LISTS OF SPECIES Check List 9(2): 329–390, 2013 © 2013 Check List and Authors ISSN 1809-127X (available at www.checklist.org.br) Chec List Journal of species lists and distribution Marine and coastal biodiversity of Oaxaca, Mexico J. Rolando Bastida-Zavala*, María del Socorro García-Madrigal, Edgar Francisco Rosas-Alquicira, Ramón Andrés López-Pérez, Francisco Benítez-Villalobos, Juan Francisco Meraz-Hernando, Ana María Torres-Huerta, Alberto Montoya-Márquez and Norma A. Barrientos-Luján Universidad del Mar, Instituto de Recursos, Campus Puerto Ángel, Ciudad Universitaria, Apdo. Postal 47, 70902, Puerto Ángel, Oaxaca, Mexico * Corresponding author. E-mail: rolando_bastida@yahoo.com.mx Abstract: The state of Oaxaca in southern Mexico harbors the highest faunistic and floristic biodiversity of the region; however, to date, research effort has focused on terrestrial flora and fauna. Despite this bias, there is a large amount of dispersed information regarding the coastal and marine biodiversity of Oaxaca. The present study aimed to: 1) update and synthesize existing information to improve understanding of coastal and marine biodiversity; and 2) provide a baseline for future biodiversity studies in Oaxaca. The review of 198 references about Oaxaca’s flora and fauna produced 2,157 species records, from 15 taxonomic groups (Division, Phyla or Class). Nevertheless, knowledge about the fauna and flora of Oaxaca remains scarce compared to other regions, such as the Gulf of California. Additionally, floristic and faunistic knowledge is heterogeneous among the taxonomic groups; for instance, 242 (11.2%) species represent macroalgae, three invertebrate groups (annelids, crustaceans and mollusks) represent about 44.0% (949 species) of all species recorded, while the vertebrates are represented by 762 species (35.3%). Similarly, many invertebrate groups recorded in adjacent regions have not yet been recorded on the Oaxaca coast, including some platyhelminthes, rotifers, nematodes, oligochaetes, sipunculids, echiurans, tardigrades, pycnogonids, some crustaceans, brachiopods, chaetognaths, ascidians and cephalochordates. The study of the marine flora and fauna is far from finished; additional effort is required to complete the marine biodiversity inventory of Oaxaca. Introduction The state of Oaxaca in southern Mexico (Figure 1) is known to harbor the highest continental faunistic and floristic biodiversity in Mexico (González-Pérez et al. 2004). While the terrestrial biodiversity of Oaxaca is wellestablished, overall knowledge is highly asymmetric when we compare the amount of work in this region devoted to terrestrial flora and fauna with that dedicated to coastal and marine environments. In fact, the most recent publication summarizing knowledge of the biodiversity of Oaxaca focuses exclusively on terrestrial flora and fauna (cf. García-Mendoza et al. 2004). In parallel, along with the relative absence of studies on the coastal and marine environments of Oaxaca, studies among taxonomic groups are not homogeneous: some taxa have been studied more intensively and for longer than others. Some of the main taxonomic groups that have been studied include macroalgae (Fragoso and Rodríguez 2002; Ávila-Ortiz 2003; Mateo-Cid et al. 2005; 2008; Aguilar-Rosas et al. 2009; Mateo-Cid and MendozaGonzález 2009), corals (Glynn and Leyte-Morales 1997; Leyte-Morales 1997; Reyes-Bonilla and Leyte-Morales 1998; Cruz-Piñón and Reyes-Bonilla 1999), polychaetes (Fernández-Álamo 1987; González-Ortiz et al. 1996; 1997; Gómez et al. 1997; Bastida-Zavala and ten Hove 2003; Bastida-Zavala 2008; Ruiz-Cancino et al. 2011), crustaceans (Howard 1952; Garth 1959; 1961; VillallobosFigueroa 1967; Haig 1968; Luke 1977; Sosa-Hernández et al. 1980; Villalobos-Hiriart and Nates-Rodríguez 1990; Carvacho and Haasmann 1984; Hendrickx et al. 1997; Hendrickx and Vázquez-Cureño 1998; Barbosa-Ledesma et al. 2000; Ramírez-Luna et al. 2002; Ayón-Parente and Hendrickx 2007; 2009; García-Madrigal 2010; García- Madrigal and Andreu-Sánchez 2010; Jarquín-González and García-Madrigal 2010), mollusks (Rodríguez-Palacios et al. 1988; Holguín-Quiñones and González-Pedraza 1989; de León-Herrera 2000; Ramírez-González and Barrientos-Luján 2007; Zamorano et al. 2008, 2010; RíosJara et al. 2009; Reyes-Gómez et al. 2010), echinoderms (Benítez-Villalobos 2001; Zamorano et al. 2006; BenítezVillalobos et al. 2008), fish (Chávez 1979; Castro-Aguirre 1982; Tapia-García et al. 1995; 1998; Castro-Aguirre Figure 1. Location of Oaxaca State in southern Mexico and main sites mentioned in the text: 1) Corralero Lagoon; 2) Chacahua-Pastoria Lagoon; 3) Puerto Escondido; 4) Agua Blanca; 5) Ventanilla-Mazunte; 6) Puerto Ángel; 7) Huatulco Bays; 8) Barra de la Cruz; 9) Morro Ayuta; 10) Salina Cruz; 11) Huave Lagoon System; 12) Mar Muerto. 329 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca and Espinosa-Pérez 1996; Tapia-García and GutiérrezDíaz 1998; Castro-Aguirre et al. 1999; Espinosa-Pérez et al. 2004; Tapia-García and Mendoza-Rodríguez 2005; Ramírez-Gutiérrez et al. 2007; Robertson and Allen 2008; López-Pérez et al. 2008, 2010, 2012; Mendoza-Vallejo et al. 2009), reptiles (Meraz-Hernando and Ávila-Nahón 2006; García-Grajales et al. 2007), coastal birds (Meraz-Hernando 2001; Navarro-Sigüenza et al. 2004; Meraz-Hernando and González-Bravo 2005), and marine mammals (MerazHernando and Sánchez-Díaz 2008). While a range of faunal prospecting has been undertaken in the area (Holguín-Quiñones and González-Pedraza 1989), these surveys have focused solely on specific localities or environments along the Oaxacan coast. For example, the rocky substrates of Huatulco, Puerto Ángel and Agua Blanca have been studied by Gómez et al. (1997), Fragoso and Rodríguez (2002) and Zamorano et al. (2006; 2008). In addition, pelagic waters have been studied by Fernández-Álamo (1987; 2000; 2002), while the soft bottoms of the continental shelf of the Gulf of Tehuantepec were prospected by Gamboa-Contreras and Tapia-García (1998) and Ríos-Jara et al. (2009). Meanwhile, floristic studies have focused on littoral and sublittoral rocky substrates; the sites that have been most extensively studied include Santa Elena (León-Tejera et al. 1993), Huatulco Bays (Mendoza-González and Mateo-Cid 1998), Salina Cruz (Galindo-Villegas et al. 1997), and the Huave Lagoon System (Dreckmann 1999). In contrast, there are several environments that have not been studied at all. For example, except for some records of polychaetes (Fauchald 1972), there is no information about the soft bottoms of most of the beaches of Oaxaca, nor of the deep-sea environments (over 200 m depth), particularly regarding the meiofauna (Bastida-Zavala and Guevara-Cruz 2012). In addition, there is a lack of knowledge about the symbiotic relationships between the shallow-water invertebrates or parasites of fish, birds, and marine mammals. It is remarkable that several invertebrate groups in Oaxaca have scarcely been studied or are even completely unknown. For instance, benthic hydrozoans, flatworms, rotifers, entoprocts (or kamptozoans), nemerteans, nematodes, sipunculids, echiurans, copepods, bryozoans and ascidians are practically unrecorded in Oaxaca. The study of some of these groups should in fact be mandatory, since some of them comprise the main components of bio-fouling fauna (e.g., sponges, benthic hydrozoans, entoproctos, amphipods, bryozoans and ascidians), and therefore have a high potential to be non-native invasive species (Okolodkov et al. 2007). Based on these accounts, it is clear that there is a major information gap in knowledge about the coastal and marine biodiversity of Oaxaca. The aim of the present study was therefore twofold: 1) to update and synthesize available information to improve understanding of the biodiversity of the region, and 2) to use this information as a baseline for future studies in Oaxaca. Materials and Methods Information was mainly gathered from 198 primary references that had been formally published as papers and monographs. Unpublished databases, technical reports and theses were avoided. In general, the taxonomic information was found to be of varying quality, depending on the taxonomic group. However, all taxa names were checked in the World Register of Marine Species (WoRMS, Appeltans et al. 2012), Integrated Taxonomic Information System (ITIS 2012), Smithsonian Tropical Research Institute (STRI, Robertson and Allen 2008), American Museum of Natural History (AMNH, Frost 2012) and American Ornithologists’ Union checklist (AOU 1998) and subsequent supplements (AOU 2000; Banks et al. 2002; 2004; 2006; 2007; 2008; Chesser et al. 2010). Questionable names were noted on the type localities. Results and Discussion A total of 2,157 species belonging to two divisions of the Kingdoms Plantae and Chromista, and ten Phyla of the Kingdom Animalia were recorded along the coastal and marine environments of Oaxaca (Table 1).This is clearly a smaller number of species when compared to the 3,112 terrestrial arthropods or the 1,431 vertebrates recorded for Oaxaca (González-Pérez et al. 2004). Although the marine mammals have a relatively low species richness along the coast of Oaxaca, it is nonetheless higher than in the Gulf of Mexico or the Caribbean (AguayoLobo et al. 1990). Reptile richness is also higher than the rest of the country, and is high in comparison to other parts of the world (CONABIO 1998). On the other hand, aquatic birds are poorly represented in Oaxaca in comparison to other regions of Mexico and abroad, possibly due to the absence of breeding areas and endemism (González-Bravo and Meraz-Hernando 2010). The best-represented group is the fish, with 594 species (27.5%), followed by mollusks (462 species, 21.4%), crustaceans (265, 12.3%), macroalgae (242, 11.2%) and annelids (222, 10.3%). In comparison, sponges (17 species, 0.8%), flatworms (8, 0.4%), nemerteans and bryozoans (1 species each, 0.1%) are the least represented marine invertebrates in Oaxaca (Table 1). The differential effort given to taxonomic groups is evident in the records derived from published papers. For example, while macroalgae, cnidarians, annelids, crustaceans and mollusks appeared in 152 papers, the remaining taxa have been recorded in less than 50 publications. Meanwhile, many groups have not yet been recorded on the coast of Oaxaca. For example, some flatworms (turbellarians and cestods), rotifers, nematodes, oligochaetes, sipunculids, echiurans, tardigrades, pycnogonids, some crustaceans (branchiopods, mysids, cumaceans and ostracods), brachiopods, chaetognaths, ascidians and cephalochordates remain unrecorded. The coastal/marine flora and fauna of Oaxaca has a low species richness in comparison to other relatively well-known ocean basins in Mexico (Table 2). For example, Oaxaca possesses only 33.5% of the species richness recorded in the Gulf of California (Hendrickx et al. 2005), but ~22% of that in the Gulf of Mexico (Felder and Camp 2009). Such differences might not only arise because of heterogeneity in the studies conducted in Oaxaca in comparison with the Gulf of California and Gulf of Mexico (Hendrickx et al. 2005; Felder and Camp 2009 and references therein), but also as a result of differences in basin size: Oaxaca represents only 10% and 1% of the littoral area of the Gulf of California and the Gulf of Mexico, 330 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca respectively. Background problems in knowledge about marine biodiversity Species from most taxonomic groups were recorded near major towns or harbors, such as Puerto Escondido, Puerto Angel, Bahias de Huatulco and Salina Cruz, while the floral and faunal composition of extensive areas of shallow littoral habitats, the continental shelf and the deep sea of Oaxaca remain understudied (López-Pérez et al. 2012). Fish may be an exception, because they have been recorded in several habitats, including coral reefs, coastal lagoons, the continental shelf and the deep sea. Records from coastal lagoons relate to Chacahua-Pastoria (Mendoza-Vallejo et al. 2009), Mar Muerto (Chávez 1979; Tapia-García et al. 1998), and the Huave Lagoon System (Castro-Aguirre 1982; Tapia-García and MendozaRodríguez 2005). However, there is still no data for the Corralero and Mar Tileme lagoons. In addition, there is a lack of information on near-shore coastal fish from southeastern Oaxaca (Armenta to Salina Cruz). A differential effort is evident in the records of species from the studied groups. For example, while records for macroalgae, crustaceans, and mollusks were published from the second half of the 19th century, studies of the remaining groups began to appear in the early or late 20th century. A problem associated with older records is that they were based on species diagnoses that may have since been updated. To resolve this issue, it is necessary to evaluate the voucher material. Nevertheless, several references that include new records lack voucher collection numbers, contrary to what has been recommended as a result of recent initiatives (e.g., Salazar-Vallejo et al. 2007). Taxonomic re-evaluations of several taxa would also improve the accuracy of the analysis of the marine species richness of Oaxaca. For example, the recent reevaluation of several Genera of macroalgae (e.g., Riosmena-Rodríguez et al. 2009) in the Gulf of California has demonstrated that species richness there had been overestimated. In the same way, the revision of serpulid polychaetes from the Eastern Pacific (Bastida-Zavala 2008) has led to a reduction in the number of species recorded. In addition, an update of the taxonomy of the mollusk bivalves is recommended based on recent publications (e.g., Coan and Valentich-Scott 2012). Table 1. Number of species recorded by taxonomic group in Oaxaca. Taxa with coastal, freshwater and/or terrestrial species, plus marine species, are marked with an asterisk (*). In fish, amphidromus and catadromus species were considered. In the case of reptiles and birds, only species associated with, or living, in adjacent areas to, aquatic bodies were considered. The rest of the taxa are exclusively marine groups. The number of references consulted was included. Taxa Number of species Macroalgae Sponges Cnidarians Flatworms Nemerteans Annelids Crustaceans* Mollusks Bryozoans Echinoderms Fish* Amphibian and reptilian* Birds* Mammals* Total 242 17 118 8 1 222 265 462 1 59 594 37 120 11 2,157 % References consulted 11.2 0.8 5.5 0.4 0.1 10.3 12.3 21.4 0.1 2.7 27.5 1.7 5.6 0.5 15 9 25 4 1 30 68 14 2 12 11 1 2 4 198 Checklists The information for each major group, at the Phylum or Class level, together with a checklist of the species recorded in Oaxaca, is summarized in the following sections. The adopted classification scheme is stated for each group. For practical purposes species and supra-specific taxa are presented in a strictly alphabetical arrangement. Macroalgae Kingdoms Plantae and Chromista A historical overview of the phycological studies focused on Oaxaca State from 1847 to 1994 has been provided by León-Álvarez and González-González (1993) and León-Tejera and González-González (1993). These authors also present the first macroalgal checklists for the region, listing a total of 178 species from 21 localities using both historical records and new collected samples. The present checklist represents an updated version based on published records (León-Tejera et al. 1993; Galindo-Villegas et al. 1997; Mateo-Cid and MendozaGonzález 1997; Mendoza-González and Mateo-Cid 1998; 1999; Dreckmann 1999; Lara-Isassi et al. 2000; Fragoso Table 2. Comparison of the number of species by selected taxonomic group in the Gulf of California, Gulf of Mexico, and Oaxaca. Sources: 1) Hendrickx et al. (2005); 2) Felder and Camp (2009); 3) Brusca et al. (2005); 4) Solís-Marín et al. (2005); 5) Guiry and Guiry (2010); 6) Fredericq et al. (2009). Taxa Fish Mollusks Crustaceans Macroalgae Annelids Cnidarians Echinoderms Sponges Bryozoans Total Oaxaca 15,000 km2 594 462 265 242 222 118 59 17 1 1,980 Gulf of California 1) 160,000 km2 891 3) 2,193 1,025 320 5) 713 253 193 4) 85 168 5,910 % in relation to the Gulf of California 66.7% 21.1% 25.9% 75.6% 31.1% 46.6% 30.6% 20.0% 0.6% 33.5% Gulf of Mexico 2) 1,600,000 km2 1,541 2,445 1,967 673 6) 854 327 522 339 266 8,934 % in relation to the Gulf of Mexico 38.6% 18.9% 13.4% 36.0% 26.0% 36.1% 11.3% 5.0% 0.4% 22.2% 331 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca and Rodríguez 2002; Ávila-Ortiz 2003; Mateo-Cid et al. 2005; 2008; Aguilar-Rosas et al. 2009; Mateo-Cid and Mendoza-González 2009). In total, 242 species were found (30 Orders, 50 Families and 103 Genera) (Table 3) and reported from 16 new localities that were not referred to in the previous accounts of León-Álvarez and GonzálezGonzález (1993) and León-Tejera and González-González (1993). The current names were included in the checklist, while synonymous names, according to Guiry and Guiry (2010), were avoided. Most of the species records in the region were the result of floristic studies. Taxonomic accounts revising recent and historical material, including analysis of information about type material, have been published for species from just 14 Genera (Dreckmann 1999; Fragoso and Rodríguez 2002; Ávila-Ortiz 2003; Mateo-Cid et al. 2005; Mateo-Cid and Mendoza-González 2009). Finally, the real marine macroalgae biodiversity in Oaxaca is still unknown, and will not be resolved until taxonomic evaluations and the revision of voucher specimens are completed. Kingdom Animalia Porifera The sponges are well studied in the Gulf of Mexico, the Caribbean (Gómez 2002), and the Gulf of California (Brusca 1980; Green and Gómez 1986; Gómez et al. 2002; Carballo et al. 2004; 2008a, b). However, studies are limited in number for this group in Oaxaca, and very recently new records of boring sponges have been reported for coral reefs. Only 17 sponge species of 11 Genera and eight Families (Table 4) have been recorded (Dickinson 1945; Gómez and Bakus 1992; Carballo et al. 2003; Bautista-Guerrero et al. 2006; Carballo et al. 2007; 2008a; Cruz-Barraza and Carballo 2008; Carballo and Cruz-Barraza 2010; Cruz-Barraza et al. 2011). GamboaContreras and Tapia-García (1998) recorded 12 species from the continental shelf of the Gulf of Tehuantepec; most of the collection sites were however from Chiapas State. Cnidaria To date, 118 species of 74 Genera and 45 Families of cnidarians have been reported to occur along the coast of Oaxaca (Table 5). Of these, 74 belong to Hydrozoa, 41 to Anthozoa, and four to Scyphozoa. Species of Cubozoa have never been recorded. The most extensively studied group of cnidarians in the Oaxaca area by far is Scleractinia. Topics as varied as the distribution (Leyte-Morales 1997; Glynn and Leyte-Morales 1997; Reyes-Bonilla and Leyte-Morales 1998; Cruz-Piñón and Reyes-Bonilla 1999; Reyes-Bonilla et al. 2005), species composition (López-Pérez et al. 2002; López-Pérez and Hernández-Ballesteros 2004), recruitment (López-Pérez et al. 2007), bioerosion (Herrera-Escalante et al. 2005), and biogeography (Reyes-Bonilla and López-Pérez 1998; Reyes-Bonilla and Cruz-Piñón 2000) of scleractinians have received the most attention. In particular, the basic nomenclature and distribution of stony coral species in the area is relatively well established (Reyes-Bonilla et al. 2005). However, new records of ahermatypic species are expected in the near future, as the bathymetric and geographic extent of ongoing studies widens to include currently unexplored areas. New records of hermatypic species are also expected to increase, although to a lesser extent, as unrecorded Central American coral species disperse into the Mexican Pacific area, as suggested by biogeographic studies (Ketchum-Mejía and Reyes-Bonilla 1997; Reyes-Bonilla and López-Pérez 1998), probably as a consequence of the prevailing current in the western Mexican Pacific (Kessler 2006). In contrast to Scleractinia, less attention has been devoted to Hydrozoa, Scyphozoa (Fraser 1938a, b; 1948; Chávez 1979; Segura-Puertas 1984; Ocaña-Luna and Gómez-Aguirre 1999; Calder et al. 2009), and is absent for Cubozoa. In particular, the taxonomy of these groups has been poorly resolved in the area. For example, three out of 118 species were identified at the supraspecific level but were included because they represent Genera or Families with unique records (Luke 1998), whereas four species are of doubtful identity and require further investigation (species inquirenda) and two more taxa possess a name of unknown or doubtful application (nomen dubium). Concurrently, it is relevant that the 78 reported hydrozoan and scyphozoan species (Table 5) have been addressed in only eight papers, in contrast to the 14 published articles on scleractinians, in which only 41 species were reported. These observations indicate that there are potentially far more cnidarians on the coast of Oaxaca in addition to the stony corals. This finding presents a clear message to institutions and taxonomists that there still remain large unexplored areas, as well as the largely ignored cnidarian groups in the Mexican Pacific. Platyhelmintes The Phylum Platyhelmintes is poorly studied in Oaxaca. Only eight species from seven Genera and seven Families of flatworms have been recorded (Table 6). Of these, two species belong to the Class Monogenea and six to Trematoda (Manter 1937; Meserve 1938; Manter 1940; Gámez-Vivaldo et al. 2006). All these species are parasites of either marine fish or sea turtles. Free-living flatworms (Class Turbellaria) and parasitic flatworms of the Class Cestoda have never been recorded. Nemertina This group is poorly studied in the Mexican Pacific. Only one nemertean species (Table 7) has been recorded on the coast of Oaxaca (Coe 1940). Annelida The annelid fauna from Oaxaca has been the subject of limited study (Bastida-Zavala and Guevara-Cruz 2012). Only 222 species of 130 Genera and 42 Families, mostly of the Polychaete Class, have been recorded in 30 papers (Table 8). This number is limited compared to the over 700 species recorded in the Gulf of California (Hendrickx 2005). Most records were taken from extensive monographs, including Chamberlin (1919), Hartman (1939; 1940; 1941; 1944; 1950; 1961), Treadwell (1941), Rioja (1962), Fauchald (1968; 1970; 1972), Bastida-Zavala and ten Hove (2003) and Bastida-Zavala (2008). In addition, a few papers have focused on particular localities in Oaxaca, such as the studies by Gómez et al. (1997), HernándezAlcántara et al. (2003) and Tovar-Hernández and SalazarVallejo (2008), or on the continental shelf of the Gulf of 332 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Tehuantepec, including studies by Hernández-Alcántara et al. (1994), González-Ortiz et al. (1996; 1997), HernándezAlcántara and Solís-Weiss (2000), Solís-Weiss et al. (2000) and Hernández-Alcántara et al. (2006). Other records were sourced from studies about pelagic polychaetes, such as Dales (1957), Fernández-Álamo (1987; 2000; 2002) and Fernández-Álamo and Sanvicente-Añorve (2005). Pelagic polychaetes comprise 20 species (9% of the polychaete records), with the remaining 201 species (91%) being benthic polychaetes. Only two polychaete Families have been reviewed for the Oaxaca region: the Eunicidae and Serpulidae (BastidaZavala 2008; Ruiz-Cancino et al. 2011). The Eunicidae Family was the highest represented, with 21 species (9.5%), followed by Serpulidae with 18 species (8.1%), and Spionidae with 17 species (7.7%). Some studies that include lists of species from the continental shelf of the Gulf of Tehuantepec, such as the study by Gamboa-Contreras and Tapia-García (1998), do not indicate whether species were recorded in Oaxaca or Chiapas States. These records were for this reason not included in the present checklist. The previous list compiled by Salazar-Vallejo (1989) was the primary basis of this updated list. However, several species names (139 species, 62.9%) are questionable, either because the records have not yet been confirmed by experts, or because the names correspond to species from areas the distribution of which is at some distance from the Tropical Eastern Pacific (according to Hartman 1959; Salazar-Vallejo and Londoño-Mesa 2004; BastidaZavala 2008; de León-González et al. 2009; Ruiz-Cancino et al. 2011). At least one species, Hydroides sanctaecrucis Krøyer in Mörch, 1863, is an exotic species (Bastida-Zavala 2008). Additionally, one record of a hirudinean ectoparasite of the olive ridley turtle, Lepidochelys olivacea (Eschscholtz, 1829), was included (Gámez-Vivaldo et al. 2006). Crustaceans Little is known about crustaceans in Oaxaca. Different checklists from the Tropical Eastern Pacific (e.g., Hendrickx 1993; Hendrickx and Harvey 1999; GarcíaMadrigal 2008) reveal a disparity between records from the Gulf of California and those from the Mexican South Pacific (García-Madrigal et al. 2012). A total of 265 species and 153 Genera from 72 Families were obtained, based on 68 references (Table 9). These records were gathered from two sources: 1) isolated records from monographs and papers mainly related to large expeditions (e.g., Velero, Zaca, Searcher, Albatross Expeditions) from the Pacific Coast of America (Benedict 1903; Rathbun 1918; 1925; 1930; 1937; Maccagno 1928; Wilson 1937; Schmitt 1940; Holthuis 1952; Barnard 1954; Holthuis and VillalobosFigueroa 1962; Garth 1958; 1959; 1961; 1966; Haig 1957; 1960; 1968; Garth and Stephenson 1966; VillalobosFigueroa 1967; Efford 1971; Ball 1972; Manning 1972; Ball and Haig 1974; Glynn et al. 1975; Crane 1947; 1975; Luke 1977; Barnard and Barnard 1982; Wicksten 1978; 1983; 1984; Poore 1984; Pérez-Farfante 1985; Kim and Abele 1988; Brusca et al. 1992; Castro 1996); and 2) papers focused on ecology, fisheries, descriptions of new species or faunal studies of particular localities in Oaxaca (Howard 1952; Sosa-Hernández et al. 1980; Carvacho and Haasmann 1984; Hendrickx and Wicksten 1989; LópezCortés 1990; Villalobos-Hiriart and Nates-Rodríguez 1990; Wicksten and Hendrickx 1992; Hendrickx 1996; 1997; 1999; 2005; Hendrickx and Esparza-Haro 1997; Hendrickx et al. 1997: Hendrickx and Espinosa-Pérez 1998; Hendrickx and Vázquez-Cureño 1998; Hernández-Aguilera 1998; Hendrickx and Harvey 1999; Barbosa-Ledesma et al. 2000; Ramírez-Luna et al. 2002; Boyko 2002; ÁlvarezSilva et al. 2003; Hernández-Álvarez and Villalobos-Hiriart 2005; Gámez-Vivaldo et al. 2006; Martínez-Guerrero 2007; Ayón-Parente and Hendrickx 2007; 2009; García-Madrigal 2010; García-Madrigal and Andreu-Sánchez 2010; JarquínGonzález and García-Madrigal 2010). Studies incorporating species lists from the continental shelf of the Gulf of Tehuantepec, such as that by GamboaContreras and Tapia-García (1998), or pelagic copepods, e.g. Fernández-Álamo et al. (2000), were not included in this checklist, because they do not indicate whether the species was recorded in Oaxaca or Chiapas State. Mollusks Mollusks are one of the best-known invertebrate groups on the Pacific coast. Taxonomic studies began in the second half of the 19th century (García-Cubas 1986; Reguero-Reza and García-Cubas 1993; González 1993; Naranjo-García 2003). Most species collected were published in a monograph by Keen (1971); there are 3,345 species of mollusks from Baja California to Peru (Keen 1971). Skoglund (2001; 2002) recorded 2,918 gastropods and 111 chitons and, recently, Coan and Valentich-Scott (2012) have recorded 890 bivalves. There have been few mollusk studies on the coast of Oaxaca. Inventories of the mollusk fauna have been carried out on the continental shelf (Gamboa-Contreras and Tapia-García 1998; Ríos-Jara et al. 2009) and on the coastline (Piña-Arce and Herrera-Peña 1980) of the Gulf of Tehuantepec, and have focused on the central and southern coast of the state (Holguín-Quiñones and González-Pedraza 1989; de León-Herrera 2000; Ramírez-González and Barrientos-Luján 2007) and on the Classes of cephalopods (Sánchez 2003) and polyplacophorans (Reyes-Gómez et al. 2010). Studies on mollusk communities have been carried out at the rocky intertidal zone (Zamorano et al. 2008), coral reefs (Rodríguez-Palacios et al. 1988; Zamorano et al. 2006) and lagoon systems (Zamorano et al. 2010). There have additionally been some studies aimed at shellfish of economic importance, such as the pen shell Atrina maura (G.B. Sowerby I, 1835) (Ahumada-Sempoal 1998; Ángel-Pérez et al. 2007) and the purple snail Plicopurpura columelaris (Lamarck, 1816) (Castillo-Rodríguez and Amezcua-Linares 1992; Vázquez-Gil et al. 2004). The studies of Gamboa-Contreras and Tapia-García (1998) and Alejo-Plata et al. (2001), based on the continental shelf of the Gulf of Tehuantepec, were not included in the list because the authors do not indicate whether the species were recorded in Oaxaca or Chiapas State. In this review, the mollusk fauna of the coast of Oaxaca yielded 109 Families, 258 Genera and 462 species (Table 10). Of these, 281 were Gastropoda, 153 Bivalvia, 22 Polyplacophora and six Cephalopoda. Mollusks that lack economic importance, and that are small-sized, 333 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca inconspicuous, rare and/or living in deep waters, such as Scaphopoda, have not yet been studied (Ríos-Jara et al. 2003). Bryozoans Only two references were found for bryozoans (Osburn 1953; Soule et al. 1995); the authors recorded the same species (Table 11). Echinoderms Although echinoderms on the Pacific coast of Mexico have been studied since the late 19th century (Buitrón and Solís-Marín 1993), these studies did not cover the entire coast. During the 1970s, data for specimens collected in Mexican waters and deposited in foreign collections were reviewed with a number of species from the coast of Oaxaca being reported (Caso 1978; 1980; 1983). Solís-Marín et al. (1993) published a taxonomic list for the asteroids and echinoids of Mexico. After a bibliographic revision and a review of the Echinoderm National Collection (Instituto de Ciencias del Mar y Limnología, UNAM), the authors reported just six species of echinoderms for Oaxaca. There are now corrected and updated regional taxonomic lists for various localities, particularly from the north of Mexico (Bautista-Romero et al. 1994; Solís-Marín et al. 1997; Cintra-Buenrostro 2001; Solís-Marín et al. 2005; 2009), and the Mexican Caribbean and Gulf of Mexico (Laguarda-Figueras et al. 2009). However, these lists remain incomplete for the rest of the country. Only a few basic studies have been carried out along the coast of Oaxaca, providing sparse data about the species inhabiting this zone. Deichmann (1938; 1941; 1958) recorded nine species of holothurians from the coast of Oaxaca. Benítez-Villalobos (2001) compared two reef communities of Bahías de Huatulco, recording six species of echinoderms. Zamorano and Leyte-Morales (2005) reported seasonal changes in echinoderm diversity in a bay of the Parque Nacional Huatulco, providing a list of 24 species. Honey-Escandón et al. (2008) compiled a taxonomic list for the echinoderms of the Mexican Pacific, mentioning 30 species from the coast of Oaxaca. Benítez-Villalobos et al. (2008) presented an updated list of echinoids and asteroids from the coast of Oaxaca, reporting 11 new records for this state. Granja-Fernández and López-Pérez (2011) recorded 13 species of ophiuroids from Puerto Escondido, including three new records for Oaxaca. For the general taxonomic organization of the list, we used the criteria suggested by the following authors: Class Asteroidea, Clark (1989; 1993; 1996); Class Ophiuroidea, Matsumoto (1915) and Fell (1960); Class Echinoidea, Mortensen (1928; 1935; 1940; 1943; 1948; 1950; 1951); Class Holothuroidea, Pawson and Fell (1965). The taxonomic list obtained from the bibliographic revision comprises 59 species from 47 Genera and 31 Families, which were distributed across four Classes (Table 12). Fish In order to review the current status of fish species recorded from the coast of Oaxaca, a search was carried out of information published in books (Allen et al. 1995; Compagno et al. 1995; McEachran and Notarbartolo-diSciara 1995; Castro-Aguirre and Espinosa-Pérez 1996; Tapia-García and Gutiérrez-Díaz 1998; Castro-Aguirre et al. 1999; Espinosa-Pérez et al. 2004), one electronic database (Robertson and Allen 2008) and journals (Chávez 1979; Castro-Aguirre 1982; Tapia-García et al. 1998; TapiaGarcía and Mendoza-Rodríguez 2005; Ramírez-Gutiérrez et al. 2007; Mendoza-Vallejo et al. 2009; López-Pérez et al. 2010). Each species was confirmed using recent data to verify synonyms. Only seven studies included lists of species from the coast of Oaxaca, mainly from coastal lagoons. These studies included: Chacahua (MendozaVallejo et al. 2009), Mar Muerto (Chávez 1979; Tapia-García et al. 1998) and the Huave Lagoon System (Castro-Aguirre 1982; Tapia-García and Mendoza-Rodríguez 2005). In addition, there are only two published lists for coral reef fishes: Ramírez-Gutiérrez et al. (2007) at a locality of San Agustin Bay, Santa María Huatulco, and López-Pérez et al. (2010) from Mazunte to Huatulco. An updated list of fish species, including teleost and elasmobranches, has been provided (Table 13). In total, 594 species were recorded off the coast of Oaxaca, which corresponds to 313 Genera and 113 Families. Perciformes is the best represented Order, with 303 species. One endemic species was recorded, Poeciliopsis fasciata (Meek, 1904), which is distributed only in Chiapas and Oaxaca. Amphibians and reptiles Five of the seven extant marine turtle species nest on the coast of Oaxaca (Table 14). La Escobilla is in fact probably the most important nesting beach in the world for the olive ridley Lepidochelys olivacea (Márquez 1990). Other important marine turtle nesting beaches are Chacahua, la Ventanilla, Morro Ayuta and Barra de la Cruz. Eight leatherback turtles Dermochelys coriacea (Vandelli, 1761) were recorded nesting in one night on Barra de la Cruz beach in the winter of 2001 (Meraz-Hernando pers. obs.). Several reports have been compiled for L. olivacea (Meraz-Hernando and Ávila-Barrientos 2007a, b) and D. coriacea (Sánchez-Díaz and Meraz-Hernando 2001). The American crocodile Crocodylus acutus Cuvier, 1807 is a resident species at several locations along the coast, and it is even possible to observe adults moving between areas by sea. Reports of this species along the coast of Oaxaca have highlighted the importance of crocodiles in the state (Meraz-Hernando and Ávila-Nahón 2006; Brandon-Pliego 2007; García-Grajales et al. 2007; Meraz-Hernando et al. 2008). There is a single record of the sea snake Pelamis platurus (Linnaeus, 1758) (Meraz-Hernando 2007b), and one of a captive iguana Ctenosaura pectinata Wiegmann, 1834 (Villegas-Zurita 1999). Oaxaca has several endemic amphibian and reptile species, with the turtle Kinosternon oaxacae Berry and Iverson, 1980 from the Colotepec and Tonameca rivers (Berry and Iverson 1980) being important for the coast, and the iguana Ctenosaura oaxacana Gray, 1842 from the Tehuantepec Isthmus (Köhler 2004). Limited attention has been given to the amphibians (Table 14) that inhabit the coast of Oaxaca. There is only one report of the cane toad Rhinella marina (Linnaeus, 1758) (Becerril-Morales 2001). In addition, there are a few records covering the herpetofauna of the state, but 334 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca these are from restricted areas (Smith 1959; Lynch and Smith 1966). The present list of species is based on a chapter from Casas-Andreu et al. (2004), and includes records obtained from several surveys along the coast of Oaxaca, as well as from captive species held at the Turtle Museum at the Mexican Center for the Study of the Turtles, on the coast of Oaxaca. In total, 37 species in 30 Genera and 15 Families were recorded (Table 14). Some reports on freshwater turtles have been included (Ippi and Flores 2001). Frost (2012) was followed for taxonomy and nomenclature of amphibians. Major changes were added for reptiles with the inclusion of the Family Colubridae and the new Family Dipsadidae from Zaher et al. (2009). Birds Waterbird richness on the coast of Oaxaca is high, due to the rich diversity of habitats. In total, 120 species of 70 Genera and 26 Families have been recorded (Table 15). Ardeidae and Laridae are amongst the Families with the highest richness; they mainly inhabit lagoons, rivers, estuaries and cliffs. Nevertheless, the number of seabird species is relatively low, maybe because of the low abundance of small pelagic fish. The present list of species was compiled from a comprehensive species account by Navarro-Sigüenza et al. (2004), in which species from the coast and the Tehuantepec Isthmus to the limits of the state were documented (Table 15). The list was restricted to species with a confirmed record in Binford (1989) and subsequent reports (Winker et al. 1992; Schaldach et al. 1997; Mellink et al. 1998; Meraz-Hernando 2001; Meraz-Hernando and González-Bravo 2005; González-Bravo and MerazHernando 2010). In addition, world distribution reports were incorporated into the list for boobies (Nelson 1978), shorebirds (Paulson 1993), gulls (Olsen 2000), and pelicans and cormorants (Nelson 2005). Due to several taxonomic changes made in recent years, the classification of the American Ornithologist’s Union was followed, and was also based on the AOU checklist (AOU 1998) and subsequent supplements (AOU 2000; Banks et al. 2002; 2004; 2006; 2007; 2008; Chesser et al. 2010). It is important to note that the present list includes only aquatic birds for which records have been confirmed. Some non-aquatic birds have been included because they were observed in close proximity to aquatic bodies during activities such as feeding, nesting or when seeking refuge. Marine mammals The Mexican coastal area is divided into four regions, with Oaxaca being part of Zone 3 or tropical Pacific (Torres et al. 1995). The tropical Mexican Pacific is estimated to include 18 marine mammals, and is therefore more diverse than the Gulf of Mexico and the Caribbean, but with a richness that is lower than the north Pacific and the Gulf of California (Aguayo-Lobo et al. 1990). The abrupt shape of the coastline in Oaxaca, including bays, cliffs and islands, is a refuge for several resident cetacean species. Furthermore, the influence of the cold California Current during winter (Pacheco-Sandoval 1991) promotes the presence of migrating species such as Megaptera novaeangliae (Borowski, 1781). Marine mammal studies have primarily been undertaken over the last two decades, with some recent records being based on occasional sightings (Sánchez-Díaz and Meraz-Hernando 2001; Meraz-Hernando 2003) and strandings (Meraz-Hernando and Becerril-Morales 2004; Meraz-Hernando 2007a; Lira-Torres 2007). Some tentative lists contain species for which there are both possible and confirmed sightings (Aurioles-Gamboa 1993; Salinas and Ladrón-de-Guevara 1993). Based on marine surveys, strandings, and sightings from the coast during a period of over five years, a detailed report on marine mammals has been compiled for the central coast of Oaxaca (Meraz-Hernando and Sánchez-Díaz 2008) and used as the main reference for the present list (Table 16). Other species that are likely to be present in the state were added from reliable reports (Carwardine 1995; Jefferson and Leatherwood 1995; Medrano-González et al. 2008) and databases of the Mexican national commission for Biodiversity (CONABIO). One species of particular interest is Zalophus californianus (Lesson, 1828), which has been sighted in Oaxaca on several occasions over a period of several years. In the case of the list derived from Medrano-González et al. (2008), species recorded for the Tropical Pacific were considered. Two species, the ziphidae Mesoplodon peruvianus Reyes, Mead and Van Waerebeek, 1991 and the otariid Arctocephalus galapagoensis Heller, 1904, are likely to occur in Oaxaca but were not considered due to the reduced number of sightings in Mexico (Urbán-Ramírez and Guerrero-Ruiz 2008). The list comprises 11 species from 10 Genera and three Families (Table 16). Conclusions The review of 198 references for the flora and fauna of Oaxaca produced 2,157 species records, corresponding to 15 taxonomic groups (Division, Phyla or Class level) (Table 1). It was evident that floristic and faunistic knowledge is heterogeneous among the taxonomic groups. Additionally, many invertebrate groups that have been recorded in adjacent regions have not yet been recorded on the coast of Oaxaca, such as some platyhelminths (turbellarians, cestods), rotifers, nematodes, oligochaetes, sipunculids, echiurans, tardigrades, pycnogonids, some crustaceans (branchiopods, mysids, cumaceans and ostracods), brachiopods, chaetognaths, ascidians and cephalochordates. The inventory of marine flora and fauna is far from complete, and additional effort is needed. The region could potentially be characterized as a marine hotspot, similar to the adjacent terrestrial biodiversity. This represents a major scientific gap and an opportunity for taxonomists. 335 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 3. Checklist of the macroalgae recorded in marine areas from Oaxaca, Mexico. When the first published record of the species was a synonymous of a currently accepted name, this was added after ‘=’ symbol. The nomenclature and classification followed Guiry and Guiry (2010). Kingdom Plantae Haeckel, 1866 Reference Division Chlorophyta Reichenbach, 1834 Class Bryopsidophyceae Bessey, 1907 Order Bryopsidales J.H. Schaffner, 1922 Family Bryopsidaceae Bory de Saint-Vincent, 1829 Bryopsis hypnoides J.V. Lamouroux, 1809 Bryopsis muscosa J.V. Lamouroux, 1809 Mendoza-González and Mateo-Cid (1998) León-Tejera and González-González (1993) Bryopsis pennata var. minor J. Agardh, 1887 =Bryopsis pennatula J. Agardh León-Tejera and González-González (1993) Caulerpa imbricata G. Murray, 1887 =Caulerpa peltata Lamouroux f. imbricata (G. Murray) Webervan Bosse 1898 Caulerpa peltata J.V. Lamouroux, 1809 León-Tejera et al. (1993) Caulerpa racemosa (Forsskal) J. Agardh, 1873 León-Tejera and González-González (1993) Bryopsis plumosa (Hudson) C. Agardh, 1823 Family Caulerpaceae Kützing, 1843 Caulerpa sertularioides (S. Gmelin) M.A. Howe, 1905 Caulerpa sertularioides f. brevipes (J. Agardh) Svedelius, 1906 Caulerpa sertularioides f. longiseta (Bory de Saint-Vincent) Svedelius, 1906 Caulerpella ambigua (Okamura) Prud’homme van Reine and Lokhorst, 1992 =Caulerpa vickersiae Borgesen 1911 Family Codiaceae Kützing, 1843 Codium dichotomum S.F. Gray, 1821 Codium edule P.C. Silva, 1952 Codium giraffa P.C. Silva, 1979 Codium setchelii N.L. Gardner, 1919 Family Derbesiaceae Hauck, 1884 Derbesia hollenbergii W.R. Taylor, 1945 Family Halimedaceae Link, 1832 León-Tejera and González-González (1993) Mendoza-González and Mateo-Cid (1998) León-Tejera et al. (1993) León-Tejera and González-González (1993) León-Tejera and González-González (1993) León-Tejera et al. (1993) León-Tejera and González-González (1993) León-Tejera et al. (1993) Mendoza-González and Mateo-Cid (1998) León-Álvarez and González-González (1993) Mendoza-González and Mateo-Cid (1999) Halimeda discoidea Decaisne, 1842 León-Tejera et al. (1993) Chlorodesmis hildenbrandtii A. Gepp and E.S. Geep, 1911 León-Tejera et al. (1993) Halimeda tuna (J. Ellis and Solander) J.V. Lamouroux, 1846 Family Udoteaceae J. Agardh, 1887 Chlorodesmis mexicana W.R. Taylor, 1945 Class Ulvophyceae K.R. Mattox and K.D. Stewart, 1978 Order Cladophorales Haeckel, 1894 Family Cladophoraceae W. Wille ex Warming, 1884 Chaetomorpha anteninna (Bory de Saint-Vincent) Kützing, 1847 Mendoza-González and Mateo-Cid (1998) Mendoza-González and Mateo-Cid (1999) León-Tejera and González-González (1993) Chaetomorpha brachygona Harvey, 1858 León-Tejera and González-González (1993) Chaetomorpha gracilis Kützing, 1845 León-Tejera and González-González (1993) Chaetomorpha californica F.S. Collins, 1900 Chaetomorpha exposita (Borgesen) E.Y. Dawson, 1954 Chaetomorpha linum (O.F. Müller) Kützing, 1845 Cladophora albida (Nees) Kützing, 1843 Cladophora brasiliana G. Martens 1868 Cladophora lehmanniana (Lindenberg) Kützing, 1843 =Cladophora utriculosa Kützing 1843 Cladophora microcladioides F.S. Collins, 1909 Cladophora nigrescens Zanardini ex Frauenfeld, 1855 Cladophora perpusilla Skottsberg and Levring ex Levring, 1941 Cladophora prolifera (Roth) Kützing, 1843 Cladophora sericea (Hudson) Kützing, 1843 Rhizoclonium africanum Kützing, 1853 León-Tejera et al. (1993) León-Tejera and González-González (1993) León-Tejera and González-González (1993) León-Tejera et al. (1993) León-Tejera and González-González (1993) León-Tejera and González-González (1993) León-Tejera et al. (1993) León-Tejera et al. (1993) León-Tejera and González-González (1993) Mendoza-González and Mateo-Cid (1998) Mateo-Cid and Mendoza-González (1997) Mateo-Cid and Mendoza-González (1997) Rhizoclonium riparium (Roth) Harvey, 1849 =Rhizoclonium implexum (Dillwyn) Kützing Mateo-Cid et al. (2005) Batophora oerstedii J. Agardh, 1854 Mendoza-González and Mateo-Cid (1998) Order Dasycladales Pascher, 1931 Family Dasycladaceae Family Polyphysaceae Kützing, 1843 Acetabularia caliculus J.V. Lamouroux, 1824 Acetabularia crenulata J.V. Lamouroux, 1816 Parvocaulis parvulus (Solms-Laubach) Berger, Fettweiss, Gleissberg, Liddle, Richter, Sawitsky and Zuccarello, 2003 =Acetabularia parvula Solms-Laubach 1895 Order Ctenocladales P.C. Silva, 1982 Family Ulvellaceae Schmidle, 1899 Acrochaete viridis (Reinke) R. Nielsen, 1979 = Entocladia viridis Reinke,1876 Order Ulvales Blackman and Tansley, 1902 León-Tejera and González-González (1993) Mendoza-González and Mateo-Cid (1998) León-Tejera et al. (1993) Mendoza-González and Mateo-Cid (1999) 336 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 3. Continued. Family Phaeophilaceae Chappell, O’Kelly, Wilcox and Floyd, 1990 Phaeophila dendroides (P.L. Crouan and H.M. Crouan) Batters, 1902 Phaeophila floridearum Hauck, 1867 = Phaeophila engleri Reinke 1889 Family Ulvaceae Lamouroux ex Dumortier, 1822 Blidingia marginata (J. Agardh) P.J.L. Dangeard, 1958 =Enteromorpha marginata J. Agardh Mendoza-González and Mateo-Cid (1998) León-Tejera and González-González (1993) León-Tejera and González-González (1993) Enteromorpha ramulosa (J.E. Smith) Carimichael, 1833 León-Tejera and González-González (1993) Ulva clathrata (Roth) C. Agardh, 1811 = Enteromorpha clathrata (Roth) Greville 1830 León-Tejera and González-González (1993) Ulva acanthophora (Kützing) Hayden, Blomster, Maggs, P.C. Silva, Stanhope and J.R. Waaland, 2003 = Enteromorpha acanthopora Kützing, 1879 Ulva compressa Linnaeus, 1753 =Enteromorpha compressa (Linnaeus) Nees 1820 Ulva flexuosa Wulfen, 1803 =Enteromorpha lingulata J. Agardh 1883 Ulva intestinalis Linnaeus, 1753 = Enteromorpha intestinalis (Linnaeus) Nees 1820 León-Tejera and González-González (1993) León-Tejera et al. (1993) León-Tejera and González-González (1993) León-Tejera and González-González (1993) Ulva lactuca Linnaeus, 1753 León-Tejera et al. (1993) Ulva spinulosa Okamura and Segawa, 1936 León-Tejera and González-González (1993) Ulva linza Linnaeus, 1753 =Enteromorpha linza (Linnaeus) J. Agardh 1883 Ulva rigida C. Agardh, 1823 Order Siphonocladales (Blackman and Tansley) Oltmanns, 1904 Family Boodleaceae Borgesen, 1925 Phyllodictyon anastomosans (Harvey) Kraft and M.J. Wynne, 1996 =Struvea anastomosans (Harv.) Piccone and Grunow ex Piccone 1884 Division Rhodophyta Wettstein 1922 León-Tejera and González-González (1993) Mendoza-González and Mateo-Cid 1998 León-Tejera et al. (1993) Class Bangiophyceae Wettstein 1901 Order Bangiales Schmitz ex Engler 1892 Family Bangiaceae Engler 1892 Bangia enteromorphides E.Y. Dawson, 1953 Class Compsopogonophyceae G.W. Saunders and Hommersand, 2004 Order Erythropeltidales Garbary, Hansen and Scagel, 1980 Family Erythrotrichiaceae G.M. Smith, 1933 Erythrocladia irregularis Ronsenvinge, 1909 Erythrocladia pinnata W.R.Taylor, 1942 Erythrotrichia carnea (Dillwyn) J. Agardh, 1883 Sahlingia subintegra (Rosenvinge) Kornmann, 1989 Class Porphyridiophyceae Kylin, 1937 Order Porphyridiales Kylin, 1937 Family Porphyridiaceae Kylin, 1937 Bangiopsis dumontoides (P.L. Crouan and H.M. Crouan) V. Krishnmurthy, 1957 Stylonema alsidii (Zanardini) K.M. Drew, 1956 =Goniotrichum alsidii (Zanardini) M.A. Howe 1914 Class Florideophyceae Cronquist, 1960 Order Acrochaetiales Feldmann, 1953 Mateo-Cid et al. (2005) León-Tejera and González-González (1993) León-Tejera and González-González (1993) León-Tejera and González-González (1993) Mendoza-González and Mateo-Cid (1998) León-Tejera and González-González (1993) León-Tejera and González-González (1993) Family Acrochaetiaceae Fritsch ex W.R. Taylor, 1957 Acrochaetium microscopicum (Nägeli ex Kützing) Nägeli ex Nägeli and Cramer, 1858 =Audouinella microscopica (Nägeli ex Kützing) Woelkerling 1971 Acrochaetium savianum (Meneghini) Nägeli, 1862 =Audouinella saviana (Meneghini) Woelkerling 1973 Order Ahnfeltiales C.A. Maggs and C.M. Pueschel, 1989 Family Ahnfeltiaceae C.A. Maggs and C.M. Pueschel, 1989 Ahnfeltia plicata (Hudson) E.M. Fries, 1836 Order Bonnemaisoniales Feldmann and Feldmann, 1942 Family Bonnemaisoniaceae Schmitz ex Engler, 1892 Asparagopsis taxiformis (Delile) Trevisan de Saint-Léon, 1845 =Falkenbergia hillebrandii (Bornet) Falkenberg 1901 Order Ceramiales Oltmanns, 1904 Family Ceramiaceae Dumortier, 1822 Anotrichium tenue (C. Agardh) Nägeli, 1862 Anotrichium tenue var. thyrsigerum (Thwaites ex Harvey) Kim and Lee, 1992 = Griffithsia tenuis C. Agardh 1828 Anthithamnionella elegans (Berthold) Price (Berthold) Price and John ex Price, John and Lawson, 1986 Bornetia binderiana (Sonder) Zanardini, 1866 Callithamnion corymbosum (Smith) Lyngbye, 1819 =Callithamnion rupicula C.L. Anderson 1894 Mateo-Cid and Mendoza-González (1997) Mateo-Cid and Mendoza-González (1997) Mendoza-González and Mateo-Cid (1998) León-Tejera and González-González (1993) León-Tejera and González-González (1993) León-Tejera and González-González (1993) Mendoza-González and Mateo-Cid (1998) Mendoza-González and Mateo-Cid (1999) Mendoza-González and Mateo-Cid (1998) Callithamnion rupinicola Anderson, 1894 =Callithamnion uncinatum E.Y. Dawson 1962 León-Tejera and González-González (1993) Ceramium camouii E.Y. Dawson, 1944 Mendoza-González and Mateo-Cid (1998) Centroceras clavulatum (C. Agardh) Montagne, 1846 Ceramium affine var. peninsularis E.Y. Dawson, 1950 León-Tejera and González-González (1993) León-Tejera and González-González (1993) 337 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 3. Continued. Ceramium caudatum Setchell and N.L. Gardner, 1924 Ceramium clarionensis Setchell and N.L. Gardner, 1930 Ceramium gracillimum f. byssoideum (Harvey) Mazoyer, 1938 Ceramium hamatispinum E.Y. Dawson, 1950 León-Tejera and González-González (1993) León-Tejera and González-González (1993) León-Tejera and González-González (1993) Mateo-Cid and Mendoza-González (1997) Ceramium macilentum J. Agardh, 1894 =Ceramium mazatlense E.Y. Dawson 1950 León-Tejera and González-González (1993) Ceramium sinicola Setchell and N.L. Gardner, 1924 Mendoza-González and Mateo-Cid (1999) Ceramium monocanthum J. Agardh, 1894 Ceramium paniculatum Okamura, 1921 Gayliella flaccida (Harvey ex Kützing) Cho and McIvor ex Cho, Boo, Hommersand, Maggs, McIvor and Fredericq, 2008 =Ceramium flaccidum (Harvey ex Kützing) Ardissone 1871 Gayliella taylorii (E.Y. Dawson) Cho and Boo ex Cho, Boo, Hommersand, Maggs, McIvor and Fredericq, 2008 =Ceramium taylorii E.Y. Dawson 1950 Mendoza-González and Mateo-Cid (1998) León-Tejera and González-González (1993) León-Tejera and González-González (1993) Griffithsia pacifica Kylin, 1925 León-Tejera and González-González (1993) Pleonosporium globuliferum Levring, 1941 León-Tejera and González-González (1993) Pleonosporium mexicanum E.Y. Dawson, 1962 Pleonosporium rhizoideum E.Y. Dawson, 1962 Spyridia filamentosa (Wulfen) Harvey ex Hooker, 1833 Family Dasyaceae Kützing, 1843 León-Tejera and González-González (1993) León-Tejera and González-González (1993) León-Tejera and González-González (1993) Mendoza-González and Mateo-Cid (1998) Dasya sinicola var. abyssicola (E.Y. Dawson) E.Y. Dawson, 1963 Mateo-Cid and Mendoza-González (1997) Hypoglossum attenuatum N.L. Gardner, 1927 Mateo-Cid and Mendoza-González (1997) Hypoglossum attenuatum N.L. Gardner. var. abyssicolum (W.R. Taylor) E.Y. Dawson, 1962 León-Tejera and González-González (1993) Family Delesseriaceae Bory, 1828 Cryptopleura peltata (Montagne) M.J.Wynne, 2002 =Cryptopleura corallinarum (Nott) N.L.Gardner 1927 Taenioma perpusillum (J. Agardh) J. Agardh, 1863 =Polysiphonia perpussilla J. Agardh 1847 Family Rhodomelaceae J.E. Areschoug, 1847 Alsidium pusillum E.Y. Dawson, 1963 Chondria arcuata Hollenberg, 1945 Chondria decipiens Kylin, 1941 Herposiphonia hollenbergii E.Y. Dawson, 1963 Herposiphonia secunda (C. Agardh) Ambronn, 1880 Herposiphonia secunda f. tenella (C. Agardh) M.J. Wynne, 1985 Herposiphonia subdisticha Okamura, 1899 Herposiphonia tenella (C.Agardh) Ambronn, 1880 Herposiphonia verticillata (Harvey) Kylin, 1925 Laurencia clarionensis Setchell and N.L. Gardner, 1937 Laurencia hancockii E.Y. Dawson, 1958 León-Tejera and González-González (1993) León-Tejera and González-González (1993) Mendoza-González and Mateo-Cid (1998) León-Tejera et al. (1993) León-Tejera and González-González (1993) León-Tejera and González-González (1993) León-Tejera and González-González (1993) Mendoza-González and Mateo-Cid (1998) León-Tejera and González-González (1993) León-Tejera and González-González (1993) Mateo-Cid et al. (2005) León-Tejera and González-González (1993) Mateo-Cid and Mendoza-González (1997) Laurencia lajolla E.Y. Dawson, 1958 León-Tejera et al. (1993) Neosiphonia simplex (Hollenberg) Y.P. Lee, 2008 =Polysiphonia simplex Hollenberg 1942 León-Tejera and González-González (1993) Laurencia subcorymbosa E.Y. Dawson, 1963 Laurencia voragina W.R. Taylor, 1945 Neosiphonia sphaerocarpa (Borgesen) M.S. Kim and I.K. Lee, 1999 =Polysiphonia sphaerocarpa Boergesen 1918 Polysiphonia confusa Hollenberg 1961 Polysiphonia decussata Hollenberg, 1942 Polysiphonia foetidissima Cocks ex Bornet, 1892 Polysiphonia japonica var. savatieri (Hariot) Yoon, 1986 =Polysiphonia savatieri Hariot 1891 Polysiphonia scoplorum var. villum (J. Agardh) Hollenberg, 1968 Polysiphonia sertularioides (Grateloup) J. Agardh, 1863 =Polysiphonia flaccidisima Hollenberg 1942 Polysiphonia sonorensis Hollenberg, 1942 Tayloriella dictyurus (J. Agardh) Kylin, 1956 Order Corallinales P.C. Silva and H.W. Johansen, 1986 Family Corallinaceae J.V.Lamouroux, 1812 Amphiroa beauvoisii J.V. Lamouroux, 1816 =Amphiroa mexicana W.R. Taylor 1945 Amphiroa crosslandii M. Lemoine, 1929 Amphiroa dimorpha M. Lemoine, 1929 Amphiroa galapaguensis W.R. Taylor, 1945 Amphiroa magdalensis E.Y. Dawson, 1953 Amphiroa minutissima W.R. Taylor, 1945 Amphiroa misakiensis Yendo, 1902 =Amphiroa brevianceps E.Y. Dawson 1953 Amphiroa rigida J.V. Lamouroux, 1816 Amphiroa taylorii E.Y. Dawson, 1953 Amphiroa valonioides Yendo, 1902 León-Tejera and González-González (1993) Mendoza-González and Mateo-Cid (1998) Mendoza-González and Mateo-Cid (1999) Mendoza-González and Mateo-Cid (1998) León-Tejera and González-González (1993) León-Tejera and González-González (1993) Mendoza-González and Mateo-Cid (1998) Mendoza-González and Mateo-Cid (1998) Mendoza-González and Mateo-Cid (1998) León-Tejera and González-González (1993) León-Tejera and González-González (1993) León-Tejera et al. (1993) León-Tejera and González-González (1993) León-Tejera et al. (1993) León-Tejera and González-González (1993) León-Tejera and González-González (1993) León-Tejera and González-González (1993) León-Tejera et al. (1993) León-Tejera and González-González (1993) León-Tejera and González-González (1993) Mendoza-González and Mateo-Cid (1998) 338 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 3. Continued. Hydrolithon farinosum J.V. Lamouroux) D. Penrose and Y.M. Chamberlain, 1993 =Fosliella farinosa (J.V. Lamouroux) M.A. Howe Fragoso and Rodríguez (2002) Hydrolithon rupestre (Foslie) Penrose, 1996 Fragoso and Rodríguez (2002) Hydrolithon onkodes (Heydrich) D. Penrose and Woelkerling, 1992 Hydrolithon samoënse (Foslie) Keats and Y.M. Chamberlain, 1994 =Lithophyllum samoënse Foslie 1906 Jania adhaerens J.V. Lamouroux, 1816 Jania capillacea Harvey, 1853 Jania pacifica Areschough, 1852 Jania pumila J.V. Lamouroux, 1816 Jania tenella (Kützing) Grunow, 1874 Lithophyllum corallinae (P.L. Crouan and H.M. Crouan) Heydrich, 1897 Lithophyllum decussatum (J. Ellis and Solander) Philippi, 1837 =Lithophyllum farlowii Heydrich 1901 Lithophyllum dispar (Foslie) Foslie, 1909 =Titanoderma dispar (Foslie) Woelkerling, Y. M. Chamberlain and P.C. Silva 1985 Lithophyllum imitans Foslie, 1909 Lithophyllum lithophylloides Heydrich, 1901 Lithophyllum stictaeforme (J.E. Areschoug) Hauck, 1877 =Lithophyllum frondosum (Dufour) G. Furnari, Cormaci and Alongi 1996 Neogoniolithon trichotomum (Heydrich) Setchell and L.R. Mason, 1943 =Lithophyllum trichotomum (Heydrich) Me. Lemoine 1929 Pneophyllum confervicola (Kützing) Y.M. Chamberlain, 1983 = Heteroderma minutulum (Foslie) Foslie 1909 Pneophyllum cornicum (E.Y. Dawson) Keats, Y.M. Chamberlain and Baba, 1997 Pneophyllum fragile Kützing, 1843 Spongites decipiens (Foslie) Y.M. Chamberlain, 1993 Spongites fructiculosa Kützing, 1842 Family Hapalidiaceae J.E. Gray, 1864 Choreonema thuretii (Bornet) F. Schmitz, 1889 Leptophytum microsporum (Foslie) Athanasiadis and Adey, 2006 =Lithothamnion microsporum (Foslie) Foslie 1929 Lithothamnion australe Foslie, 1904 Melobesia membranacea (Esper) J.V. Lamouroux, 1812 Phymatolithon lenormandii (J.E. Areschoug) W.H. Adey =Lithothamnion lenormandii (J. E. Areschoug) Foslie 1895 Order Halymeniales G.W. Saunders and G.T. Kraft, 1996 Family Halymeniaceae Kützing, 1847 Grateloupia angusta (Okamura) Kawaguchi and Wang ex Wang, Kawaguchi, Horiguchi and Masuda, 2001 =Cryptonemia angustata Okamura 1895 Grateloupia filicina (J.V. Lamouroux) C. Agardh, 1822 Grateloupia huertana Mateo-Cid, Mendoza-González and Gavio, 2005 Grateloupia maxima (Gardner) Kylin, 1941 Grateloupia prolongata J. Agardh, 1847 Grateloupia versicolor (J. Agardh) J. Agardh, 1847 Order Peyssonneliales D.M. Krayesky, Fredericq and J.N. Norris, 2009 Family Peyssonneliaceae Denizot, 1968 Peyssonelia mexicana E.Y. Dawson, 1953 Peyssonelia orientalis (Weber-van Bosse) Cormaci and G. Furnari, 1987 Peyssonelia pacifica Kylin, 1925 Order Gelidiales Kylin 1923 Family Gelidiaceae Kützing, 1843 Gelidium crinale (Hare ex Turner) Gaillon, 1828 Gelidium isabelae W.R. Taylor, 1945 Gelidium microdentatum E.Y. Dawson, 1953 Gelidium pusillum (Stackhouse) Le Jolis, 1863 Gelidium sclerophyllum W.R. Taylor, 1945 Pterocladiella bulbosa (N.H. Loomis) Santelices, 1997 =Pterocladia bulbosa N.H. Loomis 1960 Family Gelidiellaceae Fan 1961 Gelidiella acerosa (Forsskäl) Feldmann and G. Hainel, 1934 Gelidiella hancockii E.Y. Dawson, 1944 Gelidiella ligulata E.Y. Dawson, 1953 Order Gigartinales Schmitz ex Engler, 1892 Family Dicranemataceae Kylin, 1932 Dicranema rosaliae Setchell and Gardner, 1924 León-Álvarez and González-González (1993) Mendoza-González and Mateo-Cid (1998) León-Tejera and González-González (1993) León-Tejera and González-González (1993) León-Tejera et al. (1993) León-Tejera and González-González (1993) León-Tejera and González-González (1993) Fragoso and Rodríguez (2002) Mendoza-González and Mateo-Cid (1999) Mateo-Cid and Mendoza-González (1997) León-Álvarez and González-González (1993) León-Álvarez and González-González (1993) Fragoso and Rodríguez (2002) León-Álvarez and González-González (1993) León-Álvarez and González-González (1993) Mendoza-González and Mateo-Cid (1998) Mendoza-González and Mateo-Cid (1998) Mendoza-González and Mateo-Cid (1998) León-Álvarez and González-González (1993) Mendoza-González and Mateo-Cid (1998) Mendoza-González and Mateo-Cid (1998) León-Álvarez and González-González (1993) Fragoso and Rodríguez (2002) León-Álvarez and González-González (1993) Mateo-Cid et al. (2005) León-Tejera and González-González (1993) Mateo-Cid et al. (2005) León-Tejera and González-González (1993) León-Tejera and González-González (1993) León-Tejera and González-González (1993) León-Álvarez and González-González (1993) Mendoza-González and Mateo-Cid (1999) León-Tejera and González-González (1993) León-Tejera and González-González (1993) Mateo-Cid et al. (2005) Mateo-Cid and Mendoza-González (1997) León-Tejera et al. (1993) León-Tejera and González-González (1993) León-Tejera and González-González (1993) Mendoza-González and Mateo-Cid (1998) León-Tejera and González-González (1993) León-Tejera and González-González (1993) León-Tejera and González-González (1993) 339 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 3. Continued. Family Hypneaceae J. Agardh, 1851 Hypnea johnstonii Setchell and N.L. Gardner, 1924 Hypnea pannosa J. Agardh, 1847 Hypnea spinella (C. Agardh) Kützing, 1847 Family Phyllophoraceae Nägeli, 1847 Ahnfeltiopsis concinna (J. Agardh) P.C. Silva and DeCew, 1992 Ahnfeltiopsis gigartinoides (J. Agardh) P.C. Silva and DeCew, 1992 Mendoza-González and Mateo-Cid (1999) León-Tejera and González-González (1993) León-Tejera et al. (1993) León-Tejera and González-González (1993) León-Tejera and González-González (1993) Ahnfeltiopsis serenei (E.Y. Dawson) Masuda, 1993 =Gymnogongrus serenei E.Y. Dawson 1954 León-Tejera and González-González (1993) Petrocelis anastomosans E.Y. Dawson, 1961 León-Álvarez and González-González (1993) Predaea subpeltata E.Y. Dawson, 1960 León-Álvarez and González-González (1993) Gymnogongrus crustiforme E.Y. Dawson, 1961 Gymnogongrus johnstonii (Setchell and N.L. Gardner) E.Y. Dawson, 1961 Order Nemastomatales Kylin, 1925 Family Nemastomataceae Schmitz, 1892 Order Gracilariales Fredericq S. and M.H. Hommersand, 1989 Family Gracilariaceae Nägeli, 1847 Gracilaria bursa-pastoris (S.G. Gmelin) P.C. Silva, 1952 Gracilaria crispata Setchell and N.L. Gardner, 1924 Gracilaria pachydermatica Setchell and N.L. Gardner, 1924 Gracilaria parvispora I.A. Abbott, 1985 Gracilaria ramisecunda E.Y. Dawson, 1949 Gracilaria symmetrica E.Y. Dawson, 1949 Gracilaria tepocensis (E.Y. Dawson) E.Y. Dawson, 1961 Gracilaria veleroae E.Y. Dawson, 1944 Gracilaria verrucosa (Hudson) Papenfus, 1950 Gracilariopsis andersonii (Grunow) E.Y. Dawson, 1949 =Gracilariopsis sjoestedii (Kylin) E.Y. Dawson 1949 Order Halymeniales G.W. Saunders and G.T. Kraft, 1996 Family Halymeniaceae Kützing, 1867 Prionitis sternbergii (C. Agardh) J. Agardh,1851 =Zanardinula sternbergii (C. Agardh) G. De Toni 1936 Order Hildenbrandiales Pueschel and Cole, 1982 León-Tejera and González-González (1993) León-Tejera and González-González (1993) Mendoza-González and Mateo-Cid (1998) Mendoza-González and Mateo-Cid (1998) Mateo-Cid and Mendoza-González (1997) Dreckmann (1999) Mendoza-González and Mateo-Cid (1999) León-Tejera and González-González (1993) Mateo-Cid and Mendoza-González (1997) Mendoza-González and Mateo-Cid (1998) León-Tejera and González-González (1993) León-Tejera and González-González (1993) León-Tejera and González-González (1993) Family Hildenbrandiaceae Rabenhorst, 1868 Hildenbrandia rubra (Sommerfelt) Meneghini, 1841 Order Nemaliales Schmitz ex Engler, 1892 Family Galaxauraceae P.G. Parkinson, 1983 Galaxaura rugosa (J. Ellis and Solander) J.V. Lamouroux, 1816 Scinaia latifrons M.A. Howe, 1911 Tricleocarpa cylindrica (J. Ellis and Solander) Huisman and Borowitzka, 1990 =Galaxaura fastigiata Decaisne 1842 Family Liagoraceae Kützing, 1843 León-Tejera et al. (1993) León-Tejera et al. (1993) León-Tejera and González-González (1993) León-Tejera et al. (1993) Dermonema virens (J. Agardh) Pedroche and Ávila-Ortíz, 1996 =Nemalion virens J. Agardh 1847 León-Tejera and González-González (1993) Champia parvula (C. Agardh) Harvey, 1853 León-Tejera and González-González (1993) Order Rhodymeniales Schmitz ex Engler, 1892 Family Champiaceae Kützing, 1843 Family Rhodymeniaceae Harvey, 1849 Gelidiopsis variabilis (Greville ex J. Agardh) Schmitz, 1895 =Gelidiopsis tenuis Setchell and Gardner 1924 Microphyllum crispum (W.R. Taylor) E.Y. Dawson, 1963 Rhodymenia divaricata E.Y. Dawson, 1941 Kingdom Chromista Cavalier-Smith, 1981 Class Phaeophyceae Kjellman, 1891 Order Dictyotales Bory de Saint-Vincent, 1828 Family Dictyotaceae J.V. Lamouroux ex Dumortier, 1822 Dictyota binghamiae J. Agardh, 1894 =Pachydictyon binghamiae (J. Agardh) E.Y. Dawson 1950 Dictyota crenulata J. Agardh, 1847 Dictyota dichotoma (Hudson) J.V. Lamouroux, 1809 Dictyota dichotoma var. intricata (C. Agardh) Greville, 1830 =Dictyota divaricata J.V. Lamouroux 1809 Dictyota flabellata (F.S. Collins) Setchell and N.L. Gardner, 1924 Dictyota friabilis Setchell, 1926 =Dictyota pfafi Schnetter 1972 Dictyota pinnata (E.Y. Dawson) I. Hörnig, R. Schnetteret and Prud’homme van Reine, 1993 Dictyopteris delicatula J.V. Lamouroux, 1809 Lobophora variegate (J.V. Lamouroux) Womersley ex E.C. Oliveira, 1977 León-Tejera et al. (1993) Mateo-Cid and Mendoza-González (1997) León-Tejera and González-González (1993) León-Tejera and González-González (1993) León-Tejera and González-González (1993) León-Tejera et al. (1993) León-Tejera and González-González (1993) León-Tejera and González-González (1993) León-Tejera et al. (1993) Mendoza-González and Mateo-Cid (1998) Mendoza-González and Mateo-Cid (1998) León-Álvarez and González-González (1993) 340 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 3. Continued. Padina caulescens Thivy ex W.R. Taylor, 1945 Padina crispata Thivy ex W.R. Taylor, 1945 Padina durvillaei Bory Saint-Vincent, 1827 Padina gymnospora (Kützing) Sonder, 1871 Padina mexicana E.Y. Dawson, 1944 Mateo-Cid and Mendoza-González (1997) León-Tejera and González-González (1993) León-Tejera et al. (1993) León-Tejera et al. (1993) Mendoza-González and Mateo-Cid (1998) P. mexicana var. erecta Ávila-Ortiz, 2003 Ávila-Ortiz (2003) Feldmannia cylindrica (Saunders) Hollenberg and I.A. Abbott, 1966 León-Tejera et al. (1993) Order Ectocarpales Bessey, 1907 Family Acinetosporaceae Hamel ex J. Feldmann, 1937 Feldmannia indica (Sonder) (Sonder) Womersley and A. Bailey, 1970 Hincksia mitchelliae (Harvey) P.C. Silva ex Silva, Meñez and Moe, 1987 Hincksia rallsiae (Vickers) ex P.C. Silva, Meñez and Moe, 1987 Family Chordariaceae Greville, 1830 Kuetzingiella elachistaeformis (Heydrich) M. Balakrishnan and Kinkar, 1981 =Feldmannia elachistaeformis (Heydrich) Pham-Hoàng Hô 1969 Family Ectocarpaceae C. Agardh, 1828 Ectocarpus commensalis Setchell and N.L. Gardner, 1922 =Ectocarpus parvus (De A. Saunders) H. Hollenberg 1971 Ectocarpus siliculosus var. dasycarpus (Kuckuck) Gallardo, 1992 =Ectocarpus confervoides (Roth) Le Jol. var. dasycarpa (Kuckuck) Rosenvinge and Lund 1941 Ectocarpus siliculosus var. pygmaeus (J.E. Areschoug) Gallardo, 1992 =Ectocarpus confervoides (Roth) Le Jolis. var. pygmaeus (J. Areschoug) Rosenvinge 1893 Order Fucales Bory de Saint-Vincent, 1827 Family Sargassaceae Kützing, 1843 Sargassum howellii Setchell, 1937 Sargassum liebmanii J. Agardh, 1847 Order Ralfsiales Nakamura ex Lim and Kawai ex Lim, Sakaguchi, Hanyuda, Kogame, Phang and Kawai, 2007 León-Tejera and González-González (1993) León-Tejera and González-González (1993) Mateo-Cid and Mendoza-González (1997) León-Tejera et al. (1993) Mendoza-González and Mateo-Cid (1998) León-Tejera et al. (1993) León-Tejera et al. (1993) Mendoza-González and Mateo-Cid (1998) León-Tejera et al. (1993) Family Ralfsiaceae Farlow, 1881 Ralfsia confusa Hollenberg, 1969 Ralfsia hancockii E.Y. Dawson, 1944 León-Tejera et al. (1993) León-Álvarez and González-González (1993) Stragularia clavata (Harvey) G. Hamel, 1939 León-Álvarez and González-González (1993) Chnoospora minima (Hering) Papenfuss, 1956 León-Tejera and González-González (1993) Order Scytosiphonales Feldmann, 1949 Family Chnoosporaceae Setchell and N.L. Gardner, 1925 Family Scytosiphonaceae Farlow, 1881 Colpomenia ramosa W.R. Taylor, 1945 Diplura simulans Hollenberg, 1969 Rosenvingea antillarum (P.L. Crouan and H.M. Crouan) M.J. Wynne, 1997 Rosenvingea floridana (W.R. Taylor) W.R. Taylor, 1955 Order Scytothamnales A.F. Peters and M.N. Clayton, 1998 Family Scytothamnaceae Womersley, 1987 Asteronema breviarticulatum (J. Agardh) Ouriques and Bouzon, 2000 =Ectocarpus breviarticulatus J. Agardh 1847 Order Sphacelariales Migula, 1909 Family Sphacelariaceae Decaisne, 1842 Hapalospongidion gelatinosum De A. Saunders, 1899 Pseudolithoderma nigrum Hollenberg, 1969 Sphacelaria rigidula Kützing, 1843 Sphacelaria tribuloides meneghini 1840 Table 4. Checklist of the sponges recorded in the marine areas of Oaxaca, Mexico. Phylum Porifera Grant, 1836 Mateo-Cid and Mendoza-González (1997) León-Tejera et al. (1993) Aguilar-Rosas et al. (2009) Mendoza-González and Mateo-Cid (1998) León-Tejera and González-González (1993) León-Tejera et al. (1993) León-Álvarez and González-González (1993) León-Tejera et al. (1993) León-Tejera et al. (1993) Reference Class Demospongiae Sollas, 1885 Order Astrophorida Sollas, 1887 Family Thoosidae Cockerell, 1925 Thoosa calpulli Carballo, Cruz-Barraza and Gómez, 2004 Order Chondrosida Boury-Esnault and López, 1985 Family Chondrillidae Gray, 1872 Chondrilla montanusa Carballo, Gómez, Cruz-Barraza and Flores-Sánchez, 2003 Chondrosia tenochca Carballo, Gómez, Cruz-Barraza and Flores-Sánchez, 2003 Order Hadromeridae Topsent, 1894 Family Clionaidae d’Orbigny, 1851 Cliona mucronata Sollas, 1878 Carballo et al. (2008a) Carballo et al. (2003) Carballo et al. (2003) Bautista-Guerrero et al. (2006) 341 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 4. Continued. Cliona pocillopora Bautista-Guerrero, Carballo, Cruz-Barraza and Nava, 2006 Cliona tropicalis Cruz-Barraza, Carballo, Bautista-Guerrero and Nava, 2011 Cliona vermifera Hancock, 1867 Cliona sp. Carballo, Bautista-Guerrero and Leyte-Morales, 2008 Cliothosa hancocki (Topsent, 1888) Cliothosa tylostrongylata Cruz-Barraza, Carballo, Bautista-Guerrero and Nava, 2011 Pione carpenteri (Hancock, 1867) Order Haplosclerida Topsent, 1928 Family Callyspongiidae de Laubenfels, 1936 Callyspongia californica Dickinson, 1945 Family Niphatidae van Soest, 1980 Amphimedon texotli Cruz-Barraza and Carballo, 2008 Family Phloeodictyidae Carter, 1882 Aka cryptica Carballo, Hepburn, Nava, Cruz-Barraza and Bautista-Guerrero, 2007 Order Poecilosclerida Topsent, 1928 Family Mycalidae Lundbeck, 1905 Mycale (Carmia) cecilia de Laubenfels, 1936 Mycale (Carmia) aff. magnirhaphidifera van Soest, 1984 Order Verongida Bergquist, 1978 Family Aplysinidae Carter, 1875 Aplysina gerardogreeni Gómez and Bakus, 1992 Bautista-Guerrero et al. (2006) Cruz-Barraza et al. (2011) Carballo et al. (2008a) Carballo et al. (2008a) Carballo et al. (2008a) Cruz-Barraza et al. (2011) Carballo et al. (2008a) Dickinson (1945) Cruz-Barraza and Carballo (2008) Carballo et al. (2007) Carballo and Cruz-Barraza (2010) Carballo and Cruz-Barraza (2010) Gómez and Bakus (1992) Table 5. Checklist of the cnidarians recorded in marine areas from Oaxaca, Mexico. Order according to Daly et al. (2007). Phylum Cnidaria Hatschek, 1888 Reference Class Anthozoa Ehrenberg, 1834 SubClass Hexacorallia Haeckel1866 Order Scleractinia Bourne, 1900 SubOrder Astrocoeniina Vaughan and Wells, 1943 Family Pocilloporidae Gray, 1842 Madracis pharensis (Heller, 1868) Pocillopora capitata Verril, 1864 Pocillopora damicornis (Linnaeus, 1758) Reyes-Bonilla and Cruz-Piñón (2000) Palmer (1928) Squires (1959) Pocillopora effusus Veron, 2000 Glynn and Ault (2000) Pocillopora elegans Dana, 1846 Reyes-Bonilla et al. (2005) Pocillopora inflata Glynn, 1999 Leyte-Morales (1997) Pocillopora meandrina Dana, 1846 Glynn and Leyte-Morales (1997) Pocillopora eydouxi Milne-Edwards and Haime, 1860 Palmer (1928) Pocillopora verrucosa (Ellis and Solander, 1786) Leyte-Morales (1997) Gardineroseris planulata (Dana, 1846) Leyte-Morales (1997) Pavona gigantea Verril, 1869 Durham (1947) SubOrder Fungiina Verrill, 1865 Family Agariciidae Gray, 1847 Leptoseris papyracea (Dana, 1846) Leyte-Morales (1997) Pavona varians Verril, 1964 Leyte-Morales (1997) Pavona clavus (Dana, 1846) Family Fungiacyathidae Chevalier, 1987 Fungiacyathus marenzelleri (Vaughan, 1906) Family Fungiidae Dana, 1846 Fungia distorta Michelin, 1842 Leyte-Morales et al. (2001) Reyes-Bonilla and Cruz-Piñón (2000) Reyes-Bonilla et al. (2005) Family Poritidae Gray, 1842 Porites panamensis Verril, 1866 Family Siderastreidae Vaughan and Wells, 1943 Psammocora stellata (Verril, 1866) SubOrder Faviina Vaughan and Wells, 1943 Palmer (1928) Glynn and Leyte-Morales (1997) Family Oculinidae Gray, 1847 Madrepora oculata Linnaeus, 1758 Oculina profunda Cairns, 1991 Family Rhizangiidae D’Orbigny, 1851 Reyes-Bonilla and Cruz-Piñón (2000) Reyes-Bonilla and Cruz-Piñón (2000) Astrangia browni Palmer, 1928 Palmer (1928) Astrangia costata Verril, 1866 Reyes-Bonilla and Cruz-Piñón (2000) Astrangia dentata Verril, 1866 Astrangia haimei Verril, 1866 Astrangia tangolaensis Durham, 1947 Reyes-Bonilla and Cruz-Piñón (2000) Palmer (1928) Durham (1947) 342 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 5. Continued. Coenangia conferta Verril, 1870 SubOrder Caryophyllina Vaughan and Wells, 1943 Family Caryophylliidae Dana, 1846 Caryophyllia diomedeae Marenzeller, 1904 Coenocyathus bowersi Vaughan, 1906 Desmophyllum dianthus (Esper, 1794) Lophelia pertusa (Linnaeus, 1758) Paracyathus humilis Verrill, 1870 Phyllangia consagensis (Durham and Barnard, 1952) Phyllangia dispersa Verril, 1864 Family Flabellidae Bourne, 1905 Reyes-Bonilla and Cruz-Piñón (2000) Reyes-Bonilla and Cruz-Piñón (2000) Reyes-Bonilla and Cruz-Piñón (2000) Reyes-Bonilla and Cruz-Piñón (2000) Reyes-Bonilla and Cruz-Piñón (2000) Reyes-Bonilla and Cruz-Piñón (2000) Reyes-Bonilla and Cruz-Piñón (2000) Reyes-Bonilla and Cruz-Piñón (2000) Javania cailleti (Duchassaing and Michelotti, 1864) Reyes-Bonilla and Cruz-Piñón (2000) Sphenotrochus hancocki Durham and Barnard, 1952 Reyes-Bonilla and Cruz-Piñón (2000) Polymyces montereyensis (Durham, 1947) Family Turbinoliidae Milne-Edwards and Haime, 1848 SubOrder Dendrophylliina Vaughan and Wells, 1943 Family Dendrophylliidae Gray, 1847 Reyes-Bonilla and Cruz-Piñón (2000) Balanophyllia cedrosensis Durham, 1947 Reyes-Bonilla and Cruz-Piñón (2000) Endopachys grayi Milne-Edwards and Haime, 1848 Reyes-Bonilla et al. (1997) Cladopsammia eguchii (Wells, 1982) Dendrophyllia oldroydae Oldroyd, 1924 Tubastraea coccinea Lesson, 1829 SubClass Octocorallia Haeckel, 1866 Reyes-Bonilla and Cruz-Piñón (2000) Reyes-Bonilla and Cruz-Piñón (2000) Reyes-Bonilla and Cruz-Piñón (2000) Order Alcyonacea Lamouroux, 1816 Family Plexauridae Gray, 1859 Plexauriidae sp. (Luke, 1998) Luke (1998) Order Pennatulacea Verrill, 1865 Family Pennatulidae Ehrenberg, 1828 Ptilosarcus undulatus (Verrill, 1865) Luke (1998) Family Renillidae Gray, 1860 Renilla sp. (Luke, 1998) Luke (1998) Class Hydrozoa Owen, 1843 SubClass Hydroidolina Collins, 2000 Order Anthoathecata Cornelius, 1992 SubOrder Filifera Kühn, 1913 Family Bougainvilliidae Lütken, 1850 Bougainvillia crassa Fraser, 1938 Fraser (1938a) Eudendrium ramosum (Linnaeus, 1758) Fraser (1938a) Bougainvillia robusta (Fraser, 1938) (species inquirenda) Family Eudendriidae Agassiz, 1862 Eudendrium tenellum Allman, 1877 (species inquirenda) Family Hydractiniidae Agassiz, 1862 Fraser (1938a) Fraser (1938a) Hydractinia apicata (Kramp, 1959) Segura-Puertas (1984) Hydractinia carolinae Fraser, 1912 Fraser (1938a) Family Oceaniidae Eschscholtz, 1829 Garveia laxa (Fraser, 1938) (species inquirenda) Tubiclava triserialis Fraser, 1938 (nomen dubium) Turritopsis nutricula McCrady, 1857 Family Pandeidae Haeckel, 1879 Halitholus pauper Hartlaub, 1913 Fraser (1938a) Fraser (1938a) Fraser (1938a) Segura-Puertas (1984) Leuckartiara zacae Bigelow, 1940 Segura-Puertas (1984) Leuckartiara octona (Fleming, 1823) Fraser (1938a) Family Proboscidactylidae Hand and Hendrickson, 1950 Proboscidactyla ornata (McCrady, 1859) Segura-Puertas (1984) Lizzia alvarinoae Segura, 1980 Segura-Puertas (1984) Lizzia ferrarii Segura, 1980 Segura-Puertas (1984) Family Rathkeidae Russell, 1953 Lizzia gracilis (Mayer, 1900) Segura-Puertas (1984) Podocorynoides minima (Trinci, 1903) Segura-Puertas (1984) Euphysilla pyramidata Kramp, 1955 Segura-Puertas (1984) SubOrder Capitata Kühn, 1913 Family Corymorphidae Allman, 1872 343 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 5. Continued. Family Tubulariidae Fleming, 1828 Ectopleura sacculifera Kramp, 1957 Family Zancleidae Russell, 1953 Zanclea costata Gegenbaur, 1857 Segura-Puertas (1984) Segura-Puertas (1984) Order Leptothecata Cornelius, 1982 SubOrder Conica Broch, 1910 Family Aequoreidae Eschscholtz, 1829 Aequorea coerulescens (Brandt, 1838) Family Eirenidae Haeckel, 1879 Phialopsis diegensis Torrey, 1909 Segura-Puertas (1984) Segura-Puertas (1984) Family Haleciidae Hincks, 1868 Endothecium reduplicatum Fraser, 1935 Fraser (1948) Halecium halecinum (Linnaeus, 1758) Fraser (1938a) Halecium beanii (Johnston, 1838) Halecium flabellatum Fraser, 1935 Family Halopterididae Millard, 1962 Antennella secundaria (Gmelin, 1791) Monostaechas quadridens (McCrady, 1859) Family Hebellidae Fraser, 1912 Scandia expansa Fraser, 1938 (nomen dubium) Family Lafoeidae Hincks, 1868 Filellum serpens (Hassall, 1848) Family Lovenellidae Russell, 1953 Fraser (1938a) Fraser (1938a) Fraser (1938a) Fraser (1938a) Fraser (1938b) Fraser (1938a) Eucheilota comata (Bigelow, 1909) Segura-Puertas (1984) Eucheilota menoni A. Agassiz, 1862 Segura-Puertas (1984) Lovenella nodosa Fraser, 1938 Fraser (1938a) Family Malagazziidae Bouillon, 1984 Octophialucium bigelowi Kramp, 1955 Segura-Puertas (1984) Family Plumulariidae Agassiz, 1862 Plumularia adjecta Fraser, 1948 Plumularia biarmata Fraser, 1938 Plumularia floridana Nutting, 1900 Plumularia propinqua Fraser, 1938 Family Sertulariidae Linnaeus, 1758 Fraser (1948) Fraser (1938a) Fraser (1938a) Fraser (1938a) Dynamena crisioides Lamouroux, 1824 Fraser (1948) Dynamena disticha (Bosc, 1802) Fraser (1938a) Sertularella conica Allman, 1877 Sertularia desmoides Torrey, 1902 Sertularia distans (Lamouroux, 1816) Thuiaria tubuliformis Nutting, 1904 SubOrder Proboscoida Broch, 1910 Family Campanulariidae Johnston, 1836 Clytia bakeri Torrey, 1904 Clytia fascicularis Fraser, 1938 Clytia gracilis (Sars, 1850) Clytia hemisphaerica (Linnaeus, 1767) Clytia hesperia (Torrey, 1904) (species inquirenda) Clytia irregularis Fraser, 1938 Clytia kincaidi (Nutting, 1899) Clytia paulensis (Vanhöffen, 1910) Clytia seriata Fraser, 1938 Clytia simplex (Browne, 1902) Clytia universitatis Torrey, 1904 Obelia dichotoma (Linnaeus, 1758) Obelia geniculata (Linnaeus, 1758) SubClass Trachylina Parker, 1982 Order Narcomedusae Haeckel, 1879 Fraser (1938a) Fraser (1938a) Fraser (1948) Fraser (1938a) Fraser (1938b) Fraser (1938a) Fraser (1938a) Fraser (1938a) Fraser (1948) Fraser (1938a) Fraser (1938a) Fraser (1938a) Fraser (1938a) Segura-Puertas (1984) Fraser (1938b) Fraser (1938a) Fraser (1938a) Family Aeginidae Gegenbaur, 1857 Solmundella bitentaculata (Quoy and Gaimard, 1833) Family Cuninidae Bigelow, 1913 Segura-Puertas (1984) Cunina frugifera Kramp, 1948 Segura-Puertas (1984) Cunina octonaria McCrady, 1857 Segura-Puertas (1984) Cunina peregrina Bigelow, 1909 Segura-Puertas (1984) 344 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 5. Continued. Cunina tenella (Bigellow, 1909) Segura-Puertas (1984) Family Solmarisidae Haeckel, 1879 Pegantha martagon Haeckel, 1879 Segura-Puertas (1984) Pegantha triloba Haeckel, 1879 Segura-Puertas (1984) Order Trachymedusae Haeckel, 1866 Family Geryoniidae Péron and Lesueur, 1810 Geryonia proboscidalis (Huxley, 1859) Segura-Puertas (1984) Liriope tetraphylla (Chamisso and Eysenhardt, 1821) Segura-Puertas (1984) Family Rhopalonematidae Russell, 1953 Aglaura hemistoma Péron and Lesueur, 1810 Segura-Puertas (1984) Amphogona apicata Kramp, 1957 Segura-Puertas (1984) Rhopalonema velatum Gegenbaur, 1857 Segura-Puertas (1984) Sminthea eurygaster Gegenbaur, 1856 Segura-Puertas (1984) Class Scyphozoa Götte, 1887 Order Coronatae Vanhöffen, 1892 Family Nausithoidae Bigelow, 1913 Nausithoe punctata Kölliker, 1853 Segura-Puertas (1984) Pelagia noctiluca (Forsskal, 1775) Segura-Puertas (1984) Order Semaeostomeae Agassiz, 1862 Family Pelagiidae Gegenbaur, 1856 Order Rhizostomeae Cuvier, 1799 Family Cassiopeidae Agassiz, 1862 Cassiopea sp. Péron and Lesueur, 1810 Chávez (1979) Family Stomolophidae Haeckel, 1880 Chávez (1979) Stomolophus meleagris Agassiz, 1862 Table 6. Checklist of the platyhelmints recorded in marine areas from Oaxaca, Mexico. Order according to Cavalier-Smith (1998). When the speciesname of the record was changed, because it was synonymized, it was indicated with a ’=’ symbol, followed by original recorded name in parenthesis. Phylum Platyhelminthes Gegenbaur, 1859 em. Minot, 1876. Reference Class Monogenea (Van Beneden, 1858) Carus, 1863 SubClass Polyopisthocotylea Odhner, 1912 Order Mazocraeidea Price, 1936 Family Gastrocotylidae Price, 1943 Pseudaxine mexicana Meserve, 1938 Meserve (1938) Family Neothoracocotylidae Lebedev, 1969 Thoracocotyle paradoxica Meserve, 1938 Meserve (1938) Class Trematoda Rudolphi, 1808 SubClass Digenea Carus, 1863 Order Plagiorchiida La Rue, 1957 Family Acanthocolpidae Lühe, 1906 Stephanostomum casum (Linton, 1910) McFarlane, 1934 Manter (1940) Homalometron mexicanum (Manter, 1937) (=Apocreadium mexicanum) Manter (1937) Stephanostomum hispidum (Yamaguti, 1934) Manter, 1940 Manter (1940) Family Apocreadiidae Skrjabin, 1942 Family Cryptogonimidae Ward, 1917 Paracryptogonimus americanus Manter, 1940 Manter (1940) Family Plagiorchiidae Lühe, 1901 Gámez-Vivaldo et al. (2006) Enodiotrema megachondrus Looss, 1901 Family Pronocephalidae Looss, 1899 Gámez-Vivaldo et al. (2006) Adenogaster serialis Looss, 1901 Table 7. Checklist of the nemerteans recorded in marine areas from Oaxaca, Mexico. Order according to Cavalier-Smith (1998). Phylum Nemertea Minot, 1876. Reference Class Anopla Schultze, 1851 Subclass Heteronemertea Family Lineidae Euborlasia hancocki Coe, 1940 Coe (1940) Table 8. Checklist of the annelids in the marine areas of Oaxaca, Mexico. When the species-name of the record was changed, because it was synonymized, it was indicated with a ’=’ symbol, followed by original recorded name in parenthesis. EX= invasive species; Q= questionable species-names. Phylum Annelida Lamarck, 1809 Class Clitellata Lamarck, 1818 Reference Family Ozobranchidae Pinto, 1921 Ozobranchus branchiatus (Menzies, 1791) Gámez-Vivaldo et al. (2006) 345 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 8. Continued. Class Polychaeta Grube, 1850 Family Alciopidae Ehlers, 1864 Alciopina parasitica Claparède and Panceri, 1867 Q Krohnia lepidota (Krohn, 1845) Q Plotohelmis capitata (Greeff, 1876) Q Rhynchonerella gracilis Costa, 1862 Q Fernández-Álamo (1987) Fernández-Álamo and Sanvicente-Añorve (2005) Fernández-Álamo (1987) Fernández-Álamo (1987) Rhynchonerella moebii (Apstein, 1893) Q Fernández-Álamo (1987) Amphicteis scaphobranchiata Moore, 1906 Solís-Weiss et al. (2000) Isolda pulchella Müller, in Grube, 1858 Q Solís-Weiss et al. (2000) Solís-Weiss et al. (2000) Chloeia viridis Schmarda, 1861 Q Treadwell (1941) Family Ampharetidae Malmgren, 1867 Pterolysippe vanelli (Fauvel, 1936) sensu Eliason, 1955 Q Family Amphinomidae Savigny, in Lamarck, 1818 Eurythoe complanata (Pallas, 1766) Q Hartman (1940) Linopherus ambigua (Monro, 1933) Solís-Weiss et al. (2000) Notopygos ornata Grube, 1856 Gómez et al. (1997) Pareurythoe californica (Johnson, 1897) Gómez et al. (1997) Family Aphroditidae Malmgren, 1867 Aphrodita sp. Treadwell (1941) Family Capitellidae Grube, 1862 Treadwell (1941) Notomastus daueri Ewing, 1982 Q Solís-Weiss et al. (2000) Chrysopetalum occidentale Johnson, 1897 Q Gómez et al. (1997) Notomastus (Clistomastus) hemipodus Hartman, 1947 Q Family Chrysopetalidae Ehlers, 1864 Family Cirratulidae Carus, 1863 González-Ortiz et al. (1996) Aphelochaeta multifilis (Moore, 1909) Q Fauchald (1972) Chaetozone setosa Malmgren, 1867 Q Solís-Weiss et al. (2000) Cirratulus sp. Gómez et al. (1997) Monticellina serratiseta (Banse and Hobson, 1968) Q Solís-Weiss et al. (2000) M. tesselata (Hartman, 1960) Q Solís-Weiss et al. (2000) Tharyx parvus Berkeley, 1929 Q Solís-Weiss et al. (2000) Family Cossuridae Day, 1963 Cossura brunnea Fauchald, 1972 Family Dorvilleidae Chamberlin, 1919 Dorvillea cerasina (Ehlers, 1901) Q Family Eulepethidae Chamberlin, 1919 Fauchald (1972) Gómez et al. (1997) Grubeulepis mexicana (Berkeley and Berkeley, 1939) Solís-Weiss et al. (2000) Eunice cf. aphroditois Ruiz-Cancino et al. (2011) Family Eunicidae Berthold, 1827 Eunice antennata (Savigny in Lamarck, 1818) Q Hartman (1944) Eunice filamentosa Grube, 1856 Q Hartman (1944) Eunice chicasi de León-González, Rivera and Romero, 2004 Eunice cf. filamentosa Eunice gagzoi Augener, 1922 Q Eunice lucei Grube, 1856 Ruiz-Cancino et al. (2011) Ruiz-Cancino et al. (2011) Gómez et al. (1997) Gómez et al. (1997) Eunice mutilata Webster, 1884 Q Gómez et al. (1997) Eunice vittata (Delle Chiaje, 1828) Q Solís-Weiss et al. (2000) Lysidice ninetta Audouin and Milne-Edwards, 1833 Q Gómez et al. (1997) Eunice sp. 2 Lysidice cf. collaris Ruiz-Cancino et al. (2011) Ruiz-Cancino et al. (2011) Marphysa aenea (Blanchard, 1849) Q Hartman (1944) Marphysa mixta Fauchald, 1970 Fauchald (1970) Marphysa sp. Ruiz-Cancino et al. (2011) Nematonereis unicornis (Grube, 1840) Q Nematonereis cf. unicornis Palola pallidus Hartman, 1938 Q Palola siciliensis (Grube, 1840) Q Palola sp. 1 Palola sp. 2 Family Flabelligeridae Saint-Joseph, 1894 Brada villosa (Rathke, 1843) Q Ilyphagus bythincola Chamberlin, 1919 Pherusa inflata (Treadwell, 1914) Q Gómez et al. (1997) Ruiz-Cancino et al. (2011) Ruiz-Cancino et al. (2011) Rioja (1962) Ruiz-Cancino et al. (2011) Ruiz-Cancino et al. (2011) Solís-Weiss et al. (2000) Chamberlin (1919) Gómez et al. (1997) 346 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 8. Continued. Piromis gracilis Hartman, 1961 Q Hartman (1961) Family Glyceridae Grube, 1850 Glycera convoluta Keferstein, 1862 Q Solís-Weiss et al. (2000) Glycera papillosa Grube, 1857 Q Solís-Weiss et al. (2000) Glycera tesselata Grube, 1840 Q Solís-Weiss et al. (2000) Family Goniadidae Kinberg, 1866 Glycinde solitaria (Webster, 1879) Q Solís-Weiss et al. (2000) Goniada maculata Örsted, 1843 Q Solís-Weiss et al. (2000) Family Hesionidae Grube, 1850 Podarke pugettensis Johnson, 1901 Q Family Iospilidae Bergström, 1914 Phalacrophorus uniformis Reibisch, 1895 Q Family Lopadorhynchidae Claparède, 1868 Gómez et al. (1997) Fernández-Álamo (1987) Lopadorhynchus brevis Grube, 1855 Q Fernández-Álamo (1987) Lopadorhynchus henseni Reibisch, 1893 Q Fernández-Álamo (2002) Lopadorhynchus nationalis Reibisch, 1895 Q Fernández-Álamo (1987) Lopadorhynchus uncinatus Fauvel, 1915 Q Dales (1957) Lopadorhynchus krohnii (Claparède, 1870) Q Fernández-Álamo (2002) Pelagobia longicirrata Greeff, 1879 Q Fernández-Álamo (1987) Family Lumbrineridae Schmarda, 1861 Lumbrineris latreilli Audouin and Milne-Edwards, 1834 Q Hartman (1944) Lumbrineris limicola Hartman, 1944 Q Solís-Weiss et al. (2000) Lumbrineris platylobata Fauchald, 1970 Solís-Weiss et al. (2000) Ninoe foliosa Fauchald, 1972 Solís-Weiss et al. (2000) Ninoe jessicae Hernández-Alcántara, Pérez-Mendoza and Solís-Weiss, 2006 Hernández-Alcántara et al. (2006) Scoletoma crassidentata (Fauchald, 1970) Solís-Weiss et al. (2000) Scoletoma tetraura (Schmarda, 1861) Q Solís-Weiss et al. (2000) Scoletoma uncinigera (Hartmann-Schröder, 1959) Gómez et al. (1997) Scoletoma zonata (Johnson, 1901) Q Solís-Weiss et al. (2000) Family Magelonidae Cunningham and Ramage, 1888 Magelona californica Hartman, 1944 Q Solís-Weiss et al. (2000) Magelona pacifica Monro, 1933 Solís-Weiss et al. (2000) Magelona pitelkai Hartman, 1944 Q Solís-Weiss et al. (2000) Magelona tehuanensis Hernández-Alcántara and Solís-Weiss, 2000 Hernández-Alcántara and Solís-Weiss (2000) Family Maldanidae Malmgren, 1867 Maldanella fibrillata Chamberlin, 1919 Q Chamberlin (1919) Aglaophamus verrilli (McIntosh, 1885) Q Solís-Weiss et al. (2000) Maldanidae sp. Chamberlin, 1919 Family Nephtyidae Grube, 1850 Chamberlin (1919) Nephtys caecoides Hartman, 1938 Q Hartman (1950) Nephtys magellanica Augener, 1912 Q Hartman (1940) Nephtys paradoxa Malm, 1874 Q Fauchald (1972) Nephtys parva Clark and Jones, 1955 Q Solís-Weiss et al. (2000) Family Nereididae Johnston, 1865 Alitta succinea (Frey and Leuckart, 1847) Q Solís-Weiss et al. (2000) Ceratocephale oculata Banse, 1977 Q Solís-Weiss et al. (2000) Ceratonereis singularis Treadwell, 1929 Gómez et al. (1997) Gymnonereis crosslandi (Monro, 1933) Gómez et al. (1997) Nereis angelensis Fauchald, 1972 Solís-Weiss et al. (2000) Nereis callaona (Grube, 1857) Gómez et al. (1997) Nereis mediator Chamberlin, 1918 Q Nereis pelagica Linnaeus, 1758 Q Solís-Weiss et al. (2000) Nereis pseudoneanthes Hartman, 1936 (=N. mediator after Hartman 1959) Q Nicon sp. Gómez et al. (1997) Gómez et al. (1997) Perinereis elenacasoae Rioja, 1947 Gómez et al. (1997) Pseudonereis gallapagensis Kinberg, 1866 Hartman (1940) Neanthes pseudonoodti Fauchald, 1977 Solís-Weiss et al. (2000) Hernández-Alcántara et al. (2003) Hartman (1940) Family Oenonidae Kinberg, 1865 Arabella iricolor (Montagu, 1804) Q Solís-Weiss et al. (2000) Oenone fulgida (Savigny in Lamarck, 1818) Q Gómez et al. (1997) Family Onuphidae Kinberg, 1865 Diopatra obliqua Hartman, 1944 Hartman (1944) Diopatra ornata Moore, 1911 Q Treadwell (1941) 347 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 8. Continued. Diopatra splendidissima Kinberg, 1865 González-Ortiz et al. (1997) Diopatra tridentata Hartman, 1944 Q González-Ortiz et al. (1997) Kinbergonuphis cedroensis (Fauchald, 1968) Q González-Ortiz et al. (1997) Mooreonuphis nebulosa (Moore, 1911) Q Fauchald (1968) Mooreonuphis peruana (Hartman, 1944) Solís-Weiss et al. (2000) Onuphis vexillaria Moore, 1911 Q Hartman (1944) Sarsonuphis litabranchia (Chamberlin, 1919) Chamberlin (1919) Family Opheliidae Malmgren, 1867 Armandia agilis (Andrews, 1891) Q Solís-Weiss et al. (2000) Armandia brevis (Moore, 1906) Q Gómez et al. (1997) Armandia intermedia Fauvel, 1902 Q Solís-Weiss et al. (2000) Ophelia assimilis Tebble, 1953 Q Ophelia pulchella Tebble, 1953 Q González-Ortiz et al. (1996) González-Ortiz et al. (1996) Ophelina acuminata Örsted, 1843 Q Gómez et al. (1997) Leitoscoloplos mexicanus (Fauchald, 1972) Solís-Weiss et al. (2000) Leitoscoloplos panamensis (Monro, 1933) Leitoscoloplos pugettensis (Pettibone, 1957) Q Solís-Weiss et al. (2000) Scoloplos (Leodamas) ohlini (Ehlers, 1901) Q Solís-Weiss et al. (2000) Aricidea suecica Eliason, 1920 Q Solís-Weiss et al. (2000) Polyophtalmus pictus (Dujardin, 1839) Q Solís-Weiss et al. (2000) Family Orbiniidae Hartman, 1957 Scoloplos cf. capensis sensu Solís-Weiss et al. (2000) Family Paraonidae Cerruti, 1909 Solís-Weiss et al. (2000) Solís-Weiss et al. (2000) Aricidea (Acmira) lopezi Berkeley and Berkeley, 1956 Q Fauchald (1972) Aricidea (Aedicira) pacifica Hartman, 1944 Q Solís-Weiss et al. (2000) Aricidea (Aricidea) fragilis Webster, 1879 Q Solís-Weiss et al. (2000) Aricidea (Aricidea) longicirrata Hartmann-Schröder, 1965 Q Solís-Weiss et al. (2000) Cirrophorus furcatus (Hartman, 1957) Q Solís-Weiss et al. (2000) Levinsenia gracilis (Tauber, 1879) Q Solís-Weiss et al. (2000) Paradoneis lyra (Southern, 1914) Q Solís-Weiss et al. (2000) Family Pectinariidae Quatrefages, 1866 Cistenides brevicoma (Johnson, 1901) Q Family Phyllodocidae Örsted, 1843 Hartman (1941) Eteone sp. Gómez et al. (1997) Eulalia myriacyclum (Schmarda, 1861) Q Gómez et al. (1997) Nereiphylla castanea (von Marenzeller, 1879) Q Gómez et al. (1997) Phyllodoce (Anaitides) lamelligera (Linnaeus, 1791) Q Gómez et al. (1997) Phyllodoce (Anaitides) williamsi Hartman, 1936 Q Gómez et al. (1997) Sigambra tentaculata (Treadwell, 1941) Q Solís-Weiss et al. (2000) Phyllodoce (Anaitides) cortezi (Kudenov, 1975) Gómez et al. (1997) Family Pilargidae de Saint-Joseph, 1899 Hermundura salazarvallejoi (de Léon-González, 1991) Solís-Weiss et al. (2000) Poecilochaetus johnsoni Hartman, 1939 Q Solís-Weiss et al. (2000) Family Poecilochaetidae Hannerz, 1956 Family Polynoidae Malmgren, 1867 Halosydna glabra Hartman, 1939 Gómez et al. (1997) Halosydna leucohyba (Schmarda, 1861) Q Gómez et al. (1997) Harmothoe sp. Gómez et al. (1997) Lepidasthenia sp. Gómez et al. (1997) Lepidonopsis humilis (Augener, 1922) Q Gómez et al. (1997) Thormora johnstoni (Kinberg, 1855) Gómez et al. (1997) Family Sabellariidae Johnston, 1865 Idanthyrsus cretus Chamberlin, 1919 Gómez et al. (1997) Idanthyrsus saxicavus (Baird, 1863) Q Gómez et al. (1997) Jasmineira pacifica Annenkova, 1937 Q González-Ortiz et al. (1996) Phragmatopoma moerchi digitata Rioja, 1962 Rioja (1962) Family Sabellidae Malmgren, 1867 Megalomma carunculata Tovar-Hernández and Salazar-Vallejo, 2008 Megalomma mushaensis (Gravier, 1906) Q Gómez et al. (1997) Megalomma splendida (Moore, 1905) Q Rioja (1962) Notaulax nudicollis (Kröyer, 1856) Q Gómez et al. (1997) Potamethus mucronatus (Moore, 1923) Q Solís-Weiss et al. (2000) Potamilla neglecta (Sars, 1851) Q Tovar-Hernández and Salazar-Vallejo (2008) Solís-Weiss et al. (2000) 348 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 8. Continued. Pseudopotamilla debilis Bush, 1905 Q Gómez et al. (1997) Pseudopotamilla reniformes (Müller, 1771) Q Gómez et al. (1997) Hydroides brachyacanthus Rioja, 1941 Gómez et al. (1997) Family Serpulidae Johnston, 1865 Crucigera websteri Benedict, 1887 Hydroides cruciger Mörch, 1863 Hydroides glandifer Rioja, 1941 Hydroides humilis (Bush, 1905) Hydroides ochoterenai Rioja, 1941 Hydroides sanctaecrucis Krøyer in Mörch, 1863 EX Hydroides trompi Bastida-Zavala and ten Hove, 2003 Pomatostegus stellatus (Abildgaard, 1789) Pseudovermilia conchata ten Hove, 1975 Q Pseudovermilia occidentalis (McIntosh, 1885) Pyrgopolon ctenactis (Mörch, 1863) Salmacina tribranchiata (Moore, 1923) Spirobranchus cf. gaymardi Spirobranchus giganteus (Pallas, 1766) Q Spirobranchus incrassatus (Krøyer, 1856) Spirobranchus minutus (Rioja, 1941) Vermiliopsis infundibulum/glandigera complex Family Sigalionidae Malmgren, 1867 Sthenelanella uniformis Moore, 1910 Q Sthenolepis fimbriarum (Hartman, 1939) Thalenessa lewisii (Berkeley and Berkeley, 1939) Bastida-Zavala (2008) Gómez et al. (1997) Bastida-Zavala (2008) Bastida-Zavala (2008) Bastida-Zavala (2008) Bastida-Zavala and ten Hove (2003) Bastida-Zavala and ten Hove (2003) Gómez et al. (1997) Gómez et al. (1997) Gómez et al. (1997) Bastida-Zavala (2008) Bastida-Zavala (2008) Bastida-Zavala (2008) Gómez et al. (1997) Gómez et al. (1997) Bastida-Zavala (2008) Bastida-Zavala (2008) Hartman (1939) Solís-Weiss et al. (2000) Solís-Weiss et al. (2000) Family Spionidae Grube, 1850 Boccardia tricuspa (Hartman, 1939) Boccardia sp. Gómez et al. (1997) Gómez et al. (1997) Dipolydora socialis (Schmarda, 1861) Q Hernández-Alcántara et al. (1994) Dispio uncinata Hartman, 1951 Q Hernández-Alcántara et al. (1994) Minuspio delta (Hartman, 1965) Q Hernández-Alcántara et al. (1994) Paraprionospio pinnata (Ehlers, 1901) Q Hernández-Alcántara et al. (1994) Prionospio pygmaeus Hartman, 1961 Q Hernández-Alcántara et al. (1994) Prionospio lighti Maciolek, 1985 Q Solís-Weiss et al. (2000) Prionospio steenstrupi Malmgren, 1867 Q Hernández-Alcántara et al. (1994) Prionospio dayi (Foster, 1969) Q Hernández-Alcántara et al. (1994) Prionospio (Minuspio) cf. lighti Hernández-Alcántara et al. (1994) Pseudopolydora sp. Gómez et al. (1997) Scolelepis (Parascolelepis) texana Foster, 1971 Q Rhynchospio glutaea (Ehlers, 1897) Q González-Ortiz et al. (1996) González-Ortiz et al. (1996) Scolelepis (Scolelepis) squamata (O.F. Muller, 1806) Q Hernández-Alcántara et al. (1994) Spiophanes cf. kroeyeri Hernández-Alcántara et al. (1994) Spiophanes wigleyi Pettibone, 1961 Q Hernández-Alcántara et al. (1994) Family Syllidae Grube, 1850 Branchiosyllis exilis (Gravier, 1900) Q Gómez et al. (1997) Brania heterocirra Rioja, 1941 Gómez et al. (1997) Haplosyllis brevicirra Rioja, 1941 Gómez et al. (1997) Odontosyllis heterodonta Góngora-Garza and de León-González, 1993 Gómez et al. (1997) Opisthosyllis brunnea Langerhans, 1879 Q Gómez et al. (1997) Opisthosyllis japonica Imajima, 1966 Q Gómez et al. (1997) Branchiosyllis pacifica Rioja, 1941 Odontosyllis polycera (Schmarda, 1861) Q Gómez et al. (1997) Gómez et al. (1997) Parasphaerosyllis indica Monro, 1937 Q Syllis alternata Moore, 1908 Q Gómez et al. (1997) Syllis cornuta Rathke, 1843 Q Syllis elongata (Johnson, 1901) Q Gómez et al. (1997) Syllis fasciata Malmgren, 1867 Q Gómez et al. (1997) Syllis hyalina Grube, 1863 Q Gómez et al. (1997) Gómez et al. (1997) Solís-Weiss et al. (2000) Syllis prolifera Krohn, 1852 Q Solís-Weiss et al. (2000) Trypanosyllis taeniaeformis (Haswell, 1886) Q Gómez et al. (1997) Family Terebellidae Malmgren, 1867 Lanicides taboguillae (Chamberlin, 1919) Pista quadrilobata (Augener, 1918) Q Gómez et al. (1997) González-Ortiz et al. (1996) 349 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 8. Continued. Polycirrus mexicanus (Rioja, 1947) Gómez et al. (1997) Family Tomopteridae Johnston, 1865 Tomopteris (Johnstonella) dunckeri Rosa, 1908 Q Fernández-Álamo (2000) Tomopteris elegans Chun, 1888 Q Fernández-Álamo (2000) Tomopteris euchaeta Chun, 1887 Q Fernández-Álamo (2000) Tomopteris nationalis Apstein, 1900 Q Fernández-Álamo (2000) Tomopteris planktonis Apstein, 1900 Q Fernández-Álamo (1987) Family Typhloscolecidae Uljanin, 1878 Sagitella kowalewskii Wagner, 1872 Q Fernández-Álamo (1987) Travisiopsis dubia Stɸp-Bowitz, 1948 Q Fernández-Álamo (1987) Typhloscolex mülleri Busch, 1851 Q Fernández-Álamo (1987) Table 9. Checklist of the crustaceans recorded in the marine and coastal areas of Oaxaca, Mexico. The list follows the classification for higher taxa of Martin and Davis (2001), with updates to the Infraorder Brachyura by Ng et al. (2008). The arrangement of Genera and species is strictly alphabetical. Phylum Arthropoda Lamarck, 1812 Reference Subphylum Crustacea Brünnich, 1772 Class Malacostraca Latreille, 1802 Subclass Eumalacostraca Grobben, 1892 Superorder Eucarida Calman, 1904 Order Decapoda Latreille, 1802 Suborder Dendrobranchiata Bate, 1888 Family Penaeidae Rafinesque, 1815 Farfantepenaeus brevirostris (Kingsley, 1878) Sosa-Hernández et al. (1980) Farfantepenaeus californiensis (Holmes, 1900) Sosa-Hernández et al. (1980) Rimapenaeus faoe (Obarrio, 1954) Litopenaeus vannamei (Boone, 1931) Hendrickx (1996) Rimapenaeus pacificus (Burkenroad, 1934) Sosa-Hernández et al. (1980) Sicyonia aliaffinis (Burkenroad, 1934) Sosa-Hernández et al. (1980) Trachysalambria brevisuturae (Burkenroad, 1934) Sosa-Hernández et al. (1980) Sosa-Hernández et al. (1980) Family Sicyoniidae Ortmann, 1898 Sicyonia disdorsalis (Burkenroad, 1934) Sicyonia martini Pérez-Farfante and Boothe, 1981 Family Solenoceridae Wood-Mason, 1891 Solenocera florea Burkenroad, 1938 Solenocera mutator Burkenroad, 1938 Sosa-Hernández et al. (1980) Pérez-Farfante (1985) Sosa-Hernández et al. (1980) Hendrickx (1996) Suborder Pleocyemata Burkenroad, 1963 Infraorder Anomura MacLeay, 1838 Family Albuneidae Stimpson, 1858 Albunea lucasia de Saussure, 1853 Lepidopa deamae Benedict, 1903 Lepidopa haigae Efford, 1971 Family Coenobitidae Dana, 1851 Coenobita compressus H. Milne-Edwards, 1837 Family Diogenidae Ortmann, 1892 Boyko (2002) Benedict (1903) Efford (1971) Ball (1972) Calcinus obscurus Stimpson, 1859 Calcinus californiensis Bouvier, 1898 Ayón-Parente and Hendrickx (2007) Hendrickx and Harvey (1999) Dardanus nudus Ayón-Parente and Hendrickx, 2009 Clibanarius janethaigae Hendrickx and Esparza-Haro, 1997 Ayón-Parente and Hendrickx (2009) Dardanus sinistripes (Stimpson, 1859) Ball and Haig (1974) Hendrickx and Esparza-Haro (1997) Dardanus stimpsoni Ayón-Parente and Hendrickx, 2009 Ayón-Parente and Hendrickx (2009) Paguristes cf. bakeri Hendrickx et al. (1997) Petrochirus californiensis Bouvier, 1895 Hendrickx et al. (1997) Family Galatheidae Samouelle, 1819 Pleuroncodes planipes Stimpson, 1860 Hendrickx et al. (1997) Family Porcellanidae Haworth, 1825 Euceramus panatelus Glassell, 1938 Haig (1960) Megalobrachium festai (Nobili, 1901) Haig (1960) Megalobrachium garthi Haig, 1957 Haig (1957) Megalobrachium sinuimanus (Lockington, 1878) Ramírez-Luna et al. (2002) Neopisosoma dohenyi Haig, 1960 García-Madrigal and Andreu-Sánchez (2010) Neopisosoma mexicanum (Streets, 1871) Haig (1960) Neopisosoma sp. Ramírez-Luna et al. (2002) Pachycheles biocellatus (Lockington, 1878) Haig (1960) 350 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 9. Continued. Pachycheles calculosus Haig, 1960 Haig (1960) Pachycheles panamensis Faxon, 1839 Haig (1960) Pachycheles setimanus (Lockington, 1878) Hernández-Álvarez and Villalobos-Hiriart (2005) Pachycheles spinidactylus Haig, 1960 Haig (1960) Petrolisthes agassizii Faxon, 1893 Petrolisthes armatus (Gibbes, 1850) García-Madrigal and Andreu-Sánchez (2010) Petrolisthes crenulatus Lockington, 1878 Petrolisthes edwardsii (Saussure, 1853) Hernández-Álvarez and Villalobos-Hiriart (2005) García-Madrigal and Andreu-Sánchez (2010) Petrolisthes cf. edwardsii Haig (1968) Haig (1960) Petrolisthes glasselli Haig, 1957 Haig (1968) Petrolisthes gracilis Stimpson, 1858 Haig (1960) Petrolisthes haigae Chace, 1962 Haig (1960) Petrolisthes hians Nobili, 1901 Haig (1968) Petrolisthes hirtispinosus Lockington, 1878 García-Madrigal and Andreu-Sánchez (2010) Petrolisthes lewisi austrinus Haig, 1960 García-Madrigal and Andreu-Sánchez (2010) Petrolisthes lewisi lewisi (Glassell, 1936) Haig (1960) Petrolisthes nobilii Haig, 1960 Haig (1968) Petrolisthes ortmanni Nobili, 1901 Haig (1968) Petrolisthes robsonae Glassell, 1945 Petrolisthes polymitus Glassell, 1937 García-Madrigal and Andreu-Sánchez (2010) Petrolisthes tonsorius Haig, 1960 Petrolisthes sanfelipensis Glassell, 1936 Haig (1960) Pisidia magdalenensis (Glassell, 1936) Haig (1960) Petrolisthes marginatus Stimpson, 1859 Porcellana cancrisocialis Glassell, 1936 Porcellana hancocki Glassell, 1938 Porcellana paguriconviva Glassell, 1936 Ulloaia perpusillia Glassell, 1938 Infraorder Brachyura Latreille, 1802 Family Aethridae Dana, 1851 Haig (1968) Haig (1960) García-Madrigal and Andreu-Sánchez (2010) Haig (1968) Hendrickx et al. (1997) Haig (1968) Ramírez-Luna et al. (2002) Hepatella amica Smith, 1869 Garth (1966) Hepatus kossmanni Newman, 1878 Garth (1966) Osachila lata Faxon, 1893 Hendrickx et al. (1997) Family Calappidae De Haan, 1833 Calappa convexa Saussure, 1853 Calappula saussurei (Rathbun, 1898) Cryptosoma bairdii (Stimpson, 1860) Platymera gaudichaudii H. Milne-Edwards, 1837 Family Dairidae Serène, 1965 Daira americana Stimpson, 1860 Family Daldorfiidae Ng and Rodriguez, 1986 Daldorfia trigona (A. Milne-Edwards, 1869) Family Domeciidae Ortmann, 1893 Domecia hispida Eydoux and Souleyet, 1842 Family Dromiidae de Haan, 1833 Garth (1966) Hendrickx (1997) Rathbun (1937) Garth (1966) Ramírez-Luna et al. (2002) Garth (1959) Crane (1947) Hypoconcha panamensis Smith in Verrill, 1869 Sosa-Hernández et al. (1980) Moreiradromia sarraburei (Rathbun, 1910) Hendrickx et al. (1997) Family Dynomenidae Ortmann, 1892 Hirsutodynomene ursula (Stimpson, 1860) Family Epialtidae MacLeay, 1838 Rathbun (1937) Epialtus minimus Lockington, 1877 Ramírez-Luna et al. (2002) Herbstia camptacantha (Stimpson, 1871) Garth (1958) Herbstia tumida (Stimpson, 1871) Hendrickx (1999) Herbstia cf. tumida Hendrickx and Vázquez-Cureño (1998) Neodoclea boneti Buitendijk, 1950 Garth (1958) Notolopas lamellatus Stimpson, 1871 Garth (1958) Pelia pacifica A. Milne-Edwards, 1875 Garth (1959) Pitho picteti (Saussure, 1853) Family Ethusidae Guinot, 1977 Ethusa lata Rathbun, 1894 Ethusa panamensis Finnegan, 1931 Family Euryplacidae Stimpson, 1871 Euryplax polita Smith, 1870 Garth (1958) Rathbun (1937) Garth (1966) Garth (1961) 351 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 9. Continued. Family Grapsidae MacLeay, 1838 Geograpsus lividus (H. Milne-Edwards, 1837) Grapsus grapsus (Linnaeus, 1758) Pachygrapsus transversus (Gibbes, 1850) Planes minutus (Linnaeus, 1758) Family Inachidae MacLeay, 1838 Podochela cf. casoae Crane (1947) Crane (1947) Crane (1947) Rathbun (1918) Ramírez-Luna et al. (2002) Podochela veleronis Garth, 1958 Garth (1959) Podochela ziesenhennei Garth, 1940 Garth (1959) Stenorhynchus debilis (Smith, 1871) Garth (1958) Family Inachoididae Dana, 1851 Collodes tenuirostris Rathbun, 1893 Euprognatha bifida Rathbun, 1894 Garth (1959) Garth (1958) Inachoides laevis Stimpson, 1860 Garth (1959) Paradasygyius depressus (Bell, 1835) Luke (1977) Hendrickx et al. (1997) Iliacantha hancocki Rathbun, 1935 Rathbun (1937) Pyromaia tuberculata (Lockington, 1877) Family Leucosiidae Samouelle, 1819 Iliacantha schmitti Rathbun, 1935 Persephona townsendi (Rathbun, 1893) Randallia bulligera Rathbun, 1898 Uhlias ellipticus Stimpson, 1871 Family Majidae Samouelle, 1819 Rathbun (1937) Rathbun (1937) Rathbun (1937) Garth (1966) Hemus finneganae Garth, 1958 Garth (1959) Hemus sp. Ramírez-Luna et al. (2002) Microphrys platysoma (Stimpson, 1860) Ramírez-Luna et al. (2002) Microphrys triangulatus (Lockington, 1877) Garth (1959) Mithraculus denticulatus (Bell, 1835) Rathbun (1925) Mithrax armatus Saussure, 1853 Crane (1947) Mithrax pygmaeus Bell, 1835 Ramírez-Luna et al. (2002) Thoe sulcata Stimpson, 1860 Rathbun (1925) Mithrax sp. Teleophrys cristulipes Stimpson, 1860 Family Ocypodidae Rafinesque, 1815 Garth (1959) Crane (1947) Ocypode occidentalis Stimpson, 1860 Rathbun (1918) Uca brevifrons (Stimpson, 1860) Crane (1975) Uca princeps (Smith, 1870) Family Oziidae Dana, 1851 Ozius perlatus Stimpson, 1860 Ozius tenuidactylus (Lockington, 1877) Ozius verreauxii de Saussure, 1853 Family Panopeidae Ortmann, 1893 Maccagno (1928) Crane (1947) Crane (1947) Crane (1947) Chasmophora macrophthalma (Rathbun, 1898) Garth (1961) Hexapanopeus orcutti Rathbun, 1930 Garth (1961) Lophoxanthus lamellipes (Stimpson, 1860) Panopeus sp. 1 Family Parthenopidae MacLeay, 1838 Crane (1947) Ramírez-Luna et al. (2002) Heterocrypta macrobrachia Stimpson, 1871 Garth (1958) Solenolambrus arcuatus Stimpson, 1871 Garth (1958) Spinolambrus exilipes (Rathbun, 1893) Garth (1958) Family Pilumnidae Samouelle, 1819 Pilumnus gracilipes A. Milne-Edwards, 1880 Ramírez-Luna et al. (2002) Pilumnus limosus Smith, 1869 Garth (1961) Pilumnus pygmaeus Boone, 1927 Garth (1961) Pilumnus sp. 1 Ramírez-Luna et al. (2002) Pilumnus sp. 2 Ramírez-Luna et al. (2002) Pilumnus sp. 3 Ramírez-Luna et al. (2002) Pilumnus sp. 4 Family Portunidae Rafinesque, 1815 Achelous asper (A. Milne-Edwards, 1861) Ramírez-Luna et al. (2002) Garth (1961) Arenaeus mexicanus (Gerstaecker, 1856) Garth (1961) Callinectes arcuatus Ordway, 1863 Garth (1961) Cronius ruber (Lamarck, 1818) Garth and Stephenson (1966) 352 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 9. Continued. Euphylax dovii Stimpson, 1860 Luke (1977) Euphylax robustus A. Milne-Edwards, 1874 Garth (1961) Portunus acuminatus (Stimpson, 1871) Garth (1961) Portunus affinis (Faxon, 1893) Garth (1961) Portunus minimus Rathbun, 1898 Sosa-Hernández et al. (1980) Portunus tuberculatus (Stimpson, 1860) Garth (1961) Portunus xantusii (Stimpson, 1860) Hendrickx et al. (1997) Portunidae sp. Luke (1977) Family Pseudorhombilidae Alcock, 1900 Oediplax granulata Rathbun, 1893 Hendrickx et al. (1997) Trapezia corallina Gerstaecker, 1856 Castro (1996) Family Trapeziidae Miers, 1886 Quadrella nitida Smith, 1869 Trapezia bidentata (Forskål, 1775) Trapezia formosa Smith, 1869 Hendrickx and Vázquez-Cureño (1998) Crane (1947) Castro (1996) Trapezia digitalis Latreille, 1828 Ramírez-Luna et al. (2002) Trapezia sp. Ramírez-Luna et al. (2002) Family Xanthidae MacLeay, 1838 ?Actaea angusta Rathbun, 1898 Cycloxanthops vittatus (Stimpson, 1860) Edwardsium lobipes (Rathbun, 1898) Heteractaea lunata (H. Milne-Edwards and Lucas, 1844) Liomera cinctimana (White, 1847) Ramírez-Luna et al. (2002) Rathbun (1930) Hendrickx et al. (1997) Hendrickx and Vázquez-Cureño (1998) Crane (1947) Microcassiope xantusii (Stimpson, 1871) Garth (1961) Micropanope sp. Ramírez-Luna et al. (2002) Paractaea sulcata (Stimpson, 1860) Crane (1947) Platyactaea dovii (Stimpson, 1871) Ramírez-Luna et al. (2002) Platypodia sp. Ramírez-Luna et al. (2002) Xanthodius sternberghii Stimpson, 1859 Crane (1947) ? Xanthodius stimpsoni (A. Milne-Edwards, 1879) Crane (1947) Xanthoidea sp. 3 Ramírez-Luna et al. (2002) Alpheus bellimanus Lockington, 1877 Wicksten (1983) Xanthoidea sp. 1 Xanthoidea sp. 2 Infraorder Caridea Dana, 1852 Family Alpheidae Rafinesque, 1815 Alpheus galapagensis Sivertsen, 1933 Alpheus longinquus Kim and Abele, 1988 Alpheus lottini Guérin-Méneville, 1829 Alpheus normanni Kingsley, 1879 Ramírez-Luna et al. (2002) Ramírez-Luna et al. (2002) Kim and Abele (1988) Martínez-Guerrero (2007) Martínez-Guerrero (2007) Hendrickx and Vázquez-Cureño(1998) Alpheus websteri Kingsley, 1880 Martínez-Guerrero (2007) Automate dolichognatha de Man, 1888 Martínez-Guerrero (2007) Synalpheus digueti Coutiére, 1909 Martínez-Guerrero (2007) Synalpheus mexicanus Coutiére, 1909 Synalpheus nobilii Coutiére, 1909 Family Hippolytidae Dana, 1852 Heptacarpus yaldwyni Wicksten, 1984 Martínez-Guerrero (2007) Wicksten (1983) Wicksten (1984) Family Ogyrididae Holthuis, 1955 Ogyrides alphaerostris (Kingsley, 1880) Wicksten and Hendrickx (1992) Ogyrides sp. Wicksten (1983) Family Palaemonidae Rafinesque, 1815 Macrobrachium acanthochirus Villalobos-Figueroa, 1967 Villalobos-Figueroa (1967) Macrobrachium americanum Bate, 1868 Martínez-Guerrero (2007) Macrobrachium hobbsi Nates-Rodríguez and Villalobos-Hiriart, 1990 Villalobos-Hiriart and Nates-Rodríguez (1990) Macrobrachium michoacanus Nates-Rodríguez and Villalobos-Hiriart, 1990 Villalobos-Hiriart and Nates-Rodríguez (1990) Macrobrachium occidentale Holthuis, 1950 Martínez-Guerrero (2007) Macrobrachium tenellum (Smith, 1871) Holthuis (1952) Palaemon ritteri Holmes, 1895 Martínez-Guerrero (2007) Family Pandalidae Haworth, 1825 Plesionika beebei Chace, 1937 Hendrickx and Wicksten (1989) Processa aequimana (Paulson, 1875) Hendrickx et al. (1997) Plesionika mexicana Chace, 1937 Wicksten (1978) Family Processidae Ortmann, 1890 353 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 9. Continued. Infraorder Palinura Latreille, 1802 Family Palinuridae Latreille, 1802 Panulirus gracilis Streets, 1871 Holthuis and Villalobos-Figueroa (1962) Panulirus inflatus (Bouvier, 1895) Holthuis and Villalobos-Figueroa (1962) Family Scyllaridae Latreille, 1825 Evibacus princeps Smith, 1869 Infraorder Thalassinidea Latreille, 1831 Hendrickx et al. (1997) Family Axiidae Huxley, 1879 Acanthaxius caespitosa (Squires, 1979) Family Thalassinidae Latreille, 1831 Corallianassa xutha (Manning, 1988) Order Euphausiacea Dana, 1852 Family Euphausiidae Dana, 1852 Euphausia distinguenda Hansen, 1908 Euphausia diomedeae Ortmann, 1894 Euphausia eximia Hansen, 1911 Euphausia lamelligera Hansen, 1911 Euphausia tenera Hansen, 1905 Nematobrachion flexipes (Ortmann, 1893) Nematoscelis atlantica Hansen, 1916 Nematoscelis gracilis Hansen, 1910 Stylocheiron affine Hansen, 1910 Stylocheiron carinatum Sars, 1883 Superorder Peracarida Calman, 1904 Order Amphipoda Latreille, 1816 Family Ampeliscidae Costa, 1857 Hendrickx (2005) Hernández-Aguilera (1998) López-Cortés (1990) López-Cortés (1990) López-Cortés (1990) López-Cortés (1990) López-Cortés (1990) López-Cortés (1990) López-Cortés (1990) López-Cortés (1990) López-Cortés (1990) López-Cortés (1990) Ampelisca agassizi (Judd, 1896) Barnard (1954) Ampelisca cristoides Barnard, 1954 Barnard (1954) Ampelisca mexicana Barnard, 1954 Barnard (1954) Ampelisca milleri Barnard, 1954 Barnard (1954) Family Caprellidae Leach, 1814 Caprella sp. Family Maeridae Krapp-Schickel, 2008 Gámez-Vivaldo et al. (2006) Elasmopus bastidai García-Madrigal, 2010 García-Madrigal (2010) Elasmopus karlae García-Madrigal, 2010 García-Madrigal (2010) Elasmopus lecroyae García-Madrigal, 2010 García-Madrigal (2010) Elasmopus marcelae García-Madrigal, 2010 García-Madrigal (2010) Elasmopus oaxaquensis García-Madrigal, 2010 García-Madrigal (2010) Elasmopus temori Barnard, 1979 García-Madrigal (2010) Elasmopus tubar Barnard, 1979 García-Madrigal (2010) Elasmopus zoanthidea Barnard, 1979 García-Madrigal (2010) Maera umarae García-Madrigal, 2010 García-Madrigal (2010) Quadrimaera chinarra (Barnard, 1979) García-Madrigal (2010) Quadrimaera reishi (Barnard, 1979) García-Madrigal (2010) Family Melitidae Bousfield, 1973 Melita bousfieldi García-Madrigal, 2010 García-Madrigal (2010) Family Phoxocephalidae Sars, 1891 Foxiphalus obtusidens (Alderman, 1936) Barnard and Barnard (1982) Rocinela signata Schioedte and Meinert, 1879 Brusca et al. (1992) Order Isopoda Latreille, 1817 Family Aegidae White, 1850 Family Cirolanidae Dana, 1852 Anopsilana oaxaca Carvacho and Haasmann, 1984 Carvacho and Haasmann (1984) Cirolana parva Hansen, 1890 Brusca et al. (1995) Excirolana braziliensis Richardson, 1912 Glynn et al. (1975) Family Corallanidae Hansen, 1890 Excorallana bruscai Delaney, 1984 Family Holognathidae Thomson, 1904 Cleantioides planicauda (Benedict, 1899) Family Idoteidae Samouelle, 1819 Parasymmerus annamaryae Brusca and Wallerstein, 1979 Family Paranthuridae Menzies and Glynn, 1968 Califanthura squamosissima (Menzies, 1951) Hendrickx and Espinosa-Pérez (1998) Brusca and Wallerstein (1979) Carvacho and Haasmann (1984) Poore (1984) 354 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 9. Continued. Family Sphaeromatidae Latreille, 1825 Dynoides crenulatus Carvacho and Haasmann, 1984 Dynoides saldanai Carvacho and Haasmann, 1984 Striella balani Glynn, 1968 Order Tanaidacea Dana, 1849 Family Apseudidae Leach, 1814 Paradoxapseudes cf. garthi Family Leptocheliidae Lang, 1973 Leptochelia cf. dubia Carvacho and Haasmann (1984) Carvacho and Haasmann (1984) Carvacho and Haasmann (1984) Jarquín-González and García-Madrigal (2009) Jarquín-González and García-Madrigal (2009) Family Metapseudidae Lang, 1970 Apseudomorpha sp. Jarquín-González and García-Madrigal (2009) Cyclopoapseudes cf. indecorus Synapseudes sp. Howard (1952) Synapseudes sp. Jarquín-González and García-Madrigal (2009) Family Paratanaidae Lang, 1949 Paratanais cf. denticulatus Jarquín-González and García-Madrigal (2009) Family Parapseudidae Gutu, 1980 Parapseudes pedispinis (Boone, 1923) Family Tanaidae Dana, 1849 Sinelobus stanfordi (Richardson, 1901) Jarquín-González and García-Madrigal (2009) Jarquín-González and García-Madrigal (2009) Subclass Hoplocarida Calman, 1904 Order Stomatopoda Latreille, 1817 Suborder Unipeltata Latreille, 1825 Family Eurysquillidae Manning, 1977 Eurysquilla veleronis (Schmitt, 1940) Schmitt (1940) Family Gonodactylidae Giesbrecht, 1910 Neogonodactylus stanschi (Schmitt, 1940) Schmitt (1940) Neogonodactylus zacae (Manning, 1972) Manning (1972) Family Lysiosquillidae Giesbrecht, 1910 Lysiosquilla desaussurei (Stimpson, 1857) (= Cloridopsis dubia after Appeltants et al. 2012) Manning (1972) Lysiosquilla panamica Manning, 1971 Barbosa-Ledesma et al. (2000) Family Pseudosquillidae Manning, 1977 Pseudosquillisma adiastalta (Manning, 1964) Manning (1972) Family Squillidae Latreille, 1802 Meiosquilla swetti (Schmitt, 1940) Manning (1972) Squilla aculeata aculeata Bigelow, 1891 Manning (1972) Squilla biformis Bigelow, 1891 Manning (1972) Squilla hancocki Schmitt, 1940 Schmitt (1940) Squilla mantoidea Bigelow, 1893 Sosa-Hernández et al. (1980) Squilla panamensis Bigelow, 1891 Manning (1972) Squilla parva Bigelow, 1891 Schmitt (1940) Squilla tiburonensis Schmitt, 1940 Sosa-Hernández et al. (1980) Squilla sp. Luke (1977) Luke (1977) Class Maxillopoda Dahl, 1956 Subclass Copepoda Milne-Edwards, 1840 Order Calanoidea Sars, 1903 Family Pontellidae Dana, 1852 Labidocera acuta (Dana, 1849) Álvarez-Silva et al. (2003) Pontella tenuiremis Giesbrecht, 1889 Álvarez-Silva et al. (2003) Labidocera darwinii Lubbock, 1853 Pontella agassizii Giesbrecht, 1895 Pontellina plumata (Dana, 1849) Order Siphonostomatoida Thorell, 1859 Family Caligidae Burmeister, 1835 Caligus bonito Wilson, 1905 Subclass Thecostraca Gruvel, 1905 Order Lepadiformes Buckeridge and Newman, 2006 Álvarez-Silva et al. (2003) Álvarez-Silva et al. (2003) Álvarez-Silva et al. (2003) Wilson (1937) Family Lepadidae Conchoderma virgatum Spengler, 1789 Order Sessilia Lamarck, 1818 Family Chelonibiidae Pilsbry, 1916 Chelonibia testudinaria (Linnaeus, 1758) Gámez-Vivaldo et al. (2006) Gámez-Vivaldo et al. (2006) 355 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 10. Checklist of the mollusks recorded in the marine areas of Oaxaca, Mexico. The systematic classification follows that of Keen (1971), modified by Skoglund (2001, 2002). Phylum Mollusca Cuvier, 1795 Reference Class Bivalvia Linnaeus, 1758 Subclass Anomalodesmata Dall, 1889 Order Pholadomyoida Newell, 1965 Superfamily Pandoroidea Rafinesque, 1815 Family Lyonsiidae Fisher, 1887 Entodesma lucasanum (Bartch and Rehder, 1939) Keen (1971) Periploma (Cricoploma) carpenteri Dall, 1896 Skoglund (2001) Superfamily Thracioidea Stoliczka, 1870 Family Periplomatidae Dall, 1895 Family Thraciidae Stoliczka, 1870 Cyathodonta dubiosa Dall, 1915 Cyathodonta lucasana Conrad, 1849 Keen (1971) Keen (1971) Subclass Heterodonta Neumayr, 1884 Order Carditoida Dall, 1889 Superfamily Crassatelloidea Family Crassatellidae Férussac, 1822 Crassinella ecuadoriana Olsson, 1961 Rodríguez-Palacios et al. (1988) Order Myoida Stoliczka, 1870 Superfamily Gastrochaeniodea J. E. Gray, 1840 Family Gastrochaenidae J. E. Gray, 1840 Gastrochaena (Rocellaria) ovata Sowerby, 1834 Rodríguez-Palacios et al. (1988) Martesia striata (Linnaeus, 1758) Zamorano et al. (2010) Superfamily Pholadoidea Lamarck, 1809 Family Pholadidae Lamarck, 1809 Order Veneroida H. Adams and A. Adams, 1856 Superfamily Cardioidea Lamarck, 1809 Family Cardiidae (Lamarck, 1809) Americardia biangulata (Broderip and Sowerby,1829) Holguín-Quiñones and González-Pedraza (1989) Laevicardium clarionense (Hertlein and Strong, 1947) Ríos-Jara et al. (2009) Laevicardium substriatum (Conrad, 1837) Rodríguez-Palacios et al. (1988) Papyridea aspersa (G.B. Sowerby I, 1833) Trachycardium (Dallocardia) senticosum (G.B. Sowerby I, 1833) Trachycardium (Mexicardia) panamense (Sowerby, 1833) Trachycardium (Mexicardia) procerum (Sowerby, 1833) Trachycardium (Trachycardium) consors (Sowerby, 1833) Trigoniocardia (Apiocardia) obovalis (G.B. Sowerby I, 1833) Superfamily Carditoidea Férussac, 1822 Family Carditidae Férussac, 1822 Holguín-Quiñones and González-Pedraza (1989) Holguín-Quiñones and González-Pedraza (1989) Holguín-Quiñones and González-Pedraza (1989) Keen (1971) Holguín-Quiñones and González-Pedraza (1989) Ríos-Jara et al. (2009) Cardites crassicostata (Sowerby, 1825) Holguín-Quiñones and González-Pedraza (1989) Cardites grayi (Dall, 1903) Holguín-Quiñones and González-Pedraza (1989) Cardites laticostata (Sowerby, 1833) Carditamera affinis (Sowerby, 1833) Carditamera radiata (Sowerby, 1833) Cyclocardia beebei (Hertlein, 1958) Superfamily Chamoidea Lamarck, 1809 Family Chamidae Lamarck, 1809 Arcinella californica (Dall, 1903) Chama (Chama) buddiana C.B. Adams, 1852 Chama (Chama) echinata Broderip, 1835 Chama (Chama) mexicana Carpenter, 1857 Holguín-Quiñones and González-Pedraza (1989) Holguín-Quiñones and González-Pedraza (1989) Holguín-Quiñones and González-Pedraza (1989) Piña-Arce and Herrera-Peña (1980) Holguín-Quiñones and González-Pedraza (1989) Zamorano et al. (2008) Holguín-Quiñones and González-Pedraza (1989) Holguín-Quiñones and González-Pedraza (1989) Chama (Chama) sordida Broderip, 1835 Rodríguez-Palacios et al. (1988) Pseudochama panamensis (Reeve, 1847) Holguín-Quiñones and González-Pedraza (1989) Family Corbiculidae J.E. Gray, 1847 Polymesoda (Polymesoda) cordata (Martens, 1865) Skoglund (2001) Family Sportellidae Dall, 1899 Keen (1971) Pseudochama corrugata (Broderip, 1833) Pseudochama inermis (Dall 1871) Superfamily Corbiculoidea J. E. Gray, 1847 Superfamily Cyamioidea Sars, 1878 Basterotia peninsularis (Jordan, 1936) Rodríguez-Palacios et al. (1988) Holguín-Quiñones and González-Pedraza (1989) Superfamily Cyrenoidoidea H. Adams and A. Adams, 1857 356 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 10. Continued. Family Cyrenoididae H. Adams and A. Adams, 1857 Cyrenoida insula Morrison, 1946 Zamorano et al. (2008) Superfamily Galeommatoidea Gray, 1840 Family Lasaeidae Gray, 1842 Kellia suborbicularis (Montagu, 1803) Rodríguez-Palacios et al. (1988) Codakia distinguenda (Tryon, 1872) Rodríguez-Palacios et al. (1988) Ctena mexicana (Dall, 1901) Ríos-Jara et al. (2009) Superfamily Lucinoidea Fleming, 1828 Family Lucinidae Fleming, 1828 Lucina (Callucina) lampra (Dall, 1901) Keen (1971) Diplodonta sericata (Reeve, 1850) Holguín-Quiñones and González-Pedraza (1989) Pegophysema (Pegophysema) edentuloides (Verrill, 1870) Piña-Arce and Herrera-Peña (1980) Family Ungulinidae H. Adams and A. Adams, 1856 Superfamily Mactroidea Lamarck, 1809 Family Mactridae Lamarck, 1809 Harvella elegans (Sowerby, 1825) Piña-Arce and Herrera-Peña (1980) Mactrellona exoleta (Gray, 1837) Ríos-Jara et al. (2009) Mulinia pallida (Broderip and Sowerby, 1829) Ríos-Jara et al. (2009) Superfamily Solenoidea Lamarck, 1809 Family Solenidae Lamarck, 1809 Solen mexicanus Dall, 1899 Solen pazensis Lowe, 1935 Solen pfeifferi Dunker, 1862 Superfamily Tellinoidea Blainville, 1814 Family Donacidae Fleming, 1828 Donax (Chion) punctatostriatus Hanley, 1843 Keen (1971) Keen (1971) Keen (1971) Holguín-Quiñones and González-Pedraza (1989) Donax (Donax) asper Hanley, 1845 Keen (1971) Donax (Paradonax) gracilis Hanley, 1845 Zamorano et al. (2010) Family Psammobiidae Fleming, 1828 Gari fucata (Hinds, 1845) Zamorano et al. (2010) Family Semelidae Stoliczka, 1870 Semele (Amphidesma) formosa (Sowerby, 1833) Semele rosea tabogaensis (G.B. Sowerby I, 1833) Semele rubropicta Dall, 1871 Family Solecurtidae d’Orbigny, 1846 Ramírez-González and Barrientos-Luján (2007) Keen (1971) Piña-Arce and Herrera-Peña (1980) Tagelus bourgeoisae Hertlein, 1951 Keen (1971) Tagelus (Taguelus) dombeii (Lamarck, 1818) Rodríguez-Palacios et al. (1988) Tagelus (Taguelus) longisinuatus Pilsbry and Lowe, 1932 Zamorano et al. (2010) Family Tellinidae Blainville, 1814 Macoma (Psammacoma) lamproleuca (Pilsbry and Lowe, 1932) Psammotreta (Psamotreta) viridotincta (Carpenter, 1856) Strigilla (Strigilla) chroma Salisbury, 1934 Ríos-Jara et al. (2009) Zamorano et al. (2010) Ríos-Jara et al. (2009) Strigilla (Strigilla) dichotoma (Philippi, 1846) Ríos-Jara et al. (2009) Tellina (Elpidollina) decumbens Carpenter, 1865 Ríos-Jara et al. (2009) Tellina (Eurytellina) prora (Hanley, 1844) Piña-Arce and Herrera-Peña (1980) Tellina (Lyratellina) lyra Hanley, 1844 Piña-Arce and Herrera-Peña (1980) Tellina (Phyllodella) insculpta Hanley, 1844 Tellina (Moerella) amianta Dall, 1900 Piña-Arce and Herrera-Peña (1980) Tellina (Scissula) virgo Hanley, 1844 Ríos-Jara et al. (2009) Temnoconcha cognata (C.B. Adams, 1852) Piña-Arce and Herrera-Peña (1980) Petricola (Petricola) lucasana Hertlein and Strong, 1948 Keen (1971) Superfamily Veneroidea Rafinesque, 1815 Family Petricolidae d’Orbigny, 1840 Family Veneridae Rafinesque, 1815 Chione (Chione) californiensis (Broderip, 1835) Piña-Arce and Herrera-Peña (1980) Holguín-Quiñones and González-Pedraza (1989) Chione (Chione) guatulcoensis Hertlein and Strong, 1948 Keen (1971) Chione (Chione) subimbricata (Sowerby, 1835) Holguín-Quiñones and González-Pedraza (1989) Chione (?Chione) tumens (Verrill, 1870) Ramírez-González and Barrientos-Luján (2007) Chione (Chione) undatella (Sowerby, 1835) Holguín-Quiñones and González-Pedraza (1989) Chione (Chionopsis) amathusia (Philippi, 1844) Ríos-Jara et al. 2009 Chione (Chionopsis) pulicaria (Broderip, 1835) Chione (Iliochione) subrugosa (Wood, 1828) Holguín-Quiñones and González-Pedraza (1989) Cyclinella producta (Carpenter, 1856) Holguín-Quiñones and González-Pedraza (1989) Holguín-Quiñones and González-Pedraza (1989) 357 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 10. Continued. Cyclinella saccata (Gould, 1851) Ríos-Jara et al. (2009) Dosinia (Dosinia) dunkeri (Philippi, 1844) Holguín-Quiñones and González-Pedraza (1989) Dosinia (Dosinia) ponderosa (Schumacher, 1817) Piña-Arce and Herrera-Peña (1980) Megapitaria aurantiaca (Sowerby, 1831) Holguín-Quiñones and González-Pedraza (1989) Megapitaria squalida (Sowerby, 1835) Holguín-Quiñones and González-Pedraza (1989) Mercenaria kellettii (Hinds, 1845) Holguín-Quiñones and González-Pedraza (1989) Periglypta multicostata (Sowerby, 1835) Holguín-Quiñones and González-Pedraza (1989) Pitar (Pitar) consanguineus (C.B. Adams, 1852) Pitar (Hysteroconcha) lupanaria (Lesson, 1830) Keen (1971) Holguín-Quiñones and González-Pedraza (1989) Pitar (Hysteroconcha) multispinosus (Sowerby, 1851) Holguín-Quiñones and González-Pedraza (1989) Pitar (Lamelliconcha) alternatus (Broderip, 1835) Holguín-Quiñones and González-Pedraza (1989) Pitar (Lamelliconcha) concinnus (Sowerby, 1835) Holguín-Quiñones and González-Pedraza (1989) Pitar (Lamelliconcha) hesperius Berry, 1960 Piña-Arce and Herrera-Peña (1980) Pitar (Lamelliconcha) vinaceus (Olsson, 1961) Holguín-Quiñones and González-Pedraza (1989) Protothaca (Leukoma) asperrima (Sowerby, 1835) Holguín-Quiñones and González-Pedraza (1989) Protothaca (Protothaca) grata (Say, 1831) Puberella lilacina (Carpenter, 1864) Timoclea squamosa (Carpenter, 1857) Tivela (Tivela) byronensis (Gray, 1838) Tivela (Tivela) delessertii (G.B. Sowerby II, 1854 ex Deshayes MS) Transennella modesta (Sowerby, 1835) Transennella puella (Carpenter, 1864) Holguín-Quiñones and González-Pedraza (1989) Piña-Arce and Herrera-Peña (1980) Rodríguez-Palacios et al. (1988) Holguín-Quiñones and González-Pedraza (1989) Ríos-Jara et al. (2009) Holguín-Quiñones and González-Pedraza (1989) Rodríguez-Palacios et al. (1988) Subclass Protobranchia Pelseneer, 1889 Order Nuculoida Dall, 1889 Superfamily Nuculanoidea H. Adams and A. Adams, 1858 Family Nuculanidae Meek, 1864 Adrana exoptata (Pilsbry and Lowe, 1932) Keen (1971) Family Tindariidae Verril and Bush, 1897 Tindaria compressa Dall, 1908 Keen (1971) Nucula (Nucula) schencki Hertlein and Strong, 1940 Keen (1971) Superfamily Nuculoidea Gray, 1824 Family Nuculidae Gray, 1824 Subclass Pteriomorphia Beurlen, 1944 Order Arcoida Stoliczka, 1871 Superfamily Arcoidea Lamarck, 1809 Family Arcidae Lamarck, 1809 Acar bailyi (Bartsch, 1931). Zamorano et al. (2008) Acar gradata (Broderip and Sowerby, 1829) Rodríguez-Palacios et al. (1988) Acar rostae Berry, 1954 Rodríguez-Palacios et al. (1988) Arca (Arca) mutabilis (Sowerby, 1833) Ríos-Jara et al. (2009) Arca (Esmerarca) reinhardi (Lowe, 1935) Arca (Larkinia) multicostata (Sowerby, 1833) Barbatia (Barbatia) lurida (Sowerby, 1833) Barbatia (Calloarca) alternata (Sowerby, 1833) Barbatia (Cucullaearca) reeveana (d’Orbignyi, 1846) Holguín-Quiñones and González-Pedraza (1989) Ramírez-González and Barrientos-Luján (2007) Rodríguez-Palacios et al. (1988) Rodríguez-Palacios et al. (1988) Zamorano et al. (2006) Fugleria illota (Sowerby, 1833) Zamorano et al. (2008) Noetia (Noetia) reversa (G.B. Sowerby I, 1833) Piña-Arce and Herrera-Peña (1980) Lunarca brevifrons (G.B. Sowerby I, 1833) Family Noetiidae Steward, 1930 Ríos-Jara et al. (2009) Superfamily Glycymeridoidea Newton, 1916 Family Glycymerididae Dall, 1908 Glycymeris (Axinactis) delessertii (Reeve, 1843) Glycymeris (Glycymeris) gigantea (Reeve, 1846) Glycymeris (Glycymeris) maculata (Broderip, 1832) Tucetona (Bellaxinea) multicostata (Sowerby, 1833) Tucetona (Tucetona) strigilata (Sowerby, 1833) Holguín-Quiñones and González-Pedraza (1989) Piña-Arce and Herrera-Peña (1980) Piña-Arce and Herrera-Peña (1980) Holguín-Quiñones and González-Pedraza (1989) Holguín-Quiñones and González-Pedraza (1989) Order Limoida Waller, 1978 Superfamily Limoidea Rafinesque, 1815 Family Limidae Rafinesque 1815 Lima tetrica Gould, 1851 Holguín-Quiñones and González-Pedraza (1989) Limaria pacifica (d’Orbygny, 1846) Holguín-Quiñones and González-Pedraza (1989) Order Mytiloida Férrusac, 1822 Superfamily Mytiloidea Rafinesque, 1815 358 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 10. Continued. Family Mytilidae Rafinesque, 1815 Brachidontes adamsianus (Dunker, 1857) Holguín-Quiñones and González-Pedraza (1989) Brachidontes semilaevis (Menke, 1849) de León-Herrera (2000) Choromytilus palliopunctatus (Carpenter, 1857) Holguín-Quiñones and González-Pedraza (1989) Gregariella coarctata (Carpenter, 1857) Zamorano et al. (2008) Lithophaga (Myoforceps) aristata (Dillwyn, 1817) Rodríguez-Palacios et al. (1988) Lioberus splendidus (Dunker, 1857) Zamorano et al. (2008) Lithophaga (Diberus) plumula (Hanley, 1843) Rodríguez-Palacios et al. (1988) Lithophaga (Labis) attenuata (Deshayes, 1836) Modiolus (Modiolus) americanus (Leach, 1815) Rodríguez-Palacios et al. (1988) Modiolus (Modiolus) capax (Conrad, 1837) Mytella guyanensis (Lamarck, 1819) Ríos-Jara et al. (2009) Mytella speciosa (Reeve, 1857) Ríos-Jara et al. (2009) Mytella strigata (Hanley, 1843) Rodríguez-Palacios et al. (1988) Septifer zeteki Hertlein and Strong, 1946 Holguín-Quiñones and González-Pedraza (1989) Holguín-Quiñones and González-Pedraza (1989) Holguín-Quiñones and González-Pedraza (1989) Order Ostreoida Férrusac, 1822 Superfamily Anomioidea Rafinesque, 1815 Family Anomiidae Rafinesque, 1815 Anomia (Anomia) peruviana d’Orbigny, 1846 Holguín-Quiñones and González-Pedraza (1989) Pododesmus (Pododesmus) pernoides (J.E. Gray, 1853) Skoglund (2001) Hyotissa hyotis (Linnaeus, 1758) Holguín-Quiñones and González-Pedraza (1989) Superfamily Ostreoidea Rafinesque, 1815 Family Gryphaeridae Vialov, 1936 Family Ostreidae Rafinesque, 1815 Crassostrea prismatica (Gray, 1825) Crassostrea palmula (Carpenter, 1857) Holguín-Quiñones and González-Pedraza (1989) Ostrea conchaphila Carpenter, 1857 Myrakeena angelica (Rochebrune, 1895) Rodríguez-Palacios et al. (1988) Superfamily Pectinoidea Rafinesque, 1815 Family Pectinidae Rafinesque, 1815 Holguín-Quiñones and González-Pedraza (1989) Holguín-Quiñones and González-Pedraza (1989) Argopecten circularis (Sowerby, 1835) Piña-Arce and Herrera-Peña (1980) Nodipecten subnodosus (Sowerby, 1835) Ramírez-González and Barrientos-Luján (2007) Spondylus limbatus G.B. Sowerby II, 1847 Ríos-Jara et al. (2009) Plicatula penicilata Carpenter, 1857 Rodríguez-Palacios et al. (1988) Argopecten ventricosus (G.B. Sowerby II, 1842) Ríos-Jara et al. (2009) Delectopecten zacae (Hertlein, 1935) Holguín-Quiñones and González-Pedraza (1989) Spondylus calcifer Carpenter, 1857 Holguín-Quiñones and González-Pedraza (1989) Family Spondylidae Gray, 1826 Superfamily Plicatuloidea Watson, 1930 Family Plicatulidae Watson, 1930 Order Pterioida Newell, 1965 Superfamily Pinnoidea Leach, 1819 Family Pinnidae Leach, 1819 Atrina (Servatrina) maura (Sowerby, 1835) Pinna rugosa Sowerby, 1835 Holguín-Quiñones and González-Pedraza (1989) Holguín-Quiñones and González-Pedraza (1989) Superfamily Pterioidea Gray 1847 Family Isognomonidae Woodring, 1925 Isognomon (Melina) janus Carpenter, 1857 Holguín-Quiñones and González-Pedraza (1989) Pinctada mazatlanica (Hanley, 1856) Holguín-Quiñones and González-Pedraza (1989) Isognomon (Melina) recognitus (Mabille, 1895) Family Pteriidae J.E. Gray 1847 Holguín-Quiñones and González-Pedraza (1989) Class Gastropoda Cuvier, 1797 Subclass Caenogastropoda Cox 1960 Order Neotaenioglossa Haller, 1892 Superfamily Calyptraeoidea Lamarck, 1809 Family Calyptraeidae Blainville, 1824 Calyptraea (Calyptraea) conica Broderip, 1834 Calyptraea (Calyptraea) mamillaris Broderip, 1834 Holguín-Quiñones and González-Pedraza (1989) Ríos-Jara et al. (2009) Calyptraea (Trochita) spirata (Forbes, 1852) Keen (1971) Cheilea cepacea (Broderip, 1834) Rodríguez-Palacios et al. (1988) Crepidula aculeata (Gmelin, 1791) Holguín-Quiñones and González-Pedraza (1989) Crepidula excavata (Broderip, 1834) Piña-Arce and Herrera-Peña (1980) Crepidula incurva (Broderip, 1834) Holguín-Quiñones and González-Pedraza (1989) 359 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 10. Continued. Crepidula lessonii (Broderip, 1834) Ríos-Jara et al. (2009) Crepidula striolata Menke, 1851 Holguín-Quiñones and González-Pedraza (1989) Crepidula uncata Menke, 1847 Rodríguez-Palacios et al. (1988) Crepidula onyx G.B. Sowerby I, 1824 Rodríguez-Palacios et al. (1988) Crepipatella lingulata (Gould, 1846) Rodríguez-Palacios et al. (1988) Crucibulum (Crucibulum) cyclopium Berry, 1969 Holguín-Quiñones and González-Pedraza (1989) Crucibulum (Crucibulum) lignarium (Broderip, 1834) Crucibulum (Crucibulum) monticulus Berry, 1969 Keen (1971) Crucibulum (Crucibulum) personatum Keen, 1958 Ríos-Jara et al. (2009) Piña-Arce and Herrera-Peña (1980) Crucibulum (Crucibulum) scutellatum (Wood, 1928) Rodríguez-Palacios et al. (1988) Crucibulum (Crucibulum) spinosum (Sowerby, 1824) Piña-Arce and Herrera-Peña (1980) Crucibulum (Crucibulum) umbrella (Deshayes, 1830) Holguín-Quiñones and González-Pedraza (1989) Family Capulidae Fleming, 1822 Capulus sericeus J. and R. Burch, 1961 Zamorano et al. (2008) Family Cancellariidae Forbes and Hanley, 1851 Keen (1971) Crucibulum pectinatum Carpenter, 1856 Superfamily Cancellarioidea Gray, 1853 Cancellaria (Euclia) balboae Pilsbry, 1931 Superfamily Cerithioidea Férussac 1822 Family Cerithiidae Férrusac 1822 Rodríguez-Palacios et al. (1988) Cerithium (Thericium) maculosum Kiener, 1841 Holguín-Quiñones and González-Pedraza (1989) Cerithium (Thericium) stercusmuscarum Valenciennes, 1833 Holguín-Quiñones and González-Pedraza (1989) Family Modulidae Fisher, 1884 Modulus catenulatus (Philippi, 1849) Zamorano et al. (2010) Modulus cerodes (A. Adams, 1851) Rodríguez-Palacios et al. (1988) Family Turritellidae Lovén 1847 Turritella banksi Reeve, 1849 Ríos-Jara et al. (2009) Vermicularia frisbeyae McLean, 1970 Rodríguez-Palacios et al. (1988) Vermicularia pellucida eburnea (Reeve, 1842) Zamorano et al. (2008) Turritella leucostoma Valenciennes, 1832 Superfamily Conoidea Fleming, 1822 Family Conidae Fleming, 1822 Piña-Arce and Herrera-Peña (1980) Conus (Asprella) arcuatus Broderip and Sowerby, 1829 Piña-Arce and Herrera-Peña (1980) Conus baccatus Sowerby, 1877 Skoglund (2002) Conus fergusoni G.B. Sowerby II, 1873 Piña-Arce and Herrera-Peña (1980) Conus purpurascens G.B. Sowerby I, 1833 Conus (Conus) brunneus Wood, 1828 Piña-Arce and Herrera-Peña (1980) Rodríguez-Palacios et al. (1988) Conus (Conus) princeps Linnaeus, 1758 Holguín-Quiñones and González-Pedraza (1989) Conus (Leptoconus) recurvus Broderip, 1833 Holguín-Quiñones and González-Pedraza (1989) Conus (Stephanoconus) nux Broderip, 1833 Holguín-Quiñones and González-Pedraza (1989) Conus (Ximeniconus) perplexus Sowerby, 1857 Holguín-Quiñones and González-Pedraza (1989) Conus (Ximeniconus) ximenes Gray, 1839 Holguín-Quiñones and González-Pedraza (1989) Family Terebridae H. Adams and A. Adams, 1854 Hastula (Impages) cinerea (Born, 1778) Holguín-Quiñones and González-Pedraza (1989) Terebra armillata Hinds, 1844 Holguín-Quiñones and González-Pedraza (1989) Terebra elata Hinds, 1844 Holguín-Quiñones and González-Pedraza (1989) Terebra robusta Hinds, 1844 Piña-Arce and Herrera-Peña (1980) Family Turridae Swainson 1840 Agladrillia flucticulus McLean and Poorman, 1971 Keen (1971) Calliclava craneana (Hertlein and Strong, 1951) Keen (1971) Calliclava rhodina McLean and Poorman, 1971 Keen (1971) Clathurella rigida (Hinds, 1843) Rodríguez-Palacios et al. (1988) Crassispira (Crassispirella) discors (Sowerby, 1834) Zamorano et al. (2008) Eleocyma ricaudae Berry, 1969 Skoglund (2002) Crassispira (Dallspira) cerithoidea (Carpenter, 1857) Cruziturricula arcuata (Reeve, 1843) Rodríguez-Palacios et al. (1988) Keen (1971) Drillia (Clathrodrillia) allyniana (Hertlein and Strong, 1951) Keen (1971) Fusiturricula armilda (Dall, 1908) Holguín-Quiñones and González-Pedraza (1989) Gemmula hindsiana Berry, 1958 Holguín-Quiñones and González-Pedraza (1989) Knefastia funiculata (Kiener, 1840, ex Valenciennes, MS) Keen (1971) Knefastia olivacea (Sowerby, 1833) Piña-Arce and Herrera-Peña (1980) Kurzia arteaga (Dall and Bartch, 1910) Kylix contracta McLean and Poorman, 1971 Keen (1971) Keen (1971) 360 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 10. Continued. Leptadrillia elissa (Dall, 1919) Keen (1971) Pilsbryspira (Pilsbryspira) garciacubasi Shaski, 1971 Zamorano et al. (2008) Polystira picta (Reeve, 1843) Holguín-Quiñones and González-Pedraza (1989) Pyrgocythara phaethusa (Dall, 1919) Splendrillia arga McLean and Poorman, 1971 Superfamily Cypraeoidea Rafinesque 1815 Family Cypraeidae Rafinesque 1815 Keen (1971) Skoglund (2002) Erosaria (Erosaria) albuginosa Gray, 1825 Holguín-Quiñones and González-Pedraza (1989) Macrocypraea cervinetta (Kiener, 1843) Holguín-Quiñones and González-Pedraza (1989) Mauritia arabicula (Lamarck, 1811) Family Ovulidae Fleming, 1822 Holguín-Quiñones and González-Pedraza (1989) Jenneria pustulata ([Lightfoot, 1786], ex Solander MS) Rodríguez-Palacios et al. (1988) Neosimnia aequalis (Sowerby, 1832) Rodríguez-Palacios et al. (1988) Simnialena rufa (Sowerby, 1832) Superfamily Eulimoidea Philippi 1853 Family Eulimidae Philippi, 1853 Eulima (Eulima) raymondi Rivers, 1904 Melanella drangai (Hertlein and Strong, 1951) Rodríguez-Palacios et al. (1988) Skoglund (2002) Keen (1971) Superfamily Janthinoidea Lamarck, 1812 Family Epitoniidae Berry, 1910 Amaea (Scalina) ferminiana (Dall, 1908) Amaea (Scalina) tehuanarum DuShane and McLean, 1968 Holguín-Quiñones and González-Pedraza (1989) Keen (1971) Epitonium (Lamelliscala) mitraeforme (Sowerby, 1844) Keen (1971) Opalia (Nodiscala) sanjuanensis (Lowe, 1932) Rodríguez-Palacios et al. (1988) Littoraria (Littoraria) zebra Gray, 1839 Ríos-Jara et al. (2009) Superfamily Littorinoidea Children, 1834 Family Littorinidae Children, 1834 Nodilittorina (Nodilittorina) aspera (Philippi, 1846) Rodríguez-Palacios et al. (1988) Nodilittorina (Fossarillitorina) modesta (Philippi, 1846) Rodríguez-Palacios et al. (1988) Superfamily Muricoidea Rafinesque, 1815 Family Buccinidae Rafinesque, 1815 Caducifer nigricostatus (Reeve, 1846) Rodríguez-Palacios et al. (1988) Cantharus (Cantharus) panamicus (Hertlein and Strong, 1951) Rodríguez-Palacios et al. (1988) Cantharus (Pollia) gemmatus (Reeve, 1846) Holguín-Quiñones and González-Pedraza (1989) Cantharus (Pollia) ringens (Reeve, 1846) Cantharus (Pollia) sanguinolentus (Duclos, 1833) Cantharus (Solenosteira) gatesi (Berry, 1963) Colubraria procera (Sowerby, 1832) Ríos-Jara et al. (2009) Holguín-Quiñones and González-Pedraza (1989) Piña-Arce and Herrera-Peña (1980) Keen (1971) Engina fusiformis Pease, 1865 Rodríguez-Palacios et al. (1988) Engina jugosa (C.B. Adams, 1852) Zamorano et al. (2008) Engina maura (Sowerby, 1832) Engina tabogaensis Bartsch, 1931 Northia northiae (Griffith and Pidgeon, 1834, ex Gray MS) Phos (Cymatophos) crassus Hinds, 1843 Family Columbellidae Swainson 1840 Aesopus (Aesopus) sanctus Dall, 1919 Anachis (Anachis) scalarina (Sowerby, 1832) Anachis phanea Dall, 1919 Columbella fuscata Sowerby, 1832 Columbella haemastoma Sowerby, 1832 Columbella major Sowerby, 1832 Zamorano et al. (2006) Holguín-Quiñones and González-Pedraza (1989) Keen (1971) Piña-Arce and Herrera-Peña (1980) Rodríguez-Palacios et al. (1988) Zamorano et al. (2006) Keen (1971) Holguín-Quiñones and González-Pedraza (1989) Rodríguez-Palacios et al. (1988) Holguín-Quiñones and González-Pedraza (1989) Columbella socorroensis Shaski, 1970 Rodríguez-Palacios et al. (1988) Columbella sonsonatensis (Mörch, 1860) Rodríguez-Palacios et al. (1988) Columbella strombiformis Lamarck, 1822 Holguín-Quiñones and González-Pedraza (1989) Cosmiconcha palmeri (Dall, 1913) Holguín-Quiñones and González-Pedraza (1989) Costoanachis hilli (Pilsbry and Lowe, 1932) Rodríguez-Palacios et al. (1988) Costoanachis ritteri (Hertlein and Strong, 1951) Keen (1971) Costoanachis treva (Baker, Hanna and Strong, 1938) Rodríguez-Palacios et al. (1988) Costoanachis lentiginosa (Hinds, 1844) Rodríguez-Palacios et al. (1988) Cotonopsis (Cotonopsis) suteri Jung, 1989 Skoglund (2002) Mitrella baccata (Gaskoin, 1851) Holguín-Quiñones and González-Pedraza (1989) Mitrella caulerpae Keen, 1971 Zamorano et al. (2006) Mitrella lalage Pilsbry and Lowe, 1932 Keen (1971) 361 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 10. Continued. Mitrella ocellata (Gmelin, 1791) Rodríguez-Palacios et al. (1988) Mitrella santabarbarensis (Gould and Carpenter, 1857) Rodríguez-Palacios et al. (1988) Nassarina (Cigclirina) helenae Keen, 1971 Keen (1971) Nassarina (Cigclirina) perata Keen, 1971 Keen (1971) Parvanachis gaskoini (Carpenter, 1857) Rodríguez-Palacios et al. (1988) Radwinia tehuantepecensis Shaski, 1970 Keen (1971) Steironepion melanosticta Pilsbry and Lowe, 1932 Rodríguez-Palacios et al. (1988) Zafrona incerta (Stearns, 1892) Family Fasciolariidae Gray, 1853 Rodríguez-Palacios et al. (1988) Fusinus (Fusinus) dupetitthouarsi (Kiener, 1840) Piña-Arce and Herrera-Peña (1980) Fusinus ambustus (Gould, 1853) Piña-Arce and Herrera-Peña (1980) Fusinus spectrum (A. Adams and Reeve, 1848) Piña-Arce and Herrera-Peña (1980) Latirus mediamericanus Hertlein and Strong, 1951 Rodríguez-Palacios et al. (1988) Leucozonia cerata (Wood, 1828) Holguín-Quiñones and González-Pedraza (1989) Leucozonia rudis (Reeve, 1847) Rodríguez-Palacios et al. (1988) Opeatostoma pseudodon (Burrow, 1815) Holguín-Quiñones and González-Pedraza (1989) Pleuroploca granosa (Broderip, 1832) Holguín-Quiñones and González-Pedraza (1989) Pleuroploca princeps (Sowerby, 1825) Holguín-Quiñones and González-Pedraza (1989) Pleuroploca salmo (Woods, 1828) Piña-Arce and Herrera-Peña (1980) Family Harpidae Bronn, 1840 Harpa crenata Swainson, 1822 Piña-Arce and Herrera-Peña (1980) Morum (Morum) tuberculosum (Reeve, 1842, ex Sowerby, MS) Rodríguez-Palacios et al. (1988) Family Melongenidae (McLean, 1996) Melongena patula (Broderip and Sowerby, 1829) Family Muricidae Rafinesque, 1815 Piña-Arce and Herrera-Peña (1980) Acanthais brevidentata (Wood, 1828) Holguín-Quiñones and González-Pedraza (1989) Acanthais (Acanthais) pyramidalis (Broderip, 1833) Rodríguez-Palacios et al. (1988) Coralliophila (Coralliophila) neritoidea (Lamarck, 1816) Rodríguez-Palacios et al. (1988) Favartia (Favartia) erosa (Broderip, 1833) Zamorano et al. (2006) Rodríguez-Palacios et al. (1988) Hexaplex regius (Swainson, 1821) Piña-Arce and Herrera-Peña (1980) Babelomurex costata (Blainville, 1832) Favartia (Favartia) lappa (Broderip, 1833) Hexaplex erythrostomus (Swainson, 1831) Rodríguez-Palacios et al. (1988) Holguín-Quiñones and González-Pedraza (1989) Hexaplex (Trunculariopsis) brassica (Lamarck, 1822) Piña-Arce and Herrera-Peña (1980) Hexaplex (Trunculariopsis) princeps (Broderip, 1833) Rodríguez-Palacios et al. (1988) Homalocantha oxyacantha (Broderip, 1833) Mancinella speciosa (Valenciennes, 1832) Mancinella triangularis (Blainville, 1832) Muricanthus ambiguus (Reeve, 1845) Rodríguez-Palacios et al. (1988) Holguín-Quiñones and González-Pedraza (1989) Holguín-Quiñones and González-Pedraza (1989) Keen (1971) Muricanthus nigritus (Philippi, 1845) Piña-Arce and Herrera-Peña (1980) Muricanthus radix (Gmelin, 1791) Ríos-Jara et al. (2009) Muricopsis (Muricopsis) zeteki Hertlein and Strong, 1951 Rodríguez-Palacios et al. (1988) Neorapana muricata (Broderip, 1832) Holguín-Quiñones and González-Pedraza (1989) Neorapana tuberculata (Sowerby, 1835) Phyllonotus peratus Keen, 1960 Ramírez-González and Barrientos-Luján (2007) Keen (1971) Plicopurpura columellaris (Lamarck, 1822) Holguín-Quiñones and González-Pedraza (1989) Tripterotyphis fayae (Keen and Campbell, 1964) Zamorano et al. (2008) Tripterotyphis lowei lowei (Pilsbry, 1931) Rodríguez-Palacios et al. (1988) Thaisella kiosquiformis (Duclos, 1832) Ríos-Jara et al. (2009) Vasula melones (Duclos, 1832) Vitularia salebrosa (King and Broderip, 1832) Zamorano et al. (2010) Family Nassariidae Iredale 1916 Nassarius bailyi (Pilsbry and Lowe, 1932) Zamorano et al. (2010) Quoyula madreporarum (Sowerby, 1834) Stramonita biserialis (Blainville, 1832) Trachypollia lugubris (C.B. Adams, 1852) Vokesimurex recurvirostris (Broderip, 1833) Nassarius catallus (Dall, 1908) Nassarius versicolor (C.B. Adams, 1852) Superfamily Naticoidea Guilding, 1834 Family Naticidae Guilding, 1834 Natica (Naticarius) chemnitzii Récluz in Reeve, 1855 Natica (Naticarius) colima Strong and Hertlein, 1937 Rodríguez-Palacios et al. (1988) Piña-Arce and Herrera-Peña (1980) Rodríguez-Palacios et al. (1988) Keen (1971) Piña-Arce and Herrera-Peña (1980) Holguín-Quiñones and González-Pedraza (1989) Rodríguez-Palacios et al. (1988) Holguín-Quiñones and González-Pedraza (1989) Holguín-Quiñones and González-Pedraza (1989) 362 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 10. Continued. Natica (Naticarius) othello Dall, 1908 Natica (Stigmaulax) broderipiana Récluz, 1844 Polinices uber (Valenciennes, 1832) Stigmaulax elenae (Récluz, 1844) Superfamily Olivoidea Olsson, 1956 Family Cystiscidae Stimpson, 1865 Keen (1971) Holguín-Quiñones and González-Pedraza (1989) Holguín-Quiñones and González-Pedraza (1989) Piña-Arce and Herrera-Peña (1980) Persicula imbricata (Hinds, 1844) Holguín-Quiñones and González-Pedraza (1989) Persicula phrygia (Hinds, 1844) Rodríguez-Palacios et al. (1988) Family Mitridae Swainson 1831 Mitra (Fusimitra) swainsonii Broderip, 1836 Rodríguez-Palacios et al. (1988) Mitra (Nebularia) lens (Wood, 1828) Zamorano et al. (2008) Mitra (Nebularia) crenata Broderip, 1836 Mitra (Nebularia) inca d’Orbigny, 1841 Family Olivellidae Troschel, 1869 Ríos-Jara et al. (2009) Olivella (Olivella) altatae (Marratin: Sowerby, 1871) Zamorano et al. (2008) Olivella (Olivella) dama (Wood, 1828, ex Mawe MS) Holguín-Quiñones and González-Pedraza (1989) Olivella (Olivella) gracilis (Broderip and Sowerby, 1829) Holguín-Quiñones and González-Pedraza (1989) Family Olividae Latreille, 1825 Agaronia testacea (Lamarck, 1811) Holguín-Quiñones and González-Pedraza (1989) Oliva (Strephona) incrassata (Lightfoot, 1786) Holguín-Quiñones and González-Pedraza (1989) Oliva (Strephona) spicata (Röding, 1798) Oliva (Strephona) polpasta Duclos, 1833 Piña-Arce and Herrera-Peña (1980) Oliva (Strephona) splendidula Sowerby, 1825 Piña-Arce and Herrera-Peña (1980) Piña-Arce and Herrera-Peña (1980) Family Vasidae H. Adams and A. Adams, 1854 Vasum caestus (Broderip, 1833) Holguín-Quiñones and González-Pedraza (1989) Barleeia bifasciata (Carpenter, 1957) Rodríguez-Palacios et al. (1988) Superfamily Rissooidea Gray, 1847 Family Barleeidae Gray, 1857 Lirobarleeia ingrami (Hertlein and Strong, 1951) Keen (1971) Family Rissoidae Gray 1847 Rissoina (Rissoina) axeliana Hertlein and Strong, 1951 Keen (1971) Rissoina (Rissoina) stricta Menke, 1850 Rodríguez-Palacios et al. (1988) Superfamily Stromboidea Rafinesque, 1815 Family Strombidae Rafinesque, 1815 Strombus (Strombus) gracilior Sowerby, 1825 Holguín-Quiñones and González-Pedraza (1989) Strombus (Tricornis) galeatus Swainson, 1823 Holguín-Quiñones and González-Pedraza (1989) Strombus (Tricornis) peruvianus Swainson, 1823 Superfamily Tonnoidea Suter, 1913 Holguín-Quiñones and González-Pedraza (1989) Family Bursidae Thiele, 1925 Bursa (Colubrellina) corrugata corrugata (Perry, 1811) Ríos-Jara et al. (2009) Marsupina nana (Broderip and Sowerby, 1829) Piña-Arce and Herrera-Peña (1980) Family Cassidae Latreille, 1825 Cypraecassis (Levenia) coarctata (Sowerby, 1825) Holguín-Quiñones and González-Pedraza (1989) Family Ficidae Meek, 1864 Ficus ventricosa (Sowerby, 1825) Piña-Arce and Herrera-Peña (1980) Family Personidae Gray, 1854 Distorsio (Distorsio) decussata (Valencianes, 1832) Piña-Arce and Herrera-Peña (1980) Family Ranellidae Gray, 1854 Monoplex lignarium (Broderip, 1833) Holguín-Quiñones and González-Pedraza (1989) Monoplex wiegmanni (Anton, 1839) Holguín-Quiñones and González-Pedraza (1989) Malea ringens (Swainson, 1822) Holguín-Quiñones and González-Pedraza (1989) Turritriton gibbosum (Broderip, 1833) Family Tonnidae Suter, 1913 Piña-Arce and Herrera-Peña (1980) Superfamily Triphoroidea Gray, 1847 Family Cerithiopsidae H. Adams and A. Adams, 1853 Cerithiopsis guatulcoensis Hertlein and Strong, 1951 Keen (1971) Cerithiopsis oaxacana Hertlein and Strong, 1951 Keen (1971) Cerithiopsis perrini Hertlein and Strong, 1951 Keen (1971) Seila assimilata (C.B.Adams, 1852) Rodríguez-Palacios et al. (1988) Triphora chathamensis Bartch, 1907 Rodríguez-Palacios et al. (1988) Seila pulmoennsis DuShane and Draper, 1975 Rodríguez-Palacios et al. (1988) Family Triphoridae Gray, 1847 Superfamily Vanikoroidea Gray, 1840 363 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 10. Continued. Family Hipponicidae Troschel, 1861 Hipponix antiquatus panamensis C.B. Adams, 1852 Hipponix grayanus Menke, 1853 Pilosabia pilosa (Deshayes, 1832) Pilosabia planatus (Carpenter, 1857) Holguín-Quiñones and González-Pedraza (1989) Rodríguez-Palacios et al. (1988) Rodríguez-Palacios et al. (1988) Rodríguez-Palacios et al. (1988) Family Vanikoridae Gray, 1840 Vanikoro aperta (Carpenter, 1864) Rodríguez-Palacios et al. (1988) Superfamily Velutinoidea Gray 1840 Family Triviidae Troschel 1863 Trivia fusca (Sowerby, 1832, ex Gray, MS) Rodríguez-Palacios et al. (1988) Hespererato columbella (Menke, 1847) Rodríguez-Palacios et al. (1988) Hespererato scabriscula (Sowerby, 1832) Zamorano et al. (2006) Trivia pacifica (Sowerby, 1832, ex Gray, MS) Trivia (Pusula) sanguinea (Sowerby, 1832) Superfamily Vermetoidea Rafinesque, 1815 Skoglund (2002) Family Planaxidae Gray, 1850 Planaxis obsoletus Menke, 1851 Keen (1971) Planaxis planicostatus Sowerby, 1825 Holguín-Quiñones and González-Pedraza (1989) Family Potamididae H. Adams and A. Adams, 1854 Cerithidea montagnei (d’Orbigny, 1839) Cerithidea (Cerithideopsis) californica mazatlanica Carpenter, 1857 Cerithidea (Cerithideopsis) valida (C.B. Adams, 1852) Family Vermetidae Rafinesque, 1815 Zamorano et al. (2010) Holguín-Quiñones and González-Pedraza (1989) Holguín-Quiñones and González-Pedraza (1989) Eualetes centiquadra (Velencienes, 1846) Holguín-Quiñones and González-Pedraza (1989) Petaloconchus (Petaloconchus) complicatus Dall, 1908 Rodríguez-Palacios et al. (1988) Serpulorbis margaritaceus (Chenu, 1844) Ríos-Jara et al. (2009) Petaloconchus (Petaloconchus) flavescens (Carpenter, 1857) Holguín-Quiñones and González-Pedraza (1989) Petaloconchus macrophagma Carpenter, 1857 Rodríguez-Palacios et al. (1988) Serpulorbis oryzata (Mörch, 1862) Holguín-Quiñones and González-Pedraza (1989) Tripsycha (Tripsycha) tripsycha (Pilsbry and Lowe, 1932) Rodríguez-Palacios et al. (1988) Vermetus (Thylaeodus) contortus (Carpenter, 1857) Keen (1983) Vermetus (Thylaeodus) indentatus (Carpenter, 1857) Rodríguez-Palacios et al. (1988) Xenophora (Xenophora) conchyliophora (Born, 1780) Keen (1971) Superfamily Xenophoroidea Troschel, 1852 Family Xenophoridae Troschel, 1852 Subclass Heterobranchia Gray, 1840 Order Heterostropha Fischer, 1885 Superfamily Architectonicoidea J.E. Gray, 1850 Family Architectonicidae Gray, 1850 Architectonica (Architectonica) nobilis Röding, 1798 Superfamily Pyramidelloidea J.E. Gray, 1840 Holguín-Quiñones and González-Pedraza (1989) Family Pyramidellidae Gray, 1840 Evalea gallegosiana (Hertlein and Strong, 1951) Keen (1971) Evalina tehuantepecana (Hertlein and Strong, 1951) Keen (1971) Pyrgiscus tehuantepecana (Hertlein and Strong, 1951) Keen (1971) Turbonilla (Mormula) guatulcoensis Hertlein and Strong, 1951 Keen (1971) Turbonilla (Pyrgolampros) soniliana Hertlein and Strong, 1951 Keen (1971) Turbonilla (Strioturbonilla) contrerasiana Hertlein and Strong, 1951 Keen (1971) Turbonilla (Strioturbonilla) oaxacana Hertlein and Strong, 1952 Keen (1971) Subclass Neritimorpha Bandel, 1992 Superfamily Neritoidea Rafinesque, 1815 Family Neritidae Rafinesque, 1815 Nerita (Cymostyla) scabricosta Lamarck, 1822 Holguín-Quiñones and González-Pedraza (1989) Nerita (Theliostyla) funiculata Menke, 1851 Holguín-Quiñones and González-Pedraza (1989) Theodoxus (Vittoclithon) luteofasciatus (Miller, 1879) Holguín-Quiñones and González-Pedraza (1989) Subclass Opistobranchia H. Milne-Edwards, 1848 Order Anaspidea Fischer, 1883 Superfamily Aplysioidea Lamarck, 1809 Family Aplysiidae Lamarck, 1809 Aplysia (Neaplysia) californica Cooper, 1863 Dolabella auricularia (Lighfoot, 1786) Rodríguez-Palacios et al. (1988) Rodríguez-Palacios et al. (1988) Order Cephalaspidea Fischer, 1883 Superfamily Bulloidea Lamarck, 1801 364 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 10. Continued. Family Bullidae Rafinesque, 1815 Bulla (Bulla) punctulata A. Adams in: Sowerby, 1850 Holguín-Quiñones and González-Pedraza (1989) Order Gymnophila Baker 1955 Superfamily Onchidioidea Rafinesque, 1815 Family Onchidiidae Rafinesque, 1815 Hoffmannola hansi (Stearns, 1892) Rodríguez-Palacios et al. (1988) Hoffmannola lesliei (Stearns, 1892) Holguín-Quiñones and González-Pedraza (1989) Order Nudibranchia Blainville, 1814 Family Dendrodorididae O’donoghue, 1924 Dendrodoris fumata (Rüppell and Leuckart, 1830) Family Discodorididae Bergh, 1891 Dialula sandiegensis (Cooper, 1863) Rodríguez-Palacios et al. (1988) Rodríguez-Palacios et al. (1988) Family Tritoniidae H. Adams and A. Adams, 1858 Tritonia festiva (Stearns, 1873) Rodríguez-Palacios et al. (1988) Subclass Patellogastropoda Lindberg, 1988 Superfamily Acmaeoidea Forbes, 1850 Family Lottiidae Gray, 1840 Lottia acutapex (Berry, 1960) Holguín-Quiñones and González-Pedraza (1989) Lottia discors (Philippi, 1849) Holguín-Quiñones and González-Pedraza (1989) Lottia mesoleuca (Menke, 1851) Holguín-Quiñones and González-Pedraza (1989) Lottia mitella (Menke, 1847) Holguín-Quiñones and González-Pedraza (1989) Lottia pediculus (Philippi, 1846) Holguín-Quiñones and González-Pedraza (1989) Lottia strongiana (Hertlein, 1958) Rodríguez-Palacios et al. (1988) Macrarene spectabilospina Shasky, 1970 Keen (1971) Tectura fascicularis (Menke, 1851) Holguín-Quiñones and González-Pedraza (1989) Tectura ubiquita (Lindberg and McLean, 1981) Skoglund (2002) Superfamily Eoacmaeoidea Nakano and Ozawa, 2007 Family Eoacmaeidae Nakano and Ozawa, 2007 Eoacmaea semirubida (Dall, 1914) Rodríguez-Palacios et al. (1988) Superfamily Fissurelloidea Fleming 1822 Family Fissurellidae Fleming 1822 Diodora inaequalis (Sowerby, 1835) Holguín-Quiñones and González-Pedraza (1989) Diodora saturnalis (Carpenter, 1864) Rodríguez-Palacios et al. (1988) Fissurella (Cremides) decemcostata McLean, 1970 Holguín-Quiñones and González-Pedraza (1989) Fissurella (Cremides) gemmata Menke, 1847 Holguín-Quiñones and González-Pedraza (1989) Fissurella (Cremides) microtrema Sowerby, 1835 Rodríguez-Palacios et al. (1988) Fissurella (Cremides) rubropicta Pilsbry, 1890 Keen (1971) Fissurella (Cremides) spongiosa Carpenter, 1857 Holguín-Quiñones and González-Pedraza (1989) Fissurella (Cremides) asperella Sowerby, 1835 Fissurella (Cremides) nigrocincta Carpenter, 1856 Lucapinella aequalis (G.B. Sowerby, 1835) Octomarginula natlandi (Durham, 1950) Puncturella (Puncturella) punctocostata Berry, 1947 Superfamily Patelloidea Rafinesque, 1815 Family Patellidae Rafinesque, 1815 Scutellastra mexicana (Broderip and G.B. Sowerby I, 1829) Holguín-Quiñones and González-Pedraza (1989) Holguín-Quiñones and González-Pedraza (1989) Keen (1971) Rodríguez-Palacios et al. (1988) Rodríguez-Palacios et al. (1988) Holguín-Quiñones and González-Pedraza (1989) Subclass Pulmonata Cuvier, 1817 Order Hygrophila Férussac, 1822 Superfamily Siphonarioidea Gray, 1827 Family Siphonariidae Gray, 1827 Siphonaria (Heterosiphonaria) maura Sowerby, 1835 Siphonaria (Heterosiphonaria) palmata Carpenter, 1857 Willamia peltoides (Carpenter, 1864) Holguín-Quiñones and González-Pedraza (1989) Holguín-Quiñones and González-Pedraza (1989) Rodríguez-Palacios et al. (1988) Subclass Vetigastropoda Salvini-Plawen, 1980 Superfamily Trochoidea Rafinesque 1815 Family Trochidae Rafinesque 1815 Tegula (Agathistoma) corteziana McLean, 1970 de León-Herrera (2000) Tegula (Agathistoma) globulus (Carpenter, 1857) Ramírez-González and Barrientos-Luján (2007) Family Turbinidae Rafinesque 1815 Astraea (Uvanilla) olivacea (Wood, 1828) Holguín-Quiñones and González-Pedraza (1989) Astraea (Uvanilla) unguis (Wood, 1828) Holguín-Quiñones and González-Pedraza (1989) Class Polyplacophora Gray, 1821 Order Neoloricata Bergernhayn, 1955 365 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 10. Continued. Superfamily Lepidopleuroidea Pilsbry, 1892 Family Acanthochitonidae Pilsbry, 1893 Acanthochitona angelica Dall, 1919 Rodríguez-Palacios et al. (1988) Acanthochitona hirundiniformis (Carpenter, 1864) Rodríguez-Palacios et al. (1988) Callistoplax retusa (Sowerby in: Broderip and Sowerby, 1832) Reyes-Gómez et al. (2010) Chiton (Chiton) albolineatus Broderip and Sowerby, 1829 Rodríguez-Palacios et al. (1988) Chiton (Chiton) articulatus Sowerby in: Broderip and Sowerby, 1832 Rodríguez-Palacios et al. (1988) Acanthochitona arragonites (Carpenter, 1857) Reyes-Gómez et al. (2010) Acanthochitona rodhea (Pilsbry, 1893) Rodríguez-Palacios et al. (1988) Acanthochitona avicula (Carpenter, 1864) Family Callistoplacidae Pilsbry, 1893 Family Chitonidae Rafinesque, 1815 Tonicia forbesii forbesii Carpenter, 1857 Family Ischnochitonidae Dall 1889 Holguín-Quiñones and González-Pedraza (1989) Rodríguez-Palacios et al. (1988) Callistochiton colimensis (A.G. Smith, 1961) Rodríguez-Palacios et al. (1988) Callistochiton gabbi Pilsbry, 1893 Rodríguez-Palacios et al. (1988) Chaetopleura (Pallochiton) lanuginosa mixta (Dall, 1919) Zamorano et al. (2008) Ischnochiton (Ischnochiton) tenuisculptus (Carpenter, 1864) Rodríguez-Palacios et al. (1988) Chaetopleura (Chaetopleura) hanselmani (Ferreira, 1982) Reyes-Gómez et al. (2010) Chaetopleura (Chaetopleura) lurida (Sowerby in: Broderip and Sowerby, 1832) Rodríguez-Palacios et al. (1988) Lepidochitona (Dendrochiton) lirulata (Berry, 1963) Rodríguez-Palacios et al. (1988) Lepidochitona (Lepidochitona) beanii (Carpenter, 1857) Rodríguez-Palacios et al. (1988) Stenoplax (Stenoplax) limaciformis (Sowerby in Broderip and Sowerby, 1832) Reyes-Gómez et al. (2010) Lepidochitona (Lepidochitona) salvadorensis García Ríos, 2006 Reyes-Gómez et al. (2010) Lepidozona (Lepidozona) serrata (Carpenter, 1864) Reyes-Gómez et al. (2010) Stenoplax (Stenoplax) rugulata (Sowerby In: Broderip and Sowerby, 1832) Rodríguez-Palacios et al. (1988) Stenoplax (Stenoplax) mariposa (Bartsch MS, Dall, 1919) Reyes-Gómez et al. (2010) Class Cephalopoda Cuvier, 1795 Subclass Coleoidea Bather, 1888 Order Octopoda Leach, 1818 Superfamily Octopodoidea Orbigny, 1840 Family Octopodidae Orbigny, 1840 Octopus chierchiae Jatta, 1889 Octopus hubbsorum Berry, 1953 Octopus rubescens Berry, 1953 Order Myopsida Naef, 1916 Family Loliginidae Lesuer, 1821 Lolliguncula diomedeae Hoyle, 1904 Lolliguncula panamensis Berry, 1911 Order Oegopsida d’Orbigny, 1845 Family Omastrephidae Steenstrup, 1857 Dosidicus gigas (d’Orbigny, 1835) Table 11. Checklist of the bryozoans recorded in the marine areas of Oaxaca, Mexico. Sánchez (2003) Sánchez (2003) Sánchez (2003) Sánchez (2003) Sánchez (2003) Sánchez (2003) Phylum Bryozoa Ehrenberg, 1831 Reference Class Gymnolaemata Allman, 1856 Order Cheilostomatida Busk, 1852 Suborder Neocheilostomatina d’Hondt, 1985 Infraorder Ascophora Levinsen, 1909 Superfamily Schizoporelloidea Jullien, 1883 Family Cryptosulidae Vigneaux, 1949 Cryptosula pallasiana (Moll, 1803) Table 12. Checklist of the echinoderms recorded in the marine areas of Oaxaca, Mexico. Phylum Echinodermata Bruguiere, 1789 Osburn (1953) Reference Class Asteroidea Blainville, 1830 Order Paxillosida Perrier, 1884 Family Astropectinidae Gray, 1840 Astropecten armatus Gray, 1840 Family Luidiidae Sladen, 1889 Luidia bellonae Lütken, 1864 Luidia columbia (Gray, 1840) Luidia latiradiata (Gray, 1871) Honey-Escandón et al. (2008) Honey-Escandón et al. (2008) Honey-Escandón et al. (2008) Solís-Marín et al. (1993) 366 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 12. Continued. Luidia superba A.H. Clark, 1917 Honey-Escandón et al. (2008) Eremicaster crassus (Sladen, 1883) Benítez-Villalobos et al. (2008) Heliaster microbrachius Xantus, 1860 Solís-Marín et al. (1993) Asteropsis carinifera (Lamarck, 1816) Honey-Escandón et al. (2008) Luidia tessellata Lütken, 1859 Family Porcellanasteridae Sladen, 1889 Honey-Escandón et al. (2008) Order Forcipulatida Perrier, 1884 Family Heliasteridae Viguier, 1878 Order Valvatida Perrier, 1884 Family Asteropseidae Hotchkiss and A.M. Clark, 1976 Family Mithrodiidae Viguier, 1878 Mithrodia bradleyi Verrill, 1867 Benítez-Villalobos et al. (2008) Family Ophidiasteridae Verrill, 1870 Leiaster teres (Verrill, 1871) Benítez-Villalobos et al. (2008) Pharia pyramidatus (Gray, 1840) Zamorano and Leyte-Morales (2005) Phataria unifascialis (Gray, 1840) Zamorano and Leyte-Morales (2005) Family Oreasteridae Fisher, 1911 Nidorellia armata (Gray, 1840) Benítez-Villalobos et al. (2008) Pentaceraster cumingi (Gray, 1840) Benítez-Villalobos et al. (2008) Class Ophiuroidea Gray, 1840 Order Ophiurida Müller and Troschel, 1840 Family Amphiuridae Ljungman, 1867 Ophiophragmus marginatus (Lütken, 1859) Family Ophiacanthidae Perrier, 1891 Honey-Escandón et al. (2008) Ophiacantha paucispina Lütken and Mortensen, 1899 Honey-Escandón et al. (2008) Ophiactis simplex (Le Conte, 1851) Benítez-Villalobos (2001) Ophiocoma alexandri Lyman, 1860 Zamorano and Leyte-Morales (2005) Family Ophiactidae Matsumoto, 1915 Ophiactis savignyi (Müller and Troschel, 1842) Benítez-Villalobos (2001) Family Ophiocomidae Ljungman, 1867 Ophiocoma aethiops Lütken, 1859 Zamorano and Leyte-Morales (2005) Family Ophiodermatidae Ljungman, 1867 Diopederma daniana (Verrill, 1867) Honey-Escandón et al. (2008) Ophiocryptus granulosus Nielsen, 1932 Benítez-Villalobos (2001) Ophioderma sp. Granja-Fernández and López-Pérez, 2011 Ophioderma teres (Lyman, 1860) Ophioderma variegata Lütken, 1856 Family Ophionereididae Ljungman, 1867 Honey-Escandón et al. (2008) Ophionereis annulata (Le Conte, 1851) Zamorano and Leyte-Morales (2005) Ophiothrix rudis Lyman, 1874 Benítez-Villalobos (2001) Ophiothrix spiculata Le Conte, 1851 Zamorano and Leyte-Morales (2005) Family Ophiotrichidae Ljungman, 1866 Ophiothela mirabilis Verrill, 1867 Granja-Fernández and López-Pérez, 2011 Family Ophiuridae Lyman, 1865 Ophiolepis pacifica Lütken, 1856 Ophiolepis variegata Lütken, 1856 Granja-Fernández and López-Pérez, 2011 Honey-Escandón et al. (2008) Ophiomusium glabrum Lütken and Mortensen, 1899 Honey-Escandón et al. (2008) Family Cassidulidae L. Agassiz and Desor, 1846 Ryncholampas pacificus (A. Agassiz, 1863) Benítez-Villalobos et al. (2008) Meoma ventricosa grandis Gray, 1851 Caso (1983) Class Echinoidea Leske, 1778 Order Cassiduloida L. Agassiz and Desor, 1847 Order Spatangoida Claus, 1876 Family Brissidae Gray, 1825 Family Loveniidae Lambert, 1905 Lovenia cordiformis Agassiz, 1872 Family Schizasteridae Lambert, 1905 Agassizia scrobiculata Valenciennes, 1846 Order Diadematoida Duncan, 1889 Family Diadematidae Gray, 1855 Astropyga pulvinata (Lamarck, 1816) Diadema mexicanum Agassiz, 1863 Order Echinoida Claus, 1876 Caso (1983) Benítez-Villalobos et al. (2008) Caso (1978) Benítez-Villalobos (2001) 367 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 12. Continued. Family Echinometridae Gray, 1825 Echinometra vanbrunti Agassiz, 1863 Family Toxopneustidae Troschel, 1872 Tripneustes depressus Agassiz, 1863 Caso (1978) Benítez-Villalobos et al. (2008) Toxopneustes roseus (A. Agassiz, 1863) Zamorano and Leyte-Morales (2005) Encope micropora L. Agassiz, 1841 Caso (1980) Order Clypeasteroida Agassiz, 1872 Family Mellitidae Stefanini, 1911 Order Cidaroida Claus, 1880 Family Cidariidae Gray, 1825 Eucidaris thouarsii (Valenciennes, 1846) Hesperocidaris asteriscus Clark, 1948 Class Holothuroidea Blainville, 1834 Order Dendrochirotida Grube, 1840 Family Cucumariidae Ludwig, 1894 Pseudocnus californicus (Semper, 1868) Staurocucumis abyssorum (Theél, 1886) Family Psolidae Perrier, 1902 Lissothuria ornata Verril, 1867 Family Phyllophoridae Östergren, 1907 Euthyodiniella zacae (Deichmann 1938) Family Sclerodactylidae Panning, 1949 Afrocucumis ovulum (Selenka, 1867) Pentamera zacae Deichmann, 1938 Order Aspidochirotida Grube, 1840 Family Holothuriidae Ludwing, 1894 Holothuria (Stauropora) fuscocinerea Jaeger, 1833 Holothuria (Selenkothuria) lubrica Selenka, 1867 Holothuria (Selenkothuria) portovallartensis Caso, 1954 Holothuria (Selenkothuria) theeli Deichmann, 1938 Holothuria (Semperothuria) imitans Ludwig, 1875 Holothuria (Thymiosycia) arenicola Semper, 1868 Holothuria (Thymiosycia) impatiens (Forskaal, 1775) Caso (1978) Zamorano and Leyte-Morales (2005) Deichmann (1941) Deichmann (1941) Deichmann (1941) Deichmann (1938) Deichmann (1941) Deichmann (1938) Zamorano and Leyte-Morales (2005) Deichmann (1958) Deichmann (1958) Deichmann (1938) Zamorano and Leyte-Morales (2005) Zamorano and Leyte-Morales (2005) Benítez-Villalobos (2001) Family Stichopodidae Haeckel, 1896 Isostichopus fuscus (Ludwig, 1875) Honey-Escandón et al. (2008) Table 13. Checklist of the fish recorded in the marine areas of Oaxaca, Mexico. A= amphidromous; C= catadromous. Phylum Chordata Bateson, 1885 Reference Class Chondrichthyes Goodrich, 1909 Order Carcharhiniformes Compagno, 1973 Family Carcharhinidae Jordan and Evermann, 18961 Carcharhinus albimarginatus Rüppell, 1837 Carcharhinus falciformis (Müller and Henle, 1839) Carcharhinus galapagensis (Snodgrass and Heller, 1905) Carcharhinus leucas (Müller and Henle, 1839) A Carcharhinus limbatus (Müller and Henle, 1839) A Carcharhinus longimanus (Poey, 1861) Carcharhinus porosus (Ranzani, 1839) Galeocerdo cuvieri (Péron and Lesueur, 1822) Nasolamia velox (Gilbert, 1898) Negaprion brevirostris (Poey, 1868) Prionace glauca (Linnaeus, 1758) Rhizoprionodon longurio (Jordan and Gilbert, 1882) Family Sphyrnidae Gill, 1872 Sphyrna corona Springer, 1940 Sphyrna lewini (Griffith and Smith, 1834) Sphyrna media Springer, 1940 Sphyrna mokarran (Rüppell, 1837) Sphyrna tiburo (Linnaeus, 1758) Sphyrna zygaena (Linnaeus, 1758) Family Triakidae Gray, 1851 Mustelus dorsalis Gill, 1864 Mustelus lunulatus Jordan and Gilbert, 1882 Compagno et al. (1995) Compagno et al. (1995) Compagno et al. (1995) Compagno et al. (1995) Compagno et al. (1995) Compagno et al. (1995) Compagno et al. (1995) Compagno et al. (1995) Compagno et al. (1995) Compagno et al. (1995) Compagno et al. (1995) Compagno et al. (1995) Compagno et al. (1995) Tapia-García et al. (1995) Compagno et al. (1995) Compagno et al. (1995) Compagno et al. (1995) Compagno et al. (1995) Compagno et al. (1995) Tapia-García et al. (1995) 368 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 13. Continued. Order Heterodontiformes Garman, 1885 Family Heterodontidae Gray, 1851 Heterodontus francisci (Girard, 1855) Heterodontus mexicanus Taylor and Castro-Aguirre, 1972 Order Lamniformes Berg, 1940 Family Alopiidae Bonaparte, 1938 Alopias pelagicus Nakamura, 1935 Alopias superciliosus (Lowe, 1841) Family Lamnidae (Müller and Henle, 1838) Compagno et al. (1995) Compagno et al. (1995) Compagno et al. (1995) Compagno et al. (1995) Isurus oxyrhinchus Rafinesque, 1810 Compagno et al. (1995) Ginglymostoma cirratum (Bonnaterre, 1788) Compagno et al. (1995) Order Orectolobiformes Compagno, 1973 Family Ginglymostomatidae Gill, 1862 Family Rhincodontidae Müller and Henle, 1839 Rhincodon typus Smith, 1828 Order Pristiformes Buen, 1926 Family Pristidae Bonaparte, 1838 Pristis microdon Latham, 1794 A Pristis pectinata Latham, 1794 A Pristis perotteti Muller and Henle, 1841 A Order Rajiformes Berg, 1940 Family Dasyatididae Jordan, 1888 Dasyatis brevis (Garman, 1880) Compagno et al. (1995) Castro-Aguirre and Espinoza-Pérez (1996) Castro-Aguirre and Espinoza-Pérez (1996) Robertson and Allen (2008) McEachran and Notarbartolo-di-Sciara (1995) Dasyatis dipterura (Jordan and Gilbert, 1880) Robertson and Allen (2008) Dasyatis longa (Garman, 1880) McEachran and Notarbartolo-di-Sciara (1995) Himantura pacifica (Beebe and Tee-Van, 1941) Pteroplatytrygon violacea (Bonaparte, 1832) Family Gymnuridae Fowler, 1934 Gymnura crebripunctata (Peters, 1869) Gymnura marmorata (Cooper, 1864) Family Myliobatididae Bonaparte, 1838 McEachran and Notarbartolo-di-Sciara (1995) Castro-Aguirre and Espinoza-Pérez (1996) McEachran and Notarbartolo-di-Sciara (1995) McEachran and Notarbartolo-di-Sciara (1995) Aetobatus narinari (Euphrasen, 1790) McEachran and Notarbartolo-di-Sciara (1995) Mobula tarapacana (Philippi, 1892) McEachran and Notarbartolo-di-Sciara (1995) Manta birostris (Dondorff, 1798) Mobula japonica (Müller and Henle, 1841) Mobula thurstoni (Lloyd, 1908) Rhinoptera steindachneri Evermann and Jenkins, 1891 Family Rajidae Blainville, 1816 Raja cortezensis McEachran and Miyake, 1988 McEachran and Notarbartolo-di-Sciara (1995) McEachran and Notarbartolo-di-Sciara (1995) McEachran and Notarbartolo-di-Sciara (1995) McEachran and Notarbartolo-di-Sciara (1995) McEachran and Notarbartolo-di-Sciara (1995) Raja equatoralis Jordan and Bollman, 1890 Tapia-García et al. (1995) Raja velezi Chirichigno, 1973 McEachran and Notarbartolo-di-Sciara (1995) Family Rhinobatidae Müller and Henle, 1837 Rhinobatos glaucostigma Jordan and Gilbert, 1884 Rhinobatos leucorhynchus Günther, 1867 Rhinobatos planiceps Garman, 1880 McEachran and Notarbartolo-di-Sciara (1995) Tapia-García et al. (1995) Zapteryx exasperata (Jordan and Gilbert, 1880) Tapia-García et al. (1995) Zapteryx xyster Jordan and Evermann, 1896 Robertson and Allen (2008) Family Urolophidae Müller and Henle, 1841 Urobatis concentricus Osburn and Nichols, 19l6 Urobatis halleri (Cooper, 1863) Family Urotrygonidae McEachran, Dunn and Miyake, 1996 Urotrygon aspidura (Jordan and Gilbert, 1882) Urotrygon chilensis (Günther, 1871) Urotrygon munda Gill, 1863 Urotrygon nana Miyake and McEachran, 1988 Urotrygon rogersi (Jordan and Starks, 1895) McEachran and Notarbartolo-di-Sciara (1995) McEachran and Notarbartolo-di-Sciara (1995) McEachran and Notarbartolo-di-Sciara (1995) Tapia-García et al. (1995) Castro-Aguirre and Espinoza-Pérez (1996) Tapia-García et al. (1995) Tapia-García et al. (1995) Order Torpediniformes Buen, 1926 Family Narcinidae Gill, 1862 Diplobatis ommata (Jordan and Gilbert, 1890) Narcine entemedor Jordan and Starks, 1895 Narcine vermiculatus Breder, 1928 McEachran and Notarbartolo-di-Sciara (1995) Tapia-García et al. (1995) Tapia-García et al. (1995) Class Actinopterygii Klein, 1885 Order Albuliformes Greenwood, 1977 369 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 13. Continued. Family Albulidae Bleeker, 1859 Albula esuncula (Garman, 1899) Robertson and Allen (2008) Albula vulpes (Linnaeus, 1758) A Chávez (1979) Order Anguilliformes Regan, 1909 Family Chlopsidae Clausen, 1959 Chlopsis apterus (Beebe and Tee-Van, 1938) Robertson and Allen (2008) Chlopsis kazuko Lavenberg, 1988 Family Congridae Kaup, 1856 Robertson and Allen (2008) Arisoma gilberti (Ogilby, 1898) Allen et al. (1995) Bathycongrus macrurus (Gilbert, 1891) Allen et al. (1995) Bathycongrus varidens (Garman, 1899) Allen et al. (1995) Chiloconger dentatus (Garman, 1899) Robertson and Allen (2008) Chiloconger obtusus (Garman, 1899) Gnathophis cinctus (Garman, 1899) Allen et al. (1995) Gorgasia punctata Meek and Hildebrand, 1923 Allen et al. (1995) Paraconger californiensis Kanazawa, 1961 Allen et al. (1995) Heteroconger pellegrini Castle, 1999 Japonoconger proriger (Gilbert, 1891) Paraconger similis (Wade, 1946) Rhynchoconger nitens (Jordan and Bollman, 1890) Allen et al. (1995) Xenomystax atrarius Gilbert, 1891 Allen et al. (1995) Family Moringuidae Clausen, 1959 Neoconger vermiformis Gilbert, 1890 Family Muraenidae Günther, 1870 Robertson and Allen (2008) Echidna nebulosa (Ahl, 1789) Allen et al. (1995) Echidna nocturna (Cope, 1872) Allen et al. (1995) Robertson and Allen (2008) Gymnothorax castaneus (Jordan and Gilbert, 1882) Allen et al. (1995) Gymnothorax equatorialis (Hildebrand, 1946) Allen et al. (1995) Gymnomuraena zebra (Shaw, 1797) Gymnothorax eurygnathos Bohlke, 2001 Robertson and Allen (2008) Gymnothorax phalarus Bussing, 1998 Robertson and Allen (2008) Gymnothorax panamensis (Steindachner, 1876) Tapia-García et al. (1995) Muraena argus (Steindachner, 1870) Allen et al. (1995) Muraena clepsydra Gilbert, 1898 Allen et al. (1995) Muraena lentiginosa Jenyns, 1842 Allen et al. (1995) Uropterygius macrocephalus (Bleeker, 1865) Allen et al. (1995) Family Muraenesocidae Cope, 1872 Cynoponticus coniceps (Jordan and Gilbert, 1881) Family Ophichthidae Jordan and Evermann, 1896 Apterichtus equatorialis (Myers and Wade, 1941) Allen et al. (1995) Allen et al. (1995) Bascanichthys bascanoides Osborn and Nichols, 1916 Allen et al. (1995) Bascanichthys panamensis Meek and Hildebrand, 1923 Allen et al. (1995) Callechelys cliffi Böhlke and Briggs, 1954 Allen et al. (1995) Callechelys eristigma McCosker and Rosenblatt, 1972 Echiophis brunneus (Castro-Aguirre and Suárez de los Cobos, 1983) Allen et al. (1995) Ichthyapus selachops (Jordan and Gilbert, 1882) Allen et al. (1995) Letharchus rosenblatti McCosker, 1974 Allen et al. (1995) Allen et al. (1995) Myrichthys tigrinus Girard, 1859 Allen et al. (1995) Myrophis vafer Jordan and Gilbert, 1881 Allen et al. (1995) Ophichthus apachus McCosker and Rosenblatt, 1998 Ophichthus frontalis Garman, 1899 Allen et al. (1995) Ophichthus longipenis McCosker and Rosenblatt, 1998 Robertson and Allen (2008) Ophichthus triserialis (Kaup, 1856) Tapia-García et al. (1995) Ophichthus zophochir Jordan and Gilbert, 1882 Tapia-García et al. (1995) Herpetoichthys fossatus (Myers and Wade, 1941) Myrichthys aspetocheiros McCosker and Rosenblatt, 1993 Ophichthus mecopterus McCosker and Rosenblatt, 1998 Allen et al. (1995) Allen et al. (1995) Robertson and Allen (2008) Robertson and Allen (2008) Paraletharchus pacificus (Osburn and Nichols, 1916) Allen et al. (1995) Phaenomonas pinnata Myers and Wade, 1941 Allen et al. (1995) Quassiremus nothochir (Gilbert, 1890) Allen et al. (1995) Scytalichthys miurus (Jordan and Gilbert, 1882) Allen et al. (1995) Order Atheriniformes Goüan, 1770 Family Atherinopsidae Fowler, 1903 370 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 13. Continued. Atherinella eriarcha (Jordan and Gilbert, 1881) Allen et al. (1995) Atherinella guatemalensis (Günther, 1864) A Allen et al. (1995) Atherinellanepenthe (Myers and Wade, 1942) Allen et al. (1995) Atherinella nesiotes Myers and Wade, 1942 Allen et al. (1995) Order Aulopiformes Rosen, 1973 Family Synodontidae Gill, 1872 Synodus evermanni Jordan and Bollman, 1889 Tapia-García et al. (1995) Synodus lacertinus Gilbert, 1890 Allen et al. (1995) Synodus scituliceps Jordan and Gilbert, 1881 Tapia-García et al. (1995) Synodus sechurae Hildebrand, 1946 Allen et al. (1995) Order Batrachoidiformes Goodrich, 1909 Family Batrachoididae Jordan and Everman, 1898 Batrachoides boulengeri (Gilbert and Starks, 1904) Chávez (1979) Batrachoides walkeri Collette and Russo, 1981 A Porichthys ephippiatus Walker and Rosenblatt, 1988 Allen et al. (1995) Porichthys greenei Gilbert and Starks, 1904 Allen et al. (1995) Porichthys margaritatus (Richardson, 1844) Tapia-García et al. (1995) Allen et al. (1995) Order Beloniformes Berg, 1940 Family Belonidae Gill, 1832 Ablennes hians (Valenciennes, 1846) Allen et al. (1995) Platybelone argalus (Lesueur, 1821) Allen et al. (1995) Strongylura exilis (Girard, 1854) A Allen et al. (1995) Tylosurus acus pacificus (Steindachner, 1876) Allen et al. (1995) Tylosurus crocodiles fodiator Jordan and Gilbert, 1882 Allen et al. (1995) Family Exocoetidae Risso, 1827 Cheilopogon xenopterus (Gilbert, 1890) Allen et al. (1995) Cypselurus callopterus (Günther, 1866) Allen et al. (1995) Exocoetus volitans Linnaeus, 1758 Allen et al. (1995) Fodiator rostratus (Günther, 1866) Allen et al. (1995) Exocoetus monocirrhus Richardson, 1846 Allen et al. (1995) Hirundichthys marginatus (Nichols and Breder, 1928) Allen et al. (1995) Hirundichthys speculiger Valenciennes, 1846 Allen et al. (1995) Prognichthys tringa Breder, 1928 Allen et al. (1995) Family Hemirhamphidae Gill, 1859 Euleptorhamphus viridis (Van Hasselt, 1823) Allen et al. (1995) Hemiramphus saltator Gilbert and Starks, 1904 Allen et al. (1995) Hyporhamphus gilli Meek and Hildebrand, 1923 Allen et al. (1995) Hyporhamphus rosae (Jordan and Gilbert, 1880) A Allen et al. (1995) Hyporhamphus snyderi Meek and Hildebrand, 1923 Allen et al. (1995) Hyporhamphus naos Banford and Collette, 2001 Robertson and Allen (2008) Hyporhamphus unifasciatus (Ranzani, 1842) Allen et al. (1995) Oxyporhamphus micropterus micropterus (Valenciennes, 1847) Allen et al. (1995) Order Beryciformes Regan, 1909 Family Holocentridae Richardson, 1846 Myripristis leiognathus Valenciennes, 1846 Allen et al. (1995) Sargocentron suborbitalis (Gill, 1863) Allen et al. (1995) Order Clupeiformes Bleeker, 1959 Family Engraulidae Ridewood, 1905 Anchoa analis (Miller, 1945) Allen et al. (1995) Anchoa argentivittata (Regan, 1904) Allen et al. (1995) Anchoa curta (Jordan and Gilbert, 1882) A Anchoa exigua (Jordan and Gilbert, 1882) Anchoa ischana (Jordan and Gilbert, 1882) Anchoa lucida (Jordan and Gilbert, 1882) Chávez (1979) Allen et al. (1995) Chávez (1979) Chávez (1979) Anchoa mundeola (Gilbert and Pierson, 1898) Allen et al. (1995) Anchoa nasus (Kner and Steindachner, 1867) Tapia-García et al. (1995) Anchoa scofieldi (Jordan and Culver, 1895) Chávez (1979) Cetengraulis mysticetus (Günther, 1867) Tapia-García et al. (1995) Harengula thrissina (Jordan and Gilbert, 1882) Tapia-García et al. (1995) Anchoa walkeri Baldwinm and Chang, 1970 A Anchovia macrolepidota (Kner and Steindachner, 1864) Family Clupeidae Cuvier, 1817 Etrumeus teres (DeKay, 1842) Tapia-García et al. (1995) Chávez (1979) Castro-Aguirre et al. (1999) 371 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 13. Continued. Lile gracilis Castro-Aguirre and Vivero, 1990 Robertson and Allen (2008) Neoopisthopterus tropicus (Hildebrand, 1946) Robertson and Allen (2008) Lile nigrofasciata Castro-Aguirre, Ruiz-Campos and Balart, 2002 Lile stolifera (Jordan and Gilbert, 1882) Opisthonema bulleri (Regan, 1904) Opisthonema libertate (Günther, 1866) Opisthonema medirastre Berry and Barrett, 1963 Opisthopterus dovii (Günther, 1868) Allen et al. (1995) Chávez (1979) Chávez (1979) Tapia-García et al. (1995) Tapia-García et al. (1995) Tapia-García et al. (1995) Pliosteostoma lutipinnis (Jordan and Gilbert, 1881) Tapia-García et al. (1995) Sardinops caeruleus (Girard, 1854) Allen et al. (1995) Sardinops sagax (Jenyns, 1842) Family Pristigasteridae Myers, 1960 Ilisha fuerthii (Steindachner, 1875) Order Cyprinodontiformes Berg, 1940 Family Anablepidae Garman, 1895 Anableps dowei Gill, 1861 A Family Poeciliidae Swainson, 1839 Poecilia sphenops Valenciennes, 1846 A Poeciliopsis fasciata (Meek, 1904) A Poeciliopsis turrubarensis (Meek, 1912) A Order Elopiformes Greenwood, Rosen, Weitzman and Myers, 1966 Family Elopidae Whitehead, 1962 Elops affinis Regan, 1909 Order Gadiformes Goodrich, 1909 Family Bregmacerotidae Gill, 1872 Bregmaceros bathymaster Jordan and Bollman, 1890 Castro-Aguirre et al. (1999) Tapia-García et al.1995 Chávez (1979) Chávez (1979) Castro-Aguirre (1982) Allen et al. (1995) Castro-Aguirre and Espinoza-Pérez (1996) Robertson and Allen (2008) Family Moridae Moreau, 1881 Laemonema verecundum (Jordan and Cramer 1897) Allen et al. (1995) Physiculus nematopus Gilbert, 1890 Allen et al. (1995) Physiculus talarae Hildebrand and Barton, 1949 Allen et al. (1995) Family Merlucciidae Adams, 1864 Merluccius angustimanus Garman, 1899 Allen et al. (1995) Order Gonorynchiformes Berg, 1940 Family Chanidae Jordan, 1887 Chanos chanos (Forsskäl, 1775) A Order Lampriformes (Berg, 1940) Family Regalecidae Clausen, 1959 Regalecus glesne Ascanius, 1772 Chávez (1979) Allen et al. (1995) Order Lophiiformes Garman, 1899 Family Antennariidae Gill, 1863 Antennarius avalonis Jordan and Starks, 1907 Allen et al. (1995) Antennarius sanguineus Gill, 1863 Allen et al. (1995) Lophiodes caulinaris (Garman, 1899) Tapia-García et al. (1995) Lophiodes spilurus (Garman, 1899) Allen et al. (1995) Antennarius strigatus Gill, 1863 Allen et al. (1995) Family Lophiidae Moreau, 1881 Family Ogcocephalidae Jordan, 1895 Zalieutes elater (Jordan and Gilbert, 1882) Order Ophidiiformes Berg, 1937 Family Carapidae Jordan and Fowler, 1902 Tapia-García et al. (1995) Echiodon exsilium Rosenblatt, 1961 Robertson and Allen (2008) Encheliophis dubius (Putnam, 1874) Castro-Aguirre et al. 1999 Encheliophis vermicularis Müller, 1842 Family Ophidiidae Rafinesque 1810 Robertson and Allen (2008) Brotula clarkae Hubbs, 1944 Tapia-García et al. (1995) Cherublemma emmelas (Gilbert, 1890) Allen et al. (1995) Lepophidium pardale (Gilbert, 1890) Tapia-García et al. (1995) Lepophidium prorates (Jordan and Bollman, 1890) Allen et al. (1995) Neobythites stelliferoides Gilbert, 1890 Robertson and Allen (2008) Ophidion imitator Lea, 1997 Robertson and Allen (2008) Ophidion galeoides Gilbert, 1890 Allen et al. (1995) Otophidium indefatigable Jordan and Bollman, 1889 Allen et al. (1995) Order Perciformes Bleeker, 1859 372 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 13. Continued. Family Acanthuridae Bleeker, 1859 Acanthurus nigricans (Linnaeus, 1758) Allen et al. (1995) Acanthurus triostegus (Linneeus, 1758) Allen et al. (1995) Acanthurus xanthopterus Valenciennes, 1835 Prionurus laticlavius (Valenciennes, 1846) Allen et al. (1995) Prionurus punctatus Gill, 1862 Allen et al. (1995) Allen et al. (1995) Family Ammodytidae Lee, 1989 Ammodytoides gilli (Bean, 1895) Robertson and Allen (2008) Family Apogonidae Jordan and Gilbert, 1882 Apogon pacificus (Herre, 1935) Apogon retrosella (Gill, 1862) Family Blenniidae Rafinesque 1810 Robertson and Allen (2008) Robertson and Allen (2008) Ophioblennius steindachneri Jordan and Evermann, 1898 Robertson and Allen (2008) Chaenopsis coheni Böhlke, 1957 Allen et al. (1995) Plagiotremus azaleus (Jordan and Bollman, 1890) Family Chaenopsidae Bleeker, 1859 Family Callionymidae Bleeker, 1859 Synchiropus atrilabiatus (Garman, 1899) Family Carangidae Rafinesque, 1815 Robertson and Allen (2008) Robertson and Allen (2008) Alectis ciliaris (Bloch, 1788) Tapia-García et al. (1995) Caranx caballus Günther, 1868 Tapia-García et al. (1995) Caranx lugubris Poey, 1860 Allen et al. (1995) Caranx melampygus Cuvier, 1833 Allen et al. (1995) Caranx sexfasciatus Quoy and Gaimard, 1824 Allen et al. (1995) Caranx caninus Günther, 1867 Allen et al. (1995) Caranx otrynter Jordan and Gilbert, 1883 Tapia-García et al. (1995) Caranx speciosus Forsskål, 1775 Allen et al. (1995) Caranx vinctus (Jordan and Gilbert, 1882) Chávez (1979) Chloroscombrus orqueta Jordan and Gilbert, 1882 Tapia-García et al. (1995) Decapterus macrosoma Bleeker, 1851 Allen et al. (1995) Decapterus muroadsi (Temmink and Schlegel, 1844) Allen et al. (1995) Elegatis bipinnulata (Quoy and Gaimard, 1825) Allen et al. (1995) Hemicaranx leucurus (Günther, 1864) Chávez (1979) Oligoplites altus (Günther, 1868) Chávez (1979) Selar crumenophthalmus (Bloch, 1793) Tapia-García et al. (1995) Hemicaranx zelotes Gilbert, 1898 Naucrates ductor (Linnaeus, 1758) Oligoplites refulgens Gilbert and Starks, 1904 Oligoplites saurus (Bloch and Schneider, 1801) Tapia-García et al. (1995) Allen et al. (1995) Allen et al. (1995) Chávez (1979) Selene brevoortii (Gill, 1863) Tapia-García et al. (1995) Selene peruviana (Guichenot, 1866) Tapia-García et al. (1995) Selene orstedii Lütken, 1880 Tapia-García et al. (1995) Seriola peruana Steindachner, 1876 Allen et al. (1995) Seriola rivoliana Valenciennes, 1833 Allen et al. (1995) Trachinotus rhodopus Gill, 1863 Allen et al. (1995) Trachinotus kennedyi Steindachner, 1875 Allen et al. (1995) Trachinotus paitensis Cuvier, 1832 Allen et al. (1995) Family Centropomidae Poey, 1868 Centropomus armatus Gill, 1863 Chávez (1979) Centropomus medius Gunther, 1864 Allen et al. (1995) Centropomus nigrescens Günther, 1864 A Allen et al. (1995) Centropomus robalito Jordan and Gilbert, 1882 Centropomus viridis Lockington, 1877 Family Chaetodontidae Bonaparte, 1832 Chávez (1979) Allen et al. (1995) Chaetodon humeralis Günther, 1860 Tapia-García et al. (1995) Family Cirrhitidae Rafinesque, 1810 Cirrhitus oxycephalus (Bleeker, 1855) Allen et al. (1995) Cirrhitus rivulatus Valenciennes, 1846 Allen et al. (1995) Oxycirrhites typus Bleeker, 1857 Allen et al. (1995) Johnrandallia nigrirostris (Gill, 1862) Family Coryphaenidae Rafinesque, 1810 Allen et al. (1995) Coryphaena equiselis Linnaeus, 1758 Allen et al. (1995) Coryphaena hippurus Linnaeus, 1758 Allen et al. (1995) 373 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 13. Continued. Family Dactyloscopidae Bleeker, 1859 Dactyloscopus amnis Miller and Briggs, 1962 Dactylagnus mundus Gill, 1863 Robertson and Allen (2008) Dactyloscopus byersi Dawson, 1969 Robertson and Allen (2008) Dactyloscopus fimbriatus (Reid, 1935) Dactyloscopus lunaticus Gilbert, 1890 Robertson and Allen (2008) Dactyloscopus pectoralis Gill, 1861 Dactyloscopus minutus Dawson, 1975 Robertson and Allen (2008) Gillellus searcheri Dawson, 1977 Robertson and Allen (2008) Robertson and Allen (2008) Robertson and Allen (2008) Robertson and Allen (2008) Gillellus semicinctus Gilbert, 1890 Robertson and Allen (2008) Heteristius cinctus (Osburn and Nichols, 1916) Robertson and Allen (2008) Family Echeneidae Gill, 1972 Echeneis naucrates Linnaeus, 1758 Allen et al. (1995) Phtheirichthys lineatus (Menzies, 1791 Allen et al. (1995) Remora australis Bennett, 1840 Allen et al. (1995) Remora brachyptera (Lowe, 1839) Allen et al. (1995) Remora osteochir Cuvier, 1829 Remora remora (Linnaeus, 1758) Allen et al. (1995) Remorina albescens (Temminck and Schlegel, 1850) Allen et al. (1995) Family Eleotridae Bleeker, 1859 Allen et al. (1995) Dormitator latifrons (Richardson, 1845) A Allen et al. (1995) Eleotris picta Kner, 1863 A Allen et al. (1995) Gobiomorus polylepis Ginsburg, 1953 A Allen et al. (1995) Chaetodipterus zonatus (Girard, 1858) Tapia-García et al. (1995) Parapsettus panamensis (Steindachner, 1876) Allen et al. (1995) Erotelis armiger (Jordan and Richardson In: Jordan, 1895) Allen et al. (1995) Gobiomorus maculatus (Günther, 1859) A Tapia-García et al. (1998) Family Ephippidae Parenti, 1981 Family Gempylidae Lee, 1989 Gempylus serpens Cuvier, 1829 Allen et al. (1995) Lepidocybium flavobrunneum (Smith 1843) Allen et al. (1995) Diapterus aureolus (Jordan and Gilbert, 1882) Tapia-García et al. (1995) Diapterus peruvianus (Cuvier, 1830) Eucinostomus argenteus Baird and Girard, 1855 A Allen et al. (1995) Eucinostomus currani Zahuranec, 1980 A Tapia-García et al. (1995) Nealotus tripes Johnson, 1865 Allen et al. (1995) Family Gerreidae Gill, 1972 Eucinostomus dowii (Gill, 1863) Eucinostomus entomelas Zahuranec, 1980 Castro-Aguirre (1982) Tapia-García et al. (1995) Chávez (1979) Eucinostomus gracilis (Gill, 1862) A Tapia-García et al. (1995) Eugerres axillaris (Günther, 1864) Allen et al. (1995) Eugerres brevimanus (Günther, 1864) Allen et al. (1995) Eugerres lineatus (Humbolt and Valenciennes, 1821) Allen et al. (1995) Gerres cinerus (Walbaum, 1792) A Chávez (1979) Family Gobiidae Fleming, 1822 Aboma etheostoma Jordan and Starks, 1895 Robertson and Allen (2008) Awaous trasandeanus (Gunther, 1861) A Allen et al. (1995) Bathygobius ramosus Ginsburg, 1947 Allen et al. (1995) Bollmannia ocellata Gilbert, 1892 Allen et al. (1995) Bollmannia stigmatura Gilbert, 1892 Allen et al. (1995) Ctenogobius manglicola (Jordan and Starks In: Jordan, 1895) Allen et al. (1995) Ctenogobius sagittula (Günter, 1862) Allen et al. (1995) Evermannia zosterura (Jordan and Gilbert, 1882) Chávez (1979) Elacatinus puncticulatus (Ginsburg, 1938) Allen et al. (1995) Enypnias seminudus (Günther, 1861) Allen et al. (1995) Gobionellus microdon (Gilbert, 1892) A Chávez (1979) Evorthodus minutus Meek and Hildebrand, 1928 Gobiosoma nudum (Meek and Hildebrand, 1928) Gobiosoma paradoxum (Günther, 1861) Allen et al. (1995) Allen et al. (1995) Allen et al. (1995) Microgobius brevispinnis Ginsburg, 1939 Allen et al. (1995) Microgobius emblematicus (Jordan and Gilbert, 1882) Allen et al. (1995) Microgobius erectus Ginsburg, 1938 Allen et al. (1995) Microgobius cyclolepis Gilbert, 1890 Allen et al. (1995) 374 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 13. Continued. Microgobius miraflorensis Gilbert and Starks, 1904 Microgobius tabogensis Meek and Hildebrand, 1928 Parrella maxillaris Ginsburg, 1938 Chávez (1979) Allen et al. (1995) Allen et al. (1995) Family Haemulidae Richarson, 1848 Anisotremus caesius (Jordan and Gilbert, 1882) Allen et al. (1995) Anisotremus dovii (Günther, 1864) Anisotremus interruptus (Gill, 1862) Allen et al. (1995) Anisotremus pacifici (Günther, 1864) Chávez (1979) Anisotremus taeniatus Gill, 1861 Allen et al. (1995) Conodon serrifer Jordan and Gilbert, 1882 Allen et al. (1995) Haemulon flaviguttatum Gill, 1862 Allen et al. (1995) Haemulon maculicauda (Gill, 1862) Allen et al. (1995) Haemulon scudderi Gill, 1862 Allen et al. (1995) Haemulon sexfasciatum Gill, 1862 Allen et al. (1995) Allen et al. (1995) Haemulon steindachneri (Jordan and Gilbert, 1882) Allen et al. (1995) Haemulopsis axillaris ((Steindachner, 1869) Tapia-García et al. (1995) Haemulopsis elongatus (Jordan and Gilbert, 1879) Tapia-García et al. (1995) Haemulopsis leuciscus (Günther, 1864) Tapia-García et al. (1995) Microlepidotus brevipinnis (Steindachner, 1869) Allen et al. (1995) Orthopristis cantharinus (Jenyns, 1840) Allen et al. (1995) Orthopristis chalceus (Günther, 1864) Tapia-García et al. (1995) Pomadasys branickii (Steindachner, 1879) Allen et al. (1995) Pomadasys macracanthus (Günther, 1864) Chávez (1979) Xenistius californiensis (Steindachner, 1875) Allen et al. (1995) Haemulopsis nitidus (Steindachner, 1869) Tapia-García et al. (1995) Pomadasys bayanus (Jordan and Evermann, 1898) A Allen et al. (1995) Pomadasys panamensis (Steindachner, 1875) Tapia-García et al. (1995) Xenichthys xanti Gill, 1863 Family Istiophoridae Lee, 1989 Tapia-García et al. (1995) Istiophorus platypterus (Shaw, 1792) Allen et al. (1995) Makaira indica (Cuvier In: Cuvier and Valenciennes, 1832) Allen et al. (1995) Makaira mazara (Jordan and Snyder, 1901) Allen et al. (1995) Tetrapturus angustirostris Tanaka, 1915 Allen et al. (1995) Tetrapturus audax (Philippi, 1887) Allen et al. (1995) Family Kuhliidae Gill, 1972 Kuhlia mugil (Forster In: Bloch and Schneider, 1801) Robertson and Allen (2008) Family Kyphosidae Gill, 1972 Kyphosus analogus (Gill, 1862) Allen et al. (1995) Kyphosus elegans (Peters, 1869) Allen et al. (1995) Sectator ocyurus (Jordan and Gilbert, 1882) Allen et al. (1995) Family Labridae Cuvier, 1816 Bodianus diplotaenia (Gill, 1862) Allen et al. (1995) Decodon melasma Gomon, 1974 Allen et al. (1995) Halichoeres aestuaricola Bussing, 1972 Allen et al. (1995) Halichoeres chierchiae Di Caporiacco, 1947 Halichoeres dispilus (Günther, 1864) Allen et al. (1995) Halichoeres melanotis (Gilbert, 1890) Allen et al. (1995) Halichoeres nicholsi (Jordan and Gilbert, 1882) Allen et al. (1995) Halichoeres notospilus (Günther, 1864) Allen et al. (1995) Iniistius pavo (Valenciennes in: Cuvier and Valenciennes, 1840) Novaculichthys taeniourus (Lacépède, 180l) Allen et al. (1995) Ramírez-Gutiérrez et al. (2007) Allen et al. (1995) Thalassoma grammaticum Gilbert, 1890 Allen et al. (1995) Thalassoma lucasanum (Gill, 1862) Allen et al. (1995) Family Labrisomidae Bleeker, 1859 Paraclinus mexicanus (Gilbert, 1904) Robertson and Allen (2008) Family Lobotidae Springer and Raasch, (1995) Lobotes pacificus Gilbert 1898 Allen et al. (1995) Lobotes surinamensis (Bloch, 1790) Chávez (1979) Family Lutjanidae Springer and Raasch, (1995) Hoplopagrus guentherii Gill, 1862 Allen et al. (1995) Lutjanus aratus (Günther, 1864) Allen et al. (1995) Lutjanus argentiventris (Peters, 1869) Allen et al. (1995) Lutjanus colorado Jordan and Gilbert, 1882 Tapia-García et al. (1995) 375 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 13. Continued. Lutjanus guttatus (Steindachner, 1869) Tapia-García et al. (1995) Lutjanus inermis (Peters, 1869) Allen et al. (1995) Lutjanus jordani (Gilbert, 1897) Allen et al. (1995) Lutjanus novemfasciatus Gill, 1862 Allen et al. (1995) Lutjanus peru (Nichols and Murphy, 1922) Tapia-García et al. (1995) Lutjanus viridis (Valenciennes, 1846) Allen et al. (1995) Family Luvaridae Parenti, 1981 Luvarus imperialis Rafinesque, 1810 Allen et al. (1995) Caulolatilus affinis Gill, 1865 Allen et al. (1995) Caulolatilus hubbsi Dooley, 1978 Allen et al. (1995) Family Malacanthidae Springer and Raasch, (1995) Caulolatilus princeps (Jenyns, 1840) Allen et al. (1995) Family Microdesmidae Bleeker, 1859 Microdesmus affinis Meek and Hildebrand, 1928 Robertson and Allen (2008) Microdesmus dipus Günther, 1864 Castro-Aguirre et al. (1999) Microdesmus dorsipunctatus Dawson, 1968 Family Mugilidae Bonaparte, 1831 Agonostomus monticola (Bancroft, 1834) C Castro-Aguirre et al. (1999) Allen et al. (1995) Chaenomugil proboscideus (Günther, 1861) Allen et al. (1995) Mugil curema Valenciennes, 1836 C Chávez (1979) Joturus pichardi Poey, 1860 C Mugil cephalus Linnaeus, 1758 C Allen et al. (1995) Chávez (1979) Mugil hospes Jordan and Culver, 1895 Allen et al. (1995) Mulloidichthys dentatus (Gill, 1862) Tapia-García et al. (1995) Pseudupeneus grandisquamis (Gill, 1863) Tapia-García et al. (1995) Mugil setosus Gilbert, 1892 Family Mullidae Günther, 1859 Chávez (1979) Family Nematistiidae Springer and Raasch, (1995) Nemastistius pectoralis Gill, 1862 Family Nomeidae Lee, 1989 Castro-Aguirre et al. (1999) Nomeus gronovii (Gmelin, 1789) Allen et al. (1995) Psenes sio Haedrich, 1970 Allen et al. (1995) Family Opistognathidae Springer and Raasch, (1995) Lonchopisthus sinuscalifornicus (Castro-Aguirre and Villavicencio-Garayzar, 1988) Opistognathus punctatus Peters, 1869 Robertson and Allen (2008) Opistognathus scops (Jenkins and Evermann, 1889) Robertson and Allen (2008) Family Polynemidae Springer and Raasch, (1995) Polydactylus approximans (Lay and Bennett, 1839) Chávez (1979) Stereolepis gigas Ayres, 1859 Allen et al. (1995) Polydactylus opercularis (Gill, 1863) Family Polyprionidae Springer and Raasch, (1995) Family Pomacanthidae Fraser-Brunner, 1933 Tapia-García et al. (1995) Holacanthus passer Valenciennes, 1846 Allen et al. (1995) Pomacanthus zonipectus (Gill, 1862) Allen et al. (1995) Family Pomacentridae (Cuvier, 1816) Allen et al. (1995) Abudefduf concolor (Gill, 1862) Abudefduf declivifrons (Gill, 1862) Robertson and Allen (2008) Abudefduf troschelii (Gill, 1862) Allen et al. (1995) Chromis alta Greenfield and Woods, 1980 Allen et al. (1995) Chromis atrilobata Gill, 1862 Allen et al. (1995) Microspathodon bairdii (Gill, 1862) Allen et al. (1995) Microspathodon dorsalis (Gill, 1862) Allen et al. (1995) Stegastes acapulcoensis (Fowler, 1944) Allen et al. (1995) Stegastes flavilatus (Gill, 1862) Allen et al. (1995) Stegastes rectifraenum (Gill, 1862) Allen et al. (1995) Family Priacanthidae Springer and Raasch, (1995) Cookeolus japonicus (Cuvier In: Cuvier and Valenciennes, 1829) Allen et al. (1995) Heteropriacanthus cruentatus (Lacepède, 1801) Tapia-García et al. (1995) Ptereleotris carinata Bussing, 2001 Robertson and Allen (2008) Pristigenys serrula (Gilbert, 1891) Allen et al. (1995) Family Ptereleotridae Thacker, 2000 Family Scaridae Rafinesque, 1810 Nicholsina denticulata (Evermann and Radcliffe, 1917) Allen et al. (1995) 376 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 13. Continued. Scarus compressus (Osburn and Nichols, 1916) Allen et al. (1995) Scarus ghobban Forsskål, 1775 Allen et al. (1995) Scarus perrico Jordan and Gilbert, 1882 Allen et al. (1995) Scarus rubroviolaceus Bleeker, 1847 Allen et al. (1995) Family Sciaenidae Regan, 1913 Bairdiella armata Gill, 1863 Allen et al. (1995) Bairdiella ensifera (Jordan and Gilbert, 1882) Allen et al. (1995) Bairdiella icistia (Jordan and Gilbert, 1882) Allen et al. (1995) Corvula macrops (Steindachner, 1876) Allen et al. (1995) Cynoscion albus (Günther, 1864) Allen et al. (1995) Cynoscion phoxocephalus Jordan and Gilbert, 1882 Tapia-García et al. (1995) Cynoscion reticulatus (Günther, 1864) Tapia-García et al. (1995) Cynoscion nannus Castro-Aguirre and Arvizu-Martínez, 1976 Allen et al. (1995) Cynoscion squamipinnis (Günther, 1867) Allen et al. (1995) Elattarchus archidium (Jordan and Gilbert, 1882) Tapia-García et al. (1995) Isopisthus remifer Jordan and Gilbert, 1882 Tapia-García et al. (1995) Larimus acclivis Jordan and Brisol, 1898 Tapia-García et al. (1995) Larimus argenteus (Gill, 1863) Tapia-García et al. (1995) Cynoscion stolzmanni (Steindachner, 1879) Allen et al. (1995) Larimus effulgens Gilbert, 1898 Tapia-García et al. (1995) Larimus pacificus Jordan and Bollman, 1890 Tapia-García et al. (1995) Menticirrhus elongatus (Günther, 1864) Tapia-García et al. (1995) Menticirrhus nasus (Günther, 1868) Tapia-García et al. (1995) Menticirrhus panamensis (Steindachner, 1877) Tapia-García et al. (1995) Menticirrhus undulatus (Girard, 1854) Allen et al. (1995) Micropogonias altipinnis (Günther, 1864) Tapia-García et al. (1995) Menticirrhus paitensis Hildebrand, 1946 Allen et al. (1995) Micropogonias ectenes Jordan and Gilbert, 1882 Allen et al. (1995) Micropogonias megalops (Gilbert, 1890) Allen et al. (1995) Odontoscion xanthops Gilbert, 1898 Allen et al. (1995) Ophioscion imiceps (Jordan and Gilbert, 1882) Allen et al. (1995) Ophioscion scierus (Jordan and Gilbert, 1884) Allen et al. (1995) Ophioscion strabo Gilbert, 1897 Allen et al. (1995) Paralonchurus goodei Gilbert In: Jordan and Evermann, 1898 Paralonchurus rathbuni (Jordan and Bollman, 1890) Allen et al. (1995) Pareques fuscovittatus (Kendall and Radcliffe, 1912) Allen et al. (1995) Stellifer chrysoleuca (Günther, 1867) Allen et al. (1995) Tapia-García et al. (1995) Stellifer ericymba (Jordan and Gilbert, 1882) Allen et al. (1995) Stellifer illecebrosus Gilbert In: Jordan and Evermann, 1898 Tapia-García et al. (1995) Umbrina bussingi López, 1980 Allen et al. (1995) Umbrina analis Günther, 1868 Allen et al. (1995) Umbrina dorsalis Gill, 1862 Allen et al. (1995) Umbrina xanti Gill, 1862 Tapia-García et al. (1995) Family Scombridae Rafinesque, 1815 Acanthocybium solandri (Cuvier, 1832) Allen et al. (1995) Euthynnus lineatus Kishinouye, 1920 Allen et al. (1995) Katsuwonus pelamis (Linnaeus, 1758) Allen et al. (1995) Auxis rochei eudorax Collette and Aadland, 1996 Allen et al. (1995) Auxis thazard brachydorax Collette and Aadland, 1996 Allen et al. (1995) Sarda orientalis (Temminck and Schlegel, 1844) Allen et al. (1995) Scomberomorus sierra Jordan and Starks, 1895 Tapia-García et al. (1995) Thunnus albacares (Bonnaterre, 1788) Allen et al. (1995) Thunnus obesus (Lowe, 1839) Allen et al. (1995) Alphestes multiguttatus (Günther, 1867) Allen et al. (1995) Family Serranidae Swainson, 1839 Alphestes immaculatus Breder, 1936 Allen et al. (1995) Cephalopholis panamensis (Steindachner, 1877) Allen et al. (1995) Dermatolepis dermatolepis (Boulenger, 1895) Allen et al. (1995) Diplectrum eumelum Rosenblatt and Johnson, 1974 Allen et al. (1995) Diplectrum euryplectrum Jordan and Bollmann, 1890 Tapia-García et al. (1995) Diplectrum labarum Rosenblatt and Johnson, 1974 Tapia-García et al. (1995) Diplectrum macropoma (Günther, 1864) Tapia-García et al. (1995) Diplectrum maximum Hildebrand, 1946 Allen et al. (1995) 377 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 13. Continued. Diplectrum pacificum Meek and Hildebrand, 1925 Tapia-García et al. (1995) Diplectrum rostrum Bortone, 1974 Allen et al. (1995) Epinephelus acanthistius (Gilbert, 1892) Allen et al. (1995) Epinephelus analogus Gill, 1864 Tapia-García et al. (1995) Epinephelus exsul (Fowler, 1944) Allen et al. (1995) Epinephelus itajara (Lichtenstein, 1822) Allen et al. (1995) Epinephelus labriformis (Jenyns, 1840) Allen et al. (1995) Hemanthias signifer (Garman, 1899) Allen et al. (1995) Hemanthias peruanus (Steindachner, 1874) Allen et al. (1995) Hyporthodus niphobles (Gilbert and Starks, 1897) Allen et al. (1995) Mycteroperca xenarcha Jordan, 1888 Allen et al. (1995) Paralabrax loro Walford, 1936 Allen et al. (1995) Paralabrax nebulifer (Girard, 1854) Allen et al. (1995) Paranthias colonus (Valenciennes, 1846) Allen et al. (1995) Pronotogrammus eos Gilbert, 1890 Allen et al. (1995) Pronotogrammus multifasciatus Gill, 1863 Allen et al. (1995) Rypticus nigripinnis Gill, 1861 Tapia-García et al. (1995) Serranus aequidens (Gilbert, 1890) Allen et al. (1995) Serranus huascarii Boulenger, 1895 Allen et al. (1995) Serranus psittacinus Valenciennes, 1846 Allen et al. (1995) Rypticus bicolor Valenciennes, 1846 Family Sparidae Bonaparte, 1832 Calamus brachysomus (Lockington, 1880) Allen et al. (1995) Allen et al. (1995) Family Sphyraenidae Röse, 1793 Tapia-García et al. (1995) Peprilus medius (Peters, 1869) Tapia-García et al. (1995) Peprilus snyderi Gilbert and Starks, 1904 Tapia-García et al. (1995) Sphyraena ensis Jordan and Gilbert, 1882 Family Stromateidae Lee, 1989 Family Trichiuridae Rafinesque, 1810 Trichiurus lepturus Linnaeus, 1758 A Family Uranoscopidae Regan, 1913 Allen et al. (1995) Astroscopus zephyreus Gilbert and Starks In: Gilbert, 1897 Allen et al. (1995) Kathetostoma averruncus Jordan and Bollmann, 1890 Allen et al. (1995) Family Xiphiidae Regan, 1909 Xiphias gladius Linnaeus, 1758 Allen et al. (1995) Family Zanclidae Parenti, 1981 Zanclus cornutus (Linnaeus, 1758) Robertson and Allen (2008) Order Pleuronectiformes Bleeker, 1859 Family Achiridae Swainson, 1839 Achirus mazatlanus (Steindachner, 1869) A Tapia-García et al. (1995) Achirusscutum (Günther, 1862) Tapia-García et al. (1995) Achirus zebrinus Clark, 1936 Tapia-García et al. (1995) Trinectes fonsecensis (Günther, 1862) Tapia-García et al. (1995) Bothus constellatus (Jordan and Goss, 1889) Tapia-García et al. (1995) Bothus leopardinus (Günther, 1862) Allen et al. (1995) Engyphrys sanctilaurentia Jordan and Bollman, 1890 Tapia-García et al. (1995) Monolene asaedai Clark, 1936 Perissias taeniopterus (Gilbert, 1890) Allen et al. (1995) Symphurus atramentatus (Jordan and Bollman, 1890) Tapia-García et al. (1995) Symphurus callopterus Munroe and Mahadeva, 1989 Allen et al. (1995) Family Bothidae Norman, 1934 Family Cynoglossidae Swainson, 1839 Tapia-García et al. (1995) Symphurus chabanaudi Mahadeva and Munroe, 1990 Allen et al. (1995) Symphurus fasciolaris Gilbert, 1892 Allen et al. (1995) Symphurus gorgonae Chabanaud, 1948 Allen et al. (1995) Symphurus elongatus (Günther, 1868) Tapia-García et al. (1995) Symphurus leei Jordan and Bollman, 1890 Allen et al. (1995) Symphurus melanurus Clark, 1936 Allen et al. (1995) Symphurus prolatinaris Munroe, Nizinski and Mahadeva, 1991 Allen et al. (1995) Symphurus melasmatotheca Munroe and Nizinski, 1990 Allen et al. (1995) Symphurus oligomerus Mahadeva and Munroe, 1990 Allen et al. (1995) Symphurus undecimplerus Munroe and Nizinski, 1990 Allen et al. (1995) Symphurus williamsi Jordan and Culver In: Jordan, 1895 Allen et al. (1995) 378 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 13. Continued. Family Paralichthyidae Swainson, 1839 Ancylopsetta dendritica Gilbert, 1890 Tapia-García et al. (1995) Citharichthys gilberti Jenkins and Evermann, 1889 A Tapia-García et al. (1995) Citharichthys mariajorisae van der Heiden and Mussot-Pérez, (1995) Tapia-García et al. (1995) Citharichthys stigmaeus Jordan and Gilbert, 1882 Allen et al. (1995) Citharichthys xanthostigma Gilbert, 1890 Allen et al. (1995) Citharichthys platophrys Gilbert, 1891 Robertson and Allen (2008) Cyclopsetta panamensis (Steindachner, 1875) Tapia-García et al. (1995) Cyclopsetta querna (Jordan and Bollman, 1890) Tapia-García et al. (1995) Etropus crossotus Jordan and Gilbert, 1882 Tapia-García et al. (1995) Etropus peruvianus Hildebrand, 1946 Allen et al. (1995) Hippoglossina bollmani Gilbert, 1890 Allen et al. (1995) Hippoglossina tetrophthalma (Gilbert, 1890) Allen et al. (1995) Paralichthys woolmani Jordan and Williams, 1897 Tapia-García et al. (1995) Syacium latifrons Jordan and Gilbert, 1882 Tapia-García et al. (1995) Syacium ovale (Günther, 1864) Tapia-García et al. (1995) Order Scorpaeniformes Garman, 1899 Family Scorpaenidae Risso, 1827 Pontinus furcirhinus Garman, 1899 Allen et al. (1995) Pontinus sierra (Gilbert, 1890) Tapia-García et al. (1995) Scorpaena histrio Jenyns, 1840 Allen et al. (1995) Scorpaena plumieri mystes Jordan and Starks, 1895 Allen et al. (1995) Scorpaena russula Jordan and Bollman, 1889 Tapia-García et al. (1995) Scorpaenodes xyris (Jordan and Gilbert, 1882) Allen et al. (1995) Family Triglidae Regan, 1913 Bellator gymnostethus (Gilbert, 1892) Allen et al. (1995) Bellator loxias (Jordan, 1897) Tapia-García et al. (1995) Bellator xenisma (Jordan and Bollman, 1890) Allen et al. (1995) Prionotus albirostris Jordan and Bollman, 1890 Tapia-García et al. (1995) Prionotus birostratus Richardson, 1844 Tapia-García et al. (1995) Prionotus horrens Richardson, 1844 Tapia-García et al. (1995) Prionotus ruscarius Gilbert and Starks, 1904 Tapia-García et al. (1995) Prionotus stephanophrys Lockington, 1881 Tapia-García et al. (1995) Order Siluriformes Cuvier, 1817 Family Ariidae Günter, 1864 Arius guatemalensis Günter, 1864 A Allen et al. (1995) Bagre pinnimaculatus Steindachner, 1876 A Allen et al. (1995) Ariusseemanni Günther, 1864 A Chávez (1979) Cathorops fuerthii (Steindachner, 1877) A Allen et al. (1995) Cathorops liropus (Bristol, 1897) Tapia-García et al. (1998) Notarius kessleri (Steindachner, 1877) Allen et al. (1995) Notarius planiceps (Steindachner, 1877) Allen et al. (1995) Occidentarius platypogon (Günther, 1864) Allen et al. (1995) Bagre panamensis Gill, 1863 Sciades dowii (Gill, 1863) A Sciades troschelii (Gill 1863) Tapia-García et al. (1995) Allen et al. (1995) Allen et al. (1995) Order Syngnathiformes Berg, 1940 Family Fistulariidae Blainville, 1818 Fistularia commersonii Rüppell, 1838 Tapia-García et al. (1995) Hippocampus ingens Girard, 1858 Tapia-García et al. (1995) Syngnathus auliscus (Swain, 1882) Castro-Aguirre et al. (1999) Fistularia corneta Gilbert and Starks, 1904 Family Syngnathidae Bonaparte, 1831 Order Tetraodontiformes Berg, 1940 Family Balistidae Regan, 1902 Allen et al. (1995) Balistes polylepis Steindachner, 1876 Tapia-García et al. (1995) Canthidermis maculatus (Bloch, 1786) Allen et al. (1995) Melichthys niger (Bloch, 1786) Allen et al. (1995) Pseudobalistes naufragium (Jordan and Starks, 1895) Allen et al. (1995) Sufllamen verres (Gilbert and Starks, 1904) Allen et al. (1995) Family Diodontidae Bibron, 1855 Diodon eydouxii Brissout de Barneville, 1846 Diodon holocanthus Linnaeus, 1758 Robertson and Allen (2008) Tapia-García et al. (1995) 379 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 13. Continued. Diodon hystrix Linneaus, 1758 Family Molidae Parr, 1933 Tapia-García et al. (1995) Mola mola (Linnaeus, 1758) Allen et al. (1995) Ranzania laevis (Pennant, 1776) Allen et al. (1995) Family Monacanthidae Parr, 1933 Aluterus scriptus (Osbeck, 1765) Tapia-García et al. (1995) Aluterus monocerus (Linnaeus, 1758) Allen et al. (1995) Cantherhines dumerilii (Hollard, 1854) Allen et al. (1995) Family Ostraciidae Rafinesque, 1815 Ostracion meleagris Shaw, 1796 Family Tetraodontidae Bonaparte, 1832 Robertson and Allen (2008) Arothron hispidus (Linnaeus, 1758) Allen et al. (1995) Arothron meleagris (Lacépède, 1798) Canthigaster punctatissima (Günther, 1870) Allen et al. (1995) Sphoeroides lobatus (Steindachner, 1870) Tapia-García et al. (1995) Allen et al. (1995) Lagocephalus lagocephalus (Linnaeus, 1758) Allen et al. (1995) Sphoeroides annulatus (Jenyns, 1842) Chávez (1979) Sphoeroides sechurae Hildebrand, 1946 Sphoeroides trichocephalus (Cope, 1870) Allen et al. (1995) Allen et al. (1995) Table 14. Checklist of amphibians and reptiles recorded in the marine and coastal areas of Oaxaca, Mexico. Phylum Chordata Bateson, 1885 Reference Class Amphibia Linnaeus, 1758 Subclass Lisssamphibia Haeckel, 1866 Order Anura (Fischer von Waldheim, 1813) Gray 1825 Family Bufonidae Gray, 1825 Incilius marmoreus (Wiegmann, 1833) Rhinella marina (Linnaeus, 1758) Family Ranidae Rafinesque, 1814 Lithobates berlandieri (Baird, 1854) Lithobates forreri (Boulenger, 1883) Class Reptilia (Linnaeus, 1758) Order Crocodylia Owen, 1842 Family Crocodylidae Cuvier, 1807 Crocodylus acutus Cuvier, 1807 Order Squamata Oppel, 1811 Suborder Lacertilia Günther, 1867 Family Corytophanidae Fitzinger, 1843 Basiliscus vittatus Wiegmann, 1828 Family Iguanidae Frost and Etheridge, 1989 Ctenosaura oaxacana Gray, 1842 Ctenosaura pectinata Wiegmann, 1834 Iguana iguana Linnaeus, 1758 Suborder Serpentes Linnaeus, 1758 Family Boidae Gray, 1825 Boa constrictor Linnaeus, 1758 Family Colubridae Oppel, 1811 Drymarchon corais Boie, 1827 Leptophis diplotropis Günter, 1872 Oxybelis aeneus Wagler, 1824 Salvadora lemniscata Cope, 1895 Senticolis triaspis Cope, 1866 Stenorrhina freminvillei Duméril, Bibron et Duméril, 1854 Symphimus leucostomous Cope, 1870 (1869) Trimorphodon biscutatus Duméril, Bibron et Duméril, 1854 Family Dipsadidae Bonaparte, 1838 Conophis vittatus Peters, 1860 Leptodeira annulata Linnaeus, 1758 Leptodeira maculata Hallowell, 1861 Leptodeira nigrofasciata Günter, 1868 Manolepis putnami Jan, 1863 Rhadinaea decorate Günter, 1858 Family Elapidae Boie, 1827 Casas-Andreu et al. 2004 Casas-Andreu et al. 2004 Casas-Andreu et al. 2004 Casas-Andreu et al. 2004 Casas-Andreu et al. 2004 Casas-Andreu et al. 2004 Casas-Andreu et al. 2004 Casas-Andreu et al. 2004 Casas-Andreu et al. 2004 Casas-Andreu et al. 2004 Casas-Andreu et al. 2004 Casas-Andreu et al. 2004 Casas-Andreu et al. 2004 Casas-Andreu et al. 2004 Casas-Andreu et al. 2004 Casas-Andreu et al. 2004 Casas-Andreu et al. 2004 Casas-Andreu et al. 2004 Casas-Andreu et al. 2004 Casas-Andreu et al. 2004 Casas-Andreu et al. 2004 Casas-Andreu et al. 2004 Casas-Andreu et al. 2004 Casas-Andreu et al. 2004 380 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 14. Continued. Micrurus browni Schmidt et Smith, 1943 Casas-Andreu et al. 2004 Family Hydrophiidae Boie, 1827 Casas-Andreu et al. 2004 Pelamis platurus Linnaeus, 1766 Family Leptotyphlopidae Stejnejer, 1892 Leptotyphlops goudotti Duméril et Bibron, 1844 Casas-Andreu et al. 2004 Family Loxocemidae Cope, 1861 Loxocemus bicolor Cope, 1861 Casas-Andreu et al. 2004 Order Testudines Linnaeus, 1758 Family Cheloniidae Bauer, 1893 Casas-Andreu et al. 2004 Chelonia agassizi (Boucourt, 1868) Casas-Andreu et al. 2004 Caretta caretta Linnaeus, 1758 Casas-Andreu et al. 2004 Eretmochelys imbricada Linnaeus, 1776 Lepidochelys olivacea (Eschscholtz, 1829) Casas-Andreu et al. 2004 Family Dermochelyidae Fitzinger, 1843 Casas-Andreu et al. 2004 Dermochelys coriacea (Vandelli, 1761) Family Emydidae Rafinesque, 1815 Casas-Andreu et al. 2004 Rhinoclemmys pulcherrima Gray, 1856 Rhinoclemmys rubida Cope, 1869 Casas-Andreu et al. 2004 Kinosternon oaxacae Berry and Iverson, 1980 Casas-Andreu et al. 2004 Family Kinosternidae Agassiz, 1857 Kinosternon integrum Le Conte, 1854 Casas-Andreu et al. 2004 Table 15. Checklist of birds recorded in the marine and coastal areas of Oaxaca, Mexico. Class Aves Linnaeus, 1758 Order Anseriformes Wagler, 1831 Reference Family Anatidae Vigors, 1825 Cairina moschata (Linnaeus, 1758 Binford (1989) Dendrocygna autumnalis (Linnaeus, 1758) Binford (1989) Dendrocygna bicolor (Vieillot, 1816) Binford (1989) Anas acuta Linnaeus, 1758 Binford (1989) Anas americana Gmelin, 1789 Binford (1989) Anas clypeata Linnaeus, 1758 Binford (1989) Anas crecca Linnaeus, 1758 Binford (1989) Anas discors Linnaeus, 1766 Binford (1989) Anas strepera Linnaeus, 1758 Binford (1989) Aythya affinis (Eyton, 1838) Binford (1989) Aythya collaris (Donovan, 1809) Oxyura jamaicensis (Gmelin, 1789) Order Procellariiformes Fürbringer, 1888 Binford (1989) Binford (1989) Family Hydrobatidae Mathews, 1912 Oceanodroma melania (Bonaparte, 1854) Binford (1989) Oceanodroma microsoma (Coues, 1864) Binford (1989) Puffinus griseus (Gmelin, 1789) Navarro-Sigüenza et al. (2004) Family Procellariidae Boie, 1826 Puffinus auricularis Townsend, 1890 Puffinus Ilherminieri Lesson, 1839 Puffinus pacificus (Gmelin, 1789) Navarro-Sigüenza et al. (2004) Binford (1989) Binford (1989) Order Phaethontiformes Sharpe, 1891 Family Phaetodontidae Brandt, 1840 Phaeton aethereus Linnaeus, 1758 Order Suliformes Reichembach, 1849 Family Fregatidae Garrod, 1874 Fregata magnificens Mathews, 1914 Navarro-Sigüenza et al. (2004) Binford (1989) Family Sulidae Reichenbach, 1849 Sula granti Rothschild, 1902 Binford (1989) Sula leucogaster (Boddaert, 1783) Binford (1989) Sula nebouxii Milne-Edwards 1882 Sula sula (Linnaeus, 1766) Family Phalacrocoracidae Reichenbach, 1850 Phalacrocorax brasilianus (Gmelin, 1789) Family Anhingidae Reichenbach, 1849 Anhinga anhinga (Linnaeus, 1766) Navarro-Sigüenza et al. (2004) Binford (1989) Navarro-Sigüenza et al. (2004) Binford (1989) Order Pelecaniformes Sharpe, 1891 381 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 15. Continued. Family Pelecanidae Rafinesque, 1815 Pelecanus occidentalis Linnaeus, 1766 Binford (1989) Family Ardeidae Leach, 1820 Ardea alba (Linnaeus, 1758) Binford (1989) Ardea herodias Linnaeus, 1758 Binford (1989) Bubulcus ibis (Linnaeus, 1758) Binford (1989) Butorides virescens (Linnaeus, 1758) Navarro-Sigüenza et al. (2004) Cochlearius cochlearius (Linnaeus, 1766) Binford (1989) Egretta caerulea (linnaeus, 1758) Binford (1989) Egretta rufescens (Gmelin, 1789) Binford (1989) Egretta thula (Molina, 1782) Binford (1989) Egretta tricolor (Müller, 1776) Binford (1989) Tigrisoma mexicanum Swainson, 1834 Binford (1989) Nyctanassa violacea (Linnaeus, 1758) Nycticorax nycticorax (Linnaeus, 1758) Navarro-Sigüenza et al. (2004) Binford (1989) Family Threskiornithidae Richmond, 1917 Eudocimus albus (Linnaeus, 1758) Binford (1989) Platalea ajaja Linnaeus, 1758 Binford (1989) Plegadis chichi (Vieillot, 1817) Binford (1989) Order Ciconiiformes Bonaparte, 1854 Family Ciconiidae Gray, 1840 Binford (1989) Family Cathartidae Lafresnaye, 1839 Cathartes aura (Linnaeus, 1758) Binford (1989) Coragyps atratus (Bechstein, 1793) Binford (1989) Mycteria americana Linnaeus, 1758 Order Accipitriformes Vieillot, 1807 Family Pandionidae Bonaparte, 1854 Pandion haliaetus (Linnaeus, 1758) Binford (1989) Family Accipitridae Vieillot, 1816 Buteo albonotatus Kaup, 1847 Binford (1989) Buteo jamaicensis (Gmelin, 1788) Binford (1989) Buteo magnirostris (Gmelin, 1788) Binford (1989) Buteo nitidus (Latham, 1790) Binford (1989) Buteo platypterus (Vieillot, 1823) Binford (1989) Buteogallus urubitinga (Gmelin, 1788) Binford (1989) Rostrhamus sociabilis (Vieillot, 1817) Binford (1989) Buteogallus anthracinus (Deppe, 1830) Binford (1989) Order Falconiformes Sharpe, 1874 Family Falconidae Vigors, 1824 Caracara cheirway (Jacquin, 1784) Binford (1989) Falco peregrinus Tunstall, 1771 Binford (1989) Falco sparverius Linnaeus, 1758 Binford (1989) Herpetotheres cachinnans (Linnaeus, 1758) Binford (1989) Order Charadriiformes Huxley, 1867 Family Charadriidae Vigors, 1825 Charadrius alexandrinus Linnaeus, 1758 Binford (1989) Charadrius collaris Vieillot, 1816 Binford (1989) Charadrius semipalmatus Bonaparte, 1825 Binford (1989) Charadrius vociferus Linnaeus, 1758 Binford (1989) Charadrius wilsonia Ord, 1814 Binford (1989) Pluvialis dominica (Müller, 1776) Binford (1989) Haematopus palliatus Temminck, 1820. Binford (1989) Pluvialis squatarola (Linnaeus, 1758) Binford (1989) Family Haematopodidae Gray, 1840 Family Jacanidae Stejneger, 1885 Jacana spinosa (Linnaeus, 1758) Binford (1989) Family Laridae Vigors, 1825 Chlidonias niger (Linnaeus, 1758) Binford (1989) Gelochelidon nilotica Gmelin, 1789 Binford (1989) Hydroprogne caspia Pallas, 1770 Binford (1989) Larus argentatus Pontoppidan, 1763 Larus californicus Lawrence, 1854 Larus delawarensis Ord, 1815 Navarro-Sigüenza et al. (2004) Navarro-Sigüenza et al. (2004) Binford (1989) 382 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 15. Continued. Leucophaeus atricilla Linnaeus, 1758 Binford (1989) Leucophaeus pipixcan Wagler, 1831 Binford (1989) Onychoprion fuscatus Linnaeus, 1766 Binford (1989) Rhynchops niger Linnaeus, 1758 Binford (1989) Stercorarius longicaudus Vieillot, 1819 Binford (1989) Stercorarius parasiticus (Linnaeus, 1758) Binford (1989) Sterna forsteri Nuttall, 1832 Binford (1989) Sternulla antillarum (Lesson, 1847) Binford (1989) Thalasseus elegans Gambel, 1849 Binford (1989) Thalasseus maximus Boddaert, 1783 Binford (1989) Thalasseus sandvicensis Latham, 1787 Binford (1989) Family Recurvirostridae Bonaparte, 1854 Himantopus mexicanus (Müller, 1776) Binford (1989) Sterna hirundo Linnaeus 1758 Recurvirostra americana Gmelin, 1789 Binford (1989) Binford (1989) Family Scolpacidae Vigors, 1825 Actitis macularius (Linnaeus, 1766) Binford (1989) Arenaria interpres (Linnaeus, 1758) Binford (1989) Calidris alba (Pallas, 1764) Binford (1989) Calidris alpina (Linnaeus, 1758) Binford (1989) Calidris canutus (Linnaeus, 1758) Binford (1989) Calidris mauri (Cabanis, 1857) Binford (1989) Calidris minutilla (Vieillot, 1819) Binford (1989) Calidris pusilla (Linnaeus, 1766) Binford (1989) Gallinago gallinago (Linnaeus, 1758) Binford (1989) Limosa fedoa (Linnaeus, 1758) Binford (1989) Limosa haemastica (linnaeus, 1758) Binford (1989) Limnodromus griseus (Gmelin, 1789) Binford (1989) Limnodromus scolopaceus (Say, 1823) Binford (1989) Numenius phaeophus (Linnaeus, 1758) Binford (1989) Phalaropus fulicarius (Linnaeus, 1758) Binford (1989) Numenius americanus Bechstein, 1812 Binford (1989) Phalaropus lobatus (Linnaeus, 1758) Binford (1989) Phalaropus tricolor (Vieillot, 1819) Binford (1989) Tringa flavipes (Gmelin, 1789) Binford (1989) Tringa melanoleuca (Gmelin, 1789) Binford (1989) Tringa semipalmata (Gmelin, 1789) Binford (1989) Tringa solitaria Wilson, 1813 Binford (1989) Chloroceryle americana (Gmelin, 1788) Binford (1989) Megaceryle alcyon (Linnaeus, 1758) Binford (1989) Megaceryle torquata (Linnaeus, 1766) Binford (1989) Order Coraciiformes Forbes, 1884 Family Alcedinidae Rafinesque, 1815 Order Passeriformes Linnaeus, 1758 Family Hirundinidae Linnaeus, 1758 Progne chalybea (Gmelin, 1789) Binford (1989) Progne subis (Linnaeus, 1758) Binford (1989) Stelgidopteryx serripennis (Audubon, 1838) Binford (1989) Quiscalus mexicanus (Gmelin, 1788) Navarro-Sigüenza et al. (2004) Tachycineta albilinea (Lawrence, 1863) Family Icteridae Linneus, 1758 Family Tyrannidae Vigors, 1825 Myiozetetes similis (Spix, 1825) Binford (1989) Binford (1989) Pitangus sulphuratus (Linnaeus, 1766) Binford (1989) Tyrannus melancholicus Vieillot, 1819 Binford (1989) 383 Bastida-Zavala et al. | Marine and coastal biodiversity of Oaxaca Table 16. Checklist of marine mammals recorded in the marine areas of Oaxaca, Mexico. Phylum Chordata Bateson, 1885 Reference Class Mammalia Linnaeus, 1758 Order Cetacea Brisson, 1762 Suborder Mysticeti Cope, 1891 Family Balaenopteridae Gray, 1864 Balaenoptera musculus (Linnaeus, 1758) Lira-Torres (2007) Megaptera novaeangliae (Borowski, 1781) Salinas and Ladrón-de-Guevara (1993) Suborder Odontoceti Flower, 1869 Family Delphinidae Gray, 1821 Feresa attenuata Gray, 1875 Globicephala macrorhynchus Gray, 1843 Grampus griseus (Cuvier, 1812) Orcinus orca (Linnaeus, 1758) Pseudorca crassidens (Owen, 1846) Stenella attenuata (Gray, 1846) Stenella longirostris (Gray, 1828) Tursiops truncatus (Montagu, 1821) Order Carnivora Bowdich, 1821 Family Otariidae (Gray, 1825) Gill, 1866 Zalophus californianus (Lesson, 1828) Acknowledgments: We appreciate the support of the many people who provided bibliographic data, records in databases, or help in the field. Regarding invertebrates, we wish to thank Jani Jarquín-González (ECOSUR, Chetumal), Zyanya Gema Mora-Vallín (UMAR, Puerto Ángel), and Citlalli Guevara-Cruz (Instituto Tecnológico de Bahía de Banderas, Nayarit) for finding several published records of crustaceans, hydrozoans and polychaetes, and Jose Luis Carballo and Jose Maria Aguilar-Camacho (ICML, UNAM, Mazatlán) for providing their papers on sponges. We thank Alejandra Lizárraga Rodríguez for assistance on echinoderms and Blanca Sánchez, Felipe Becerril (INECOL, Xalapa), Benjamin Lobato, Virgilio Antonio, Betzabeth González and Víctor Sánchez (UMAR, Puerto Ángel) for tetrapods. Special thanks are given to the Biodiversity Heritage Library (BHL) for making available many of the Allan Hancock Foundation Publications. We thank Lucero Piña-Mejía and Cotsikayala PachecoRamírez (UMAR, Puerto Ángel) for their help in references checking. We appreciate the comments and corrections of Ross MacCulloch, Rafael Riosmena, Sergio Salazar-Vallejo, José Luis Carballo, Francisco Solís, Luis Medrano and three anonymous referees. The projects FOSEMARNAT (2004-01-92) and CONABIO (HJ029 and HJ031) partially supported the compilation of the invertebrate and fish checklists. We thank Carmen Esqueda and Eduardo Ríos-Jara (Universidad de Guadalajara) for the review of the mollusk section. Literature Cited Aguayo-Lobo, A., S. Gaona, G. López-Ortega and M. Salinas-Zacarías. 1990. Mamíferos marinos dulceacuícolas, semiacuáticos y con tendencia al agua; p. 8-9 In M.T. Gutiérrez-de Mac Gregor, A. Coll-Hurtado and A. García-de Fuentes (coord.). Atlas Nacional de México. Volume II, section IV. México: Instituto de Geografía, UNAM. Aguilar-Rosas, R., M.J. Ochoa-Izaguire, L.E. Aguilar-Rosas, O.A. TovalínHernández and F. Páez-Osuna. 2009. Nuevos registros de macroalgas marinas para el estado de Sinaloa, México. Polibotánica 28(1): 1-14. 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