Tropical Bryology 25: 35-69, 2004 A revised checklist of Hawaiian mosses G. W. Staples, C. T. Imada, W. J. Hoe, and C. W. Smith Correction for Page 38: Hawaiian Moss Flora in Summary Families Genera Species Total named taxa Endemic Indigenous Alien Total 0 2 75 90 41 126 166 169 1 7 14 14 42 135 255 273 Correction for Page 39: Frahm et al. 2000, cited under Amphidium tortuosum, should correctly refer to the following: Frahm, J.-P., T. Klöcker, R. Schmidt, and C. Schöter. 2000. Revision der Gattung Amphidium (Musci, Dicranaceae). Tropical Bryology 18: 173-184. 35 Revised checklist of Hawaiian mosses Tropical Bryology 25: 35-69, 2004 A revised checklist of Hawaiian mosses1 G. W. StaplesA, C. T. ImadaA, W. J. Hoe,B and C. W. SmithC A – Hawaii Biological Survey, Bishop Museum, 1525 Bernice St., Honolulu, Hawaii 96817 U.S.A. B – deceased; C – Department of Botany, The Natural History Museum, Cromwell Road, London, SW7 5BD, U.K. Abstract. A revised and updated literature-based checklist of Hawaiian mosses is presented. Geographic coverage includes the eight main Hawaiian Islands; the Northwestern Hawaiian Islands are excluded. The checklist is alphabetically ordered by scientific names; the family is noted for each genus. Synonyms and misapplied names are cross-referenced to the accepted names. A bibliography of supporting references is included. Introduction The Hawaii Biological Survey (HBS) was established as a program of Bishop Museum by the Hawaii State Legislature in 1992, specifically to create and maintain accurate inventories of all species present in the Hawaiian Islands, and to document their presence with voucher specimen collections and bibliographies of relevant scientific literature. In the ensuing eleven years, comprehensive taxonomic inventories have been compiled for many groups of organisms, both native and alien, that comprise the Hawaiian biota. Vascular plants and marine algae have been the primary focus of botanical inventory efforts; research on non-marine algae, terrestrial cryptogamic plants, fungi, and lichens has lagged behind. This checklist is intended to fill one of the gaps in our knowledge of Hawaiian terrestrial 1 Contribution No. 2004–001 to the Hawaii Biological Survey. TROPICAL BRYOLOGY 25 (2004) cryptogamic plants—the mosses. Also in preparation by HBS staff members are checklists and bibliographies for the Hawaiian anthocerotes (hornworts) and hepatics (liverworts). A nonmarine algal checklist for the Hawaiian Islands is also in an advanced stage of preparation (A. Sherwood, pers. comm.). Eventually a checklist of Hawaiian lichens is planned. These ongoing inventories will help the Hawaii Biological Survey to fulfill its mission by systematically compiling literature-based inventories for each of these groups of organisms. Genesis of the current checklist This checklist of Hawaiian mosses begins where its predecessor—the checklist compiled by Hoe (1974)—left off, and updates that document by revising the nomenclature, adding new taxa and distributional records, and providing a bibliography of literature that supports the names in current use. The checklist 36 and bibliography were compiled by a collaborative effort between HBS staff members (Staples, Imada) working in partnership with two bryological specialists (Hoe, Smith), who had assembled a cache of literature citations and new distribution records pertinent to Hawaiian mosses. Funding from the USDA–PLANTS database project in 1998 provided the impetus for Staples and Imada to undertake conversion of Hoe’s 1974 moss checklist to an electronic format and then begin to update the taxonomy and distribution information it contained. Hoe and Smith generously shared their preliminary cache of information in order to facilitate the update process. Sadly, in May 2000, Hoe died suddenly, a great loss to the botanical community in Hawai‘i and to the world bryological community. After some reorganization the project resumed and was completed by Staples and Imada. In the final version of the checklist, taxonomic decisions were made by Staples, with review and critical input from Smith and two independent reviewers. Coverage It must be emphasized that this is a literature-based taxonomic checklist, not a specimen-based inventory. It is compiled from literature published from the mid-1970s to the present (2003). The starting point is the checklist published by Hoe (1974), who summarized the literature and reconciled the scientific names for Hawaiian mosses in use up to that time. The present checklist attempts to account for all scientific names in use for Hawaiian mosses in the latter decades of the twentieth century; it is not a comprehensive nomenclator for all names that have ever been published for, or applied to, Hawaiian mosses. No nomenclatural novelties or taxonomic changes are initiated in this checklist. Geographically, coverage focuses on the eight main Hawaiian Islands. There are, as yet, no published reports for bryophytes from the uninhabited Northwestern Hawaiian Islands, although a few bryophyte collections were made there in the last decades of the twentieth century. It had been Bill Hoe’s intention to identify these voucher specimens and write them up for publication but the specimens remain unstudied as this checklist goes to print. They await Staples et al. inclusion in a future iteration of the Hawaiian moss checklist. In the last quarter of the twentieth century there was a renaissance of bryological research worldwide and as a result many revisions, floristic accounts, and several monographs have been published that impact Hawaiian moss nomenclature. These regional and global studies resulted in significant taxonomic changes, as well as reporting discovery of several taxa new to the Islands, either rare native mosses or newly naturalized alien species. This checklist aims to summarize all this information in a concise format, as follows. Format Accepted scientific name The main body of the checklist is alphabetically organized by genus, species, and infraspecific taxon. Accepted names appear in bold italic type. Authors are included for the genera, species, and infraspecific epithets. Authors of moss genera have been standardized against the classification proposed in Bryophyte Biology (Shaw & Goffinet 2000); authors for five genera not found in that classification have been checked on the MOST website (cited below). All author names are abbreviated according to Authors of Plant Names style (Brummitt & Powell 1992). References that explain the taxonomic concept or provide geographic distribution data are cited after the accepted taxon name. The first mention of a genus includes the family assignment on the same line. Most genera are still placed in the same families recognized by Hoe (1974), but a few new family placements have been made since then. In such cases, we provide first the family assignment used by Hoe (1974), and mention alternative family placements afterward, with a reference to the source. Synonyms and misapplied names (in italics) follow the accepted name in paragraph style. These are limited primarily to names in use for Hawaiian mosses in the latter part of the twentieth century, although a few names from other parts of the world that enter into the nomenclature for particular taxa have been included to enable tracking names through the literature. TROPICAL BRYOLOGY 25 (2004) Revised checklist of Hawaiian mosses Status Following the taxonomy is a column indicating status of the taxon in the Hawaiian flora: end indicates an endemic, ind indicates indigenous, and nat signifies naturalized. A ? following any of these signifies uncertainty about the status of the taxon in the Hawaiian flora. While some research has been devoted to the mode of introduction of pteridophytes and angiosperms to Hawai‘i, resulting in a list of species that are likely to have been introduced by Polynesian voyagers that settled the archipelago and another (much larger) list of plants that arrived following European contact, little attention has been devoted to bryophytes in this regard. The authors have used their best judgment in cases where the status is not clear and appended a ? to whatever status we assigned, to draw attention to the species and invite investigation from specialists. One case worth noting is Racopilum cuspidigerum, which is widespread in the Southern Hemisphere but is found only in the Hawaiian Islands in the Northern Hemisphere. It is a prime candidate as a Polynesian introduction to Hawai‘i, and indeed to many of the other Polynesian islands in the South Pacific (S. Olson, pers. comm.). Island distribution Following the status category is a summary of geographic distribution on the eight main Hawaiian Islands. The island name abbreviations are explained at the foot of each page. An island abbreviation appears in the row to the right of the taxon name to signify that the taxon is reported in the literature from that island. Absence of the island name abbreviation indicates the moss is not (yet) known from that island. Given the inadequate state of moss collecting in the Hawaiian Islands, many new island distribution records are sure to be found in the future. Synonym list and Literature Consulted Following the main body of the checklist, an alphabetical list of all synonyms and misapplied names is provided. This list crossreferences every synonym or misapplied name to the main entry found under the scientific name TROPICAL BRYOLOGY 25 (2004) 37 accepted in this checklist. The checklist concludes with a bibliography of the references cited in the checklist; more than 130 bryological titles were consulted in the course of the project, but only those providing taxonomic or distribution data pertinent to this checklist are included here. Anomalies and incomplete records A few moss names without any island distribution are included, based on (mostly) older literature that reports the species from the Hawaiian (or sometimes Sandwich) Islands without stating on which island the species was found. Two examples in the list are Dicranoweisia cirrata and Plagiothecium denticulatum. Their presence in the Hawaiian Islands needs to be confirmed and documented with contemporary collections. Another anomaly is Mniobryoides degeneri, an invalid name published without a description or indication of a type specimen by H. Hörmann. It has been pointed out (S. Olson, pers. comm.) that this is probably a misidentification for some well-known species. However, M. degeneriae has never been assessed by a competent bryologist to ascertain its true identity. We leave the name in the checklist to point out that more study is required. A note about family circumscriptions The past 25 years has brought an explosion in research on bryophyte systematics and a great deal of new information has accrued, which has led to many changes in moss classification. Family classification in particular has become increasingly problematic as new data provide insights into relationships between genera. Several new families have been erected and other genera have been reassigned. We have indicated the family where Hoe (1974) assigned each genus, followed by alternative family assignments where these were discovered. Two principle reference works were consulted for family assignments: first, Bryophyte Biology (Shaw & Goffinet 2000), and in a few cases the bryological nomenclator included in the W 3 MOST database of mosses (http:// mobot.mobot.org/W3T/Search/most.html) on the TROPICOS website. Staples et al. 38 Hawaiian Moss Flora in Summary Families0 Genera Species Total named taxa Endemic 41 2 75 90 The way forward It is hoped that by bringing together the literature and providing an updated checklist of names in current use, one or more competent bryologists might be inspired to undertake preparation of a moss flora for the Hawaiian Islands. The last floristic account of the Hawaiian mosses is that by Bartram (1933, 1939). Bill Hoe intended to write a modern flora of the Hawaiian mosses, but his untimely death prevented that. It is our hope that professional bryologists will take up the challenge to complete this much needed reference work. Those who would study Hawaiian mosses will find type material and historic specimens widely distributed in herbaria around the world. In the last decades of the twentieth century, D. Vitt and his students as well as a team of cryptogamic botanists from the National Museum of Tokyo collected mosses throughout the main Hawaiian Islands. But by far the most comprehensive late twentieth century collections of Hawaiian mosses are those made by Hoe between 1965 and 1995. Today the first set of Hoe’s collections resides in two herbaria: the Herbarium Pacificum, Bishop Museum, Honolulu (BISH) and the Claude Philipps Herbarium, Delaware State University, Dover (DOV), with duplicates widely distributed in other herbaria. Hoe donated his personal herbarium to BISH in 1983. His residual duplicates and some new material acquired after 1983 were deposited at DOV after the sale of Hoe’s bryological library and herbarium. The Hoe collections in BISH and DOV provide an excellent resource for bryological researchers and afford a solid foundation for any floristic account of Hawaiian mosses. Indigenous 1 126 166 169 Alien 42 7 14 14 Total 135 255 273 Acknowledgments Funding for initial data compilation was made available by the USDA-PLANTS program; we thank Scott Petersen and Mark Skinner for their support and assistance. Our Museum colleagues Katie Anderson and Karin Kami assisted with data entry and bibliographic compilation. Bishop Museum librarian Patrice Belcher obtained numerous critical references through interlibrary loan services. Reviews of the draft checklist by Storrs Olson, U.S. National Museum, and James Shevock, U.S. Geological Survey, brought to light several bryological articles overlooked during the compilation as well as many suggestions for its improvement. 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Genera of Pottiaceae: mosses of harsh environments. Bulletin of the Buffalo Society of Natural Sciences 32: i–vi, 1–378. Zander, R.H., and P.M. Eckel. 1982. Hymenostylium recurvirostrum var. insigne and Barbula amplexifolia in British Columbia, Canada. Canadian Journal of Botany 60: 1596– 1600. Zander, R.H., and W.J. Hoe. 1979. Geographic disjunction and heterophylly in Tortella fragilis var. tortelloides (= Sarconeurum tortelloides). Bryologist 82(1): 84–87. Zander, R.H., and D.H. Vitt. 1979. Gametophytic distinctions of Zygodon (Orthotrichaceae) and Anoectangium and Leptodontium (Pottiaceae) and the status of Anoectangium rubrigemmium of Hawaii. Canadian Journal of Botany 57(3): 292–296. Zanten, B.O. van. 1959. Trachypodaceae—a critical revision. Blumea 9(2): 477–575. Zanten, B.O. van. 1971. Musci. Pp. 173–227. In Zinderen-Bakker, E.M. van, J.M. Winterbottom, and R.A. Dyer (eds.), Marion and Prince Edward Islands. A. A. Balkema, Cape Town. Zilstra, G. 1990. Report of the Committee for Bryophyta: I. Taxon 39(2): 289–292. Island abbreviations: Ni Ni‘ihau; K Kaua‘i; O O‘ahu; Mo Moloka‘i; M Maui; Ka Kahoolawe; H Hawai‘i. TROPICAL BRYOLOGY 25 (2004) 43 Revised checklist of Hawaiian mosses FullName ACROPORIUM Mitt. (Sematophyllaceae) Acroporium fuscoflavum (Paris) Broth. [Hoe 1974, Miller et al. 1978] Acroporium fuscoflavum (Paris) Broth. var. baldwinii (Paris) Broth. ?Acroporium sigmatodontium misapplied AEROBRYOPSIS M.Fleisch. (Meteoriaceae) Aerobryopsis subdivergens (Broth.) Broth. ssp. scariosa (E.B.Bartram) Nog. [Hoe 1974, Noguchi 1976] Aerobryopsis scariosa E.B.Bartram Ptychomnion aciculare misapplied Aerobryopsis wallichii (Brid.) M.Fleisch. [Hoe 1974, Noguchi 1976] Aerobryopsis longissima (Dozy & Molk.) M.Fleisch. Aerobryopsis longissima (Dozy & Molk.) M.Fleisch. var. dozyana M.Fleisch. AMPHIDIUM Schimp. (Dicranaceae, or Rhabdoweisiaceae [Shaw & Goffinet 2000]) Amphidium lapponicum (Hedw.) Schimp. [Hoe 1979] Amphidium tortuosum (Hornsch.) Cufod. [Frahm et al. 2000, Muñoz & Pando 2000] Amphidium cyathicarpum (Mont.) Broth. Grimmia haleakalae Reichardt Grimmia haliacalae Reichardt ANDREAEA Hedw. (Andreaeaceae) Andreaea acutifolia Hook. F. & Wilson ssp. acuminata (Mitt.) Vitt [Vitt 1980a, Vitt & Hoe 1980] Andreaea acuminata Mitt. Andreaea rupestris Hedw. [Hoe 1977, Hoe 1979, Vitt 1980a, Vitt & Hoe 1980] Andreaea perpapillosa Broth. ANOECTANGIUM Schwägr. (Pottiaceae) Anoectangium aestivum (Hedw.) Mitt. [Hoe 1974, Zander 1977, Zander & Vitt 1979] Anoectangium euchloron (Schwägr.) Mitt. Anoectangium haleakalae (Müll.Hal.) Paris var. haleakalae Anoectangium haleakalae (Müll.Hal.) Paris var. laxulum E.B.Bartram Anoectangium harttiae E.B.Bartram ANOMOBRYUM Schimp. (Bryaceae) Anomobryum angustirete Broth. [Hoe 1974] TROPICAL BRYOLOGY 25 (2004) STATUS NI K O MO L M KA H ind K O Mo L M H ind K O Mo ind K O Mo L M H ind ind M M H ind M H ind M ind K O Mo L M end K M L M H Staples et al. 44 FullName BALDWINIELLA M.Fleisch. (Neckeraceae) Baldwiniella kealeensis (Reichardt) E.B.Bartram [Hoe 1974, Hoe 1975] BARBELLOPSIS Broth. (Meteoriaceae) Barbellopsis trichophora (Mont.) W.R.Buck [Hoe 1974, Streimann 1993, Buck 1998] Barbella trichophora (Mont.) M.Fleisch. BARBULA Hedw. (Pottiaceae) Barbula indica (Hook.) Spreng. [Hoe 1978, Zander 1993, Darigo 1996] Barbula cruegeri Sond. ex Müll.Hal. Barbula unguiculata misapplied BARTRAMIA Hedw. (Bartramiaceae) Bartramia baldwinii Müll.Hal. [Hoe 1974] Bartramia halleriana Hedw. [Hoe 1974] BRACHYMENIUM Schwägr. (Bryaceae) Brachymenium exile (Dozy & Molk.) Bosch & Sande Lac. [Hoe 1974] Brachymenium indicum (Dozy & Molk.) Bosch & Sande Lac. [Hoe 1979] BRACHYTHECIUM Schimp. (Brachytheciaceae) Brachythecium hawaiicum E.B.Bartram [Hoe 1974] Brachythecium lamprocarpum (Müll.Hal.) A.Jaeger [Hoe 1974] Brachythecium plumosum (Hedw.) B.S.G. [Hoe 1974] Brachythecium oxyrrhynchum (Dozy & Molk.) A.Jaeger Cirriphyllum oxyrrhynchum (Dozy & Molk.) M.Fleisch. Brachythecium rutabulum (Hedw.) B.S.G. [Hoe 1974] BREUTELIA (Bruch & Schimp.) Schimp. (Bartramiaceae) Breutelia affinis (Hook.) Mitt. [Hoe 1974, Virtanen 1997] Breutelia kilaueae (Müll.Hal.) Broth. Breutelia arundinifolia (Duby) M.Fleisch. [Hoe 1974, Virtanen 1997] Breutelia crassicaulis (Müll.Hal.) Paris [Virtanen 1997] BRYOERYTHROPHYLLUM P.C.Chen (Pottiaceae) Bryoerythrophyllum recurvirostre (Hedw.) P.C.Chen [Hoe 1974] Didymodon recurvirostris (Hedw.) Jenn. STATUS NI K O MO L M KA H end K O Mo L M H ind K O Mo H nat O end ind ind M M M M K O Mo ind M O H H end ind K M M H H ind K O Mo M H ind M ind ind H K O M H ind M H ind M TROPICAL BRYOLOGY 25 (2004) 45 Revised checklist of Hawaiian mosses FullName BRYUM Hedw. (Bryaceae) Bryum argenteum Hedw. [Hoe 1974, Miller et al. 1978, Tan & Koponen 1989] Bryum argenteum Hedw. var. lanatum (P. Beauv.) Hampe Bryum leucophyllum Dozy & Molk. Bryum atrovirens Brid. [Hoe 1974] Bryum erythrocarpum Schwägr. Bryum baldwinii Broth. [Hoe 1974] Bryum caespiticium Hedw. [Hoe 1974] Bryum chrysoneuron Müll.Hal. [Miller et al. 1978] Bryum conicum Hornsch. [Ochi 1980] Bryum ceramiocarpum Müll.Hal. Bryum flavituber R.Wilczek & Demaret [Wilczek & Demaret 1976] Bryum hawaiicum Hoe [Hoe 1974] Bryum crassicostatum Broth. Bryum klinggraeffii Schimp. [Hoe 1979] Bryum lisae De Not. var. cuspidatum (Bruch & Schimp.) Margad. [Olson 1999a] Bryum mildeanum Jur. [Hoe 1974, Ochi 1974, Hoe 1979, Ignatov & Afonina 1992] Bryum apiculatum Schwägr. Bryum mauiensis Broth. Bryum megalostegium Sull. Bryum nitens Hook. Bryum pseudocapillare Besch. [Hoe 1975] Bryum radiculosum Brid. [Olson 1999a] BUCKIELLA Ireland (Hypnaceae) Buckiella draytonii (Sull.) Ireland [Hoe 1974, Ireland 2001] Hypnum draytoni(i) Sull. Hypnum eudorae Sull. Plagiothecium draytonii (Sull.) Müll.Hal. CALYMPERES Sw. (Calymperaceae) Calymperes fasciculatum Dozy & Molk. [Hoe 1974, Reese et al. 1986] Calymperes hawaiiense E.B.Bartram Calymperes tenerum Müll.Hal. [Hoe 1974] CAMPTOCHAETE Reichardt (Lembophyllaceae) Camptochaete pulvinata (Hook.f. & Wilson) A.Jaeger [Hoe 1974, Tangney 1997] CAMPYLOPODIUM (Müll.Hal.) Besch. (Dicranaceae) Campylopodium medium (Duby) Giese & J.-P.Frahm TROPICAL BRYOLOGY 25 (2004) STATUS NI K O MO L M KA H ind K O M H ind O M H end ind ind ind M M M nat O end M nat nat O O ind K O nat? nat K end K O Mo ind ind H L M O M H M H O K O Mo ind ind H H M K O Mo M H Staples et al. 46 FullName [Hoe 1974, Giese & Frahm 1985] Campylopodium euphorocladum (Müll.Hal.) Besch. CAMPYLOPUS Brid. (Dicranaceae) Campylopus exasperatus (Nees & Blume) Brid. [Frahm 1991] Campylopus fragilis (Brid.) Bruch. & Schimp. ssp. zollingerianus (Müll.Hal.) J.-P.Frahm [Hoe 1974, Frahm 1991, Frahm 1992] Campylopus boswellii (Müll.Hal.) Paris Campylopus boswellii (Müll.Hal.) Paris var. capitulatus E.B.Bartram Campylopus hawaiicus (Müll.Hal.) A.Jaeger var. densifolius (Ångstr.) J.-P.Frahm [Frahm 1978, Frahm 1991] Campylopus densifolius Ångstr. Campylopus densiretis Broth. Campylopus wawreanus Reichardt Campylopus hawaiicus (Müll.Hal.) A.Jaeger var. hawaiicoflexuosus (Müll.Hal.) J.-P.Frahm [Frahm 1978, Frahm 1991] Campylopus densifolius Ångstr. var. falcatus Broth. Campylopus densifolius Ångstr. var. hawaiicoflexuosus (Müll.Hal.) H.A.Mill. Campylopus hawaiicoflexuosus (Müll.Hal.) Müll.Hal. Campylopus hawaiicoflexuosus (Müll.Hal.) Paris Campylopus perangustifolius Müll.Hal. Campylopus purpureoflavescens Hampe, in part Dicranum hawaiicoflexuosum Müll.Hal. Campylopus hawaiicus (Müll.Hal.) A.Jaeger var. hawaiicus [Frahm 1978, Frahm 1991] Campylopus densifolius Ångstr. var. purpureoflavescens (Hampe ex Müll.Hal.) H.A.Mill. Campylopus purpureoflavescens Hampe, in part Dicranum hawaiicum Müll.Hal. Campylopus incurvatus J.-P.Frahm & Hoe [Frahm & Hoe 1978, Frahm 1991] Campylopus laxitextus Sande Lac. [Hoe 1974, Frahm 1991] Campylopus fumarioli Müll.Hal. Campylopus praemorsus (Müll.Hal.) A.Jaeger [Frahm 1976, Hoe 1979, Frahm 1991] Campylopus exasperatus (Nees & Blume) Brid. Campylopus microcephalus (Müll.Hal.) Paris Dicranum microcephalum Müll.Hal. Dicranum praemorsum Müll.Hal. STATUS NI K O MO L M KA H ind K ind K O Mo M end K O Mo M H end K O Mo L M H ind K O Mo L M H end M H ind K O Mo L M H end K O H L M TROPICAL BRYOLOGY 25 (2004) 47 Revised checklist of Hawaiian mosses FullName STATUS NI K O MO L M KA H ind KO M H Campylopus schmidii (Müll.Hal.) A.Jaeger ssp. hemitrichius (Müll.Hal.) J.-P.Frahm [Hoe 1974, Frahm 1991] Campylopus skottsbergii Broth. Campylopus tubulosus E.B.Bartram Dicranum skottsbergii Broth. ind K O M H Campylopus schmidii (Müll.Hal.) A.Jaeger ssp. schmidii [Hoe 1974, Hoe 1977, Frahm 1991] Campylopus aureus Bosch & Sande Lac. Campylopus introflexus misapplied Campylopus introflexus (Hedw.) Brid. var. polytrichoides misapplied Campylopus polytrichoides misapplied Campylopus umbellatus (Arn.) Paris var. pterotoneuron ind O (Müll.Hal.) J.-P.Frahm [Hoe 1974, Frahm 1991] Campylopus geniculatus Ångstr. ind K O L M H Campylopus umbellatus (Arn.) Paris var. umbellatus [Hoe 1974, Frahm 1991] ?Campylopus didrichsenii (Müll.Hal.) A.Jaeger ?Campylopus pertristis (Müll.Hal.) Paris Campylopus richardii misapplied Campylopus sandwicensis Paris Campylopus umbellatus (Arn.) E.B.Bartram Dicranum didrichsenii Müll.Hal. Dicranum pertriste Müll.Hal. Thysanomitrion hawaiicum Müll.Hal. Thysanomitrion umbellatum Arn. end K M H Campylopus wheeleri (Müll.Hal.) Paris [Hoe 1974, Frahm 1991] Dicranoloma wheeleri (Müll.Hal.) Paris Dicranum wheeleri Hampe ex Müll.Hal. CERATODON Brid. (Ditrichaceae) Ceratodon purpureus (Hedw.) Brid. ssp. purpureus [Hoe 1974, Hoe 1977, Miller et al. 1978, Burley & Pritchard 1990] Ceratodon purpureus (Hedw.) Brid. fma. purpureus Ceratodon purpureus (Hedw.) Brid. fma. xanthopus (Sull.) E.Britton ex Grout CHENIA R.H.Zander (Pottiaceae) Chenia leptophylla (Müll.Hal.) R.H.Zander [Hoe 1974, Zander 1993] Tortula rhizophylla (Sakurai) Z.Iwats. & K.Saito Tortula vectensis E.F.Warb. & Crundw. CLAOPODIUM (Lesq. & James) Renauld & Cardot (Thuidiaceae, or Leskeaceae [Shaw & Goffinet 2000]) Claopodium prionophyllum (Müll. Hal.) Broth. [Hoe 1974] TROPICAL BRYOLOGY 25 (2004) ind K O nat O ind O Mo M H M H Staples et al. 48 FullName Claopodium amblystegioides Dixon Claopodium nervosum M.Fleisch. Claopodium whippleanum (Sull.) Renauld & Cardot [Hoe 1974] Claopodium hawaiiense R.S.Williams CLASTOBRYOPSIS M.Fleisch. (Sematophyllaceae) Clastobryopsis robusta (Broth.) M.Fleisch. [Hoe 1974, Tan & Jia 1999] Aptychella robusta (Broth.) M.Fleisch. CTENIDIUM (Schimp.) Mitt. (Hypnaceae) Ctenidium elegantulum Broth. [Hoe 1974, Nishimura 1985] STATUS NI K O MO L M KA ind K O M ind K M end O M CYRTOPUS (Brid.) Hook.f. (Cyrtopodaceae) Cyrtopus setosus (Hedw.) Hook.f. [Hoe 1974] ind DALTONIA Hook. & Taylor (Hookeriaceae, or Daltoniaceae [Shaw & Goffinet 2000]) Daltonia baldwinii Broth. [Hoe 1974] Daltonia contorta Müll.Hal. [Hoe 1974, Hoe 1979] Daltonia pseudostenophylla E.B.Bartram [Hoe 1974] Daltonia rufescens Broth. [Hoe 1974] end ind ind ind M K O Mo L M O M O M end end M M DICRANELLA (Müll.Hal.) Schimp. (Dicranaceae) Dicranella exilis Sull. [Hoe 1974] Dicranella hawaiica (Müll.Hal.) Broth. var. hawaiica [Hoe 1974] Microdus hawaiicus (Müll.Hal.) Paris Dicranella hawaiica (Müll.Hal.) Broth. var. tomentella E.B.Bartram [Hoe 1974] Dicranella hochreutineri Cardot [Hoe 1974] Dicranella hochreuteri misapplied Dicranella integrifolia E.B.Bartram [Hoe 1974] Dicranella rigidula E.B.Bartram [Hoe 1974] DICRANODONTIUM Bruch & Schimp. (Dicranaceae) Dicranodontium denudatum (Brid.) E.Britton [Hoe 1974] Dicranodontium falcatum Broth. var. atrovirens (Broth.) E.B.Bartram [Hoe 1974] Dicranodontium porodictyon Cardot & Thér. [Hoe 1974, Hoe 1979, Frahm 1997] Dicranodontium falcatum Broth. var. falcatum Dicranodontium hawaiicum Broth. DICRANOLOMA (Renauld) Renauld (Dicranaceae) Dicranoloma gracile Broth. ex E.B.Bartram [Hoe 1974] H H H K Mo H H H H end O M ind K O L M H end end O Mo K O M H H ind end ind end K O Mo M M H M H M TROPICAL BRYOLOGY 25 (2004) 49 Revised checklist of Hawaiian mosses FullName DICRANOWEISIA Milde (Dicranaceae, or Rhabdoweisiaceae [Shaw & Goffinet 2000]) Dicranoweisia cirrata (Hedw.) Lindb. [Hoe 1974, Miller et al. 1978] DICRANUM Hedw. (Dicranaceae) Dicranum speirophyllum Mont. var. breviflagellare (Müll.Hal.) E.B.Bartram [Hoe 1974] Dicranum speirophyllum Mont. var. condensatum (Sull.) Wijk & Margad. [Hoe 1974] Dicranum speirophyllum Mont. var. elongatum (Sull.) Wijk & Margad. [Hoe 1974] Dicranum speirophyllum Mont. var. speirophyllum [Hoe 1974] DIDYMODON Hedw. (Pottiaceae) Didymodon insulanus (De Not.) M.O.Hill [Hoe 1974, Zander 1978, Corley et al. 1981] Barbula vinealis Brid. var. cylindrica (Taylor) Boulay Barbula vinealis Brid. var. flaccida B.S.G. Didymodon vinealis (Brid.) R.H.Zander var. flaccidus (Bruch & Schimp.) R.H.Zander DISTICHIUM Bruch & Schimp. (Ditrichaceae) Distichium capillaceum (Hedw.) B.S.G. [Hoe 1974] DISTICHOPHYLLUM Dozy & Molk. (Hookeriaceae, or Daltoniaceae [Shaw & Goffinet 2000]) Distichophyllum freycinetii (Schwägr.) Mitt. var. crasseturgescens (Müll.Hal.) E.B.Bartram ex Hoe [Hoe 1974] Mniadelphus freycineti (Schwägr.) Müll.Hal. var. crasseturgescens Müll.Hal. Distichophyllum freycinetii (Schwägr.) Mitt. var. freycinetii [Hoe 1974] Distichophyllum paradoxum (Mont.) Mitt. [Hoe 1974] ECTROPOTHECIUM Mitt. (Hypnaceae) Ectropothecium arcuatum Mitt. [Hoe 1974] Ectropothecium decurrens (Sull.) Nishimura [Hoe 1974, Nishimura 1985] Ctenidium decurrens (Sull.) Broth. Ctenidium pulcherrimum Broth. Ctenidium submalacobolum Broth. Cupressina decurrens (Sull.) Müll.Hal. Ectropothecium rockii Broth. Hypnum decurrens Sull. TROPICAL BRYOLOGY 25 (2004) STATUS NI K O MO L M KA ind end H? O M end ind H H end end H M K O Mo O M H M ind H end end K O Mo L M H end K O Mo L M H end end K O K O Mo H H M M Staples et al. 50 FullName STATUS NI K Stereodon decurrens (Sull.) Mitt. Taxiphyllum gracile Broth. Ectropothecium sandwichense (Hook. & Arn.) Mitt. ind K [Hoe 1974, Hoe 1979] ind Ectropothecium tutuilum (Sull.) Mitt. [Miller et al. 1978] Ectropothecium viridifolium E.B.Bartram [Hoe 1974, end Ni Wichman & St. John 1990] Ectropothecium zollingeri (Müll. Hal.) A.Jaeger var. ind K filicaule (M.Fleisch.) H.A.Mill., H.Whittier & B.Whittier [Hoe 1974, Hoe 1975, Miller et al. 1978] Glossadelphus zollingeri (Müll.Hal.) M.Fleisch. var. filicaulis (M.Fleisch.) M.Fleisch. ENCALYPTA Hedw. (Encalyptaceae) Encalypta ciliata Hedw. [Horton 1983] Encalypta scabrata E.B.Bartram Encalypta rhabdocarpa Schwägr. [Hoe 1974] Encalypta rhaptocarpa misapplied Encalypta sandwicensis Sull. [Hoe 1974, Hoe 1977] ENTODON Müll.Hal. (Entodontaceae) Entodon plicatus Müll.Hal. [Hoe 1974, Buck 1990] Entodon hillebrandii Müll.Hal. Entodon solanderi (Ångstr.) A.Jaeger Entodon subcuspidatus (Müll.Hal.) E.B.Bartram [Hoe 1974, Hoe 1979] ENTODONTOPSIS Broth. (Stereophyllaceae) Entodontopsis anceps (Bosch & Sande Lac.) W.R.Buck & Ireland [Hoe 1974, Buck & Ireland 1985] Stereophyllum oahuense Broth. ENTOSTHODON Schwägr. (Funariaceae) Entosthodon subintegrus (Broth.) H.A.Mill., H.Whittier & B.Whittier [Miller et al. 1978] Funaria subintegra Broth. FABRONIA Raddi (Fabroniaceae) Fabronia ciliaris (Brid.) Brid. [Hoe 1977] Fabronia degeneri E.B.Bartram [Hoe 1974] Fabronia nietneri Müll.Hal. [Hoe 1974, Hoe 1975] FISSIDENS Hedw. (Fissidentaceae) Fissidens aphelotaxifolius Pursell [Pursell & Hoe 1977] Fissidens bryoides Hedw. [Pursell & Hoe 1977] Fissidens hawaiicus E.B.Bartram Fissidens insularis E.B.Bartram Fissidens oahuensis E.B.Bartram O MO L M KA H O Mo L M H H O L M O Mo L M H M H ind ind H ind ind K O ind K ind Mo M H M H M O end K O Mo L M H ind end ind O Mo K O M H H ind ind M K O Mo L M H M TROPICAL BRYOLOGY 25 (2004) 51 Revised checklist of Hawaiian mosses FullName Fissidens delicatulus Ångstr. [Pursell & Hoe 1977] Fissidens elegans Brid. [Pursell & Hoe 1977] Fissidens baldwinii Broth. Fissidens bishopii Hoe Fissidens hoei Pursell [Pursell & Hoe 1977] Fissidens kilaueae Hoe & H.A.Crum [Pursell & Hoe 1977] Fissidens lancifolius E.B.Bartram [Pursell & Hoe 1977] Fissidens lindbergii Mitt. [Pursell & Hoe 1977, Darigo 1996, Pursell 1997] Fissidens crumii Hoe Fissidens nothotaxifolius Pursell & Hoe [Pursell & Hoe 1977] Fissidens pacificus Ångstr. [Hoe 1974, Pursell & Hoe 1977] Fissidens mauiensis Müll.Hal. Fissidens submarginatus Bruch [Pursell & Hoe 1977, Hoe 1979, Bruggeman-Nannenga & Pursell 1995] Fissidens intermedius Müll.Hal. Fissidens taxifolius Hedw. [Pursell & Hoe 1977] FLORIBUNDARIA M.Fleisch. (Meteoriaceae) Floribundaria floribunda (Dozy & Molk.) M.Fleisch. [Hoe 1974, Tan 1984] Floribundaria baldwinii Broth. FUNARIA Hedw. (Funariaceae) Funaria hygrometrica Hedw. [Hoe 1974, Hoe 1975] GAROVAGLIA Endl. (Pterobryaceae, or Garovagliaceae [Shaw & Goffinet 2000]) Garovaglia powellii Mitt. var. haleakalae (Broth.) During [During 1977] Garovaglia haleakalae Broth. GLOSSADELPHUS M.Fleisch.1 (Sematophyllaceae) Glossadelphus acutifolius E.B.Bartram [Hoe 1974, Tixier 1988] Glossadelphus limnobioides Broth. [Hoe 1974, Tixier 1988] Glossadelphus mauiensis Broth. [Hoe 1974, Tixier 1988] Glossadelphus ogatae Broth. & Yasuda [Hoe 1974, Hoe 1979, Tixier 1988] Glossadelphus chrysobasilaris Broth. 1 STATUS NI K O MO L M KA end K O L M ind K O Mo L M L L M H H H end end O K O end nat K O Mo L M O M H end K O Mo L M H end K O M H ind K O Mo L M H ind O M ind O M ind O M end M end ind O H H ind end H M H M M H Buck (1987b) reduced the type species of Glossadelphus to Phyllodon so the genus name is not usable. The correct placement for the 4 species left here remains to be determined. TROPICAL BRYOLOGY 25 (2004) Staples et al. 52 FullName STATUS NI K O MO L M KA GRIMMIA Hedw. (Grimmiaceae) Grimmia anodon Bruch & Schimp. [Hoe 1977, Hoe ind M 1979, Muñoz & Pando 2000] Grimmia apocarpa Hedw. var. pulvinata (Hedw.) G.N.Jones Gymnostomum pulvinatum Hedw. Grimmia laevigata (Brid.) Brid. [Hoe 1974, Hoe 1979, ind M Muñoz & Pando 2000] Grimmia longirostris Hook. [Hoe 1977,Muñoz 1998] ind M Grimmia pilifera misapplied end M Grimmia mauiense Greven [Greven 2003] end M Grimmia maunakeaense Greven [Greven 2003] Grimmia pulvinata (Hedw.) Sm. [Muñoz & Pando 2000] ind M Leersia pulvinata Hedw. Grimmia trichophylla Grev. [Hoe 1974, Muñoz & Pando ind M 2000] HAGENIELLA Broth. (Sematophyllaceae) Hageniella micans (Mitt.) B.C.Tan & Y.Jia [Hoe 1974, Tan & Jia 1999] Hageniella pacifica Broth. HAPLOCLADIUM Müll.Hal. (Leskeaceae) Haplocladium microphyllum (Hedw.) Broth. [Hoe 1974, Watanabe & Iwatuski 1981, Zilstra 1990] Bryohaplocladium microphyllum (Hedw.) R.Watan. & Z.Iwats. HAPLOHYMENIUM Dozy & Molk. (Thuidiaceae, or Anomodontaceae [Shaw & Goffinet 2000]) Haplohymenium triste (Ces.) Kindb. [Hoe 1974, Hoe 1979] HETEROPHYLLIUM (Schimp.) Kindb. (Sematophyllaceae) Heterophyllium subauriculatum (Müll.Hal.) Broth. [Hoe 1974] HOLOMITRIUM Brid. (Dicranaceae) Holomitrium seticalycinum Müll.Hal. [Hoe 1974, Hoe 1979] Holomitrium squarrifolium E.B.Bartram [Hoe 1974] Holomitrium ferriei misapplied HOMALIODENDRON M.Fleisch. (Neckeraceae) Homaliodendron flabellatum (Sm.) M.Fleisch. [Hoe 1974] Homalia densa Bosw. Homalia praelonga Reichardt ind K H H H H H H M H M H end M H end K O Mo L M H ind O ind K O end ind O K O Mo L M H H TROPICAL BRYOLOGY 25 (2004) 53 Revised checklist of Hawaiian mosses FullName HOOKERIA Sm. (Hookeriaceae) Hookeria acutifolia Hook. & Grev. [Hoe 1974, Hoe 1975] HYMENOSTYLIUM Brid. (Pottiaceae) Hymenostylium recurvirostrum (Hedw.) Dixon var. cylindricum (E.B.Bartram) R.H.Zander [Zander & Eckel 1982] Merceyopsis crassinervis (Broth.) E.B.Bartram Molendoa crassinervis (Broth.) E.B.Bartram Hymenostylium recurvirostrum (Hedw.) Dixon var. recurvirostrum [Zander 1993] Hymenostylium firmum (Müll.Hal.) Broth. ex E.B.Bartram STATUS NI K O MO L M KA ind O L M ind H H H ind K O M H HYOPHILA Brid. (Pottiaceae) Hyophila involuta (Hook.) A.Jaeger [Hoe 1974] Hyophila dozy-molkenboeri M.Fleisch. ind K O Mo M H HYPNUM Hedw. (Hypnaceae) Hypnum plumaeforme Wilson [Crum & MuellerDombois 1968, Hoe 1974] nat M H HYPOPTERYGIUM Brid. (Hypopterygiaceae) Hypopterygium sandwicense Broth. [Hoe 1974, Kruijer 2002] end M H ISOPTERYGIUM Mitt. (Hypnaceae) Isopterygium albescens (Hook.) A.Jaeger [Hoe 1974] Isopterygium vineale E.B.Bartram [Hoe 1974] ind end K O Mo L M K O Mo M H H LEPTOBRYUM (Bruch & Schimp.) Wilson (Bryaceae, or Meesiaceae [Shaw & Goffinet 2000]) Leptobryum pyriforme (Hedw.) Wilson [Hoe 1974] ind M H ind M H end end K O Mo L M K O Mo M H end end K O Mo L M LEPTODONTIUM (Müll.Hal.) Lindb. (Pottiaceae) Leptodontium flexifolium (Dicks. ex With.) Hampe [Hoe 1974, Hoe 1977] Leptodontium brevicaule E.B.Bartram LEUCOBRYUM Hampe (Leucobryaceae) Leucobryum gracile Sull. var. gracile [Hoe 1974] Leucobryum gracile Sull. var. hamatum E.B.Bartram [Hoe 1974] Leucobryum pachyphyllum Müll.Hal. [Hoe 1974] Leucobryum seemannii Mitt. var. fumarioli (Müll.Hal.) H.A.Mill. [Hoe 1974] TROPICAL BRYOLOGY 25 (2004) H Staples et al. 54 FullName STATUS NI K O MO L M KA Leucobryum hawaiense (Reichardt) E.B.Bartram var. fumarioli (Müll.Hal.) E.B.Bartram Leucobryum solfatarae Müll.Hal. var. fumarioli (Müll.Hal.) Wijk & Margad. end K O Mo L M Leucobryum seemannii Mitt. var. seemannii [Hoe 1974, Hoe 1975] Leucobryum hawaiense (Reichardt) E.B.Bartram Leucobryum solfatarae Müll.Hal. var. hawaiense (Reichardt) Wijk & Marg. end K Mo M Leucobryum seemannii Mitt. var. solfatarae (Müll.Hal.) H.A.Mill. [Hoe 1974] Leucobryum hawaiense (Reichardt) E.B.Bartram var. solfatarae (Müll.Hal.) E.B.Bartram Leucobryum solfatarae Müll.Hal. LEUCOLOMA Brid. (Dicranaceae) Leucoloma molle (Müll.Hal.) Mitt. [Hoe 1974] Leucoloma scaberulum E.B.Bartram [Hoe 1974] ind end O O LEUCOPHANES Brid. (Leucobryaceae, or Calymperaceae [Shaw & Goffinet 2000]) Leucophanes glaucum (Schwägr.) Mitt. [Hoe 1974] ind O LIMBELLA (Müll.Hal.) Müll.Hal. (Amblystegiaceae) Limbella tricostata (Sull.) Müll.Hal. [Hoe 1974, Christy 1987] Sciaromium tricostatum (Sull.) Mitt. MACROCOMA (Müll.Hal.) Grout (Orthotrichaceae) Macrocoma tenue (Hook. & Grev.) Vitt ssp. sullivantii (Müll.Hal.) Vitt [Hoe 1974, Vitt 1980b, Vitt 1981] Macrocoma intricata (Müll.Hal.) H.A.Miller, H.Whittier & B.Whittier Macromitrium intricatum Müll.Hal. MACROMITRIUM Brid. (Orthotrichaceae) Macromitrium brevisetum Mitt. [Hoe 1974, Hoe 1979] Macromitrium emersulum Müll.Hal. [Hoe 1974, Hoe 1975] Macromitrium incurvifolium (Hook. & Grev.) Schwägr. [Hoe 1974] Macromitrium cumingii Müll.Hal. Macromitrium microstomum (Hook. & Grev.) Schwägr. [Hoe 1974, Vitt 1983] Macromitrium owahiense Müll.Hal. Macromitrium reinwardtii Schwägr. Macromitrium piliferum Schwägr. [Hoe 1974] H H H M H end K O Mo M H ind K O Mo M H end end K O Mo L M K O Mo L M H H ind K M H ind K O Mo L M H end K O Mo L M H TROPICAL BRYOLOGY 25 (2004) 55 Revised checklist of Hawaiian mosses FullName MACROTHAMNIUM M.Fleisch. (Hylocomiaceae) Macrothamnium macrocarpum (Reinw. & Hornsch.) M.Fleisch. [Hoe 1974] MICRODUS Schimp. ex Besch. (Dicranaceae) Microdus hillebrandii (Müll.Hal.) Paris [Hoe 1974] Dicranella hillebrandii (Müll.Hal.) Broth. MNIOBRYOIDES Hörmann (Bryaceae) Mniobryoides degeneriae Hörmann [Hoe 1974] MNIUM Hedw. (Mniaceae) Mnium marginatum (With.) P.Beauv. [Hoe 1974] Mnium serratum Schrad. ex Brid. NECKERA Hedw. (Neckeraceae) Neckera hawaiicopennata Müll.Hal. [Hoe 1974] NECKEROPSIS Reichardt (Neckeraceae) Neckeropsis lepineana (Mont.) M.Fleisch. [Hoe 1974, Enroth 1989] Neckeropsis obtusata (Mont.) Broth. [Olson 1999b] OCTOBLEPHARUM Hedw. (Leucobryaceae, or Calymperaceae [Shaw & Goffinet 2000]) Octoblepharum albidum Hedw. [Hoe 1974, Hoe 1975] ORTHODONTIUM Wilson (Bryaceae, or Orthodontiaceae [Shaw & Goffinet 2000]) Orthodontium pellucens (Hook.) B.S.G. [Hoe 1974, Hoe 1977] ORTHOTRICHUM Hedw. (Orthotrichaceae) Orthotrichum alpestre Hornsch. var. berggrenii (E.B.Bartram) Lewinsky [Hoe 1974, Lewinksy 1993] Orthotrichum berggrenii E.B.Bartram Orthotrichum diaphanum Schrad. ex Brid. [Hoe 1974, Hoe 1977, Lewinsky 1993] Orthotrichum hawaiicum Müll.Hal. [Hoe 1974] Orthotrichum incrassatum Lewinsky [Lewinsky 1993] Orthotrichum lyellii Hook. & Taylor [Hoe 1974, Lewinsky 1993] Orthotrichum verrucatum Müll.Hal. Orthotrichum rupestre Schleich. & Schwägr. [Hoe 1974, Lewinsky 1993] Orthotrichum hillebrandii Müll.Hal. TROPICAL BRYOLOGY 25 (2004) STATUS NI K O MO L M KA ind M ind O Mo end H ind M end M ind K O nat K ind K O Mo M ind K M end H M O? H H ind M H end end ind M H H M H ind O Staples et al. 56 FullName STATUS NI K O MO L M KA OXYRRHYNCHIUM (Schimp.) Warnst. (Brachytheciaceae) ind K O Mo L M Oxyrrhynchium vagans (A.Jaeger) Ignatova & Huttunen [Hoe 1974, Hoe 1975, Ignatov & Huttunen 2002] Eurhynchium vagans (A.Jaeger) E.B.Bartram Rhynchostegium vagans A.Jaeger OXYSTEGUS (Limpr.) Hilp. (Pottiaceae) Oxystegus tenuirostris (Hook. & Taylor) A.J.E.Sm. var. gemmiparus (Schimp.) R.H.Zander [Hoe 1977, Hoe 1979, Zander 1982] Trichostomum tenuirostre (Hook. & Taylor) Lindb. PALAMOCLADIUM Müll.Hal. (Brachytheciaceae) Palamocladium wilkesianum (Sull.) Müll.Hal. [Hoe 1974, Hofmann 1997] Hypnum wilkesianum Sull. Meteorium pallidovirens Müll.Hal. Meteorium sciuroides Müll.Hal. Palamocladium aptychoides Müll.Hal. Palamocladium sciurellum Müll.Hal. Palamocladium wilkesianum (Sull.) Müll.Hal. var. altisetum Müll.Hal. Palamocladium wilkesianum (Sull.) Müll.Hal. var. sciuroides (Müll.Hal.) Wijk & Margad. Pleuropus aptychoides (Müll.Hal.) Broth. Pleuropus pallidovirens (Müll.Hal.) Broth. Pleuropus sciurellus (Müll.Hal.) Broth. Pleuropus wilkesianus (Sull.) Broth. Pleuropus wilkesianus (Sull.) Broth. var. sciuroides (Müll.Hal.) E.B.Bartram PELEKIUM Mitt. (Thuidiaceae) Pelekium versicolor (Hornsch. ex Müll.Hal.) Touw [Hoe 1974, Touw 2001] Thuidium crenulatum Mitt. Thuidium nanophyllum Müll.Hal. PHILONOTIS Brid. (Bartramiaceae) Philonotis falcata (Hook.) Mitt. [Hoe 1974] Philonotis hastata (Duby) Wijk & Margad. [Hoe 1974] Philonotis laxissima Mitt. Philonotis hawaica (Müll.Hal.) Broth. [Hoe 1974] Philonotis turneriana (Schwägr.) Mitt. var. sullivantii (Müll.Hal.) E.B.Bartram [Hoe 1974] Philonotis turneriana (Schwägr.) Mitt. var. turneriana [Hoe 1974] ind K O end K O Mo L M ind O H H M H M H ind ind K O Mo O M M H end end K O Mo L M O M H H ind K O Mo L M H TROPICAL BRYOLOGY 25 (2004) 57 Revised checklist of Hawaiian mosses FullName PILOTRICHELLA (Müll.Hal.) Besch. (Meteoriaceae, or Lembophyllaceae [Shaw & Goffinet 2000]) Pilotrichella mauiensis (Sull.) A.Jaeger [Hoe 1974] PLAGIOMNIUM T.J.Kop. (Mniaceae) Plagiomnium rostratum (Schrad.) T.J.Kop. [Hoe 1974] Mnium rostratum Schrad. Plagiomnium prorepens (Müll.Hal.) T.J.Kop. Plagiomnium rhynchosporum (Hook.) T.J.Kop. PLAGIOPUS Brid. (Bartramiaceae) Plagiopus oederiana (Sw.) H.A.Crum & L.E.Anderson [Hoe 1974] Plagiopus longisetus (Brid.) E.B.Bartram Plagiopus oederi (Brid.) Limpr. PLAGIOTHECIUM Bruch & Schimp. (Plagiotheciaceae) Plagiothecium cavifolium (Brid.) Z.Iwats. [Hoe 1977] Plagiothecium denticulatum (Hedw.) Schimp. [Hoe 1974] Plagiothecium mauiense Broth. [Hoe 1974] STATUS NI K O MO L M KA end ind O K O Mo H M H M H ind M ind ind M end M PLATYHYPNIDIUM M.Fleisch. (Amblystegiaceae, or Brachytheciaceae [Shaw & Goffinet 2000]) Platyhypnidium muelleri (A.Jaeger) M.Fleisch. [Hoe 1974] Eurhynchium mulleri misapplied ind K O Mo POGONATUM P. Beauv. (Polytrichaceae) Pogonatum tahitense Schimp. ex Besch. [Hoe 1974] Pogonatum baldwinii (Müll.Hal.) Paris ind K O Mo L M end M ind M H M H M H POHLIA Hedw. (Bryaceae, or Mniaceae [Shaw & Goffinet 2000]) Pohlia baldwinii (Broth. ex E.B.Bartram) W.SchultzeMotel [Hoe 1974] Webera baldwinii Broth. ex E.B.Bartram Pohlia cruda (Hedw.) Lindb. [Hoe 1974, Hoe 1979] Webera cruda (Hedw.) Fürnr. Pohlia flexuosa Hook. [Hoe 1974, Gangulee 1974] Pohlia gracilescens (E.B.Bartram) E.B.Bartram Pohlia gracillima (Cardot) Horik. & Ochi Pohlia leucostoma (Bosch & Sande Lac.) M.Fleisch. Pohlia leucostomoides (Broth.) E.B.Bartram Webera gracilescens E.B.Bartram Webera leucostomoides Broth. Pohlia mauiensis (Broth. ex E.B.Bartram) W.SchultzeMotel [Hoe 1974] TROPICAL BRYOLOGY 25 (2004) ind end K O Mo M H Staples et al. 58 FullName POLYTRICHUM Hedw. (Polytrichaceae) Polytrichum commune Hedw. [Hoe 1974] Polytrichum juniperinum Hedw. [Hoe 1974, Hyvönen 1986] Polytrichum piliferum Hedw. [Hoe 1974] PSEUDOSCLEROPODIUM (Limpr.) M.Fleisch. (Brachytheciaceae) Pseudoscleropodium purum (Hedw.) M.Fleisch. [Hoe 1974] PSEUDOSYMBLEPHARIS Broth. (Pottiaceae) Pseudosymblepharis angustata (Mitt.) Hilp. [Hoe 1974, Hoe 1979, Norris & Koponen 1989] Pseudosymblepharis mauiensis (Mull.Hal.) Broth. PSEUDOTAXIPHYLLUM Z.Iwats. (Hypnaceae) Pseudotaxiphyllum elegans (Brid.) Z.Iwats. [Hoe 1979, Iwatsuki 1987] Isopterygium elegans (Brid.) Lindb. PTEROGONIDIUM Broth. (Sematophyllaceae) Pterogonidium pulchellum (Hook.) Müll.Hal. [Hoe 1979] PTYCHOMITRIUM Fürnr. (Ptychomitriaceae) Ptychomitrium lepidomitrium (Müll.Hal.) Schimp. ex Besch. [Hoe 1974, Cao et al. 2003] Ptychomitrium mauiense Broth. PYLAISIADELPHA Broth. (Meteoriaceae) Pylaisiadelpha opaeodon (Sull.) W.R. Buck [Hoe 1974, Buck 1984] Brotherella opaeodon (Sull.) Broth. Hypnum opaeodon Sull. PYRRHOBRYUM Mitt. (Rhizogoniaceae) Pyrrhobryum spiniforme (Hedw.) Mitt. [Hoe 1974, Koponen et al. 1986] Rhizogonium spiniforme (Hedw.) Bruch RACOMITRIUM Brid. (Grimmiaceae) Racomitrium crispulum (Hook.f. & Wilson) Hook.f. & Wilson [Hoe 1974] Racomitrium fasciculare (Schrad. ex Hedw.) Brid. var. erosum Broth. [Hoe 1974] Racomitrium lanuginosum (Hedw.) Brid. var. lanuginosum [Hoe 1974] STATUS NI K O MO L M KA ind ind H H ind M nat ind H H H K O Mo ind M H M nat? O ind M H ind K O Mo M H ind K O Mo L M H ind K H M end M ind M TROPICAL BRYOLOGY 25 (2004) 59 Revised checklist of Hawaiian mosses FullName Racomitrium lanuginosum (Hedw.) Brid. var. pruinosum Wilson [Hoe 1974] Racomitrium lanuginosum (Hedw.) Brid. var. sandvicense Reichardt RACOPILUM P.Beauv. (Racopilaceae) Racopilum cuspidigerum (Schwägr.) Ångstr. [Hoe 1974] Rhacomitrium cuspidigerum misapplied RADULINA W.R.Buck & B.C.Tan (Sematophyllaceae) Radulina hamata (Dozy & Molk.) W.R.Buck & B.C.Tan [Hoe 1974, Buck & Tan 1989] Trichosteleum hamatum (Dozy & Molk.) A.Jaeger REMYELLA Müll.Hal. (Brachytheciaceae) Remyella hawaica Müll.Hal. [Hoe 1974, Ignatov & Huttunen 2002] Rhynchostegiella hawaica (Müll.Hal.) Broth. RHABDOWEISIA Bruch & Schimp. (Dicranaceae, or Rhabdoweisiaceae [Shaw & Goffinet 2000]) Rhabdoweisia crenulata (Mitt.) H.Jameson [Hoe 1975, Frahm et al. 2000] Rhabdoweisia crispata (With.) Lindb. [Hoe 1974, Hoe 1977, Hoe 1979] Rhabdoweisia denticulata B.S.G. STATUS NI K O MO L M KA ind K O Mo M nat? K O Mo L M H ind K O Mo L M H end O ind M ind M H H RHAPHIDOSTICHUM M.Fleisch. (Sematophyllaceae) Rhaphidostichum pustulatum Hoe [Hoe 1974] end RHIZOGONIUM Brid. (Rhizogoniaceae) Rhizogonium pungens Sull. [Hoe 1974] end K O Mo L M ind K O M ind K O Mo M RHODOBRYUM (Schimp.) Limpr. (Bryaceae) Rhodobryum giganteum (Schwägr.) Paris [Hoe 1974, Crosby et al. 1983, Tan & Iwatsuki 1991] Bryum giganteum (Schwägr.) Arn. RHYNCHOSTEGIUM Bruch & Schimp. (Brachytheciaceae) Rhynchostegium celebicum (Sande Lac.) A.Jaeger [Hoe 1974, Hoe 1977] Eurhynchium celebicum (Sande Lac.) E.B.Bartram Rhynchostegium gaudichaudii (Mont.) A.Jaeger [Hoe 1974] Rhynchostegium selaginellifolium Müll.Hal. var. recurvirameum (Müll.Hal.) Wijk & Margad. [Hoe 1974] Eurhynchium selaginellifolium var. recurvirameum TROPICAL BRYOLOGY 25 (2004) H H O ind end K O H Staples et al. 60 FullName (Müll.Hal.) E.B.Bartram Rhynchostegium selaginellifolium Müll.Hal. var. selaginellifolium [Hoe 1974] Eurhynchium selaginellifolium (Müll.Hal.) E.B.Bartram ROSULABRYUM J.R.Spence (Bryaceae) Rosulabryum billardierei (Schwägr.) J.R.Spence [Mohamed 1979, Anderson et al. 1990, Spence 1996] Bryum andicola Hook. Bryum decaisnei Dozy & Molk. Bryum limbatomarginatum Müll.Hal. Bryum truncorum misapplied Rosulabryum capillare (Hedw.) J.R.Spence [Hoe 1974, Hoe 1977, Spence 1996] Bryum capillare Hedw. Bryum vinoviride E.B.Bartram SAELANIA Lindb. (Ditrichaceae) Saelania glaucescens (Hedw.) Broth. [Hoe 1974] SCHISTIDIUM Bruch & Schimp. (Grimmiaceae) Schistidium apocarpum (Hedw.) Bruch & Schimp. [Schultze-Motel 1963, Muñoz & Pando 2000] Grimmia apocarpa Hedw. Schistidium strictum (Turner) Loeske ex Mårtensson [Hoe 1974, Cao & Churchill 1995, Muñoz & Pando 2000] Grimmia apocarpa Hedw. var. stricta (Turner) Hook. & Taylor Grimmia scabrifolia Broth. Grimmia stricta Turner SCHIZYMENIUM Harv. (Bryaceae, or Mniaceae [Shaw & Goffinet 2000]) Schizymenium campylocarpum (Hook. & Arn.) A.J.Shaw var. campylocarpum [Zanten 1971, Shaw 1985] Mielichhoferia campylocarpa (Hook. & Arn.) Mitt. Weissia campylocarpa Hook. & Arn. in Hook. Schizymenium nealiae (E.B.Bartram) A.J.Shaw [Hoe 1974, Shaw 1985] Mielichhoferia nealiae E.B.Bartram Schizymenium pulvinatum (Müll.Hal.) A.J.Shaw [Hoe 1974, Shaw 1985] Mielichhoferia pulvinata Müll.Hal. SCOPELOPHILA (Mitt.) Lindb. (Pottiaceae) Scopelophila infericola Hoe [Hoe 1974] STATUS NI K O MO L M KA end K O Mo H M H ind O M H ind K O M H ind M ind M ind M ind H end H end end M H H TROPICAL BRYOLOGY 25 (2004) 61 Revised checklist of Hawaiian mosses FullName SEMATOPHYLLUM Mitt. (Sematophyllaceae) Sematophyllum hawaiiense (Broth.) Broth. [Hoe 1974, Hoe 1979] Sematophyllum subpinnatum (Brid.) E.Britton [Hoe 1974, Hoe 1979, Buck 1983] Sematophyllum caespitosum misapplied SPHAGNUM L. (Sphagnaceae) Sphagnum palustre L. [Hoe 1978, Karlin & Andrus 1995] Sphagnum henryense misapplied Sphagnum lonchocladum Müll.Hal. Sphagnum perichaetiale misapplied Sphagnum vitjianum misapplied Sphagnum wheeleri Müll.Hal. [Hoe 1974, Karlin & Andrus 1995, Karlin 2001] Sphagnum compactum misapplied Sphagnum rigidulum Warnst. Sphagnum vulcanicum Warnst. SPLACHNOBRYUM Müll.Hal. (Splachnobryaceae, or Pottiaceae [Shaw & Goffinet 2000]) Splachnobryum obtusum (Brid.) Müll.Hal. [Hoe 1974, Arts 2001] STREPTOPOGON Mitt. (Pottiaceae) Streptopogon erythrodontus (Taylor) Wilson [Hoe 1974] SYMPHYODON Mont. (Symphyodontaceae) Symphyodon pygmaeus (Broth.) S.He & Snider [Hoe 1974, Hoe 1977, He & Snider 1992] Glossadelphus abortivapicus Hoe SYNTRICHIA Brid. (Pottiaceae) Syntrichia fragilis (Taylor) Ochyra [Hoe 1974, Kürschner 2000] Tortula alpina (Brid.) Arn. var. inermis (Milde) De Not. Tortula fragilis Taylor Syntrichia princeps (De Not.) Mitt. [Hoe 1974, Kürschner 2000] Tortula princeps De Not. SYRRHOPODON Schwägr. (Calymperaceae) Syrrhopodon armatus Mitt. [Hoe 1974, Orbán & Reese 1986] Syrrhopodon oahuensis Broth. Syrrhopodon hawaiicus Müll.Hal. [Hoe 1974] Syrrhopodon prolifer Schwägr. [Hoe 1974, Reese et al. TROPICAL BRYOLOGY 25 (2004) STATUS NI K O MO L M KA ind K O ind O L M H end M O ind ind H H ind nat H M O M ind M H ind M H M H M M H H ind end end O K O Mo O Mo Staples et al. 62 FullName 1986] Syrrhopodon kilaueae Müll.Hal. Syrrhopodon semiliber (Mitt.) Besch. [Hoe 1974, Reese et al. 1986] Calymperes semiliberum Mitt. Calymperopsis semiliber (Mitt.) M.Fleisch. TAXIPHYLLUM M.Fleisch. (Hypnaceae) Taxiphyllum laevifolium (Mitt.) W.R.Buck [Hoe 1974, Buck 1987b, Tixier 1988] Glossadelphus baldwinii Broth. Glossadelphus laevifolius (Mitt.) E.B.Bartram Taxiphyllum torrentium (Besch.) W.R.Buck [Hoe 1974, Buck 1987b, Tixier 1988] Glossadelphus irroratus E.B.Bartram Glossadelphus torrentium (Besch.) M.Fleisch. TAXITHELIUM Mitt. (Sematophyllaceae) Taxithelium mundulum (Sull.) E.B.Bartram [Hoe 1974] Taxithelium undulatum misapplied TAYLORIA Hook. (Splachnaceae) Tayloria sandwicensis (Müll.Hal.) Broth. [Hoe 1974] THAMNIOPSIS (Mitt.) Fleisch. (Hookeriaceae, or Callicostaceae [Buck 1987a]) Thamniopsis utacamundiana (Mont.) W.R Buck [Hoe 1974, Buck 1987a] Hookeria purpurea Müll.Hal. Hookeriopsis purpurea (Müll.Hal.) Broth. Hookeriopsis purpurea (Müll.Hal.) Broth. var. acuminatula (Müll.Hal.) E.B.Bartram Hookeriopsis purpurea (Müll.Hal.) Broth. var. ligulacea (Müll.Hal.) E.B.Bartram THAMNOBRYUM Nieuwl. (Neckeraceae) Thamnobryum speciosum (Broth.) Hoe [Hoe 1974] Thamnium speciosum Broth. THUIDIUM Bruch & Schimp. (Thuidiaceae) Thuidium cymbifolium (Dozy & Molk.) Dozy & Molk. [Hoe 1974, Touw 2001] Thuidium glabratum Broth. ?Thuidium havaiense Reichardt Thuidium oahuense Broth. Thuidium plicatum Mitt. Thuidium plicatum Mitt. var. brevifolium E.B.Bartram STATUS NI K O MO L M KA ind ind O K O Mo H M ind M ind K O Mo L M end M ind K O Mo H H H M H end M H ind K O Mo L M H TROPICAL BRYOLOGY 25 (2004) 63 Revised checklist of Hawaiian mosses FullName TIMMIA Hedw. (Timmiaceae) Timmia megapolitana Hedw. ssp. bavarica (Hessl.) Brassard [Brassard 1984] Timmia bavarica Hessl. TORTELLA (Lindb.) Limpr. (Pottiaceae) Tortella fragilis (Hook. & Wilson) Limpr. var. tortelloides (S.W.Greene) R.H.Zander & Hoe [Zander & Hoe 1979] Sarconeurum tortelloides S.W.Greene Tortella tortelloides (S.W.Greene) H.Rob. Tortella humilis (Hedw.) Jenn. [Hoe 1974] Tortella caespitosa (Schwägr.) Limpr. Tortella tortuosa (Hedw.) Limpr. [Hoe 1979] TORTULA Hedw. (Pottiaceae) Tortula atrovirens (Sm.) Lindb. [Hoe 1974, Hoe 1979, Corley et al. 1981, Zander 1993] Desmatodon convolutus (Brid.) Grout TRACHYLOMA Brid. (Pterobryaceae, or Trachylomataceae [Shaw & Goffinet 2000]) Trachyloma indicum Mitt. [Hoe 1974] Trachyloma tahitense Besch. TRACHYPODOPSIS M.Fleisch. (Trachypodaceae, or Meteoriaceae [Shaw & Goffinet 2000]) Trachypodopsis auriculata (Mitt.) M.Fleisch. [Hoe 1974] Trachypodopsis ornans (Reichardt) M.Fleisch. Trachypus ornans (Reichardt) Müll.Hal. fma. simplex Broth. Trachypodopsis serrulata (P.Beauv.) M.Fleisch. var. crispatula (Hook.) Zanten [Hoe 1979] TRACHYPUS Reinw. & Hornsch. (Trachypodaceae, or Meteoriaceae [Shaw & Goffinet 2000]) Trachypus bicolor Reinw. & Hornsch. [Hoe 1974, Hoe 1979] Trachypus humilis Lindb. var. flagellifer (Broth. ex Herzog) H.A.Mill. [Zanten 1959, Miller 1967, Hoe 1974] Trachypus humilis Lindb. var. tenerrimus (Herzog) Zanten Trachypus mauiensis Broth. Trachypus humilis Lindb. var. humilis [Hoe 1974, Hoe 1977] TREMATODON Michx. (Dicranaceae, or Bruchiaceae [Shaw & Goffinet 2000]) TROPICAL BRYOLOGY 25 (2004) STATUS NI K O MO L M KA ind H ind ind M K O ind L M H L ind M ind M ind H K O Mo M ind H H H ind K M ind K O M ind K M H Staples et al. 64 FullName Trematodon latinervis Müll.Hal. [Hoe 1974] Trematodon longicollis misapplied TRICHOSTOMUM Bruch (Pottiaceae) Trichostomum brachydontium Bruch [Sollman 1984] Trichostomum oblongifolium E.B.Bartram Trichostomum crispulum Bruch [Hoe 1974, Hoe 1975, Norris & Koponen 1989] Trichostomum bartramii H.A.Mill. Trichostomum mauiense Broth. ULOTA D.Mohr (Orthotrichaceae) Ulota cervina Hoe & H.A.Crum [Hoe 1974] VESICULARIA (Müll.Hal.) Müll.Hal. (Hypnaceae) Vesicularia inflectens (Brid.) Müll.Hal. [Hoe 1979] Vesicularia perviridis (Ångstr.) Müll.Hal. [Hoe 1974] Vesicularia graminicolor Müll.Hal. WEISSIA Hedw. (Pottiaceae) Weissia controversa Hedw. [Hoe 1974, Wichman & St. John 1990] Weisia viridula Hedw. Weissia ovalis (R.S.Williams) E.B.Bartram [Hoe 1974] ZYGODON Hook. & Taylor (Orthotrichaceae) Zygodon reinwardtii (Hornsch.) A.Braun [Hoe 1974] Zygodon rubrigemmius (Hoe & H.A.Crum) R.H.Zander & Vitt [Hoe 1974, Zander & Vitt 1979] Anoectangium rubrigemmium Hoe & H.A.Crum Zygodon tetragonostomus A.Braun ex B.S.G. [Hoe 1974] STATUS NI K O MO L M KA ind K O Mo M H H ind K O Mo M H ind K O Mo L M H end M ind end M K O Mo L M ind Ni K O H end K O Mo L M ind end M M ind K O H M H H TROPICAL BRYOLOGY 25 (2004) Revised checklist of Hawaiian mosses CROSS-REFERENCED LIST OF SYNONYMS AND MISAPPLIED NAMES Acroporium fuscoflavum var. baldwinii See Acroporium fuscoflavum ?Acroporium sigmatodontium misapplied See Acroporium fuscoflavum Aerobryopsis longissima See Aerobryopsis wallichii Aerobryopsis longissima var. dozyana See Aerobryopsis wallichii Aerobryopsis scariosa See Aerobryopsis subdivergens ssp. scariosa Amphidium cyathicarpum See Amphidium tortuosum Andreaea acuminata See Andreaea acutifolia ssp. acuminata Andreaea perpapillosa See Andreaea rupestris Anoectangium euchloron See Anoectangium aestivum Anoectangium haleakalae var. haleakalae See Anoectangium aestivum Anoectangium haleakalae var. laxulum See Anoectangium aestivum Anoectangium harttiae See Anoectangium aestivum Anoectangium rubrigemmium See Zygodon rubrigemmius Aptychella robusta See Clastobryopsis robusta Barbella trichophora See Barbellopsis trichophora Barbula cruegeri See Barbula indica Barbula unguiculata misapplied See Barbula indica Barbula vinealis var. cylindrica See Didymodon insulanus Barbula vinealis var. flaccida See Didymodon insulanus Brachythecium oxyrrhynchum See Brachythecium plumosum Breutelia kilaueae See Breutelia affinis Brotherella opaeodon See Pylaisiadelpha opaeodon Bryohaplocladium microphyllum See Haplocladium microphyllum Bryum andicola Hook. See Rosulabryum billardierei Bryum apiculatum See Bryum mildeanum Bryum argenteum var. lanatum See Bryum argenteum Bryum capillare See Rosulabryum capillare Bryum ceramiocarpum See Bryum conicum Bryum crassicostatum See Bryum hawaiicum Bryum decaisnei See Rosulabryum billardierei Bryum erythrocarpum See Bryum atrovirens Bryum giganteum See Rhodobryum giganteum Bryum leucophyllum See Bryum argenteum Bryum limbatomarginatum See Rosulabryum billardierei Bryum mauiensis See Bryum mildeanum Bryum megalostegium See Bryum mildeanum Bryum nitens See Bryum mildeanum Bryum truncorum misapplied See Rosulabryum billardierei Bryum vinoviride See Rosulabryum capillare Calymperes hawaiiense See Calymperes fasciculatum Calymperes semiliberum See Syrrhopodon semiliber Calymperopsis semiliber See Syrrhopodon semiliber Campylopodium euphorocladum See Campylopodium medium Campylopus aureus See Campylopus schmidii ssp. schmidii Campylopus boswellii See Campylopus fragilis ssp. zollingerianus Campylopus boswellii var. capitulatus See Campylopus fragilis ssp. zollingerianus Campylopus densifolius var. densifolius See Campylopus hawaiicus var. densifolius Campylopus densifolius var. falcatus See Campylopus hawaiicus var. hawaiicoflexuosus TROPICAL BRYOLOGY 25 (2004) 65 66 Staples et al. Campylopus densifolius var. hawaiicoflexuosus See Campylopus hawaiicus var. hawaiicoflexuosus Campylopus densifolius var. purpureoflavescens See Campylopus hawaiicus var. hawaiicus Campylopus densiretis See Campylopus hawaiicus var. densifolius ?Campylopus didrichsenii See Campylopus umbellatus var. umbellatus Campylopus exasperatus See Campylopus praemorsus Campylopus fumarioli See Campylopus laxitextus Campylopus geniculatus See Campylopus umbellatus var. pterotoneuron Campylopus hawaiicoflexuosus See Campylopus hawaiicus var. hawaiicoflexuosus Campylopus introflexus misapplied See Campylopus schmidii ssp. schmidii Campylopus introflexus var. polytrichoides misapplied See Campylopus schmidii ssp. schmidii Campylopus microcephalus See Campylopus praemorsus Campylopus perangustifolius See Campylopus hawaiicus var. hawaiicoflexuosus ?Campylopus pertristis See Campylopus umbellatus var. umbellatus Campylopus polytrichoides misapplied See Campylopus schmidii ssp. schmidii Campylopus purpureoflavescens, in part See Campylopus hawaiicus var. hawaiicoflexuosus, C. hawaiicus var. hawaiicus Campylopus richardii misapplied See Campylopus umbellatus var. umbellatus Campylopus sandwicensis See Campylopus umbellatus var. umbellatus Campylopus skottsbergii See Campylopus schmidii ssp. hemitrichius Campylopus tubulosus See Campylopus schmidii ssp. hemitrichius Campylopus wawreanus See Campylopus hawaiicus var. densifolius Ceratodon purpureus fma. xanthopus See Ceratodon purpureus ssp. purpureus Cirriphyllum oxyrrhynchum See Brachythecium plumosum Claopodium amblystegioides See Claopodium prionophyllum Claopodium hawaiiense See Claopodium whippleanum Claopodium nervosum See Claopodium prionophyllum Ctenidium decurrens See Ectropothecium decurrens Ctenidium pulcherrimum See Ectropothecium decurrens Ctenidium submalacobolum See Ectropothecium decurrens Cupressina decurrens See Ectropothecium decurrens Desmatodon convolutus See Tortula atrovirens Dicranella hillebrandii See Microdus hillebrandii Dicranella hochreuteri misapplied See Dicranella hochreutineri Dicranodontium falcatum var. falcatum See Dicranodontium porodictyon Dicranodontium hawaiicum See Dicranodontium porodictyon Dicranoloma wheeleri See Campylopus wheeleri Dicranum didrichsenii See Campylopus umbellatus var. umbellatus Dicranum hawaiicoflexuosum See Campylopus hawaiicus var. hawaiicoflexuosus Dicranum hawaiicum See Campylopus hawaiicus var. hawaiicus Dicranum microcephalum See Campylopus praemorsus Dicranum pertriste See Campylopus umbellatus var. umbellatus Dicranum praemorsum See Campylopus praemorsus Dicranum skottsbergii See Campylopus schmidii ssp. hemitrichius Dicranum wheeleri See Campylopus wheeleri Didymodon recurvirostris See Bryoerythrophyllum recurvirostre Didymodon vinealis var. flaccidus See Didymodon insulanus Ectropothecium rockii See Ectropothecium decurrens Encalypta rhaptocarpa misapplied See Encalypta rhabdocarpa Encalypta scabrata See Encalypta ciliata Entodon hillebrandii See Entodon plicatus TROPICAL BRYOLOGY 25 (2004) Revised checklist of Hawaiian mosses Entodon solanderi See Entodon plicatus Eurhynchium celebicum See Rhynchostegium celebicum Eurhynchium muelleri misapplied See Platyhypnidium muelleri Eurhynchium selaginellifolium See Rhynchostegium selaginellifolium var. selaginellifolium Eurhynchium selaginellifolium var. recurvirameum See Rhynchostegium selaginellifolium var. recurvirameum Eurhynchium vagans See Oxyrrhynchium vagans Fissidens baldwinii See Fissidens elegans Fissidens bishopii See Fissidens elegans Fissidens crumii See Fissidens lindbergii Fissidens hawaiicus See Fissidens bryoides Fissidens insularis See Fissidens bryoides Fissidens intermedius See Fissidens submarginatus Fissidens mauiensis See Fissidens pacificus Fissidens oahuensis See Fissidens bryoides Floribundaria baldwinii See Floribundaria floribunda Funaria subintegra See Entosthodon subintegrus Garovaglia haleakalae See Garovaglia powellii var. haleakalae Glossadelphus abortivapicus See Symphyodon pygmaeus Glossadelphus baldwinii See Taxiphyllum laevifolium Glossadelphus chrysobasilaris See Glossadelphus ogatae Glossadelphus irroratus See Taxiphyllum torrentium Glossadelphus laevifolius See Taxiphyllum laevifolium Glossadelphus torrentium See Taxiphyllum torrentium Glossadelphus zollingeri var. filicaulis See Ectropothecium zollingeri var. filicaule Grimmia apocarpa See Schistidium apocarpum Grimmia apocarpa var. pulvinata See Grimmia anodon Grimmia apocarpa var. stricta See Schistidium strictum Grimmia haleakalae See Amphidium tortuosum Grimmia haliacalae See Amphidium tortuosum Grimmia pilfera See Grimmia longirostris Grimmia scabrifolia See Schistidium strictum Grimmia stricta See Schistidium strictum Gymnostoma pulvinata See Grimmia anodon Hageniella pacifica See Hageniella micans Holomitrium ferriei misapplied See Holomitrium squarrifolium Homalia densa See Homaliodendron flabellatum Homalia praelonga See Homaliodendron flabellatum Hookeria purpurea See Thamniopsis utacamundiana Hookeriopsis purpurea See Thamniopsis utacamundiana Hookeriopsis purpurea var. acuminatula See Thamniopsis utacamundiana Hookeriopsis purpurea var. ligulacea See Thamniopsis utacamundiana Hymenostylium firmum See Hymenostylium recurvirostrum var. recurvirostrum Hyophila dozy-molkenboeri See Hyophila involuta Hypnum decurrens See Ectropothecium decurrens Hypnum draytoni(i) See Buckiella draytonii Hypnum eudorae See Buckiella draytonii Hypnum opaeodon See Pylaisiadelpha opaeodon Hypnum wilkesianum See Palamocladium wilkesianum Isopterygium elegans See Pseudotaxiphyllum elegans TROPICAL BRYOLOGY 25 (2004) 67 Staples et al. 68 Leptodontium brevicaule See Leptodontium flexifolium Leucobryum hawaiense See Leucobryum seemannii var. seemannii Leucobryum hawaiense var. fumarioli See Leucobryum seemannii var. fumarioli Leucobryum hawaiense var. solfatarae See Leucobryum seemannii var. solfatarae Leucobryum solfatarae See Leucobryum seemannii var. solfatarae Leucobryum solfatarae var. fumarioli See Leucobryum seemannii var. fumarioli Leucobryum solfatarae var. hawaiense See Leucobryum seemannii var. seemannii Macrocoma intricata See Macrocoma tenue ssp. sullivantii Macromitrium cumingii See Macromitirum incurvifolium Macromitrium intricatum See Macrocoma tenue ssp. sullivantii Macromitrium owahiense See Macromitrium microstomum Macromitrium reinwardtii See Macromitrium microstomum Merceyopsis crassinervis See Hymenostylium recurvirostrum var. cylindricum Meteorium pallidovirens See Palamocladium wilkesianum Meteorium sciuroides See Palamocladium wilkesianum Microdus hawaiicus See Dicranella hawaiica var. hawaiica Mielichhoferia campylocarpa See Schizymenium campylocarpum var. campylocarpum Mielichhoferia nealiae See Schizymenium nealiae Mielichhoferia pulvinata See Schizymenium pulvinatum Mniadelphus freycineti var. crasseturgescens See Distichophyllum freycinetii var. crasseturgescens Mnium rostratum See Plagiomnium rostratum Mnium serratum See Mnium marginatum Molendoa crassinervis See Hymenostylium recurvirostrum var. cylindricum Orthotrichum berggrenii See Orthotrichum alpestre var. berggrenii Orthotrichum hillebrandii See Orthotrichum rupestre Orthotrichum verrucatum See Orthotrichum lyellii Palamocladium aptychoides See Palamocladium wilkesianum Palamocladium sciurellum See Palamocladium wilkesianum Palamocladium wilkesianum var. altisetum See Palamocladium wilkesianum Palamocladium wilkesianum var. sciuroides See Palamocladium wilkesianum Philonotis laxissima See Philonotis hastata Plagiomnium prorepens See Plagiomnium rostratum Plagiomnium rhynchosporum See Plagiomnium rostratum Plagiopus longisetus See Plagiopus oederiana Plagiopus oederi See Plagiopus oederiana Plagiothecium draytonii See Buckiella draytonii Pleuropus aptychoides See Palamocladium wilkesianum Pleuropus pallidovirens See Palamocladium wilkesianum Pleuropus sciurellus See Palamocladium wilkesianum Pleuropus wilkesianus See Palamocladium wilkesianum Pleuropus wilkesianus var. sciuroides See Palamocladium wilkesianum Pogonatum baldwinii See Pogonatum tahitense Pohlia gracilescens See Pohlia flexuosa Pohlia gracillima See Pohlia flexuosa Pohlia leucostoma See Pohlia flexuosa Pohlia leucostomoides See Pohlia flexuosa Pseudosymblepharis mauiensis See Pseudosymblepharis angustata Ptychomitrium mauiense See Ptychomitrium lepidomitrium Ptychomnion aciculare misapplied See Aerobryopsis subdivergens ssp. scariosa TROPICAL BRYOLOGY 25 (2004) Revised checklist of Hawaiian mosses Racomitrium lanuginosum var. sandvicense See Racomitrium lanuginosum var. pruinosum Rhabdoweisia denticulata See Rhabdoweisia crispata Rhacomitrium cuspidigerum misapplied See Racopilum cuspidigerum Rhizogonium spiniforme See Pyrrhobryum spiniforme Rhynchostegiella hawaica See Remyella hawaica Rhynchostegium vagans See Oxyrrhynchium vagans Sarconeurum tortelloides See Tortella fragilis var. tortelloides Sciaromium tricostatum See Limbella tricostata Sematophyllum caespitosum See Sematophyllum subpinnatum Sphagnum compactum misapplied See Sphagnum wheeleri Sphagnum henryense misapplied See Sphagnum palustre Sphagnum lonchocladum See Sphagnum palustre Sphagnum perichaetiale misapplied See Sphagnum palustre Sphagnum rigidulum See Sphagnum wheeleri Sphagnum vitjianum misapplied See Sphagnum palustre Sphagnum vulcanicum See Sphagnum wheeleri Stereodon decurrens See Ectropothecium decurrens Stereophyllum oahuense See Entodontopsis anceps Syrrhopodon kilaueae See Syrrhopodon prolifer Syrrhopodon oahuensis See Syrrhopodon armatus Taxiphyllum gracile See Ectropothecium decurrens Taxithelium undulatum misapplied See Taxithelium mundulum Thamnium speciosum See Thamnobryum speciosum Thuidium crenulatum See Pelekium versicolor Thuidium glabratum See Thuidium cymbifolium ?Thuidium havaiense See Thuidium cymbifolium Thuidium nanophyllum See Pelekium versicolor Thuidium oahuense See Thuidium cymbifolium Thuidium plicatum See Thuidium cymbifolium Thuidium plicatum var. brevifolium See Thuidium cymbifolium Thysanomitrion hawaiicum See Campylopus umbellatus var. umbellatus Thysanomitrion umbellatum See Campylopus umbellatus var. umbellatus Timmia bavarica See Timmia megapolitana ssp. bavarica Tortella caespitosa See Tortella humilis Tortella tortelloides See Tortella fragilis var. tortelloides Tortula alpina var. inermis See Syntrichia fragilis Tortula fragilis See Syntrichia fragilis Tortula princeps See Syntrichia princeps Tortula rhizophylla See Chenia leptophylla Tortula vectensis See Chenia leptophylla Trachyloma tahitense See Trachyloma indicum Trachypodopsis ornans See Trachypodopsis auriculata Trachypus humilis var. tenerrimus See Trachypus humilis var. flagellifer Trachypus mauiensis See Trachypus humilis var. flagellifer Trachypus ornans fma. simplex See Trachypodopsis auriculata Trematodon longicollis misapplied See Trematodon latinervis Trichosteleum hamatum See Radulina hamata Trichostomum bartramii See Trichostomum crispulum Trichostomum mauiense See Trichostomum crispulum Trichostomum oblongifolium See Trichostomum brachydontium TROPICAL BRYOLOGY 25 (2004) 69 Staples et al. 70 Trichostomum tenuirostre See Oxystegus tenuirostris var. gemmiparus Vesicularia graminicolor See Vesicularia perviridis Webera baldwinii See Pohlia baldwinii Webera cruda See Pohlia cruda Webera gracilescens See Pohlia flexuosa Webera leucostomoides See Pohlia flexuosa Weisia viridula See Weissia controversa Weissia campylocarpa See Schizymenium campylocarpum var. campylocarpum TROPICAL BRYOLOGY 25 (2004)