. 130 m (NW), 1073.
– TAI LO SHAN. Kwun Yam Shan. 145 m (N), 955. – TAI MO SHAN. 800 m (So and Zhu 1996c).
HONG KONG IS. MOUNT PARKER. 340 m (NW), 2389.
Habitat and Substrate: All 3 were collected in forest. 2 were found on rocks, and 1 on a trunk.
Distribution: China (Guizhou, Hainan, Hong Kong, Hubei, Hunan, Jiangsu, Taiwan, Xizang). Japan,
Thailand, Indonesia (He, 1997; Rao et al., 1997; Redfearn et al., 1996).
Taxiphyllum pilosum (Broth. & Yas.) Iwats. – So and Zhu (1996c).
Locality: HONG KONG (Locality unknown). 300 m (So and Zhu, 1996c).
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42 Memoirs of the Hong Kong Natural History Society 26, 2003
Habitat and Substrate: On soil (So and Zhu, 1996c).
Distribution: China (Hong Kong, Taiwan, Zhejiang). Japan (Matsui, 1995; Redfearn et al., 1996).
Taxiphyllum taxirameum (Mitt.) Fleisch.
Localities: NEW TERRITORIES. HUNG FA LENG. Muk Min Tau Stream. 20 m (E), 477. – TAI
MO SHAN. Ng Tung Chai Falls. 175 m (N), 1773; 380 m (NW), 1406. Shing Mun Reservoir. 175 m,
741. – TAI TO YAN. Lam Tsuen Stream. 225 m (E), 716. KOWLOON. TSIM SHA TSUI. 5 (W),
25(5). HONG KONG IS. D’AGUILAR PEAK. 110 m (W), 141(with Brotherella erythrocaulis); 150
m (W), 153. – NAN FUNG ROAD FENG SHUI WOOD. 135 m (S), 1711. – VICTORIA PEAK. 405
m, 1505; 525 m, Iwatsuki 1, 2 (IBSC, NICH). LANTAU IS. LUK TEI TONG FENG SHUI WOOD.
70 m (NE), 28(1). – SUNSET PEAK. San Shek Wan Stream. 185 m (S), 2025. OUTLYING
ISLANDS. LAMMA IS. So Kwu Wan. 70 m, Lin et al. 1992 (IBSC, KFBG). – PING CHAU. 5 m,
2274; 40 m, 2278.
Habitat and Substrate: There were 17 specimens studied, of which 13 were collected in forest, 1 in an
urban area, 1 by a road, and 2 unknown. 11 were found on rocks, 3 on soil, 1 on a tree base, 1 on steps,
and 1 on a concrete wall.
Distribution: China (Anhui, Fujian, Guangdong, Guizhou, Hainan, Henan, Hong Kong, Hubei, Hunan,
Inner Mongolia, Jiangsu, Jiangxi, Jilin, Liaoning, Shaanxi, Shandong, Shanghai, Sichuan, Taiwan,
Xizang, Yunnan, Zhejiang). Korea, Japan, Russian Far East, India, Nepal, Bhutan, Bangladesh,
Myanmar, Laos, Vietnam, Thailand, Malaysia, Singapore, Philippines, Australia, Vanuatu, North,
Central and South America (Anderson, Crum and Buck, 1990; Bai, 1996; Buck, 1998; Fang et al.,
1998; Gangulee, 1969-1980; He, 2000; Higuchi, 1996; Mohamed and Tan, 1988; Rao et al., 1997,
Redfearn et al., 1996; Tan and Iwatsuki, 1991; Yano, 1995; Zhao and Cao, 1998).
Vesicularia ferriei (Card. & Ther.) Broth.
Localities: NEW TERRITORIES. PAT SIN LENG. Pin Shan Tsai. 125 m, 1555. – TAI LAM. Lin Fa
Shan. 40 m (SW), 647. – TAI MO SHAN. Chuen Lung Stream. 520 m (SW), 2424. Tai Yuen Stream.
455 m (S), 1852. – WU KAU TANG. Mirror Pool Feng Shui Wood. 60 m, 1751. HONG KONG IS.
HIGH WEST. 475 m (N), 1731. – VICTORIA PEAK. 295 m (SW), 1496; 475 m, 1743. – WEST
POINT. Herklots 296H (BM).
Habitat and Substrate: Eight of 9 specimens were collected in forest, and 1 unknown. 8 were found on
rocks, and 1 unknown.
Distribution: China (Fujian, Guangxi, Hong Kong, Jiangsu, Jiangxi, Shanghai, Xizang, Yunnan,
Zhejiang). Japan (Fang et al., 1998; Noguchi, 1994; Redfearn et al., 1996).
Vesicularia inflectens (Brid.) C. Muell. (Ectropothecium inflectens Jaeg.)
Locality: HONG KONG (Locality unknown) (Salmon 1990).
Habitat and Substrate: Unknown.
Distribution: China (Hong Kong). Borneo, ?Australia, Pacific Islands (Streimann and Curnow, 1989;
Touw, 1978; Whittier, 1976).
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Zhang Li: Hong Kong Bryophytes 43
Vesicularia montagnei (Bel.) Broth.
Species new to Hong Kong.
Localities: NEW TERRITORIES. KOWLOON PEAK. Ho Chung Stream. 150 m (E), 1519. – TAI
MO SHAN. Ng Tung Chai Falls. 300 m (N), 5(13) (with Riccardia multifida). – TAI TO YAN. San
Lung Wai Stream. 280 m (W), 764. HONG KONG (Locality unknown). Wright s.n. (with Callicostella
papillata, FH).
Habitat and Substrate: Three of 4 specimens were collected on rocks in forest, 1 unknown.
Distribution: China(Guangxi, Hong Kong, Sichuan, Taiwan, Yunnan). Japan, Bangladesh, Sri Lanka,
Myanmar, Vietnam, Thailand, Malaysia, Singapore, Borneo, Indonesia, Australia, Africa (Gangulee,
1969-1980; Mohamed and Tan, 1988; Noguchi, 1994; O’Shea, 1999; Redfearn et al., 1996;
Stoneburner et al., 1993; Tan and Iwatsuki, 1991, 1993).
Vesicularia reticulata (Dozy & Molk.) Broth.
Localities: NEW TERRITORIES. TAI MO SHAN. Ng Tung Chai Falls. 300 m (N), 5(15). HONG
KONG IS. D’AGUILAR PEAK. 235 m (W), 166. LANTAU IS. SUNSET PEAK. Wong Lung Hang
(East Lung Stream). 700 m (N), 38(13), 38(23). HONG KONG (Locality unknown). Wright s.n. (FH).
Habitat and Substrate: Four of 5 specimens were collected in forest, and 1 unknown. All were found on
rocks.
Distribution: China (Anhui, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Shandong, Taiwan,
Xizang, Yunnan). Japan, India, Nepal, Myanmar, Vietnam, Thailand, Cambodia, Malaysia, Singapore,
Philippines, Borneo, Indonesia, Australia (Gangulee, 1969-1980; He, 2000; Mohamed and Tan, 1988;
Noguchi, 1994; Redfearn et al., 1996; Streimann and Curnow, 1989; Tan and Iwatsuki, 1991, 1993;
Touw, 1978; Zhao and Cao, 1998).
HYPOPTERYGIACEAE
Cyathophorella tonkinensis (Broth. & Par.) Broth.
Localities: NEW TERRITORIES. TAI LO SHAN. Kwun Yam Shan. 430 m (N), 973. – TAI MO
SHAN. Shek Kong Stream. 500 m (NW), 1981. – TAI TO YAN. Lam Tsuen Stream. 225 m (E), 1260;
250 m (E), 721; 285 m (E), 723. LANTAU IS. SUNSET PEAK. Wong Lung Hang (East Lung
Stream). 740 m (N), 1294.
Habitat and Substrate: On rocks in forest.
Distribution: China (Guangdong, Guizhou, Hainan, Hong Kong, Sichuan, Taiwan, Yunnan, Zhejiang).
Japan, Vietnam, Thailand, Borneo (He, 2000; Noguchi, 1991; Redfearn et al.,1996).
Hypopterygium tenellum C. Muell. (Hypopterygium sinicum Mitt., fide Z.-H. Li, pers com.).
Localities: NEW TERRITORIES. KOWLOON PEAK. Ho Chung Stream. 130 m (E), 1516; 300 m,
1524. – MA ON SHAN. Mui Tsz Lam Tsuen. 225 m, 2218. – TAI TO YAN. Lam Tsuen Stream. 140
m (N), 947; 170 m (E), 710; 240 m (E), 720. – TAI LO SHAN. Kwun Yam Shan. 160 m (N), 951. –
TAI MO SHAN. Ng Tung Chai Falls. 265 m (NW), 1401; 630 m (N), 1191. Shek Kong Stream. 750 m
(NW), 2298. HONG KONG IS. D’AGUILAR PEAK. 235 m (E), 178; 245 m (E), 510; 270 m (E),
181; 305 m (E), 172. – MOUNT PARKER. 270 m (NW), 2384. LANTAU IS. LANTAU PEAK. 305
m (N), 1305. – LIN FA SHAN. 385 m (NE), 625; 555 m (NE), 637. – SUNSET PEAK. Wong Lung
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44 Memoirs of the Hong Kong Natural History Society 26, 2003
Hang (East Lung Stream). 400 m (N), 35(17); 400 m (N), 1272; 500 m (N), 36(20). Wong Lung Hang
(Main Streams). 390 m (N), 2108; 540 m (N), 79.
Habitat and Substrate: Twenty-two of 23 specimens were collected in forest, and 1 unknown. 19 were
found on rocks, 1 on a tree base, 1 on a tree root, 1 on a rotten twig, and 1 on a concrete wall.
Distribution: China (Hainan, Hong Kong, Taiwan, Zhejiang). Japan, Sri Lanka, Vietnam, Thailand,
Malaysia, Philippines, Borneo, Indonesia, Papua New Guinea, Australia, Vanuatu, Africa (He, 2000;
Higuchi, 1996; Mohamed and Tan, 1988; Noguchi, 1991; O’Shea, 1999; Redfearn et al., 1996;
Streimann and Curnow ,1989; Tan and Iwatsuki, 1991).
Lopidium nazeense (Ther.) Broth.
Locality: HONG KONG IS. D’AGUILAR PEAK. 210 m (E), 187.
Habitat and Substrate: On rocks in forest.
Distribution: China (Hainan, Hong Kong, Taiwan). Japan (Noguchi, 1991; Redfearn et al., 1996).
LEPTODONTACEAE
Caduciella guangdongensis Enroth
Locality: NEW TERRITORIES. TAI MO SHAN. 880 m, So 95501 (HKBU, IBSC).
Habitat and Substrate: Unknown.
Distribution: Endemic to South China (Guangdong, Hainan, Hong Kong) (Zhang 1994).
LESKEACEAE
Pseudoleskeopsis zippelii (Dozy & Molk.) Broth.
Localities: NEW TERRITORIES. GRASS HILL. Fo Tan Stream. 205 m (SE), 226. – HO PUI
RESERVOIR. 175 m (N), 800. – HUNG FA LENG. Muk Min Tau Stream. 70 m (E), 589. – TAI
LAM. Kat Hing Bridge to Tsing Fai Tong. 125 m (NW), 1056. – TAI MO SHAN. Ng Tung Chai Falls.
200 m (N), 16(7); 450 m (N), 1349; 500 m (N), 3(6). Tai Shing Stream. 200 m (SE), 17(18); 500 m
(SE), 20(17); 700 m (SE), 6(3) (with Thuidium pristocalyx). – TAI TO YAN. Lam Tsuen Stream. 65 m
(E), 692; 85 m (E), 684. HONG KONG IS. D’AGUILAR PEAK. 250 m (W), 230. LANTAU IS. LIN
FA SHAN. 250 m (NE), 605; 495 m (NE), 633. – SUNSET PEAK. San Shek Wan Stream. 350 m (S),
2031; 400 m (S), 33(17). Wong Lung Hang (East Lung Stream). 500 m (N), 36(17).
Habitat and Substrate: Seventeen of 18 specimens examined were collected in forest, and 1 by a
waterfall. 16 were found on rocks, and 2 on leaves.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hebei, Heilongjiang,
Henan, Hong Kong, Hubei, Hunan, Jiangsu, Jiangxi, Jilin, Liaoning, Shandong, Shanghai, Taiwan,
Yunnan, Zhejiang). Korea, Japan, Russia, India, Sri Lanka, Vietnam, Thailand, Malaysia, Philippines,
Borneo, Indonesia, Papua New Guinea, Australia (Fang et al., 1998; Ignatov and Afonina, 1992;
Koponen et al., 2000; Redfearn et al., 1996; Touw, 1978; Zhao and Cao, 1998).
LEUCOBRYACEAE
Leucobryum aduncum Dozy & Molk.
Localities: LANTAU IS. SUNSET PEAK. Wong Lung Hang (East Lung Stream). 200 m (N), 40(1).
OUTLYING ISLANDS. TSING YI. 300 m, 978.
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Zhang Li: Hong Kong Bryophytes 45
Habitat and Substrate: On rocks in forest.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hunan,
Jiangxi, Yunnan). India, Nepal, Sri Lanka, Vietnam, Thailand, Cambodia, Malaysia, Singapore,
Borneo, Philippines, Borneo, Indonesia, Papua New Guinea, Australia, Vanuatu (Gao and Crosby,
1999; Higuchi, 1996; Mohamed and Tan, 1988; Rao et al., 1997; Streimann and Curnow, 1989; Tan
and Iwatsuki, 1991; Touw, 1978; Yamaguchi, 1993).
Leucobryum boninense Sull. & Lesq. (L. scaberulum Card. In Ren. & Card., L. scaberulum var.
divaricatum Dix.) – Gao and Crosby (1999)
Localities: NEW TERRITORIES. CASTLE PEAK. 385 m (NE), 1392. – HUNG FA LENG. Muk
Min Tau Stream. 395 m (E), 503. San Kwai Tin Stream. 370 m (N), 1152. – MA ON SHAN. Tai Tung
Wo Liu Stream. 170 m (NE), 1576. Tiu Shau Ngam. 115 m (NW), 246; 215 m (NW), 738 (Confirmed
by T. Yamaguchi, HKU, HIRO, IBSC, KFBG). – PAT SIN LENG. Pin Fung Shan. 480 m (SW), 395.
Wang Shan Keuk Sheung Tsuen. 445 m (NW), 1172. – SAI KUNG. Kap Man Hang (Left Stream). 70
m (NE), 1227. Nam She Tsim. 445 m (N), 1937. Tai Long. 80 m (S), 1388. – TAI LAM. Hung Shui
Shan. 415 m (N), 418. Kiu Keng Shan. 230 m (NW), 343. – TAI MO SHAN. Shek Kong Stream. 770
m (NW), 2291. Tai Shing Stream. 600 m (SE), 21(9); 900 m (NE), 8(17). HONG KONG IS.
D’AGUILAR PEAK. 305 m (E), 173. – VICTORIA PEAK. 540 m, 1741. LANTAU IS. LANTAU
PEAK. 400 m (S), 1665; 615 m (S), 1689. – LIN FA SHAN. 190 m (NE), 599; 365 m (NE), 616; 410
m (NE), 626; 620 m (E), 1456 (Det. By T. Yamaguchi, HKU, HIRO, IBSC, KFBG). – SHUI LO CHO.
205 m (N), 556. – SUNSET PEAK. San She Wan Stream. 500 m (S), 34(2). Wong Lung Hang (East
Lung Stream). 700 m (N), 38(2). Wong Lung Hang (Main Streams). 305 m (N), 2101.
Habitat and Substrate: Twenty-five of 28 specimens were collected in forest, 2 in grassland, and 1 in
shrubland. 22 were found on rocks, 4 on soil, 1 on a twig, and 1 on a root and a trunk.
Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hunan, Sichuan,
Taiwan, Zhejiang). Japan, Philippines (Gao and Crosby, 1999; Redfearn et al., 1996; Tan, Lubos and
Schwarz, 2000).
Leucobryum bowringii Mitt. – Gao and Crosby (1999).
Localities: NEW TERRITORIES. CASTLE PEAK. 300 m (NE), 1389. – GRASS HILL. Fo Tan
Stream. 640 m (SE), 190. – HUNG FA LENG. Muk Min Tau Stream. 60 m (E), 488. San Kwai Tin
Stream. 380 m (N), 1150. – KAI KUNG LENG. Ksiu Keng Pang Uk Stream. 420 m (NE), 1869. –
LAM TSUEN. She Shan Tsuen Feng Shui Wood. 110 m, 1362. – PAT SIN LENG. Pin Fung Shan. 470
m (SW), 389; 475 m (SW), 387. Wang Shan Keuk Sheung Tsuen. 420 m (NW), 1170. – SAI KUNG.
Kap Man Hang (Left Stream). 40 m (NE), 1217. Nam She Tsim. 445 m (N), 1938. – TAI LAM. Kiu
Keng Shan. 415 m (NW), 353. – TAI MO SHAN. Ng Tung Chai Falls. 395 m (N), 4(14); 470 m (NW),
1411; 500 m (N), 3(8); 600 m (N), 2(15); 700 m (N), 1(9), 1(19). Tai Shing Stream. 200 m (SE), 17(4);
300 m (SE), 18(8); 400 m (SE), 19(6); 500 m (SE), 20(1); 600 m (SE), 21(2); 700 m (SE), 6(5), 6(24);
830 m (NE), 18; 900 m (NE), 8(12), 8(21) (Confirmed by T. Yamaguchi, HKU, HIRO, IBSC, KFBG).
Water-pumping House Stream. 700 m (E), S6, S9, S17, S59. – TAI TO YAN. San Lung Wai Stream.
315 m (W), 760; 335 m (W), 757. – WU KAU TANG. Fan Kei Tok Summit. 355 m, 1593. HONG
KONG IS. HIGH WEST. 460 m (N), 1737. LANTAU IS. LANTAU PEAK. 685 m (N), 1330; 735 m
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46 Memoirs of the Hong Kong Natural History Society 26, 2003
(S), 1703. – LIN FA SHAN. 470 m (NE), 631. – SUNSET PEAK. San She Wan Stream. 500 m (S),
34(3), 34(12); 600 m (S), 26(4); 700 m (SE), 27(1). Wong Lung Hang (East Lung Stream). 100 m (N),
39(6); 400 m (N), 35(1); 500 m (N), 36(8); 600 m (N), 37(9); 615 m (N), 1284; 700 m (N), 38(20);
715 m (N), 1288; 800 m (N), 42(2). Wong Lung Hang (Main Streams). 545 m (N), 70; 645 m (N),
2122. OUTLYING ISLANDS. KAT O. 10 m, 2531. – KAU SAI CHAU. Tai Leng Stream. 130 m
(NW), 275.
Habitat and Substrate: Fifty-four of 56 specimens were collected in forest, 1 by a waterfall, and 1 on a
summit. 31 were found on rocks, 7 on trunks, 4 on rocks and trunks, 4 on soil, 3 on rocks and tree base,
2 on tree bases, 2 on rocks and lianas, 1 on a rotten wood, 1 on rocks and on soil, and 1 on a twig.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hubei,
Hunan, Jiangxi, Shandong, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Japan, Himalaya, India, Sri
Lanka, Laos, Vietnam, Thailand, Cambodia, Malaysia, Singapore, Philippines, Borneo, Indonesia,
Papua New Guinea, Solomon Islands, Vanuatu, Mexico, Costa Rica, Jamica (Gao and Crosby, 1999;
Higuchi, 1996; Mohamed and Tan, 1988; Redfearn et al., 1996; Sharp, Crum and Eckel, 1994; Tan and
Iwatsuki, 1991, 1993; Yamaguchi, 1993).
Leucobryum humillimum Card. – Gao and Crosby (1999)
Locality: NEW TERRITORIES. TAI MO SHAN. Ng Tung Chai Falls. 675 m (N), 1236 (Det. By T.
Yamaguchi, HKU, HIRO, IBSC, KFBG).
Habitat and Substrate: On trunks in forest.
Distribution: China (Anhui, Guangdong, Hong Kong, Jiangsu, Taiwan). Korea, Japan, Himalaya, India,
Sri Lanka (Gao and Crosby, 1999; Matsui, 1995; Redfearn et al., 1996; Yamaguchi, 1993).
Leucobryum javense (Brid.) Mitt.
Localities: NEW TERRITORIES. TAI LAM. Lin Fa Shan. 235 m (SW), 662. – SHEUNG WO
HANG FENG SHUI WOOD. 135 m (NW), 1077 (Det. By T. Yamaguchi, HKU, HIRO, IBSC,
KFBG). HONG KONG IS. MOUNT BULTER. 310 m (N), 1723. LANTAU IS. SUNSET PEAK.
Wong Lung Hang (Main Streams). 625 m (N), 51.
Habitat and Substrate: Three of 4 specimens were collected in forest, and 1 in grassland. All were
found on rocks.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hunan,
Jiangxi, Taiwan, Yunnan, Zhejiang). Japan, Himalaya, India, Nepal, Sri Lanka, Myanmar, Laos,
Vietnam, Thailand, Cambodia, Malaysia, Singapore, Philippines, Borneo, Indonesia, Papua New
Guinea (Gangulee, 1969-1980; Gao and Crosby, 1999; Mohamed and Tan, 1988; Tan and Iwatsuki,
1991, 1993; Yamaguchi, 1993).
Leucobryum juniperoideum (Brid.) C. Muell. (L. neilgherrense C. Muell.) – Gao and Crosby (1999)
Localities: NEW TERRITORIES. HUNG FA LENG. Muk Min Tau Stream. 375 m (E), 502. San
Kwai Tin Stream. 350 m (N), 1154. – PLOVER COVE RESERVOIR. Ngor Kai Teng. 215 m, 128. –
SAI KUNG. Shek Uk Shan. 460 m (NW), 308. – TAI LAM. Hung Shui Shan. 415 m (N), 420. Kat
Hing Bridge to Tsing Fai Tong. 120 m (NW), 1055. Lin Fa Shan. 130 m (SW), 656. – TAI MO SHAN.
Ng Tung Chai Falls. 200 m (N), 16(1). Tai Shing Stream. 800 m (NE), 7(4); 900 m (NE), 1122; 900 m
46
Zhang Li: Hong Kong Bryophytes 47
(NE), 8(3). Water-pumping House Stream. 700 m (E), S14. HONG KONG IS. D’AGUILAR PEAK.
10 m (W), 1257; 160 m (W), 160; 220 m (E), 522, 524; 225 m (E), 538; 235 m (W), 164. – VICTORIA
PEAK (Gao and Crosby 1999). LANTAU IS. LIN FA SHAN. 630 m (NE), 643. – SUNSET PEAK.
San She Wan Stream. 400 m (S), 33(14). Wong Lung Hang (East Lung Stream). 320 m (N), 41(15);
800 m (N), 42(26). OUTLYING ISLANDS. TSING YI. 285 m, 980. – FU WONG CHAU. 10 m,
2516. – MA SHI CHAU. 15 m, 380, 383. – WU CHAU. 5 m, 371.
Habitat and Substrate: All specimens were collected in forest. 20 were found on rocks, 4 on trunks, 2
on tree bases, and 1 on soil.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Hainan, Hong Kong, Hubei, Hunan, Jiangsu,
Jiangxi, Shanghai, Sichuan, Taiwan, Xinjiang, Xizang, Yunnan, Zhejiang). Korea, Japan, Himalaya,
India, Sri Lanka, Laos, Myanmar, Vietnam, Cambodia, Thailand, Malaysia, Philippines, Borneo,
Indonesia, Papua New Guinea, Macronesia, Russian Caucasus, Turkey, Southern Europe, Madagascar,
Brazil (Churchill, 1998; Gao and Crosby, 1999; Li, Zeng and Su, 1995; Mohamed and Tan, 1988;
Redfearn et al., 1996; Tan and Iwatsuki, 1991, 1993; Yamaguchi, 1993).
Leucobryum scabrum Sande Lac.
Localities: NEW TERRITORIES. CASTLE PEAK. 350 m (NE), 1394. – SAI KUNG. Nam Shek
Tsim. 405 m (N), 1945. Shek Uk Shan Summit. 480 m, 303. – TAI LAM. Kat Hing Bridge to Tsing
Fai Tong. 110 m (NW), 1052. – TAI MO SHAN. Tai Shing Stream. 800 m (NE), 7(3); 835 m (NE),
1423. Water-pumping House Stream. 700 m (E), S44. HONG KONG IS. HIGH WEST. 470 m (N),
1736. LANTAU IS. LIN FA SHAN. 570 m (E), 1450; 615 m (NE), 642; 620 m (E), 1455. – SUNSET
PEAK. Wong Lung Hang (East Lung Stream). 315 m (N), 1268; 320 m (N), 41(2), 41(7); 400 m (N),
35(14); 600 m (N), 37(17); 700 m (N), 38(19); 800 m (N), 42(9).
Habitat and Substrate: There were 17 specimens examined, of which 15 were collected in forest, 1 in
grassland, and 1 unknown. 14 were found on rocks, 2 on soil, 1 on a tree base, and 1 on a trunk and on
rocks.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Jiangxi,
Shanghai, Sichuan, Taiwan, Yunnan, Zhejiang). Japan, Thailand, Malaysia (Gao and Crosby, 1999;
Redfearn et al., 1996; Tan and Iwatsuki, 1991).
Leucophanes glaucum (Schwaegr.) Mitt. (L. albescens C. Muell.)
Species new to mainland China and genus and species new to Hong Kong.
Localities: NEW TERRITORIES. PAT SIN LENG. Sze Tau Leng. 240 m (NW), 990. – SHEUNG
WO HANG FENG SHUI WOOD. 170 m (NW), 1762, 1763. – TAI MO SHAN. Tai Yuen Stream. 265
m (S), 1843. – WU KAU TANG. Wu Kau Tang Feng Shui Wood. 75 m, 1752. HONG KONG IS.
D’AGUILAR PEAK. 220 m (W), 161; 220 (E), 525; 290 m (E), 177. – MOUNT NICHOLSON. 260 m
(N), 2392. LANTAU IS. SUNSET PEAK. San Shek Wan Stream. 375 m (S), 2043. OUTLYING
ISLANDS. MA SHI CHAU. 15 m, 381, 382. – SIU NIM CHAU. 10 m, 2508.
Habitat and Substrate: All 13 specimens were collected in forest. 7 were found on trunks, and 6 on
rocks.
Distribution: China (Hainan, Hong Kong, Taiwan). Thailand, Cambodia, Malaysia, Singapore,
Philippines, Borneo, Papua New Guinea, Australia, Polynesia, Society Islands, Vanuatu, Africa (He,
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48 Memoirs of the Hong Kong Natural History Society 26, 2003
2000; Higuchi, 1996; Mohamed and Tan, 1988; O’Shea, 1999; Tan and Iwatsuki, 1991, 1993; Touw,
1978; Whittier, 1976).
Octoblepharum albidum (L.) Hedw. – Gao and Crosby (1999).
Localities: NEW TERRITORIES. HO CHUNG. 10 m, 1510. – MA ON SHAN. Tai Tung Wo Liu
Stream. 150 m (NE), 1564; 265 m (NE), 1579. Tiu Shau Ngam. 195 m (NW), 260. – TAI MO SHAN.
Chuen Lung Stream. 400 m (SW), 2421. Tai Yuen Stream. 235 m (S), 1841; 330 m (S), 1846. Yin
Ngam Stream. 365 m (NE), 1890. – TAI PO KAU. 155 m (NE), 2328. HONG KONG IS. VICTORIA
PEAK. 415 m, 1506. LANTAU IS. LANTAU PEAK. 615 m (S), 1688. – SUNSET PEAK. San Shek
Wan Stream. 290 m (S), 2032.
Habitat and Substrate: Eleven of 12 specimens examined were collected in forest, and 1 by a road. 8
were found on trunks, 3 on rocks, and 1 on a twig.
Distribution: China (Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Taiwan, Yunnan). India,
Vietnam, Myanmar, Thailand, Philippines, Malesia, Borneo, Indonesia, Australia, Fiji, Vanuatu,
Hawaii, North America, Mexico, Central and South America, Brazil (Gao and Crosby, 1999; Higuchi,
1996; Li and Piippo, 1994; Redfearn et al., 1996; Sharp, Crum and Eckel, 1994; Touw, 1978; Yano,
1995).
METEORIACEAE
Aerobryidium wallichii (Brid.) Fleisch.
Locality: LANTAU IS. LANTAU PEAK. 580 m (N), 2451.
Habitat and Substrate: On rocks in forest.
Distribution: China (Fujian, Guangdong, Guizhou, Hainan, Hong Kong, Hubei, Jiangxi, Taiwan,
Yunnan, Zhejiang). India, Sri Lanka, Nepal, Vietnam, Thailand, Cambodia, Malaysia, Philippines,
Borneo, Indonesia, Papua New Guinea, Australia, Polynesia (Gangulee, 1969-1980; He, 2000;
Noguchi, 1976; Redfearn et al.,1996; Streimann and Curnow, 1989; Tan and Iwatsuki,1993; Touw,
1978).
Aerobryopsis praisii (Card.) Broth.
Localities: NEW TERRITORIES. TAI LO SHAN. Kwun Yam Shan. 160 m (N), 950; 210 m (N),
957. – TAI MO SHAN. Kadoorie Farm. 290 m (NW), 827. Ng Tung Chai Falls. 205 m (N), 1766. –
TAI PO KAU. 180 m (NE), 2333.
Habitat and Substrate: All 5 specimens were collected in forest. 3 were found on rocks, and 2 on
trunks.
Distribution: China (Fujian, Guangdong, Hong Kong, Hunan, Taiwan). Japan, Philippines (Koponen et
al., 2000; Noguchi,1989; Redfearn et al., 1996; Tan and Iwatsuki, 1991).
Aerobryopsis subdivergens (Broth.) Broth.
Species new to Hong Kong.
Localities: NEW TERRITORIES. TAI MO SHAN. Ng Tung Chai Falls. 700 m (N), 1243. Yin Ngam
Stream. 495 m (NE), 1899. LANTAU IS. LANTAU PEAK. 635 m (N), 1327. – SUNSET PEAK.
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Zhang Li: Hong Kong Bryophytes 49
Wong Lung Hang (Main Streams). 605 m (N), 53; 580 m (N), 2118; 555 m (N), 2114 (with Meteorium
subpolytrichum); 455 m (NW), 2407.
Habitat and Substrate: All 7 specimens were collected in forest. 6 were found on rocks, and 1 on a
trunk.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hubei,
Hunan, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Japan, Vietnam, Philippines, Hawaii (Koponen
et al., 2000; Noguchi, 1989; Redfearn et al.,1996).
Barbella convolvens (Mitt.) Broth.
Species new to China.
Locality: NEW TERRITORIES. TAI MO SHAN. Tai Shing Stream. 860 m (NE), 1137 (Confirmed
by P. C. Wu, HKU, IBSC, KFBG, PE).
Habitat and Substrate: On a rotten twig in forest.
Distribution: China (Hong Kong). India, Sri Lanka, Laos, Thailand, Philippines, Borneo, Indonesia
(Noguchi, 1976; Tan and Iwatsuki, 1991, 1993).
Barbella flagellifera (Card.) Nog.
Localities: NEW TERRITORIES. TAI MO SHAN. Shek Kong Stream. 825 m (NW), 2013. Tai
Shing Stream. 900 m (NE), 1142; 920 m (NE), 1110; 830 m (NE), 1419; 895 m (NE), 1131, 900 m
(NE), 1118; 920 m (NE), 1437; 900 m (NE), 8(2). Water-pumping House Stream. 770 m (E), 1968.
LANTAU IS. LANTAU PEAK. 675 m (N), 1331; 680 m (S), 1694; 730 m (S), 1708 (with
Clastobryopsis planula), 1709.
Habitat and Substrate: Twelve of 13 specimens were collected in forest, and 1 in shrubland. 7 were
found on twigs, 3 on trunks, 2 on rotten twigs, and 1 on leaves.
Distribution: China (Anhui, Fujian, Guangxi, Guizhou, Hainan, Hong Kong, Hubei, Sichuan, Taiwan,
Xizang, Yunnan, Zhejiang). Japan, India, Sri Lanka, Myanmar, Vietnam, Thailand, Philippines,
Borneo, Indonesia (Noguchi, 1976; Redfearn et al., 1996; Tan and Iwatsuki, 1993).
Duthiella wallichii (Mitt.) C. Muell.
Genus and species new to Hong Kong.
Locality: NEW TERRITORIES. TAI MO SHAN. Shek Kong Stream. 440 m (NW), 1976.
Habitat and Substrate: On rocks in forest.
Distribution: China (Guangxi, Guizhou, Hainan, Hong Kong, Sichuan, Taiwan, Xizang, Yunnan,
Zhejiang). Japan, India, Nepal, Vietnam, Thailand, Malaysia, Philippines, Indonesia (He, 2000; Li,
Zeng and Su, 1995; Mohamed and Tan, 1988; Noguchi, 1991; Tan and Iwatsuki, 1991, 1993).
Floribundaria floribunda (Dozy & Molk.) Fleisch.
Genus and species new to Hong Kong.
Localities: NEW TERRITORIES. TAI MO SHAN. Ng Tung Chai Falls. 200 m (N), 1767, 1768,
16(11), 16(12).
Habitat and Substrate: All 4 specimens were collected in forest. 3 were found on trunks, and 1 on bark.
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50 Memoirs of the Hong Kong Natural History Society 26, 2003
Distribution: China (Fujian, Guangxi, Guizhou, Hong Kong, Taiwan, Xizang, Yunnan, Zhejiang).
Japan, India, Nepal, Bhutan, Myanmar, Vietnam, Thailand, Cambodia, Malaysia, Philippines, Borneo,
Indonesia, Papua New Guinea, Australia, Vanuatu, Hawaii, Africa, Madagascar (Higuchi, 1996;
Noguchi, 1976; Redfearn et al., 1996; Tan and Iwatsuki, 1993; Touw, 1978).
Meteorium subpolytrichum (Besch.) Broth.
Localities: NEW TERRITORIES. MA ON SHAN. Ngong Ping. 345 m, 125. – PAT SIN LENG.
Hsien Ku Fung. 350 m (E), 1010; 380 m (NE), 1015; 390 m (NE), 1017; 410 m (NE), 1022. – TAI MO
SHAN. Chuen Lung Stream. 625 m (SW), 2433; 660 m (SW), 2431. – TAI TO YAN. Lam Tsuen
Stream. 505 m (E), 726. LANTAU IS. LANTAU PEAK. 430 m (N), 2445; 520 m (S), 1674; 575 m
(S), 1684. – SUNSET PEAK. Wong Lung Hang (East Lung Stream). 615 m (N), 2305. Wong Lung
Hang (Main Streams). 455 m (NW), 2408; 580 m (N), 2115; 580 m (S), 1685; 595 m (N), 65; 605 m
(N), 90; 610 m (N), 54.
Habitat and Substrate: All specimens were collected in forest. 14 were found on rocks, 3 on trunks, and
1 on a rotten twig.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hubei,
Jiangsu, Jiangxi, Shaanxi, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Japan, Nepal, Bhutan,
Philippines (Fang et al., 1998; Noguchi, 1976; Redfearn et al., 1996; Tan and Iwatsuki, 1993; Zhang,
1996).
Pseudobarbella angustifolia Nog. – So and Zhu (1996c).
Locality: HONG KONG (Locality unknown). 400 m (So and Zhu, 1996c).
Habitat and Substrate: On bark (So and Zhu, 1996c).
Distribution: Endemic to China (Anhui, Guangxi, Guizhou, Hong Kong, Taiwan, Xizang, Yunnan,
Zhejiang) (Noguchi, 1976; Redfearn et al., 1996).
Pseudobarbella attenuata (Thwait. & Mitt.) Nog.
Localities: NEW TERRITORIES. GRASS HILL. Fo Tan Stream. 450 m (SE), 205. – HUNG FA
LENG. San Kwai Tin Stream. 395 m (N), 1148. – KAU LUNG HANG SHAN. 60 m (NW), 1537. –
KOWLOON PEAK. Ho Chung Stream. 130 m, 1518. Pak Fa Lam. 160 m (E), 1533. – MA ON SHAN.
Baffalo Hill. 240 m (NE), 2224. – PAT SIN LENG. Hsien Ku Fung. 415 m (NE), 1023. Wang Shan
Keuk Sheung Tsuen. 295 m (NW), 1161. – SAI KUNG. Shek Uk Shan. 460 m (NW), 311. – TAI LO
SHAN. Kwun Yam Shan. 100 m (N), 934; 135 m (N), 944. – TAI MO SHAN. 800 m, Siu 16. Chuen
Lung Stream. 740 m (SW), 2440. Ng Tung Chai Falls. 200 m (N), 16(2); 395 m (N), 4(3); 600 m (N),
2(5), 2(29); 610 m (N), 1245. Shek Kong Stream. 320 m (NW), 1970; 635 m (NW), 1990; 785 m
(NW), 2285. Shing Mun Reservoir. 195 m, 747. Tai Shing Stream. 905 m (NE), 1117. Yin Ngam
Stream. 300 m (NE), 1885; 525 m (NE), 1900. – TAI PO KAU. 175 m (NE), 2330. – TAI TO YAN.
Lam Tsuen Stream. 80 m (E), 695. San Lung Wai Stream. 235 m (W), 768; 275 m (W), 765. – WU
KAU TANG. Fan Kei Tok Feng Shui Wood. 345 m (N), 1607. HONG KONG IS. D’AGUILAR
PEAK. 215 m (E), 534; 220 m (E), 532; 255 m (E), 184; 290 m (E), 175. LANTAU IS. LANTAU
PEAK. 530 m (S), 1673; 645 m (S), 1691. – LIN FA SHAN. 375 m (NE), 617. – SUNSET PEAK.
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Zhang Li: Hong Kong Bryophytes 51
Wong Lung Hang (East Lung Stream). 100 m (N), 39(3); 400 m (N), 35(6). Wong Lung Hang (Main
Streams). 590 m (N), 84.
Habitat and Substrate: Thirty-nine of 40 specimens were collected in forest, and 1 by a stream. 25 were
found on rocks, 9 on trunks, 3 on twigs, 1 on a rotten twig, 1 on rotten wood, and 1 on a tree base.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hubei,
Jiangsu, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Japan, Sri Lanka, Vietnam, Thailand, Malaysia,
Philippines, Borneo, Indonesia (Jia and Wang, 1993; Noguchi, 1976; Redfearn et al., 1996).
Pseudobarbella leveri (Ren. & Card.) Nog.
Species new to Hong Kong.
Locality: LANTAU IS. LANTAU PEAK. 760 m (N), 2462.
Habitat and Substrate: On rocks in forest.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Jiangxi,
Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Japan, India, Thailand (Gangulee, 1969-1980; He, 2000;
Li, Zeng and Su, 1995; Noguchi, 1989; Redfearn et al., 1996).
MNIACEAE
Plagiomnium cuspidatum (Hedw.) T. Kop.
Localities: NEW TERRITORIES. HUNG FA LENG. San Kwai Tin Stream. 395 m (N), 1147. – KAI
KUNG LENG. Ha Che Stream. 85 m (S), 1862. – KAU LUNG HANG SHAN. 30 m (NW), 1535. –
TAI LO SHAN. Kwun Yam Shan. 100 m (N), 933. – TAI MO SHAN. Shing Mun Reservoir. 190 m,
743. Tai Shing Stream. 910 m (NE), 1115; 920 m (NE), 1108. Tai Yuen Stream. 325 m (S), 1845. –
TAI TO YAN. Lam Tsuen Stream. 85 m (E), 683.
Habitat and Substrate: Eight of 9 specimens were collected in forest, and 1 in bamboo forest. 7 were
found on rocks, and 2 on soil.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hebei, Heilongjiang, Henan, Hong
Kong, Hubei, Inner Mongolia, Jiangsu, Jilin, Liaoning, Shaanxi, Shandong, Shanghai, Shanxi, Sichuan,
Taiwan, Xinjiang, Yunnan, Zhejiang). Korea, Japan, Mongolia, Russian Far East and Siberia, West
Asia, Thailand, Europe, Africa, North America, Mexico, Cuba (Bai, 1996; He, 2000; Redfearn et al.,
1996; Sharp, Crum and Eckel, 1994; Zhao and Cao, 1998).
Plagiomnium succulentum (Mitt.) T. Kop.
Localities: NEW TERRITORIES. GRASS HILL. Fo Tan Stream. 295 m (SE), 221. – HO PUI
RESERVOIR. 310 m, 818. – HUNG FA LENG. Muk Min Tau Stream. 70 m (E), 491. – KAU LUNG
HANG SHAN. 60 m (NW), 1540. – KOWLOON PEAK. Ho Chung Stream. 200 m, 1521. – TAI
LAM. Lin Fa Shan. 335 m (SW), 674. – TAI LO SHAN. Kwun Yam Shan. 185 m (N), 953. – TAI MO
SHAN. Guard Post. 700 m (W), Choi 15. Ng Tung Chai Falls. 400 m (N), Choi 14; 600 m (N), 2(26).
Shek Kong Stream. 610 m (NW), 1988; 720 m (NW), 1997. Shing Mun Reservoir. 180 m, 744. Tai
Shing Stream. 900 m (NE), 8(1). – TAI PO KAU. 190 m (NE), 2334. – TAI TO YAN. Lam Tsuen
Stream. 120 m (E), 706. – WU KAU TANG. So Lo Pun Back Stream. 125 m (NE), 1625. LANTAU
IS. SUNSET PEAK. San Shek Wan Stream. 350 m (S), 2040. Wong Lung Hang (East Lung Stream).
700 m (N), 38(11).
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52 Memoirs of the Hong Kong Natural History Society 26, 2003
Habitat and Substrate: All specimens were collected in forest. 17 were found on rocks, 1 on a trunk,
and 1 on soil.
Distribution: China (Anhui, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Heilongjiang, Henan,
Hong Kong, Hubei, Jiangsu, Jiangxi, Jilin, Liaoning, Shaanxi, Shandong, Shanxi, Sichuan, Taiwan,
Xizang, Yunnan, Zhejiang). Korea, Japan, India, Nepal, Bhutan, Myanmar, Vietnam, Thailand,
Cambodia, Malaysia, Singapore, Philippines, Borneo, Indonesia, Papua New Guinea, Vanuatu (Eddy,
1996; Fang et al., 1998; Higuchi, 1996; Koponen et al., 2000; Mohamed and Tan, 1988; Redfearn et
al., 1996; Tan and Iwatsuki, 1993; Touw, 1978; Zhao and Cao, 1998).
Plagiomnium vesicatum (Besch.) T. Kop.
Localities: NEW TERRITORIES. TAI MO SHAN. Chuen Lung Stream. 355 m (SW), 2420. Ng
Tung Chai Falls. 700 m (N), 1(23).
Habitat and Substrate: On rocks in forest.
Distribution: China (Anhui, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hebei, Heilongjiang,
Henan, Hong Kong, Hubei, Inner Mongolia, Jiangsu, Jiangxi, Jilin, Liaoning, Shaanxi, Shanxi,
Shandong, Sichuan, Taiwan, Xinjiang, Yunnan, Zhejiang). Korea, Japan, Russian Far East and Siberia
(Bai, 1996; Jia and Wang, 1993; Redfearn et al., 1996; Zhao and Cao, 1998).
Trachycystis microphylla (Dozy & Molk.) Lindb.
Genus and species new to Hong Kong.
Localities: NEW TERRITORIES. TAI MO SHAN. Tai Shing Stream. 860 m (NE), 1354; 870 m
(NE), 1134a.
Habitat and Substrate: On soil and trunks in forest.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Heilongjiang, Henan, Hong Kong,
Hubei, Hunan, Jiangsu, Jiangxi, Jilin, Liaoning, Shaanxi, Shandong, Shanghai, Sichuan, Taiwan,
Xinjiang, Yunnan, Zhejiang). Korea, Japan, Russian Far East and Siberia (Fang et al., 1998; Noguchi,
1989; Redfearn et al., 1996; Zhao and Cao, 1998).
MYURIACEAE
Oedicladium fragile Card.
Localities: NEW TERRITORIES. MA ON SHAN. Ma On Shan Tsuen Stream. 335 m (SW), 2153;
360 m (SW), 2157; 620 m (SW), 2177. Tiu Shau Ngam. 470 m (NW), 270. – TAI MO SHAN. Chuen
Lung Stream. 610 m (SW), 2430. Shek Kong Stream. 770 m (NW), 2003, 2288. Tai Shing Stream. 600
m (SE), 21(1); 780 m (NE), 23, 24; 795 m (NE), 1432; 865 m (NE), 1136. Water-pumping House
Stream. 700 m (E), S34. – WU KAU TANG. Fan Kei Tok Summit. 345 m, 1600. LANTAU IS.
LANTAU PEAK. 305 m (N), 1304; 660 m (S), 1686; 710 m (S), 1699. – LIN FA SHAN. 595 m (E),
1453. – SUNSET PEAK. Wong Lung Hang (East Lung Stream). 200 m (N), 40(15); 320 m (N),
41(12); 335 m (N), 2241; 600 m (N), 37(12); 720 m (N), 1291; 800 m (N), 42(22). Wong Lung Hang
(Main Streams). 305 m (N), 2100; 390 m (NW), 2402.
Habitat and Substrate: All specimens were collected in forest. 25 were found on rocks, and 1 on a tree
base.
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Zhang Li: Hong Kong Bryophytes 53
Distribution: China (Guangdong, Guizhou, Hainan, Hong Kong, Taiwan). Japan, Vietnam, Thailand,
Philippines (He, 2000; Noguchi, 1994; Redfearn et al., 1996; Tan and Iwatsuki, 1991).
Oedicladium rufescens (Reinw. & Hornsch.) Mitt.
Species new to Hong Kong.
Localities: NEW TERRITORIES. TAI MO SHAN. Shek Kong Stream. 790 m (NW), 2284.
LANTAU IS. LANTAU PEAK. 735 m (N), 1339; 860 m (N), 2477.
Habitat and Substrate: Two of 3 studied specimens were collected in forest, and 1 by a waterfall. 2
were found on rocks, and 1 on rotten wood.
Distribution: China (Anhui, Guangdong, Guangxi, Hong Kong, Zhejiang). Japan, Malaysia, Singapore,
Borneo (Jia and Wang, 1993; Mohamed and Tan, 1988; Noguchi, 1994; Redfearn et al., 1996; Touw,
1978; Wu et al., 1992).
Oedicladium serricuspe (Broth.) Nog. & Iwats. – So and Zhu (1996c).
Localities: NEW TERRITORIES. MA ON SHAN. Tiu Shau Ngam. 135 m (NW), 250. – TAI MO
SHAN. Chuen Lung Stream. 715 m (SW), 2434. Ng Tung Chai Falls. 700 m (N), 1244. HONG
KONG IS. HIGH WEST. 480 m (N), 1734. HONG KONG (Locality unknown). 800 m (So and Zhu,
1996c).
Habitat and Substrate: On rocks in forest.
Distribution: China (Hong Kong, Taiwan). Japan (Iwatsuki, 1979).
Palisadula chrysophylla (Card.) Toy
Genus and species new to Hong Kong.
Localities: NEW TERRITORIES. SHA TAU KOK. Muk Min Tau Feng Shui Wood. 10 m, 2134. –
TAI MO SHAN. Chuen Lung Stream. 810 m (SW), 2444. Shek Kong Stream. 750 m (NW), 2301; 755
m (NW), 2294; 810 m (NW), 2283. Tai Shing Stream. 600 m (SE), 21(18), 21(25); 745 m (SE), 26;
780 m (NE), 1429; 800 m (NE), 7(11); 865 m (NE), 15. Water-pumping House Stream. 700 m (E),
S35. LANTAU IS. LANTAU PEAK. 630 m (N), 1325; 700 m (S), 1697; 715 m (S), 1706. – SUNSET
PEAK. San She Wan Stream. 700 m (SE), 27(16). Wong Lung Hang (Main Streams). 700 m (N), 2127.
Habitat and Substrate: Sixteen of 17 specimens were collected in forest, and 1 unknown. 7 were found
on rocks, 2 on twigs, 2 on rotten wood, 2 on trunks, 1 on a trunk and on rocks, 1 on a trunk and rotten
wood, 1 on a rotten twig, and 1 on a tree base.
Distribution: China (Guangdong, Hong Kong, Taiwan). Japan (Iwatsuki, 1979; Li and Piippo, 1994).
NECKERACEAE
Himantocladium plumula (Nees) Fleisch.
Species new to Hong Kong. It was reported as H. cyclophyllum (C. Muell.) Fleisch. Previously (So and
Yip, 1995b; Zhang and Lin, 1997).
Localities: NEW TERRITORIES. MA ON SHAN. Baffalo Hill. 235 m (NE), 2220. – TAI MO
SHAN. Ng Tung Chai Falls. 180 m (N), 1772; 395 m (N), 4(13); 600 m (N), 2(28); 645 m (N), 1232;
700 m (N), 1(13). LANTAU IS. SUNSET PEAK. Wong Lung Hang (East Lung Stream). 385 m (N),
2242; 405 m (N); 1275; 415 m (N), 1274.
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54 Memoirs of the Hong Kong Natural History Society 26, 2003
Habitat and Substrate: All 9 specimens were collected in forest. 8 were found on rocks, and 1 on a tree
base.
Distribution: China (Guizhou, Hainan, Hong Kong, Taiwan). Japan, India, Bangladesh, Myanmar,
Laos, Vietnam, Thailand, Cambodia, Malaysia, Philippines, Borneo, Indonesia, Papua New Guinea,
Australia, New Caledonia, Vanuata, Fiji, Society Islands (Enroth, 1992; Higuchi, 1996; Redfearn et al.,
1996; Tan and Iwatsuki, 1991, 1993; Touw, 1978).
Homalia trichomanoides (Hedw.) B.S.G.
Localities: NEW TERRITORIES. KAI KUNG LENG. Ha Che Stream. 40 m (S), 1857. – KAU
LUNG HANG SHAN. 60 m (NW), 1538. – PAT SIN LENG. Sze Tau Leng. 65 m (NW), 987. – TAI
LAM. Kat Hing Bridge to Tsing Fai Tong. 140 m (NW), 1064. Lin Fa Shan. 280 m (SW), 668; 305 m
(SW), 670. – TAI MO SHAN. Ng Tung Chai Falls. 310 m (NW), 1405. – TAI TO YAN. Lam Tsuen
Stream. 70 m (E), 693. HONG KONG IS. D’AGUILAR PEAK. 160 m (E), 188; 210 m (E), 536; 225
m (E), 529; 275 m (E), 180.
Habitat and Substrate: All specimens were collected on rocks in forest.
Distribution: China (Anhui, Fujian, Gansu, Guangdong, Guizhou, Heilongjiang, Hong Kong, Hunan,
Inner Mongolia, Jiangsu, Jiangxi, Jilin, Liaoning, Shaanxi, Shanghai, Shandong, Shanxi, Sichuan,
Taiwan, Xizang, Zhejiang). Korea, Japan, India, Bhutan, Russia, Europe, North America, Mexico (Bai,
1996; He, 1997; Ignatov and Afonina, 1992; Li, Liao and Huang, 1998; Redfearn et al., 1996; Zhao
and Cao, 1998).
Homaliodendron exiguum (Bosch & Sande Lac.) Fleisch.
Localities: NEW TERRITORIES. GRASS HILL. Fo Tan Stream. 335 m (SE), 214. – HO PUI
RESERVOIR. 305 m, 815. – HUNG FA LENG. Muk Min Tau Stream. 35 m (E), 483. – KAU LUNG
HANG SHAN. 320 m (NW), 1552. – KOWLOON PEAK. Ho Chung Stream. 305 m, 1527. – SAI
KUNG. Shek Uk Shan. 280 m (NW), 323. – TAI LO SHAN. Kwun Yam Shan. 120 m (N), 939; 160 m
(N), 949. – TAI MO SHAN. Ng Tung Chai Falls. 395 m (N), 4(2); 610 m (N), 1246. Water-pumping
House Stream. 700 m (E), S40, S53. – WU KAU TANG. So Lo Pun Back Stream. 70 m (NE), 1622.
HONG KONG IS. D’AGUILAR PEAK. 240 m (W), 169. LANTAU IS. LANTAU PEAK. 520 m (S),
1672.
Habitat and Substrate: All specimens were collected in forest. 12 were found on rocks, 2 on trunks,
and 1 on rotten bark.
Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hunan, Jiangsu,
Taiwan, Xizang, Yunnan, Zhejiang). Japan, India, Nepal, Bhutan, Sri Lanka, Myanmar, Vietnam,
Thailand, Malaysia, Philippines, Borneo, Indonesia, Papua New Guinea, Australia, Fiji, Society
Islands, Africa (Koponen et al., 2000; O’Shea, 1999; Redfearn et al., 1996; Tan and Iwatsuki, 1991,
1993; Touw, 1978; Whittier, 1976).
Homaliodendron flabellatum (Sm.) Fleisch. (H. scapellifolium (Mitt.) Fleisch., H. squarrulosum
Fleisch., fide Enroth (1989)).
Localities: NEW TERRITORIES. TAI LO SHAN. Kwun Yam Shan. 390 m (N), 970. – TAI MO
SHAN. Chuen Lung Stream. 750 m (SW), 2441. Ng Tung Chai Falls. 645 m (N), 1233; 655 m (N),
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Zhang Li: Hong Kong Bryophytes 55
1235; 700 m (N), 1(16). Shek Kong Stream. 515 m (NW), 1982; 615 m (NW), 1989; 635 m (NW),
1991; 675 m (NW), 1994; 785 m (NW), 2286. Tai Shing Stream. 500 m (SE), 20(18); 755 m (NE),
1357; 800 m (NE), 7(1). Water-pumping House Stream. 700 m (E), S5. Yin Ngam Stream. 485 m
(NE), 1897. LANTAU IS. LANTAU PEAK. 475 m (N), 1315; 580 m (N), 2450; 730 m (S), 1704. –
LIN FA SHAN. 465 m (NE), 628. – SUNSET PEAK. Wong Lung Hang (East Lung Stream). 320 m
(N), 41(14); 390 m (N), 1270; 400 m (N), 35(15); 535 m (N), 1279; 550 m (N), 2245; 600 m (N),
37(8); 680 m (N), 2307; 700 m (N), 38(6); 800 m (N), 42(15). Wong Lung Hang (Main Streams). 280
m (NW), 2401; 470 m (N), 2112; 625 m (N), 50.
Habitat and Substrate: Thirty of 31 specimens were collected in forest, and 1 in grassland. 30 were
found on rocks, and 1 on trunks.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hubei,
Hunan, Jiangsu, Jiangxi, Liaoning, Shaanxi, Shanghai, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang).
Japan, India, Nepal, Bhutan, Sri Lanka, Laos, Myanmar, Vietnam, Thailand, Malaysia, Philippines,
Borneo, Indonesia, Papua New Guinea, Australia, New Caledonia, Vanuatu, Hawaii, Africa, Mexico,
Central and South America (Buck, 1998; Fang et al., 1998; Higuchi, 1996; Koponen et al., 2000;
O’Shea, 1999; Redfearn et al., 1996; Tan and Iwatsuki, 1991, 1993; Touw, 1978).
Homaliodendron microdendrom (Mont.) Fleisch.
Locality: HONG KONG (unknown) (So and Yip, 1995b).
Habitat and Substrate: On tree base or moist rock surfaces (So and Yip, 1995b).
Distribution: China (Guangdong, Guizhou, Hainan, Hong Kong, Hubei, Sichuan, Taiwan, Xizang,
Yunnan). Japan, India, Nepal, Bhutan, Laos, Myanmar, Vietnam, Thailand, Cambodia, Philippines,
Borneo, Vanuatu (Gangulee, 1969-1980; He, 2000; Higuchi, 1996; Noguchi, 1989; Redfearn et al.,
1996; Tan and Iwatsuki, 1991, 1993).
Neckeropsis obtusata (Mont.) Fleisch. Ex Broth. – Zhang et al. (1998a).
Localities: NEW TERRITORIES. CASTLE PEAK. 195 m (NE), 1398. – KOWLOON PEAK. Ho
Chung Stream. 300 m (E), 1523. – MA ON SHAN. Baffalo Hill. 235 m (NE), 2221. – TAI MO SHAN.
Kadoorie Farm. 160 m (NW), Lin et al. 715 (IBSC, KFBG); 170 m (NW), Lin et al. 400 (IBSC,
KFBG); 190 m (NW), Lin et al. 721, 722 (IBSC, KFBG). – WU KAU TANG. Mirror Pool Feng Shui
Wood. 50 m, 1748. HONG KONG IS. D’AGUILAR PEAK. 215 m (E), 535. LANTAU IS.
LANTAU PEAK. 340 m (S), 2214. – SUNSET PEAK. San Shek Wan Stream. 210 m (S), 2026.
Habitat and Substrate: Ten of 11 specimens examined were collected in forest, and 1 by a waterfall. All
were found on rocks.
Distribution: China (Guangdong, Guizhou, Hong Kong, Taiwan, Zhejiang). Japan, Vietnam, Hawaii
(Zhang et al., 1998a).
Pinnatella anacamptolepis (C. Muell.) Broth.
Localities: NEW TERRITORIES. GRASS HILL. Fo Tan Stream. 305 m (SE), 218; 315 m (SE), 220;
455 m (SE), 202. – KOWLOON PEAK. Pak Fa Lam. 225 m (E), 1531. – MA ON SHAN. Baffalo Hill.
215 m (NE), 2219. – TAI LO SHAN. Kwun Yam Shan. 430 m (N), 971. – TAI MO SHAN. Chuen
Lung Stream. 740 m (SW), 2442. Ng Tung Chai Falls. 265 m (NW), 1403; 380 m (N), 1351; 600 m
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56 Memoirs of the Hong Kong Natural History Society 26, 2003
(N), 2(13); 630 m (N), 1187; 650 m (N), 1234 (with Thamnobryum plicatulum). Tai Shing Stream. 460
m (SE), 1853. Yin Ngam Stream. 305 m (NE), 1886. HONG KONG IS. MOUNT PARKER. 325 m
(NW), 2388. LANTAU IS. LIN FA SHAN. 265 m (NE), 607; 330 m (NE), 612; 345 m (NE), 615. –
SUNSET PEAK. Wong Lung Hang (Main Streams). 380 m (N), 2105; 400 m (N), 2109; 560 m (N),
80.
Habitat and Substrate: All specimens were collected on rocks in forest.
Distribution: China (Guangdong, Guangxi, Hainan, Hong Kong, Hubei, Taiwan). Japan, ?India, Sri
Lanka, Vietnam, Thailand, Malaysia, Philippines, Borneo, Indonesia, Papua New Guinea (Enroth,
1994; Redfearn et al., 1996; Tan and Iwatsuki, 1991; Touw, 1978; Zhang, 1994).
Thamnobryum plicatulum (Sande Lac.) Iwats.
Localities: NEW TERRITORIES. TAI MO SHAN. Ng Tung Chai Falls. 455 m (N), 1196; 545 m
(N), 1194; 630 m (N), 1192. Shek Kong Stream. 495 m (NW), 1980; 750 m (NW), 2302. HONG
KONG IS. MOUNT PARKER. 335 m (NW), 2386; 375 m (NW), 2390. LANTAU IS. SUNSET
PEAK. San Shek Wan Stream. 395 m (S), 2045. Wong Lung Hang (Main Streams). 500 m (N), 2113;
615 m (N), 55.
Habitat and Substrate: All specimens were collected on rocks in forest.
Distribution: China (Anhui, Guizhou, Hong Kong, Hubei, Sichuan, Taiwan, Yunnan). Korea, Japan,
Russian Far East (Noguchi, 1989; Redfearn et al., 1996).
ORTHOTRICHACEAE
Macromitrium brevituberculatum Dix.
Localities: NEW TERRITORIES. GRASS HILL. Fo Tan Stream. 215 m (SE), 224. – LAM TSUEN.
Tai Om Feng Shui Wood. 110 m, 1909. – MA ON SHAN. Tiu Shau Ngam. 410 m (NW), 117. – PAT
SIN LENG. Nam Chung Stream. 295 m (NE), 1000. – TAI LAM. Kat Hing Bridge to Tsing Fai Tong.
135 m (NW), 1060. – TAI LO SHAN. Kwun Yam Shan. 350 m (N), 966. – TAI MO SHAN. Ng Tung
Chai Falls. 520 m (N), 1346. Shek Kong Stream. 715 m (NW), 1996. Yin Ngam Stream. 190 m (NE),
1876; 310 m (NE), 1884; 335 m (NE), 1887; 375 m (NE), 1891; 495 m (NE), 1898. – TAI PO KAU.
170 m (NE), 2329; 180 m (NE), 2331; 200 m (NE), 2335. HONG KONG IS. D’AGUILAR PEAK.
110 m (W), 140; 230 m (E), 514, 516. – SHAUKIWAN. Ah Nin H.8. (BM). – STANLEY. Near sea
level. Ah Nin H.43A (BM). LANTAU IS. LANTAU PEAK. 300 m (N), 1302. – LIN FA SHAN. 425
m (NE), 627. – PAK KUNG AU. 340 m (S), 2148. – SUNSET PEAK. Wong Lung Hang (East Lung
Stream). 385 m (N), 2304. Wong Lung Hang (Main Streams). 275 m (N), 2091; 395 m (NW), 2403;
420 m (NW), 2405; 550 m (N), 2116; 570 m (N), 86; 605 m (N), 88.
Habitat and Substrate: There were 31 specimens examined, of which 28 were collected in forest, and 3
unknown. 16 were found on trunks, 9 on rocks, 3 on rotten wood, 1 on a tree base, 1 on a twig, and 1
unknown.
Distribution: Endemic to Hong Kong.
Macromitrium ferriei Card. & Ther.
Locality: HONG KONG (Locality unknown). Wright s.n. (FH).
Habitat and Substrate: On rocks.
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Zhang Li: Hong Kong Bryophytes 57
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hubei,
Jiangsu, Jiangxi, Liaoning, Shaanxi, Shanxi, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Korea,
Japan, Vietnam, Thailand (Fang et al., 1998; He, 2000; Noguchi, 1989; Redfearn et al., 1996; Tan and
Iwatsuki, 1993).
Macromitrium heterodictyon Dix.
Localities: NEW TERRITORIES. MA ON SHAN. Ma On Shan Tsuen Stream. 440 m (SW), 2162;
450 m (SW), 2164. Tiu Shau Ngam. 590 m (NW), 267. – PAT SIN LENG. Hsien Ku Fung. 405 m
(NE), 1020; 420 m (NE), 1025. – TAI TO YAN. Summit. 540 m, 32, 33. – TSUEN WAN. Ah Nin
H.18, H.41 (BM). HONG KONG IS. MOUNT PARKER. Summit. 515 m, 2391. LANTAU IS.
LANTAU PEAK. Herklots 359 (BM). – NEI LAK SHAN. 455 m (NE), 2078. – SUNSET PEAK. San
Shek Wan Stream. 220 m (S), 2030; 360 m (S), 2147; 400 m (S), 33(9). Wong Lung Hang (Main
Streams). 455 m (NW), 2406. HONG KONG (Locality unknown). Wright s.n. (FH).
Habitat and Substrate: There were 17 specimens studied, of which 13 were collected in forest, 1 near
sea level, and 3 unknown. 7 were found on rocks, 6 on trunks, 1 on a rotten wood, 1 on a tree base, and
2 unknown.
Distribution: Endemic to Southern China (Fujian, Hong Kong) (Dixon, 1933).
Macromitrium japonicum Dozy & Molk.
Localities: NEW TERRITORIES. SAI KUNG. Kap Man Hang (Left Stream). 70 m (NE), 1226. –
WU KAU TANG. Lai Chi Wo Mangrove. But and Gao 90102 (originally det. As M. ferriei, CUHK).
HONG KONG IS. D’AGUILAR PEAK. 215 m (E), 186; 230 m (E), 519; 250 m (E), 511.
OUTLYING ISLANDS. AP CHAU. 10 m, 2497. – TAP MUN CHAU. 50 m, Lin et al. 1849 (IBSC,
KFBG).
Habitat and Substrate: Five of 7 specimens studied were collected in forest, and 2 unknown. 4 were
found on trunks, 2 on rocks, and 1 on rotten wood.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hebei, Heilongjiang,
Henan, Hong Kong, Hubei, Hunan, Inner Mongolia, Jiangsu, Jiangxi, Jilin, Liaoning, Shaanxi,
Shandong, Shanghai, Sichuan, Taiwan, Xizang, Zhejiang). Korea, Japan, Russian Far East, Vietnam,
Thailand (Bai, 1996; Fang et al., 1998; He, 2000; Jia and Wang, 1993; Noguchi, 1989; Rao et al.,
1997; Redfearn et al., 1996; Tan and Iwatsuki, 1993; Zhao and Cao, 1998).
Macromitrium microstomum (Hook. & Grev.) Schwaegr. (M. reinwardtii Schwaegr., fide Vitt (1983))
– So and Zhu (1996c) as M. reinwardtii
Locality: HONG KONG (Locality unknown). 400 m (So and Zhu, 1996c).
Habitat and Substrate: On a dry rock surface (So and Zhu, 1996c).
Distribution: China (Hainan, Hong Kong, Taiwan). Cambodia, Philippines, Malaysia, Borneo,
Australia, New Zealand, Southern Pacific, Polynesia, Vanuatu, Society Islands, Mexico, Costa Rica,
Cuba, Jamica, Brazil (Eddy, 1996; Fife, 1995; Higuchi, 1996; Mohamed and Tan, 1988; Sharp, Crum
and Eckel, 1994; Tan and Iwatsuki, 1991, 1993; Touw, 1978; Vitt, 1983; Whittier, 1976; Yano, 1995).
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58 Memoirs of the Hong Kong Natural History Society 26, 2003
Macromitrium nepalense (Hook. & Grev.) Schwaegr.
Localities: NEW TERRITORIES. TAI MO SHAN. Kadoorie Farm. 440 m (NW), 1071. HONG
KONG (Locality unknown). Wright s.n. (two bags, FH) as M. moorcroftii?.
Habitat and Substrate: One on shade rocks of mountain sides, and the other on a concrete wall in forest.
Distribution: China (Guangdong, Hainan, Hong Kong, Yunnan). India, Nepal, Laos, Myanmar,
Vietnam, Thailand, Cambodia, Malaysia, Philippines, Indonesia (Eddy, 1996; Gangulee, 1969-1980; Li
and Piippo, 1994; Redfearn et al., 1996; Tan and Iwatsuki, 1991, 1993).
Macromitrium subincurvum Card. & Ther.
Locality: HONG KONG (Locality unknown) (Theriot, 1906).
Habitat and Substrate: Unknown.
Distribution: Endemic to Hong Kong.
Macromitrium tuberculatum Dix.
Localities: NEW TERRITORIES. TAI MO SHAN. Herklots 297.O (BM). Tai Shing Stream. 815 m
(NE), 1426; 790 m (NE), 22. LANTAU IS. SUNSET PEAK. Wong Lung Hang (Main Streams). 595
m (N), 60.
Habitat and Substrate: Three of 4 specimens examined were collected in forest, and 1 unknown. 2 were
found on trunks, 1 on rocks, and 1 unknown.
Distribution: Endemic to Hong Kong.
Schlotheimia grevilleana Mitt.
Localities: NEW TERRITORIES. TAI MO SHAN. Ng Tung Chai Falls. 680 m (N), 1238. LANTAU
IS. LANTAU PEAK. 585 m (S), 1682. – SUNSET PEAK. Wong Lung Hang (East Lung Stream). 605
m (N), 2306. Wong Lung Hang (Main Streams). 595 m (N), 64.
Habitat and Substrate: All specimens were collected in forest. 2 were found on rocks, and 2 on trunks.
Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Taiwan, Yunnan,
Zhejiang). Japan, India, Sri Lanka, Vietnam, Thailand, Philippines, Indonesia, Central and South
Africa (Eddy, 1996; Gangulee, 1969-1980; Koponen and Enroth, 1992; Redfearn et al., 1996; Tan and
Iwatsuki, 1993).
Schlotheimia pungens Bartr.
Species new to Hong Kong.
Localities: LANTAU IS. LANTAU PEAK. 580 m (S), 1681; 730 m (N), 1337; 730 m (S), 1710; 775
m (N), 2467; 785 m (N), 2468. – SUNSET PEAK. Wong Lung Hang (East Lung Stream). 600 m (N),
1282. Wong Lung Hang (Main Streams). 385 m (N), 2107.
Habitat and Substrate: All specimens were collected in forest. 5 were found on rocks, 1 on a trunk, and
1 on a twig.
Distribution: Endemic to China (Anhui, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Jiangxi,
Sichuan, Taiwan, Zhejiang) (Fang et al., 1998; Redfearn et al., 1996).
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Zhang Li: Hong Kong Bryophytes 59
PLAGIOTHECIACEAE
Plagiothecium cavifolium (Brid.) Iwats. – So and Zhu (1996c).
Localities: NEW TERRITORIES. TAI MO SHAN. 900 m (So and Zhu, 1996c). Tai Shing Stream.
780 m (NE), 1431 (Det. By D.-K. Li, HKU, IBSC, KFBG, SHM). LANTAU IS. SUNSET PEAK. San
She Wan Stream. 700 m (SE), 27(23). Wong Lung Hang (Main Streams). 620 m (N), 2121 (Confirmed
by D.-K. Li, HKU, IBSC, KFBG, SHM).
Habitat and Substrate: On rocks in forest.
Distribution: China (Anhui, Guizhou, Heilongjiang, Henan, Hong Kong, Inner Mongolia, Jiangsu,
Jiangxi, Jilin, Liaoning, Shaanxi, Shandong, Shanghai, Sichuan, Yunnan, Zhejiang). Korea, Japan,
Mongolia, Russian Far East and Siberia, Nepal, Bhutan, Europe, North America (Anderson, Crum and
Buck, 1990; Bai, 1996; Fang et al., 1998; Redfearn et al., 1996).
Plagiothecium euryphyllum (Card. & Ther.) Iwats. – So and Zhu (1996c).
Localities: NEW TERRITORIES. TAI MO SHAN. 900 m (So and Zhu, 1996c). LANTAU IS.
LANTAU PEAK. 660 m (N), 2454 (Confirmed by D.-K. Li, HKU, IBSC, KFBG, SHM).
Habitat and Substrate: On soil in forest.
Distribution: China (Guangxi, Guizhou, Hainan, Hong Kong, Liaoning, Shandong, Taiwan, Xizang,
Zhejiang). Korea, Japan, Vietnam (Redfearn et al., 1996; Tan and Iwatsuki, 1993; Zhao and Cao,
1998).
Plagiothecium nemorale (Mitt.) Jaeg. – So and Zhu (1996c).
Localities: NEW TERRITORIES. TAI MO SHAN. 900 m (So and Zhu 1996c). LANTAU IS.
LANTAU PEAK. 725 m (N), 2459. – SUNSET PEAK. Wong Lung Hang (East Lung Stream). 700 m
(N), 38(10), 38(24).
Habitat and Substrate: On soil in forest.
Distribution: China (Anhui, Fujian, Guizhou, Heilongjiang, Hong Kong, Inner Mongolia, Jiangsu, Jilin,
Liaoning, Shaanxi, Shandong, Sichuan, Taiwan, Xizang, Zhejiang). Korea, Japan, India, Nepal,
Bhutan, Myanmar, Russia, Europe, Africa (Noguchi, 1994; Redfearn et al., 1996; Tan and Iwatsuki,
1993; Zhao and Cao, 1998).
POLYTRICHACEAE
Atrichum undulatum (Hedw.) P. Beauv. Var. gracilisetum Besch.
Variety new to Hong Kong. It is difficult to distinguish var. gracilisetum from var. undulatum in sterile
condition. Luo and Koponen (1984)’s treatment is followed.
Localities: NEW TERRITORIES. TAI MO SHAN. Ng Tung Chai Falls. 435 m (N), 1348. Shek
Kong Stream. 320 m (NW), 1972. Tai Shing Stream. 880 m (NE), 1129; 900 m (NE), 8(7). Yin Ngam
Stream. 385 m (NE), 1894. LANTAU IS. LIN FA SHAN. 645 m (E), 1459. – SUNSET PEAK. San
She Wan Stream. 700 m (SE), 27(6), 27(14).
Habitat and Substrate: There were 8 specimens studied, of which 6 were found in forest, 1 in grassland,
and 1 by a waterfall. 5 were found on soil, and 3 on rocks.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Heilongjiang, Henan, Hong Kong,
Hubei, Inner Mongolia, Jiangsu, Jiangxi, Jilin, Liaoning, Shaanxi, Shandong, Sichuan, Taiwan, Xizang,
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60 Memoirs of the Hong Kong Natural History Society 26, 2003
Yunnan, Zhejiang). Japan, Myanmar, Philippines, Russia, Europe (Lou and Koponen, 1986; Noguchi,
1987b; Redfearn et al., 1996; Tan and Iwatsuki, 1991, 1993)
Pogonatum camusii (Ther.) Touw
Species new to Hong Kong.
Localities: NEW TERRITORIES. PAT SIN LENG. Vicinity of Ngau Au Stream. 130 m (SE), 1031;
135 m (SE), 1030. LANTAU IS. SUNSET PEAK. Wong Lung Hang (East Lung Stream). 485 m (N),
1276
Habitat and Substrate: All specimens were collected in forest. 2 were found on sandy rocks, and 1 on
rocks.
Distribution: China (Guizhou, Hainan, Hong Kong, Taiwan). Japan, Vietnam, Thailand, Philippines,
Borneo, Indonesia (Hyvonen, 1989; Redfearn et al., 1996; Touw, 1978).
Pogonatum contortum (Brid.) Lesq.
Localities: NEW TERRITORIES. FAN LING. Kei Lak Tsai. 335 m, 35. – GRASS HILL. Fo Tan
Stream. 435 m (SE), 207; 475 m (SE), 201; 495 m (SE), 200. – HUNG FA LENG. San Kwai Tin
Stream. 360 m (N), 1156. – KOWLOON PEAK. Ho Chung Stream. 300 m, 1526. – MA ON SHAN.
Ma On Shan Tsuen Stream. 400 m (SW), 2160. Tai Tung Wo Liu Stream. 430 m (NE), 1584. Tiu Shau
Ngam. 165 m (NW), 255; 185 m (NW), 733; 340 m (NW), 103; 500 m (NW), Ng 8; 585 m (NW),
1587. – SAI KUNG. Nam Shek Tsim. 295 m (N), 1954. – TAI MO SHAN. Ng Tung Chai Falls. 360 m
(N), 1198; 500 m (N), 3(10). Shek Kong Stream. 520 m (NW), 1983; 745 m (NW), 1999; 770 m (NW),
2290. Tai Shing Stream. 200 m (SE), 17(8); 870 m (NE), 1182. Water-pumping House Stream. 700 m
(E), S55. – WU KAU TANG. So Lo Pun Back Stream. 50 m (NE), 1616. LANTAU IS. LANTAU
PEAK. 430 m (N), 1311; 440 m (S), 1668; 590 m (N), 1321. – LIN FA SHAN. 235 m (NE), 603. –
SUNSET PEAK. San Shek Wan Stream. 680 m (S), 2059. Wong Lung Hang (East Lung Stream). 770
m (N), 1301. Wong Lung Hang (Main Streams). 370 m (N), 2103.
Habitat and Substrate: Twenty-eight of 30 specimens examined were collected in forest, 1 by a
waterfall, and 1 by a stream. 25 were found on soil, and 5 on rocks.
Distribution: China (Anhui, Guangxi, Guizhou, Heilongjiang, Hong Kong, Jilin, Shaanxi, Shandong,
Sichuan, Taiwan). Korea, Japan, Russian Far East and Siberia, India, Nepal, North America (Gangulee,
1969-1980; Hyvonen, 1989; Redfearn et al., 1996).
Pogonatum neesii (C. Muell.) Dozy
Localities: NEW TERRITORIES. TAI MO SHAN. Ng Tung Chai Falls. 600 m (N), 2(8). Waterpumping House Stream. 835 m (E), 1967.
Habitat and Substrate: On soil in forest.
Distribution: China (Anhui, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hubei, Hunan,
Jiangsu, Jiangxi, Sichuan, Taiwan, Xinjiang, Xizang, Yunnan, Zhejiang). Korea,
Japan, India, Nepal, Bhutan, Sri Lanka, Myanmar, Vietnam, Thailand, Cambodia, Malaysia,
Philippines, Borneo, Indonesia, Papua New Guinea, Australia, Fiji, Vanuatu, New Caledonia, Samoa,
Russian Caucasus (Fang et al., 1998; Higuchi, 1996; Ignatov and Afonina, 1992; Koponen et al., 2000;
Mohamed and Tan, 1988; Redfearn et al.,1996; Touw, 1978).
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Zhang Li: Hong Kong Bryophytes 61
Polytrichastrum formosum (Hedw.) G. L. Smith (Polytrichum formosum Hedw.)
Localities: NEW TERRITORIES. TAI MO SHAN. Shek Kong Stream. 915 m (NW), 1466, 1467.
Tai Shing Stream. 845 m (NE), 16; 860 m (NE), 14.
Habitat and Substrate: On rocks in grassland.
Distribution: China (Anhui, Fujian, Guangdong, Guizhou, Heilongjiang, Hong Kong, Hunan, Inner
Mongolia, Jiangsu, Jiangxi, Jilin, Liaoning, Shanghai, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang).
Korea, Japan, Russian Siberia, Nepal, Syria, Australia, New Zealand, Europe, Africa, North America
(Fang et al., 1998; Fife, 1995; Gangulee, 1969-1980; Koponen et al., 2000; Redfearn et al., 1996;
Streimann and Curnow, 1989).
POTTIACEAE
Anoectangium thomsonii Mitt. – So and Zhu (1996c)
Locality: HONG KONG (Locality unknown). 400 m (So and Zhu, 1996c).
Habitat and Substrate: On a splash zone near a small waterfall (So and Zhu, 1996c).
Distribution: China (Anhui, Fujian, Guangdong, Hebei, Heilongjiang, Henan, Hong Kong, Jilin,
Liaoning, Shaanxi, Sichuan, Taiwan, Yunnan, Xizang, Zhejiang). Japan, Russian Far East, India,
Nepal, Myanmar (Li and Iwatsuki, 1997).
Barbula arcuata Griff. (Hydrogonium arcuatum (Griff.) Wijk. & Marg.)
Species new to Hong Kong.
Specimens examined: NEW TERRITORIES. TAI LAM. Kat Hing Bridge to Tsing Fai Tong. 110 m
(NW), 1049. – TAI MO SHAN. Summit. 915 m, 1175. – SHATIN. Sui Wo Road. 50 m, So 95515
(HKBP, IBSC).
Habitat and Substrate: Two of 3 specimens were collected in urban environment, and 1 in forest. 1 was
found on a concrete wall, 1 on rocks, and 1 unknown.
Distribution: China (Fujian, Guizhou, Henan, Hong Kong, Hunan, Inner Mongolia, Shaanxi, Shanxi,
Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Japan, India, Nepal, Myanmar, Thailand, Malaysia,
Singapore, Philippines, Indonesia, Papua New Guinea, Vanuatu, Western Oceania, West Indies,
Mexico, Venezuela, Brazil, Guatemala (He, 2000; Higuchi, 1996; Koponen et al., 2000; Li and Crosby,
2001; Mohamed and Tan, 1988; Noguchi ,1988; Redfearn et al., 1996; Tan and Iwatsuki, 1993).
Barbula indica (Hook.) Spreng. In Steud. (Semibarbula orientalis (Web.) Wijk & Marg., fide Saito
1975) – Gao (1996) as S. orientalis, Li and Crosby (2001)
Localities: NEW TERRITORIES. HUNG FA LENG. San Kwai Tin Stream. 420 m (N), 1159. –
PLOVER COVE RESERVOIR. Lo Fu Wat. 5 m, 127. HONG KONG IS. VICTORIA PEAK (Li and
Crosby, 2001). OUTLYING ISLANDS. LAMMA IS. So Kwu Wan. 30 m, Lin et al. 1983 (IBSC,
KFBG); 10 m, Lin et al. 1984 (IBSC, KFBG). – MA WAN. 5 m, 838. – PAK KWO CHAU. 60 m,
888. – PING CHAU. 5m, 2265, 2266, 2273; 10 m, 2267; 15 m, 2270. – TAI TAU CHAU. 5 m, 437. –
TAP MUN CHAU. 50 m, Lin et al. 1833, Lin et al. 1834 (KFBG, IBSC). HONG KONG (Locality
unknown). Wright 54, 78 (FH).
Habitat and Substrate: Most specimens were collected by the seashore. 10 were found on soil, 2 on
concrete steps, 1 on stone, 1 on a damp wall, 1 on a trunk, and 1 on rocks.
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62 Memoirs of the Hong Kong Natural History Society 26, 2003
Distribution: China (Beijing, Fujian, Guangdong, Hainan, Henan, Hong Kong, Inner Mongolia,
Jiangsu, Shandong, Shanxi, Sichuan, Taiwan, Xizang, Yunnan). Japan, India, Nepal, Myanmar,
Vietnam, Thailand, Malaysia, Singapore, Philippines, Borneo, Indonesia, Papua New Guinea,
Australia, Vanuatu, Hawaii, Africa, North America, Mexico, West Indies, Belize, Brazil (Anderson,
Crum and Buck, 1990; Eddy, 1990; Gao, 1996; He, 2000; Higuchi, 1996; Mohamed and Tan, 1988;
Noguchi,1988; Redfearn et al., 1996; Sharp, Crum and Eckel, 1994; Stoneburner et al., 1993; Tan and
Iwatsuki, 1993; Yano, 1995).
Barbula javanica Dozy & Molk. (Hydrogonium javanicum (Dozy & Molk.) Hilp. Var. convolutifolium
(Dix.) Chen, fide Saito 1975) – Gao (1996) as H. javanicum var. convolutifolium
Localities: NEW TERRITORIES. TAI LAM. Kiu Keng Shan. 235 m (SW), 339. – TAI MO SHAN.
Tai Yuen Stream. 350 m (S), 1847. – TUEN MUN. Tai Shui Hang. 170 m (NW), 549. LANTAU IS.
LANTAU PEAK. 150 m (S), 1658. – SZE PAK WAN. 5 m (S), 2130.
Habitat and Substrate: Three of 5 specimens were collected in forest, and 2 unknown. 3 were found on
rocks, and 2 on soil.
Distribution: China (Anhui, Fujian, Guangdong, Hainan, Henan, Hong Kong, Jiangsu, Shanxi, Taiwan,
Xizang, Yunnan). Japan, Pakistan, India, Nepal, Myanmar, Vietnam, Thailand, Cambodia, Malaysia,
Singapore, Philippines, Borneo, Indonesia, Papua New Guinea (Akiyama, 1996; Gao, 1996; He, 2000;
Li and Crosby, 2001; Mohamed and Tan, 1988; Noguchi, 1988; Norris and Koponen, 1989; Redfearn
et al., 1996; Tan and Iwatsuki, 1991, 1993; Touw, 1978).
Barbula unguiculata Hedw.
Locality: HONG KONG (Locality unknown) (Salmon, 1900).
Habitat and Substrate: Unknown (Salmon, 1900).
Distribution: Cosmopolitan.
Hyophila involuta (Hook.) Jaeg. – Li and Crosby (2001)
Localities: NEW TERRITORIES. SAI KUNG. Kap Man Hang (Left Stream). 60 m (NE), 1221. Kap
Man Hang Pool. 15 m (E), 1209. Tam Chai Shan. 35 m (E), 1036. – TAI LAM. Kiu Keng Shan. 290 m
(NW), 348. Lin Fa Shan. 335 m (SW), 676. – TAI MO SHAN. Shing Mun Reservoir. 190 m, 742. Tai
Shing Stream. 700 m (SE), 6(27). KOWLOON. TSIM SHA TSUI. 5 m (W), 25(3). HONG KONG
IS. CENTRAL. 45 m (NE), 23(1). – CONDUIT ROAD. 140 m (N), 22(1). – D’AGUILAR PEAK. 75
m (W), 1250. – VICTORIA PEAK. 295 m (SW), 1502. LANTAU IS. SUNSET PEAK. San She Wan
Stream. 100 m (S), 30(1). OUTLYING ISLANDS. LAMMA IS. So Kwu Wan. 50 m, Lin et al. 1975
(IBSC, KFBG). – PING CHAU. 5 m, 2276; 10 m, 2275. – TAI A CHAU. 10 m, 853. – SIU AP
CHAU. 5 m, 2492. HONG KONG (Locality unknown). Eddie M.27 (HKU, IBSC).
Habitat and Substrate: Fifteen of 19 specimens were collected in forest, 3 in urban areas, and 1 by a
path. 15 were found on rocks, 2 on soil, 1 on a tree base, and 1 on a concrete wall.
Distribution: Cosmopolitan.
Hyophila javanica (Nees & Blume) Brid. – Li and Crosby (2001)
Locality: HONG KONG IS. VICTORIA PEAK (Li and Crosby, 2001).
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Zhang Li: Hong Kong Bryophytes 63
Habitat and Substrate: Unknown
Distribution: Chnia (Beijing, Fujian, Hainan, Hong Kong, Shanghai, Sichuan, Yunnan). Indonesia (Li
and Crosby, 2001).
Hyophila setschwanica (Broth.) Hilp. Ex Chen – Li and Crosby (2001)
Locality: NEW TERROTORIES. TAI MO SHAN (Li and Crosby (2001) as Tai-Mo Shan (Mt.)).
Habitat and Substrate: Unknown
Distribution: Chnia (Hainan, Hong Kong, Sichuan, Yunnan) (Li and Crosby, 2001).
Luisierella barbula (Schwaegr.) Steere – So and Iwatsuki (1996)
Locality: NEW TERRITORIES. KOWLOON PEAK. 400 m, So 95802 (IBSC, HKBU).
Habitat and Substrate: Unknown.
Distribution: China (Fujian, Hong Kong, Jiangsu, Xizang, Yunnan). Japan, Indonesia, North America,
West Indies, Brazil (Anderson, Crum and Buck, 1990; Eddy, 1996; Li and Crosby, 2001; So and
Iwatsuki, 1996; Yano, 1995).
Scopelophila cataractae (Mitt.) Broth. (Merceyopsis sikkimensis (C. Muell.) Broth. & Dix.).
Genus and species new to Hong Kong.
Localities: NEW TERRITORIES. HUNG FA LENG. Muk Min Tau Stream. 395 m (E), 504. – MA
ON SHAN. Ma On Shan Tsuen Stream. 405 m (SW), 2161. Tai Tung Wo Liu Stream. 170 m (NE),
1575; 175 m (NE), 1574; 420 m (NE), 1582. – TAI MO SHAN. Kadoorie Farm. 300 m, So 94905
(HKBU, IBSC). Ng Tung Chai Falls. 170 m (N), 1775; 190 m (N), 1770; 530 m (N), 1193. Summit.
910 m, 1179.
Habitat and Substrate: Eight of 10 specimens were collected in forest, 1 by a road, and 1 unknown. 6
were found on rocks, 2 on soil, 1 on a concrete wall, and 1 unknown.
Distribution: China (Fujian, Guangdong, Guangxi, Gansu, Hong Kong, Hunan, Jiangsu, Jiangxi,
Liaoning, Shaanxi, Shandong, Sichuan, Taiwan, Xizang, Yunnan). Korea, Japan, India, Nepal, Bhutan,
Philippines, Indonesia, Papua New Guinea, Africa, North America, Mexico, Guatemala, Ecuador
(Anderson, Crum and Buck, 1990; Eddy, 1990; Gao, 1996; Long, 1994; Norris and Koponen, 1989;
O’Shea, 1999; Sharp, Crum and Eckel, 1994; Tan and Iwatsuki, 1991; Zhao and Cao, 1998).
Trichostomum hattorianum Tan & Iwats. (T. involutum Broth.)
Localities: NEW TERRITORIES. TAI MO SHAN. Tai Shing Stream. 900 m (NE), 8(15), 8(19).
Habitat and Substrate: On rocks in forest.
Distribution: Endemic to China (Fujian, Guangdong, Guizhou, Henan, Hong Kong, Jiangxi, Ningxia,
Shanghai, Taiwan, Yunnan, Zhejiang) (Li and Iwatsuki, 1996).
Weissia controversa Hedw.
Localities: HONG KONG IS. REPULSE BAY GAP. 190 m, 1370. OUTLYING ISLANDS. NGA
YING CHAU. 5 m, 464. – NGO CHAU TSAI (Not found on the map). 20 m, 927. – PAK KWO
CHAU. 75 m, 889; 90 m, 891 (with Brachymenium exile). – PING CHAU. 5 m, 2263. – SHEK CHAU.
10 m, 872, 874. – SUNG KONG. 55 m, 909. – TUNG KWO CHAU. 55 m, 893. – WAGLAN IS. 45
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64 Memoirs of the Hong Kong Natural History Society 26, 2003
m, 895; 60 m, 896. – WAN HAU CHAU. 10 m, 876. – YEUNG CHAU (near Kat O). 20 m, 2529. –
YING CHAU (Not found on the map, probably it is Nga Ying Chau). 5 m, 461. – YUEN CHAU. 5 m,
871. – YUEN KONG CHAU. 10 m, 847, 848 (with B. exile).
Habitat and Substrate: Most specimens were collected by the seashore. 11 were found on soil, and 7 on
rocks.
Distribution: Subcosmopolitan.
Weissia edentula Mitt.
Species new to Hong Kong.
Localities: NEW TERRITORIES. SAI KUNG. Hoi Ha. 60 m, 1039. OUTLYING ISLANDS. TIT
LO CHAU (Not found on the map). 40 m, 923.
Habitat and Substrate: One was collected on soil in forest, the other on soil.
Distribution: China (Anhui, Fujian, Guangdong, Guizhou, Heilongjiang, Henan, Hong Kong, Hunan,
Inner Mongolia, Jiangsu, Jilin, Liaoning, Shaanxi, Shandong, Shanghai, Sichuan, Taiwan, Xizang,
Yunnan, Zhejiang). Japan, India, Sri Lanka, Vietnam, Thailand, Cambodia, Malaysia, Philippines,
Indonesia, Papua New Guinea, Australia, Oceania, Africa (Akiyama, 1996; Bai, 1996; Eddy, 1990;
Gao, 1996; He, 2000; Mohamed and Tan, 1988; Noguchi, 1988; Norris and Koponen, 1989; O’Shea,
1999; Redfearn et al., 1996; Tan and Iwatsuki, 1991, 1993; Zhao and Cao, 1998).
Weissia longifolia Mitt. (W. crispa (Hedw.) Mitt., fide Li and Crosby (2001))
Locality: LANTAU IS. (Sullivant and Lesquereux 1978 as L’am Tong Island).
Habitat and Substrate: In small dense patches on hillsides among bushes and rocks (Sullivant and
Lesquereux, 1978).
Distribution: China (Anhui, Fujian, Guangdong, Guizhou, Hainan, Heilongjiang, Henan, Hong Kong,
Hunan, Inner Mongolia, Jiangsu, Jilin, Liaoning, Ningxia, Shaanxi, Shandong, Shanghai, Shanxi,
Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Japan, Russian Far East, West Asia, Pakistan, India,
Laos, Vietnam, Europe, North Africa (Gao (1996), Li and Crosby (2001) and Zhao and Cao (1998)
respectively included Americas and North America within its distribution ranges, but I can’t find any
reliable recording about this species occurred in Americas or North America; Matsui,1995; Redfearn et
al., 1996; Tan and Iwatsuki, 1993; Zhao and Cao, 1998).
PTEROBRYACEAE
Calyptothecium wightii (Mitt.) Fleisch.
Locality: HONG KONG (Locality unknown). 600 m, So 94520A (HKBU, IBSC).
Habitat and Substrate: On rocks.
Distribution: China (Hong Kong, Sichuan, Taiwan, Yunnan, Zhejiang). India, Nepal, Bangladesh, Sri
Lanka, Myanmar, Laos, Vietnam, Thailand (Gangulee, 1969-1980; Redfearn et al., 1996; Noguchi,
1987a).
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Zhang Li: Hong Kong Bryophytes 65
RACOPILACEAE
Racopilum aristatum Mitt.
Localities: NEW TERRITORIES. KAU LUNG HANG SHAN. 30 m (NW), 1536. – MA ON SHAN.
Ngong Ping. 385 m, 121. – TAI MO SHAN. Ng Tung Chai Falls. 200 m (N), 16(4), 16(8), 16(21); 600
m (N), 2(17). Shing Mun Reservoir. 200 m, 749. Water-pumping House Stream. 700 m (E), S41. – TAI
TO YAN. Lam Tusen Stream. 120 m (E), 707. HONG KONG IS. D’AGUILAR PEAK. 225 m (E),
530. LANTAU IS. LANTAU PEAK. 550 m (S), 1677.
Habitat and Substrate: All specimens were collected in forest. 8 were found on rocks, 2 on trunks, and
1 on a tree base.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Jiangsu,
Jiangxi, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Japan, Thailand, Philippines (He, 2000; Fang et
al., 1998; Redfearn et al., 1996; Tan and Iwatsuki, 1991).
RHIZOGONIACEAE
Pyrrhobryum spiniforme (Hedw.) Mitt.
Localities: NEW TERRITORIES. MA ON SHAN. Tiu Shau Ngam. 580 m (NW), 1588; 590 m
(NW), 268. – PAT SIN LENG. Nam Chung Stream. 260 m (NE), 1002. Pin Fung Shan. 470 m (SW),
385. Wang Shan Keuk Sheung Tsuen. 250 m, Ng 10. LANTAU IS. LANTAU PEAK. 715 m (N),
1335; 730 m (N), 2461; 735 m (N), 1342. – SHUI LO CHO. 250 m (N), 568; 260 m (N), 578. –
SUNSET PEAK. San Shek Wan Stream. 510 m (S), 2052. Wong Lung Hang (Main Streams). 290 m
(N), 2098.
Habitat and Substrate: All specimens were collected in forest. 5 were found on soil, 3 on humus, 3 on
rocks, and 1 on a trunk.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Hainan, Hong Kong, Hunan, Taiwan,
Zhejiang). Korea, Japan, India, Nepal, Sri Lanka, Myanmar, Vietnam, Thailand, Cambodia, Malaysia,
Singapore, Philippines, Borneo, Indonesia, Papua New Guinea, Australia, New Zealand, Society
Islands, Vanuatu, Hawaii, Africa, Madagascar, North America, Brazil (Anderson, Crum and Buck,
1990; Eddy, 1996; Gangulee, 1969-1980; He, 2000; Higuchi, 1996; Mohamed and Tan, 1988;
Redfearn et al., 1996; Sharp, Crum and Eckel, 1994; Tian, 1999; Touw, 1978; Whittier, 1976; Yano,
1995).
SEMATOPHYLLACEAE
Acroporium secundum (Reinw. & Hornsch.) Fleisch.
Species new to mainland China and genus and species new to Hong Kong.
Localities: NEW TERRITORIES. WU KAU TANG. Fan Kei Tok Summit. 345 m, 1601. Fan Kei
Tok Feng Shui Wood. 350 m (N), 1603, 1605. LANTAU IS. LANTAU PEAK. 860 m (N), 2475.
Habitat and Substrate: All 4 specimens were collected in forest. 2 were found on rocks, 1 on a tree
base, and 1 on soil.
Distribution: China (Hainan, Hong Kong, Taiwan). Japan, Vietnam, Thailand, Malaysia, Singapore,
Philippines, Borneo, Indonesia (Frahm et al.,1990; He, 2000; Mohamed and Tan, 1988; Tan and
Iwatsuki, 1993; Tan and Jia,1999).
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66 Memoirs of the Hong Kong Natural History Society 26, 2003
Brotherella erythrocaulis (Mitt.) Fleisch. – Tan and But (1997), Tan and Jia (1999)
Localities: NEW TERRITORIES. MA ON SHAN. Tiu Shau Ngam. 190 m (NW), 259; 340 m (NW),
100. – PAT SIN LENG. Pin Fung Shan. 430 m (SW), 403; 470 m (SW), 388. – SAI KUNG. Nam She
Tsim. 380 m (N), 1948. – TAI MO SHAN. Shek Kong Stream. 750 m (NW), 2300. KOWLOON (Tan
and But 1997). LANTAU IS. SUNSET PEAK. Wong Lung Hang (East Lung Stream). 600 m (N),
37(15).
Habitat and Substrate: All specimens were collected in forest. 4 were found on rocks, and 3 on trunks.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Hainan, Hong Kong, Inner Mongolia,
Jiangxi, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). India, Bhutan, Myanmar, Thailand (Bai, 1996;
Fang et al., 1998; Gangulee, 1969-1980; He, 2000; Tan and Iwatsuki, 1993; Tan and But, 1997; Tan
and Jia, 1999).
Brotherella falcata (Dozy & Molk.) Fleisch.
Species new to Hong Kong.
Specimen examined: LANTAU IS. LANTAU PEAK. 875 m (N), 2480.
Habitat and Substrate: On humus in forest.
Distribution: China (Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hunan, Taiwan, Yunnan,
Zhejiang). Japan, Laos, Thailand, Borneo, Malesia (He, 2000; Redfearn et al., 1996; Tan and Jia, 1999;
Touw, 1978).
Clastobryopsis planula (Mitt.) Fleisch.
Genus and species new to Hong Kong.
Localities: NEW TERRITORIES. MA ON SHAN. Tai Tung Wu Liu Stream. 410 m (NE), 1581. –
TAI MO SHAN. Tai Shing Stream. 600 m (SE), 21(6); 640 m (SE), 1918; 780 m (NE), 1428; 835 m
(NE), 1420, 1424; 855 m (NE), 1415; 880 m (NE), 1127; 885 m (NE), 1352; 900 m (NE), 1120, 8(5),
8(18); 925 m (NE), 1107. LANTAU IS. LANTAU PEAK. 570 m (S), 1671; 575 m (S), 1680; 735 m
(N), 1341; 755 m (N), 2463. – LIN FA SHAN. 675 m (E), 1463, 1465. – SUNSET PEAK. Wong Lung
Hang (East Lung Stream). 600 m (N), 37(7); 700 m (N), 38(22); 770 m (N), 2249; 800 m (E), 42(11).
Habitat and Substrate: All 23 specimens were collected in forest. 8 were found on twigs, 7 on trunks, 5
on rocks, 2 on rotten twigs, and 1 on a rotten trunk.
Distribution: China (Guangdong, Guangxi, Guizhou, Hong Kong, Sichuan, Taiwan, Xizang, Yunnan).
India, Nepal, Bhutan (Gangulee, 1969-1980; Li, Liao and Huang, 1998; Tan and Jia, 1999).
Isocladiella surcularis (Dix.) Tan & Mohamed – Tan and But (1997), Tan and Jia (1999).
Localities: NEW TERRITORIES. HUNG FA LENG. San Kwai Tin Stream. 380 m (N), 1151. – MA
ON SHAN. Tiu Shau Ngam. 500 m (NW), Ng 7. – PAT SIN LENG. Pin Fung Shan. 415 m (SW), 402;
445 m (SW), 400. – PLOVER COVE RESERVOIR. Ngor Kai Teng. 215 m, 130. – SAI KUNG. Nam
She Tsim. 435 m (N), 1944; 450 m, Choi 3. – TAI MO SHAN. Kadoorie Farm (Tan and But 1997, Tan
and Jia 1999). Ng Tung Chai Falls. 500 m (N), 3(14); 700 m (N), 1(18). Shek Kong Stream. 755 m
(NW), 2296; 785 m (NW), 2318. Tai Shing Stream. 500 m (SE), 20(10); 600 m (SE), 21(5); 700 m
(SE), 6(11); 800 m (NE), 7(10). Water-pumping House Stream. 700 m (E), S3, S8, S31, S58. – TAI TO
YAN. Summit, 475 m, 30. – WU KAU TANG. Fan Kei Tok Summit. 350 m, 1595. Wu Kau Tang
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Zhang Li: Hong Kong Bryophytes 67
Feng Shui Wood. 100 m, 1789. HONG KONG IS. D’AGUILAR PEAK. 275 m (E), 239. – HIGH
WEST. 485 m (N), 1732. LANTAU IS. LIN FA SHAN. 135 m (NE), 596; 610 m (NE), 641. –
SUNSET PEAK. San She Wan Stream. 600 m (S), 26(15); 700 m (SE), 27(11). Wong Lung Hang
(East Lung Stream). 200 m (N), 40(6). OUTLYING ISLANDS. KAU SAI CHAU. Tai Leng Stream.
140 m (NW), 288.
Habitat and Substrate: Twenty-eight of 30 specimens were collected in forest, and 2 unknown. 11 were
found on trunks, 7 on rocks, 4 on twigs, 3 on leaves, 2 on rotten wood, 2 on rotten twigs, and 1 on a
tree base.
Distribution: China (Guangdong, Guangxi, Hainan, Hong Kong, Taiwan). Japan, Sri Lanka, Indochina,
Thailand, Malesia, Australia (He, 2000; Tan and Jia, 1999).
Papillidiopsis complanata (Dix.) Buck & Tan
Species new to Hong Kong.
Localities: NEW TERRITORIES. MA ON SHAN. Tai Tung Wo Liu Stream. 155 m (NE), 1565. –
PAT SIN LENG. Hsien Ku Fung. 370 m (NE), 1011; 395 m (NE), 1018. Wang Chung Stream. 390 m
(E), 3. – TAI MO SHAN. Tai Shing Stream. 400 m (SE), 19(13). – WU KAU TANG. Fan Kei Tok
Summit. 345 m, 1596; 350 m, 1594. LANTAU IS. LANTAU PEAK. 480 m (N), 2447; 635 m (N),
2452; 760 m (N), 2465. – POR KAI SHAN. 330 m (NE), 2412. – SUNSET PEAK. Wong Lung Hang
(East Lung Stream). 235 m (N), 2239; 500 m (N), 36(14); 600 m (N), 37(2); 700 m (N), 38(14). Wong
Lung Hang (Main Streams). 285 m (NW), 2400; 585 m (N), 87; 595 m (N), 62; 645 m (N), 2123.
Habitat and Substrate: Eighteen of 19 specimens examined were collected in forest, and 1 by a
waterfall. 15 were found on rocks, 2 on twigs, 1 on a rotten wood, and 1 on a trunk.
Distribution: China (Guangdong, Hainan, Hong Kong). Japan, Indochina, Thailand, Malaysia, Africa
(He, 2000; Tan and Jia, 1999).
Papillidiopsis macrosticta (Broth. & Par.) Buck & Tan – Tan and But (1997), Tan and Jia (1999).
Localities: NEW TERRITORIES. GRASS HILL. Fo Tan Stream. 445 m (SE), 206. – HUNG FA
LENG. San Kwai Tin Stream. 405 m (N), 1145. – MA ON SHAN. Tai Tung Wo Liu Stream. 80 m
(NE), 1563. Tiu Shau Ngam. 55 m (NW), 242; 125 m (NW), 245; 145 m (NW), 730; 415 m (NW),
116. – PAT SIN LENG. Hsien Ku Fung. 310 m (E), 2. Pin Fung Shan. 470 m (SW), 386. – SAI
KUNG. Yung Shue O (Jacob’s Ladder). 175 m (W), 296. – SHATIN (Tan and But 1997). – TAI MO
SHAN. Ng Tung Chai Falls. 500 m (N), 3(4); 700 m (N), 1(26). Tai Shing Stream. 200 m (SE), 17(1);
400 m (SE), 19(5). – TUEN MUN. Tai Shui Hang. 65 m (NW), 540, 543. – WU KAU TANG. Bride’s
pool (Tan and But 1997). LANTAU IS. LANTAU PEAK. 120 m (S), 1657. – SHUI LO CHO. 210 m
(N), 559, 561. OUTLYING ISLANDS. KAU SAI CHAU. Tai Leng Stream. 130 m (NW), 278.
Habitat and Substrate: There were 20 specimens, of which 16 were collected in forest, 3 by streams,
and 1 in bamboo forest. 18 were found on rocks, and 2 on soil.
Distribution: China (Guangdong, Hong Kong, Taiwan). Japan, Indochina (Tan and Jia, 1999).
Sematophyllum subhumile (C. Muell.) Fleisch. (S. subhumile (C. Muell.) Fleisch. Ssp. Japonicum
(Broth.) Broth.) – Tan and But (1997), Tan and Jia (1999)
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68 Memoirs of the Hong Kong Natural History Society 26, 2003
Localities: NEW TERRITORIES. FAN LING. Kei Lak Tsai. 205 m, 36. – HO PUI RESERVOIR.
180 m, 801. – MA ON SHAN. Ngong Ping. 345 m, 124. – PLOVER COVE RESERVOIR. Ngor Kai
Teng. 210 m, 136. – TAI LAM. Kat Hing Bridge to Tsing Fai Tong. 90 m (NW), 1043. – TAI LO
SHAN. Kwun Yam Shan. 90 m (N), 928. – TAI MO SHAN. Ng Tung Chai Falls. 300 m (N), 5(10);
680 m (N), 1237. Tai Shing Stream. 300 m (SE), 18(3); 700 m (SE), 6(1). – TAI PO (Tan and But,
1997). HONG KONG IS. HIGH WEST. 490 m (N), 1729. – MOUNT PARKER (Tan and But, 1997).
LANTAU IS. LANTAU PEAK. 430 m (N), 2446. – SUNSET PEAK. San She Wan Stream. 400 m
(S), 33(10); 500 m (S), 34(13), 34(20); 600 m (S), 26(9). Wong Lung Hang (East Lung Stream). 400 m
(N), 35(8). OUTLYING ISLANDS. PING CHAU. 40 m, 2279. HONG KONG (Locality unknown).
Herklots H.57 (BM).
Habitat and Substrate: Eighteen of 19 specimens were collected in forest, and 1 unknown. 13 were
found on rocks, 2 on trunks, 1 on a tree base, 1 on a twig, 1 on a liana, and 1 unknown.
Distribution: China (Guangdong, Guizhou, Hong Kong, Hunan , Jiangsu, Liaoning, Shaanxi, Taiwan,
Xizang, Yunnan). Korea, Japan, Himalaya, Myanmar, Laos, Vietnam, Thailand, Cambodia,
Philippines, Indonesia, Caroline Islands (He, 2000; Rao et al., 1997; Tan and Jia, 1999; Tan and
Iwatsuki, 1993).
Sematophyllum subpinnatum (Brid.) Britt. (S. demissum (Wils.) Mitt.?, S. robustulum (Card.) Broth.,
fide Tan and Jia (1999)) – Tan and But (1997) as S. demissum and S. robustulum, Tan and Jia (1999)
Localities: NEW TERRITORIES. FAN LING. Kei Lak Tsai. 205 m, 38. – GRASS HILL. Fo Tan
Stream. 520 m (SE), 195; 525 m (SE), 192. – HUNG FA LENG. Muk Min Tau Stream. 55 m (E), 490.
– KAU LUNG HANG SHAN. 100 m (NW), 1544. – MA ON SHAN. Ngong Ping. 345 m, 123. –
SHATIN (Tan and But 1997). – TAI LAM (Tan and But 1997 as Tai Nam Chung and Tai Nan). – TAI
LO SHAN. Kwun Yam Shan. 90 m (N), 929. – TAI MO SHAN. Kadoorie Farm (Tan and But, 1997).
Ng Tung Chai Falls. 300 m (N), 5(16); 600 m (N), 2(7); 645 m (N), 1229; 685 m (N), 1240; 700 m (N),
1(15), 1(20). Shek Kong Stream. 815 m (NW), 2282. Tai Shing Stream. 200 m (SE), 17(17), 17(19);
300 m (SE), 18(1); 500 m (SE), 20(6); 915 m (NE), 1121. Tai Yuen Stream. 250 m (S), 1842. – TAI
PO (Tan and But, 1997). – TAI TO YAN. Lam Tsuen Stream. 110 m (E), 703. – WU KAU TANG.
Bride’s Pool (Tan and But, 1997). KOWLOON (Locality unknown). Ah Nin H.38 (BM). HONG
KONG IS. VICTORIA PEAK. 415 m, 1507. LANTAU IS. LANTAU PEAK (Tan and But 1997). –
SUNSET PEAK. San She Wan Stream. 200 m (S), 31(3), 31(12); 500 m (S), 34(8); 600 m (S), 26(13).
Wong Lung Hang (East Lung Stream). 100 m (N), 39(14). OUTLYING ISLANDS. TSING YI. 270
m, 982; 300 m, 979. – KIU TSUI CHAU. 5 m, 474. HONG KONG (Locality unknown). Herklots
H.56A (BM).
Habitat and Substrate: Twenty-seven of 30 specimens were collected in forest, 1 by a stream, 1 near
sea level, and 1 unknown. 19 were found on rocks, 6 on trunks, 3 on twigs, 1 on a tree, and 1 unknown.
Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Taiwan, Yunnan,
Zhejiang). Japan, Malaysia, Singapore, Africa, North, Central and South America (Anderson, Crum
and Buck, 1990; Buck, 1998; Mohamed and Tan, 1988; O’Shea, 1999; Tan and Jia, 1999).
Sematophyllum subpinnatum (Brid.) Britt. F. tristiculum (Mitt.) Tan & Jia (S. tristiculum (Mitt.)
Fleisch.)
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Zhang Li: Hong Kong Bryophytes 69
Locality: HONG KONG (Locality unknown) (Noguchi and Shin, 1952).
Habitat and Substrate: Unknown (Noguchi and Shin, 1952).
Distribution: China (Guangdong, Hong Kong, Taiwan, Yunnan, Zhejiang). India, Sri Lanka, Myanmar,
Vietnam, Thailand, Philippines, Indonesia (He, 2000; Noguchi and Shin, 1952; Redfearn et al., 1996;
Tan and Iwatsuki, 1993).
Taxithelium instratum (Brid.) Broth. In Ren. & Card. – Tan and But (1997).
Locality: NEW TERRITORIES. Sai Kung (Tan and But, 1997).
Habitat and Substrate: Unknown (Tan and But, 1997).
Distribution: China (Anhui, Hong Kong, Taiwan). Myanmar, Vietnam, Thailand, Cambodia, Malaysia,
Singapore, Philippines, Borneo, Australia, Africa (He, 2000; Mohamed and Tan, 1988; O’Shea, 1999;
Tan and Iwatsuki, 1991, 1993; Streimann and Curnow, 1989; Touw, 1978).
Taxithelium nepalense (Schwaegr.) Broth. – Tan and But (1997).
Locality: HONG KONG (Locality unknown) (Tan and But, 1997).
Habitat and Substrate: Unknown (Tan and But, 1997).
Distribution: China (Guangdong, Hainan, Hong Kong, Yunnan). India, Nepal, Bangladesh, Myanmar,
Laos, Vietnam, Thailand, Cambodia, Philippines, Borneo, Australia (He, 2000; Redfearn et al., 1996;
Streimann and Curnow, 1989; Tan and Iwatsuki, 1991, 1993; Touw, 1978).
Taxithelium oblongifolium (Sull. & Lesq.) Iwats. (T. batanense Bartr.) – Tan and But (1997).
Localities: NEW TERRITORIES. HUNG FA LENG. San Kwai Tin Stream. 375 m (N), 1149. –
KAU LUNG HANG SHAN. 95 m (NW), 1546. – LAM TSUEN. She Shan Tsuen Feng Shui Wood.
100 m, 1365. Tai Om Feng Shui Wood. 110 m, 1907. – SAI KUNG. Kap Man Hang (Left Stream). 40
m (NE), 1218. Tam Chai Shan. 35 m (E), 1035. Yung Shue O (Jacob’s Ladder). 265 m (W), 299. –
SHATIN (Tan and But, 1997). – TAI LAM. Hung Shui Shan. 315 (N), 431. Kiu Keng Shan. 265 m
(NW), 346. Lin Fa Shan. 305 m (SW), 671. – TAI LO SHAN. Kwun Yam Shan. 140 m (N), 945. –
TAI MO SHAN. Lead Mine Pass. 455 m (SW), 1475, 1477. Ng Tung Chai Falls. 200 m (N), 16(17);
395 m (N), 4(7). Shing Mun Reservoir. 205 m, 750. Tai Shing Stream. 300 m (SE), 18(6). – TAI PO
(Tan and But, 1997). – TAI TO YAN. Lam Tsuen Stream. 85 m (E), 686, 696; 240 m (E), 718. San
Lung Wai Stream. 315 m (W), 761. – WU KAU TANG. Fan Kei Tok Feng Shui Wood. 280 m (N),
1610. Mirror Pool Feng Shui Wood. 60 m, 1746. HONG KONG IS. D’AGUILAR PEAK. 75 m (W),
1249; 110 m (W), 145, 147; 135 m (W), 149; 235 m (W), 163; 250 m (E), 508. – NAM FUNG ROAD
FENG SHUI WOOD. 140 m (S), 14(1), 14(2). – TAI TAM RESERVOIR. 90 m, 1371; 145 m, 1379. –
VICTORIA PEAK. 325 m (SW), 1503, 1504. LANTAU IS. LIN FA SHAN. 280 m (NE), 608. –
SHUI LO CHO. 230 m (N), 566. – SUNSET PEAK. San Shek Wan Stream. 100 m (S), 30(5); 130 m
(S), 2022; 200 m (S), 31(10); 500 m (S), 34(5). Wong Lung Hang (East Lung Stream). 100 m (N),
39(5); 320 m (N), 41(13); 400 m (N), 35(3); 500 m(N), 36(13). OUTLYING ISLANDS. LAMMA IS.
Yung Shue Ha. 80 m, Lin et al. 2007 (IBSC, KFBG). – WONG YI CHAU. 25 m, 456.
Habitat and Substrate: All 47 specimens were collected in forest. 32 were found on rocks, 5 on trunks,
4 on rotten wood, 1 on a tree base, 1 on a tree base and rocks, 1 on a rotten twig, 1 on a liana, 1 on
bark, and 1 on soil.
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70 Memoirs of the Hong Kong Natural History Society 26, 2003
Distribution: China (Hainan, Hong Kong, Taiwan, Zhejiang). Japan (Matsui, 1995; Redfearn et al.,
1996; Tan and But, 1997).
Taxithelium parvulum (Broth. & Par.) Broth. – Tan and But (1997).
Localities: NEW TERRITORIES. CASTLE PEAK. 130 m (NE), 1399. – HUNG FA LENG. Muk
Min Tau Stream. 30 m (E), 479. – KAU LUNG HANG SHAN. 160 m (NW), 1549. – PAT SIN LENG.
Hsien Ku Fung. 455 m (NE), 1028. – SAI KUNG. Yung Shue O (Jacob’s Ladder). 180 m (W), 297.
OUTLYING ISLANDS. LAMMA IS. So Kwu Wan. 70 m, Lin et al. 2001.
Habitat and Substrate: All specimens were collected in forest. 4 were found on rocks, 1 on a trunk, and
1 on a rotten trunk.
Distribution: China (Hong Kong, Taiwan). Japan (Noguchi, 1994; Redfearn et al., 1996).
Trichosteleum boschii (Dozy & Molk.) Jaeg. – Tan and But (1997)
Localities: NEW TERRITORIES. SAI KUNG. Kap Man Hang (Left Stream). 110 m (NE), 1228. –
TAI LAM. Hung Shui Shan. 280 m (N), 434. – TAI MO SHAN. Lead Mine Pass. 455 m (SW), 1476.
Water-pumping House Stream. 700 m (E), S13. HONG KONG IS. VICTORIA PEAK. 270 m (SW),
1493. – TAI TAM RESERVOIR (Tan and But, 1997). LANTAU IS. LIN FA SHAN. 665 m (E), 1461.
– SUNSET PEAK. Wong Lung Hang (East Lung Stream). 320 m (N), 41(8). OUTLYING ISLANDS.
KAU SAI CHAU. Tai Leng Stream. 130 m (NW), 274; 135 m (NW), 279, 281; 140 m (NW), 287. –
TAI TAU CHAU. 30 m, 93. – TSING YI. 170 m, 984.
Habitat and Substrate: Twelve of 13 specimens were collected in forest, and 1 in bamboo forest. 9 were
found on rocks, 3 on soil, and 1 on a twig.
Distribution: China (Guangdong, Hainan, Hong Kong, Taiwan, Yunnan). Japan, India, Nepal,
Myanmar, Laos, Vietnam, Thailand, Cambodia, Malaysia, Singapore, Philippines, Borneo, Australia,
Oceania (Gangulee, 1969-1980; Mohamed and Tan, 1988; Noguchi, 1994; Redfearn et al., 1996; Tan
and Iwatsuki, 1991, 1993; Tan and Jia, 1999; Tan, Ramsay and Schofield, 1996; Touw, 1978).
SPLACHNACEAE
Gymnostomiella longinervis Broth.
Li and Crosby (2001) placed this genus in the Pottiaceae, and they recognized only one taxon of G.
vernicosa (Hook.) Fleisch. Var. tenerum (C. Muell. Ex Dus.) Arts, as occurring in China.
Locality: NEW TERRITORIES. TAI MO SHAN. Kadoorie Farm. 550 m (NW), Lin et al. 93 (IBSC,
KFBG).
Habitat and Substrate: On rocks under Rhododendron shrubs.
Distribution: China (Guangdong, Guangxi, Hainan, Hong Kong, Jiangxi, Taiwan, Xizang, Yunnan,
Zhejiang). Japan, Myanmar, Philippines (Li, 2000; Redfearn, 1991, Redfearn et al. 1996, Tan and
Iwatsuki 1993).
STEREODONTACEAE
Entodontopsis anceps (Bosch & Sande Lac.) Buck & Ireland
Locality: NEW TERRITORIES. TAI MO SHAN. Foot. Iwatsuki 80 (IBSC, NICH).
Habitat and Substrate: On tree.
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Zhang Li: Hong Kong Bryophytes 71
Distribution: China (Anhui, Hainan, Hong Kong, Taiwan, Yunnan). India, Bangladesh, Myanmar,
Vietnam, Thailand, Philippines (Gangulee, 1969-1980; He, 2000; Redfearn et al., 1996; Tan and
Iwatsuki,1991, 1993).
THELIACEAE
Fauriella tenerrima Broth. – So and Zhu (1996c).
Localities: NEW TERRITORIES. TAI MO SHAN. Shek Kong Stream. 730 m (NW), 1998. Tai
Shing Stream. 300 m (SE), 18(2); 500 m (SE), 20(11); 700 m (SE), 6(4); 865 m (NE), 1135. LANTAU
IS. LANTAU PEAK. 475 m (S), 1669.
Habitat and Substrate: All 6 specimens were collected in forest. 3 specimens were found on rocks, 2 on
a trunk, and 1 on a twig.
Distribution: Endemic to China (Anhui, Guangxi, Guizhou, Hong Kong, Hunan, Jiangxi). (Fang et al.,
1998; Gao and Cao, 1992; Jia and Wang, 1993; Rao et al., 1997).
THUIDIACEAE
Claopodium aciculum (Broth.) Broth.
Localities: NEW TERRITORIES. HO PUI RESERVOIR. 280 m, 813. – HUNG FA LENG. Mu Min
Tau Stream. 35 m (E), 485. – SAI KUNG. Yung Shue O (Jacob’s Ladder). 180 m (W), 298. – TAI MO
SHAN. Water-pumping House Stream. 700 m (E), S10. – WU KAU TANG. So Lo Pun Back Stream.
60 m (NE), 1619. HONG KONG IS. D’AGUILAR PEAK. 160 m (W), 159. LANTAU IS. SUNSET
PEAK. Wong Lung Hang (East Lung Stream). 100 m (N), 39(4).
Habitat and Substrate: All specimens were collected in forest. 5 specimens were found on rocks, 1 on
soil, and 1 on a concrete wall.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Heilongjiang, Henan,
Hong Kong, Hunan, Jiangsu, Jilin, Shandong, Shanghai, Sichuan, Taiwan, Yunnan, Zhejiang). Korea,
Japan, Laos, Vietnam (Jia and Wang, 1993; Koponen et al., 2000; Redfearn et al., 1996).
Claopodium assurgens (Sull. & Lesq.) Card.
Localities: NEW TERRITORIES. GRASS HILL. Fo Tan Stream. 455 m (SE), 203. – HUNG FA
LENG. San Kwai Tin Stream. 410 m (N), 1146. Muk Min Tau Stream. 35 m (E), 484; 75 m (E), 492;
90 m (E), 495. – KAU LUNG HANG SHAN. 85 m (NW), 1541; 100 m (NW), 1543. – KOWLOON
PEAK. Ho Chung Stream. 115 m (E), 1515. – LAM TSUEN. Tai Om Feng Shui Wood. 110 m, 1905. –
SAI KUNG. Yung Shue O to Pat Tam Chung. 100 m, 1384. Vicinity of High Land Reservoir Dam.
100 m, Xing 9. – TAI LAM. Kat Hing Bridge to Tsing Fai Tong Stream. 135 m (NW), 1061. – TAI LO
SHAN. Kwun Yam Shan. 185 m (N), 952. – TAI MO SHAN. Ng Tung Chai Falls. 395 m (N), 4(5);
700 m (N), 1(6), 1(25). Water-pumping House Stream. 700 m (E), S42. – TAI TO YAN. Lam Tsuen
Stream. 75 m (E), 689; 110 m (E), 719. HONG KONG IS. D’AGUILAR PEAK. 230 m (E), 518; 260
m (E), 183. LANTAU IS. SUNSET PEAK. San Shek Wan Stream. 220 m (S), 2029. Wong Lung
Hang (East Lung Stream). 400 m (N), 35(18). OUTLYING ISLANDS. KAU PEI CHAU. 45 m, 911.
Habitat and Substrate: Twenty-two of 24 specimens were collected in forest, 1 by a road, and 1
unknown. 18 were found on rocks, 3 on trunks, 1 on a tree base, and 2 on rotten twigs.
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Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Hainan, Henan, Hong Kong, Hubei,
Hunan, Jiangsu, Sichuan, Taiwan, Yunnan, Zhejiang). Korea, Japan, India, Laos, Vietnam, Thailand,
Cambodia, Philippines, Indonesia, Australia (He, 2000; Koponen et al., 2000; Redfearn et al., 1996;
Streimann and Curnow, 1989; Tan and Iwatsuki, 1991).
Haplocladium angustifolium (Hampe & C. Muell.) Broth. (Bryohaplocladium angustifolium (Hampe
& C. Muell.) Watan. & Iwats.)
Localities: NEW TERRITORIES. FAN LING. Kei Lak Tsai. 205 m, 41. – SHA TAU KOK. Muk
Min Tau Feng Shui Wood. 10 m (S), 29(5). OUTLYING ISLANDS. KAT O. 20 m, 2534. – YEUNG
CHAU (near Ma Shi Chau). 15 m, 376.
Habitat and Substrate: All 4 specimens were collected in forest. 3 were found on rocks, and 1 on rotten
wood.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hong Kong, Hubei, Hunan, Inner
Mongolia, Jiangsu, Jiangxi, Jilin, Liaoning, Shaanxi, Shandong, Taiwan, Xizang, Yunnan, Zhejiang).
Korea, Japan, Russian Siberia, Pakistan, India, Nepal, Bhutan, Myanmar, Vietnam, Southern Europe,
Africa, North America, Mexico, Jamica, Haiti, Dominica (Buck, 1998; Gangulee, 1969-1980; Koponen
et al., 2000; Redfearn et al., 1996; Tan and Iwatsuki, 1993; Zhao and Cao, 1998).
Haplocladium microphyllum (Hedw.) Broth. (Bryohaplocladium microphyllum (Hedw.) Watan. &
Iwats.)
Localities: OUTLYING ISLANDS. LAMMA IS. So Kwu Wan. 50 m, Lin et al. 1989 (IBSC, KFBG).
– PO TAI IS. 150 m, Lin et al. 1915 (IBSC, KFBG). – TAP MUN CHAU. 30 m, Lin et al. 1850
(IBSC, KFBG).
Habitat and Substrate: All 3 specimens were collected in forest. 2 were found on rocks, and 1 on soil.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Heilongjiang, Henan,
Hong Kong, Hubei, Hunan, Inner Mongolia, Jiangsu, Jiangxi, Jilin, Liaoning, Shaanxi, Shandong,
Shanghai, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Korea, Japan, Russian Siberia, India, Bhutan,
Vietnam, Thailand, Europe, North, Central and South America (He, 2000; Koponen et al., 2000; Li,
Zeng and Su, 1995; Long, 1994; Redfearn et al., 1996; Zhao and Cao, 1998).
Haplohymenium pseudo-triste (C. Muell.) Broth.
Genus and species new to Hong Kong.
Locality: NEW TERRITORIES. KAI KUNG LENG. Ha Che Stream. 310 m (S), 1867.
Habitat and Substrate: On a trunk in forest.
Distribution: China (Anhui, Guangdong, Hainan, Hong Kong, Liaoning, Sichuan, Taiwan, Zhejiang).
Korea, Japan, Sri Lanka, Vietnam, Thailand, Philippines, Australia, New Zealand, South Africa (He,
2000; Redfearn et al., 1996; Tan and Iwatsuki, 1991; Watanabe, 1972; Zhang, 1996).
Herpetineuron toccoae (Sull. & Lesq.) Card.
Localities: NEW TERRITORIES. MA ON SHAN. Ngong Ping. 385 m, 120, 122. – TAI LAM. Kat
Hing Bridge to Tsing Fai Tong. 125 m (NW), 1057. – TAI LO SHAN. Kwun Yam Shan. 100 m (N),
935. – TAI MO SHAN. Ng Tung Chai Falls. 200 m (N), 16(6); 700 m (N), 1(14) (with Bazzania
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Zhang Li: Hong Kong Bryophytes 73
japonica). Shek Kong Stream. 770 m (NW), 2004. Shing Mun Reservoir. 200 m, 748. Yin Ngam
Stream. 410 m (NE), 1895. – TAI TO YAN. Lam Tsuen Stream. 75 m (E), 687. HONG KONG IS.
D’AGUILAR PEAK. 225 m (E), 531; 235 m (E), 513; 245 m (W), 170. LANTAU IS. LANTAU
PEAK. 535 m (S), 1675. – LIN FA SHAN. 535 m (NE), 636. – NEI LAK SHAN. 205 m (NE), 2074. –
SUNSET PEAK. San Shek Wan Stream. 115 m (S), 2015; 125 m (S), 2020; 200 m (S), 31(1); 600 m
(S), 26(6). Wong Lung hang (East Lung Stream). 400 m (N), 35(13). OUTLYING ISLANDS. TAP
MUN CHAU. 50 m, Lin et al. 1836 (IBSC, KFBG).
Habitat and Substrate: All 22 specimens were collected in forest. 21 were found on rocks, and 1 on a
trunk.
Distribution: China (Anhui, Beijing, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hebei,
Heilongjiang, Henan, Hong Kong, Hubei, Hunan, Inner Mongolia, Jiangsu, Jiangxi, Jilin, Liaoning,
Shaanxi, Shandong, Shanghai, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Korea, Japan, Russian
Siberia, Pakistan, India, Nepal, Bhutan, Sri Lanka, Laos, Vietnam, Thailand, Philippines, Indonesia,
New Caledonia, Vanuatu, Australia, Africa, North and South America (Bai, 1996; Fang et al., 1998;
Gangulee, 1969-1980; Higuchi, 1996; Long, 1994; Rao et al., 1997; Redfearn et al., 1996; Streimann
and Curnow, 1989; Watanabe, 1972; Zhao and Cao, 1998).
Thuidium cymbifolium (Dozy & Molk.) Dozy & Molk.
Localities: NEW TERRITORIES. MA ON SHAN. Baffalo Hill. 235 m (NE), 2223. – TAI MO
SHAN. Ng Tung Chai Falls. 400 m (N), Choi 12; 600 m (N), 2(3); 700 m (N), 1(23). Shek Kong
Stream. 320 m (NW), 1971; 520 m (NW), 1985; 785 m (NW), 2287. HONG KONG IS. MOUNT
PARKER. 320 m (NW), 2387. LANTAU IS. LANTAU PEAK. 730 m (S), 1707. – NEI LAK SHAN.
250 m (NE), 2076. – SUNSET PEAK. San Shek Wan Stream. 350 m (S), 2039. Wong Lung Hang
(East Lung Stream). 270 m (N), 2240; 600 m (N), 37(23); 700 m (N), 38(9); 800 m (N), 42(14). Wong
Lung Hang (Main Streams). 455 m (N), 2111.
Habitat and Substrate: Fifteen of 16 specimens were collected in forest, and 1 in grassland. 15 were
found on rocks, and 1 on soil.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Heilongjiang, Henan,
Hong Kong, Hubei, Hunan, Jiangsu, Jiangxi, Liaoning, Shaanxi, Shandong, Shanghai, Sichuan,
Taiwan, Xinjiang, Xizang, Yunnan, Zhejiang). Korea,
Japan, Russia, Pakistan, India, Nepal, Bhutan, Myanmar, Laos, Vietnam, Thailand, Malaysia,
Philippines, Borneo, Indonesia, Papua New Guinea, Australia, New Zealand, Caroline Islands,
Micronesia, Fiji, New Caledonia, Samoa, Vanuatu, Cuba, Venezuela (Fang et al., 1998; Higuchi, 1996;
Ignatov and Afonina, 1992; Koponen et al., 2000; Redfearn et al., 1996; Touw, 1978; Watanabe, 1991;
Zhao and Cao, 1998).
Thuidium glaucinoides Broth.
Localities: NEW TERRITORIES. HUNG FA LENG. Ma Tseuk Leng Stream. 275 m (S), 2141. San
Kwai Tin Stream. 365 m (N), 1157. – KAI KUNG LENG. Ha Che Stream. 295 m (S), 1866. – KAU
LUNG HANG SHAN. 100 m (NW), 1542. – LAM TSUEN. Tai Om Feng Shui Wood. 110 m, 1906. –
MA ON SHAN. Ma On Shan Tsuen Stream. 230 m (SW), 2150; 355 m (SW), 2155; 620 m (SW),
2175. – TAI MO SHAN. Shek Kong Stream. 815 m (NW), 2011. Tai Shing Stream. 300 m (SE), 18(4);
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74 Memoirs of the Hong Kong Natural History Society 26, 2003
500 m (SE), 20(2); 785 m (NE), 1922. Water-pumping House Stream. 700 m (E), S21 (with
Taxiphyllum arcuatum). Yin Ngam Stream. 380 m (NE), 1893. – TAI PO KAU. 180 m (NE), 2332. –
TAI TO YAN. San Lung Wai Stream. 330 m (W), 759. HONG KONG IS. D’AGUILAR PEAK. 270
m (E), 182. LANTAU IS. LANTAU PEAK. 345 m (S), 2215; 520 m (SW), 2260. – POR KAI SHAN.
325 m (NE), 2411. – SUNSET PEAK. San Shek Wan Stream. 500 m (S), 2051.
Habitat and Substrate: All 21 specimens were collected in forest. 16 were found on rocks, 3 on trunks,
1 on a liana, and 1 on rotten wood.
Distribution: China (Anhui, Guangdong, Guangxi, Guizhou, Hong Kong, Jiangxi, Taiwan, Xizang,
Yunnan, Zhejiang). Japan, India, Sri Lanka, Myanmar, Laos, Vietnam, Thailand, Cambodia, Malaysia,
Philippines, Borneo, Indonesia, Papua New Guinea, Vanuatu, Micronesia (Fang et al., 1998; Higuchi,
1996; Redfearn et al., 1996; Watanabe, 1991).
Thuidium pristocalyx (C. Muell.) Jaeg. (T. glaucinum (Mitt.) Bosch & Sande Lac., fide Watanabe
1991)
Localities: NEW TERRITORIES. GRASS HILL. Fo Tan Stream. 495 m (SE), 199. – SAI KUNG.
Nam She Tsim. 400 m (N), 1946. – TAI LO SHAN. Kwun Yam Shan. 130 m (N), 941. – TAI MO
SHAN. Chuen Lung Stream. 570 m (SW), 2423. Ng Tung Chai Falls. 200 m (N), 16(10); 500 m (N),
3(1); 700 m (N), 1(10). Shek Kong Stream. 750 m (NW), 2297. Tai Shing Stream. 200 m (SE), 17(5);
400 m (SE), 19(4); 600 m (SE), 21(24). Water-pumping House Stream. 700 m (E), S61, S62.
LANTAU IS. LIN FA SHAN. 555 m (NE), 635. – SUNSET PEAK. San She Wan Stream. 500 m (S),
34(1); 700 m (SE), 27(17). Wong Lung Hang (East Lung Stream). 320 m (N), 41(10); 400 m (N),
35(11).
Habitat and Substrate: All specimens were collected in forest. 16 were found on rocks, 1 on rocks and a
tree base, and 1 on soil.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hunan,
Jiangsu, Jiangxi, Jilin, Liaoning, Shaanxi, Shandong, Shanghai, Sichuan, Taiwan, Xizang, Yunnan,
Zhejiang). Korea, Japan, Russian Far East, India, Nepal, Bhutan, Myanmar, Laos, Vietnam, Thailand,
Cambodia, Malaysia, Philippines, Borneo, Indonesia, Papua New Guinea, Vanuatu (Fang et al., 1998;
Higuchi, 1996; Ignatov and Afonina, 1992; Jia and Wang, 1993; Koponen et al., 2000; Touw, 1978;
Watanabe, 1991; Zhao and Cao, 1998).
Thuidium recognitum (Hedw.) Lindb. Var. delicatulum (Hedw.) Warnst. (T. delicatulum (Hedw.)
Mitt.)
Locality: HONG KONG (Locality unknown) (Salmon, 1900).
Habitat and Substrate: Unknown.
Distribution: China (Anhui, Fujian, Guizhou, Guangxi, Hainan, Hebei, Hong Kong, Hunan, Jilin,
Liaoning, Shaanxi, Sichuan, Taiwan, Yunnan, Zhejiang). Korea, Japan, Russian Siberia, Europe,
North, Central and South America (Anderson, Crum and Buck, 1990; Bai, 1996; Koponen et al., 2000;
Redfearn et al., 1996; Watanabe, 1972; Zhao and Cao, 1998)
TRACHYPODACEAE
Trachypus humilis Lindb.
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Zhang Li: Hong Kong Bryophytes 75
Localities: NEW TERRITORIES. TAI MO SHAN. Ng Tung Chai Falls. 645 m (N), 1231. HONG
KONG (Locality unknown). So 95704A (IBSC, HKBU).
Habitat and Substrate: One on rocks in forest, and the other unknown.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hunan,
Jiangxi, Shaanxi, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Korea, Japan, India, Bhutan, Sri
Lanka, Vietnam, Thailand, Cambodia, Malaysia, Philippines, Borneo, Indonesia, Papua New Guinea,
Australia, New Caledonia, Hawaii (Fang et al., 1998; Koponen et al., 2000; Redfearn et al., 1996;
Touw, 1978).
Excluded species of mosses
Bryum capillare Hedw. Ssp. Torquescens (De Not.) Kindb. (B. torquescens Bruch ex De Not.)
Zhang and Lin (1997) based on Sullivant and Lesquereux (1978) included this subspecies in their
checklist. The voucher specimen is B. coronatum (Wright s.n., 6-Apr-1854, FH).
Calymperes serratum A. Braun. Ex C. Muell.
This is one of the species which Hance (1872) compiled based on Wilson (1848) and Mitten’s “various
important contributions to bryology inserted in the Proceeding of the Linnean Society”. Unfortunately,
I cannot get any reliable information about the occurrence of this species in Hong Kong from their
works.
Calymperes taitense (Sull.) Mitt.
Although Gao and But (1992) excluded this species from Hong Kong, Gao (1996) still included Hong
Kong within its distribution regions. Zhang and Lin (1997) also listed it in their checklist. The
specimen Gao (1996) cited (Iwatsuki 33, IBSC, NICH) was listed twice under two names C. taitense
and C. molluccense (C. palisotii ssp. Molluccense). After carefully checking the specimen, it should be
C. palisotii ssp. Molluccense, and C. taitense should be excluded.
Campylopodium proscriptum (Hornsch.) Broth.
Zhang and Lin (1997) treated it as a doubtful species occurring in Hong Kong. According to Salmon
(1900), the specimens on which the name was based were Dicranella obscura ( = D. coarctata).
Campylopus comosus (Reinw. & Hornsch.) Bosch. & Sande Lac.
Gao and Crosby (1999) reported this species occurred in Hong Kong based on C. scabridorsus Dix.
Which was synonymized with C. comosus. Previously, Frahm (1987) treated C. scabridorsus as the
same as C. serratus Sande Lac. After consulting the present opinion of Frahm, I accept the latter’s
treatment (1987). Therefore, the occurrence of C. comosus should be rejected.
Didymodon vineralis (Brid.) Zand.
Zhu and So (1996) reported this species occurred in Hong Kong but cited no specimen. Detailed study
of a specimen determined as this name in HKBU (So 95515) revealed that it is Barbula arcuata.
Therefore the occurrence of D. vineralis in Hong Kong should be excluded.
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76 Memoirs of the Hong Kong Natural History Society 26, 2003
Ectropothecium monumentorum (Duby) Jaeg. F. macromorpha Fleisch.
This form was overlooked by Wijk, Margadant and Florschuetz (1959-1969), Crosby, Magill and
Bauer (1992), Crosby and Magill (1994, 1997), and Redfearn et al. (1996). I cannot find any reliable
distinctions between the specimens bearing this name and E. monumentorum cited by Dixon (1933)
(Ah Nin H39A, H39C, BM). Therefore the occurrence of this form in Hong Kong should be excluded.
Ectropothecium nervosum Dix.
This species is treated as the same as E. zollingeri (see explanation under E. zollingeri).
Himantocladium cyclophyllum (C. Muell.) Fleisch.
The authentic specimen on which So and Yip (1995b) based this record belongs to H. plumula (So and
Yip 94830, HKBU, IBSC).
Macromitrium moorcroftii (Hook. & Grev.) Schwaegr.
The specimens studied by Sullivant and Lesquereux (1978) with a question mark consisted of two bags
of specimens and they turn out to be M. nepalense (Wright. S.n., two bags, FH).
Macromitrium sinense Bartr.
Zhu and So (1996) reported this species occurred in Hong Kong and listed no specimen. I have seen
one specimen with this name in HKBU (So 95130), but it is M. heterodictyon. Therefore, M. sinense
should be excluded from the local bryoflora.
Orthomnion loheri Broth. In Ofvers.
So and Zhu (1996c) reported this species and genus new for Hong Kong. Careful study of the specimen
on which this record was based showed that it belongs to an unidentified species of Plagiomnium (So
941018, IBSC, HKBP).
Rhamphidium vaginatum (Dozy & Molk.) Broth.
I have checked the voucher specimen (Bowring s.n., BM No. 00576125), but unfortunately there are
only several poorly developed plants in the bag. My preliminary opinion is that the plants are
Holomitrium vaginatum.
Syrrhopodon albovaginatus Schwaegr.
The occurrence of this species in Hong Kong as well as China was reported by Sullivant and
Lesquereux (1978) only once. In fact, the authentic specimen deposited in FH included two species, i.e.
Calymperes erosum and C. palisotii ssp. Molluccense (Wright s.n., August 14, 1854, FH). Therefore, it
must be excluded from the bryoflora of China as well as of Hong Kong. Gao (1996), Li and Crosby
(2001), Redfearn et al. (1996), and Reese and Lin (1991) failed to discuss its occurrence in China.
Trichostomum inflexum Bruch in F. Muell.
Iwatsuki (1966) was not certain about the identity of this specimen (Wright 78, Feb. 25, 1855, FH).
After carefully checking this material, I think it should be Barbula indica.
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Zhang Li: Hong Kong Bryophytes 77
Vesicularia aperta (Sull.) Ther.
The species was firstly reported by Sullivant and Lesqueuerx (1978). There are two bags of specimens
bearing the above name in FH. After checking the respective specimens, it is clear that they were
misidentified. The first one collected on Mar. 23, 1854 is V. reticulata, and the second collected on
Feb. 1, 1855 included two species, V. montagnei and Callicostella papillata. The occurrence of V.
aperta should therefore be excluded from the local flora. Redfearn et al. (1996) did not include this
name in their catalogue.
Vesicularia dubyana (C. Muell.) Broth.
Dixon (1933) reported this species occurred in Hong Kong. The specimen on which this record based
is V. ferriei (Herklots 296H, BM)
Doubtful species of mosses
Atrichum undulatum (Hedw.) P. Beauv. Var. undulatum
So and Yip (1995a) reported this species new to Hong Kong. According to Lou and Koponen (1986),
no reliable specimens of this taxon have been found in China and all are var. gracilisetum Besch..
Campylopus subulatus Schimp. In Rabenh.
So (1995a) reported that this species occurs in Hong Kong but did not cite any specimens.
Diphyscium fulvifolium Mitt.
This species was first reported for Hong Kong by But and Gao (1991a). Studies of the specimen on
which this report was based showed that it belongs to D. involutum (Ip, Gao and Woo 623, CUHK). In
addition, I have made extensive surveys from various places in Hong Kong, and found only D.
involutum. Probably all the previous reports are mis-identified to D. fulvifolium.
Distichophyllum obtusifolium Ther.
So (1995a) reported that this species occurs in Hong Kong but did not cite any specimens. I have been
unable to borrow the specimen on which this record was based from HKBU for confirmation. The
occurrence of this species in China is also uncertain (Lin and Tan, 1995).
Funaria physcomitrioides Mont. (Entosodon javanensis Dozy & Molk.).
Zhu and So (1996) reported that this species occurs in Hong Kong but did not cite any specimens.
Hyophila rosea Williams
Zhu and So (1996) reported that this species occurs in Hong Kong but did not cite any specimens.
Isopterygium tenerum (Schwaegr.) Mitt.
Zhang and Lin (1997) listed this species as a doubtful record for Hong Kong based on the report of
Salmon (1900). Redfearn et al. (1996) doubted this species occurred in China. The occurrence of this
species in Hong Kong needs further confirmation.
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78 Memoirs of the Hong Kong Natural History Society 26, 2003
Myurella julacea (Schwaegr.) B.S.G.
So (1995a) reported that this species occurs in Hong Kong but did not cite any specimens.
Ochrobryum gardneri (C. Muell.) Lindb.
This species was first reported by Hance (1872) as Schistomitrium gardnerianum Mitt. The known
distribution of this species is Mexico, Costa Rica and Brazil (Sharp, Crum and Eckel 1994), and the
possibility of this species occurring in Hong Kong is low.
Palamocladium nilgheriense (Mont.) C. Muell.
Zhu and So (1996) reported that this species occurs in Hong Kong but did not cite any specimens.
Pelekium velatum Mitt.
But and Gao (1991b) reported this species new to Hong Kong. Zhang and Lin (1997) included it in
their checklist as Lorentzia velata (Mitt.) Buck & Crum. Unfortunately, I have been unable to borrow
the specimen (Ip, Gao and But 721, CUHK) on which this record was based from CUHK for
confirmation.
Philonotis mollis (Dozy & Molk.) Mitt. (Bartramia mollis Dozy & Molk.)
I followed Sullivant and Lesquereux (1978) who treated it as a doubtful species.
Taxiphyllum inundatum Reim.
Zhu and So (1996) reported that this species occurs in Hong Kong but did not cite any specimens.
Trichosteleum mammosum (C. Muell.) Jaeg.
So and Yip (1995a) reported this species occurred in Hong Kong. According to Tan and Jia (1999), all
published records of this species in China should be accepted with doubt unless re-confirmed.
List of accepted liverworts and hornworts
ANEURACEAE
Aneura pinguis (L.) Dum.
Locality: HONG KONG (unknown). 200 m (So and Zhu, 1995).
Habitat and Substrate: On wet soil (So and Zhu, 1995).
Distribution: Cosmopolitan.
Riccardia jackii Schiffn. – So and Zhu (1995c), Zhu and So (1996)
Locality: NEW TERRITORIES. TAI PO (Zhu and So, 1996).
Habitat and Substrate: Near a stream (Zhu and So, 1996).
Distribution: China (Hainan, Hong Kong, Taiwan). Southeast Asia (Piippo, 1990; Zhu and So, 1996).
Riccardia kodame Mizut. & Hatt. – So and Zhu (1996b), Zhu and So (1996)
Locality: NEW TERRITORIES. TAI MO SHAN. 950 m (So and Zhu, 1996b).
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Zhang Li: Hong Kong Bryophytes 79
Habitat and Substrate: On rocks (So and Zhu, 1996b).
Distribution: China (Hong Kong). Japan (Furuki and Mizutani, 1994; So and Zhu, 1996b).
Riccardia latifrons (Lindb.) Lindb.
Localities: NEW TERRITORIES. TAI MO SHAN. Tai Shing Stream. 700 m (SE), 6(26). LANTAU
IS. SUNSET PEAK. Wong Lung Hang (East Lung Stream). 200 m (N), 40(10); 800 m (N), 42(20).
Habitat and Substrate: All 3 specimens were collected in forest. 1 was found on rocks, 1 on soil, and 1
on a tree root.
Distribution: China (Guangdong, Heilongjiang, Hong Kong, Jilin, Taiwan, Xinjiang, Yunnan,
Zhejiang). Japan, Russian Far East, Nepal, Europe, North America (Bai, 1996; Furuki and Higuchi,
1995; Furuki and Mizutani, 1994; Gao and Cao, 2000; Whittemore et al., 1998; Zhu, So and Ye, 1998).
Riccardia multifida (L.) S. Gray
Localities: NEW TERRITORIES. TAI MO SHAN. Ng Tung Chai Falls. 700 m (N), 1(27). Tai Shing
Stream. 400 m (SE), 19(15); 600 m (SE), 21(21). LANTAU IS. SUNSET PEAK. Wong Lung Hang
(East Lung Stream). 500 m (N), 36(18); 600 m (N), 37(20).
Habitat and Substrate: All 5 specimens were collected in forest. 4 were found on rocks, and 1 on soil.
Distribution: China (Fujian, Guangdong, Guizhou, Hainan, Heilongjiang, Hong Kong, Jilin, Taiwan,
Xinjiang, Yunnan, Zhejiang). Korea, Japan, Papua New Guinea, New Zealand, Samoa, Hawaii, Russia,
Europe, East African Islands, Greenland, North America (Constantinova, Potemkin and Schljakov,
1992; Furuki and Mizutani, 1994; Grolle, 1995; Grolle and Piippo, 1984; Grolle and Schultze-Motel,
1972; Hamlin, 1972; Li, Liao and Huang, 1998; Mizutani and Hattori, 1957; Piippo, 1990; Yamada and
Choe, 1997; Zhu, So and Ye, 1998).
Riccardia palmata (Hedw.) Carruth.
Locality: HONG KONG (unknown). Above 500 m (So and Zhu, 1995c).
Habitat and Substrate: On wet rock surfaces (So and Zhu, 1995c).
Distribution: China (Fujian, Hong Kong, Hunan, Jilin, Shaanxi, Shandong, Sichuan, Taiwan, Xinjiang,
Yunnan, Zhejiang). Japan, Russian Siberia, Europe, North America (Furuki and Mizutani, 1994;
Piippo, 1990; Piippo, He and Koponen, 1997; Rao et al., 1997; Whittemore et al., 1998; Zhao and Cao,
1998).
ANTHOCEROTACEAE
Anthoceros punctatus L. – (So, 1996), Zhu and So (1996)
Locality: NEW TERRITORIES. TAI PO. 50 m (So, 1996).
Habitat and Substrate: On soil (So, 1996).
Distribution: China (Anhui, Guangdong, Heilongjiang, Hong Kong, Jiangxi, Jilin, Liaoning, Taiwan,
Yunnan, Zhejiang). Japan, India, Australia, New Caledonia, Russia, Europe, Africa, North and South
America (Asthana and Srivastava, 1991; Constantinova, Potemkin and Schljakov, 1992; Fang et al.,
1998; Furuki and Mizutani, 1994; Gao and Cao, 2000; So, 1996; Piippo, 1990; Scott and Bradshaw,
1985; Zhu, So and Ye, 1998).
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80 Memoirs of the Hong Kong Natural History Society 26, 2003
Folioceros fuciformis (Mont.) Bharadw. – (So 1996), Zhu and So (1996)
Locality: NEW TERRITORIES. TAI MO SHAN. Ng Tung Chai. 400 m (So, 1996)
Habitat and Substrate: On wet rocks and soil banks (So, 1996; Zhu and So, 1996).
Distribution: China (Fujian, Hong Kong, Hunan, Taiwan, Yunnan). Japan, Philippines, Borneo,
Indonesia, Papua New Guinea, Africa, East African Islands (Furuki and Mizutani, 1994; Gao and Cao,
2000; Grolle, 1995; Hasegawa, 1984; Koponen et al., 2000; So, 1996; Tan and Engel, 1986; Wigginton
and Grolle, 1996).
Megaceros flagellaris (Mitt.) Steph. – So (1996), Zhu and So (1996)
Localities: NEW TERRITORIES. TAI MO SHAN. 800 m. – TAI PO KAU. 100 m (So, 1996).
Habitat and Substrate: On wet rocks (So, 1996).
Distribution: China (Fujian, Hong Kong, Hunan, Taiwan, Yunnan). Japan, India, Thailand, Philippines,
Borneo, Indonesia, Papua New Guinea, New Caledonia, Samoa, Society Islands, Hawaii, Africa
(Furuki and Mizutani, 1994; Gao and Cao, 2000; Grolle and Piippo, 1984; Hasegawa, 1984; Koponen
et al., 2000; Menzel, 1988; Wigginton and Grolle, 1996).
Phaeoceros laevis (L.) Prosk. – So (1996)
Localities. NEW TERRITORIES. SHA TAU KOK. Tsiu Hang. 10 m, 45(2). TAI PO. 400 m (So,
1996).
Habitat and Substrate: On soil in agricultural land.
Distribution: China (Fujian, Guangdong, Guangxi, Heilongjiang, Hong Kong, Hebei, Jiangxi, Jilin,
Liaoning, Taiwan, Yunnan, Zhejiang). Korea, Japan, India, Philippines, Borneo, Australia, New
Zealand, Russia, Europe, North America, Brazil (Asthana and Srivastava, 1991; Constantinova,
Potemkin and Schljakov, 1992; Fang et al., 1998; Furuki and Mizutani, 1994; Hamlin, 1972; Jia and
Wang, 1993; Menzel, 1988; Piippo, 1990; So, 1996; Scott and Bradshaw, 1985; Tan and Engel, 1986;
Wu et al., 1992; Yamada and Choe, 1997; Yano, 1995).
AYTONIACEAE
Reboulia hemisphaerica (L.) Raddi – So and Zhu (1996b)
Locality: NEW TERRITORIES. WU KAU TANG. Lai Chi Wo. 100 m (So and Zhu, 1996b).
Habitat and Substrate: On soil (So and Zhu, 1996b).
Distribution: Cosmopolitan.
CALYPOGEIACEAE
Calypogeia arguta Mont. & Nees – So and Zhu (1996b)
Localities: NEW TERRITORIES. SHATIN. Campus of the Chinese University (So and Zhu, 1996b).
– TAI MO SHAN. Ng Tung Chai Falls. 600 m (N), 2(4); 700 m (N), 1(27). Tai Shing Stream. 200 m
(SE), 17(6) (with Heteroscyphus argutus). – VICINITY OF HO PUI RESERVOIR. 260 m (NW),
43(1). HONG KONG IS. VICTORIA PEAK. 400 m (So and Zhu 1996b). LANTAU IS. SUNSET
PEAK. San She Wan Stream. 600 m (S), 26(10). Wong Lung Hang (East Lung Stream). 200 m (N),
40(12).
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Zhang Li: Hong Kong Bryophytes 81
Habitat and Substrate: Five of 6 specimens were collected in forest, and 1 in shrubland. 5 were
collected on soil, and 1 on rocks.
Distribution: China (Anhui, Guangdong, Guangxi, Hainan, Hong Kong, Jiangsu, Jiangxi, Shanghai,
Taiwan, Yunnan, Zhejiang). Korea, Japan, Papua New Guinea, Solomon Islands, Russian Caucasus,
Africa (Constantinova, Potemkin and Schljakov, 1992; Fang et al., 1998; Furuki and Mizutani, 1994;
Gao and Cao, 2000; Grolle, 1995; Grolle and Piippo, 1984; Jia and Wang, 1993; Piippo, 1993; So and
Zhu, 1996b; Wigginton and Grolle, 1996; Yamada and Choe, 1997).
Calypogeia tosana (Steph.) Steph.
Localities: NEW TERRITORIES. SHATIN. Campus of the Chinese University. – TAI PO KAU
(Zhu and So, 2001). HONG KONG IS. VICTORIA PEAK. 400 m (So and Zhu, 1996b).
Habitat and Substrate: On leaves (Zhu and So, 2001).
Distribution: China (Anhui, Fujian, Guangdong, Hainan, Hong Kong, Jiangsu, Jiangxi, Shanghai,
Taiwan, Yunnan, Zhejiang). Korea, Japan (Furuki and Mizutani, 1994; Gao and Cao, 2000; Piippo,
1990; Yamada and Choe, 1997; Zhu and So, 2001; Zhu, So and Ye, 1998).
CEPHALOZIACEAE
Cephalozia hamatiloba Steph. – Zhu and So (1996)
Locality: HONG KONG (Locality unknown).
Habitat and Substrate: On moist soils (Zhu and So, 1996).
Distribution: China (Hainan, Hong Kong, Taiwan). Japan, Sakhalin, probably Southeast Asia (Zhu and
So, 1996).
Cephalozia macounii (Aust.) Aust.
Locality. NEW TERRITORIES. WU KAU TANG. A Ma Wat. 240 m (But and Gao, 1991a).
Habitat and Substrate: On leaves.
Distribution: China (Heilongjiang, Hong Kong, Jiangxi, Jilin, Yunnan). Russia, North America
(Constantinova, Potemkin and Schljakov, 1992; Fang et al., 1998; Gao and Cao, 2000; Piippo, 1990;
Stotler and Crandall-Stotler, 1977).
Cephalozia otaruensis Steph. Var. setiloba (Steph.) Amak. – So and Zhu (1996b)
Locality: NEW TERRITORIES. TAI MO SHAN. 300 m, 700 m (So and Zhu, 1996b).
Habitat and Substrate: On soil (So and Zhu 1996b).
Distribution: China (Hainan, Hong Kong). Insufficient data on its worldwide ranges.
CEPHALOZIELLACEAE
Cephaloziella microphylla (Steph.) Douin – Zhu and So (1996)
Locality: NEW TERRITORIES. TAI MO SHAN. 650 m (Gao and But, 1988).
Habitat and Substrate: On leaves (Gao and But, 1988) and on moist soils (Zhu and So, 1996).
Distribution: China (Guangdong, Hainan, Hong Kong, Hunan, Inner Mongolia, Jiangxi, Yunnan,
Zhejiang). Korea, Japan, Nepal, Bhutan, Thailand (Bai, 1996; Fang et al., 1998; Furuki and Mizutani,
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82 Memoirs of the Hong Kong Natural History Society 26, 2003
1994; Long and Grolle, 1990; Piippo, 1990; Rao et al., 1997; Yamada and Choe, 1997; Zhu, So and
Ye, 1998).
Cylindrocolea tagawae (N. Kitag.) Schust. – So and Zhu (1997), Zhu and So (1996)
Locality: NEW TERRITORIES. WU KAU TANG. 50 m (So and Zhu, 1997).
Habitat and Substrate: On rocks (So and Zhu, 1997).
Distribution: China (Hong Kong). Japan, Thailand, Malaysia, Solomon Islands (Furuki and Mizutani,
1994; Piippo, 1993).
CONOCEPHALACEAE
Conocephalum conicum (L.) Dum.
Localities: NEW TERRITORIES. WU KAU TANG. A Ma Wat. HONG KONG IS. ZOOLOGICAL
and BOTANICAL GARDENS. – POKFULAM RESERVOIR (Gao and But, 1992).
Habitat and Substrate: On rocks and a flower garden wall.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hebei, Heilongjiang, Hong Kong,
Hunan, Inner Mongolia, Jiangsu, Jiangxi, Jilin, Liaoning, Shaanxi, Shandong, Sichuan, Taiwan,
Xinjiang, Xizang, Yunnan, Zhejiang). Korea, Japan, Russian Far East, India, Bhutan, Europe, Africa,
North America (Fang et al., 1998; Furuki and Mizutani, 1994; Jia and Wang, 1993; Koponen et al.,
2000; Li, Liao and Huang, 1998; Piippo, 1990; Piippo, He and Koponen, 1997; Whittemore et al.,
1998; Zhao and Cao, 1998).
Conocephalum japonicum (Thunb.) Grolle
Localities. NEW TERRITORIES. TAI MO SHAN. Kadoorie Farm. Ng Tung Chai Falls (Gao and
But, 1992).
Habitat and Substrate: On soil.
Distribution: China (Fujian, Guangdong, Guangxi, Hong Kong, Hunan, Jiangsu, Jiangxi, Liaoning,
Shaanxi, Shanxi, Taiwan, Xizang, Yunnan, Zhejiang). Korea, Japan, Russian Siberia and Kamchatka,
India, Nepal, Bhutan, Philippines, Hawaii (Fang et al., 1998; Furuki and Higuchi, 1995; Furuki and
Mizutani, 1994; Jia and Wang, 1993; Koponen et al., 2000; Li, Liao and Huang, 1998; Long and
Grolle, 1990; Piippo, 1990; Zhu, So and Ye, 1998).
FOSSOMBROBIACEAE
Fossombronia cristula Aust. – So and Zhu (1996b), Zhu and So (1996)
Locality: NEW TERRITORIES. SAI KUNG. 50 m (So and Zhu, 1996b).
Habitat and Substrate: On soil (So and Zhu, 1996b).
Distribution: China (Hong Kong, Taiwan). Korea, Japan, India, Indonesia, Papua New Guinea, North
America (Piippo, 1990; So and Zhu, 1996b; Yamada and Choe, 1997).
FRULLANIACEAE
Frullania ericoides (Nees) Mont.
Localities: HONG KONG (Locality unknown) (Hattori and Lin, 1985).
Habitat and Substrate: Unknown.
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Zhang Li: Hong Kong Bryophytes 83
Distribution: China (Guangdong, Guangxi, Gansu, Hainan, Hong Kong, Hunan, Jiangsu, Sichuan,
Taiwan, Xizang, Yunnan, Zhejiang). Korea, Japan, Philippines, Borneo, Papua New Guinea, Africa,
East African Islands, Brazil (Chao and Lin, 1992; Furuki and Mizutani, 1994; Grolle, 1995; Grolle and
Piippo, 1984; Long and Grolle, 1990; Piippo, He and Koponen, 1997; Menzel, 1988; Piippo, 1990; Rao
et al., 1997; Tan and Engel, 1986; Wigginton and Grolle, 1996; Yamada and Choe, 1997; Yano, 1995;
Zhu, So and Ye, 1998).
Frullania moniliata (Reinw. Et al.) Mont. – Zhu and So (2001)
Localities: NEW TERRITORIES. TAI MO SHAN (Zhu and So, 2001). LANTAU IS. SUNSET
PEAK. Wong Lung Hang (East Lung Stream). 100 m (N), 39(13). HONG KONG (Locality unknown)
(Hattori and Lin, 1985).
Habitat and Substrate: In forest. On rocks.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Heilongjiang, Hong Kong,
Hubei, Hunan, Jiangxi, Shaanxi, Shandong, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Korea,
Russian Far East, India, Sri Lanka, Indochina (Fang et al., 1998; Piippo, 1990; Piippo, He and
Koponen, 1997; Rao et al., 1997; Zhao and Cao, 1998; Zhu and So, 2001; Zhu, So and Ye, 1998).
Frullania motoyata Steph.
Locality: HONG KONG (Locality unknown) (Hattori and Lin, 1985).
Habitat and Substrate: Unknown.
Distribution: China (Guangdong, Guangxi, Hainan, Hong Kong, Taiwan, Yunnan, Zhejiang). Japan
(Furuki and Mizutani, 1994; Piippo, 1990).
Frullania muscicola Steph.- Zhu and So (1996, 2001)
Localities: NEW TERRITORIES. MA ON SHAN. Kak Hang Tun. 120 m (But and Gao, 1991a). –
TAI PO KAU (Zhu and So, 2001). LANTAU IS. SUNSET PEAK. San She Wan Stream. 100 m (S),
30(4); 200 m (S), 31(7).
Habitat and Substrate: On rocks and a tree base in forest.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Heilongjiang, Hong Kong,
Hubei, Hunan, Inner Mongolia, Jiangsu, Jiangxi, Jilin, Shaanxi, Shandong, Shanxi, Sichuan, Taiwan,
Xizang, Yunnan, Zhejiang). Korea, Japan, Mongolia, Russian Far East and Sakhalin, Pakistan, India
(Bai, 1996; Constantinova, Potemkin and Schljakov, 1992; Chao, Lin and Chan, 1992; Fang et al.,
1998; Furuki and Mizutani, 1994; Piippo, 1990; Piippo, He and Koponen, 1997; Zhu, So and Ye, 1998;
Rao et al., 1997; Zhao and Cao, 1998).
Frullania nepalensis (Spreng.) Lehm. & Lindenb.
Locality: HONG KONG (Locality unknown) (Verdoorn, 1937)
Habitat and Substrate: Unknown.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hong Kong, Hunan, Shaanxi,
Shandong, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Korea, Japan, India, Nepal, Bhutan,
Philippines, Indonesia, Papua New Guinea (Chao and Lin, 1992; Furuki and Mizutani, 1994; Grolle
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84 Memoirs of the Hong Kong Natural History Society 26, 2003
and Piippo, 1984; Long and Grolle, 1990; Piippo, 1990; Piippo, He and Koponen. 1997; Rao et al.,
1997; Tan and Engel, 1986; Yamada and Choe, 1997; Zhao and Cao, 1998; Zhu, So and Ye, 1998).
Frullania tamarisci (L.) Dum.
Localities: NEW TERRITORIES. PAT SIN LENG. Wang Shan Keuk. 250 m. Ha Tsat Muk Kiu. 230
m. – MA ON SHAN. Kei Ling Ha. 60 m (But and Gao, 1991a). LANTAU IS. SUNSET PEAK. Wong
Lung Hang (East Lung Stream). 320 m (NE), 41(16); 500 m (N), 36(21).
Habitat and Substrate: Two specimens were collected in forest. 1 was found on a tree base and 1 on a
trunk.
Distribution: China (Anhui, Guangxi, Hong Kong, Hunan, Jiangsu, Shaanxi, Sichuan, Taiwan, Yunnan,
Zhejiang). Japan, Russian Sakhalin, India, Nepal, Bhutan, Europe, Africa, North America, Brazil
(Chao, Lin and Chan, 1992; Furuki and Mizutani, 1994; Jia and Wang, 1993; Long and Grolle, 1990;
Piippo et al., 1998; Piippo, He and Koponen, 1997; Tian, 1999; Wigginton and Grolle, 1996; Yano,
1995).
Frullania tamsuina Steph.
Locality: HONG KONG (Locality unknown) (Hattori, 1974).
Habitat and Substrate: Unknown.
Distribution: China (Guangdong, Hong Kong, Taiwan). Japan, Myanmar (Furuki and Mizutani, 1994;
Hattori and Lin, 1985; Piippo 1990).
Frullania trichodes Mitt. – So and Zhu (1996b), Zhu and So (1996)
Localities: NEW TERRITORIES. TAI MO SHAN. 800 m (So and Zhu, 1996b). Ng Tung Chai Falls.
395 m (N), 4(1b) (Det. By P.-C. Wu, HKU, IBSC, KFBG, PE); 500 m (N), 3(13). LANTAU IS.
LANTAU PEAK. 800 m (So and Zhu, 1996b).
Habitat and Substrate: Both two specimens were found in forest. 1 on trunks and 1 on rocks.
Distribution: China (Guangdong, Hainan, Hong Kong, Taiwan, Yunnan). Japan, Myanmar, Borneo,
Indonesia, Papua New Guinea, Solomon Islands, Fiji (Chao and Lin, 1992; Furuki and Mizutani, 1994;
Hattori and Piippo, 1986; Menzel, 1988; Piippo, 1990; Piippo et al., 1998; So and Zhu, 1996b).
GEOCALYCACEAE
Chiloscyphus minor (Nees) Engel & Schust. – So and Zhu (1996b), Zhu and So (1996)
Localities: NEW TERRITORIES. TAI PO KAU. 200 m. HONG KONG IS. Old Peak Road. 300 m
(So and Zhu, 1996b).
Habitat and Substrate: On rock surface and on soil (So and Zhu, 1996b).
Distribution: China (Anhui, Guangdong, Heilongjiang, Hong Kong, Hunan, Inner Mongolia, Jiangsu,
Jilin, Liaoning, Shandong, Taiwan, Xinjiang, Xizang, Yunnan, Zhejiang). Japan, Mongolia, Himalayas,
Russia, Europe, North America (Bai, 1996; Li, Liao and Huang, 1998; Piippo, 1990; Piippo et al.,
1998; Rao et al., 1997; Whittemore et al., 1998; Zhao and Cao, 1998).
Chiloscyphus polyanthos (L.) Corda – Zhu and So (1996) as C. polyanthus
Locality: NEW TERRITORIES. TAI MO SHAN. Ng Tung Chai (Zhu and So, 1996).
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Zhang Li: Hong Kong Bryophytes 85
Habitat and Substrate: Unknown.
Diatribution: China (Hebei, Heilongjiang, Jiangsu, Jilin, Liaoning, Sichuan, Taiwan, Xizang, Yunnan,
Zhejiang). Korea, Japan, Russia Far East, Siberia, Himalaya, Europe, North America (Gao and Cao,
2000; Piippo, 1990; Piippo et al., 1998).
Heteroscyphus argutus (Reinw. Et al.) Schiffn. – Zhu and So (2001)
Localities: NEW TERRITORIES. KOWLOON PEAK. 110 m. – TAI MO SHAN. Ng Tung Chai
Falls. 300 m (N), 5(14); 500 m (N), 3(9). Tai Shing Stream. 200 m (SE), 17(9), 17(14); 400 m (SE),
19(8); 500 m (SE), 20(8); 600 m (SE), 21(15); 700 m (SE), 6(14) (with Pallavicinia subciliata); 800 m
(NE), 7912). Tsing Tam Lower Irrigation Reservoir. 140 m (But and Gao 1991a). – TAI PO KAU (Zhu
and So, 2001). LANTAU IS. SUNSET PEAK. San She Wan Stream. 600 m (S), 26(12); 700 m (SE),
27(19). Wong Lung Hang (East Lung Stream). 100 m (N), 39(10); 500 m (N), 36(12); 700 m (N),
38(18).
Habitat and Substrate: All specimens were collected in forest. 10 were found on rocks, 2 on soil, 1 on a
tree base, and 1 on a fern root.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Hainan, Hong Kong, Hunan, Jiangxi,
Shaanxi, Taiwan, Xizang, Yunnan, Zhejiang). Japan, India, Nepal, Bhutan, Vietnam, Thailand,
Malaysia, Philippines, Borneo, Indonesia, Papua New Guinea, Australia, New Zealand, Samoa,
Solomon Islands, Oceania, Mauritius, Canary (Fang et al., 1998; Furuki and Mizutani, 1994; Grolle
and Piippo, 1984; Grolle and Schultze-Motel, 1972; Hamlin, 1972; Long and Grolle, 1990; Menzel,
1988; Piippo, 1985b, 1990, 1993; Rao et al., 1997; Scott and Bradshaw, 1985; Tan and Engel, 1986;
Zhu and So, 2001; Zhu, So and Ye, 1998).
Heteroscyphus coalitus (Hook.) Schiffn. (H. bescherellei (Steph.) Hatt.).
Localities: NEW TERRITORIES. TAI MO SHAN. 650 m (Gao and But, 1988). LANTAU IS.
SUNSET PEAK. Wong Lung Hang (East Lung Stream). 800 m (N), 42(5).
Habitat and Substrate: On soil in forest.
Distribution: China (Anhui, Guizhou, Hainan, Hong Kong, Jiangxi, Taiwan, Yunnan, Zhejiang). Korea,
Japan, Nepal, Bhutan, Indochina, Malaysia, Philippines, Borneo, Indonesia, Papua New Guinea,
Australia, New Zealand, Caroline Islands, Fiji, New Caledonia, Samoa, Solomon Islands (Fang et al.,
1998; Furuki and Mizutani, 1994; Gao and Cao, 2000; Grolle and Piippo, 1984; Grolle and SchultzeMotel, 1972; Hamlin, 1972; Long and Grolle, 1990; Menzel, 1988; Piippo, 1985b, 1993; Tan and
Engel, 1986; Yamada and Choe, 1997; Zhu and So, 2001; Zhu, So and Ye, 1998).
Heteroscyphus planus (Mitt.) Schiffn.
Locality: NEW TERRITORIES. TAI MO SHAN. Tsing Tam Lower Irrigation Reservoir. 140 m (But
and Gao, 1991a).
Habitat and Substrate: On leaves.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Henan, Hong Kong,
Hubei, Hunan, Jiangsu, Jiangxi, Shanghai, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Korea, Japan,
Philippines (Fang et al., 1998; Furuki and Mizutani, 1994; Li, Liao and Huang, 1998; Piippo, He and
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86 Memoirs of the Hong Kong Natural History Society 26, 2003
Koponen, 1997; Piippo, 1990; Rao et al., 1997; Tan and Engel, 1986; Yamada and Choe, 1997; Zhu
and So, 2001; Zhu, So and Ye, 1998).
HAPLOMITRIACEAE
Haplomitrium mnioides (Lindb.) Schust.
Localities: NEW TERRITORIES. TAI MO SHAN. Ng Tung Chai Falls. 150 m (N), 1779; 600 m
(N), 2(20) (with Thuidium pristocalyx); 700 m (NE), 1(4). – TAI PO KAU. 300 m (NE), 2338. HONG
KONG IS. MOUNT PARKER. 310 m (NW), 2385. – ABERDEEN (Gao and But, 1992). LANTAU
IS. LANTAU PEAK. 430 m (S), 1667. – SUNSET PEAK. Wong Lung Hang (Main Streams). 435 m
(N), 2110; 520 m (N), 76; 530 m (N), 73.
Habitat and Substrate: All specimens were collected in forest. 6 were found on rocks, and 3 on soil.
Distribution: China (Fujian, Guangxi, Hong Kong, Taiwan, Yunnan). Japan, Thailand (Furuki and
Mizutani,1994; Gao and Cao, 2000; Kitagawa,1978; Jia and Wang, 1993; Piippo, 1990).
HERBERTACEAE
Herbertus aduncus (Dicks.) Gray – So and Zhu (1996b), Zhu and So (1996)
Localities: NEW TERRITORIES. TAI MO SHAN. 800 m (So and Zhu, 1996b). LANTAU IS.
LANTAU PEAK. 760 m (N), 2464; 785 m (N), 2469.
Habitat and Substrate: On rocks and twigs in forest.
Distribution: China (Anhui, Heilongjiang, Hong Kong, Hunan, Shandong, Taiwan, Xizang, Yunnan,
Zhejiang). Korea, Japan, Russian Far East, Philippines, Borneo, Europe, North America
(Constantinova, Potemkin and Schljakov, 1992; Furuki and Mizutani, 1994; Gao and Cao, 2000;
Menzel,1988; Piippo, 1990; Rao et al., 1997; So and Zhu, 1996b; Tan and Engel, 1986; Yamada and
Choe, 1997; Zhao and Cao, 1998; Zhu, So and Ye, 1998).
JACKIELLACEAE
Jackiella javanica Schiffn. – So and Zhu (1996b)
Locality: NEW TERRITORIES. TAI MO SHAN. 700 m, 800 m (So and Zhu, 1996b).
Habitat and Substrate: On soil (So and Zhu, 1996b).
Distribution: China (Fujian, Hainan, Hong Kong, Taiwan, Yunnan). Japan, India, Bhutan, Sri Lanka,
Thailand, Philippines, Borneo, Indonesia, Papua New Guinea, New Caledonia, Society Islands, Samoa
(Furuki and Mizutani, 1994; Gao and Cao, 2000; Grolle and Piippo, 1984; Long and Grolle, 1990;
Menzel, 1988).
JUBULACEAE
Jubula hutchinsiae (Hook.) Dum. Ssp. Javanica (Steph.) Verd. – So and Zhu (1996b)
Locality: NEW TERRITORIES. TAI MO SHAN. 800 m (So and Zhu, 1996b).
Habitat and Substrate: On moist rock (So and Zhu, 1996b).
Distribution: China (Anhui, Fujian, Guangxi, Hainan, Hong Kong, Taiwan, Yunnan, Zhejiang). Japan,
Russian Far East, India, Sri Lanka, Bhutan, Philippines, Borneo, Indonesia, Papua New Guinea,
Samoa, Solomon Islands, Hawaii, Europe (Constantinova, Potemkin and Schljakov, 1992; Furuki and
Mizutani, 1994; Gao and Cao, 2000; Grolle and Piippo, 1984; Grolle and Schultze-Motel, 1972; Jia
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Zhang Li: Hong Kong Bryophytes 87
and Wang,1993; Lin et al., 1994; Long and Grolle, 1990; Menzel, 1988; Piippo, 1985a, 1990, 1993;
Zhu, So and Ye, 1998).
JUNGERMANNIACEAE
Jungermannia infusca (Mitt.) Steph. – So and Zhu (1996b), Zhu and So (1996)
Locality: NEW TERRITORIES. TAI MO SHAN. 700 m (So and Zhu, 1996b).
Habitat and Substrate: On soil (So and Zhu, 1996b).
Distribution: China (Guangdong, Guangxi, Hong Kong, Taiwan, Yunnan, Zhejiang). Korea, Japan,
Russian Far East (Constantinova, Potemkin and Schljakov, 1992; Furuki and Mizutani 1994; Gao and
Cao, 2000; Yamada and Choe, 1997; Zhu, So and Ye, 1998).
Jungermannia tetragona Lindenb. – So and Zhu (1996b)
Locality: NEW TERRITORIES. TAI PO. 300 m. Sha Lo Tung. 300 m (So and Zhu, 1996b).
Habitat and Substrate: On soil (So and Zhu, 1996b).
Distribution: China (Hong Kong, Jiangxi, Taiwan, Yunnan). Japan, India, Sri Lanka, Bhutan, Thailand,
Malaysia, Philippines, Borneo, Indonesia, Papua New Guinea, Australia, Oceania (Fang et al., 1998;
Furuki and Mizutani, 1994; Gao and Cao, 2000; Grolle and Piippo, 1984; Long and Grolle, 1990;
Menzel, 1988; Vana, 1972).
Jungermannia truncata Nees – So and Zhu (1996b)
Locality: NEW TERRITORIES. TAI MO SHAN. 700 m. Shing Mun Reservoir (Jubilee Reservoir).
100 m (So and Zhu, 1996b).
Habitat and Substrate: On soil (So and Zhu 1996b).
Distribution: China (Guangdong, Guizhou, Hainan, Hong Kong, Taiwan, Yunnan). Korea, Japan,
India, Myanmar, Cambodia, Malaysia, Philippines, Borneo, Indonesia, Papua New Guinea, Australia,
Oceania (Furuki and Mizutani 1994; Grolle and Piippo, 1984; Liu et al., 1999; Menzel, 1988; Piippo,
1990; Tan and Engel, 1986; Vana and Piippo, 1989).
Nardia assamica (Mitt.) Amak. – So and Zhu (1996b)
Locality: NEW TERRITORIES. TAI PO. 500 m (So and Zhu, 1996b).
Habitat and Substrate: On moist soil (So and Zhu, 1996b).
Distribution: China (Anhui, Hong Kong, Hunan, Jiangxi, Taiwan, Yunnan, Zhejiang). Korea, Japan,
India, Russian Caucasus (Fang et al., 1998; Furuki and Mizutani, 1994; Gao and Cao, 2000; Piippo,
1990; Rao et al., 2000; Vana, 1972; Zhu, So and Ye, 1998).
Notoscyphus lutescens (Lehm. & Lindenb.) Mitt. – So and Zhu (1996b)
Locality: NEW TERRITORIES. TAI MO SHAN. 980 m (The altitude in So and Zhu (1996b) must
be wrong because the highest point of Tai Mo Shan is only 957 m). TAI PO. 300 m (So and Zhu,
1996b).
Habitat and Substrate: On soil (So and Zhu, 1996b).
Distribution: China (Guangdong, Hainan, Hong Kong, Taiwan, Yunnan, Zhejiang). Japan, Philippines,
Papua New Guinea, New Caledonia, Samoa, Hawaii, Africa, East African Islands (Furuki and
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88 Memoirs of the Hong Kong Natural History Society 26, 2003
Mizutani, 1994; Gao and Cao, 2000; Grolle, 1995; Grolle and Piippo, 1984; Grolle and SchultzeMotel, 1972; So and Zhu, 1996b; Tan and Engel, 1986; Wigginton and Grolle, 1996).
LEJEUNEACEAE
Acrolejeunea pusilla (Steph.) Grolle & Gradst. – So and Zhu (1996b), Zhu and So (1996)
Localities: NEW TERRITORIES. MA ON SHAN. Tai Lam Wu. 130 m (So and Zhu, 1996b).
HONG KONG IS. Conduit Road. 140 m (N), 22(2).
Habitat and Substrate: On rocks.
Distribution: China (Fujian, Guangdong, Hainan, Hong Kong, Taiwan, Zhejiang). Japan (Furuki and
Mizutani, 1994; So and Zhu, 1996b).
Archilejeunea planiuscula (Mitt.) Steph. – Zhu and So (1996)
Locality: HONG KONG (Locality unknown)
Habitat and Substrate: On wet rocks and tree trunks (Zhu and So, 1996).
Distribution: China (Hong Kong, Yunnan, Zhejiang). Southeast Asia, Australia.
Archilejeunea polymorpha (Sande Lac.) B. Thiers & Gradst. – Zhu and So (1999a)
Localities: NEW TERRITORIES. PAT SIN LENG. 250 m (Zhu and So, 1999a). LANTAU IS.
SUNSET PEAK. Wong Lung Hang (East Lung Stream). 400 m (N), 35(4) (Det by P.-C. Wu as
Spruceanthus polymorphus (Sande Lac.) Verd.).
Habitat and Substrate: On trunks.
Distribution: China (Guangxi, Hainan, Hong Kong, Taiwan, Yunnan). Japan, India, Philippines,
Borneo, Indonesia, Papua New Guinea, Australia, New Caledonia, Somoa, Solomons, Tahiti, Hawaii
(Zhu and So, 1999a).
Cheilolejeunea imbricata (Nee.) Hatt. – So and Zhu (1996a)
Localities. NEW TERRITORIES. PAT SIN LENG. Wang Shan Keuk. 250 m (But and Gao, 1991). –
SAI KUNG. Yung Shue O. – TAI MO SHAN (So and Zhu, 1996a).
Habitat and Substrate: On leaves (But and Gao, 1991) and tree bark as well as rock surface (So and
Zhu, 1996a).
Distribution: China (Anhui, Fujian, Guangdong, Guizhou, Guangxi, Guizhou, Hainan, Hong Kong,
Hunan, Jiangxi, Taiwan, Xizang, Yunnan, Zhejiang). Korea, Japan, India, Nepal, Bhutan, Philippines,
Borneo, Indonesia, Papua New Guinea, Samoa (Furuki and Mizutani, 1994; Grolle and Piippo, 1984;
Jia and Wang, 1993; Koponen et al., 2000; Long and Grolle, 1990; Menzel, 1988; Piippo, 1990; So and
Zhu, 1996a; Yamada and Choe, 1997; Zhu and So, 2001).
Cheilolejeunea intertexta (Lindb.) Steph. – So and Zhu (1996a), Zhu and So (1996)
Locality. NEW TERRITORIES. SAI KUNG (So and Zhu, 1996a).
Habitat and Substrate: On tree-bark and rock in partial shade at low altitude (So and Zhu, 1996a).
Distribution: China (Hong Kong, Taiwan, Yunnan, ). Japan, India, Sri Lanka, Philippines, Borneo,
Indonesia, Papua New Guinea, Micronesia, Somoa, Society Islands, Africa (Furuki and Mizutani,
1994; Gao and Cao, 2000; Grolle and Piippo, 1984; Menzel, 1988; Piippo, 1990; So and Zhu, 1996a).
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Zhang Li: Hong Kong Bryophytes 89
Cheilolejeunea osumiensis (Hatt.) Mizut. – So and Zhu (1996a), Zhu and So (1996, 2001)
Locality. NEW TERRITORIES. TAI MO SHAN. 880 m (Zhu and So, 2001). Peak. 950 m (So and
Zhu, 1996a).
Habitat and Substrate: On rocks, tree bark and decaying logs (So and Zhu, 1996a).
Distribution: China (Gaungdong, Hainan, Hong Kong). Japan (Furuki and Mizutani, 1994).
Cheilolejeunea ryukyuensis Mizut. – So and Zhu (1996a), Zhu and So (1996, 2001)
Localities. NEW TERRITORIES. MA ON SHAN. Tai Lam Wu. – TAI MO SHAN (So and Zhu
1996a). Ng Tung Chai Falls. 200 m (N), 16(18); 300 m (N), 5(17). Tai Shing Stream. 200 m (SE),
17(16) (Det. By P.-C. Wu, HKU, KFBG, IBSC, PE); 500 m (SE), 20(12) (Det. By P.-C. Wu, HKU,
KFBG, IBSC, PE). Tai Wai feng shui wood. 225 m (W), 12(1) (Det. By P.-C. Wu, HKU, KFBG,
IBSC, PE). – TAI PO KAU (Zhu and So, 2001).
Habitat and Substrate: On rotten wood, liana, and trunks.
Distribution: China (Hong Kong). Japan (Furuki and Mizutani 1994).
Cheilolejeunea trifaria (Reinw. Et al.) Mizut. – So and Zhu (1996a), Zhu and So (1996, 2001)
Localities. NEW TERRITORIES. TAI MO SHAN. North of Kadoorie Farm (So and Zhu 1996a).
LANTAU IS. SUNSET PEAK. Wong Lung Hang (East Lung Stream). 100 m (N), 39(7) (Confirmed
by P.-C. Wu, HKU, KFBG, IBSC, PE).
Habitat and Substrate: On rocks in forest.
Distribution: China (Guangxi, Hainan, Hong Kong, Taiwan, Yunnan). Japan, Sri Lanka, Thailand,
Philippines, Borneo, Indonesia, Papua New Guinea, Australia, Society Islands, Africa, Brazil, Central
and South America (Furuki and Mizutani, 1994; Gao and Cao, 2000; Menzel, 1988; Piippo, 1990;
Scott and Bradshaw, 1985; So and Zhu, 1996a; Yano, 1995; Zhu and So, 2001).
Cheilolejeunea ventricosa (Schiffn.) X.-L. He – Zhu and So (1999a)
Locality: NEW TERRITORIES. TAI PO KAU (Zhu and So, 1999a).
Habitat and Substrate: On tree trunk (Zhu and So, 1999a).
Distribution: China (Hong Kong, Yunnan). Malaysia, Singapore, Indonesia, Papua New Guinea (Zhu
and So, 1999a).
Cololejeunea appressa (A. Evans) Bened. – Zhu and So (2001)
Locality: HONG KONG IS. JARDINE MT. (Zhu and So, 2001).
Habitat and Substrate: Unknown.
Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Hong Kong, Taiwan, Xizang, Yunnan,
Zhejiang). Pantropics (Zhu and So, 2001)
Cololejeunea ceratilobula (Chen) Schust. (C. reineckeana Steph., fide Zhu and So 1998). – Zhu and
So (1996 as C. reinecheana, 1998, 2001)
Localities: NEW TERRITORIES. MA ON SHAN. Mau Ping. 200 m. Tai Lam Wu. 130 m – SAI
KUNG. Yung Shue O. 30 m. (But and Gao, 1991a). – TAI MO SHAN. Ng Tung Chai Falls. 500 m
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90 Memoirs of the Hong Kong Natural History Society 26, 2003
(Zhu and So, 1998). – WU KAU TANG. Lai Chi Wo. 50 m, 100m. Wu Kau Tang. 140 m (But and
Gao, 1991a).
Habitat and Substrate: On leaves (But and Gao, 1991a).
Distribution: China (Fujian, Guangdong, Guizhou, Hainan, Hong Kong, Jiangxi, Taiwan, Xizang,
Yunnan). Japan, India, Sri Lanka, Cambodia, Vietnam, Malaysia, Indonesia, Samoa (Grolle and
Schultze-Motel, 1972; Piippo, 1990; Zhu and So, 1998).
Cololejeunea floccosa (Lehm. & Lindenb.) Schiffn.
Localities: NEW TERRITORIES. TAI MO SHAN. 730 m. – WU KAU TANG. Lai Chi Wo. 50 m.
Mui Tsz Lam. 150 m (But and Gao, 1991a).
Habitat and Substrate: On leaves.
Distribution: China (Anhui, Fujian, Guangdong, Hainan, Hong Kong, Taiwan, Xizang, Yunnan).
Japan, Indochina, Malaysia, Philippines, Borneo, Indonesia, Papua New Guinea, New Caledonia, Fiji,
Africa, Madagascar (Furuki and Mizutani, 1994; Gao and Cao, 2000; Grolle, 1995; Grolle and Piippo,
1984; Piippo, 1990; Pocs and Streimann, 1999; Wigginton and Grolle, 1996; Zhu, 1995).
Cololejeunea goebelii (Gott. Ex Schiffn.) Schiffn.- Zhu and So (2001)
Localities: NEW TERRITORIES. KAM SHAN. 240 m. – MA ON SHAN. Kak Hang Tun. 120 m.
Kei Ling Ha. 60 m. Mau Ping. 200 m. – SAI KUNG. Tai Mong Tsai (Ping Tun). 120 m. – TAI MO
SHAN. Ng Tung Chai Falls. 100 m. – TAI PO KAU. 150 m. – WU KAU TANG. A Ma Wat. 240 m.
Lai Chi Wo. 50 m, 100 m. Mui Tsz Lam. 150 m. (But and Gao, 1991). HONG KONG IS. VICTORIA
PEAK (Zhu and So, 2001).
Habitat and Substrate: On leaves.
Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Sichuan, Taiwan,
Xizang, Yunnan, Zhejiang). Korea, Japan, Nepal, Indochina, Malaysia, Philippines, Borneo, Indonesia,
Australia (Furuki and Mizutani, 1994; Mizutani, 1978; Piippo, 1990).
Cololejeunea hasskarliana (Lehm. & Lindenb.) Schiffn. – So and Zhu (1996b), Zhu and So (1996,
2001)
Locality: NEW TERRITORIES. TAI MO SHAN. 700 m, 880 m (So and Zhu, 1996b; Zhu and So,
2001).
Habitat and Substrate: On rotten branches (So and Zhu, 1996b).
Distribution: China (Hainan, Hong Kong, Taiwan, Yunnan). Japan, Bhutan, India, Sri Lanka,
Cambodia, Vietnam, Malaysia, Philippines, Borneo, Indonesia, New Caledonia, Renuion, Seychelles
(Furuki and Mizutani, 1994; Grolle, 1995; Zhu, 1995; Zhu and So, 2001).
Cololejeunea hispidissima (Steph.) Pande et al.
Locality: NEW TERRITORIES. TAI MO SHAN. 650 m (Gao and But, 1988).
Habitat and Substrate: On leaves.
Distribution: China (Guangdong, Hong Kong, Yunnan). Borneo (Menzel, 1988; Piippo, 1990).
Insufficient data on its worldwide ranges.
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Zhang Li: Hong Kong Bryophytes 91
Cololejeunea lanciloba Steph. – Zhu and So (1996, 2001)
Locality: NEW TERRITORIES. WU KAU TANG. 140 m (But and Gao 1991a). HONG KONG IS.
ROBINSON RD. 50 m (Zhu and So, 2001).
Habitat and Substrate: On leaves.
Distribution: China (Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Jiangxi, Taiwan, Yunnan).
Japan, India, Bangladesh, Sri Lanka, Nicobar Islands, Cambodia, Thailand, Philippines, Malaysia,
Borneo, Indonesia, Papua New Guinea, Australia, New Caledonia, Polynesia, Hawaii, Africa (Li, Liao
and Huang, 1998; Piippo, 1990; Pocs, Mizutani and Piippo, 1994; Pocs and Streimann, 1999; Rao et
al., 1997).
Cololejeunea latilobula (Herz.) Tix. (C. himalayensis (Pande & Misra) Schust., fide Grolle 1995) –
Zhu and So (2001)
Localities: NEW TERRITORIES. KAM SHAN. 240 m. – MA ON SHAN. Hang Cho Shui. 120 m.
Kak Hang Tun.120 m. Kei Ling Ha. 60 m. Mau Ping. 200 m. Tai Lam Wu. 130 m. Wong Che. 120 m.
– PAT SIN LENG. Hok Tau Reservoir. 140 m. – SAI KUNG. Tai Mong Tsai (Ping Tun). 120 m. Yung
Shue O. 30 m. – TAI MO SHAN. Ng Tung Chai Falls. 200 m (N), 16(15). Tsing Lam Lower Irrigation
Reservoir. 140 m. – TAI PO KAU. 150 m. – WU KAU TANG. 140 m (But and Gao, 1991). LANTAU
IS. (Zhu and So, 2001).
Habitat and Substrate: On leaves.
Distribution: Distribution: China (Anhui, Guangdong, Guizhou, Hainan, Hong Kong, Hunan, Jiangxi,
Taiwan, Xizang, Yunnan, Zhejiang). Japan, India, Myanmar, Vietnam, Africa, Seychelles (Furuki and
Mizutani, 1994; Grolle, 1995; Piippo, 1990; Pocs, 1992; Wigginton and Grolle, 1996; Zhu and So,
2001; Zhu, So and Ye, 1998).
Cololejeunea ocellata (Horik.) Bened.
Localities: NEW TERRITORIES. SAI KUNG. Tai Lam Wu. 130 m. – SHATIN. Wong Chuk Yeung.
220 m. – TAI PO KAU. 150 m. – WU KAU TANG. Lai Chi Wo. 50 m. Mui Tsz Lam. 150 m. Wu Kau
Tang. 140 m (But and Gao, 1991a).
Habitat and Substrate: On leaves.
Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Hong Kong, Jiangxi, Taiwan, Xizang,
Yunnan, Zhejiang). Japan, Bhutan, Vietnam, Thailand (Furuki and Mizutani, 1994; Jia and Wang,
1993; Zhu, 1995; Zhu and So, 2001).
Cololejeunea ocelloides (Horik.) S. Hatt. – Zhu and So (2001)
Locality: HONG KONG IS. JARDINE MT. (Zhu and So, 2001).
Habitat and Substrate: Unknown.
Distribution: China (Guangdong, Guangxi, Hainan, Hong Kong, Taiwan, Yunnan). Japan, Cambodia,
Vietnam, Thailand, Malaysia, Philippines, Indonesia (Zhu and So, 2001).
Cololejeunea planissima (Mitt.) Abeyw. – So and Zhu (1996b), Zhu and So (2001)
Localities: NEW TERRITORIES. SHA TAU KOK. Luk Keng (Zhu and So, 2001).- TAI MO SHAN.
700 m. HONG KONG IS. VICTORIA PEAK. 500 m (So and Zhu, 1996b).
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92 Memoirs of the Hong Kong Natural History Society 26, 2003
Habitat and Substrate: On decaying logs and on tree trunks (So and Zhu, 1996b).
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hunan,
Jiangxi, Sichuan, Xizang, Taiwan, Yunnan, Zhejiang). Korea, Japan, India, Sri Lanka, Cambodia,
Laos, Malaysia, Borneo, Indonesia, Micronesia, Tanzania (Furuki and Mizutani, 1994; Mizutani, 1984;
Piippo, 1990; Piippo et al., 1998; Rao et al., 1997; Zhu, So and Ye, 1998).
Cololejeunea pseudocristallina Chen & Wu
Localities: NEW TERRITORIES. SAI KUNG. Tai Mong Tsai (Shek Hang). 90 m. Tai Lam Wu, 130
m. – WU KAU TANG. Mui Tsz Lam. 150 m. (But and Gao, 1991a).
Habitat and Substrate: On leaves.
Distribution: Endemic to China (Guangdong, Guizhou, Hong Kong, Yunnan) (Li and Wu, 1992;
Piippo, 1990).
Cololejeunea raduliloba Steph. – So and Zhu (1996b), Zhu and So (2001)
Localities: NEW TERRITORIES. TAI MO SHAN. 700 m. – WU KAU TAN. 50 m. HONG KONG
IS. VICTORIA PEAK. 500 m (So and Zhu, 1996b; Zhu and So, 2001).
Habitat and Substrate: On living leaves and rocks (So and Zhu, 1996b).
Distribution: China (Fujian, Hainan, Hong Kong, Jiangxi, Taiwan, Yunnan, Zhejiang). Korea, Japan,
Nepal, India, Vietnam, Borneo, Indonesia, Australia, Micronesia, Madagascar, Seychelles (Furuki and
Mizutani, 1994; Grolle, 1995; Menzel, 1988; Piippo, 1990; Zhu, 1995; Zhu and So, 2001).
Cololejeunea schmidtii Steph. – Zhu and So (1996 as C. schmdtii, 2001)
Localities: NEW TERRITORIES. MA ON SHAN. Mui Tsz Lam (Meizhilin). WU KAU TANG. 140
m (But and Gao, 1991a; Zhu and So, 2001).
Habitat and Substrate: On leaves.
Distribution: China (Fujian, Hainan, Hong Kong, Taiwan). Japan, Sri Lanka, Laos, Vietnam, Thailand,
Cambodia, Malaysia, Philippines, Indonesia, Papua New Guinea (Pocs, Mizutani and Piippo, 1994;
Zhu, 1995).
Cololejeunea schwabei Herz. – So and Zhu (1996b), Zhu and So (1996)
Localities: NEW TERRITORIES. TAI MO SHAN. Kadoorie Farm. 300 m. – WU KAU TANG. Lai
Chi Wo. 150 m. HONG KONG IS. VICTORIA PEAK. 500 m (So and Zhu, 1996b). LANTAU IS.
SUNSET PEAK. San She Wan Stream. 100 m (S), 30(8).
Habitat and Substrate: On rocks.
Distribution: China (Guangdong, Hainan, Hong Kong, Taiwan). Japan (Furuki and Mizutani, 1994; Liu
et al., 1999; Zhu, 1995).
Cololejeunea spinosa (Horik.) Pande & Misra. – Zhu and So (1996, 2001)
Localities: NEW TERRITORIES. SAI KUNG. Yung Shue O. 30 m. Tai Lam Wu. 130 m. WU KAU
TANG. Lai Chi Wo. 50 m (But and Gao, 1991a).
Habitat and Substrate: On leaves (But and Gao, 1991a).
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Zhang Li: Hong Kong Bryophytes 93
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Jiangxi,
Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Korea, Japan, Nepal, India, Philippines (Fang et al.,
1998; Furuki and Mizutani, 1994; Jia and Wang, 1993; Zhu, 1995; Zhu and So, 2001).
Cololejeunea stylosa (Steph.) Steph. Ex Mizut. – So and Zhu (1996b), Zhu and So (2001)
Locality: NEW TERRITORIES. SAI KUNG. 150 m (So and Zhu, 1996b).
Habitat and Substrate: On living leaves (So and Zhu, 1996b).
Distribution: China (Guangdong, Hong Kong, Taiwan, ). Japan, Laos, Vietnam, Malaysia, Philippines,
Micronesia (Furuki and Mizutani, 1994; Mizutani, 1984; Li, Liao and Huang, 1998).
Drepanolejeunea angustifolia (Mitt.) Grolle – So and Zhu (1996b), Zhu and So (1996)
Locality: NEW TERRITORIES. TAI MO SHAN. 650 m, 800 m, 950 m (So and Zhu, 1996b; Zhu
and So, 1996).
Habitat and Substrate: On rocks and trunks (So and Zhu, 1996b).
Distribution: China (Anhui, Fujian, Guangxi, Guizhou, Hong Kong, Hunan, Jiangxi, Sichuan, Taiwan,
Xizang, Yunnan, Zhejiang). Japan, India, Nepal, Bhutan, Cambodia, Vietnam, Thailand, Philippines,
Borneo, Indonesia, Papua New Guinea, New Caledonia (Fang et al., 1998; Furuki and Mizutani, 1994;
Grolle and Piippo, 1984; Jia and Wang, 1993; Long and Grolle, 1990; Piippo, 1990; Piippo et al.,
1998; Zhu, So and Ye, 1998).
Drepanolejeunea erecta (Steph.) Mizut. – Zhu and So (2001)
Locality: NEW TERRITORIES. TAI MO SHAN. 800 m (Zhu and So, 2001).
Habitat: Unknown.
Distribution: China (Fujian, Guangdong, Guizhou, Hainan, Hong Kong, Jiangxi, Taiwan, Xizang,
Yunnan, Zhejiang). Japan, Bhutan, Nepal, India, Laos, Vietnam (Zhu and So, 2001).
Drepanolejeunea spicata (Steph.) Grolle & R. L. Zhu (Rhaphidolejeunea spicata (Steph.) Grolle) –
But and Gao (1991a) as R. spicata, Zhu and So (2001)
Locality: NEW TERRITORIES. PAT SIN LENG. Ha Tsat Muk Kiu. 230 m (But and Gao, 1991a).
Habitat and Substrate: On leaves.
Distribution: China (Hainan, Hong Kong, Yunnan). Japan, India, Laos, Vietnam, Thailand, Cambodia,
Malaysia, Borneo, Indonesia (Zhu, Long and Yamaguchi, 1999; Zhu and So, 2001).
Drepanolejeunea ternatensis (Gott.) Steph. – So and Zhu (1996b), Zhu and So (1996)
Locality: NEW TERRITORIES. TAI MO SHAN. 800 m (So and Zhu, 1996b).
Habitat and Substrate: On rocks (So and Zhu, 1996b).
Distribution: China (Fujian, Guangdong, Hainan, Hong Kong, Taiwan, Zhejiang). Japan, India,
Philippines, Malaysia, Borneo, Indonesia, Papua New Guinea, Australia, Caroline Is., Fiji, Micronesia,
Samoa, Seychelles (Furuki and Mizutani, 1994; Grolle, 1995; Grolle and Piippo, 1984; Grolle and
Schultze-Motel, 1972; Menzel, 1988; Pocs and Streimann, 1999; Scott and Bradshaw, 1985; Tan and
Engel, 1986; Zhu and So, 2001).
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94 Memoirs of the Hong Kong Natural History Society 26, 2003
Lejeunea anisophylla Mont. (L. catanduana (Steph.) Miller et al.) – Zhu and So (2001)
Localities: NEW TERRITORIES. PAT SIN LENG. Ha Tsat Muk Kiu. 230 m. – SAI KUNG. Tai
Lam Wu. 130 m. Wang Che. 120 m (But and Gao, 1991a). – SHA TAU KOK. Muk Min Tau Feng
Shui Wood. 10 m (S), 29(6) (Det. By P.-C. Wu, HKU, KFBG, IBSC, PE).- TAI PO KAU (Zhu and So,
2001).
Habitat and Substrate: On leaves.
Distribution: China (Anhui, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong,
Jiangxi, Sichuan, Taiwan, Yunnan, Zhejiang). Sri Lanka, Japan, Vietnam, Malaysia, Philippines,
Borneo, Papua New Guinea, Australia, Micronesia, New Caledonia, Samoa, Polynesia, Tahiti, Hawaii,
Tonga (Grolle and Piippo, 1984; Li and Piippo, 1994; Menzel, 1988; Piippo, 1990; Pocs, Piippo and
Mizutani, 1995; Pocs and Streimann, 1999; Zhu and So, 2001).
Lejeunea cocoes Mitt. – So and Zhu (1998)
Locality. NEW TERRITORIES. TAI MO SHAN. 700 m (So and Zhu, 1998).
Habitat and Substrate: HONG KONG (Locality unknown) (So and Zhu, 1998).
Distribution: China (Anhui, Guangxi, Hainan, Hong Kong, Taiwan, Yunnan, Zhejiang). Sri Lanka,
Malaysia, Indonesia, Papua New Guinea (Mizutani, 1963; Piippo, 1990; So and Zhu, 1998; Zhu and
So, 1999a).
Lejeunea curviloba Steph. – Zhu and So (1996, 2001)
Localities: NEW TERRITORIES. TAI MO SHAN. 880 m (Zhu and So, 2001). LANTAU IS.
SUNSET PEAK. San She Wan Stream. 700 m (SE), 27(5).
Habitat and Substrate: On trunks.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hong Kong, Hunan, Jiangxi,
Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Japan, India, Bhutan (Furuki and Mizutani, 1994; Long
and Grolle, 1990; Zhu and So, 2001; Zhu, So and Ye, 1998).
Lejeunea flava (Swaegr.) Nees – Zhu and So (2001)
Localities: NEW TERRITORIES. KOWLOON PEAK. 110 m. – TAI MO SHAN. 500 m (N). 730 m.
– SAI KUNG. Tai Mong Tsai (Ping Tun). 120 m. Tai Mong Tsai (Shek Hang). 90 m (But and Gao,
1991a; Zhu and So, 2001).
Habitat and Substrate: On leaves.
Distribution: China (Fujian, Guangdong, Guizhou, Hainan, Hong Kong, Hunan, Jiangxi, Sichuan,
Taiwan, Xizang, Yunnan, Zhejiang). Korea, Japan, India, Himalaya, Indochina, Philippines, Borneo,
Indonesia, Papua New Guinea, Australia, New Zealand, Samoa, Europe (Ireland), Africa, East African
Islands, North, Central and South America (Fang et al., 1998; Furuki and Mizutani, 1994; Grolle,
1995; Grolle and Piippo, 1984; Grolle and Schultze-Motel, 1972; Hamlin, 1972; Li and Piippo, 1994;
Liu et al., 1999; Mizutani, 1977; Piippo, 1990; Piippo, He and Koponen, 1997; Pocs and Streimann,
1999; Wigginton and Grolle, 1996; Yamada and Choe, 1997; Yano, 1995; Zhu and So, 2000b, 2001;
Zhu, So and Ye, 1998).
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Zhang Li: Hong Kong Bryophytes 95
Lejeunea japonica Mitt.
Localities: NEW TERRITORIES. TAI MO SHAN. Ng Tung Chai Falls. 300 m (N), 5(12) (Det. By
P.-C. Wu, HKU, KFBG, IBSC, PE). Tai Shing Stream. 900 m (NE), 8(25) (Det. By P.-C. Wu, HKU,
KFBG, IBSC, PE).
Habitat and Substrate: On rocks and tree base.
Distribution: China (Guangdong, Guizhou, Hong Kong, Shandong, Taiwan, Zhejiang). Korea, Japan
(Furuki and Mizutani, 1994; Piippo, 1990; Zhao and Cao, 1998, Zhu and So, 2001; Zhu, So and Ye,
1998)
Lejeunea magohukui Mizut. – So and Zhu (1998), Zhu and So (2001)
Localities: NEW TERRITORIES. TAI PO KAU. 200 m (So and Zhu, 1998). – WU KAU TANG. 50
m (Zhu and So, 2001).
Habitat and Substrate: Unknown (So and Zhu, 1998).
Distribution: China (Anhui, Fujian, Hong Kong, Yunnan). Japan (Furuki and Mizutani, 1994; Gao and
Cao, 2000; So and Zhu, 1998; Zhu and So, 2001).
Lejeunea otiana Hatt. – Zhu and So (1996)
Locality: NEW TERRITORIES. SHA TAU KOK. Muk Min Tau Feng Shui Wood. 10 m (S), 29(3).
Habitat and Substrate: In forest. On tree base.
Distribution: China (Hong Kong). Japan (Furuki and Mizutani, 1994; Zhu and So, 1996)
Lejeunea parva (Hatt.) Mizut. – So and Zhu (1998), Zhu and So (1996, 2001)
Localities. NEW TERRITORIES. TAI MO SHAN. 880 m (So and Zhu, 1998; Zhu and So, 2001). Ng
Tung Chai Falls. 600 m (N), 2(12) (Confirmed by P.-C. Wu, HKU, KFBG, IBSC, PE). Tai Shing
Stream. 700 m (SE), 6(28) (Det. By P.-C. Wu, HKU, KFBG, IBSC, PE); 900 m (NE), 8(10) (Det. By
P.-C. Wu, HKU, KFBG, IBSC, PE).
Habitat and Substrate: All in forest. Two on rocks and 1 on trunks.
Distribution: China (Anhui, Fujian, Guangdong, Hainan, Hong Kong, Hunan, Jiangxi, Liaoning,
Shandong, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Korea, Japan, Samoa (Furuki and Mizutani,
1994; Grolle and Schultze-Motel, 1972; Piippo, 1990; Piippo et al., 1998; Piippo, He and Koponen,
1997).
Lejeunea tuberculosa Steph. – So and Zhu (1998).
Locality. NEW TERRITORIES. TAI MO SHAN. 800 m (So and Zhu, 1998).
Habitat and Substrate: Unknown (So and Zhu, 1998).
Distribution: China (Guangdong, Hainan, Hong Kong, Yunnan). India, Nepal, Bhutan, Philippines,
Africa (Gao and Cao, 2000; So and Zhu,1998; Zhu and So, 2001).
Lejeunea ulicina (Tayl.) Gott. (L. punctiformis Taylor, Microlejeunea punctiformis (Tayl.) Steph.) –
Zhu and So (1996), Zhu and So (2001) as L. punctiformis
Localities: NEW TERRITORIES. TAI MO SHAN. 730 m (But and Gao, 1991a). – WU KAU
TANG. 50 m (Zhu and So, 2001).
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96 Memoirs of the Hong Kong Natural History Society 26, 2003
Habitat and Substrate: On leaves (But and Gao, 1991a).
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Jiangxi,
Taiwan, Xizang, Yunnan, Zhejiang). Korea, Japan, Bhutan, Nepal, India, Sri Lanka, Vietnam,
Thailand, Europe, Africa, North America (Furuki and Mizutani, 1994; Mizutani, 1978; Piippo, 1990;
Stotler and Crandall-Stotler, 1977; Yamada and Choe, 1997; Zhu and So, 2001; Zhu, So and Ye, 1998).
Leptolejeunea elliptica (Lehm. & Lindenb.) Schiffn. – Zhu and So (2001)
Localities: NEW TERRITORIES. MA ON SHAN. Kak Hang Tun. 120 m. Mau Ping. 200 m. – PAT
SIN LENG. Ha Tsat Muk Kiu. 230 m. – SAI KUNG. Tai Mong Tsai (Shek Hang). 90 m. Yung Shue
O. 30 m. – TAI MO SHAN. Ng Tung Chai Falls. 200 m (N), 16(14) (Det. By P.-C. Wu, HKU, KFBG,
IBSC, PE), 16(16); 300 m. – TAI PO KAU. 150 m. – WU KAU TANG. Lai Chi Wo. 50 m, 100 m. Wu
Kau Tang. 140 m (But and Gao, 1991).
Habitat and Substrate: On leaves.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hunan,
Jiangxi, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Japan, India, Nepal, Bhutan, Philippines,
Borneo, New Zealand, East African Islands, North America, Brazil (Fang et al., 1998; Furuki and
Mizutani, 1994; Grolle, 1995; Hamlin, 1972; Jia and Wang, 1993; Lin et al., 1994; Long and Grolle,
1990; Menzel, 1988; Stotler and Crandall-Stotler, 1977; Tan and Engel, 1986; Yano, 1995; Zhu, So
and Ye, 1998).
Leucolejeunea xanthocarpa (Lehm. & Lindenb.) Evans – So and Zhu (1996b), Zhu and So (1996,
1999)
Locality: NEW TERRITORIES. TAI MO SHAN. 880 m (So and Zhu, 1996b; Zhu and So, 1999).
Habitat and Substrate: On tree-trunks (So and Zhu,1996b; Zhu and So, 1999).
Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Jiangxi, Sichuan,
Taiwan, Zhejiang). Japan, Philippines, Borneo, Papua New Guinea, Australia, Africa, North America,
Brazil (Furuki and Mizutani, 1994; Grolle and Piippo, 1984; Menzel, 1988; Scott and Bradshaw, 1985;
Stotler and Crandall-Stotler, 1977; Yano, 1995; Zhu and So, 1999b; Zhu, So and Ye, 1998).
Lopholejeunea applanata (Reinw. Et al.) Steph.
Localities: NEW TERRITORIES. PAT SIN LENG. Hok Tau Reservoir. 140 m (But and Gao,
1991a). TAI MO SHAN. Tai Shing Stream, 900 m (NE), 8(8).
Habitat and Substrate: On soil.
Distribution: China (Guangdong, Hong Kong, Shaanxi, Taiwan, Xizang, Zhejiang). Thailand,
Philippines, Borneo, Indonesia, Papua New Guinea (Grolle and Piippo, 1984; Menzel, 1988; Mizutani,
1979; Piippo, 1990; Zhu, So and Ye, 1998).
Lopholejeunea eulopha (Tayl.) Schiffn. – So and Zhu (1996b), Zhu and So (1996)
Locality: NEW TERRITORIES. TAI PO. 400 m (So and Zhu, 1996b).
Habitat and Substrate: On moist rock surface (So and Zhu, 1996b).
Distribution: China (Guangdong, Hainan, Hong Kong, Taiwan). Japan, Philippines, Borneo, Indonesia,
Papua New Guinea, Australia, Samoa, Africa, Central and South America (Furuki and Mizutani, 1994;
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Zhang Li: Hong Kong Bryophytes 97
Grolle and Piippo, 1984; Grolle and Schultze-Motel, 1972; Mizutani, 1979; Piippo, 1990; Tan and
Engel, 1986; Zhu and So, 2001).
Lopholejeunea nicobarica Steph. – So and Zhu (1996b), Zhu and So (1996)
Locality: NEW TERRITORIES. TAI MO SHAN. Ng Tung Chai Falls. 400 m (So and Zhu, 1996b).
Habitat and Substrate: In splash zone (So and Zhu, 1996b).
Distribution: China (Hong Kong, Taiwan). Japan, Thailand, Philippines, Nicobar Islands (Furuki and
Mizutani, 1994; Mizutani, 1985; Piippo, 1990,).
Lopholejeunea subfusca (Nees) Steph.
Localities: NEW TERRITORIES. MA ON SHAN. Mau Ping. 220 m (But and Gao, 1991a). TAI MO
SHAN. Tai Shing Stream. 300 m (SE), 18(11); 500 m (SE), 20(5).
Habitat and Substrate: On liana and on rocks.
Distribution: China (Anhui, Fujian, Hainan, Hong Kong, Jiangxi, Taiwan, Xizang, Yunnan, Zhejiang).
Korea, Japan, Philippines, Borneo, Indonesia, Papua New Guinea, Africa, East African Islands, North
America, Brazil (Furuki and Mizutani, 1994; Gao and Cao, 2000; Grolle, 1995; Grolle and Piippo,
1984; Menzel, 1988; Piippo, 1990; Pocs and Streimann, 1999; Stotler and Crandall-Stotler, 1977; Tan
and Engel, 1986; Wigginton and Grolle, 1996; Yamada and Choe, 1997; Yano, 1995; Zhu and So,
2001; Zhu, So and Ye, 1998).
Ptychanthus striatus (Lehm & Lindenb.) Nees.
Genus and species new to Hong Kong.
Locality: LANTAU IS. SUNSET PEAK. Wong Lung Hang (East Lung Stream). 700 m (N), 38(7).
Habitat and Substrate: On rocks in forest.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hunan,
Jiangxi, Shaanxi, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Japan, India, Nepal, Bhutan,
Philippines, Borneo, Indonesia, Papua New Guinea, Australia, Pacific Islands, Africa (Furuki and
Mizutani, 1994; Grolle and Piippo, 1984; Long and Grolle, 1990; Menzel, 1988; Piippo, 1990; Piippo,
He and Koponen, 1997; Rao et al., 1997; Zhu, So and Ye, 1998).
Spruceanthus semirepandus (Nees) Verd. – So and Zhu (1996b), Zhu and So (1996)
Locality: NEW TERRITORIES. TAI MO SHAN. 800 m (So and Zhu, 1996b).
Habitat and Substrate: On rocks (So and Zhu, 1996b).
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hunan,
Jiangxi, Sichuan, Taiwan, Xinjiang, Xizang, Yunnan, Zhejiang). Japan, Philippines, Borneo (Furuki
and Mizutani, 1994; Koponen et al., 2000; Jia and Wang, 1993; Menzel, 1988; Piippo, 1990; Tan and
Engel, 1986; Zhu and So, 2001).
Stenolejeunea apiculata (Sande Lac.) R. M. Schust. (Drepanolejeunea formosana Horik.) – Gao and
But (1988) as D. formosana, Zhu and So (2001)
Locality: NEW TERRITORIES. TAI MO SHAN. 650 m (Gao and But, 1988).
Habitat and Substrate: On leaves (Gao and But, 1988)
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98 Memoirs of the Hong Kong Natural History Society 26, 2003
Distribution: China (Guangdong, Hainan, Hong Kong, Taiwan). Japan, Sri Lanka, Cambodia, Vietnam,
Malaysia, Philippines, Indonesia, New Caledonia, Australia, New Zealand (Piippo, 1990; Tan and
Engel, 1986).
Thysananthus flavescens (Hatt.) Gradst.
Locality: NEW TERRITORIES. TAI MO SHAN. Ng Tung Chai. 400 m (Zhu and So, 1996).
Habitat and Substrate: On soil (Zhu and So, 1996).
Distribution: China (Hong Kong). Japan (Zhu and So, 1996).
Trocholejeunea sandvicensis (Gott.) Mizut. – So and Zhu (1996b)
Localities: NEW TERRITORIES. MA ON SHAN. Tai Lam Wu. 130 m (So and Zhu, 1996b).
LANTAU IS. SUNSET PEAK. San She Wan Stream. 500 m (S), 34(15).
Habitat and Substrate: On trunks in forest.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Heilongjiang, Hong Kong,
Hunan, Inner Mongolia, Jiangsu, Jiangxi, Jilin, Liaoning, Shaanxi, Shandong, Sichuan, Taiwan,
Xizang, Yunnan, Zhejiang). Korea, Japan, Russian Far East, India, Nepal, Bhutan, Vietnam, Borneo,
Samoa, Society Islands, Hawaii (Bai, 1996; Fang et al., 1998; Furuki and Mizutani, 1994; Grolle and
Schultze-Motel, 1972; Long and Grolle, 1990; Menzel, 1988; Piippo, 1990; Piippo, He and Koponen,
1997; Rao et al., 1997; Zhao and Cao, 1998; Zhu, So and Ye, 1998).
Tuyamaella molischii (Schiffn.) Hatt. – Zhu and So (1996, 2000a, 2001)
Locality: NEW TERRITORIES. TAI MO SHAN. 880 m, 950 m (Zhu and So, 2001).
Habitat and Substrate: Unknown.
Distribution: China (Fujian, Guangdong, Hainan, Hong Kong, Jiangxi, Zhejiang). Japan, Vietnam,
Malaysia (Furuki and Mizutani, 1994; Zhu and So, 2000a).
LEPICOLEACEAE
Mastigophora diclados (Brid.) Nees – So and Zhu (1996b)
Localities: NEW TERRITORIES. MA ON SHAN. Tiu Shau Ngam. 565 m (NW), 119. – TAI MO
SHAN. 800 m (So and Zhu 1996b). Tai Shing Stream. 780 m (NE), 1440. Shek Kong Stream. 770 m
(NW), 2289. LANTAU IS. LANTAU PEAK. 705 m (N), 2455. – LIN FA SHAN. 645 m (E), 1458. –
SUNSET PEAK. San Shek Wan Stream. 690 m (S), 2062. Wong Lung Hang (Main Streams). 590 m
(N), 2120; 595 m (N), 68; 610 m (N), 56, 57.
Habitat and Substrate: Nine of 10 specimens were collected in forest, and 1 in grassland. 8 were found
on rocks, 1 on a rotten trunk, and 1 on soil.
Distribution: China (Guangdong, Hainan, Hong Kong, Taiwan). Japan, Philippines, Borneo, Indonesia,
Papua New Guinea, Australia, Pacific Islands, Samoa, Solomon Islands, East African Islands (Furuki
and Mizutani, 1994; Grolle, 1995; Grolle and Piippo, 1984; Grolle and Schultze-Motel, 1972;
Kitagawa, 1978; Menzel, 1988; Piippo, 1993; Scott and Bradshaw, 1985; Tan and Engel, 1986).
LEPIDOZIACEAE
Bazzania fauriana (Steph.) Hatt. – So and Zhu (1996b), Zhu and So (1996)
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Zhang Li: Hong Kong Bryophytes 99
Localities: NEW TERRITORIES. TAI MO SHAN. 800 m. Ng Tung Chai. 400 m (So and Zhu,
1996b). LANTAU IS. SUNSET PEAK. San She Wan Stream. 700 m (SE), 27(4).
Habitat and Substrate: On rocks in forest.
Distribution: China (Anhui, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Yunnan, Taiwan,
Zhejiang). Japan (Furuki and Mizutani, 1994; Li, Liao and Huang, 1998; Piippo, 1990; Zhu, So and
Ye, 1998).
Bazzania japonica (Sande Lac.) Lindb. – So and Zhu (1996b)
Localities: NEW TERRITORIES. TAI MO SHAN. 700 m (So and Zhu, 1996b). Tai Shing Stream.
800 m (NE), 7(5). LANTAU IS. SUNSET PEAK. San She Wan Stream. 500 m (S), 34(7). Wong Lung
Hang (East Lung Stream). 800 m (N), 42(6).
Habitat and Substrate: On rocks in forest.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hunan,
Jiangxi, Taiwan, Yunnan, Zhejiang). Japan, Russian Far East, Vietnam, Thailand, Philippines,
Indonesia, Samoa (Constantinova, Potemkin and Schljakov, 1992; Fang et al., 1998; Furuki and
Mizutani, 1994; Grolle and Schultze-Motel, 1972, Mizutani and Chang, 1986; Piippo, 1990; Rao et al.,
1997; Tan and Engel, 1986; Zhu, So and Ye, 1998).
Bazzania sikkimensis (Steph.) Herz. – So and Zhu (1996b), Zhu and So (1996)
Locality: NEW TERRITORIES. TAI MO SHAN. 800 m (So and Zhu 1996b).
Habitat and Substrate: On rocks (So and Zhu 1996b).
Distribution: China (Guangxi, Hainan, Hong Kong, Taiwan, Xizang, Yunnan, Zhejiang). Japan, India,
Nepal, Bhutan, Thailand, Philippines (Furuki and Mizutani, 1994; Gao and Cao, 2000; Long and
Grolle, 1990; Piippo, 1990; Zhu, So and Ye, 1998).
Bazzania tricrenata (Wahl.) Trev.
Locality: NEW TERRITORIES. KOWLOON PEAK. 110 m (But and Gao 1991).
Habitat and Substrate: On leaves.
Distribution: China (Hong Kong, Inner Mongolia, Jiangxi, Jilin, Sichuan, Taiwan, Yunnan, Xizang).
Korea, Japan, South India, Nepal, Bhutan, Europe, North America, Guatemala (Bai, 1996; Fang et al.,
1998; Furuki and Mizutani, 1994; Long and Grolle, 1990; Mizutani and Chang, 1986; Piippo, 1990;
Piippo, He and Koponen, 1997; Yamada and Choe, 1997).
Bazzania tridens (Reinw. Et al.) Trev.
Localities: NEW TERRITORIES. TAI MO SHAN. Ng Tung Chai Falls. 500 m (N), 3(11); 600 m
(N), 2(23), 2(25); 700 m (N), 1(3), 1(5), 1(11). Tai Shing Stream. 200 m (SE), 17(3); 300 m (SE),
18(9); 500 m (SE), 20(3); 600 m (SE), 21(3); 700 m (SE), 6(18). Tsing Tam Lower Irrigation
Reservoir. 140 m (But and Gao 1991). HONG KONG IS. VICTORIA PEAK. 540 m (E), 15(3).
LANTAU IS. SUNSET PEAK. San She Wan Stream. 700 m (SE), 27(2). Wong Lung Hang (East
Lung Stream). 100 m (N), 39(15); 200 m (N), 40(3); 320 m (N), 41(4); 400 m (N), 35(10); 500 m (N),
36(10); 600 m (N), 37(10); 700 m (N), 38(4), 38(16); 800 m (N), 42(3).
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100 Memoirs of the Hong Kong Natural History Society 26, 2003
Habitat and Substrate: Twenty-one of 22 specimens were collected in forest, and 1 in grassland. 16
were found on rocks, 4 on soil, 2 on trunks.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hubei,
Hunan, Inner Mongolia, Jiangsu, Jiangxi, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Korea, Japan,
India, Nepal, Bhutan, Philippines, Borneo, Papua New Guinea, Samoa (Bai, 1996; Fang et al., 1998;
Furuki and Mizutani,1994; Grolle and Piippo; 1984; Long and Grolle, 1990; Menzel, 1988; Piippo,
1990; Rao et al., 1997; Tan and Engel, 1986; Yamada and Choe, 1997; Zhu, So and Ye, 1998).
Kurzia gonyotricha (Sande Lac.) Grolle – So and Zhu (1996b)
Locality: NEW TERRITORIES. TAI MO SHAN. 700 m (So and Zhu, 1996b).
Habitat and Substrate: On soil (So and Zhu, 1996b).
Distribution: China (Fujian, Guangdong, Guangxi, Hainan, Hong Kong, Hunan, Taiwan, Yunnan).
Japan, Philippines, Malaysia, Borneo, Indonesia, Papua New Guinea (Furuki and Mizutani, 1994; Gao
and Cao, 2000; Menzel, 1988; Piippo, 1985c, 1990; Rao et al., 1997).
Lepidozia fauriana Steph.
Locality: NEW TERRITORIES. TAI MO SHAN. 650 m (Gao and But, 1988).
Habitat and Substrate: Unknown.
Distribution: China (Fujian, Guangdong, Guangxi, Hainan, Hong Kong, Hunan, Taiwan, Yunnan,
Zhejiang). Korea, Japan, Philippines, Borneo (Furuki and Mizutani, 1994; Menzel, 1988; Piippo, 1990;
Rao et al., 1997; Yamada and Choe, 1997; Zhu, So and Ye, 1998).
Lepidozia vitrea Steph. – So and Zhu (1996b)
Localities: NEW TERRITORIES. TAI MO SHAN. Ng Tung Chai Falls. 600 m (N), 2(22); 700 m
(N), 1(1). Tai Shing Stream. 400 m (SE), 19(10); 800 m (NE), 7(8). Water-pumping House Stream.
700 m (E), S24. LANTAU IS. SUNSET PEAK. Wong Lung Hang (East Lung Stream). 200 m (N),
40(13); 320 m (N), 41(6).
Habitat and Substrate: All specimens were collected in forest. 4 were found on soil, and 3 on rocks.
Distribution: China (Guangdong, Hong Kong, Jiangxi, Taiwan, Yunnan, Zhejiang). Korea, Japan, East
African Islands (Furuki and Mizutani, 1994; Gao and Cao, 2000; Grolle, 1995; Yamada and Choe,
1997).
LOPHOZIACEAE
Chandonanthus birmensis Steph. – So and Zhu (1996b)
Localities: NEW TERRITORIES. TAI MO SHAN. Tai Shing Stream. 600 m (SE), 21(7); 780 m
(NE), 25; 790 m (NE), 1433; 830 m (NE), 17; 835 m (NE), 1418; 885 m (NE), 11; 900 m (NE), 1125.
LANTAU IS. LANTAU PEAK. 800 m (N), 2471; 840 m (N), 2474. – SUNSET PEAK. Wong Lung
Hang (East Lung Stream), 800 m (N), 2312.
Habitat and Substrate: There were 10 specimens studied, of which 9 were collected in forest, and 1 in
grassland. 9 were found on rocks, and 1 on soil.
Distribution: China (Fujian, Guangdong, Guangxi, Hong Kong, Jiangxi, Taiwan, Xizang, Yunnan).
Korea, Japan, Russian Far East, Africa, Madagascar (Constantinova, Potemkin and Schljakov, 1992;
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Zhang Li: Hong Kong Bryophytes 101
Fang et al., 1998; Furuki and Mizutani, 1994; Grolle, 1995; Kitagawa, 1978; Piippo, 1990; Yamada
and Choe, 1997; Wigginton and Grolle, 1996; Zhu, So and Ye, 1998).
MAKINOACEAE
Makinoa crispata (Steph.) Miyak.
Localities: NEW TERRITORIES. TAI MO SHAN. Ng Tung Chai Falls. 700 m (N), 1(8), 1(29). Tai
Shing Stream. 700 m (SE), 6(17).
Habitat and Substrate: On rocks in forest.
Distribution: China (Anhui, Guangdong, Guangxi, Hong Kong, Hunan, Liaoning, Taiwan, Yunnan,
Zhejiang). Korea, Japan, Philippines, Indonesia, Papua New Guinea (Furuki and Mizutani, 1994; Gao
and Cao, 2000; Grolle and Piippo, 1984; Jia and Wang, 1993; Koponen et al., 2000; Yamada and
Choe, 1997).
MARCHANTIACEAE
Marchantia emarginata Reinw. Et al. ssp. Tosana (Steph.) Bischl.
Locality: HONG KONG (Locality unknown) (So and Zhu, 1995c).
Habitat and Substrate: Unknown.
Distribution: China (Guangdong, Guangxi, Guizhou, Hong Kong, Jiangxi, Sichuan, Taiwan, Yunnan,
Zhejiang). Korea, Japan, Thailand, Vietnam (Bischler-Causse, 1989; Fang et al., 1998; Furuki and
Mizutani, 1994; Yamada and Choe, 1997; Zhu, So and Ye, 1998).
Marchantia polymorpha L.
Locality: HONG KONG (Locality unknown) (So and Zhu, 1995c).
Habitat and Substrate: Unknown.
Distribution: Nearly cosmopolitan. (Bischler-Causse, 1989).
METZGERIACEAE
Metzgeria conjugata Lindb.
Localities: NEW TERRITORIES. MA ON SHAN. Wang Che. 120 m. – PAT SIN LENG. Ha Tsat
Muk Kiu. 230 m. – TAI MO SHAN. 730 m. – WU KAU TANG. A Ma Wat. 240 m (But and Gao,
1991a).
Habitat and Substrate: On leaves (But and Gao,1991a).
Distribution: China (Fujian, Guangdong, Guangxi, Hainan, Heilongjiang, Hong Kong, Inner Mongolia,
Jiangsu, Jiangxi, Jilin, Shandong, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Korea, Japan, Russian
Far East, Nepal, Papua New Guinea, Australia, Samoa, East African Islands, Europe, North and South
America (Bai, 1996; Fang et al., 1998; Furuki and Mizutani, 1994; Grolle, 1995; Grolle and Piippo,
1984; Grolle and Schultze-Motel, 1972; Jia and Wang, 1993; Lin et al., 1994; Piippo, 1990; Piippo, He
and Koponen, 1997; Scott and Bradshaw, 1985; Yamada and Choe, 1997; Zhao and Cao, 1998; Zhu,
So and Ye, 1998).
Metzgeria consanguinea Schiffn. – Zhu and So (2001)
Locality: NEW TERRITORIES. TAI MO SHAN. 880 m (Zhu and So, 2001).
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102 Memoirs of the Hong Kong Natural History Society 26, 2003
Habitat: Unknown.
Distribution: China (Hong Kong, Zhejiang). Pantropics (cf. Zhu and So (2001)).
Metzgeria furcata (L.) Dum.
Localities: NEW TERRITORIES. TAI MO SHAN. Tai Shing Stream. 700 m (SE), 6(23); 900 m
(NE), 8(14). LANTAU IS. SUNSET PEAK. San She Wan Stream. 500 m (S), 34(10); 700 m (SE),
27(9), 27(20). Wong Lung Hang (East Lung Stream). 500 m (N), 36(22). HONG KONG (Locality
unknown). 970 m (So and Zhu, 1995) (This altitude probably was wrong because the highest point of
Hong Kong is 957 m).
Habitat and Substrate: All 6 specimens were collected in forest. 5 were found on trunks, and 1 on
rocks.
Distribution: China (Anhui, Guangdong, Guangxi, Heilongjiang, Hong Kong, Hunan, Inner Mongolia,
Jilin, Shaanxi, Taiwan, Yunnan, Zhejiang). Korea, Japan, Mongolia, Russian Far East, Philippines,
Australia, New Zealand, Europe, East African Islands, North America, Brazil (Bai, 1996; Furuki and
Mizutani, 1994; Grolle, 1995; Hamlin, 1972; Jia and Wang, 1993; Li, Liao and Huang, 1998; Piippo,
1990; Rao et al., 1997; Scott and Bradshaw,1985; Tan and Engel, 1986; Yamada and Choe, 1997;
Yano, 1995; Zhu, So and Ye, 1998).
PALLAVICINIACEAE
Pallavicinia lyellii (Hook.) Carruth.
Localities: NEW TERRITORIES. TAI MO SHAN. Ng Tung Chai Falls. 300 m (N), 5(7). Tai Shing
Stream. 200 m (SE), 17(13); 400 m (SE), 19(2); 500 m (SE), 20(15); 700 m (SE), 6(19); 800 m (NE),
7(13). – TAI PO KAU. Tsung Tsai Yuen. – WU KAU TANG. Lai Chi Wo. HONG KONG IS.
ABERDEEN. – POKFULAM (Gao and But, 1992).
Habitat and Substrate: All specimens were collected in forest. 5 were found on rocks, and 1 on soil.
Distribution: China (Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hunan, Jiangxi, Liaoning,
Shandong, Sichuan, Taiwan, Yunnan, Zhejiang). Japan, Philippines, Borneo, Indonesia, Papua New
Guinea, Australia, New Zealand, Russia, Africa, East African Islands, North America, Brazil
(Constantinova, Potemkin and Schljakov, 1992; Fang et al., 1998; Furuki and Mizutani, 1994; Grolle,
1995; Grolle and Piippo, 1984; Hamlin, 1972; Jia and Wang,1993; Menzel, 1988; Piippo, He and
Koponen, 1997; Rao et al., 1997; Scott and Bradshaw, 1985; Stotler and Crandall-Stotler, 1977;
Wigginton and Grolle, 1996; Yano, 1995; Zhao and Cao, 1998).
Pallavicinia subciliata (Aust.) Steph. (P. longispina Steph.)
Localities: NEW TERRITORIES. TAI MO SHAN. Ng Tung Chai Falls. 700 m (N), 1(4). Tai Shing
Stream. 200 m (SE), 17(6); 500 m (SE), 20(4). LANTAU IS. SUNSET PEAK. Wong Lung Hang (East
Lung Stream). 100 m (N), 39(1); 200 m (N), 40(9); 500 m (N), 36(1).
Habitat and Substrate: All specimens were collected in forest. 3 were found on rocks, 2 on soil, and 1
on a fern root.
Distribution: China (Guangdong, Hainan, Hong Kong, Hunan, Jiangxi, Taiwan, Yunnan, Zhejiang).
Japan (Fang et al., 1998; Furuki and Mizutani, 1994; Gao and Cao, 2000; Koponen et al., 2000; Lin et
al., 1994).
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Zhang Li: Hong Kong Bryophytes 103
PLAGIOCHILACEAE
Plagiochila chinensis Steph.
Locality: LANTAU IS. SUNSET PEAK. Wong Lung Hang (East Lung Stream). 800 m (N), 42(17).
Habitat and Substrate: On rocks in forest.
Distribution: China (Fujian, Guangxi, Guizhou, Hebei, Hong Kong, Hunan, Jiangxi, Sichuan, Taiwan,
Xizang, Yunnan, Zhejiang). India, Pakistan, Nepal, Bhutan, Vietnam, Thailand (Grolle and So, 1998;
Long and Grolle, 1990; Piippo, 1990; Piippo, He and Koponen, 1997; So, 2001; Zhu, So and Ye,
1998).
Plagiochila fordiana Steph. – Grolle and So (1999b), So (2001), Zhu and So (2001)
Locality: NEW TERRITORIES. TAI MO SHAN (Grolle and So, 1999b; So, 2001).
Habitat and Substrate: Unknown (Grolle and So, 1999b).
Distribution: China (Fujian, Guangdong, Hainan, Hong Kong, Taiwan, Yunnan). Japan, India, Bonin
Island, Vietnam, Thailand (Grolle and So, 1999b; So, 2001).
Plagiochila javanica (Sw.) Nees & Mont. (P. tjibodensis Schiffn., fide Grolle and So (1999b)) – Grolle
and So (1999b), So and Zhu (1997), Zhu and So (1996) as P. tjibodensis
Localities: NEW TERRITORIES. TAI MO SHAN. 600 m (So and Zhu, 1997). HONG KONG
(Locality unknown). 800 m (Grolle and So, 1999b).
Habitat and Substrate: On wet rocks (Grolle and So, 1999b, So and Zhu, 1997).
Distribution: China (Guangdong, Hong Kong, Taiwan, Yunnan). India, Sri Lanka, Bhutan, Nepal,
Thailand, Philippines, Indonesia, Papua New Guinea, Australia, Somoa, Solomon Islands (Grolle and
So, 1999a; Piippo, 1989, 1993; Scott and Bradshaw, 1985).
Plagiochila parvifolia Lindenb. (P. yokogurensis Steph., fide Grolle and So (1999b)) – Grolle and So
(1999b), So (2001); Zhu and So (1996) as P. yokogurensis
Locality. NEW TERRITORIES. TAI MO SHAN. 600 m (Grolle and So, 1999b; So, 2001).
Habitat and Substrate: On rock surface beside stream (Grolle and So, 1999b).
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hunan,
Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Korea, Japan, India, Bangladesh, Sri Lanka, Bhutan,
Nepal, Myanmar, Vietnam, Thailand, Philippines, Indonesia, Papua New Guinea (Furuki and Mizutani,
1994; Grolle and So, 1999b; So, 2001).
Plagiochila peculiaris Schiff. – So (2001)
Localities: NEW TERRITORIES. TAI MO SHAN (So, 2001).
Habitat and Substrate: Unknown.
Distribution: China (Fujian, Guangdong, Guangxi, Hainan, Hong Kong, Jiangxi, Taiwan, Yunnan,
Zhejiang). Japan, India, Bhutan, Nepal, Vietnam, Thailand, Borneo, Indonesia) (So, 2001).
Plagiochila sciophila Nees – So (2001), Zhu and So (2001)
Localities: NEW TERRITORIES. MA ON SHAN. Tai Lam Wu, 130 m (Zhu and So (2001),
originally reported as P. fruticosa by But and Gao (1991a)). -. TAI MO SHAN (So, 2001). Ng Tung
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104 Memoirs of the Hong Kong Natural History Society 26, 2003
Chai Falls. 200 m (N), 16(20); 300 m (N), 5(6); 395 m (N), 4(19); 600 m (N), 2(14). LANTAU IS.
SUNSET PEAK. Wong Lung Hang (East Lung Stream). 400 m (N), 35(12); 500 m (N), 36(9); 700 m
(N), 38(25).
Habitat and Substrate: All 7 specimens examined were collected in forest. 5 were found on rocks, and 2
on tree bases.
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Jiangsu,
Jiangxi, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Korea, Japan, India, Nepal, Pakistan, Bhutan,
Vietnam, Thailand, Philippines, Borneo, Indonesia, Papua New Guinea, Eastern North America
(Arkansas) (Fang et al., 1998; Furuki and Mizutani, 1994; Jia and Wang, 1993; Long and Grolle, 1990;
Menzel, 1988; Piippo, 1989, 1990; Piippo et al., 1998; So, 2001; Yamada and Choe, 1997; Zhu and So,
2001; Zhu, So and Ye, 1998).
Plagiochila semidecurrens (Lehm. & Lindenb.) Lindenb. – So (2001), Zhu and So (1996)
Locality: NEW TERRITORIES. TAI MO SHAN. 650 m or above (Zhu and So, 1996).
Habitat and Substrate: On rock and soil surfaces (Zhu and So, 1996).
Distribution: China (Anhui, Fujian, Guangxi, Guizhou, Hong Kong, Jiangxi, Sichuan, Taiwan, Xizang,
Yunnan, Zhejiang). Korea, Japan, India, Sri Lanka, Bhutan, Nepal, Vietnam, Thailand, Philippines,
Borneo, Indonesia, Canada) (So, 2001).
PORELLACEAE
Porella campylophylla (Lehm. & Lindenb.) Trev. – So and Zhu (1996b)
Locality: NEW TERRITORIES. TAI MO SHAN. 800 m (So and Zhu, 1996b).
Habitat and Substrate: On submerged rocks (So and Zhu, 1996b).
Distribution: China (Guangdong, Guangxi, Hong Kong, Xizang, Yunnan, Zhejiang). India, Nepal,
Bhutan, Sri Lanka, Myanmar, Vietnam (Long and Grolle, 1990; Piippo, 1990).
Porella perrottetiana (Mont.) Trev. – So and Zhu (1996b), Zhu and So (1996)
Locality: NEW TERRITORIES. TAI MO SHAN. Ng Tung Chai. 400 m (So and Zhu, 1996b).
Habitat and Substrate: On rocks (So and Zhu, 1996b).
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hong Kong, Hunan, Jiangsu,
Shaanxi, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang). Korea, Japan, India, Bhutan, Sri Lanka,
Myanmar, Thailand, Philippines (Furuki and Mizutani, 1994; Hattori, 1978; Long and Grolle, 1990;
Piippo, 1990; Piippo, He and Koponen, 1997; Yamada and Choe, 1997).
RADULACEAE
Radula apiculata Sande Lac. – So and Zhu (1996b), Zhu and So (1996)
Locality: NEW TERRITORIES. TAI MO SHAN. 200-850 m (Zhu and So, 1996).
Habitat and Substrate: On trunks and moist rocks (Zhu and So, 1996).
Distribution: China (Guangdong, Hong Kong, Taiwan, Zhejiang). Japan, Thailand, Philippines,
Nicobar Islands, Indonesia, Borneo, Papua New Guinea, Caroline Islands, Samoa, Society Islands
(Furuki and Mizutani, 1994; Grolle and Schultze-Motel, 1972; Li and Piippo, 1994; Menzel, 1988;
Yamada and Piippo, 1989; Zhu, So and Ye, 1998).
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Zhang Li: Hong Kong Bryophytes 105
Radula cavifolia Hampe – So and Zhu (1996b), Zhu and So (1996)
Locality: NEW TERRITORIES. TAI MO SHAN. 800 m (N), 950 m (So and Zhu, 1996b; Zhu and
So, 1996).
Habitat and Substrate: On trunks (So and Zhu, 1996b).
Distribution: China (Anhui, Fujian, Guangxi, Hong Kong, Jiangxi, Sichuan, Taiwan, Yunnan,
Zhejiang). Korea, Japan, Vietnam, Malaysia, Philippines, Borneo, Indonesia (Fang et al., 1998; Furuki
and Mizutani, 1994; Gao and Cao, 2000; Jia and Wang, 1993; Menzel, 1988; Piippo, 1990; Piippo, He
and Koponen, 1997; Yamada, 1989; Yamada and Choe, 1997; Zhu, So and Ye, 1998).
Radula japonica Gott. Ex Steph. – Zhu and So (1996)
Locality: NEW TERRITORIES. TAI MO SHAN. 800 m, 900 m (So and Zhu, 1996b; Zhu and So,
1996).
Habitat and Substrate: On trunks and moist rocks (Zhu and So, 1996).
Distribution: China (Fujian, Hong Kong, Jiangsu, Shandong, Taiwan, Yunnan, Zhejiang). Korea, Japan
(Furuki and Mizutani, 1994; Gao and Cao, 2000; Yamada and Choe, 1997; Zhao and Cao, 1998).
Radula javanica Gott. In Gott. Et al. – So and Zhu (1996b)
Localities: NEW TERRITORIES. TAI MO SHAN. 700 m (So and Zhu, 1996b). – TAI PO KAU
(Zhu and so, 2001).
Habitat and Substrate: On rocks (So and Zhu, 1996b).
Distribution: China (Fujian, Guangdong, Guangxi, Hainan, Hong Kong, Jiangxi, Taiwan, Xizang,
Yunnan, Zhejiang). Japan, India, Sri Lanka, Vietnam, Thailand, Malaysia, Philippines, Borneo,
Indonesia, Papua New Guinea, Australia, Society Islands, Samoa (Fang et al., 1998; Furuki and
Mizutani, 1994; Gao and Cao, 2000; Grolle and Piippo, 1984; Menzel, 1988; Yamada, 1989; Yamada
and Piippo, 1985; Zhu and So, 2001).
Radula kojana Steph. – Zhu and So (2001)
Locality: Unknown (Zhu and So, 2001).
Habitat: Unknown (Zhu and So, 2001).
Diatribution: China (Anhui, Fujian, Guangdong, Guangxi, Hong Kong, Hunan, Jiangxi, Sichuan,
Taiwan, Yunnan). Korea, Japan, Philippines (Zhu and So, 2001).
Radula oyamensis Steph. – So and Zhu (1996b), Zhu and So (1996)
Locality: NEW TERRITORIES. TAI MO SHAN. 700 m (So and Zhu, 1996b).
Habitat and Substrate: On trunks, rotten logs, moist rocks, bark (So and Zhu, 1996b; Zhu and So,
1996).
Distribution: China (Fujian, Hong Kong, Taiwan, Zhejiang). Japan (Furuki and Mizutani, 1994; So and
Zhu, 1996b).
Radula tokiensis Steph. – So and Zhu (1996b), Zhu and So (1996)
Locality: NEW TERRITORIES. TAI MO SHAN. 700 m (So and Zhu, 1996b).
Habitat and Substrate: On wet rocks and bark (So and Zhu, 1996b; Zhu and So, 1996).
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106 Memoirs of the Hong Kong Natural History Society 26, 2003
Distribution: China (Hong Kong, Sichuan, Taiwan, Zhejiang). Korea, Japan (Furuki and Mizutani,
1994; Piippo, He and Koponen, 1997; Zhu, So and Ye, 1998).
RICCIACEAE
Riccia fluitans L. – So and Zhu (1996b)
Localities: NEW TERRITORIES. FANLING (So and Zhu, 1996b). – SHA TAU KOK. Tsiu Hang.
10 m, 45(1). HONG KONG IS. ZOOLOGOCAL AND BOTANICAL GARDENS (So and Zhu,
1996b).
Habitat and Substrate: On soil in agricultural land.
Distribution: Cosmopolitan.
Riccia glauca L.
Locality: HONG KONG (Locality unknown) (So and Zhu, 1995c).
Habitat and Substrate: On wet soil along river banks and vegetable fields (So and Zhu, 1995c).
Distribution: Cosmopolitan.
SCAPANIACEAE
Scapania stephanii K. Muell. – Zhu and So (1996)
Locality: NEW TERRITORIES. TAI MO SHAN. 900 m (Zhu and So, 1996).
Habitat and Substrate: Unknown.
Distribution: China (Anhui, Fujian, Guangdong, Guizhou, Hunan, Jiangxi, Liaoning, Sichuan, Taiwan,
Yunnan, Zhejiang). Japan, Nepal (Gao and Cao, 2000; Piippo, 1990).
TRICHOCOLEACEAE
Trichocolea tomentella (Ehrh.) Dum., s. lat. – Zhang (1998a)
Localities: NEW TERRITORIES. MA ON SHAN. Ma On Shan Tsuen Stream. 580 m (SW), 2167. –
TAI MO SHAN. Tai Shing Stream. 700 m (SE), 6(13). Water-pumping House Stream. 550 m (E), Lin
et al. 2064 (IBSC, KFBG); 680 m (E), Lin et al. 2034 (IBSC, KFBG). LANTAU IS. LANTAU PEAK.
Wong Lung Hang (Main Streams). 590 m (N), 69.
Habitat and Substrate: All 5 specimens were collected in forest. 2 were found on rocks, 2 on humus,
and 1 on a tree base and rocks.
Distribution: China (Anhui, Guangdong, Guangxi, Guizhou, Hainan, Heilongjiang, Hong Kong,
Hunan, Jiangxi, Shaanxi, Shandong, Sichuan, Taiwan, Xinjiang, Xizang, Yunnan, Zhejiang). Korea,
Japan, Bhutan, Philippines, Borneo, Indonesia, Papua New Guinea, Caroline Islands, Somoa, Fiji,
Solomon Islands, Russia, Europe, North America (Constantinova, Potemkin and Schljakov, 1992; Fang
et al., 1998; Furuki and Mizutani, 1994; Jia and Wang, 1993; Li, Liao and Huang, 1998; Long and
Grolle, 1990; Menzel, 1988; Piippo, 1993; Piippo, He and Koponen, 1997; Rao et al., 1997; Tan and
Engel, 1986; Yamada and Choe, 1997; Zhang et al., 1998a; Zhao and Cao, 1998).
WIESNERELLACEAE
Dumortiera hirsuta (Swaegr.) Nees
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Zhang Li: Hong Kong Bryophytes 107
Localities: NEW TERRITORIES. TAI MO SHAN. Ng Tung Chai Falls. 300 m (N), 5(11)
(Confirmed by P.-C. Wu); 395 m (N), 4(9); 600 m (N), 2(1); 700 m (N), 1(21).
Habitat and Substrate: All 4 specimens were collected in forest. 2 were found on rocks, and 2 on soil.
Distribution: Cosmopolitan.
4.6
Excluded species of liverworts
Marchantia emarginata Reinw. Et al.
Zhang and Lin (1997) errorously listed this species as occurring in Hong Kong based on So and Zhu
(1995c).
Plagiochila fruticosa Mitt.
According to Zhu and So (2001), the voucher of the present species is P. sciophila.
4.6
Doubtful species of liverworts
Archilejeunea mariana (Gott.) B. Thiers & Gradst.
Zhu and So (2000b) excluded this species from China, but a specimen from Hong Kong was quite
similar to the present species.
Specimen examined: LANTAU IS. SUNSET PEAK. San She Wan Stream. 300 m (S), on tree base,
32(8) (Det. By P.-C. Wu as Spruceanthus mariana).
Cephalozia otaruensis Steph.
So (1995a) reported that this species occurs in Hong Kong but did not cite any specimens. Zhang and
Lin (1997) and Zhu and So (1996) included it in their lists.
Cololejeunea amoena Bened.
According to Zhu and So (2001), the occurrence of this species in China remained doubtful.
Marchantia paleacea Bertol.
So (1995a) reported that this species occurs in Hong Kong but did not cite any specimens. Zhang and
Lin (1997) and Zhu and So (1996) included it in their lists.
Mastigolejeunea auriculata (Wils.) Schiffn.
So (1995a) reported that this species occurs in Hong Kong but did not cite any specimens.
Plagiochila ovalifolia Mitt.
Zhu and So (1996) included this species in their list, but Zhang and Lin (1997) could not locate the
source reference on which the record was based.
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ACKNOWLEDGEMENTS
First of all, I would like to express my sincere thanks to my supervisor, Dr. Richard T. Corlett, for his
guidance, encouragement, inspiration and patience throughout the study (November 1997-June 2001).
Special thanks are extended to Dr. Lawrence K. C. Chau and Ms. Gloria Siu for their invaluable
support during the course of study, and Ms. Laura Wong for her technical help.
Thanks are also extended to Dr. Ng Sai Chit, Prof. F.-W. Xing, Mr. Wicky Lee, Mr. Benson Lee, Mr.
Choi Kin-Hung, Mr. Ken Chu, and Mr. Ken So for their help during field surveys, Prof. P. P.-H. But
for reviewing the manuscript, Prof. I. J. Hodgkiss for the arrangement of publication of this paper, Prof.
P.-C. Wu, Dr. B. C. Tan, Prof. W. D. Reese, Dr. T. Yamaguchi, Prof. Y.-F. Wang, Prof. D.-K. Li, Prof.
118
Zhang Li: Hong Kong Bryophytes 119
Z.-H. Li, Prof. J.-P. Frahm, Dr. M. L. So, Dr. R.-L. Zhu, Dr. R. M. K. Saunders, Dr. Si He, Dr. H.
Akiyama, Prof. P.-J. Lin, Prof. T. Cao, Mr. Z. Magombo, and Mr. R.-J. Wang for their various help.
I wish to thank The University of Hong Kong for granting me a postgraduate studentship for
supporting my study, and Kadoorie Farm & Botanic Garden for supplying me the working and living
convenience from November 1997 to April 2000.
I am also grateful to the Department of Ecology & Biodiversity, The University of Hong Kong, and
Karoorie Farm & Botanic Garden for their joint support for the publication of this inventory.
This paper is mainly based on Chapter 4 of my Ph.D. thesis with updated information considered.
119
120 Memoirs of the Hong Kong Natural History Society 26, 2003
INDEX TO SCIENTIFIC NAMES
Family name is in Roman type. Accepted names are in boldface and italic, and excluded, doubtful or
synonyms are in italic type.
88
1, 4, 65
48
48
48
5
78
78
61
79
79
5
5
107
88
88
59, 77
59, 77
80
1, 4, 49
49
61, 75
61, 76
62
62
78
5
98
73, 99
99
99
99
8, 63, 64
6
6
6
7
Acrolejeunea pusilla
Acroporium secundum
Aerobryidium wallichii
Aerobryopsis praisii
Aerobryopsis subdivergens
Amblystegiaceae
Aneura pinguis
Aneuraceae
Anoectangium thomsonii
Anthoceros punctatus
Anthocerotaceae
Archidiaceae
Archidium ohioense
Archilejeunea mariana
Archilejeunea planiuscula
Archilejeunea polymorpha
Atrichum undulatum var. gracilisetum
Atrichum undulatum var. undulatum
Aytoniaceae
Barbella convolvens
Barbella flagellifera
Barbula arcuata
Barbula indica
Barbula javanica
Barbula unguiculata
Bartramia mollis
Bartramiaceae
Bazzania fauriana
Bazzania japonica
Bazzania sikkimensis
Bazzania tricrenata
Bazzania tridens
Brachymenium exile
Brachytheciaceae
Brachythecium garovaglioides
Brachythecium moriense
Brachythecium plumosum
120
Zhang Li: Hong Kong Bryophytes 121
Brachythecium wichurae
Brotherella erythrocaulis
Brotherella falcata
Bryaceae
Bryohaplocladium angustifolium
Bryohaplocladium microphyllum
Bryum argenteum
Bryum atrovirens
Bryum billardieri
Bryum caespiticium
Bryum capillare
Bryum capillare ssp. torquescens
Bryum cellulare
Bryum coronatum
Bryum erythrocarpum
Bryum torquescens
Caduciella guangdongensis
Callicostella papillata
Calymperaceae
Calymperes afzelii
Calymperes erosum
Calymperes fasciculatum
Calymperes graeffeanum
Calymperes lonchophyllum
Calymperes molluccense
Calymperes palisotii ssp. molluccense
Calymperes serratum
Calymperes taitense
Calymperes tenerum
Calypogeia arguta
Calypogeia tosana
Calypogeiaceae
Calyptothecium wightii
Campylopodium proscriptum
Campylopus atrovirens
Campylopus comosus
Campylopus ericoides
Campylopus japonicus
Campylopus laxitextus
Campylopus scabridorsus
Campylopus serratus
Campylopus subulatus
6
42, 66
66
8
72
72
8
8
8
9
9
75
9
9, 75
8
75
44
35, 43, 77
10
10
11, 76
12
12
12, 14
12, 75
12, 75, 76
75
75
12
32, 80
81
80
64
75
16
17, 75
16
17
17
17, 75
17, 75
77
121
122 Memoirs of the Hong Kong Natural History Society 26, 2003
17
18
81
81
107
81
81
81
81
100
88
88
89
89
89
89
84
84
84
71
71
49, 66
107
89
89
90
90
90
91
90
91
91
91
91
91
92
92
89
92
92
92
93
Campylopus taiwanensis
Campylopus umbellatus
Cephalozia hamatiloba
Cephalozia macounii
Cephalozia otaruensis
Cephalozia otaruensis var. setiloba
Cephaloziaceae
Cephaloziella microphylla
Cephaloziellaceae
Chandonanthus birmensis
Cheilolejeunea imbricata
Cheilolejeunea intertexta
Cheilolejeunea osumiensis
Cheilolejeunea ryukyuensis
Cheilolejeunea trifaria
Cheilolejeunea ventricosa
Chiloscyphus minor
Chiloscyphus polyanthos
Chiloscyphus polyanthus
Claopodium aciculum
Claopodium assurgens
Clastobryopsis planula
Cololejeunea amoena
Cololejeunea appressa
Cololejeunea ceratilobula
Cololejeunea floccosa
Cololejeunea goebelii
Cololejeunea haskarliana
Cololejeunea himalayensis
Cololejeunea hispidissima
Cololejeunea lanciloba
Cololejeunea latilobula
Cololejeunea ocellata
Cololejeunea ocelloides
Cololejeunea planissima
Cololejeunea pseudocristallina
Cololejeunea raduliloba
Cololejeunea reineckeana
Cololejeunea schmidtii
Cololejeunea schwabei
Cololejeunea spinosa
Cololejeunea stylosa
122
Zhang Li: Hong Kong Bryophytes 123
Conocephalaceae
Conocephalum conicum
Conocephalum japonicum
Cryphaeaceae
Ctenidium andoi
Ctenidium capillifolium
Ctenidium serratifolium
Cyathophorella tonkinensis
Cylindrocolea tagawae
Dicranaceae
Dicranella coarctata var. coarctata
Dicranella coarctata var. obscura
Dicranella coarctata var. torrentium
Dicranella obscura
Dicranella varia
Dicranodontium asperulum
Dicranodontium didictyon
Dicranoloma assimile
Dicranoloma cylindrothecium
Dicranoloma fragiliforme
Dicranum sericifolium
Didymodon vineralis
Diphysciaceae
Diphyscium fulvifolium
Diphyscium involutum
Distichophyllum collenchymatosum
Distichophyllum maibarae
Distichophyllum obtusifolium
Ditrichaceae
Ditrichum pallidum
Drepanolejeunea angustifolia
Drepanolejeunea erecta
Drepanolejeunea formosana
Drepanolejeunea spicata
Drepanolejeunea ternatensis
Dumortiera hirsuta
Duthiella wallichii
Eccremidium brisbanicum
Ectropothecium dealbatum
Ectropothecium inflectens
Ectropothecium monumentorum
Ectropothecium monumentorum f. macromorpha
82
82
82
16
37
37
37
43
82
16
18, 75
18
19
75
19
19
19
20
20
20
20
75
22
77
22, 30, 77
36
36
77
22
22
93
93
97
93
93
106
49
23, 24
38
42
38, 76
76
123
124 Memoirs of the Hong Kong Natural History Society 26, 2003
Ectropothecium nervosum
Ectropothecium obtusulum
Ectropothecium planulum
Ectropothecium zollingeri
Entodon cladorrizans
Entodon concinnus
Entodon macropodus
Entodon obtusatus
Entodon viridulus
Entodontaceae
Entodontopsis anceps
Entosodon javanensis
Ephemeraceae
Eurhynchium asperisetum
Eurhynchium hians
Eurhynchium savatieri
Fabroniaceae
Fauriella tenerrima
Fissidens adelphinus
Fissidens areolatus
Fissidens brevinervis
Fissidens bryoides var. schmidii
Fissidens ceylonensis
Fissidens cf. incognitus
Fissidens crenulatus
Fissidens crispulus var. cripulus
Fissidens crispulus var. robinsonii
Fissidens cristatus
Fissidens curvatus
Fissidens dubius
Fissidens flaccidus
Fissidens gardneri
Fissidens geminiflorus
Fissidens geppii
Fissidens guangdongensis
Fissidens gymnogynus
Fissidens hollianus
Fissidens javanicus
Fissidens jungermannioides
Fissidens kinabaluensis
Fissidens laxus
Fissidens linearis var. obscurirete
1, 4, 38, 76
38
38
1, 4, 38
23
23
24
24
24
23
70
77
24
7
7
7
25
71
33
33
28
25
25
30
26
26
27
27
27
27
28
28
28
29
29
29
29
30
30
30
32
31
124
Zhang Li: Hong Kong Bryophytes 125
Fissidens maceratus
Fissidens minutus
Fissidens nobilis
Fissidens oblongifolius
Fissidens obscurirete
Fissidens papillous
Fissidens pellucidus
Fissidens polypodioides
Fissidens serratus
Fissidens splachnobryoides
Fissidens strictulus
Fissidens teysmannianus
Fissidens tosanensis
Fissidens zippelianus
Fissidens zollingeri
Fissidentaceae
Floribundaria floribunda
Folioceros fuciformis
Fossombrobiaceae
Fossombronia cristula
Frullania ericoides
Frullania moniliata
Frullania motoyata
Frullania muscicola
Frullania nepalensis
Frullania tamarisci
Frullania tamsuina
Frullania trichodes
Frullaniaceae
Funaria hygrometrica
Funaria physcomitrioides
Funariaceae
Garckea flexuosa
Geocalycaceae
Glossadelphus lingulatus
Grimmiaceae
Gymnostomiella longinervis
Gymnostomiella vernicosa var. tenerum
Haplocladium angustifolium
Haplocladium microphyllum
Haplohymenium pseudo-triste
Haplomitriaceae
28
31, 32
31
31, 32
31
33
32
33
33
28
27
33
34
26
34
25
49
80
82
82
82
83
83
83
83
84
84
84
82
34
77
34
23
84
40
35
70
70
72
72
72
86
125
126 Memoirs of the Hong Kong Natural History Society 26, 2003
86
86
86
72
37, 80, 85
85
85
85
53, 76
53, 76
20
20
20, 76
54
54
54
55
54
54
36
35
37
61
62
37
62
62
77
63
37
39
39
43
43
43
66
40
40
77
86
86
86
Haplomitrium mnioides
Herbertaceae
Herbertus aduncus
Herpetineuron toccoae
Heteroscyphus argutus
Heteroscyphus bescherellei
Heteroscyphus coalitus
Heteroscyphus planus
Himantocladium cyclophyllum
Himantocladium plumula
Holomitrium cylindraceum
Holomitrium densifolium
Holomitrium vaginatum
Homalia trichomanoides
Homaliodendron exiguum
Homaliodendron flabellatum
Homaliodendron microdendrom
Homaliodendron scapellifolium
Homaliodendron squarrulosum
Hookeria acutifolia
Hookeriaceae
Hookeriopsis utacamundiana
Hydrogonium arcuatum
Hydrogonium javanicum var. convolutifolium
Hylocomiaceae
Hyophila involuta
Hyophila javanica
Hyophila rosea
Hyophila setschwanica
Hypnaceae
Hypnum plumaeformae var. minus
Hypnum plumaeformae var. plumaeformae
Hypopterygiaceae
Hypopterygium sinicum
Hypopterygium tenellum
Isocladiella surcularis
Isopterygium albescens
Isopterygium minutirameum
Isopterygium tenerum
Jackiella javanica
Jackiellaceae
Jubula hutchinsiae ssp. javanica
126
Zhang Li: Hong Kong Bryophytes 127
Jubulaceae
Jungermannia infusca
Jungermannia tetragona
Jungermannia truncata
Jungermanniaceae
Kurzia gonyotricha
Lejeunea anisophylla
Lejeunea catanduana
Lejeunea cocoes
Lejeunea curviloba
Lejeunea flava
Lejeunea japonica
Lejeunea magohukui
Lejeunea otiana
Lejeunea parva
Lejeunea punctiformis
Lejeunea tuberculosa
Lejeunea ulicina
Lejeuneaceae
Lepicoleaceae
Lepidozia fauriana
Lepidozia vitrea
Lepidoziaceae
Leptodontaceae
Leptolejeunea elliptica
Leskeaceae
Leucobryaceae
Leucobryum aduncum
Leucobryum boninense
Leucobryum bowringii
Leucobryum humillimum
Leucobryum javense
Leucobryum juniperoideum
Leucobryum neilgherrense
Leucobryum scaberulum
Leucobryum scaberulum var. divaricatum
Leucobryum scabrum
Leucolejeunea xanthocarpa
Leucoloma molle
Leucoloma walkeri
Leucophanes albescens
Leucophanes glaucum
86
87
87
87
87
100
94
94
94
94
94
95
95
95
95
95
95
95
88
98
100
100
98
44
96
44
44
44
45
45
46
46
46
46
45
45
47
96
21
21
47
1, 4, 47
127
128 Memoirs of the Hong Kong Natural History Society 26, 2003
96
96
97
97
100
44
78
63
56
56, 57
57, 76
57
57
58, 76
58, 76
57
76
58
58
37
101
101
107
101
107
101
101
107
98
80
63
48
49, 50
101
101
102
101
21
95
23, 24
1, 4, 13
13
Lopholejeunea applanata
Lopholejeunea eulopha
Lopholejeunea nicobarica
Lopholejeunea subfusca
Lophoziaceae
Lopidium nazeense
Lorentzia velata
Luisierella barbula
Macromitrium brevituberculatum
Macromitrium ferriei
Macromitrium heterodictyon
Macromitrium japonicum
Macromitrium microstomum
Macromitrium moorcroftii
Macromitrium nepalense
Macromitrium reinwardtii
Macromitrium sinense
Macromitrium subincurvum
Macromitrium tuberculatum
Macrothamnium macrocarpum
Makinoa crispata
Makinoaceae
Marchantia emarginata
Marchantia emarginata ssp. tosana
Marchantia paleacea
Marchantia polymorpha
Marchantiaceae
Mastigolejeunea auriculata
Mastigophora diclados
Megaceros flagellaris
Merceyopsis sikkimensis
Meteoriaceae
Meteorium subpolytrichum
Metzgeria conjugata
Metzgeria consanguinea
Metzgeria furcata
Metzgeriaceae
Microdus brasiliensis
Microlejeunea punctiformis
Micromitrium tenerum
Mitthyridium fasciculatum
Mitthyridium flavum
128
Zhang Li: Hong Kong Bryophytes 129
Mniaceae
Myurella julacea
Myuriaceae
Nardia assamica
Neckeraceae
Neckeropsis obtusata
Notoscyphus lutescens
Ochrobryum gardneri
Octoblepharum albidum
Oedicladium fragile
Oedicladium rufescens
Oedicladium serricuspe
Orthomnion loheri
Orthotrichaceae
Palamocladium nilgheriense
Palisadula chrysophylla
Pallavicinia longispina
Pallavicinia lyellii
Pallavicinia subciliata
Pallaviciniaceae
Papillidiopsis complanata
Papillidiopsis macrosticta
Pelekium velatum
Phaeoceros laevis
Philonotis appressifolia
Philonotis hastata
Philonotis mollis
Philonotis thwaitesii
Philonotis turneriana
Phyllodon lingulatus
Physcomitrium eurystomum
Physcomitrium japonicum
Physcomitrium sphaericum
Pilotrichopsis dentata
Pinnatella anacamptolepis
Plagiochila chinensis
Plagiochila fordiana
Plagiochila fruticosa
Plagiochila javanica
Plagiochila ovalifolia
Plagiochila parvifolia
Plagiochila peculiaris
51
78
52
87
53
55
87
78
48
52
53
53
76
56
78
53
102
102
85, 102
102
67
67
78
80
6
5
78
6
6
40
34
35
35
16
55
103
103
103, 107
103
107
103
103
129
130 Memoirs of the Hong Kong Natural History Society 26, 2003
103, 107
104
103
103
103
51
51
52
59
59
59
59
5
60
60
60
10
10
59
61
61
104
104
104
61, 70
50
50
51
26, 44
41
64
97
65
35
65
65
104
105
105
105
105
105
Plagiochila sciophila
Plagiochila semidecurrens
Plagiochila tjibodensis
Plagiochila yokogurensis
Plagiochilaceae
Plagiomnium cuspidatum
Plagiomnium succulentum
Plagiomnium vesicatum
Plagiotheciaceae
Plagiothecium cavifolium
Plagiothecium euryphyllum
Plagiothecium nemorale
Platyhypnidium riparioides
Pogonatum camusii
Pogonatum contortum
Pogonatum neesii
Pohlia camptotrachela
Pohlia proligera
Polytrichaceae
Polytrichastrum formosum
Polytrichum formosum
Porella campylophylla
Porella perrottetiana
Porellaceae
Pottiaceae
Pseudobarbella angustifolia
Pseudobarbella attenuata
Pseudobarbella leveri
Pseudoleskeopsis zippelii
Pseudotaxiphyllum pohliaecarpum
Pterobryaceae
Ptychanthus striatus
Pyrrhobryum spiniforme
Racomitrium fasciculare
Racopilaceae
Racopilum aristatum
Radula apiculata
Radula cavifolia
Radula japonica
Radula javanica
Radula kojana
Radula oyamensis
130
Zhang Li: Hong Kong Bryophytes 131
105
104
80
76
93
65
10
8
5
78
78
79
43, 79
79
106
106
106
106
106
78
58
58
25
63
65
68
68
67
67
68
68
68
61
70
88
107
97
97
70
41
76
13
Radula tokiensis
Radulaceae
Reboulia hemisphaerica
Rhamphidium vaginatum
Rhaphidolejeunea spicata
Rhizogoniaceae
Rhodobryum giganteum
Rhynchostegium pallidifolium
Rhynchostegium riparioides
Riccardia jackii
Riccardia kodame
Riccardia latifrons
Riccardia multifida
Riccardia palmata
Riccia fluitans
Riccia glauca
Ricciaceae
Scapaniaceae
Scapania stephanii
Schistomitrium gardnerianum
Schlotheimia grevilleana
Schlotheimia pungens
Schwetschkeopsis fabronia
Scopelophila cataractae
Sematophyllaceae
Sematophyllum demissum
Sematophyllum robustulum
Sematophyllum subhumile
Sematophyllum subhumile ssp. japonicum
Sematophyllum subpinnatum
Sematophyllum subpinnatum f. tristiculum
Sematophyllum tristiculum
Semibarbula orientalis
Splachnaceae
Spruceanthus polymorphus
Spruceanthus mariana
Spruceanthus semirepandus
Stenolejeunea apiculata
Stereodontaceae
Stereodontopsis pseudorevoluta
Syrrhopodon albovaginatus
Syrrhopodon armatus
131
132 Memoirs of the Hong Kong Natural History Society 26, 2003
14
14
14
14
15
15
1, 4, 16
16
41, 74
2, 78
40
41
41
42
69
69
69
69
70
56
71
71
73
74
73
74
25, 44, 74, 86
74
98
52
74
74
21
106
106
70
78
63
76
63
98
98
Syrrhopodon chenii
Syrrhopodon flammeonnervis
Syrrhopodon gardneri
Syrrhopodon hongkongensis
Syrrhopodon japonicus
Syrrhopodon prolifer var. tosaensis
Syrrhopodon spiculosus
Syrrhopodon trachyphyllus
Taxiphyllum arcuatum
Taxiphyllum inundatum
Taxiphyllum minutirameum
Taxiphyllum pilosum
Taxiphyllum subarcuatum
Taxiphyllum taxirameum
Taxithelium batanense
Taxithelium instratum
Taxithelium nepalense
Taxithelium oblongifolium
Taxithelium parvulum
Thamnobryum plicatulum
Theliaceae
Thuidiaceae
Thuidium cymbifolium
Thuidium delicatulum
Thuidium glaucinoides
Thuidium glaucinum
Thuidium pristocalyx
Thuidium recognitum var. delicatulum
Thysananthus flavescens
Trachycystis microphylla
Trachypodaceae
Trachypus humilis
Trematodon longicollis
Trichocolea tomentella s. lat.
Trichocoleaceae
Trichosteleum boschii
Trichosteleum mammosum
Trichostomum hattorianum
Trichostomum inflexum
Trichostomum involutum
Trocholejeunea sandvicensis
Tuyamaella molischii
132
Zhang Li: Hong Kong Bryophytes 133
Vesicularia aperta
Vesicularia dubyana
Vesicularia ferriei
Vesicularia inflectens
Vesicularia montagnei
Vesicularia reticulata
Weissia controversa
Weissia crispa
Weissia edentula
Weissia longifolia
Wiesnerellaceae
77
77
42, 77
42
43, 77
43, 77
63
64
64
64
106
133